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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.127.1748" ID-GBIF-Dataset="5c4f147a-0e68-4abd-8b74-d15416f61f4d" ID-PMC="PMC3175127" ID-Pensoft-Pub="1313-2970-127-1" ID-Pensoft-UUID="FFF2FFC95E2C4223FFFFFFBCFFDAFF8F" ID-PubMed="21998544" ID-Zenodo-Dep="577174" ID-ZooBank="D170BE13535B467899997BA431A1AD0A" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-127-1" ModsDocOrigin="ZooKeys 127" ModsDocTitle="Halocoryza Alluaud 1919, sea-side beetles of the Indian, Atlantic (sensu lato), and Pacific Oceans: a generic synopsis and description of a remarkable new species from Baja California Sur, México (Coleoptera, Carabidae, Scaritini, Clivinina)" checkinTime="1451249931271" checkinUser="pensoft" docAuthor="Erwin, Terry L." docDate="2011" docId="C4881D51D1D9D7E852050143CA0A9778" docLanguage="en" docName="ZooKeys 127: 1-13" docOrigin="ZooKeys 127" docSource="http://dx.doi.org/10.3897/zookeys.127.1748" docTitle="Halocoryza arenaria Darlington 1939" docType="treatment" docVersion="4" id="FFF2FFC95E2C4223FFFFFFBCFFDAFF8F" lastPageNumber="5" masterDocId="FFF2FFC95E2C4223FFFFFFBCFFDAFF8F" masterDocTitle="Halocoryza Alluaud 1919, sea-side beetles of the Indian, Atlantic (sensu lato), and Pacific Oceans: a generic synopsis and description of a remarkable new species from Baja California Sur, Mexico (Coleoptera, Carabidae, Scaritini, Clivinina)" masterLastPageNumber="13" masterPageNumber="1" pageNumber="4" updateTime="1668152055969" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Halocoryza Alluaud 1919, sea-side beetles of the Indian, Atlantic (sensu lato), and Pacific Oceans: a generic synopsis and description of a remarkable new species from Baja California Sur, Mexico (Coleoptera, Carabidae, Scaritini, Clivinina)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Erwin, Terry L.</mods:namePart>
<mods:affiliation>Hyper-diversity Group, Department of Entomology, MRC- 187, National Museum of Natural History, Smithsonian Institution, Washington, P. O. Box 37012, DC 20013 - 7012, USA</mods:affiliation>
<mods:nameIdentifier type="email">erwint@si.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2011</mods:date>
<mods:detail type="volume">
<mods:number>127</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>13</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.127.1748</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.127.1748</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-127-1</mods:identifier>
<mods:identifier type="ZooBank">D170BE13535B467899997BA431A1AD0A</mods:identifier>
<mods:identifier type="Pensoft-UUID">FFF2FFC95E2C4223FFFFFFBCFFDAFF8F</mods:identifier>
<mods:identifier type="Zenodo-Dep">577174</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152030631" LSID="urn:lsid:plazi:treatment:C4881D51D1D9D7E852050143CA0A9778" httpUri="http://treatment.plazi.org/id/C4881D51D1D9D7E852050143CA0A9778" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<subSubSection pageId="3" pageNumber="4" type="nomenclature">
<paragraph pageId="3" pageNumber="4">
<taxonomicName LSID="C4881D51-D1D9-D7E8-5205-0143CA0A9778" authority="(Darlington, 1939)" authorityName="Darlington" authorityYear="1939" baseAuthorityName="Darlington" baseAuthorityYear="1939" class="Insecta" family="Carabidae" genus="Halocoryza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Halocoryza arenaria" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="arenaria">Halocoryza arenaria (Darlington, 1939)</taxonomicName>
<figureCitation captionStart="" captionStartId="F1" captionText=". Figure 1 - 2. 1 Color image, showing habitus of Halocoryza acapulcana Whitehead, 1966, dorsal aspect, ABL = 2.4 mm, ADP 127171; Acapulco, Mexico 2 Color image, showing habitus of Halocoryza arenaria (Darlington, 1939), dorsal aspect, ABL = 2.9 mm, ADP 116798; St. Johns, Virgin Islands." httpUri="https://binary.pensoft.net/fig/17265" pageId="3" pageNumber="4">Figs 2</figureCitation>
