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<document id="98D3C8920BF954829CB710A249AB7F92" ID-CLB-Dataset="302398" ID-DOI="10.11646/phytotaxa.364.2.8" ID-GBIF-Dataset="1a0e5142-858d-4e9c-8c1c-cbb4f569a3bd" ID-ISSN="1179-3163" ID-Zenodo-Dep="13704134" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725582439378" checkinUser="felipe" docAuthor="Averyanov, Leonid V., Hans. - Jürgen, Nguyen, Khang Sinh &amp; Maisak, Tatiana V." docDate="2018" docId="3439F83CFF95FFF34ACEFC34FD8AD3CE" docLanguage="en" docName="phytotaxa.364.2.8.pdf" docOrigin="Phytotaxa 364 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.364.2.8" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Aspidistra bella Aver., Tillich &amp; K. S. Nguyen 2018, sp. nov." docType="treatment" docVersion="3" lastPageNumber="207" masterDocId="C8008044FF95FFF14A46FFDEFF9BD42C" masterDocTitle="Aspidistra bella (Asparagaceae), a new species from northern Vietnam" masterLastPageNumber="208" masterPageNumber="205" pageNumber="205" updateTime="1725594367310" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="9BB3C49A2977FF8845E75A423D6B7575">
<mods:title id="17BEBC1CD34AE993839AEFA245F880D2">Aspidistra bella (Asparagaceae), a new species from northern Vietnam</mods:title>
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</mods:titleInfo>
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<mods:name id="93A75DC2B3F2A111E97B76E5B760E261" type="personal">
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</mods:role>
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<mods:namePart id="91B473F4D159B859D726239F0B679279">Averyanov, Leonid V.</mods:namePart>
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</mods:name>
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<mods:name id="CE67D4DD7A0CC0A0BC6D5B82B3FD1015" type="personal">
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</mods:role>
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<mods:namePart id="ABCE73DB9A2E7487A7DDF02DEBA0CE1A">Hans. - Jürgen</mods:namePart>
<mods:affiliation id="7B907B27005A08131F6BD94026B0513C">Ludwig-Maximilians-University, Institute of Systematic Botany, Munich, Germany.</mods:affiliation>
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</mods:name>
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<mods:name id="8C267F7FF7554A154CDA28991295D743" type="personal">
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</mods:role>
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<mods:namePart id="8BE49C7A2E82BE46DE91908DA459C94F">Nguyen, Khang Sinh</mods:namePart>
<mods:affiliation id="48C6116D8636EB2D65683535B33C9F10">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, Cau Giay, Ha Noi, Vietnam</mods:affiliation>
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</mods:name>
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<mods:name id="E68667B7A8B5CCDC30A7040A1FE20802" type="personal">
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</mods:role>
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<mods:namePart id="089D27BA0BE6AEF783F548041F6D43FC">Maisak, Tatiana V.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="8A7CBE87A52D9BCA1CC07D40B56020E8">text</mods:typeOfResource>
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<mods:title id="6A151D4C9FD85D81769DD85E24971937">Phytotaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="EC5367279B4485086851D8804C71355D">
<mods:date id="9E4B4A58A8F4F1DBCF873A98C5CE370A">2018</mods:date>
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<mods:number id="B33D8FE307D60AA55E85DEC8604BA32F">2018-08-09</mods:number>
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</mods:detail>
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<mods:detail id="25494EDC294F535ADE9B105ACA42F8F2" type="volume">
<mods:number id="67ABED8F828B75FAE0702ADC48D0003F">364</mods:number>
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</mods:detail>
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<mods:detail id="460ACA0AD59E963CB31CC4877D473243" type="issue">
<mods:number id="0FCC92EC3ACE75738BF81B6DDE2C43A0">2</mods:number>
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</mods:detail>
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<mods:extent id="B52C9E981326A31B50F67E6C0FB5168D" unit="page">
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<mods:end id="9A2EF1959ABB63D3BD3FEBD317B38C1C">208</mods:end>
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</mods:extent>
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<mods:location id="F81E5760B4AB6498138C219796E1B29C">
<mods:url id="079BC4167E37A1E6EE381C18A49900B4">http://dx.doi.org/10.11646/phytotaxa.364.2.8</mods:url>
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</mods:location>
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<mods:classification id="88ED1847301A70182B7E1BD3BEDEBE47">journal article</mods:classification>
<mods:identifier id="D2364DA1F88C720EB2E760AE3754F905" type="CLB-Dataset">302398</mods:identifier>
<mods:identifier id="413DC89D1B1F7FCB9C46A82A500AC493" type="DOI">10.11646/phytotaxa.364.2.8</mods:identifier>
<mods:identifier id="C2450FF59E97EB43F313A843E3D6AED8" type="GBIF-Dataset">1a0e5142-858d-4e9c-8c1c-cbb4f569a3bd</mods:identifier>
<mods:identifier id="F72280FA3AEC211ED37235AD1B6656BF" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="F004A11AA9B2EE3928A72171E483D2F1" type="Zenodo-Dep">13704134</mods:identifier>
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</mods:mods>
