285 lines
31 KiB
XML
285 lines
31 KiB
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<document id="1BBE46AE975A9F5382EA3AFFE0200919" ID-CLB-Dataset="31827" ID-DOI="10.11646/zootaxa.4337.1.9" ID-GBIF-Dataset="2ee3c954-03da-4e1b-8fde-66de4737b120" ID-ISSN="1175-5326" ID-Zenodo-Dep="1014115" ID-ZooBank="E2606CA3-079D-4D8E-81A2-7E3489E47F37" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1508235764385" checkinUser="plazi" docAuthor="Steiner, Helmut" docDate="2017" docId="DB4D8789FFA2914F57B026A8FD2E47F2" docLanguage="en" docName="zootaxa.4337.1.9.pdf" docOrigin="Zootaxa 4337 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mimogonellus dreybrodti Steiner 2017, new species" docType="treatment" docVersion="7" masterDocId="2774FFF1FFA3914C57272612FF964033" masterDocTitle="Mimogonellus dreybrodti sp. n., a new cave-inhabiting Osoriinae from Laos (Coleoptera: Staphylinidae)" updateTime="1698469669963" updateUser="ExternalLinkService">
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<mods:mods id="366A479670B6A8F6B4C7D27EE1D6D45A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="3FCDDE45A4B685582DA18311A9A81A73">
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<mods:title id="B86B731DDDDA5C2C3EDDA3175A729FE6">Mimogonellus dreybrodti sp. n., a new cave-inhabiting Osoriinae from Laos (Coleoptera: Staphylinidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="D8F8D3BF1847EA758CA9A35FECEF35A6" type="personal">
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<mods:roleTerm id="19393006DF30355A9E67E761849DC1CD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="158723EDB4D501C51CC033DA9305476D">Steiner, Helmut</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="1B7F5869B4576825B40724EA1A629A3B">text</mods:typeOfResource>
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<mods:relatedItem id="B4B1683930348966BB8DAF97E4B93C7E" type="host">
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<mods:titleInfo id="5CF9BD76AA5E4A3CCDAE5E31DC800B0E">
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<mods:title id="6EDEDF0365FB6C5D47C04C173AD29320">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="576130866843F9F7A3EC41D7EC3C5EDC">
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<mods:date id="2E6E7F35D609EFBE2B65666A2FCA50A8">2017</mods:date>
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<mods:detail id="854BF4FE099F83273774EDDA862D6A6F" type="pubDate">
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<mods:number id="38D8F2D7B327BBB3B83448A221210B55">2017-10-17</mods:number>
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</mods:detail>
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<mods:detail id="5977E240D38D5762001B9A7F2578C396" type="volume">
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<mods:number id="C6DB453D68A31B9BE38DAA993C635B8C">4337</mods:number>
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</mods:detail>
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<mods:detail id="F22B2B572F35C5361419122E22E6900E" type="issue">
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<mods:number id="5535B96CF607C3783F392A26209353AC">1</mods:number>
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</mods:detail>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="918EA1549AFCCB1CE08A1B4490E91979">journal volume</mods:classification>
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<mods:identifier id="D6E93DAEF1FE17BDE7F67FCAF503CB69" type="CLB-Dataset">31827</mods:identifier>
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<mods:identifier id="24068558834FE34F4C3A38D95578936C" type="DOI">10.11646/zootaxa.4337.1.9</mods:identifier>
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<mods:identifier id="31F7315D4556346462BD74CF16828BCD" type="GBIF-Dataset">2ee3c954-03da-4e1b-8fde-66de4737b120</mods:identifier>
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<mods:identifier id="5D9E3F623421A5705DAF6779C2DEC091" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="AFA971E1EF28305CA1D2DFDABBF9A8BF" type="Zenodo-Dep">1014115</mods:identifier>
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<mods:identifier id="376C8EBC54025AFCBD7CBAD2305D63E0" type="ZooBank">E2606CA3-079D-4D8E-81A2-7E3489E47F37</mods:identifier>
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</mods:mods>
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<treatment id="DB4D8789FFA2914F57B026A8FD2E47F2" ID-DOI="http://doi.org/10.5281/zenodo.6026605" ID-GBIF-Taxon="135450166" ID-Zenodo-Dep="6026605" LSID="urn:lsid:plazi:treatment:DB4D8789FFA2914F57B026A8FD2E47F2" httpUri="http://treatment.plazi.org/id/DB4D8789FFA2914F57B026A8FD2E47F2" lastPageId="3" pageId="1">
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<subSubSection id="1BFE6514FFA2914D57B026A8FD7F40E7" box="[151,745,186,212]" pageId="1" type="nomenclature">
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<paragraph id="535B369FFFA2914D57B026A8FD7F40E7" blockId="1.[151,1436,152,729]" box="[151,745,186,212]" pageId="1">
