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<document id="72607FF21162564F323C790FB091FE4C" ID-CLB-Dataset="8712" ID-DOI="10.11646/zootaxa.4908.4.5" ID-GBIF-Dataset="6925e575-3634-4492-8be6-b422b73fe888" ID-ISSN="1175-5326" ID-Zenodo-Dep="4447214" ID-ZooBank="FD97747A-BF20-4820-B35E-67B432FF9987" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610980108797" checkinUser="plazi" docAuthor="Sikorski, Andrey V., Pavlova, Lyudmila V., Sardá, Rafael, Langeneck, Joachim, Gil, João &amp; Ravara, Ascensão" docDate="2021" docId="03C687E2FFF0EE03F0CB57FFDB6A1DD4" docLanguage="en" docName="zootaxa4908.4.5.pdf" docOrigin="Zootaxa 4908 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Laonice (Sarsiana) alberti Sikorski, Langeneck &amp; Pavlova 2021, sp. nov." docType="treatment" docUuid="4D214346-F738-43B2-AEE0-A6D82B53EDA8" docUuidSource="ZooBank" docVersion="7" lastPageNumber="518" masterDocId="FFFFFF9AFFF2EE07F05C5600DA261D01" masterDocTitle="Two new deep-sea species of Laonice (Annelida: Spionidae) from the Mediterranean Sea" masterLastPageNumber="526" masterPageNumber="515" pageNumber="517" updateTime="1698924590607" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="95D94A0C6B81E1BFA79575A4384A3E48">Two new deep-sea species of Laonice (Annelida: Spionidae) from the Mediterranean Sea</mods:title>
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<mods:namePart id="C2ACDA28D18BA77E53377E6CFAEFDA04">Sikorski, Andrey V.</mods:namePart>
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<mods:affiliation id="251F7624FA74A5C8F0F1FBC82E470132">Akvaplan-niva AS, Fram Centre, 9296 Troms, Norway. as @ akvaplan. niva. no; https: // orcid. org / 0000 - 0001 - 8073 - 0027</mods:affiliation>
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<mods:namePart id="786197123320656095E0DAE06CCBCE82">Pavlova, Lyudmila V.</mods:namePart>
<mods:affiliation id="77ED8278C53F8FE288294C83DB016A9E">Russian Academy of Sciences, Kola Science Centre, Murmansk Marine Biological Institute, Vladimirskaya str. 17, Murmansk 183010, Russia. sea 1234 @ mail. ru; http: // orcid. org / 0000 - 0003 - 4422 - 0366</mods:affiliation>
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<mods:namePart id="407168C4BDA0C425D49050831D5AE62E">Sardá, Rafael</mods:namePart>
<mods:affiliation id="300032EF9BA0086EEE4A3FB1C3E3422E">Centre d’Estudis Avançats de Blanes (CEAB-CSIC), Carrer d’accès a la Cala St. Francesc, 14, Blanes-Girona- 17300, Spain. sarda @ ceab. csic. es; http: // orcid. org / 0000 - 0003 - 2059 - 9017</mods:affiliation>
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<mods:namePart id="94A25833308CDA79976CD31821FD28DA">Langeneck, Joachim</mods:namePart>
<mods:affiliation id="4A0127181DBB095F16559F986BB63F17">Dipartimento di Biologia, Universit di Pisa, via Derna 1, 56126 Pisa, Italy. jlangeneck @ biologia. unipi. it; http: // orcid. org / 0000 - 0003 - 3665 - 8683</mods:affiliation>
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<mods:namePart id="03AD98DEF2639C18A05794793549C19C">Gil, João</mods:namePart>
<mods:affiliation id="98E506CE00FFA361B28D724B0892F74D">Centre of Marine Sciences (CCMAR), Universidade do Algarve, Campus de Gambelas, 8005 - 139 Faro, Portugal. joaocfgil @ gmail. com; http: // orcid. org / 0000 - 0002 - 1856 - 3333</mods:affiliation>
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<mods:namePart id="149174724505AF9C816779491A7D7B4A">Ravara, Ascensão</mods:namePart>
<mods:affiliation id="A85F8A556449B42CC616D2CE883D6E06">CESAM - Centro de Estudos do Ambiente e do Mar, Dep. Biologia, Universidade de Aveiro, Campus Universitário de Santiago, 3810 - 193 Aveiro, Portugal. aravara @ ua. pt; http: // orcid. org / 0000 - 0002 - 1689 - 2985</mods:affiliation>
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<treatment id="03C687E2FFF0EE03F0CB57FFDB6A1DD4" ID-DOI="http://doi.org/10.5281/zenodo.4455620" ID-GBIF-Taxon="177014015" ID-Zenodo-Dep="4455620" LSID="urn:lsid:zoobank.org:act:4D214346-F738-43B2-AEE0-A6D82B53EDA8" httpUri="http://treatment.plazi.org/id/03C687E2FFF0EE03F0CB57FFDB6A1DD4" lastPageId="4" lastPageNumber="518" pageId="2" pageNumber="517">
<subSubSection id="C375657FFFF0EE05F0CB57FFD9EA1F18" box="[151,972,511,538]" pageId="2" pageNumber="517" type="nomenclature">
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<taxonomicName id="4C6F4D77FFF0EE05F0CB57FFD9401F18" ID-CoL="85RGT" authority="Sikorski, Langeneck &amp; Pavlova" authorityName="Sikorski, Langeneck &amp; Pavlova" authorityYear="2021" box="[151,870,511,538]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="2" pageNumber="517" phylum="Annelida" rank="species" species="alberti" status="sp. nov." subGenus="Sarsiana">