<treatment id="3439F83CFF95FFF34ACEFC34FD8AD3CE" LSID="urn:lsid:plazi:treatment:3439F83CFF95FFF34ACEFC34FD8AD3CE" httpUri="http://treatment.plazi.org/id/3439F83CFF95FFF34ACEFC34FD8AD3CE" lastPageId="2" lastPageNumber="207" pageId="0" pageNumber="205">
<subSubSection id="F48A1AA1FF95FFF14ACEFC34FCE1D029" box="[136,890,1002,1029]" pageId="0" pageNumber="205" type="nomenclature">
<paragraph id="BC2F492AFF95FFF14ACEFC34FCE1D029" blockId="0.[136,890,1002,1029]" box="[136,890,1002,1029]" pageId="0" pageNumber="205">
<heading id="E767FE46FF95FFF14ACEFC34FCE1D029" box="[136,890,1002,1029]" fontSize="11" level="2" pageId="0" pageNumber="205" reason="3">
<taxonomicName id="7B9032A9FF95FFF14ACEFC34FD10D029" ID-CoL="8WRF9" authority="Aver., Tillich &amp; K.S. Nguyen" authorityName="Aver., Tillich &amp; K. S. Nguyen" authorityYear="2018" box="[136,651,1002,1029]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="0" pageNumber="205" phylum="Tracheophyta" rank="species" species="bella" status="sp. nov.">
<emphasis id="8EE49538FF95FFF14ACEFC34FEA7D029" bold="true" box="[136,316,1002,1029]" italics="true" pageId="0" pageNumber="205">Aspidistra bella</emphasis>
Aver., Tillich &amp; K.S. Nguyen
</taxonomicName>
<emphasis id="8EE49538FF95FFF148D7FC32FCE1D029" bold="true" box="[657,890,1004,1029]" pageId="0" pageNumber="205">
<taxonomicNameLabel id="95D72843FF95FFF148D7FC32FD70D029" box="[657,747,1004,1029]" pageId="0" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
(
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<figureCitation id="24AB55AFFF95FFF148BCFC32FCA5D029" box="[762,830,1004,1029]" captionStart="FIGURE 1" captionStartId="1.[136,229,1916,1937]" captionTargetBox="[317,1271,457,1887]" captionTargetId="figure-143@1.[317,1271,457,1887]" captionTargetPageId="1" captionText="FIGURE 1. Aspidistra bella Aver., Tillich &amp; K.S. Nguyen corresponding to Averyanov, T. Maisak CPC 7484 / 14369. VIETNAM, Ha Giang province, Bac Me district, plant flowered under cultivation on 10 March 2018. Photos, correction and design by L. Averyanov." figureDoi="http://doi.org/10.5281/zenodo.13704136" httpUri="https://zenodo.org/record/13704136/files/figure.png" pageId="0" pageNumber="205">Fig. 1</figureCitation>
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&amp;
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<figureCitation id="24AB55AFFF95FFF14922FC32FCE8D029" box="[868,883,1004,1029]" captionStart="FIGURE 2" captionStartId="2.[136,229,1420,1441]" captionTargetBox="[215,1372,187,1392]" captionTargetId="figure-17@2.[215,1372,187,1393]" captionTargetPageId="2" captionText="FIGURE 2. Aspidistra bella Aver., Tillich &amp; K.S. Nguyen. Paratype and paratype specimen detail. Averyanov et al., CPC 7484 (LE01042158). Photos, correction and design by L. Averyanov." figureDoi="http://doi.org/10.5281/zenodo.13704138" httpUri="https://zenodo.org/record/13704138/files/figure.png" pageId="0" pageNumber="205">2</figureCitation>
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)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F48A1AA1FF95FFF14ACEFBEBFA3ED108" pageId="0" pageNumber="205" type="materials_examined">
<paragraph id="BC2F492AFF95FFF14ACEFBEBFB8AD05C" blockId="0.[136,1452,1077,2072]" pageId="0" pageNumber="205">
<emphasis id="8EE49538FF95FFF14ACEFBEBFF78D060" bold="true" box="[136,227,1077,1100]" pageId="0" pageNumber="205">Type:—</emphasis>
<collectingCountry id="C48709BAFF95FFF14AA5FBEBFEFAD060" box="[227,353,1077,1100]" name="Vietnam" pageId="0" pageNumber="205">VIETNAM</collectingCountry>
,
<collectingRegion id="7E5487C8FF95FFF14B2AFBEBFDA1D060" box="[364,570,1077,1100]" country="Vietnam" name="Ha Giang" pageId="0" pageNumber="205">Ha Giang province</collectingRegion>
, Bac Me district, ex hort. on
<date id="C82E6FEAFF95FFF14935FBEBFB82D060" box="[883,1049,1077,1100]" pageId="0" pageNumber="205" value="2018-03-10">10 March 2018</date>
,
<emphasis id="8EE49538FF95FFF14E62FBEBFF50D05C" italics="true" pageId="0" pageNumber="205">
<collectionCode id="DA81D1EFFF95FFF14E62FBEBFBADD060" box="[1060,1078,1077,1100]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="0" pageNumber="205" type="Herbarium">L</collectionCode>
. Averyanov,
<collectionCode id="DA81D1EFFF95FFF14EFBFBEBFB56D060" box="[1213,1229,1077,1100]" name="Tavera, Department of Geology and Geophysics" pageId="0" pageNumber="205">T</collectionCode>
. Maisak
<collectionCode id="DA81D1EFFF95FFF14F6BFBEBFAFAD060" box="[1325,1377,1077,1100]" country="Netherlands" httpUri="http://grbio.org/cool/3snx-0r7t" name="Culture collection of Pedro Crous" pageId="0" pageNumber="205">CPC</collectionCode>
7484 / 14369
</emphasis>
(
<typeStatus id="632BF788FF95FFF14A9FFB87FEA7D05C" box="[217,316,1113,1136]" pageId="0" pageNumber="205" type="holotype">holotype</typeStatus>
:
<collectionCode id="DA81D1EFFF95FFF14B00FB87FEFCD05C" box="[326,359,1113,1136]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="0" pageNumber="205">LE</collectionCode>
01042153,
<typeStatus id="632BF788FF95FFF14B98FB87FDA1D05C" box="[478,570,1113,1136]" pageId="0" pageNumber="205" type="isotype">isotypes</typeStatus>
:
<collectionCode id="DA81D1EFFF95FFF14802FB87FDFED05C" box="[580,613,1113,1136]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="0" pageNumber="205">LE</collectionCode>
01042154,
<collectionCode id="DA81D1EFFF95FFF1489AFB87FD66D05C" box="[732,765,1113,1136]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="0" pageNumber="205">LE</collectionCode>
01042155,
<collectionCode id="DA81D1EFFF95FFF14933FB87FC0CD05C" box="[885,919,1113,1136]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="0" pageNumber="205">LE</collectionCode>
01042156).