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<heading id="081381F3FFA2914D57B026A8FD7F40E7" bold="true" box="[151,745,186,212]" fontSize="11" level="1" pageId="1" reason="1">
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<emphasis id="6190EA8DFFA2914D57B026A8FD7F40E7" bold="true" box="[151,745,186,212]" pageId="1">
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<taxonomicName id="94E44D1CFFA2914D57B026A8FE5140E7" ID-CoL="43FKD" authority="Steiner, 2017" authorityName="Steiner" authorityYear="2017" box="[151,455,186,212]" class="Insecta" family="Staphylinidae" genus="Mimogonellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="dreybrodti" status="sp. nov.">
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<emphasis id="6190EA8DFFA2914D57B026A8FE5140E7" bold="true" box="[151,455,186,212]" italics="true" pageId="1">Mimogonellus dreybrodti</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="7AA357F6FFA2914D56F126A8FDF040E7" box="[470,614,186,212]" pageId="1" rank="species">new species</taxonomicNameLabel>
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(
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<figureCitation id="CBDF2A1AFFA2914D555126A8FD4940E7" box="[630,735,186,212]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="1.[151,244,2002,2024]" captionStartId-1="2.[151,244,1980,2002]" captionTargetBox-0="[220,1385,801,1980]" captionTargetBox-1="[182,1417,697,1963]" captionTargetId-0="figure@1.[210,1385,786,1987]" captionTargetId-1="figure@2.[173,1417,677,1971]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURE 1. Mimogonellus dreybrodti, male. A. Dorsal habitus. B. Forebody. C. Left antenna. Scale bars: 2 mm in A; 0.5 mm in B – C." captionText-1="FIGURE 2. Mimogonellus dreybrodti, male. A. Protibia. B. Mesotibia. C. Tergites IX – X. D. Sternite VIII. E. Sternite IX. F – G. Aedeagus, in lateral (F), and dorsal (G) views. Scale bars: 0.3 mm in A – D; 0.2 mm in E – G." httpUri-0="https://zenodo.org/record/1014117/files/figure.png" httpUri-1="https://zenodo.org/record/1014119/files/figure.png" pageId="1">Figs 1–2</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="1BFE6514FFA2914D57B026CFFBDB410A" pageId="1" type="materials_examined">
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<paragraph id="535B369FFFA2914D57B026CFFBDB410A" blockId="1.[151,1436,152,729]" pageId="1">
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<materialsCitation id="E38C3CC2FFA2914D57B026CFFD664124" ID-GBIF-Occurrence="1675163016" collectingDate="2009-01-13" collectorName="Tham Nam Lot, S & Vieng Thong, H & Prov." country="Laos" latitude="20.248945" location="Ban Boumfart Village" longLatPrecision="14" longitude="103.396385" pageId="1" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<emphasis id="6190EA8DFFA2914D57B026CFFD8A40C7" bold="true" box="[151,540,221,244]" pageId="1">
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<typeStatus id="8C5F883DFFA2914D57B026CFFF5940C7" box="[151,207,221,244]" pageId="1">Type</typeStatus>
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material.
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<typeStatus id="8C5F883DFFA2914D566A26CFFE2F40C7" box="[333,441,221,244]" pageId="1" type="holotype">Holotype</typeStatus>
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:
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<collectingCountry id="2BF3760FFFA2914D56ED26CFFD8140C7" box="[458,535,221,244]" name="Laos" pageId="1">LAOS</collectingCountry>
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:
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</emphasis>
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<specimenCount id="45E2FD16FFA2914D550026CFFDA840C7" box="[551,574,221,244]" pageId="1" type="male">♂</specimenCount>
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: ‘111/09-,
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<collectingDate id="371EE9B7FFA2914D559C26CFFCA040C7" box="[699,822,221,244]" pageId="1" value="2009-01-13">
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<date id="275A105FFFA2914D559C26CFFCA040C7" box="[699,822,221,244]" pageId="1" value="2009-01-13">13.01.2009</date>
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</collectingDate>
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,
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<geoCoordinate id="36D05058FFA2914D546126CFFC4C40C7" box="[838,986,221,244]" direction="north" orientation="latitude" pageId="1" precision="1" value="20.248945">N20°14'56.2"</geoCoordinate>
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,
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<geoCoordinate id="36D05058FFA2914D54CD26CFFBE240C7" box="[1002,1140,221,244]" direction="east" orientation="longitude" pageId="1" precision="15" value="103.396385">E103°23'47"</geoCoordinate>
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, (
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<collectorName id="FE115349FFA2914D53AB26CFFAF640C7" box="[1164,1376,221,244]" pageId="1">Tham Nam Lot, S</collectorName>
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.
