321 lines
57 KiB
XML
321 lines
57 KiB
XML
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<document id="1AD6C46CDC5602EAE0FB37CB34BE3D81" ID-CLB-Dataset="3396" ID-DOI="10.11646/zootaxa.5068.4.2" ID-GBIF-Dataset="45ebb61b-259d-4ac3-bd7c-3a7da0f644f5" ID-ISSN="1175-5326" ID-Zenodo-Dep="5709510" ID-ZooBank="F769B986-8F7B-4ABF-A7EF-58A813718760" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1637219239794" checkinUser="plazi" docAuthor="Golovatch, Sergei I., Aswathy, Mathilakath Dasan, Bhagirathan, Usha & Sudhikumar, Ambalaparambil Vasu" docDate="2021" docId="3A1C87D9970DFFCCFF1F2A8CBDD8A0CE" docLanguage="en" docName="zootaxa.5068.4.2.pdf" docOrigin="Zootaxa 5068 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Delarthrum anomalans Golovatch & Aswathy & Bhagirathan & Sudhikumar 2021, sp. nov." docType="treatment" docVersion="6" lastPageNumber="511" masterDocId="C625FFA19715FFD6FF882F49B964A65C" masterDocTitle="Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae)" masterLastPageNumber="516" masterPageNumber="485" pageNumber="509" updateTime="1699468853807" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="245F2F048449BA3BD76CB28F049D754B">
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<mods:title id="FA67DA23EEE8D51A3BEDA52A4989A85C">Review of the millipede tribe Polydrepanini, with the description of a new species from Kerala state, southern India (Diplopoda, Polydesmida, Paradoxosomatidae, Alogolykinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="0E4E7AF2067EB42BF1A6B2178BA1B060" type="personal">
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<mods:roleTerm id="2FE0A33FBC6EE3451404BA2F2D871999">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="412E2CAF3D0E35B41B5D72FAEDBA4C01">Golovatch, Sergei I.</mods:namePart>
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<mods:affiliation id="ADBEA1B9BDECD00EE119DFE04EC4C809">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 11907, Russia.</mods:affiliation>
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</mods:name>
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<mods:name id="A112F5F73E7B65270094A6A3535248F5" type="personal">
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<mods:role id="9DE2F720ABF7485BFA05F3BC33A40E37">
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<mods:roleTerm id="BBEFE147EBBA2C601FDA1605D62B79BB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="35A4DCA60C49CA19473816BE387A480B">Aswathy, Mathilakath Dasan</mods:namePart>
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<mods:nameIdentifier id="68A4B3F6A74EE51210F034BE21B148B8" type="ORCID">0000-0002-3904-8066</mods:nameIdentifier>
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<mods:affiliation id="D2EC1BCCEEB06E842A8E86BBABCF3C6D">Centre for Animal Taxonomy and Ecology (CATE), Christ College, Irinjalakuda, Kerala, India. & aswathym. das 94 @ gmail. com; https: // orcid. org / 0000 - 0002 - 3904 - 8066</mods:affiliation>
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<mods:nameIdentifier id="8430D4821773FB1B796BCDEAADB1E597" type="email">aswathym.das94@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="F183C988D5275AD9BBD7AF6FD3946799" type="personal">
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<mods:role id="1780124460F8E5CDCECC2057406BC7D7">
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<mods:roleTerm id="5D8C61C617B6FB7C70BECD16E65BC32B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BAD303FB1262462D0C79AAC7612D3188">Bhagirathan, Usha</mods:namePart>
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</mods:name>
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<mods:name id="342300383D138E89D11AAD9B0192E0D5" type="personal">
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<mods:roleTerm id="8318B381010BC2E618995D5C65C26CD6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2B1DC235541DDC9902F1E4AEFA80926C">Sudhikumar, Ambalaparambil Vasu</mods:namePart>
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<mods:nameIdentifier id="98D727AE01AA70BEE6D9CC8481D56D19" type="ORCID">0000-0002-4479-4995</mods:nameIdentifier>
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<mods:affiliation id="66E9378EADDFDEA3E053E502698F8258">Centre for Animal Taxonomy and Ecology (CATE), Christ College, Irinjalakuda, Kerala, India. & spidersudhi @ gmail. com; https: // orcid. org / 0000 - 0002 - 4479 - 4995</mods:affiliation>
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<mods:nameIdentifier id="556F25034C29B0745EE3B9F5965AD211" type="email">spidersudhi@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="96B3DF6ABA81FD7E96627828DC76C21A">text</mods:typeOfResource>
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<mods:title id="D1BD587493900F0E7A08255E4705A985">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="2AA1476F9B7E9A88932487A5DFBEBE6C">
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<mods:date id="26FE42133BF0BCC07AF6FF11FE4884F9">2021</mods:date>
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<mods:number id="883A37C4CD5CACFFBBFE1B6562E4C4AE">2021-11-18</mods:number>
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<mods:detail id="9D56948827EA1757147601FFD0DDD546" type="volume">
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<mods:number id="3B12B28270BDD96E09F517DD9D68D3FE">5068</mods:number>
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</mods:detail>
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<mods:detail id="5DA4DA1C3C300E12F5E5145ED8218347" type="issue">
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<mods:number id="4CEF833B5ED65D204D22EF606C0A39A2">4</mods:number>
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<mods:start id="C805CBEEC78FACB75076382CEA0EB071">485</mods:start>
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<mods:end id="BDDF7925B606ABED686D024F98B9A807">516</mods:end>
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<mods:classification id="7179B519610354FC00200F68372A39DD">journal article</mods:classification>
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<mods:identifier id="FAEAD352D4DE600B9422BA7F0C717DA1" type="CLB-Dataset">3396</mods:identifier>
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<mods:identifier id="A1B19C7B1754C4E2C08E3F3F07F3FF1D" type="DOI">10.11646/zootaxa.5068.4.2</mods:identifier>
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<mods:identifier id="07C4BA2D963DE803BECF6C96684E160B" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="4DBBF16884F8BDB10EB7749D6E8EDD81" type="Zenodo-Dep">5709510</mods:identifier>
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</mods:mods>
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<treatment id="3A1C87D9970DFFCCFF1F2A8CBDD8A0CE" ID-DOI="http://doi.org/10.5281/zenodo.5709520" ID-GBIF-Taxon="190309938" ID-Zenodo-Dep="5709520" LSID="urn:lsid:plazi:treatment:3A1C87D9970DFFCCFF1F2A8CBDD8A0CE" httpUri="http://treatment.plazi.org/id/3A1C87D9970DFFCCFF1F2A8CBDD8A0CE" lastPageId="26" lastPageNumber="511" pageId="24" pageNumber="509">
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<subSubSection id="FAAF6544970DFFCEFF1F2A8CBB7DA3BC" box="[151,537,1477,1504]" pageId="24" pageNumber="509" type="nomenclature">
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<paragraph id="B20A36CF970DFFCEFF1F2A8CBB7DA3BC" blockId="24.[151,537,1477,1540]" box="[151,537,1477,1504]" pageId="24" pageNumber="509">
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<heading id="E94281A3970DFFCEFF1F2A8CBB7DA3BC" bold="true" box="[151,537,1477,1504]" fontSize="11" level="1" pageId="24" pageNumber="509" reason="1">
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<emphasis id="80C1EADD970DFFCEFF1F2A8CBB7DA3BC" bold="true" box="[151,537,1477,1504]" pageId="24" pageNumber="509">
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<taxonomicName id="75B54D4C970DFFCEFF1F2A8CB8D0A3BC" authority="Golovatch & Aswathy & Bhagirathan & Sudhikumar, 2021" authorityName="Golovatch & Aswathy & Bhagirathan & Sudhikumar" authorityYear="2021" box="[151,436,1477,1504]" class="Diplopoda" family="Paradoxosomatidae" genus="Delarthrum" kingdom="Animalia" order="Polydesmida" pageId="24" pageNumber="509" phylum="Arthropoda" rank="species" species="anomalans" status="sp. nov.">
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<emphasis id="80C1EADD970DFFCEFF1F2A8CB8D0A3BC" bold="true" box="[151,436,1477,1504]" italics="true" pageId="24" pageNumber="509">Delarthrum anomalans</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="9BF257A6970DFFCEFE332A8FBB7DA3BC" box="[443,537,1478,1504]" pageId="24" pageNumber="509" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="FAAF6544970DFFCEFF1F2AA3B878A058" box="[151,284,1514,1540]" pageId="24" pageNumber="509" type="description">
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<paragraph id="B20A36CF970DFFCEFF1F2AA3B878A058" blockId="24.[151,537,1477,1540]" box="[151,284,1514,1540]" pageId="24" pageNumber="509">