</paragraph>
<paragraph id="BC2F492AFF95FFF14AFBFBA3FA3ED108" blockId="0.[136,1452,1077,2072]" pageId="0" pageNumber="205">
<emphasis id="8EE49538FF95FFF14AFBFBA3FEC8D0B8" bold="true" box="[189,339,1149,1172]" pageId="0" pageNumber="205">
<typeStatus id="632BF788FF95FFF14AFBFBA3FEA9D0B8" box="[189,306,1149,1172]" pageId="0" pageNumber="205" type="paratype">Paratypes</typeStatus>
:—
</emphasis>
<collectingCountry id="C48709BAFF95FFF14B15FBA3FE4AD0B8" box="[339,465,1149,1172]" name="Vietnam" pageId="0" pageNumber="205">VIETNAM</collectingCountry>
,
<collectingRegion id="7E5487C8FF95FFF14B9DFBA3FD33D0B8" box="[475,680,1149,1172]" country="Vietnam" name="Ha Giang" pageId="0" pageNumber="205">Ha Giang province</collectingRegion>
, Bac Me district, Thuong Tan municipality, primary broad-leaved humid evergreen forest on very steep slopes and along rocky ridge composed with solid crystalline highly eroded limestone at elevation
<quantity id="7B68E4CFFF95FFF14AB7FB1BFE1ED0F0" box="[241,389,1221,1244]" metricMagnitude="3" metricUnit="m" metricValue="1.2425" metricValueMax="1.285" metricValueMin="1.2" pageId="0" pageNumber="205" unit="m" value="1242.5" valueMax="1285.0" valueMin="1200.0">1200–1285 m</quantity>
around point
<geoCoordinate id="D9A42FEDFF95FFF1485AFB1BFD2DD0F0" box="[540,694,1221,1244]" degrees="22" direction="north" minutes="38" orientation="latitude" pageId="0" pageNumber="205" precision="1" seconds="19.0" value="22.63861">22°38’19.0’’N</geoCoordinate>
<geoCoordinate id="D9A42FEDFF95FFF148FBFB1BFCFED0F0" box="[701,869,1221,1244]" degrees="105" direction="east" minutes="17" orientation="longitude" pageId="0" pageNumber="205" precision="1" seconds="06.5" value="105.28514">105°17’06.5’’E</geoCoordinate>
, terrestrial herb on rocky slope, leaves uniformly dark green, fruits globular, about
<quantity id="7B68E4CFFF95FFF14B85FB37FE60D12C" box="[451,507,1257,1280]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="0" pageNumber="205" unit="cm" value="1.0">1 cm</quantity>
in diam., densely haired with soft spine-like hairs, stalk
<quantity id="7B68E4CFFF95FFF14E32FB37FB41D12C" box="[1140,1242,1257,1280]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="0" pageNumber="205" unit="cm" value="3.25" valueMax="4.0" valueMin="2.5">2.5–4 cm</quantity>
long, common,
<date id="C82E6FEAFF95FFF14FD7FB37FEACD108" pageId="0" pageNumber="205" value="2014-11-16">16 November 2014</date>
,
<emphasis id="8EE49538FF95FFF14B04FAD3FBC1D108" box="[322,1114,1293,1316]" italics="true" pageId="0" pageNumber="205">
<collectionCode id="DA81D1EFFF95FFF14B04FAD3FECFD108" box="[322,340,1293,1316]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="0" pageNumber="205" type="Herbarium">L</collectionCode>
. Averyanov,
<collectionCode id="DA81D1EFFF95FFF14B9CFAD3FE74D108" box="[474,495,1293,1316]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="0" pageNumber="205" type="Herbarium">N</collectionCode>
.