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<location id="563B6044FFA2914D525626CFFECE4124" LSID="urn:lsid:plazi:treatment:DB4D8789FFA2914F57B026A8FD2E47F2:563B6044FFA2914D525626CFFECE4124" country="Laos" latitude="20.248945" longLatPrecision="14" longitude="103.396385" name="Ban Boumfart Village" pageId="1">Ban Boumfart Village</location>
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),
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<collectorName id="FE115349FFA2914D564A2712FDB74124" box="[365,545,256,279]" pageId="1">Vieng Thong, H</collectorName>
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(o)uaphan(h)
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<collectorName id="FE115349FFA2914D55922712FD664124" box="[693,752,256,279]" pageId="1">Prov.</collectorName>
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</materialsCitation>
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,
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<materialsCitation id="E38C3CC2FFA2914D55DA2712FEAE410A" ID-GBIF-Occurrence="1675163015" collectionCode="SNUC" collectorName="H. Steiner" country="Laos" location="Col" municipality="Northern Laos-European Cave Project" pageId="1" specimenCount="1" typeStatus="holotype">
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<collectingCountry id="2BF3760FFFA2914D55DA2712FCA54124" box="[765,819,256,279]" name="Laos" pageId="1">Laos</collectingCountry>
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,
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<location id="563B6044FFA2914D54652712FCFB4124" LSID="urn:lsid:plazi:treatment:DB4D8789FFA2914F57B026A8FD2E47F2:563B6044FFA2914D54652712FCFB4124" box="[834,877,256,279]" country="Laos" municipality="Northern Laos-European Cave Project" name="Col" pageId="1">Col</location>
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:
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<collectorName id="FE115349FFA2914D545D2712FC7C4124" box="[890,1002,256,279]" pageId="1">H. Steiner</collectorName>
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,
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<collectingMunicipality id="B33FACE5FFA2914D54DF2712FA0D4124" box="[1016,1435,256,279]" pageId="1">Northern Laos-European Cave Project</collectingMunicipality>
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2009’ (
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<collectionCode id="35F5AE5AFFA2914D57C42730FEA6410A" box="[227,304,290,313]" pageId="1">SNUC</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="E38C3CC2FFA2914D56652730FBDF410A" ID-GBIF-Occurrence="1675163017" box="[322,1097,290,313]" collectionCode="SNUC" country="Laos" location="LAOS" pageId="1" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
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<emphasis id="6190EA8DFFA2914D56652730FD83410A" bold="true" box="[322,533,290,313]" pageId="1">
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<typeStatus id="8C5F883DFFA2914D56652730FE2E410A" box="[322,440,290,313]" pageId="1" type="paratype">Paratypes</typeStatus>
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:
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<collectingCountry id="2BF3760FFFA2914D56E42730FD86410A" box="[451,528,290,313]" name="Laos" pageId="1">LAOS</collectingCountry>
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:
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</emphasis>
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<specimenCount id="45E2FD16FFA2914D553B2730FDCF410A" box="[540,601,290,313]" pageId="1" type="male">2 ♂♂</specimenCount>
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, same collecting data as the
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<typeStatus id="8C5F883DFFA2914D54AE2730FC70410A" box="[905,998,290,313]" pageId="1" type="holotype">holotype</typeStatus>
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(
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<collectionCode id="35F5AE5AFFA2914D54D22730FBD4410A" box="[1013,1090,290,313]" pageId="1">SNUC</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="1BFE6514FFA2914D57E02757FAA94192" pageId="1" type="diagnosis">
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<paragraph id="535B369FFFA2914D57E02757FAA94192" blockId="1.[151,1436,152,729]" pageId="1">
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<emphasis id="6190EA8DFFA2914D57E02757FE42416F" bold="true" box="[199,468,325,348]" pageId="1">Comparative diagnosis.</emphasis>
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<taxonomicName id="94E44D1CFFA2914D56FC2756FD7F416F" authorityName="Steiner" authorityYear="2017" box="[475,745,324,348]" class="Insecta" family="Staphylinidae" genus="Mimogonellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="dreybrodti">
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<emphasis id="6190EA8DFFA2914D56FC2756FD7F416F" box="[475,745,324,348]" italics="true" pageId="1">Mimogonellus dreybrodti</emphasis>
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</taxonomicName>
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can be readily separated from the two Asian congeners,
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<emphasis id="6190EA8DFFA2914D52772757FAE6416F" box="[1360,1392,325,348]" italics="true" pageId="1">i.e.</emphasis>
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,
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<taxonomicName id="94E44D1CFFA2914D52582757FEC3414C" authority="Naomi" authorityName="Naomi" class="Insecta" family="Staphylinidae" genus="Mimogonellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="japonicus">
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<emphasis id="6190EA8DFFA2914D52582757FA03416F" box="[1407,1429,325,348]" italics="true" pageId="1">M</emphasis>
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.
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<emphasis id="6190EA8DFFA2914D57B0277AFF69414C" box="[151,255,360,383]" italics="true" pageId="1">japonicus</emphasis>
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Naomi
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</taxonomicName>
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, and
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<taxonomicName id="94E44D1CFFA2914D56B1277AFD17414C" authority="Naomi" authorityName="Naomi" box="[406,641,359,383]" class="Insecta" family="Staphylinidae" genus="Mimogonellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="yasutoshii">
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<emphasis id="6190EA8DFFA2914D56B1277AFE3A414C" box="[406,428,360,383]" italics="true" pageId="1">M</emphasis>
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.