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<figureCitation id="2A8E2A4A970DFFCEFF1F2AA3B87CA058" box="[151,280,1514,1540]" captionStart-0="FIGURES 69–73" captionStart-1="FIGURES 74–80" captionStart-2="FIGURES 81–87" captionStartId-0="25.[151,264,1581,1605]" captionStartId-1="27.[151,264,1943,1967]" captionStartId-2="28.[151,264,1943,1967]" captionTargetBox-0="[170,1417,181,1557]" captionTargetBox-1="[280,1307,181,1919]" captionTargetBox-2="[291,1296,181,1919]" captionTargetId-0="figure-19@25.[170,1417,181,1557]" captionTargetId-1="figure-19@27.[280,1307,181,1919]" captionTargetId-2="figure-22@28.[291,1296,181,1919]" captionTargetPageId-0="25" captionTargetPageId-1="27" captionTargetPageId-2="28" captionText-0="FIGURES 69–73. Delarthrum anomalans sp. nov., paratypes. 69. Live male in the field, dorsal view (photographed not to scale). 70, 71. Habitus, fixed male and fixed female, respectively, lateral views. 72, 73. Fixed male, anterior and posterior parts of body, respectively, dorsal view." captionText-1="FIGURES 74–80. Delarthrum anomalans sp. nov., male paratypes. 74–76. Habitus, anterior, middle and posterior parts of body, respectively, lateral views. 77. Head, frontal view. 78. Male body rings 5–8 with legs, ventral view. 79. Telson, ventrocaudal view (photographed not to scale). 80. Both gonopods in situ, ventral view (photographed not to scale)." captionText-2="FIGURES 81–87. Delarthrum anomalans sp. nov., male paratypes. 81. Legs 1, lateral view. 82. Legs 2, lateral view. 83. Leg 26 (anterior pair on body ring 16), lateral view. 84–87. Entire right gonopod, mesal, submesal, mesal and subventral views, respectively. Abbreviations: a and b, distofemoral processes; fe, femorite; sl, solenomere; sph, solenophore; z, dorsobasal peg." figureDoi-0="http://doi.org/10.5281/zenodo.5709564" figureDoi-1="http://doi.org/10.5281/zenodo.5709566" figureDoi-2="http://doi.org/10.5281/zenodo.5709568" httpUri-0="https://zenodo.org/record/5709564/files/figure.png" httpUri-1="https://zenodo.org/record/5709566/files/figure.png" httpUri-2="https://zenodo.org/record/5709568/files/figure.png" pageId="24" pageNumber="509">Figs 69–87</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="FAAF6544970DFFCEFF1F2965BDCAA0A6" pageId="24" pageNumber="509" type="materials_examined">
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<paragraph id="B20A36CF970DFFCEFF1F2965BBF0A08A" blockId="24.[151,1436,1580,2039]" pageId="24" pageNumber="509">
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<emphasis id="80C1EADD970DFFCEFF1F2965B8E7A01A" bold="true" box="[151,387,1580,1606]" pageId="24" pageNumber="509">Material examined.</emphasis>
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<materialsCitation id="02DD3C92970DFFCEFE042965B98FA0D2" ID-GBIF-Occurrence="3409009301" country="India" elevation="29" latitude="10.355278" location="Thrissur district" longLatPrecision="21" longitude="76.21333" pageId="24" pageNumber="509" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Kerala" typeStatus="holotype">
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Male
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<typeStatus id="6D0E886D970DFFCEFE472965BB55A01A" box="[463,561,1580,1606]" pageId="24" pageNumber="509" type="holotype">holotype</typeStatus>
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(CATE-61604A),
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<specimenCount id="A4B3FD46970DFFCEFC802965BA31A01A" box="[776,853,1580,1606]" pageId="24" pageNumber="509" type="male">2 male</specimenCount>
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(CATE-61604B and CATE-61604C) and
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<specimenCount id="A4B3FD46970DFFCEFAB12965BCF8A01A" box="[1337,1436,1580,1606]" pageId="24" pageNumber="509" type="female">2 female</specimenCount>
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<typeStatus id="6D0E886D970DFFCEFF1F2918B866A037" box="[151,258,1617,1643]" pageId="24" pageNumber="509" type="paratype">paratypes</typeStatus>
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(CATE-61604D and CATE-61604E),
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<collectingCountry id="CAA2765F970DFFCEFD392919BB89A036" box="[689,749,1616,1642]" name="India" pageId="24" pageNumber="509">India</collectingCountry>
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,
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<collectingRegion id="7071F82D970DFFCEFD712919BAE5A037" box="[761,897,1616,1643]" country="India" name="Kerala" pageId="24" pageNumber="509">Kerala state</collectingRegion>
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,
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<location id="B76A6014970DFFCEFC042919BD25A036" LSID="urn:lsid:plazi:treatment:3A1C87D9970DFFCCFF1F2A8CBDD8A0CE:B76A6014970DFFCEFC042919BD25A036" box="[908,1089,1616,1642]" country="India" latitude="10.355278" longLatPrecision="21" longitude="76.21333" name="Thrissur district" pageId="24" pageNumber="509" stateProvince="Kerala">Thrissur district</location>
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,
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<geoCoordinate id="D7815008970DFFCEFBC42919BDB8A036" box="[1100,1244,1616,1642]" degrees="10" direction="north" minutes="21" orientation="latitude" pageId="24" pageNumber="509" precision="15" seconds="19" value="10.355278">10°21’19’’N</geoCoordinate>
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,
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<geoCoordinate id="D7815008970DFFCEFB602919BC10A036" box="[1256,1396,1616,1642]" degrees="76" direction="east" minutes="12" orientation="longitude" pageId="24" pageNumber="509" precision="15" seconds="48" value="76.21333">76°12’48’’E</geoCoordinate>
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,
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<elevation id="3998D1FC970DFFCEFA082919B98FA0D2" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="24" pageNumber="509" unit="m" value="29.0">
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<quantity id="754D9B2A970DFFCEFA082919B9C9A0D3" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="24" pageNumber="509" unit="m" value="29.0">29 m</quantity>
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a.s.l.
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</elevation>
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</materialsCitation>
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,
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<materialsCitation id="02DD3C92970DFFCEFF71293DBBF0A08A" ID-GBIF-Occurrence="3409009302" collectingDate="2021-06" collectorName="M. D. Aswathy" country="India" county="Valliyur" elevation="716" latitude="11.802222" location="Wayanad district" longLatPrecision="21" longitude="76.031944" municipality="Manathavady" pageId="24" pageNumber="509" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Kerala" typeStatus="paratype">
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in
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<date id="C60B100F970DFFCEFE9F293CB8EFA0D2" box="[279,395,1652,1679]" pageId="24" pageNumber="509" value="2021-06">
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<collectingDate id="D64FE9E7970DFFCEFE9F293CB8EFA0D2" box="[279,395,1652,1679]" pageId="24" pageNumber="509" value="2021-06">June 2021</collectingDate>
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</date>
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and
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<specimenCount id="A4B3FD46970DFFCEFE4D293DBB75A0D2" box="[453,529,1652,1678]" pageId="24" pageNumber="509" type="male">1 male</specimenCount>
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(CATE-61604J) and
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<specimenCount id="A4B3FD46970DFFCEFC8F293DBA0DA0D2" box="[775,873,1652,1678]" pageId="24" pageNumber="509" type="female">2 female</specimenCount>
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<typeStatus id="6D0E886D970DFFCEFCFA293CBAB9A0D3" box="[882,989,1653,1679]" pageId="24" pageNumber="509" type="paratype">paratypes</typeStatus>
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(CATE-61604K and CATE-61604L all in 5302B–CATE-5302I) from the sacred groves of
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<collectingCounty id="5B6B4E43970DFFCEFD6E29D1BA24A0EE" box="[742,832,1688,1714]" pageId="24" pageNumber="509">Valliyur</collectingCounty>
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kavu,
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<collectingMunicipality id="526EACB5970DFFCEFC1A29D1BD48A0EE" box="[914,1068,1688,1714]" pageId="24" pageNumber="509">Manathavady</collectingMunicipality>
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,
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<location id="B76A6014970DFFCEFBB129D1BD9FA0EE" LSID="urn:lsid:plazi:treatment:3A1C87D9970DFFCCFF1F2A8CBDD8A0CE:B76A6014970DFFCEFBB129D1BD9FA0EE" box="[1081,1275,1688,1714]" country="India" county="Valliyur" latitude="11.802222" longLatPrecision="21" longitude="76.031944" municipality="Manathavady" name="Wayanad district" pageId="24" pageNumber="509" stateProvince="Kerala">Wayanad district</location>
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,
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<geoCoordinate id="D7815008970DFFCEFA8129D1BCF3A0EE" box="[1289,1431,1688,1714]" degrees="11" direction="north" minutes="48" orientation="latitude" pageId="24" pageNumber="509" precision="15" seconds="08" value="11.802222">11°48’08’’N</geoCoordinate>
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,
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<geoCoordinate id="D7815008970DFFCEFF1F29F5B847A08B" box="[151,291,1724,1751]" degrees="76" direction="east" minutes="01" orientation="longitude" pageId="24" pageNumber="509" precision="15" seconds="55" value="76.031944">76°01’55’’E</geoCoordinate>
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,
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|
<elevation id="3998D1FC970DFFCEFEA729F5B8D7A08A" box="[303,435,1724,1751]" metricMagnitude="2" metricUnit="m" metricValue="7.159999999999999" pageId="24" pageNumber="509" unit="m" value="716.0">
|
|||
|
<quantity id="754D9B2A970DFFCEFEA729F5B812A08B" box="[303,374,1724,1751]" metricMagnitude="2" metricUnit="m" metricValue="7.159999999999999" pageId="24" pageNumber="509" unit="m" value="716.0">716 m</quantity>
|
|||
|
a.s.l.