<collectionCode id="DA81D1EFFF95FFF14BBFFAD3FD92D108" box="[505,521,1293,1316]" name="Tavera, Department of Geology and Geophysics" pageId="0" pageNumber="205">T</collectionCode>
. Hiep,
<collectionCode id="DA81D1EFFF95FFF14812FAD3FDF2D108" box="[596,617,1293,1316]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="0" pageNumber="205" type="Herbarium">N</collectionCode>
.
<collectionCode id="DA81D1EFFF95FFF14835FAD3FD18D108" box="[627,643,1293,1316]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="0" pageNumber="205" type="Herbarium">S</collectionCode>
. Khang,
<collectionCode id="DA81D1EFFF95FFF148A4FAD3FD69D108" box="[738,754,1293,1316]" name="Tavera, Department of Geology and Geophysics" pageId="0" pageNumber="205">T</collectionCode>
. Maisak,
<collectionCode id="DA81D1EFFF95FFF1491EFAD3FCF1D108" box="[856,874,1293,1316]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="0" pageNumber="205" type="Herbarium">L</collectionCode>
. Osinovetz,
<collectionCode id="DA81D1EFFF95FFF149ADFAD3FB84D108" box="[1003,1055,1293,1316]" country="Netherlands" httpUri="http://grbio.org/cool/3snx-0r7t" name="Culture collection of Pedro Crous" pageId="0" pageNumber="205">CPC</collectionCode>
7484
</emphasis>
(
<collectionCode id="DA81D1EFFF95FFF14E37FAD3FB09D108" box="[1137,1170,1293,1316]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="0" pageNumber="205">LE</collectionCode>
01042157,
<collectionCode id="DA81D1EFFF95FFF14F4CFAD3FAB7D108" box="[1290,1324,1293,1316]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="0" pageNumber="205">LE</collectionCode>
01042158).
</paragraph>
</subSubSection>
<subSubSection id="F48A1AA1FF95FFF14AFBFAEFFC11D1BC" pageId="0" pageNumber="205" type="diagnosis">
<paragraph id="BC2F492AFF95FFF14AFBFAEFFC11D1BC" blockId="0.[136,1452,1077,2072]" pageId="0" pageNumber="205">
<emphasis id="8EE49538FF95FFF14AFBFAEFFED5D164" bold="true" box="[189,334,1329,1352]" pageId="0" pageNumber="205">Diagnosis:—</emphasis>
<taxonomicName id="7B9032A9FF95FFF14B09FAEFFE67D164" box="[335,508,1329,1352]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="0" pageNumber="205" phylum="Tracheophyta" rank="species" species="bella">
<emphasis id="8EE49538FF95FFF14B09FAEFFE67D164" box="[335,508,1329,1352]" italics="true" pageId="0" pageNumber="205">Aspidistra bella</emphasis>
</taxonomicName>
differs from closest
<taxonomicName id="7B9032A9FF95FFF148AEFAEFFBFBD164" authority="Chen (1982: 77)" authorityName="Chen" authorityPageNumber="77" authorityYear="1982" box="[744,1120,1329,1352]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="0" pageNumber="205" phylum="Tracheophyta" rank="species" species="dolichanthera">
<emphasis id="8EE49538FF95FFF148AEFAEFFC04D164" box="[744,927,1329,1352]" italics="true" pageId="0" pageNumber="205">A. dolichanthera</emphasis>
<bibRefCitation id="D80134DBFF95FFF149ECFAEFFBFBD164" author="Chen, X. - X. &amp; Fang, D." box="[938,1120,1329,1352]" pageId="0" pageNumber="205" pagination="77 - 79" refId="ref2245" refString="Chen, X. - X. &amp; Fang, D. (1982) Two new species of the Aspidistra (Liliaceae) from Guangxi. Guihaia 2: 77 - 79." type="journal article" year="1982">Chen (1982: 77)</bibRefCitation>
</taxonomicName>
by the creeping rhizome; the almost naked peduncle
<quantity id="7B68E4CFFF95FFF14BC6FA8BFE6CD140" box="[384,503,1365,1388]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="3.5" metricValueMin="1.4" pageId="0" pageNumber="205" unit="cm" value="2.45" valueMax="3.5" valueMin="1.4">1.4–3.5 cm</quantity>
long and curved downward; the numerous flowers
<quantity id="7B68E4CFFF95FFF14E53FA8BFB0FD140" box="[1045,1172,1365,1388]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.5" metricValueMin="5.5" pageId="0" pageNumber="205" unit="mm" value="6.5" valueMax="7.5" valueMin="5.5">5.5–7.5 mm</quantity>
in diameter, the triangular perigone lobes
<quantity id="7B68E4CFFF95FFF14B6DFAA7FE30D1BC" box="[299,427,1401,1424]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.5" metricValueMin="4.5" pageId="0" pageNumber="205" unit="mm" value="5.5" valueMax="6.5" valueMin="4.5">4.5–6.5 mm</quantity>
long and the short anthers
<quantity id="7B68E4CFFF95FFF1488AFAA7FCD7D1BC" box="[716,844,1401,1424]" metricMagnitude="-3" metricUnit="m" metricValue="4.199999999999999" metricValueMax="4.6" metricValueMin="3.8" pageId="0" pageNumber="205" unit="mm" value="4.199999999999999" valueMax="4.6" valueMin="3.8">3.8–4.6 mm</quantity>
long.