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<emphasis id="6190EA8DFFA2914D56982775FDBD414C" box="[447,555,359,383]" italics="true" pageId="1">yasutoshii</emphasis>
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Naomi
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</taxonomicName>
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, by the much larger body size (
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<quantity id="941C9B7AFFA2914D54D7277AFB07414C" box="[1008,1169,360,383]" metricMagnitude="-3" metricUnit="m" metricValue="5.045" metricValueMax="5.13" metricValueMin="4.96" pageId="1" unit="mm" value="5.045" valueMax="5.13" valueMin="4.96">4.96–5.13 mm</quantity>
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<emphasis id="6190EA8DFFA2914D53BA277AFB25414C" box="[1181,1203,360,383]" italics="true" pageId="1">vs</emphasis>
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. 3.0–
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<quantity id="941C9B7AFFA2914D53D3277AFAD8414C" box="[1268,1358,360,383]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="1" unit="mm" value="3.2">3.2 mm</quantity>
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), more strongly curved aedeagal median lobe, and much longer parameres each exceeding the apex of the median lobe.
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</paragraph>
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</subSubSection>
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<subSubSection id="1BFE6514FFA2914F57E027BFFE05476A" lastPageId="3" pageId="1" type="description">
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<paragraph id="535B369FFFA2914D57E027BFFC2541D4" blockId="1.[151,1436,152,729]" pageId="1">
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<emphasis id="6190EA8DFFA2914D57E027BFFEC441F7" bold="true" box="[199,338,429,452]" pageId="1">Description.</emphasis>
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Male. Body (
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<figureCitation id="CBDF2A1AFFA2914D56C927BFFDD541F7" box="[494,579,429,452]" captionStart="FIGURE 1" captionStartId="1.[151,244,2002,2024]" captionTargetBox="[220,1385,801,1980]" captionTargetId="figure@1.[210,1385,786,1987]" captionTargetPageId="1" captionText="FIGURE 1. Mimogonellus dreybrodti, male. A. Dorsal habitus. B. Forebody. C. Left antenna. Scale bars: 2 mm in A; 0.5 mm in B – C." httpUri="https://zenodo.org/record/1014117/files/figure.png" pageId="1">Fig. 1A</figureCitation>
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) elongate, cylindrical; length
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|
|
<quantity id="941C9B7AFFA2914D54B727BFFBA441F7" box="[912,1074,429,452]" metricMagnitude="-3" metricUnit="m" metricValue="5.045" metricValueMax="5.13" metricValueMin="4.96" pageId="1" unit="mm" value="5.045" valueMax="5.13" valueMin="4.96">4.96–5.13 mm</quantity>
|
|||
|
|
; forebody length
|
|||
|
|
<quantity id="941C9B7AFFA2914D53D127BFFA0141F7" box="[1270,1431,429,452]" metricMagnitude="-3" metricUnit="m" metricValue="2.41" metricValueMax="2.42" metricValueMin="2.4" pageId="1" unit="mm" value="2.41" valueMax="2.42" valueMin="2.4">2.40–2.42 mm</quantity>
|
|||
|
|
; reddish brown to dark reddish brown, tarsi and mouthparts lighter in color.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="535B369FFFA2914D57E027E0FE6A42EA" blockId="1.[151,1436,152,729]" pageId="1">
|
|||
|
|
Head (
|
|||
|
|
<figureCitation id="CBDF2A1AFFA2914D562827E0FEF7423A" box="[271,353,498,521]" captionStart="FIGURE 1" captionStartId="1.[151,244,2002,2024]" captionTargetBox="[220,1385,801,1980]" captionTargetId="figure@1.[210,1385,786,1987]" captionTargetPageId="1" captionText="FIGURE 1. Mimogonellus dreybrodti, male. A. Dorsal habitus. B. Forebody. C. Left antenna. Scale bars: 2 mm in A; 0.5 mm in B – C." httpUri="https://zenodo.org/record/1014117/files/figure.png" pageId="1">Fig. 1B</figureCitation>
|
|||
|
|
) longer than wide, length
|
|||
|
|
<quantity id="941C9B7AFFA2914D55A627E0FCB6423A" box="[641,800,498,521]" metricMagnitude="-4" metricUnit="m" metricValue="7.85" metricValueMax="8.1" metricValueMin="7.6" pageId="1" unit="mm" value="0.785" valueMax="0.81" valueMin="0.76">0.76–0.81 mm</quantity>
|
|||
|
|
, width
|
|||