<figureCitation captionStart="Figure 4–8" captionStartId="F3" captionText="Figure 4 - 8. 4 Line drawing, showing habitus of Halocoryza acapulcana Whitehead, 1966, dorsal aspect, ABL = 2.4 mm; Acapulco, Mexico; modified from Whitehead 1966 5 Line drawing, showing habitus of Halocoryza arenaria (Darlington, 1939), dorsal aspect, ABL = 2.9 mm; Rio Piedras, Puerto Rico; modified from Whitehead, 1966 6 Line drawing, showing habitus of Halocoryza whiteheadiana sp. n., dorsal aspect, ABL = 2.9 mm; ABL = 2.8 mm, ADP 127139; Isla Del Carmen, Mexico 7 Line drawing, showing male genitalia, median lobe, of Halocoryza acapulcana Whitehead, 1966, left lateral aspect, 0.41 mm (dorsal margin of basal lobe to apex); Acapulco, Mexico; modified from Whitehead 1966 8 Line drawing, showing male genitalia, median lobe, Halocoryza arenaria (Darlington, 1939), left lateral aspect with internal sac everted, 0.41 mm (dorsal margin of basal lobe to apex); Rio Piedras, Puerto Rico; modified from Whitehead, 1966." httpUri="https://binary.pensoft.net/fig/17267" pageId="3" pageNumber="4">5, 8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="4" type="reference_group">
<paragraph pageId="3" pageNumber="4">
<taxonomicName class="Insecta" family="Carabidae" genus="Schizogenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="arenaria">Schizogenius arenaria</taxonomicName>
Darlington, 1939: 84
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="4" type="common name">
<paragraph pageId="3" pageNumber="4">Common name.</paragraph>
<paragraph pageId="3" pageNumber="4">
<emphasis bold="true" pageId="3" pageNumber="4">Sand Saline Catarrh Beetle</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="4" type="geographic distribution">
<paragraph pageId="3" pageNumber="4">Geographic Distribution.</paragraph>
<paragraph pageId="3" pageNumber="4">
Native, New World. Barbados; Brazil; Dominican Republic; Grenada; Guadeloupe; Jamaica;
<normalizedToken originalValue="México">Mexico</normalizedToken>
- QR, YC;
<normalizedToken originalValue="Panamá">Panama</normalizedToken>
; Puerto Rico; USA - FL; Virgin Islands - St. John, St. Thomas; natural invasive, Africa - Cameroon.
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="way of life">
<paragraph pageId="3" pageNumber="4">Way of Life.</paragraph>
<paragraph lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<emphasis bold="true" pageId="3" pageNumber="4">Macrohabitat:</emphasis>
Lowlands, sea level - 1 meter altitude, on sea beaches and in the intertidal area, at or near the high tide line, and in mangrove swamps.
<emphasis bold="true" pageId="4" pageNumber="5">
<pageBreakToken pageId="4" pageNumber="5" start="start">Microhabitat</pageBreakToken>
:
</emphasis>
Adults are ground-dwelling on exposed wet substrate consisting of coquina-coral cemented by very fine silt or sand and covered with seaweed mats.
<emphasis bold="true" pageId="4" pageNumber="5">Dispersal abilities:</emphasis>
Wing-polymorphic: macropterous form probably capable of flight; brachypterous form, consequently flightless thus vagility limited to walking or running; both forms slow runners.
<emphasis bold="true" pageId="4" pageNumber="5">Seasonal occurrence:</emphasis>
Adults have been found in March - April, July, and October.
<emphasis bold="true" pageId="4" pageNumber="5">Behavior:</emphasis>
Adults are nocturnal predaceous halobionts and take cover in the sand or under drift and piles of seaweed on the beach. Populations of this species are associated with the centipede
<taxonomicName authorityName="Crabill" authorityYear="1959" class="Chilopoda" family="Schendylidae" genus="Pectiniunguis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Geophilomorpha" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="halirrhytus">
<emphasis italics="true" pageId="4" pageNumber="5">Pectiniunguis halirrhytus</emphasis>
</taxonomicName>
Crabill. In the northern part of their range, adults overwinter in the substrate; in the southern part, they likely aestivate during the dry season in the substrate.