|
|||
|
</elevation>
|
|||
|
,
|
|||
|
<collectorName id="1F405319970DFFCEFE3729F4BB00A08A" box="[447,612,1724,1751]" pageId="24" pageNumber="509">M.D. Aswathy</collectorName>
|
|||
|
leg.
|
|||
|
</materialsCitation>
|
|||
|
</paragraph>
|
|||
|
<paragraph id="B20A36CF970DFFCEFF4F29A9BDCAA0A6" blockId="24.[151,1436,1580,2039]" box="[199,1198,1760,1787]" pageId="24" pageNumber="509">
|
|||
|
<emphasis id="80C1EADD970DFFCEFF4F29A9B869A0A6" bold="true" box="[199,269,1760,1786]" pageId="24" pageNumber="509">Name</emphasis>
|
|||
|
. To emphasize the anomalous absence of adenostyles from both male legs 1 and 2.
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection id="FAAF6544970DFFCEFF4F284DBB9FA1AA" pageId="24" pageNumber="509" type="diagnosis">
|
|||
|
<paragraph id="B20A36CF970DFFCEFF4F284DBB9FA1AA" blockId="24.[151,1436,1580,2039]" pageId="24" pageNumber="509">
|
|||
|
<emphasis id="80C1EADD970DFFCEFF4F284DB85EA142" bold="true" box="[199,314,1796,1822]" pageId="24" pageNumber="509">Diagnosis</emphasis>
|
|||
|
. Differs from congeners by the absence of adenostyles from both male legs 1 and 2, coupled with the sexually dimorphic colour pattern, the transverse tergal sulci starting with the collum, and the peculiar gonopodal conformation, primarily the presence of a small, thumb-shaped, distoventral projection (
|
|||
|
<emphasis id="80C1EADD970DFFCEFBF32805BDEDA13A" bold="true" box="[1147,1161,1868,1894]" pageId="24" pageNumber="509">a</emphasis>
|
|||
|
) basal to the division of the postfemoral part into (1) the main, dorsal piece of the solenophore (
|
|||
|
<emphasis id="80C1EADD970DFFCEFC3E2839BA84A1D6" bold="true" box="[950,992,1904,1930]" pageId="24" pageNumber="509">sph</emphasis>
|
|||
|
), which is long, obliquely truncate dorsad at the base, bifid in the distal third and slightly curved mesad, and (2) a dorsal process that is split immediately at the base into a longer, stalked, leaf-shaped outgrowth (
|
|||
|
<emphasis id="80C1EADD970DFFCEFCAF28F1BA53A18E" bold="true" box="[807,823,1976,2002]" pageId="24" pageNumber="509">b</emphasis>
|
|||
|
) with peculiar, ragged, saw-like distal margins, and a much shorter, simple, dorsobasal peg (
|
|||
|
<emphasis id="80C1EADD970DFFCEFDCF2895BB37A1AA" bold="true" box="[583,595,2012,2038]" pageId="24" pageNumber="509">z</emphasis>
|
|||
|
) (
|
|||
|
<figureCitation id="2A8E2A4A970DFFCEFDE42895BB8BA1AA" box="[620,751,2012,2038]" captionStart="FIGURES 81–87" captionStartId="28.[151,264,1943,1967]" captionTargetBox="[291,1296,181,1919]" captionTargetId="figure-22@28.[291,1296,181,1919]" captionTargetPageId="28" captionText="FIGURES 81–87. Delarthrum anomalans sp. nov., male paratypes. 81. Legs 1, lateral view. 82. Legs 2, lateral view. 83. Leg 26 (anterior pair on body ring 16), lateral view. 84–87. Entire right gonopod, mesal, submesal, mesal and subventral views, respectively. Abbreviations: a and b, distofemoral processes; fe, femorite; sl, solenomere; sph, solenophore; z, dorsobasal peg." figureDoi="http://doi.org/10.5281/zenodo.5709568" httpUri="https://zenodo.org/record/5709568/files/figure.png" pageId="24" pageNumber="509">Figs 84–87</figureCitation>
|
|||
|
).
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<caption id="E6CA6647970CFFCFFF1F2964B895A0D1" ID-DOI="http://doi.org/10.5281/zenodo.5709564" ID-Zenodo-Dep="5709564" httpUri="https://zenodo.org/record/5709564/files/figure.png" pageId="25" pageNumber="510" startId="25.[151,264,1581,1605]" targetBox="[170,1417,181,1557]" targetPageId="25">
|
|||
|
<paragraph id="B20A36CF970CFFCFFF1F2964B895A0D1" blockId="25.[151,1436,1581,1677]" pageId="25" pageNumber="510">
|
|||
|
<emphasis id="80C1EADD970CFFCFFF1F2964B832A019" bold="true" box="[151,342,1581,1605]" pageId="25" pageNumber="510">FIGURES 69–73.</emphasis>
|
|||
|
<taxonomicName id="75B54D4C970CFFCFFED72964BB2AA019" authority="Golovatch & Aswathy & Bhagirathan & Sudhikumar, 2021" authorityName="Golovatch & Aswathy & Bhagirathan & Sudhikumar" authorityYear="2021" box="[351,590,1581,1605]" class="Diplopoda" family="Paradoxosomatidae" genus="Delarthrum" kingdom="Animalia" order="Polydesmida" pageId="25" pageNumber="510" phylum="Arthropoda" rank="species" species="anomalans" status="sp. nov.">
|
|||
|
<emphasis id="80C1EADD970CFFCFFED72964BB2AA019" box="[351,590,1581,1605]" italics="true" pageId="25" pageNumber="510">Delarthrum anomalans</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
<emphasis id="80C1EADD970CFFCFFDDF2964BBD4A019" bold="true" box="[599,688,1581,1605]" pageId="25" pageNumber="510">
|
|||
|
<taxonomicNameLabel id="9BF257A6970CFFCFFDDF2964BBC8A019" box="[599,684,1581,1605]" pageId="25" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
,
|
|||
|
</emphasis>
|
|||
|
paratypes.
|
|||
|
<emphasis id="80C1EADD970CFFCFFCA12964BA26A019" bold="true" box="[809,834,1581,1605]" pageId="25" pageNumber="510">69</emphasis>
|
|||
|
. Live male in the field, dorsal view (photographed not to scale).
|
|||
|
<emphasis id="80C1EADD970CFFCFFF572918B87AA035" bold="true" box="[223,286,1617,1641]" pageId="25" pageNumber="510">70, 71</emphasis>
|
|||
|
. Habitus, fixed male and fixed female, respectively, lateral views.
|
|||
|
<emphasis id="80C1EADD970CFFCFFC492918BD64A035" bold="true" box="[961,1024,1617,1641]" pageId="25" pageNumber="510">72, 73</emphasis>
|
|||
|
. Fixed male, anterior and posterior parts of body, respectively, dorsal view.