|
|
<quantity id="941C9B7AFFA2914D545727E0FB86423A" box="[880,1040,498,521]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" metricValueMax="7.4" metricValueMin="7.199999999999999" pageId="1" unit="mm" value="0.73" valueMax="0.74" valueMin="0.72">0.72–0.74 mm</quantity>
|
|||
|
|
; dorsum sparsely covered with large and shallow punctures, with long and yellow setae directed posteriorly, surface of disc between punctures polished and lacking microsculpture; mediobasal portion, area posterior to eyes, genae, and gular area with dense reticulate microsculpture; gular suture divergent for short distance at base, fused anteriorly; eyes slightly prominent, reniform in lateral view, each eye composed of about 130 ommatidia. Antennae (
|
|||
|
|
<figureCitation id="CBDF2A1AFFA2914D545E246FFC5C42A7" box="[889,970,637,660]" captionStart="FIGURE 1" captionStartId="1.[151,244,2002,2024]" captionTargetBox="[220,1385,801,1980]" captionTargetId="figure@1.[210,1385,786,1987]" captionTargetPageId="1" captionText="FIGURE 1. Mimogonellus dreybrodti, male. A. Dorsal habitus. B. Forebody. C. Left antenna. Scale bars: 2 mm in A; 0.5 mm in B – C." httpUri="https://zenodo.org/record/1014117/files/figure.png" pageId="1">Fig. 1C</figureCitation>
|
|||
|
|
) reaching 1/3 elytral length when stretched posteriorly; antennomeres I–III distinctly elongate, III longer than II and IV, IV–X successively stouter, moniliform, XI longer than X, narrowed apically.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="079B6617FFA2914D57B021C0FA0A47DB" box="[151,1436,2002,2024]" httpUri="https://zenodo.org/record/1014117/files/figure.png" pageId="1" targetBox="[220,1385,801,1980]" targetPageId="1">
|
|||
|
|
<paragraph id="535B369FFFA2914D57B021C0FA0A47DB" blockId="1.[151,1436,2002,2024]" box="[151,1436,2002,2024]" pageId="1">
|
|||
|
|
<emphasis id="6190EA8DFFA2914D57B021C0FE9A47DB" bold="true" box="[151,268,2002,2024]" pageId="1">FIGURE 1.</emphasis>
|
|||
|
|
<taxonomicName id="94E44D1CFFA2914D563521C1FD9247DB" authorityName="Steiner" authorityYear="2017" box="[274,516,2003,2024]" class="Insecta" family="Staphylinidae" genus="Mimogonellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="dreybrodti">
|
|||
|
|
<emphasis id="6190EA8DFFA2914D563521C1FD9247DB" box="[274,516,2003,2024]" italics="true" pageId="1">Mimogonellus dreybrodti</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, male.
|
|||
|
|
<emphasis id="6190EA8DFFA2914D556D21C0FDF447DB" bold="true" box="[586,610,2002,2024]" pageId="1">A.</emphasis>
|
|||
|
|
Dorsal habitus.
|
|||
|
|
<emphasis id="6190EA8DFFA2914D55D921C0FC8247DB" bold="true" box="[766,788,2002,2024]" pageId="1">B.</emphasis>
|
|||
|
|
Forebody.
|
|||
|
|
<emphasis id="6190EA8DFFA2914D54A721C0FC0147DB" bold="true" box="[896,919,2002,2024]" pageId="1">C.</emphasis>
|
|||
|
|
Left antenna. Scale bars: 2 mm in A; 0.5 mm in B–C.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="535B369FFFA1914E57E0268AFB7440C7" blockId="2.[151,1436,152,625]" pageId="2">
|
|||
|
|
Pronotum (
|
|||
|
|
<figureCitation id="CBDF2A1AFFA1914E5618268AFE06409C" box="[319,400,152,175]" captionStart="FIGURE 1" captionStartId="1.[151,244,2002,2024]" captionTargetBox="[220,1385,801,1980]" captionTargetId="figure@1.[210,1385,786,1987]" captionTargetPageId="1" captionText="FIGURE 1. Mimogonellus dreybrodti, male. A. Dorsal habitus. B. Forebody. C. Left antenna. Scale bars: 2 mm in A; 0.5 mm in B – C." httpUri="https://zenodo.org/record/1014117/files/figure.png" pageId="2">Fig. 1B</figureCitation>
|
|||
|
|
) about as broad as elytra, transverse, widest at middle, strongly constricted at base; length
|
|||
|
|
<quantity id="941C9B7AFFA1914E5278268AFF6F40E2" metricMagnitude="-4" metricUnit="m" metricValue="7.1" metricValueMax="7.3" metricValueMin="6.8999999999999995" pageId="2" unit="mm" value="0.71" valueMax="0.73" valueMin="0.69">0.69– 0.73 mm</quantity>
|
|||
|
|
, width
|
|||
|
|
<quantity id="941C9B7AFFA1914E566E26A8FE7F40E2" box="[329,489,186,209]" metricMagnitude="-4" metricUnit="m" metricValue="9.25" metricValueMax="9.6" metricValueMin="8.9" pageId="2" unit="mm" value="0.925" valueMax="0.96" valueMin="0.89">0.89–0.96 mm</quantity>
|
|||
|
|