<emphasis id="B91BEAE6FFF0EE05F0CB57FFDADF1F18" bold="true" box="[151,249,511,537]" italics="true" pageId="2" pageNumber="517">Laonice</emphasis>
(
<emphasis id="B91BEAE6FFF0EE05F15657FFDB501F18" bold="true" box="[266,374,511,537]" italics="true" pageId="2" pageNumber="517">Sarsiana</emphasis>
)
<emphasis id="B91BEAE6FFF0EE05F1DB57FFDBF31F1B" bold="true" box="[391,469,511,538]" italics="true" pageId="2" pageNumber="517">alberti</emphasis>
Sikorski, Langeneck &amp; Pavlova
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(
<figureCitation id="13542A71FFF0EE05F0C35423DB2C1F3C" box="[159,266,547,573]" captionStart="FIGURE 1" captionStartId="3.[151,250,1232,1256]" captionTargetBox="[387,1200,181,1208]" captionTargetId="figure-18@3.[387,1200,181,1208]" captionTargetPageId="3" captionText="FIGURE 1. Laonice (Sarsiana) alberti sp. nov. Anterior end, dorsal view. Holotype MOM INV-0022694. Scale 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4447216" httpUri="https://zenodo.org/record/4447216/files/figure.png" pageId="2" pageNumber="517">Figures 1</figureCitation>
,
<figureCitation id="13542A71FFF0EE05F1445423DB031F3C" box="[280,293,547,573]" captionStart="FIGURE 2" captionStartId="4.[151,250,1538,1562]" captionTargetBox="[157,1430,244,1506]" captionTargetId="figure-49@4.[151,1436,237,1514]" captionTargetPageId="4" captionText="FIGURE 2. Laonice (Sarsiana) alberti sp. nov. A. Anterior end, dorsal view. B. Anterior end, side view. C. Chaetigers 47–50, dorsal view. D. Right parapodium, chaetiger 3, anterior view. E. Right parapodium, chaetiger 12, anterior view. F. Right parapodium, chaetiger 50, anterior view. G. Neuropodial hooded hook, chaetiger 50, lateral view. H. Neuropodial hooded hook, chaetiger 50, three-quarter view. Material: A, D–F, H—paratype (MOM INV-0022695); B–C, G—holotype MOM INV-0022694. Scales (mm): A–C = 0.5; D–F = 0.4; G–H = 0.0165." figureDoi="http://doi.org/10.5281/zenodo.4447220" httpUri="https://zenodo.org/record/4447220/files/figure.png" pageId="2" pageNumber="517">2</figureCitation>
A–H, 4)
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LSID:
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urn:lsid:zoobank.org:act:
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<treatmentCitationGroup id="AB7F11DAFFF0EE05F0CB548CD8671FA5" box="[151,577,652,676]" pageId="2" pageNumber="517">
<taxonomicName id="4C6F4D77FFF0EE05F0CB548CDACF1FA2" ID-CoL="84PJQ" authorityName="Malmgren" authorityYear="1867" box="[151,233,652,675]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="2" pageNumber="517" phylum="Annelida" rank="genus">
<emphasis id="B91BEAE6FFF0EE05F0CB548CDACF1FA2" box="[151,233,652,675]" italics="true" pageId="2" pageNumber="517">Laonice</emphasis>
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sp.—
<treatmentCitation id="0ACE10E5FFF0EE05F17E548CD81B1FA5" author="Langeneck, J. &amp; Busoni, G. &amp; Aliani, S. &amp; Castelli, A." box="[290,573,652,676]" page="143" pageId="2" pageNumber="517" year="2017">
<bibRefCitation id="EFFE4B05FFF0EE05F17E548CD81B1FA5" author="Langeneck, J. &amp; Busoni, G. &amp; Aliani, S. &amp; Castelli, A." box="[290,573,652,676]" pageId="2" pageNumber="517" pagination="142 - 152" refId="ref7585" refString="Langeneck, J., Busoni, G., Aliani, S. &amp; Castelli, A. (2017) Deep-sea polychaetes (Annelida) from the Malta Escarpment (western Ionian Sea). The European Zoological Journal, 84 (1), 142 - 152. https: // doi. org / 10.1080 / 24750263.2017.1287964" type="journal article" year="2017">
Langeneck
<emphasis id="B91BEAE6FFF0EE05F1C7548CDBE11FA5" box="[411,455,652,676]" italics="true" pageId="2" pageNumber="517">et al</emphasis>
. 2017: 143
</bibRefCitation>
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.
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</paragraph>
<paragraph id="8BD036F4FFF0EE05F0CB54ABD89A1FC3" blockId="2.[151,700,652,706]" box="[151,700,682,706]" pageId="2" pageNumber="517">
<taxonomicName id="4C6F4D77FFF0EE05F0CB54ABDB7C1FC3" ID-CoL="85KJH" authority="" authorityName="Sikorski, Gunton &amp; Pavlova" authorityYear="2017" box="[151,346,683,706]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="2" pageNumber="517" phylum="Annelida" rank="subGenus" subGenus="Sarsiana">
<emphasis id="B91BEAE6FFF0EE05F0CB54ABDB7C1FC3" box="[151,346,683,706]" italics="true" pageId="2" pageNumber="517">Laonice (Sarsiana)</emphasis>
</taxonomicName>
sp. A—Langeneck
<emphasis id="B91BEAE6FFF0EE05F27F54ABD8761FC3" box="[547,592,682,706]" italics="true" pageId="2" pageNumber="517">et al</emphasis>
. 2019: 87.