|
|||
|
</paragraph>
|
|||
|
</caption>
|
|||
|
<subSubSection id="FAAF6544970CFFCCFF4F29F2BDD8A0CE" lastPageId="26" lastPageNumber="511" pageId="25" pageNumber="510" type="description">
|
|||
|
<paragraph id="B20A36CF970CFFCFFF4F29F2BC69A13A" blockId="25.[151,1437,1723,2038]" pageId="25" pageNumber="510">
|
|||
|
<emphasis id="80C1EADD970CFFCFFF4F29F2B835A089" bold="true" box="[199,337,1723,1749]" pageId="25" pageNumber="510">Description</emphasis>
|
|||
|
. Length of
|
|||
|
<typeStatus id="6D0E886D970CFFCFFE5529F2BB26A089" box="[477,578,1723,1749]" pageId="25" pageNumber="510" type="holotype">holotype</typeStatus>
|
|||
|
, 17.9 mm, of
|
|||
|
<typeStatus id="6D0E886D970CFFCFFD6029F5BA32A08A" box="[744,854,1724,1750]" pageId="25" pageNumber="510" type="paratype">paratypes</typeStatus>
|
|||
|
, 17.0–17.6 mm (male) or 14.0–14.8 mm (female). Width of midbody rings, 2.6 mm (
|
|||
|
<typeStatus id="6D0E886D970CFFCFFD9E2996BB1AA0A6" box="[534,638,1759,1786]" pageId="25" pageNumber="510" type="holotype">holotype</typeStatus>
|
|||
|
), 1.9–2.1 mm (male
|
|||
|
<typeStatus id="6D0E886D970CFFCFFCEC29A9BAB5A0A6" box="[868,977,1760,1786]" pageId="25" pageNumber="510" type="paratype">paratypes</typeStatus>
|
|||
|
) or 2.0–2.3 mm (female
|
|||
|
<typeStatus id="6D0E886D970CFFCFFB6029A9BC3DA0A6" box="[1256,1369,1760,1786]" pageId="25" pageNumber="510" type="paratype">paratypes</typeStatus>
|
|||
|
). Coloration sex-dimorphic:
|
|||
|
<emphasis id="80C1EADD970CFFCFFE13284DB888A141" box="[411,492,1796,1821]" italics="true" pageId="25" pageNumber="510">in vivo,</emphasis>
|
|||
|
dorsum dark brown to black, venter chestnut brown (male) (
|
|||
|
<figureCitation id="2A8E2A4A970CFFCFFB01284ABC62A141" box="[1161,1286,1795,1821]" captionStart="FIGURES 69–73" captionStartId="25.[151,264,1581,1605]" captionTargetBox="[170,1417,181,1557]" captionTargetId="figure-19@25.[170,1417,181,1557]" captionTargetPageId="25" captionText="FIGURES 69–73. Delarthrum anomalans sp. nov., paratypes. 69. Live male in the field, dorsal view (photographed not to scale). 70, 71. Habitus, fixed male and fixed female, respectively, lateral views. 72, 73. Fixed male, anterior and posterior parts of body, respectively, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5709564" httpUri="https://zenodo.org/record/5709564/files/figure.png" pageId="25" pageNumber="510">Figs 69, 70</figureCitation>
|
|||
|
) to yellowish white (female) (
|
|||
|
<figureCitation id="2A8E2A4A970CFFCFFEC1286EB8F8A11D" box="[329,412,1831,1857]" captionStart="FIGURES 69–73" captionStartId="25.[151,264,1581,1605]" captionTargetBox="[170,1417,181,1557]" captionTargetId="figure-19@25.[170,1417,181,1557]" captionTargetPageId="25" captionText="FIGURES 69–73. Delarthrum anomalans sp. nov., paratypes. 69. Live male in the field, dorsal view (photographed not to scale). 70, 71. Habitus, fixed male and fixed female, respectively, lateral views. 72, 73. Fixed male, anterior and posterior parts of body, respectively, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5709564" httpUri="https://zenodo.org/record/5709564/files/figure.png" pageId="25" pageNumber="510">Fig. 71</figureCitation>
|
|||
|
); paraterga mostly contrasting pale. Legs light brown (male) or whitish (female). Coloration after one month of preservation in 96% ethanol somewhat darkened, but pattern evident (
|
|||
|
<figureCitation id="2A8E2A4A970CFFCFFBF62802BC65A13A" box="[1150,1281,1867,1894]" captionStart-0="FIGURES 69–73" captionStart-1="FIGURES 74–80" captionStartId-0="25.[151,264,1581,1605]" captionStartId-1="27.[151,264,1943,1967]" captionTargetBox-0="[170,1417,181,1557]" captionTargetBox-1="[280,1307,181,1919]" captionTargetId-0="figure-19@25.[170,1417,181,1557]" captionTargetId-1="figure-19@27.[280,1307,181,1919]" captionTargetPageId-0="25" captionTargetPageId-1="27" captionText-0="FIGURES 69–73. Delarthrum anomalans sp. nov., paratypes. 69. Live male in the field, dorsal view (photographed not to scale). 70, 71. Habitus, fixed male and fixed female, respectively, lateral views. 72, 73. Fixed male, anterior and posterior parts of body, respectively, dorsal view." captionText-1="FIGURES 74–80. Delarthrum anomalans sp. nov., male paratypes. 74–76. Habitus, anterior, middle and posterior parts of body, respectively, lateral views. 77. Head, frontal view. 78. Male body rings 5–8 with legs, ventral view. 79. Telson, ventrocaudal view (photographed not to scale). 80. Both gonopods in situ, ventral view (photographed not to scale)." figureDoi-0="http://doi.org/10.5281/zenodo.5709564" figureDoi-1="http://doi.org/10.5281/zenodo.5709566" httpUri-0="https://zenodo.org/record/5709564/files/figure.png" httpUri-1="https://zenodo.org/record/5709566/files/figure.png" pageId="25" pageNumber="510">Figs 69–76</figureCitation>
|
|||
|
).
|
|||
|
</paragraph>
|
|||
|
<paragraph id="B20A36CF970CFFCCFF4F2826BBEBA741" blockId="25.[151,1437,1723,2038]" lastBlockId="26.[151,1437,150,1682]" lastPageId="26" lastPageNumber="511" pageId="25" pageNumber="510">
|
|||
|
Body with 20 rings in both sexes. In width, collum <head <ring 2<3<4<5<6<7<8<9=10, body gradually tapering thereafter (
|
|||
|
<figureCitation id="2A8E2A4A970CFFCFFEB628DAB8A6A1F1" box="[318,450,1939,1965]" captionStart="FIGURES 69–73" captionStartId="25.[151,264,1581,1605]" captionTargetBox="[170,1417,181,1557]" captionTargetId="figure-19@25.[170,1417,181,1557]" captionTargetPageId="25" captionText="FIGURES 69–73. Delarthrum anomalans sp. nov., paratypes. 69. Live male in the field, dorsal view (photographed not to scale). 70, 71. Habitus, fixed male and fixed female, respectively, lateral views. 72, 73. Fixed male, anterior and posterior parts of body, respectively, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5709564" httpUri="https://zenodo.org/record/5709564/files/figure.png" pageId="25" pageNumber="510">Figs 72, 73</figureCitation>
|
|||
|
). Vertex glossy and bare; stipes, cardo and clypeus moderately setose, epicranial sulcus distinct and ending before the level of antennal sockets. Clypeal region with
|
|||
|
<emphasis id="80C1EADD970CFFCFFC7A28F1BD68A18D" box="[1010,1036,1976,2001]" italics="true" pageId="25" pageNumber="510">ca</emphasis>
|
|||
|
5+5 setiferous pits. Antennae moderately long, only slightly clavate, extending back to ring 4 when stretched caudolaterally (
|
|||
|
<figureCitation id="2A8E2A4A970CFFCFFB2C2892BD99A1A9" box="[1188,1277,2011,2037]" captionStart="FIGURES 69–73" captionStartId="25.[151,264,1581,1605]" captionTargetBox="[170,1417,181,1557]" captionTargetId="figure-19@25.[170,1417,181,1557]" captionTargetPageId="25" captionText="FIGURES 69–73. Delarthrum anomalans sp. nov., paratypes. 69. Live male in the field, dorsal view (photographed not to scale). 70, 71. Habitus, fixed male and fixed female, respectively, lateral views. 72, 73. Fixed male, anterior and posterior parts of body, respectively, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5709564" httpUri="https://zenodo.org/record/5709564/files/figure.png" pageId="25" pageNumber="510">Figs 72</figureCitation>
|
|||
|
,
|
|||
|
<figureCitation id="2A8E2A4A970CFFCFFA822892BC4EA1A9" box="[1290,1322,2011,2037]" captionStart="FIGURES 74–80" captionStartId="27.[151,264,1943,1967]" captionTargetBox="[280,1307,181,1919]" captionTargetId="figure-19@27.[280,1307,181,1919]" captionTargetPageId="27" captionText="FIGURES 74–80. Delarthrum anomalans sp. nov., male paratypes. 74–76. Habitus, anterior, middle and posterior parts of body, respectively, lateral views. 77. Head, frontal view. 78. Male body rings 5–8 with legs, ventral view. 79. Telson, ventrocaudal view (photographed not to scale). 80. Both gonopods in situ, ventral view (photographed not to scale)." figureDoi="http://doi.org/10.5281/zenodo.5709566" httpUri="https://zenodo.org/record/5709566/files/figure.png" pageId="25" pageNumber="510">74</figureCitation>