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/17266" pageId="4" pageNumber="5" start="Figure 3" startId="F2">
<paragraph pageId="4" pageNumber="5">
<emphasis bold="true" pageId="4" pageNumber="5">Figure 3.</emphasis>
Color image, showing habitus of
<taxonomicName authorityName="Erwin" authorityYear="2011" class="Insecta" family="Carabidae" genus="Halocoryza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="whiteheadiana">
<emphasis italics="true" pageId="4" pageNumber="5">Halocoryza whiteheadiana</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="4" pageNumber="5">sp. n.</emphasis>
, 3a, left lateral aspect, 3b, dorsal aspect, ABL = 2.9mm; Holotype: Isla Del Carmen, BJ,
<normalizedToken originalValue="México">Mexico</normalizedToken>
.
</paragraph>
</caption>
<caption httpUri="https://binary.pensoft.net/fig/17267" pageId="4" pageNumber="5" start="Figure 4–8" startId="F3">
<paragraph pageId="4" pageNumber="5">
<emphasis bold="true" pageId="4" pageNumber="5">Figure 4-8.</emphasis>
<emphasis bold="true" pageId="4" pageNumber="5">4</emphasis>
Line drawing, showing habitus of
<taxonomicName authorityName="Whitehead" authorityYear="1966" class="Insecta" family="Carabidae" genus="Halocoryza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="acapulcana">
<emphasis italics="true" pageId="4" pageNumber="5">Halocoryza acapulcana</emphasis>
</taxonomicName>
Whitehead, 1966, dorsal aspect, ABL = 2.4mm; Acapulco,
<normalizedToken originalValue="México">Mexico</normalizedToken>
; modified from
<bibRefCitation author="Whitehead, DR" journalOrPublisher="Psyche" pageId="8" pageNumber="9" pagination="217 - 228" refId="B19" refString="Whitehead, DR, 1966. A review of Halocoryza Alluaud, with notes on its relationship to Schizogenius Putzeys (Coleoptera, Carabidae). Psyche 73: 217 - 228" title="A review of Halocoryza Alluaud, with notes on its relationship to Schizogenius Putzeys (Coleoptera, Carabidae)." volume="73" year="1966">Whitehead 1966</bibRefCitation>
<emphasis bold="true" pageId="4" pageNumber="5">5</emphasis>
Line drawing, showing habitus of
<taxonomicName baseAuthorityName="Darlington" baseAuthorityYear="1939" class="Insecta" family="Carabidae" genus="Halocoryza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="arenaria">
<emphasis italics="true" pageId="4" pageNumber="5">Halocoryza arenaria</emphasis>
</taxonomicName>
(Darlington, 1939), dorsal aspect, ABL = 2.9mm; Rio Piedras, Puerto Rico; modified from Whitehead, 1966
<emphasis bold="true" pageId="4" pageNumber="5">6</emphasis>
Line drawing, showing habitus of
<taxonomicName authorityName="Erwin" authorityYear="2011" class="Insecta" family="Carabidae" genus="Halocoryza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="whiteheadiana">
<emphasis italics="true" pageId="4" pageNumber="5">Halocoryza whiteheadiana</emphasis>
</taxonomicName>
sp. n., dorsal aspect, ABL = 2.9mm; ABL = 2.8mm,ADP127139; Isla Del Carmen,
<normalizedToken originalValue="México">Mexico</normalizedToken>
<emphasis bold="true" pageId="4" pageNumber="5">7</emphasis>
Line drawing, showing male genitalia, median lobe, of
<taxonomicName authorityName="Whitehead" authorityYear="1966" class="Insecta" family="Carabidae" genus="Halocoryza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="acapulcana">
<emphasis italics="true" pageId="4" pageNumber="5">Halocoryza acapulcana</emphasis>
</taxonomicName>
Whitehead, 1966, left lateral aspect, 0.41mm (dorsal margin of basal lobe to apex); Acapulco,
<normalizedToken originalValue="México">Mexico</normalizedToken>
; modified from