</paragraph>
</subSubSection>
<subSubSection id="C375657FFFF0EE05F0CB54F0DBCB192A" pageId="2" pageNumber="517" type="materials_examined">
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<emphasis id="B91BEAE6FFF0EE05F0CB54F0DBA01E0B" bold="true" box="[151,390,752,778]" pageId="2" pageNumber="517">Material examined.</emphasis>
<materialsCitation id="3B073CA9FFF0EE05F1CD54F0DFBF1E2E" ID-GBIF-Occurrence="3023040301" collectingDate="2009-05-19" collectionCode="MOM, R, V" country="Malta" elevation="1200" latitude="36.531" location="Escarpment" longLatPrecision="6" longitude="15.434" municipality="Central Mediterranean Sea" pageId="2" pageNumber="517" specimenCode="INV-0022694, INV-0022695" specimenCount="2" typeStatus="holotype">
<typeStatus id="54D48856FFF0EE05F1CD54F0DBDF1E0B" box="[401,505,752,778]" pageId="2" pageNumber="517" type="holotype">Holotype</typeStatus>
(
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) and
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(
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<specimenCode id="DBC99E8FFFF0EE05F45054F0DE8B1E0A" box="[1036,1197,752,779]" collectionCode="INV-" pageId="2" pageNumber="517">INV-0022695</specimenCode>
):
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,
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,
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/
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, Project BIOFUN, St.
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1,
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,
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, 8–
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</date>
</materialsCitation>
.
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(
<specimenCode id="DBC99E8FFFF0EE05F1555539DB811E52" box="[265,423,824,851]" collectionCode="MNCN" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35275" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="2" pageNumber="517" type="Museum">MNCN 16.01</specimenCode>
/18547):
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slope,
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,
<collectionCode id="ED7EAE31FFF0EE05F3845539D9CC1E52" box="[984,1002,825,851]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="517">R</collectionCode>
/
<collectionCode id="ED7EAE31FFF0EE05F3AF5539DE201E52" box="[1011,1030,825,851]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="517" type="Museum">V</collectionCode>
<taxonomicName id="4C6F4D77FFF0EE05F4535539DF7B1E52" authority=", Project BIOFUN, St." authorityName="Project BIOFUN, St." box="[1039,1373,824,851]" class="Insecta" family="Uraniidae" genus="Urania" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAE6FFF0EE05F4535539DE461E53" box="[1039,1120,825,850]" italics="true" pageId="2" pageNumber="517">Urania</emphasis>
, Project BIOFUN, St.
</taxonomicName>
28 B 2400,
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<geoCoordinate id="EE5B5033FFF0EE05F13C555CDBEA1E77" box="[352,460,860,886]" degrees="3.6469" direction="east" orientation="longitude" pageId="2" pageNumber="517" precision="5" value="3.6469">3.6469°E</geoCoordinate>
,
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</quantity>
, clayish silt,
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<collectingDate id="EF95E9DCFFF0EE05F298555CD9631E76" box="[708,837,860,887]" pageId="2" pageNumber="517" value="2010-07-05">07.05.2010</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B073CA9FFF0EE05F30C555DD8F41EBE" ID-GBIF-Occurrence="3023040307" collectingDate="2012-10-10" collectionCode="R, V, DOS, MARES" collectorName="Blanes &amp; NW Mediterranean Sea" country="Malta" elevation="1200" latitude="41.5055" location="Garca del Cid" longLatPrecision="6" longitude="2.85" municipality="Cruise" pageId="2" pageNumber="517" specimenCode="DBUA0002347.01, DM3" specimenCount="1" typeStatus="paratype">
<typeStatus id="54D48856FFF0EE05F30C555DD9941E76" box="[848,946,861,887]" pageId="2" pageNumber="517" type="paratype">Paratype</typeStatus>
(
<specimenCode id="DBC99E8FFFF0EE05F39D555DDEB81E76" box="[961,1182,861,887]" collectionCode="DBUA" country="Portugal" httpUri="http://grbio.org/cool/p469-157j" name="Zoological Collection of the Biology Department, University of the Azores" pageId="2" pageNumber="517">DBUA0002347.01</specimenCode>
): open slope of
<collectorName id="269A5322FFF0EE05F50C555CDFBA1E77" box="[1360,1436,860,886]" pageId="2" pageNumber="517">Blanes</collectorName>
submarine canyon,
<collectorName id="269A5322FFF0EE05F12A5581D8A31E9A" box="[374,645,896,923]" pageId="2" pageNumber="517">NW Mediterranean Sea</collectorName>
,
<collectionCode id="ED7EAE31FFF0EE05F2CF5581D8801E9A" box="[659,678,897,923]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="517">R</collectionCode>
/
<collectionCode id="ED7EAE31FFF0EE05F2F25581D8E41E9A" box="[686,706,897,923]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="517" type="Museum">V</collectionCode>
<emphasis id="B91BEAE6FFF0EE05F2945581D94D1E9B" box="[712,875,896,923]" italics="true" pageId="2" pageNumber="517">
<location id="8EB0602FFFF0EE05F2945581D94D1E9B" LSID="urn:lsid:plazi:treatment:03C687E2FFF0EE03F0CB57FFDB6A1DD4:8EB0602FFFF0EE05F2945581D94D1E9B" box="[712,875,896,923]" country="Malta" latitude="41.5055" longLatPrecision="6" longitude="2.85" municipality="Cruise" name="Garca del Cid" pageId="2" pageNumber="517">Garca del Cid</location>
</emphasis>
, Project
<collectionCode id="ED7EAE31FFF0EE05F3895581DE5F1E9A" box="[981,1145,897,923]" pageId="2" pageNumber="517">DOS MARES</collectionCode>
,