|
|||
|
). Relative antennomere lengths: 1<6<4<2<5<3. Antennomere 6 broadest apically, post-antennal groove shallow, diameter of antennal socket and isthmus between sockets subequal, 0.24 and 0.23 mm, respectively (
|
|||
|
<figureCitation id="2A8E2A4A970FFFCCFB072FF2BD81A689" box="[1167,1253,187,213]" captionStart="FIGURES 74–80" captionStartId="27.[151,264,1943,1967]" captionTargetBox="[280,1307,181,1919]" captionTargetId="figure-19@27.[280,1307,181,1919]" captionTargetPageId="27" captionText="FIGURES 74–80. Delarthrum anomalans sp. nov., male paratypes. 74–76. Habitus, anterior, middle and posterior parts of body, respectively, lateral views. 77. Head, frontal view. 78. Male body rings 5–8 with legs, ventral view. 79. Telson, ventrocaudal view (photographed not to scale). 80. Both gonopods in situ, ventral view (photographed not to scale)." figureDoi="http://doi.org/10.5281/zenodo.5709566" httpUri="https://zenodo.org/record/5709566/files/figure.png" pageId="26" pageNumber="511">Fig. 77</figureCitation>
|
|||
|
). Antennomeres 5–7 each with a distinct dorso-apical group of basiconic sensilla (both male and female), these being especially conspicuous on antennomere 5 (
|
|||
|
<figureCitation id="2A8E2A4A970FFFCCFD882E4ABB3CA741" box="[512,600,259,285]" captionStart="FIGURES 69–73" captionStartId="25.[151,264,1581,1605]" captionTargetBox="[170,1417,181,1557]" captionTargetId="figure-19@25.[170,1417,181,1557]" captionTargetPageId="25" captionText="FIGURES 69–73. Delarthrum anomalans sp. nov., paratypes. 69. Live male in the field, dorsal view (photographed not to scale). 70, 71. Habitus, fixed male and fixed female, respectively, lateral views. 72, 73. Fixed male, anterior and posterior parts of body, respectively, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5709564" httpUri="https://zenodo.org/record/5709564/files/figure.png" pageId="26" pageNumber="511">Figs 72</figureCitation>
|
|||
|
,
|
|||
|
<figureCitation id="2A8E2A4A970FFFCCFDEB2E4ABBE7A741" box="[611,643,259,285]" captionStart="FIGURES 74–80" captionStartId="27.[151,264,1943,1967]" captionTargetBox="[280,1307,181,1919]" captionTargetId="figure-19@27.[280,1307,181,1919]" captionTargetPageId="27" captionText="FIGURES 74–80. Delarthrum anomalans sp. nov., male paratypes. 74–76. Habitus, anterior, middle and posterior parts of body, respectively, lateral views. 77. Head, frontal view. 78. Male body rings 5–8 with legs, ventral view. 79. Telson, ventrocaudal view (photographed not to scale). 80. Both gonopods in situ, ventral view (photographed not to scale)." figureDoi="http://doi.org/10.5281/zenodo.5709566" httpUri="https://zenodo.org/record/5709566/files/figure.png" pageId="26" pageNumber="511">74</figureCitation>
|
|||
|
).
|
|||
|
</paragraph>
|
|||
|
<paragraph id="B20A36CF970FFFCCFF4F2E6EB85FA4D9" blockId="26.[151,1437,150,1682]" pageId="26" pageNumber="511">
|
|||
|
Paraterga mostly well developed, keel-shaped, set at about upper third of body height (
|
|||
|
<figureCitation id="2A8E2A4A970FFFCCFB262E6EBC50A71E" box="[1198,1332,295,322]" captionStart-0="FIGURES 69–73" captionStart-1="FIGURES 74–80" captionStartId-0="25.[151,264,1581,1605]" captionStartId-1="27.[151,264,1943,1967]" captionTargetBox-0="[170,1417,181,1557]" captionTargetBox-1="[280,1307,181,1919]" captionTargetId-0="figure-19@25.[170,1417,181,1557]" captionTargetId-1="figure-19@27.[280,1307,181,1919]" captionTargetPageId-0="25" captionTargetPageId-1="27" captionText-0="FIGURES 69–73. Delarthrum anomalans sp. nov., paratypes. 69. Live male in the field, dorsal view (photographed not to scale). 70, 71. Habitus, fixed male and fixed female, respectively, lateral views. 72, 73. Fixed male, anterior and posterior parts of body, respectively, dorsal view." captionText-1="FIGURES 74–80. Delarthrum anomalans sp. nov., male paratypes. 74–76. Habitus, anterior, middle and posterior parts of body, respectively, lateral views. 77. Head, frontal view. 78. Male body rings 5–8 with legs, ventral view. 79. Telson, ventrocaudal view (photographed not to scale). 80. Both gonopods in situ, ventral view (photographed not to scale)." figureDoi-0="http://doi.org/10.5281/zenodo.5709564" figureDoi-1="http://doi.org/10.5281/zenodo.5709566" httpUri-0="https://zenodo.org/record/5709564/files/figure.png" httpUri-1="https://zenodo.org/record/5709566/files/figure.png" pageId="26" pageNumber="511">Figs 69–76</figureCitation>
|
|||
|
). Collum ovoid, paraterga small, subtriangular and broadly rounded. Anterior margin of collum with a single transverse row of fragile short setae (
|
|||
|
<figureCitation id="2A8E2A4A970FFFCCFE1A2E26B88CA7D5" box="[402,488,367,393]" captionStart="FIGURES 69–73" captionStartId="25.[151,264,1581,1605]" captionTargetBox="[170,1417,181,1557]" captionTargetId="figure-19@25.[170,1417,181,1557]" captionTargetPageId="25" captionText="FIGURES 69–73. Delarthrum anomalans sp. nov., paratypes. 69. Live male in the field, dorsal view (photographed not to scale). 70, 71. Habitus, fixed male and fixed female, respectively, lateral views. 72, 73. Fixed male, anterior and posterior parts of body, respectively, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5709564" httpUri="https://zenodo.org/record/5709564/files/figure.png" pageId="26" pageNumber="511">Fig. 72</figureCitation>
|
|||
|
). Anterior and posterior margins of following rings rather straight and parallel. Anterior margin with a transverse row of 2+2 short setae similar to those on collum. Paraterga 2 as usual, set clearly lower than 3
|
|||
|
<superScript id="45C09B87970FFFCCFEAC2EFFB856A798" attach="none" box="[292,306,438,452]" fontSize="6" pageId="26" pageNumber="511">rd</superScript>
|
|||
|
, squarish in shape, anterior corner/lobe drawn anteriad. Following paraterga 3–17 regularly rounded anterolaterally and increasingly drawn caudolaterad into acute triangles, but still lying within rear tergal margins (
|
|||
|
<figureCitation id="2A8E2A4A970FFFCCFF172EB6B844A446" box="[159,288,511,538]" captionStart-0="FIGURES 69–73" captionStart-1="FIGURES 74–80" captionStartId-0="25.[151,264,1581,1605]" captionStartId-1="27.[151,264,1943,1967]" captionTargetBox-0="[170,1417,181,1557]" captionTargetBox-1="[280,1307,181,1919]" captionTargetId-0="figure-19@25.[170,1417,181,1557]" captionTargetId-1="figure-19@27.[280,1307,181,1919]" captionTargetPageId-0="25" captionTargetPageId-1="27" captionText-0="FIGURES 69–73. Delarthrum anomalans sp. nov., paratypes. 69. Live male in the field, dorsal view (photographed not to scale). 70, 71. Habitus, fixed male and fixed female, respectively, lateral views. 72, 73. Fixed male, anterior and posterior parts of body, respectively, dorsal view." captionText-1="FIGURES 74–80. Delarthrum anomalans sp. nov., male paratypes. 74–76. Habitus, anterior, middle and posterior parts of body, respectively, lateral views. 77. Head, frontal view. 78. Male body rings 5–8 with legs, ventral view. 79. Telson, ventrocaudal view (photographed not to scale). 80. Both gonopods in situ, ventral view (photographed not to scale)." figureDoi-0="http://doi.org/10.5281/zenodo.5709564" figureDoi-1="http://doi.org/10.5281/zenodo.5709566" httpUri-0="https://zenodo.org/record/5709564/files/figure.png" httpUri-1="https://zenodo.org/record/5709566/files/figure.png" pageId="26" pageNumber="511">Figs 72–76</figureCitation>
|
|||
|
),
|
|||
|
<emphasis id="80C1EADD970FFFCCFEBA2EB6B835A444" box="[306,337,511,536]" italics="true" pageId="26" pageNumber="511">vs.</emphasis>
|
|||
|
small, spiniform and slightly drawn past rear tergal margin on rings 18 and 19 (
|
|||
|