</paragraph>
</subSubSection>
<subSubSection id="F48A1AA1FF95FFF14AFBFA43FF4FD1F4" pageId="0" pageNumber="205" type="etymology">
<paragraph id="BC2F492AFF95FFF14AFBFA43FF4FD1F4" blockId="0.[136,1452,1077,2072]" pageId="0" pageNumber="205">
<emphasis id="8EE49538FF95FFF14AFBFA43FEC7D198" bold="true" box="[189,348,1437,1460]" pageId="0" pageNumber="205">Etymology:—</emphasis>
The species name refers to the attractive beautiful white flowers, clustering at the apex of individual shoots.
</paragraph>
</subSubSection>
<subSubSection id="F48A1AA1FF95FFF04AFBFA3BFC6ED59F" lastPageId="1" lastPageNumber="206" pageId="0" pageNumber="205" type="description">
<paragraph id="BC2F492AFF95FFF04AFBFA3BFC6ED59F" blockId="0.[136,1452,1077,2072]" lastBlockId="1.[136,1452,160,435]" lastPageId="1" lastPageNumber="206" pageId="0" pageNumber="205">
<emphasis id="8EE49538FF95FFF14AFBFA3BFEFFD1D0" bold="true" box="[189,356,1509,1532]" pageId="0" pageNumber="205">Description:—</emphasis>
Terrestrial rhizomatous herb. Rhizome plagiotropic, creeping, terete, branching, (0.8)1–1.4(1.6) cm in diameter, densely nodal, distal part ascending to erect, terete, (3)4–6(7) cm long, (1)1.2–1.8(2) cm in diameter, simple, densely nodal, grassy green, with many cataphylls and 2–5(6) leaves; basal part with thick, distant, rigid, light grey, semiwoody, straight spreading roots distally richly branching. Cataphylls convolute, cuneate, acute, dark purple-brown and tubular when young, later light dull yellowish, almost flat, papyraceous, (1)2–8(10) cm long and (4)5–8(10) mm wide, splitting into irregular fibrous–papyraceous remains with age. Leaves petiolate, (0.5)0.6–0.9(1.2) m long. Petiole stiff, erect, straight, or slightly curved, (25)30–40(50) cm long. Leaf blade upright to somewhat arching, elliptic, attenuate at base and apex, (25)30– 45(50) cm long, (6)8–16(18) cm wide, plicate, uniformly grassy green above and below, with prominent midvein and several secondary veins well seen on both sides, arising from midvein in basal half of leaf blade. Flowers odorless, numerous, arising by (1)2–3(5) from each axil, nodding, pure milky-white, widely open, (1.3)1.5–1.8(2) cm in diameter. Peduncles appearing in groups, successively, one by one, dirty purple, finely streaked with white, (1.4)1.8–3(3.5) cm long, (1.6)1.8–2(2.2) mm in diameter, curved downward, with 4–6 sterile bracts. Peduncle bracts broadly ovate to reniform, concave, scarious, pure white or slightly tinged with violet, roundish at apex, (5)6–7(7.5) mm long and (6)7–11(12) mm wide (being flattened), upper three approximate near flower base. Perigone tube campanulate, (4)4.5–5.5(6) mm long, (5.5)6–7(7.5) mm wide, glossy. Perigone lobes (4)6(7), subequal, triangular broadly sagittate, (4.5)5–6(6.5) mm long, (4.5)5–8(8.5) mm wide, slightly convex, almost round at apex, fleshy, glossy, smooth, with irregularly recurved and often incised margin, at broadest part auriculate, narrowing adaxially into rather narrow rectangular base. Stamens (5)6(7), anthers brightly yellow, sessile, erect (parallel to style), arising near the base of the tube, narrowly ovoid, (3.8)4–4.4(4.6) mm long, (1)1.2–1.4(1.6) mm wide, with narrow connective, pollen sacs facing the style; pollen brightly yellow. Pistil cylindric, erect, (5.6)5.8–6.2(6.4) mm tall, slightly broadening at apex and base; ovary inconspicuous, white to white tinged with violet, (1)1.2–1.4(1.6) mm long and wide; style white, cylindrical, (4)4.2–4.4(4.6) mm long, (0.8)0.9–1(1.2) mm in diameter; stigma truncate, almost flat, pure white or white with very light indistinct violet irregular spots, subcircular to rounded triangular, sometimes with inconspicuous irregular lobes, (1)1.2–1.4(1.6) mm across, its upper surface smooth. Fruits greenish, globular, (0.7)0.8–1(1.1) cm long and wide, densely hairy with long soft spine like papillae, stalk (2.5)3–4(4.5) cm long.