<collectingMunicipality id="6BB4AC8EFFF0EE05F4DB5580DEF61E9B" box="[1159,1232,896,922]" pageId="2" pageNumber="517">Cruise</collectingMunicipality>
<specimenCode id="DBC99E8FFFF0EE05F4855581DF311E9A" box="[1241,1303,897,923]" collectionCode="DM" country="New Zealand" name="Dominion Museum" pageId="2" pageNumber="517">DM3</specimenCode>
, trawl code
<specimenCode id="DBC99E8FFFF0EE05F0CB55A5DAF21EBE" box="[151,212,933,959]" collectionCode="DM" country="New Zealand" name="Dominion Museum" pageId="2" pageNumber="517">DM3</specimenCode>
P2,
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<geoCoordinate id="EE5B5033FFF0EE05F12855A4DBC51EBE" box="[372,483,932,959]" degrees="02.8500" direction="east" orientation="longitude" pageId="2" pageNumber="517" precision="5" value="2.85">02.8500E</geoCoordinate>
,
<quantity id="4C979B11FFF0EE05F1B255A5D8631EBE" box="[494,581,933,959]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="517" unit="m" value="1200.0">
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</quantity>
,
<date id="FFD11034FFF0EE05F20D55A5D8F41EBE" box="[593,722,933,959]" pageId="2" pageNumber="517" value="2012-10-10">
<collectingDate id="EF95E9DCFFF0EE05F20D55A5D8F41EBE" box="[593,722,933,959]" pageId="2" pageNumber="517" value="2012-10-10">10.10.2012</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BD036F4FFF0EE05F09B55C8DBCB192A" blockId="2.[151,1437,752,2039]" pageId="2" pageNumber="517">
<materialsCitation id="3B073CA9FFF0EE05F09B55C8DBCF192A" ID-GBIF-Occurrence="3023040305" collectingDate="2006-06-05" collectionCode="R, V" country="Morocco" county="Northeast Atlantic Ocean" elevation="350" latitude="35.2978" location="Mercator" longLatPrecision="6" longitude="-6.6468" municipality="Gulf of Cadiz" pageId="2" pageNumber="517" specimenCode="DBUA0002235.03" specimenCount="1">
Additional non
<typeStatus id="54D48856FFF0EE05F12B55C9DB811EE2" box="[375,423,969,995]" pageId="2" pageNumber="517">type</typeStatus>
material, possibly belonging to the same species (
<specimenCode id="DBC99E8FFFF0EE05F38655C8DE901EE2" box="[986,1206,968,995]" collectionCode="DBUA" country="Portugal" httpUri="http://grbio.org/cool/p469-157j" name="Zoological Collection of the Biology Department, University of the Azores" pageId="2" pageNumber="517">DBUA0002235.03</specimenCode>
):
<location id="8EB0602FFFF0EE05F49455C9DF091EE2" LSID="urn:lsid:plazi:treatment:03C687E2FFF0EE03F0CB57FFDB6A1DD4:8EB0602FFFF0EE05F49455C9DF091EE2" box="[1224,1327,969,995]" country="Morocco" county="Northeast Atlantic Ocean" latitude="35.2978" longLatPrecision="6" longitude="-6.6468" municipality="Gulf of Cadiz" name="Mercator" pageId="2" pageNumber="517">Mercator</location>
mud volcano,
<collectingMunicipality id="6BB4AC8EFFF0EE05F08555ECDB521907" box="[217,372,1004,1030]" pageId="2" pageNumber="517">Gulf of Cadiz</collectingMunicipality>
, off
<collectingCountry id="F3787664FFF0EE05F1F955EDD82B1906" box="[421,525,1005,1031]" name="Morocco" pageId="2" pageNumber="517">Morocco</collectingCountry>
,
<collectingCounty id="62B14E78FFF0EE05F24B55ECD9151906" box="[535,819,1004,1031]" pageId="2" pageNumber="517">Northeast Atlantic Ocean</collectingCounty>
,
<collectionCode id="ED7EAE31FFF0EE05F36255EDD9761906" box="[830,848,1005,1031]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="517">R</collectionCode>
/
<collectionCode id="ED7EAE31FFF0EE05F30455EDD94D1906" box="[856,875,1005,1031]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="517" type="Museum">V</collectionCode>
<taxonomicName id="4C6F4D77FFF0EE05F32E55EDDE591906" authority="S. Merian, MSM" authorityName="S. Merian, MSM" box="[882,1151,1005,1031]" class="Insecta" family="Asilidae" genus="Maria" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="B91BEAE6FFF0EE05F32E55EDDE0D1907" box="[882,1067,1005,1031]" italics="true" pageId="2" pageNumber="517">Maria S. Merian</emphasis>
, MSM
</taxonomicName>
01-03, St. 242,
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<geoCoordinate id="EE5B5033FFF0EE05F0CB5210DB2E192B" box="[151,264,1040,1066]" degrees="6.6468" direction="west" orientation="longitude" pageId="2" pageNumber="517" precision="5" value="-6.6468">6.6468ºW</geoCoordinate>
,
<quantity id="4C979B11FFF0EE05F14F5210DB7A192A" box="[275,348,1040,1067]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="517" unit="m" value="350.0">
<elevation id="0042D1C7FFF0EE05F14F5210DB7A192A" box="[275,348,1040,1067]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="517" unit="m" value="350.0">350 m</elevation>
</quantity>
,
<date id="FFD11034FFF0EE05F1345210DBCF192A" box="[360,489,1040,1067]" pageId="2" pageNumber="517" value="2006-06-05">
<collectingDate id="EF95E9DCFFF0EE05F1345210DBCF192A" box="[360,489,1040,1067]" pageId="2" pageNumber="517" value="2006-06-05">06.05.2006</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C375657FFFF0EE04F09B5234D98B1879" lastPageId="3" lastPageNumber="518" pageId="2" pageNumber="517" type="description">
<paragraph id="8BD036F4FFF0EE05F09B5234DB2E1973" blockId="2.[151,1437,752,2039]" pageId="2" pageNumber="517">
<emphasis id="B91BEAE6FFF0EE05F09B5234DB7F194F" bold="true" box="[199,345,1076,1102]" pageId="2" pageNumber="517">Description.</emphasis>
All specimens incomplete (50 chaetigers in
<typeStatus id="54D48856FFF0EE05F3055234D999194E" box="[857,959,1076,1103]" pageId="2" pageNumber="517" type="holotype">holotype</typeStatus>
), consisting of anterior fragments
<quantity id="4C979B11FFF0EE05F5145235DAE51972" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="2" pageNumber="517" unit="mm" value="0.85" valueMax="0.9" valueMin="0.8">0.8–0.9 mm</quantity>
wide.