<figureCitation id="2A8E2A4A970FFFCCFB332EB6BC74A445" box="[1211,1296,511,537]" captionStart="FIGURES 69–73" captionStartId="25.[151,264,1581,1605]" captionTargetBox="[170,1417,181,1557]" captionTargetId="figure-19@25.[170,1417,181,1557]" captionTargetPageId="25" captionText="FIGURES 69–73. Delarthrum anomalans sp. nov., paratypes. 69. Live male in the field, dorsal view (photographed not to scale). 70, 71. Habitus, fixed male and fixed female, respectively, lateral views. 72, 73. Fixed male, anterior and posterior parts of body, respectively, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5709564" httpUri="https://zenodo.org/record/5709564/files/figure.png" pageId="26" pageNumber="511">Figs 73</figureCitation>
|
|||
|
,
|
|||
|
<figureCitation id="2A8E2A4A970FFFCCFA932EB6BC5FA446" box="[1307,1339,511,538]" captionStart="FIGURES 74–80" captionStartId="27.[151,264,1943,1967]" captionTargetBox="[280,1307,181,1919]" captionTargetId="figure-19@27.[280,1307,181,1919]" captionTargetPageId="27" captionText="FIGURES 74–80. Delarthrum anomalans sp. nov., male paratypes. 74–76. Habitus, anterior, middle and posterior parts of body, respectively, lateral views. 77. Head, frontal view. 78. Male body rings 5–8 with legs, ventral view. 79. Telson, ventrocaudal view (photographed not to scale). 80. Both gonopods in situ, ventral view (photographed not to scale)." figureDoi="http://doi.org/10.5281/zenodo.5709566" httpUri="https://zenodo.org/record/5709566/files/figure.png" pageId="26" pageNumber="511">76</figureCitation>
|
|||
|
). Lateral calluses of paraterga thin, only slightly thicker on pore-bearing rings than on poreless ones (
|
|||
|
<figureCitation id="2A8E2A4A970FFFCCFB222D6ABC4AA461" box="[1194,1326,547,573]" captionStart="FIGURES 69–73" captionStartId="25.[151,264,1581,1605]" captionTargetBox="[170,1417,181,1557]" captionTargetId="figure-19@25.[170,1417,181,1557]" captionTargetPageId="25" captionText="FIGURES 69–73. Delarthrum anomalans sp. nov., paratypes. 69. Live male in the field, dorsal view (photographed not to scale). 70, 71. Habitus, fixed male and fixed female, respectively, lateral views. 72, 73. Fixed male, anterior and posterior parts of body, respectively, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5709564" httpUri="https://zenodo.org/record/5709564/files/figure.png" pageId="26" pageNumber="511">Figs 69, 70</figureCitation>
|
|||
|
), smooth, delimited by distinct sulci both dorsally and, to a lesser degree, ventrally; usually with a few short setae retained at lateral margin.
|
|||
|
</paragraph>
|
|||
|
<paragraph id="B20A36CF970FFFCCFF4F2DC6BDEEA502" blockId="26.[151,1437,150,1682]" pageId="26" pageNumber="511">
|
|||
|
Tegument generally smooth and shining, prozonae very delicately shagreened, metazonae mostly striate/striolate at bases of paraterga and near transverse sulci, surface being slightly undulate (
|
|||
|
<figureCitation id="2A8E2A4A970FFFCCFBBA2DFABDD7A491" box="[1074,1203,691,717]" captionStart="FIGURES 69–73" captionStartId="25.[151,264,1581,1605]" captionTargetBox="[170,1417,181,1557]" captionTargetId="figure-19@25.[170,1417,181,1557]" captionTargetPageId="25" captionText="FIGURES 69–73. Delarthrum anomalans sp. nov., paratypes. 69. Live male in the field, dorsal view (photographed not to scale). 70, 71. Habitus, fixed male and fixed female, respectively, lateral views. 72, 73. Fixed male, anterior and posterior parts of body, respectively, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5709564" httpUri="https://zenodo.org/record/5709564/files/figure.png" pageId="26" pageNumber="511">Figs 72, 73</figureCitation>
|
|||
|
). Tergal setae mostly short and inconspicuous, present both on paraterga and in anterior half of metaterga. Two deep transverse sulci present on collum (
|
|||
|
<figureCitation id="2A8E2A4A970FFFCCFEE72DB2B8A6A549" box="[367,450,763,789]" captionStart="FIGURES 69–73" captionStartId="25.[151,264,1581,1605]" captionTargetBox="[170,1417,181,1557]" captionTargetId="figure-19@25.[170,1417,181,1557]" captionTargetPageId="25" captionText="FIGURES 69–73. Delarthrum anomalans sp. nov., paratypes. 69. Live male in the field, dorsal view (photographed not to scale). 70, 71. Habitus, fixed male and fixed female, respectively, lateral views. 72, 73. Fixed male, anterior and posterior parts of body, respectively, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5709564" httpUri="https://zenodo.org/record/5709564/files/figure.png" pageId="26" pageNumber="511">Fig. 72</figureCitation>
|
|||
|
), followed by one prominent sulcus on metazonae 2–18, smooth at bottom and reaching the base of paraterga (
|
|||
|
<figureCitation id="2A8E2A4A970FFFCCFE052C56BB6BA565" box="[397,527,799,825]" captionStart="FIGURES 69–73" captionStartId="25.[151,264,1581,1605]" captionTargetBox="[170,1417,181,1557]" captionTargetId="figure-19@25.[170,1417,181,1557]" captionTargetPageId="25" captionText="FIGURES 69–73. Delarthrum anomalans sp. nov., paratypes. 69. Live male in the field, dorsal view (photographed not to scale). 70, 71. Habitus, fixed male and fixed female, respectively, lateral views. 72, 73. Fixed male, anterior and posterior parts of body, respectively, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5709564" httpUri="https://zenodo.org/record/5709564/files/figure.png" pageId="26" pageNumber="511">Figs 72, 73</figureCitation>
|
|||
|
). A faint axial line traceable on most metazonae until 18
|
|||
|
<superScript id="45C09B87970FFFCCFB0B2C57BDF4A570" attach="left" box="[1155,1168,798,812]" fontSize="6" pageId="26" pageNumber="511">th</superScript>
|
|||
|
(
|
|||
|
<figureCitation id="2A8E2A4A970FFFCCFB132C56BC7AA565" box="[1179,1310,799,825]" captionStart="FIGURES 69–73" captionStartId="25.[151,264,1581,1605]" captionTargetBox="[170,1417,181,1557]" captionTargetId="figure-19@25.[170,1417,181,1557]" captionTargetPageId="25" captionText="FIGURES 69–73. Delarthrum anomalans sp. nov., paratypes. 69. Live male in the field, dorsal view (photographed not to scale). 70, 71. Habitus, fixed male and fixed female, respectively, lateral views. 72, 73. Fixed male, anterior and posterior parts of body, respectively, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5709564" httpUri="https://zenodo.org/record/5709564/files/figure.png" pageId="26" pageNumber="511">Figs 72, 73</figureCitation>
|
|||
|
). Strictures between pro- and metazonae narrow, nearly smooth, at most faintly striolate (
|
|||
|
<figureCitation id="2A8E2A4A970FFFCCFC722C0ABD1AA502" box="[1018,1150,835,862]" captionStart-0="FIGURES 69–73" captionStart-1="FIGURES 74–80" captionStartId-0="25.[151,264,1581,1605]" captionStartId-1="27.[151,264,1943,1967]" captionTargetBox-0="[170,1417,181,1557]" captionTargetBox-1="[280,1307,181,1919]" captionTargetId-0="figure-19@25.[170,1417,181,1557]" captionTargetId-1="figure-19@27.[280,1307,181,1919]" captionTargetPageId-0="25" captionTargetPageId-1="27" captionText-0="FIGURES 69–73. Delarthrum anomalans sp. nov., paratypes. 69. Live male in the field, dorsal view (photographed not to scale). 70, 71. Habitus, fixed male and fixed female, respectively, lateral views. 72, 73. Fixed male, anterior and posterior parts of body, respectively, dorsal view." captionText-1="FIGURES 74–80. Delarthrum anomalans sp. nov., male paratypes. 74–76. Habitus, anterior, middle and posterior parts of body, respectively, lateral views. 77. Head, frontal view. 78. Male body rings 5–8 with legs, ventral view. 79. Telson, ventrocaudal view (photographed not to scale). 80. Both gonopods in situ, ventral view (photographed not to scale)." figureDoi-0="http://doi.org/10.5281/zenodo.5709564" figureDoi-1="http://doi.org/10.5281/zenodo.5709566" httpUri-0="https://zenodo.org/record/5709564/files/figure.png" httpUri-1="https://zenodo.org/record/5709566/files/figure.png" pageId="26" pageNumber="511">Figs 72–76</figureCitation>
|
|||
|
).