; with punctures and setae similar to those on head, with reticulate microsculpture along posterolateral and posterior margins. Mesoscutellum roundly triangular, with reticulate microsculpture.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="535B369FFFA1914E57E02712FAAA410A" blockId="2.[151,1436,152,625]" pageId="2">
|
|||
|
|
Elytra (
|
|||
|
|
<figureCitation id="CBDF2A1AFFA1914E56312712FEF04124" box="[278,358,256,279]" captionStart="FIGURE 1" captionStartId="1.[151,244,2002,2024]" captionTargetBox="[220,1385,801,1980]" captionTargetId="figure@1.[210,1385,786,1987]" captionTargetPageId="1" captionText="FIGURE 1. Mimogonellus dreybrodti, male. A. Dorsal habitus. B. Forebody. C. Left antenna. Scale bars: 2 mm in A; 0.5 mm in B – C." httpUri="https://zenodo.org/record/1014117/files/figure.png" pageId="2">Fig. 1B</figureCitation>
|
|||
|
|
) about as long as wide, length
|
|||
|
|
<quantity id="941C9B7AFFA1914E55882712FCDA4124" box="[687,844,256,279]" metricMagnitude="-4" metricUnit="m" metricValue="9.15" metricValueMax="9.5" metricValueMin="8.8" pageId="2" unit="mm" value="0.915" valueMax="0.95" valueMin="0.88">0.88–0.95 mm</quantity>
|
|||
|
|
, width
|
|||
|
|
<quantity id="941C9B7AFFA1914E54BE2712FBA14124" box="[921,1079,256,279]" metricMagnitude="-4" metricUnit="m" metricValue="9.25" metricValueMax="9.3" metricValueMin="9.2" pageId="2" unit="mm" value="0.925" valueMax="0.93" valueMin="0.92">0.92–0.93 mm</quantity>
|
|||
|
|
; parallel sided; sparsely punctate, punctures shallower than those of pronotum; basal area lateral to mesoscutellum with reticulate microsculpture.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="535B369FFFA1914E57E02757FF7741F7" blockId="2.[151,1436,152,625]" pageId="2">
|
|||
|
|
Legs short; profemora projected ventrally at basal half; protibiae (
|
|||
|
|
<figureCitation id="CBDF2A1AFFA1914E54AA2757FC74416F" box="[909,994,325,348]" captionStart="FIGURE 2" captionStartId="2.[151,244,1980,2002]" captionTargetBox="[182,1417,697,1963]" captionTargetId="figure@2.[173,1417,677,1971]" captionTargetPageId="2" captionText="FIGURE 2. Mimogonellus dreybrodti, male. A. Protibia. B. Mesotibia. C. Tergites IX – X. D. Sternite VIII. E. Sternite IX. F – G. Aedeagus, in lateral (F), and dorsal (G) views. Scale bars: 0.3 mm in A – D; 0.2 mm in E – G." httpUri="https://zenodo.org/record/1014119/files/figure.png" pageId="2">Fig. 2A</figureCitation>
|
|||
|
|
) curved at basal 1/3, with seven external and two apical spines, with well-developed ctenidium along mesal margin; mesotibia (
|
|||
|
|
<figureCitation id="CBDF2A1AFFA1914E5347277AFB20414C" box="[1120,1206,360,383]" captionStart="FIGURE 2" captionStartId="2.[151,244,1980,2002]" captionTargetBox="[182,1417,697,1963]" captionTargetId="figure@2.[173,1417,677,1971]" captionTargetPageId="2" captionText="FIGURE 2. Mimogonellus dreybrodti, male. A. Protibia. B. Mesotibia. C. Tergites IX – X. D. Sternite VIII. E. Sternite IX. F – G. Aedeagus, in lateral (F), and dorsal (G) views. Scale bars: 0.3 mm in A – D; 0.2 mm in E – G." httpUri="https://zenodo.org/record/1014119/files/figure.png" pageId="2">Fig. 2B</figureCitation>
|
|||
|
|
) with lateral row of macrosetae, with two external and two apical spines; metatibia with apicolateral row of macrosetae, and two apical spines.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="535B369FFFA1914E57E027C2FB324242" blockId="2.[151,1436,152,625]" pageId="2">
|
|||
|
|
Abdomen slightly broadened posteriorly; punctation and setation similar to those on elytra, but sparser and finer, also with dense reticulate microsculpture. Paired tergite IX (
|
|||
|
|
<figureCitation id="CBDF2A1AFFA1914E540F27E0FCEA423A" box="[808,892,498,521]" captionStart="FIGURE 2" captionStartId="2.[151,244,1980,2002]" captionTargetBox="[182,1417,697,1963]" captionTargetId="figure@2.[173,1417,677,1971]" captionTargetPageId="2" captionText="FIGURE 2. Mimogonellus dreybrodti, male. A. Protibia. B. Mesotibia. C. Tergites IX – X. D. Sternite VIII. E. Sternite IX. F – G. Aedeagus, in lateral (F), and dorsal (G) views. Scale bars: 0.3 mm in A – D; 0.2 mm in E – G." httpUri="https://zenodo.org/record/1014119/files/figure.png" pageId="2">Fig. 2C</figureCitation>
|
|||
|
|
) with several exceptionally elongate lateral setae; tergite X (
|
|||
|
|