</paragraph>
<paragraph id="8BD036F4FFF0EE05F09B527CD8F81893" blockId="2.[151,1437,752,2039]" pageId="2" pageNumber="517">
Prostomium longer than wide, triangular (
<typeStatus id="54D48856FFF0EE05F2FE527CD9211996" box="[674,775,1148,1175]" pageId="2" pageNumber="517" type="holotype">holotype</typeStatus>
) to bell-shaped (
<typeStatus id="54D48856FFF0EE05F399527DDE011996" box="[965,1063,1149,1175]" pageId="2" pageNumber="517" type="paratype">paratype</typeStatus>
MOM INV-0022695), with very small apical incision (
<figureCitation id="13542A71FFF0EE05F1D152A0DBF219BA" box="[397,468,1184,1211]" captionStart="FIGURE 1" captionStartId="3.[151,250,1232,1256]" captionTargetBox="[387,1200,181,1208]" captionTargetId="figure-18@3.[387,1200,181,1208]" captionTargetPageId="3" captionText="FIGURE 1. Laonice (Sarsiana) alberti sp. nov. Anterior end, dorsal view. Holotype MOM INV-0022694. Scale 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4447216" httpUri="https://zenodo.org/record/4447216/files/figure.png" pageId="2" pageNumber="517">Figs 1</figureCitation>
and
<figureCitation id="13542A71FFF0EE05F25752A1D81719BA" box="[523,561,1185,1211]" captionStart="FIGURE 2" captionStartId="4.[151,250,1538,1562]" captionTargetBox="[157,1430,244,1506]" captionTargetId="figure-49@4.[151,1436,237,1514]" captionTargetPageId="4" captionText="FIGURE 2. Laonice (Sarsiana) alberti sp. nov. A. Anterior end, dorsal view. B. Anterior end, side view. C. Chaetigers 47–50, dorsal view. D. Right parapodium, chaetiger 3, anterior view. E. Right parapodium, chaetiger 12, anterior view. F. Right parapodium, chaetiger 50, anterior view. G. Neuropodial hooded hook, chaetiger 50, lateral view. H. Neuropodial hooded hook, chaetiger 50, three-quarter view. Material: A, D–F, H—paratype (MOM INV-0022695); B–C, G—holotype MOM INV-0022694. Scales (mm): A–C = 0.5; D–F = 0.4; G–H = 0.0165." figureDoi="http://doi.org/10.5281/zenodo.4447220" httpUri="https://zenodo.org/record/4447220/files/figure.png" pageId="2" pageNumber="517">2A</figureCitation>
); antero-lateral corners connecting to peristomium by very thin and nearly inconspicuous ventral folds hidden in the groove between prostomium and peristomium (
<figureCitation id="13542A71FFF0EE05F43D52C4DE8119DE" box="[1121,1191,1220,1247]" captionStart="FIGURE 1" captionStartId="3.[151,250,1232,1256]" captionTargetBox="[387,1200,181,1208]" captionTargetId="figure-18@3.[387,1200,181,1208]" captionTargetPageId="3" captionText="FIGURE 1. Laonice (Sarsiana) alberti sp. nov. Anterior end, dorsal view. Holotype MOM INV-0022694. Scale 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4447216" httpUri="https://zenodo.org/record/4447216/files/figure.png" pageId="2" pageNumber="517">Figs 1</figureCitation>
and
<figureCitation id="13542A71FFF0EE05F48052C5DF2619DE" box="[1244,1280,1221,1247]" captionStart="FIGURE 2" captionStartId="4.[151,250,1538,1562]" captionTargetBox="[157,1430,244,1506]" captionTargetId="figure-49@4.[151,1436,237,1514]" captionTargetPageId="4" captionText="FIGURE 2. Laonice (Sarsiana) alberti sp. nov. A. Anterior end, dorsal view. B. Anterior end, side view. C. Chaetigers 47–50, dorsal view. D. Right parapodium, chaetiger 3, anterior view. E. Right parapodium, chaetiger 12, anterior view. F. Right parapodium, chaetiger 50, anterior view. G. Neuropodial hooded hook, chaetiger 50, lateral view. H. Neuropodial hooded hook, chaetiger 50, three-quarter view. Material: A, D–F, H—paratype (MOM INV-0022695); B–C, G—holotype MOM INV-0022694. Scales (mm): A–C = 0.5; D–F = 0.4; G–H = 0.0165." figureDoi="http://doi.org/10.5281/zenodo.4447220" httpUri="https://zenodo.org/record/4447220/files/figure.png" pageId="2" pageNumber="517">2B</figureCitation>
). Prostomium narrowing backwards at first gradually until the fore level of the first pair of parapodia, and then abruptly, merging posteriorly with the caruncle. One pair of faint eyespots (
<typeStatus id="54D48856FFF0EE05F377530DD9AB1826" box="[811,909,1293,1319]" pageId="2" pageNumber="517" type="paratype">paratype</typeStatus>
MOM INV-0022695). Occipital antenna very short, erect and finger-like, inserted at the end of the prostomium, just at the rear level of the first pair of parapodia. Palps lost from all specimens. Nuchal organs as double U-shaped ciliary bands extending posteriorly to chaetigers 5–7 (
<figureCitation id="13542A71FFF0EE05F08D5378DB0D1892" box="[209,299,1400,1427]" captionStart="FIGURE 2" captionStartId="4.[151,250,1538,1562]" captionTargetBox="[157,1430,244,1506]" captionTargetId="figure-49@4.[151,1436,237,1514]" captionTargetPageId="4" captionText="FIGURE 2. Laonice (Sarsiana) alberti sp. nov. A. Anterior end, dorsal view. B. Anterior end, side view. C. Chaetigers 47–50, dorsal view. D. Right parapodium, chaetiger 3, anterior view. E. Right parapodium, chaetiger 12, anterior view. F. Right parapodium, chaetiger 50, anterior view. G. Neuropodial hooded hook, chaetiger 50, lateral view. H. Neuropodial hooded hook, chaetiger 50, three-quarter view. Material: A, D–F, H—paratype (MOM INV-0022695); B–C, G—holotype MOM INV-0022694. Scales (mm): A–C = 0.5; D–F = 0.4; G–H = 0.0165." figureDoi="http://doi.org/10.5281/zenodo.4447220" httpUri="https://zenodo.org/record/4447220/files/figure.png" pageId="2" pageNumber="517">Fig. 2A</figureCitation>
). Body width uniform along its length.