<bibRefCitation author="Whitehead, DR" journalOrPublisher="Psyche" pageId="8" pageNumber="9" pagination="217 - 228" refId="B19" refString="Whitehead, DR, 1966. A review of Halocoryza Alluaud, with notes on its relationship to Schizogenius Putzeys (Coleoptera, Carabidae). Psyche 73: 217 - 228" title="A review of Halocoryza Alluaud, with notes on its relationship to Schizogenius Putzeys (Coleoptera, Carabidae)." volume="73" year="1966">Whitehead 1966</bibRefCitation>
<emphasis bold="true" pageId="4" pageNumber="5">8</emphasis>
Line drawing, showing male genitalia, median lobe,
<taxonomicName baseAuthorityName="Darlington" baseAuthorityYear="1939" class="Insecta" family="Carabidae" genus="Halocoryza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="arenaria">
<emphasis italics="true" pageId="4" pageNumber="5">Halocoryza arenaria</emphasis>
</taxonomicName>
(Darlington, 1939), left lateral aspect with internal sac everted, 0.41mm (dorsal margin of basal lobe to apex); Rio Piedras, Puerto Rico; modified from Whitehead, 1966.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="4" pageNumber="5" type="references">
<paragraph pageId="4" pageNumber="5">References.</paragraph>
<paragraph pageId="4" pageNumber="5">
<bibRefCitation author="Bruneau de Mire, P" editor="Erwin, TL" journalOrPublisher="Proceedings of the First International Symposium of Carabidology. Dr. W. Junk b. v. Publishers, The Hague" pageId="8" pageNumber="9" pagination="327 - 330" refId="B3" refString="Bruneau de Mire, P, 1979. Trans-Atlantic dispersal: Several examples of colonization of the Gulf of Biafra by Middle American stocks of Carabidae. In: Erwin, TL, Ball, GE, Whitehead, DR, Halpern, AL, Eds., Carabid Beetles: Their evolution, natural history, and classification. Proceedings of the First International Symposium of Carabidology. Dr. W. Junk b.v. Publishers, The Hague: 327 - 330" title="Trans-Atlantic dispersal: Several examples of colonization of the Gulf of Biafra by Middle American stocks of Carabidae." volumeTitle="Carabid Beetles: Their evolution, natural history, and classification." year="1979">
Bruneau de
<normalizedToken originalValue="Miré">Mire</normalizedToken>
(1979)
</bibRefCitation>
,
<bibRefCitation author="Nichols, SW" journalOrPublisher="Cornell University" pageId="8" pageNumber="9" refId="B14" refString="Nichols, SW, 1988. Systematics and biogeography of West Indian Scaritinae (Coleoptera: Carabidae). Cornell University" title="Systematics and biogeography of West Indian Scaritinae (Coleoptera: Carabidae)" year="1988">Nichols (1988</bibRefCitation>
, Ph.D. dissertation),
<bibRefCitation author="Peck, SB" journalOrPublisher="Arthropods of Florida and Neighboring Land Areas" pageId="8" pageNumber="9" refId="B16" refString="Peck, SB, Thomas, MC, 1998. A distributional checklist of the beetles (Coleoptera) of Florida. Arthropods of Florida and Neighboring Land Areas 16, 180 pp." title="A distributional checklist of the beetles (Coleoptera) of Florida" volume="16" year="1998">Peck and Thomas (1998)</bibRefCitation>
, (
<bibRefCitation author="Whitehead, DR" journalOrPublisher="Psyche" pageId="8" pageNumber="9" pagination="217 - 228" refId="B19" refString="Whitehead, DR, 1966. A review of Halocoryza Alluaud, with notes on its relationship to Schizogenius Putzeys (Coleoptera, Carabidae). Psyche 73: 217 - 228" title="A review of Halocoryza Alluaud, with notes on its relationship to Schizogenius Putzeys (Coleoptera, Carabidae)." volume="73" year="1966">Whitehead (1966</bibRefCitation>
, 1969).
</paragraph>
</subSubSection>
</treatment>
</document>
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