</paragraph>
</subSubSection>
Add updates up until 2024-09-06 03:56:04
2024-09-06 05:58:16 +02:00
<caption id="E8EF19A2FF94FFF04ACEF8A2FA13D399" ID-DOI="http://doi.org/10.5281/zenodo.13704136" ID-Zenodo-Dep="13704136" httpUri="https://zenodo.org/record/13704136/files/figure.png" pageId="1" pageNumber="206" startId="1.[136,229,1916,1937]" targetBox="[317,1271,457,1887]" targetPageId="1" targetType="figure">
Add updates up until 2024-09-06 03:34:29
2024-09-06 05:36:38 +02:00
<paragraph id="BC2F492AFF94FFF04ACEF8A2FA13D399" blockId="1.[136,1451,1916,1973]" pageId="1" pageNumber="206">
<emphasis id="8EE49538FF94FFF04ACEF8A2FF64D3BD" bold="true" box="[136,255,1916,1937]" pageId="1" pageNumber="206">FIGURE 1.</emphasis>
<taxonomicName id="7B9032A9FF94FFF04B40F8A2FD5ED3BD" authority="Aver., Tillich &amp; K. S. Nguyen" authorityName="Aver., Tillich &amp; K. S. Nguyen" box="[262,709,1916,1937]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="206" phylum="Tracheophyta" rank="species" species="bella">
<emphasis id="8EE49538FF94FFF04B40F8A2FE3BD3BD" box="[262,416,1916,1937]" italics="true" pageId="1" pageNumber="206">Aspidistra bella</emphasis>
Aver., Tillich &amp; K.S. Nguyen
</taxonomicName>
corresponding to
<emphasis id="8EE49538FF94FFF04931F8A2FA96D3BD" box="[887,1293,1916,1937]" italics="true" pageId="1" pageNumber="206">Averyanov, T. Maisak CPC 7484 / 14369.</emphasis>
VIETNAM, Ha Giang province, Bac Me district, plant flowered under cultivation on 10 March 2018. Photos, correction and design by L. Averyanov.
</paragraph>
</caption>
Add updates up until 2024-09-06 03:56:04
2024-09-06 05:58:16 +02:00
<caption id="E8EF19A2FF97FFF34ACEFA52FD72D1E9" ID-DOI="http://doi.org/10.5281/zenodo.13704138" ID-Zenodo-Dep="13704138" httpUri="https://zenodo.org/record/13704138/files/figure.png" pageId="2" pageNumber="207" startId="2.[136,229,1420,1441]" targetBox="[215,1372,187,1392]" targetPageId="2" targetType="figure">
Add updates up until 2024-09-06 03:34:29
2024-09-06 05:36:38 +02:00
<paragraph id="BC2F492AFF97FFF34ACEFA52FD72D1E9" blockId="2.[136,1451,1420,1477]" pageId="2" pageNumber="207">
<emphasis id="8EE49538FF97FFF34ACEFA52FE98D18D" bold="true" box="[136,259,1420,1441]" pageId="2" pageNumber="207">FIGURE 2.</emphasis>
<taxonomicName id="7B9032A9FF97FFF34B4BFA52FD72D18D" authority="Aver., Tillich &amp; K. S. Nguyen." authorityName="Aver., Tillich &amp; K. S. Nguyen." box="[269,745,1420,1441]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="207" phylum="Tracheophyta" rank="species" species="bella">
<emphasis id="8EE49538FF97FFF34B4BFA52FE30D18D" box="[269,427,1420,1441]" italics="true" pageId="2" pageNumber="207">Aspidistra bella</emphasis>
Aver., Tillich &amp; K.S. Nguyen.
</taxonomicName>
Paratype and paratype specimen detail. Averyanov
<emphasis id="8EE49538FF97FFF34EBAFA52FAAAD18D" box="[1276,1329,1420,1441]" italics="true" pageId="2" pageNumber="207">et al.</emphasis>
,
<emphasis id="8EE49538FF97FFF34F04FA52FA30D18D" box="[1346,1451,1420,1441]" italics="true" pageId="2" pageNumber="207">CPC 7484</emphasis>
(LE01042158). Photos, correction and design by L. Averyanov.
</paragraph>
</caption>
<subSubSection id="F48A1AA1FF97FFF34AFBFA29FBADD222" box="[189,1078,1527,1550]" pageId="2" pageNumber="207" type="distribution">
<paragraph id="BC2F492AFF97FFF34AFBFA29FBADD222" blockId="2.[136,1452,1527,2018]" box="[189,1078,1527,1550]" pageId="2" pageNumber="207">
<emphasis id="8EE49538FF97FFF34AFBFA29FEF7D222" bold="true" box="[189,364,1527,1550]" pageId="2" pageNumber="207">Distribution:—</emphasis>
Northern
<collectingCountry id="C48709BAFF97FFF34B95FA29FDB6D222" box="[467,557,1527,1550]" name="Vietnam" pageId="2" pageNumber="207">Vietnam</collectingCountry>
(
<collectingRegion id="7E5487C8FF97FFF3487DFA29FC91D222" box="[571,778,1527,1550]" country="Vietnam" name="Ha Giang" pageId="2" pageNumber="207">Ha Giang province</collectingRegion>
, Bac Me district). Endemic.
</paragraph>
</subSubSection>
<subSubSection id="F48A1AA1FF97FFF34AFBF9C5FD8AD3CE" pageId="2" pageNumber="207" type="biology_ecology">
<paragraph id="BC2F492AFF97FFF34AFBF9C5FCCBD256" blockId="2.[136,1452,1527,2018]" pageId="2" pageNumber="207">
<emphasis id="8EE49538FF97FFF34AFBF9C5FEA0D21E" bold="true" box="[189,315,1563,1586]" pageId="2" pageNumber="207">Ecology:—</emphasis>
Very steep rocky slopes and mountain summits covered by primary broad-leaved evergreen humid forests on marble-like, highly eroded limestone at elevations of
<quantity id="7B68E4CFFF97FFF348AAF9E1FC1BD27A" box="[748,896,1599,1622]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="2" pageNumber="207" unit="m" value="1250.0" valueMax="1300.0" valueMin="1200.0">1200–1300 m</quantity>
a.s.l. Flowering in cultivation in March–April. The species is considered to be of “Least Concern” (LC) (IUCN 2012).