</paragraph>
<paragraph id="8BD036F4FFF0EE05F09B539CDE5C1B23" blockId="2.[151,1437,752,2039]" pageId="2" pageNumber="517">
Branchiae from chaetiger 2, continuing posteriorly to the end of fragments (chaetiger
<quantity id="4C979B11FFF0EE05F421539DDE9318B7" box="[1149,1205,1436,1463]" metricMagnitude="-1" metricUnit="m" metricValue="9.652" pageId="2" pageNumber="517" unit="in" value="38.0">38 in</quantity>
<typeStatus id="54D48856FFF0EE05F4E7539CDF0718B7" box="[1211,1313,1436,1462]" pageId="2" pageNumber="517" type="holotype">holotype</typeStatus>
; chaetigers
<quantity id="4C979B11FFF0EE05F0CB53C1DADF18DB" box="[151,249,1472,1499]" metricMagnitude="-1" metricUnit="m" metricValue="8.89" metricValueMax="9.906" metricValueMin="7.874" pageId="2" pageNumber="517" unit="in" value="35.0" valueMax="39.0" valueMin="31.0">31–39 in</quantity>
<typeStatus id="54D48856FFF0EE05F0A353C1DB5418DA" box="[255,370,1473,1499]" pageId="2" pageNumber="517" type="paratype">paratypes</typeStatus>
,). Branchiae free, initially slightly longer than the notopodial postchaetal lamellae, increasing its length significantly just after the end of the nuchal organs and becoming twice as long as the notopodial postchaetal lamellae after chaetiger 10–12. Length of the longest branchiae up to half of body width.
</paragraph>
<paragraph id="8BD036F4FFF0EE05F09B502CDB7F1BB2" blockId="2.[151,1437,752,2039]" pageId="2" pageNumber="517">
Notopodial prechaetal lamellae not particularly conspicuous. Longest notopodial postchaetal lamellae at chaetigers 5–12, gradually diminishing posteriorly. Notopodial postchaetal lamellae of the first 3 segments with moderately pronounced upper tips (
<figureCitation id="13542A71FFF0EE05F1A45074D8751B8E" box="[504,595,1652,1679]" captionStart="FIGURE 2" captionStartId="4.[151,250,1538,1562]" captionTargetBox="[157,1430,244,1506]" captionTargetId="figure-49@4.[151,1436,237,1514]" captionTargetPageId="4" captionText="FIGURE 2. Laonice (Sarsiana) alberti sp. nov. A. Anterior end, dorsal view. B. Anterior end, side view. C. Chaetigers 47–50, dorsal view. D. Right parapodium, chaetiger 3, anterior view. E. Right parapodium, chaetiger 12, anterior view. F. Right parapodium, chaetiger 50, anterior view. G. Neuropodial hooded hook, chaetiger 50, lateral view. H. Neuropodial hooded hook, chaetiger 50, three-quarter view. Material: A, D–F, H—paratype (MOM INV-0022695); B–C, G—holotype MOM INV-0022694. Scales (mm): A–C = 0.5; D–F = 0.4; G–H = 0.0165." figureDoi="http://doi.org/10.5281/zenodo.4447220" httpUri="https://zenodo.org/record/4447220/files/figure.png" pageId="2" pageNumber="517">Fig. 2D</figureCitation>
), upper margins of all subsequent notopodial postchaetal lamellae broadly oval (
<figureCitation id="13542A71FFF0EE05F08B5098DB3F1BB2" box="[215,281,1688,1715]" captionStart="FIGURE 2" captionStartId="4.[151,250,1538,1562]" captionTargetBox="[157,1430,244,1506]" captionTargetId="figure-49@4.[151,1436,237,1514]" captionTargetPageId="4" captionText="FIGURE 2. Laonice (Sarsiana) alberti sp. nov. A. Anterior end, dorsal view. B. Anterior end, side view. C. Chaetigers 47–50, dorsal view. D. Right parapodium, chaetiger 3, anterior view. E. Right parapodium, chaetiger 12, anterior view. F. Right parapodium, chaetiger 50, anterior view. G. Neuropodial hooded hook, chaetiger 50, lateral view. H. Neuropodial hooded hook, chaetiger 50, three-quarter view. Material: A, D–F, H—paratype (MOM INV-0022695); B–C, G—holotype MOM INV-0022694. Scales (mm): A–C = 0.5; D–F = 0.4; G–H = 0.0165." figureDoi="http://doi.org/10.5281/zenodo.4447220" httpUri="https://zenodo.org/record/4447220/files/figure.png" pageId="2" pageNumber="517">Fig. 2</figureCitation>
E–F).