|
|||
|
</paragraph>
|
|||
|
<paragraph id="B20A36CF970FFFCCFF4F2C2EBAE2A5B1" blockId="26.[151,1437,150,1682]" pageId="26" pageNumber="511">
|
|||
|
Pore formula normal (5, 7, 9, 10, 12, 13, 15–19), ozopores small, circular, opening at
|
|||
|
<emphasis id="80C1EADD970FFFCCFB142C2EBDD2A5DC" box="[1180,1206,871,896]" italics="true" pageId="26" pageNumber="511">ca</emphasis>
|
|||
|
1/3 off caudolateral corner of pore-bearing calluses (
|
|||
|
<figureCitation id="2A8E2A4A970FFFCCFD8A2CC2BBE2A5FA" box="[514,646,907,934]" captionStart="FIGURES 74–80" captionStartId="27.[151,264,1943,1967]" captionTargetBox="[280,1307,181,1919]" captionTargetId="figure-19@27.[280,1307,181,1919]" captionTargetPageId="27" captionText="FIGURES 74–80. Delarthrum anomalans sp. nov., male paratypes. 74–76. Habitus, anterior, middle and posterior parts of body, respectively, lateral views. 77. Head, frontal view. 78. Male body rings 5–8 with legs, ventral view. 79. Telson, ventrocaudal view (photographed not to scale). 80. Both gonopods in situ, ventral view (photographed not to scale)." figureDoi="http://doi.org/10.5281/zenodo.5709566" httpUri="https://zenodo.org/record/5709566/files/figure.png" pageId="26" pageNumber="511">Figs 74–76</figureCitation>
|
|||
|
). Pleurosternal carinae decreasing in size from 2 to 4. Epiproct conical and flattened dorsoventrally, extending well beyond anal valves; tip truncate, subapical lateral incisions evident; hypoproct subtriangular, with 1+1 setae on minute knobs (
|
|||
|
<figureCitation id="2A8E2A4A970FFFCCFCAD2C9ABA1DA5B1" box="[805,889,979,1005]" captionStart="FIGURES 74–80" captionStartId="27.[151,264,1943,1967]" captionTargetBox="[280,1307,181,1919]" captionTargetId="figure-19@27.[280,1307,181,1919]" captionTargetPageId="27" captionText="FIGURES 74–80. Delarthrum anomalans sp. nov., male paratypes. 74–76. Habitus, anterior, middle and posterior parts of body, respectively, lateral views. 77. Head, frontal view. 78. Male body rings 5–8 with legs, ventral view. 79. Telson, ventrocaudal view (photographed not to scale). 80. Both gonopods in situ, ventral view (photographed not to scale)." figureDoi="http://doi.org/10.5281/zenodo.5709566" httpUri="https://zenodo.org/record/5709566/files/figure.png" pageId="26" pageNumber="511">Fig. 79</figureCitation>
|
|||
|
).
|
|||
|
</paragraph>
|
|||
|
<paragraph id="B20A36CF970FFFCCFF4F2CBEBB76A29F" blockId="26.[151,1437,150,1682]" pageId="26" pageNumber="511">
|
|||
|
Legs moderately long and slender, clearly longer and thickened in male compared to female (
|
|||
|
<figureCitation id="2A8E2A4A970FFFCCFB412CBEBC2CA24D" box="[1225,1352,1015,1041]" captionStart="FIGURES 69–73" captionStartId="25.[151,264,1581,1605]" captionTargetBox="[170,1417,181,1557]" captionTargetId="figure-19@25.[170,1417,181,1557]" captionTargetPageId="25" captionText="FIGURES 69–73. Delarthrum anomalans sp. nov., paratypes. 69. Live male in the field, dorsal view (photographed not to scale). 70, 71. Habitus, fixed male and fixed female, respectively, lateral views. 72, 73. Fixed male, anterior and posterior parts of body, respectively, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5709564" httpUri="https://zenodo.org/record/5709564/files/figure.png" pageId="26" pageNumber="511">Figs 70, 71</figureCitation>
|
|||
|
), rather densely setose ventrally and with short claws; male legs with tarsal brushes absent only from last two leg-pairs (
|
|||
|
<figureCitation id="2A8E2A4A970FFFCCFAE22B52B86EA204" captionStart="FIGURES 74–80" captionStartId="27.[151,264,1943,1967]" captionTargetBox="[280,1307,181,1919]" captionTargetId="figure-19@27.[280,1307,181,1919]" captionTargetPageId="27" captionText="FIGURES 74–80. Delarthrum anomalans sp. nov., male paratypes. 74–76. Habitus, anterior, middle and posterior parts of body, respectively, lateral views. 77. Head, frontal view. 78. Male body rings 5–8 with legs, ventral view. 79. Telson, ventrocaudal view (photographed not to scale). 80. Both gonopods in situ, ventral view (photographed not to scale)." figureDoi="http://doi.org/10.5281/zenodo.5709566" httpUri="https://zenodo.org/record/5709566/files/figure.png" pageId="26" pageNumber="511">Figs 74–76, 78</figureCitation>
|
|||
|
,
|
|||
|
<figureCitation id="2A8E2A4A970FFFCCFE902B77B853A204" box="[280,311,1086,1112]" captionStart="FIGURES 81–87" captionStartId="28.[151,264,1943,1967]" captionTargetBox="[291,1296,181,1919]" captionTargetId="figure-22@28.[291,1296,181,1919]" captionTargetPageId="28" captionText="FIGURES 81–87. Delarthrum anomalans sp. nov., male paratypes. 81. Legs 1, lateral view. 82. Legs 2, lateral view. 83. Leg 26 (anterior pair on body ring 16), lateral view. 84–87. Entire right gonopod, mesal, submesal, mesal and subventral views, respectively. Abbreviations: a and b, distofemoral processes; fe, femorite; sl, solenomere; sph, solenophore; z, dorsobasal peg." figureDoi="http://doi.org/10.5281/zenodo.5709568" httpUri="https://zenodo.org/record/5709568/files/figure.png" pageId="26" pageNumber="511">83</figureCitation>
|
|||
|
). Prefemora and coxae each bearing a moderately long and stiff distoventral seta. Podomere length ratios: coxa <prefemur <tibia = postfemur <tarsus <femur (
|
|||
|
<figureCitation id="2A8E2A4A970FFFCCFCDC2B28BACCA220" box="[852,936,1121,1148]" captionStart="FIGURES 81–87" captionStartId="28.[151,264,1943,1967]" captionTargetBox="[291,1296,181,1919]" captionTargetId="figure-22@28.[291,1296,181,1919]" captionTargetPageId="28" captionText="FIGURES 81–87. Delarthrum anomalans sp. nov., male paratypes. 81. Legs 1, lateral view. 82. Legs 2, lateral view. 83. Leg 26 (anterior pair on body ring 16), lateral view. 84–87. Entire right gonopod, mesal, submesal, mesal and subventral views, respectively. Abbreviations: a and b, distofemoral processes; fe, femorite; sl, solenomere; sph, solenophore; z, dorsobasal peg." figureDoi="http://doi.org/10.5281/zenodo.5709568" httpUri="https://zenodo.org/record/5709568/files/figure.png" pageId="26" pageNumber="511">Fig. 83</figureCitation>
|
|||
|
). Neither male leg-pair 1 nor 2 with any adenostyles, only coxae 2 as usual, with gonopores on small knobs (
|
|||
|
<figureCitation id="2A8E2A4A970FFFCCFCF52BCCBD65A2C3" box="[893,1025,1157,1183]" captionStart="FIGURES 81–87" captionStartId="28.[151,264,1943,1967]" captionTargetBox="[291,1296,181,1919]" captionTargetId="figure-22@28.[291,1296,181,1919]" captionTargetPageId="28" captionText="FIGURES 81–87. Delarthrum anomalans sp. nov., male paratypes. 81. Legs 1, lateral view. 82. Legs 2, lateral view. 83. Leg 26 (anterior pair on body ring 16), lateral view. 84–87. Entire right gonopod, mesal, submesal, mesal and subventral views, respectively. Abbreviations: a and b, distofemoral processes; fe, femorite; sl, solenomere; sph, solenophore; z, dorsobasal peg." figureDoi="http://doi.org/10.5281/zenodo.5709568" httpUri="https://zenodo.org/record/5709568/files/figure.png" pageId="26" pageNumber="511">Figs 81, 82</figureCitation>
|
|||
|
). Sternal lobe between male coxae 4 small, conical and densely setose.