</paragraph>
<paragraph id="BC2F492AFF97FFF34AFBF959FD8AD3CE" blockId="2.[136,1452,1527,2018]" pageId="2" pageNumber="207">
<emphasis id="8EE49538FF97FFF34AFBF959FE6CD2B2" bold="true" box="[189,503,1671,1694]" pageId="2" pageNumber="207">Taxonomic relationships:—</emphasis>
<taxonomicName id="7B9032A9FF97FFF34BBEF959FD39D2B2" box="[504,674,1671,1694]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="207" phylum="Tracheophyta" rank="species" species="bella">
<emphasis id="8EE49538FF97FFF34BBEF959FD39D2B2" box="[504,674,1671,1694]" italics="true" pageId="2" pageNumber="207">Aspidistra bella</emphasis>
</taxonomicName>
belongs to a group of white or light yellowish-flowering species, which includes in
<collectingCountry id="C48709BAFF97FFF34B46F975FEC1D2EE" box="[256,346,1707,1730]" name="Vietnam" pageId="2" pageNumber="207">Vietnam</collectingCountry>
and allied countries species such as
<taxonomicName id="7B9032A9FF97FFF34891F975FA93D2EE" authority="Averyanov &amp; Tillich (2016: 141)" authorityName="Averyanov &amp; Tillich" authorityPageNumber="141" authorityYear="2016" box="[727,1288,1707,1730]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="207" phylum="Tracheophyta" rank="species" species="anomala">
<emphasis id="8EE49538FF97FFF34891F975FC3CD2EE" box="[727,935,1707,1730]" italics="true" pageId="2" pageNumber="207">Aspidistra anomala</emphasis>
Averyanov &amp; Tillich (2016: 141)
</taxonomicName>
,
<taxonomicName id="7B9032A9FF97FFF34F55F975FE45D2CA" authority="Averyanov &amp; Tillich (2008: 37)" authorityName="Averyanov &amp; Tillich" authorityPageNumber="37" authorityYear="2008" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="207" phylum="Tracheophyta" rank="species" species="brachystyla">
<emphasis id="8EE49538FF97FFF34F55F975FA30D2EE" box="[1299,1451,1707,1730]" italics="true" pageId="2" pageNumber="207">A. brachystyla</emphasis>
Averyanov &amp;
<bibRefCitation id="D80134DBFF97FFF34B59F911FE45D2CA" author="Tillich, H. - J. &amp; Averyanov, L. V." box="[287,478,1743,1766]" pageId="2" pageNumber="207" pagination="37 - 41" refId="ref2531" refString="Tillich, H. - J. &amp; Averyanov, L. V. (2008) Two new species and one new subspecies of Aspidistra Ker-Gawl. (Ruscaceae) from Vietnam. Feddes Repertorium 119 (1 - 2): 37 - 41. https: // doi. org / 10.1002 / fedr. 200711143" type="journal article" year="2008">Tillich (2008: 37)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="7B9032A9FF97FFF34BACF911FCFFD2CA" authority="Tillich (2007: 337)" authorityName="Tillich" authorityPageNumber="337" authorityYear="2007" box="[490,868,1743,1766]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="207" phylum="Tracheophyta" rank="species" species="campanulata">
<emphasis id="8EE49538FF97FFF34BACF911FD09D2CA" box="[490,658,1743,1766]" italics="true" pageId="2" pageNumber="207">A. campanulata</emphasis>
<bibRefCitation id="D80134DBFF97FFF348DFF911FCFFD2CA" author="Tillich, H. - J. &amp; Averyanov, L. V. &amp; Dzu, N. V. L." box="[665,868,1743,1766]" pageId="2" pageNumber="207" pagination="335 - 344" refId="ref2590" refString="Tillich, H. - J., Averyanov, L. V. &amp; Dzu, N. V. L. (2007) Six new species of Aspidistra (Ruscaceae) from Northern Vietnam. Blumea 52: 335 - 344. https: // doi. org / 10.3767 / 000651907 X 609070" type="journal article" year="2007">Tillich (2007: 337)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="7B9032A9FF97FFF349DEF911FBD7D2CA" authorityName="Chen" authorityYear="1982" box="[920,1100,1743,1766]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="207" phylum="Tracheophyta" rank="species" species="dolichanthera">
<emphasis id="8EE49538FF97FFF349DEF911FBD7D2CA" box="[920,1100,1743,1766]" italics="true" pageId="2" pageNumber="207">A. dolichanthera</emphasis>
</taxonomicName>
. Among the mentioned taxa, our species is most similar to
<taxonomicName id="7B9032A9FF97FFF34BDBF92DFDCAD326" authorityName="Chen" authorityYear="1982" box="[413,593,1779,1802]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="207" phylum="Tracheophyta" rank="species" species="dolichanthera">
<emphasis id="8EE49538FF97FFF34BDBF92DFDCAD326" box="[413,593,1779,1802]" italics="true" pageId="2" pageNumber="207">A. dolichanthera</emphasis>
</taxonomicName>
, described in detail by