</paragraph>
<paragraph id="8BD036F4FFF0EE05F09B50BCDB001BFA" blockId="2.[151,1437,752,2039]" pageId="2" pageNumber="517">
Neuropodial prechaetal lamellae not visible. Neuropodial postchaetal lamellae triangular throughout the body (
<figureCitation id="13542A71FFF0EE05F0FC50E0DAC41BFA" box="[160,226,1760,1787]" captionStart="FIGURE 2" captionStartId="4.[151,250,1538,1562]" captionTargetBox="[157,1430,244,1506]" captionTargetId="figure-49@4.[151,1436,237,1514]" captionTargetPageId="4" captionText="FIGURE 2. Laonice (Sarsiana) alberti sp. nov. A. Anterior end, dorsal view. B. Anterior end, side view. C. Chaetigers 47–50, dorsal view. D. Right parapodium, chaetiger 3, anterior view. E. Right parapodium, chaetiger 12, anterior view. F. Right parapodium, chaetiger 50, anterior view. G. Neuropodial hooded hook, chaetiger 50, lateral view. H. Neuropodial hooded hook, chaetiger 50, three-quarter view. Material: A, D–F, H—paratype (MOM INV-0022695); B–C, G—holotype MOM INV-0022694. Scales (mm): A–C = 0.5; D–F = 0.4; G–H = 0.0165." figureDoi="http://doi.org/10.5281/zenodo.4447220" httpUri="https://zenodo.org/record/4447220/files/figure.png" pageId="2" pageNumber="517">Fig. 2</figureCitation>
D–E).
</paragraph>
<paragraph id="8BD036F4FFF0EE05F09B5104DF341A42" blockId="2.[151,1437,752,2039]" pageId="2" pageNumber="517">
Continuous transverse dorsal crests absent in midbody segments, where the notopodial postchaetal lamellae are fused with quite long crests on the dorsal surface, however without connecting mid-dorsally (
<figureCitation id="13542A71FFF0EE05F4F15128DF201A42" box="[1197,1286,1832,1859]" captionStart="FIGURE 2" captionStartId="4.[151,250,1538,1562]" captionTargetBox="[157,1430,244,1506]" captionTargetId="figure-49@4.[151,1436,237,1514]" captionTargetPageId="4" captionText="FIGURE 2. Laonice (Sarsiana) alberti sp. nov. A. Anterior end, dorsal view. B. Anterior end, side view. C. Chaetigers 47–50, dorsal view. D. Right parapodium, chaetiger 3, anterior view. E. Right parapodium, chaetiger 12, anterior view. F. Right parapodium, chaetiger 50, anterior view. G. Neuropodial hooded hook, chaetiger 50, lateral view. H. Neuropodial hooded hook, chaetiger 50, three-quarter view. Material: A, D–F, H—paratype (MOM INV-0022695); B–C, G—holotype MOM INV-0022694. Scales (mm): A–C = 0.5; D–F = 0.4; G–H = 0.0165." figureDoi="http://doi.org/10.5281/zenodo.4447220" httpUri="https://zenodo.org/record/4447220/files/figure.png" pageId="2" pageNumber="517">Fig. 2C</figureCitation>
).
</paragraph>
<paragraph id="8BD036F4FFF0EE05F09B514CDF251A67" blockId="2.[151,1437,752,2039]" box="[199,1283,1868,1895]" pageId="2" pageNumber="517">Inter-parapodial pouches present from chaetigers 21–34, until the posterior end of the fragments.</paragraph>
<paragraph id="8BD036F4FFF0EE05F09B5170D9BD1AF6" blockId="2.[151,1437,752,2039]" pageId="2" pageNumber="517">
Anterior parapodia with capillary chaetae arranged in two vertical rows both in notopodia and neuropodia. Neuropodial hooded hooks appearing from chaetiger 35–38, numbering 8–10 per bundle, appearing bidentate in the lateral view (
<figureCitation id="13542A71FFF0EE05F17651B8DBA21AD2" box="[298,388,1976,2003]" captionStart="FIGURE 2" captionStartId="4.[151,250,1538,1562]" captionTargetBox="[157,1430,244,1506]" captionTargetId="figure-49@4.[151,1436,237,1514]" captionTargetPageId="4" captionText="FIGURE 2. Laonice (Sarsiana) alberti sp. nov. A. Anterior end, dorsal view. B. Anterior end, side view. C. Chaetigers 47–50, dorsal view. D. Right parapodium, chaetiger 3, anterior view. E. Right parapodium, chaetiger 12, anterior view. F. Right parapodium, chaetiger 50, anterior view. G. Neuropodial hooded hook, chaetiger 50, lateral view. H. Neuropodial hooded hook, chaetiger 50, three-quarter view. Material: A, D–F, H—paratype (MOM INV-0022695); B–C, G—holotype MOM INV-0022694. Scales (mm): A–C = 0.5; D–F = 0.4; G–H = 0.0165." figureDoi="http://doi.org/10.5281/zenodo.4447220" httpUri="https://zenodo.org/record/4447220/files/figure.png" pageId="2" pageNumber="517">Fig. 2G</figureCitation>
), but with paired apical teeth above main fang (
<figureCitation id="13542A71FFF0EE05F3C651B8D9D31AD2" box="[922,1013,1976,2003]" captionStart="FIGURE 2" captionStartId="4.[151,250,1538,1562]" captionTargetBox="[157,1430,244,1506]" captionTargetId="figure-49@4.[151,1436,237,1514]" captionTargetPageId="4" captionText="FIGURE 2. Laonice (Sarsiana) alberti sp. nov. A. Anterior end, dorsal view. B. Anterior end, side view. C. Chaetigers 47–50, dorsal view. D. Right parapodium, chaetiger 3, anterior view. E. Right parapodium, chaetiger 12, anterior view. F. Right parapodium, chaetiger 50, anterior view. G. Neuropodial hooded hook, chaetiger 50, lateral view. H. Neuropodial hooded hook, chaetiger 50, three-quarter view. Material: A, D–F, H—paratype (MOM INV-0022695); B–C, G—holotype MOM INV-0022694. Scales (mm): A–C = 0.5; D–F = 0.4; G–H = 0.0165." figureDoi="http://doi.org/10.5281/zenodo.4447220" httpUri="https://zenodo.org/record/4447220/files/figure.png" pageId="2" pageNumber="517">Fig. 2H</figureCitation>
). Single sabre chaeta appearing in the lower part of the neuropodial bundle from chaetiger 20–22 (
<figureCitation id="13542A71FFF0EE05F36451DCD9A81AF6" box="[824,910,2012,2039]" captionStart="FIGURE 2" captionStartId="4.[151,250,1538,1562]" captionTargetBox="[157,1430,244,1506]" captionTargetId="figure-49@4.[151,1436,237,1514]" captionTargetPageId="4" captionText="FIGURE 2. Laonice (Sarsiana) alberti sp. nov. A. Anterior end, dorsal view. B. Anterior end, side view. C. Chaetigers 47–50, dorsal view. D. Right parapodium, chaetiger 3, anterior view. E. Right parapodium, chaetiger 12, anterior view. F. Right parapodium, chaetiger 50, anterior view. G. Neuropodial hooded hook, chaetiger 50, lateral view. H. Neuropodial hooded hook, chaetiger 50, three-quarter view. Material: A, D–F, H—paratype (MOM INV-0022695); B–C, G—holotype MOM INV-0022694. Scales (mm): A–C = 0.5; D–F = 0.4; G–H = 0.0165." figureDoi="http://doi.org/10.5281/zenodo.4447220" httpUri="https://zenodo.org/record/4447220/files/figure.png" pageId="2" pageNumber="517">Fig. 2F</figureCitation>
).