<figureCitation id="CBDF2A1AFFA1914E56232407FECE421F" box="[260,344,533,556]" captionStart="FIGURE 2" captionStartId="2.[151,244,1980,2002]" captionTargetBox="[182,1417,697,1963]" captionTargetId="figure@2.[173,1417,677,1971]" captionTargetPageId="2" captionText="FIGURE 2. Mimogonellus dreybrodti, male. A. Protibia. B. Mesotibia. C. Tergites IX – X. D. Sternite VIII. E. Sternite IX. F – G. Aedeagus, in lateral (F), and dorsal (G) views. Scale bars: 0.3 mm in A – D; 0.2 mm in E – G." httpUri="https://zenodo.org/record/1014119/files/figure.png" pageId="2">Fig. 2C</figureCitation>
|
|||
|
|
) strongly protruding posterolaterally, with two small denticles at posterior margin. Sternites VII–VIII lacking median impression; sternite VIII (
|
|||
|
|
<figureCitation id="CBDF2A1AFFA1914E5543242AFD2C427C" box="[612,698,568,591]" captionStart="FIGURE 2" captionStartId="2.[151,244,1980,2002]" captionTargetBox="[182,1417,697,1963]" captionTargetId="figure@2.[173,1417,677,1971]" captionTargetPageId="2" captionText="FIGURE 2. Mimogonellus dreybrodti, male. A. Protibia. B. Mesotibia. C. Tergites IX – X. D. Sternite VIII. E. Sternite IX. F – G. Aedeagus, in lateral (F), and dorsal (G) views. Scale bars: 0.3 mm in A – D; 0.2 mm in E – G." httpUri="https://zenodo.org/record/1014119/files/figure.png" pageId="2">Fig. 2D</figureCitation>
|
|||
|
|
) subtriangular, each side with few long setae and two small, acute denticles; sternite IX (
|
|||
|
|
<figureCitation id="CBDF2A1AFFA1914E56A42448FE424242" box="[387,468,602,625]" captionStart="FIGURE 2" captionStartId="2.[151,244,1980,2002]" captionTargetBox="[182,1417,697,1963]" captionTargetId="figure@2.[173,1417,677,1971]" captionTargetPageId="2" captionText="FIGURE 2. Mimogonellus dreybrodti, male. A. Protibia. B. Mesotibia. C. Tergites IX – X. D. Sternite VIII. E. Sternite IX. F – G. Aedeagus, in lateral (F), and dorsal (G) views. Scale bars: 0.3 mm in A – D; 0.2 mm in E – G." httpUri="https://zenodo.org/record/1014119/files/figure.png" pageId="2">Fig. 2E</figureCitation>
|
|||
|
|
) elongate, weakly sclerotized, apex rounded and finely denticulate.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="079B6617FFA1914E57B021AEFB9547DD" httpUri="https://zenodo.org/record/1014119/files/figure.png" pageId="2" targetBox="[182,1417,697,1963]" targetPageId="2">
|
|||
|
|
<paragraph id="535B369FFFA1914E57B021AEFB9547DD" blockId="2.[151,1436,1980,2030]" pageId="2">
|
|||
|
|
<emphasis id="6190EA8DFFA1914E57B021AEFE8647E1" bold="true" box="[151,272,1980,2002]" pageId="2">FIGURE 2.</emphasis>
|
|||
|
|
<taxonomicName id="94E44D1CFFA1914E563E21AEFD9947E2" authorityName="Steiner" authorityYear="2017" box="[281,527,1980,2001]" class="Insecta" family="Staphylinidae" genus="Mimogonellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="dreybrodti">
|
|||
|
|
<emphasis id="6190EA8DFFA1914E563E21AEFD9947E2" box="[281,527,1980,2001]" italics="true" pageId="2">Mimogonellus dreybrodti</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, male.
|
|||
|
|
<emphasis id="6190EA8DFFA1914E557B21AEFDE247E1" bold="true" box="[604,628,1980,2002]" pageId="2">A.</emphasis>
|
|||
|
|
Protibia.
|
|||
|
|
<emphasis id="6190EA8DFFA1914E55FF21AEFD7847E1" bold="true" box="[728,750,1980,2002]" pageId="2">B.</emphasis>
|
|||
|
|
Mesotibia.
|
|||
|
|
<emphasis id="6190EA8DFFA1914E544021AEFCE847E1" bold="true" box="[871,894,1980,2002]" pageId="2">C.</emphasis>
|
|||
|
|
Tergites IX–X.
|
|||
|
|
<emphasis id="6190EA8DFFA1914E530421AEFBAD47E1" bold="true" box="[1059,1083,1980,2002]" pageId="2">D.</emphasis>
|
|||
|
|
Sternite VIII.
|
|||
|
|
<emphasis id="6190EA8DFFA1914E53F721AEFB7047E1" bold="true" box="[1232,1254,1980,2002]" pageId="2">E.</emphasis>
|
|||
|
|
Sternite IX.
|
|||
|
|
<emphasis id="6190EA8DFFA1914E524B21AEFA0A47E1" bold="true" box="[1388,1436,1980,2002]" pageId="2">F–G.</emphasis>
|
|||
|
|
Aedeagus, in lateral (F), and dorsal (G) views. Scale bars: 0.3 mm in A–D; 0.2 mm in E–G.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="079B6617FFA0914F57B0202EFC9C46BF" httpUri="https://zenodo.org/record/1014121/files/figure.png" pageId="3" targetBox="[151,1436,214,1555]" targetPageId="3">
|
|||
|
|
<paragraph id="535B369FFFA0914F57B0202EFC9C46BF" blockId="3.[151,1436,1596,1676]" pageId="3">
|
|||
|
|
<emphasis id="6190EA8DFFA0914F57B0202EFE9B4661" bold="true" box="[151,269,1596,1618]" pageId="3">FIGURE 3.</emphasis>
|
|||
|
|
Collecting site and environment.