<bibRefCitation id="D80134DBFF97FFF3490DF92DFB8BD326" author="Lang, K. - Y. &amp; Li, G. - Z. &amp; Wei, Y. - G. &amp; Wang, R. - X." box="[843,1040,1779,1802]" pageId="2" pageNumber="207" pagination="468 - 508" refId="ref2279" refString="Lang, K. - Y., Li, G. - Z., Wei, Y. - G. &amp; Wang, R. - X. (1999) Taxonomic and phytogeographic studies on the genus Aspidistra Ker-Gawl. (Liliaceae) in China. Acta Phytotaxonomica Sinica 37: 468 - 508." type="journal article" year="1999">
Lang
<emphasis id="8EE49538FF97FFF349CFF92DFC24D326" box="[905,959,1779,1802]" italics="true" pageId="2" pageNumber="207">et al.</emphasis>
(1999)
</bibRefCitation>
. From
<taxonomicName id="7B9032A9FF97FFF34E1AF92DFA8BD326" authorityName="Chen" authorityYear="1982" box="[1116,1296,1779,1802]" class="Liliopsida" family="Asparagaceae" genus="Aspidistra" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="207" phylum="Tracheophyta" rank="species" species="dolichanthera">
<emphasis id="8EE49538FF97FFF34E1AF92DFA8BD326" box="[1116,1296,1779,1802]" italics="true" pageId="2" pageNumber="207">A. dolichanthera</emphasis>
</taxonomicName>
our plant well differs in long creeping rhizome (vs. short ascending rhizome), descending, curved downward, purplish peduncles
<quantity id="7B68E4CFFF97FFF34F1DF8C9FF32D37E" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="3.5" metricValueMin="1.4" pageId="2" pageNumber="207" unit="cm" value="2.45" valueMax="3.5" valueMin="1.4">1.4–3.5 cm</quantity>
long, with only a few white, distant broadly ovate bracts
<quantity id="7B68E4CFFF97FFF34946F8E5FCF0D37E" box="[768,875,1851,1874]" metricMagnitude="-3" metricUnit="m" metricValue="6.25" metricValueMax="7.5" metricValueMin="5.0" pageId="2" pageNumber="207" unit="mm" value="6.25" valueMax="7.5" valueMin="5.0">5–7.5 mm</quantity>
long, roundish at apex, leaving peduncle almost naked (vs. stiff erect, white peduncles
<quantity id="7B68E4CFFF97FFF34B93F881FDA4D35A" box="[469,575,1887,1910]" metricMagnitude="-1" metricUnit="m" metricValue="1.45" metricValueMax="1.7" metricValueMin="1.2" pageId="2" pageNumber="207" unit="cm" value="14.5" valueMax="17.0" valueMin="12.0">12–17 cm</quantity>
long, nearly completely covered by acute-lanceolate, brown bracts
<quantity id="7B68E4CFFF97FFF34F43F881FAF3D35A" box="[1285,1384,1887,1910]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="2" pageNumber="207" unit="cm" value="3.25" valueMax="3.5" valueMin="3.0">3–3.5 cm</quantity>
long), perigone tubular campanulate,
<quantity id="7B68E4CFFF97FFF34B96F85DFDBCD3B6" box="[464,551,1923,1946]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="2" pageNumber="207" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
long,
<quantity id="7B68E4CFFF97FFF3482CF85DFD73D3B6" box="[618,744,1923,1946]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.5" metricValueMin="5.5" pageId="2" pageNumber="207" unit="mm" value="6.5" valueMax="7.5" valueMin="5.5">5.5–7.5 mm</quantity>
wide (vs. perigone tube widely campanulate to funnel shaped,
<quantity id="7B68E4CFFF97FFF34FC2F85DFF2AD392" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="2" pageNumber="207" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">6–8 mm</quantity>
long,
<quantity id="7B68E4CFFF97FFF34AB1F879FEAED392" box="[247,309,1959,1982]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="207" unit="mm" value="8.0">8 mm</quantity>
wide), perigone lobes triangular,
<quantity id="7B68E4CFFF97FFF348E7F879FCB9D392" box="[673,802,1959,1982]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.5" metricValueMin="4.5" pageId="2" pageNumber="207" unit="mm" value="5.5" valueMax="6.5" valueMin="4.5">4.5–6.5 mm</quantity>
long (vs. perigone lobes elongate,
<quantity id="7B68E4CFFF97FFF34EDBF879FA8BD392" box="[1181,1296,1959,1982]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="2" pageNumber="207" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10–12 mm</quantity>
long), anthers
<quantity id="7B68E4CFFF97FFF34ACEF815FE93D3CE" box="[136,264,1995,2018]" metricMagnitude="-3" metricUnit="m" metricValue="4.199999999999999" metricValueMax="4.6" metricValueMin="3.8" pageId="2" pageNumber="207" unit="mm" value="4.199999999999999" valueMax="4.6" valueMin="3.8">3.8–4.6 mm</quantity>
long (vs.
<quantity id="7B68E4CFFF97FFF34B35F815FE50D3CE" box="[371,459,1995,2018]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="2" pageNumber="207" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
long).
</paragraph>
</subSubSection>
</treatment>
</document>
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