|
|||
|
</paragraph>
|
|||
|
<paragraph id="B20A36CF970FFFCCFF4F2B85BDD8A0CE" blockId="26.[151,1437,150,1682]" pageId="26" pageNumber="511">
|
|||
|
Gonopodal aperture transverse and suboval in shape, about 2/3 as wide as prozonite 7, its rim being not elevated, but thickened laterally (
|
|||
|
<figureCitation id="2A8E2A4A970FFFCCFE722BA6BB2AA356" box="[506,590,1263,1290]" captionStart="FIGURES 74–80" captionStartId="27.[151,264,1943,1967]" captionTargetBox="[280,1307,181,1919]" captionTargetId="figure-19@27.[280,1307,181,1919]" captionTargetPageId="27" captionText="FIGURES 74–80. Delarthrum anomalans sp. nov., male paratypes. 74–76. Habitus, anterior, middle and posterior parts of body, respectively, lateral views. 77. Head, frontal view. 78. Male body rings 5–8 with legs, ventral view. 79. Telson, ventrocaudal view (photographed not to scale). 80. Both gonopods in situ, ventral view (photographed not to scale)." figureDoi="http://doi.org/10.5281/zenodo.5709566" httpUri="https://zenodo.org/record/5709566/files/figure.png" pageId="26" pageNumber="511">Fig. 78</figureCitation>
|
|||
|
). Gonopods high, complex,
|
|||
|
<emphasis id="80C1EADD970FFFCCFC042BB9BAB5A355" box="[908,977,1264,1289]" italics="true" pageId="26" pageNumber="511">in situ</emphasis>
|
|||
|
held parallel to each other (
|
|||
|
<figureCitation id="2A8E2A4A970FFFCCFA842BA6BCF4A356" box="[1292,1424,1263,1290]" captionStart="FIGURES 74–80" captionStartId="27.[151,264,1943,1967]" captionTargetBox="[280,1307,181,1919]" captionTargetId="figure-19@27.[280,1307,181,1919]" captionTargetPageId="27" captionText="FIGURES 74–80. Delarthrum anomalans sp. nov., male paratypes. 74–76. Habitus, anterior, middle and posterior parts of body, respectively, lateral views. 77. Head, frontal view. 78. Male body rings 5–8 with legs, ventral view. 79. Telson, ventrocaudal view (photographed not to scale). 80. Both gonopods in situ, ventral view (photographed not to scale)." figureDoi="http://doi.org/10.5281/zenodo.5709566" httpUri="https://zenodo.org/record/5709566/files/figure.png" pageId="26" pageNumber="511">Figs 78, 80</figureCitation>
|
|||
|
). Coxite darkened, about as long as femorite, subcylindrical, setose distoventrally (
|
|||
|
<figureCitation id="2A8E2A4A970FFFCCFBC42A5ABDB0A371" box="[1100,1236,1299,1325]" captionStart="FIGURES 81–87" captionStartId="28.[151,264,1943,1967]" captionTargetBox="[291,1296,181,1919]" captionTargetId="figure-22@28.[291,1296,181,1919]" captionTargetPageId="28" captionText="FIGURES 81–87. Delarthrum anomalans sp. nov., male paratypes. 81. Legs 1, lateral view. 82. Legs 2, lateral view. 83. Leg 26 (anterior pair on body ring 16), lateral view. 84–87. Entire right gonopod, mesal, submesal, mesal and subventral views, respectively. Abbreviations: a and b, distofemoral processes; fe, femorite; sl, solenomere; sph, solenophore; z, dorsobasal peg." figureDoi="http://doi.org/10.5281/zenodo.5709568" httpUri="https://zenodo.org/record/5709568/files/figure.png" pageId="26" pageNumber="511">Figs 84–87</figureCitation>
|
|||
|
); cannula a short curved tube, as usual. Prefemoral part short, globose, densely setose as usual, with a particularly long seta distoventrally. Femorite (
|
|||
|
<emphasis id="80C1EADD970FFFCCFEDE2A13B808A328" bold="true" box="[342,364,1370,1396]" pageId="26" pageNumber="511">fe</emphasis>
|
|||
|
) relatively stout and short, somewhat enlarged distad, subclavate, almost as long as acropodite (= postfemoral part), subcylindrical, without evidence of torsion; a small, thumb-shaped, distoventral projection (
|
|||
|
<emphasis id="80C1EADD970FFFCCFA0D2A34BCF7A3CB" bold="true" box="[1413,1427,1405,1431]" pageId="26" pageNumber="511">a</emphasis>
|
|||
|
) basal to acropodite bifurcation into a longer, larger, more slender, subacuminate, ventral, solenophore branch (
|
|||
|
<emphasis id="80C1EADD970FFFCCFAEA2AE8BCE8A3E7" bold="true" box="[1378,1420,1441,1467]" pageId="26" pageNumber="511">sph</emphasis>
|
|||
|
), rather slightly curved dorsad and bearing both a strong spine (
|
|||
|
<emphasis id="80C1EADD970FFFCCFCE92A8DBA15A382" bold="true" box="[865,881,1476,1502]" pageId="26" pageNumber="511">k</emphasis>
|
|||
|
) in distal third and a free solenomere (
|
|||
|
<emphasis id="80C1EADD970FFFCCFABB2A8DBC22A382" bold="true" box="[1331,1350,1476,1502]" pageId="26" pageNumber="511">sl</emphasis>
|
|||
|
), and a shorter, more elaborate, dorsal process divided immediately at base into a longer, stalked, leaf-shaped outgrowth (
|
|||
|
<emphasis id="80C1EADD970FFFCCFA0B2AA1BCF7A05E" bold="true" box="[1411,1427,1512,1538]" pageId="26" pageNumber="511">b</emphasis>
|
|||
|
) with peculiar, ragged, saw-like distal margins, and a much shorter, simple, dorsobasal peg (
|
|||
|
<emphasis id="80C1EADD970FFFCCFB1C2945BDC4A07A" bold="true" box="[1172,1184,1548,1574]" pageId="26" pageNumber="511">z</emphasis>
|
|||
|
). Seminal groove first running all along mesal side of femorite, then turning slowly towards bifurcation point before moving onto a free, flagelliform
|
|||
|
<emphasis id="80C1EADD970FFFCCFEAB291DB852A032" bold="true" box="[291,310,1620,1646]" pageId="26" pageNumber="511">sl</emphasis>
|
|||
|
tightly attached to mesal side of
|
|||
|
<emphasis id="80C1EADD970FFFCCFD2A291DBBA8A032" bold="true" box="[674,716,1620,1646]" pageId="26" pageNumber="511">sph</emphasis>
|
|||
|
, with only a short tip of
|
|||
|
<emphasis id="80C1EADD970FFFCCFC53291DBA8AA032" bold="true" box="[987,1006,1620,1646]" pageId="26" pageNumber="511">sl</emphasis>
|
|||
|
remaining exposed over
|
|||
|
<emphasis id="80C1EADD970FFFCCFA8D291DBC4BA032" bold="true" box="[1285,1327,1620,1646]" pageId="26" pageNumber="511">sph</emphasis>
|
|||
|
. Division of
|
|||
|
<emphasis id="80C1EADD970FFFCCFF3E293EB9A8A0CD" bold="true" box="[182,204,1655,1681]" pageId="26" pageNumber="511">fe</emphasis>
|
|||
|
and acropodite through a transverse sulcus, or of
|
|||
|
<emphasis id="80C1EADD970FFFCCFD722931BA40A0CE" bold="true" box="[762,804,1656,1682]" pageId="26" pageNumber="511">sph</emphasis>
|
|||
|
into laminae, wanting (
|
|||
|
<figureCitation id="2A8E2A4A970FFFCCFBA52931BDD4A0CE" box="[1069,1200,1656,1682]" captionStart="FIGURES 81–87" captionStartId="28.[151,264,1943,1967]" captionTargetBox="[291,1296,181,1919]" captionTargetId="figure-22@28.[291,1296,181,1919]" captionTargetPageId="28" captionText="FIGURES 81–87. Delarthrum anomalans sp. nov., male paratypes. 81. Legs 1, lateral view. 82. Legs 2, lateral view. 83. Leg 26 (anterior pair on body ring 16), lateral view. 84–87. Entire right gonopod, mesal, submesal, mesal and subventral views, respectively. Abbreviations: a and b, distofemoral processes; fe, femorite; sl, solenomere; sph, solenophore; z, dorsobasal peg." figureDoi="http://doi.org/10.5281/zenodo.5709568" httpUri="https://zenodo.org/record/5709568/files/figure.png" pageId="26" pageNumber="511">Figs 84–87</figureCitation>
|
|||
|
).
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
</treatment>
|
|||
|
</document>
|