|
|||
|
|
<emphasis id="6190EA8DFFA0914F5571202EFDF84661" bold="true" box="[598,622,1596,1618]" pageId="3">A.</emphasis>
|
|||
|
|
Area map, showing the location of Tham Nam Lot.
|
|||
|
|
<emphasis id="6190EA8DFFA0914F5349202EFB124661" bold="true" box="[1134,1156,1596,1618]" pageId="3">B.</emphasis>
|
|||
|
|
Map of Tham Nam Lot (red arrow indicates the locality where the beetles were collected).
|
|||
|
|
<emphasis id="6190EA8DFFA0914F55CC204BFC95465C" bold="true" box="[747,771,1625,1647]" pageId="3">C.</emphasis>
|
|||
|
|
One of the entrances of the cave.
|
|||
|
|
<emphasis id="6190EA8DFFA0914F536F204BFBF6465C" bold="true" box="[1096,1120,1625,1647]" pageId="3">D.</emphasis>
|
|||
|
|
General environment around the cave. (Fig. 3A–B reproduced from Dreybrodt & Laumanns 2010)
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="535B369FFFA0914F57E020A5FE044705" blockId="3.[151,1436,1719,1985]" pageId="3">
|
|||
|
|
Aedeagus (
|
|||
|
|
<figureCitation id="CBDF2A1AFFA0914F561820A5FE3946FD" box="[319,431,1719,1742]" captionStart="FIGURE 2" captionStartId="2.[151,244,1980,2002]" captionTargetBox="[182,1417,697,1963]" captionTargetId="figure@2.[173,1417,677,1971]" captionTargetPageId="2" captionText="FIGURE 2. Mimogonellus dreybrodti, male. A. Protibia. B. Mesotibia. C. Tergites IX – X. D. Sternite VIII. E. Sternite IX. F – G. Aedeagus, in lateral (F), and dorsal (G) views. Scale bars: 0.3 mm in A – D; 0.2 mm in E – G." httpUri="https://zenodo.org/record/1014119/files/figure.png" pageId="3">Fig. 2F–G</figureCitation>
|
|||
|
|
) elongate, length of median lobe
|
|||
|
|
<quantity id="941C9B7AFFA0914F543220A5FCE146FD" box="[789,887,1719,1742]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="3" unit="mm" value="0.56">0.56 mm</quantity>
|
|||
|
|
, length including parameres
|
|||
|
|
<quantity id="941C9B7AFFA0914F538C20A5FA9846FD" box="[1195,1294,1719,1742]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" pageId="3" unit="mm" value="0.64">0.64 mm</quantity>
|
|||
|
|
; median lobe weakly sclerotized at basal half, apical half strongly curved dorsally and narrowed apically; parameres long, exceeding apex of median lobe, each paramere with two long macrosetae at apex; endophallus with three sclerites, elongate, twisted, and spiral-like.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="535B369FFFA0914F57E02150FE05476A" blockId="3.[151,1436,1719,1985]" box="[199,403,1858,1881]" pageId="3">Female. Unknown.</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="1BFE6514FFA0914F57E02176FD1C4748" box="[199,650,1892,1915]" pageId="3" type="distribution">
|
|||
|
|
<paragraph id="535B369FFFA0914F57E02176FD1C4748" blockId="3.[151,1436,1719,1985]" box="[199,650,1892,1915]" pageId="3">
|
|||
|
|
<emphasis id="6190EA8DFFA0914F57E02176FECF4748" bold="true" box="[199,345,1892,1915]" pageId="3">Distribution.</emphasis>
|
|||
|
|
Northern
|
|||
|
|
<collectingCountry id="2BF3760FFFA0914F56E12176FE684748" box="[454,510,1892,1915]" name="Laos" pageId="3">Laos</collectingCountry>
|
|||
|
|
:
|
|||
|
|
<collectingRegion id="9120F87DFFA0914F552F2176FD104748" box="[520,646,1892,1915]" country="Laos" name="Houaphan" pageId="3">Houaphanh</collectingRegion>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="1BFE6514FFA0914F57E02195FD2E47F2" pageId="3" type="etymology">
|
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|
|
<paragraph id="535B369FFFA0914F57E02195FD2E47F2" blockId="3.[151,1436,1719,1985]" pageId="3">
|
|||
|
|
<emphasis id="6190EA8DFFA0914F57E02195FED147AD" bold="true" box="[199,327,1927,1950]" pageId="3">Etymology.</emphasis>
|
|||
|
|
We dedicate this new species to Dr. Joerg Dreybrodt (
|
|||
|
|
<collectingCountry id="2BF3760FFFA0914F54BA2195FB9047AD" box="[925,1030,1927,1950]" name="Germany" pageId="3">Germany</collectingCountry>
|
|||
|
|
), a specialist in survey and mapping, and the coordinator of the
|
|||
|
|
<date id="275A105FFFA0914F569621B8FDBE47F2" box="[433,552,1962,1985]" pageId="3" value="2009" valueMax="2010">2009–2010</date>
|
|||
|
|
survey team.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|