</paragraph>
<caption id="DF10667CFFF1EE04F0CB52D0DF4319E9" ID-DOI="http://doi.org/10.5281/zenodo.4447216" ID-Zenodo-Dep="4447216" box="[151,1381,1232,1256]" httpUri="https://zenodo.org/record/4447216/files/figure.png" pageId="3" pageNumber="518" startId="3.[151,250,1232,1256]" targetBox="[387,1200,181,1208]" targetPageId="3">
<paragraph id="8BD036F4FFF1EE04F0CB52D0DF4319E9" blockId="3.[151,1381,1232,1256]" box="[151,1381,1232,1256]" pageId="3" pageNumber="518">
<emphasis id="B91BEAE6FFF1EE04F0CB52D0DB3519E9" bold="true" box="[151,275,1232,1256]" pageId="3" pageNumber="518">FIGURE 1.</emphasis>
<taxonomicName id="4C6F4D77FFF1EE04F14552D1D80119E9" authority="Sikorski &amp; Pavlova &amp; Sardá &amp; Langeneck &amp; Gil &amp; Ravara, 2021" authorityName="Sikorski &amp; Pavlova &amp; Sardá &amp; Langeneck &amp; Gil &amp; Ravara" authorityYear="2021" box="[281,551,1232,1256]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="3" pageNumber="518" phylum="Annelida" rank="species" species="alberti" status="sp. nov." subGenus="Sarsiana">
<emphasis id="B91BEAE6FFF1EE04F14552D1DB4D19E9" box="[281,363,1233,1256]" italics="true" pageId="3" pageNumber="518">Laonice</emphasis>
(
<emphasis id="B91BEAE6FFF1EE04F12652D1DBF219E9" box="[378,468,1233,1256]" italics="true" pageId="3" pageNumber="518">Sarsiana</emphasis>
)
<emphasis id="B91BEAE6FFF1EE04F1BF52D0D80119E9" box="[483,551,1232,1256]" italics="true" pageId="3" pageNumber="518">alberti</emphasis>
</taxonomicName>
<emphasis id="B91BEAE6FFF1EE04F27152D0D85B19E9" bold="true" box="[557,637,1232,1256]" pageId="3" pageNumber="518">
<taxonomicNameLabel id="A228579DFFF1EE04F27152D0D85B19E9" box="[557,637,1232,1256]" pageId="3" pageNumber="518" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Anterior end, dorsal view. Holotype MOM INV-0022694. Scale 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BD036F4FFF1EE04F09B5316DB8D1831" blockId="3.[151,1437,1302,2013]" box="[199,427,1302,1328]" pageId="3" pageNumber="518">Pygidium unknown.</paragraph>
<paragraph id="8BD036F4FFF1EE04F09B533BD8701855" blockId="3.[151,1437,1302,2013]" box="[199,598,1338,1365]" pageId="3" pageNumber="518">No specific pigmentation observed.</paragraph>
<paragraph id="8BD036F4FFF1EE04F09B535ED98B1879" blockId="3.[151,1437,1302,2013]" box="[199,941,1374,1401]" pageId="3" pageNumber="518">
<emphasis id="B91BEAE6FFF1EE04F09B535EDBD81879" bold="true" box="[199,510,1374,1401]" pageId="3" pageNumber="518">Staining in Methyl Green.</emphasis>
No specific staining pattern observed.
</paragraph>
</subSubSection>
<subSubSection id="C375657FFFF1EE04F09B5383D95818E5" pageId="3" pageNumber="518" type="etymology">
<paragraph id="8BD036F4FFF1EE04F09B5383D95818E5" blockId="3.[151,1437,1302,2013]" pageId="3" pageNumber="518">
<emphasis id="B91BEAE6FFF1EE04F09B5383DB68189C" bold="true" box="[199,334,1411,1437]" pageId="3" pageNumber="518">Etymology.</emphasis>
The species is dedicated to Professor Alberto Castelli (University of Pisa,
<collectingCountry id="F3787664FFF1EE04F4D25382DEE3189C" box="[1166,1221,1410,1437]" name="Italy" pageId="3" pageNumber="518">Italy</collectingCountry>
), who provided the material that enabled the establishment of the new species, in recognition of his numerous and important contributions to the knowledge of the Mediterranean fauna of polychaetes.
</paragraph>
</subSubSection>
<subSubSection id="C375657FFFF1EE04F09B53EEDEEB1B2C" pageId="3" pageNumber="518" type="distribution">
<paragraph id="8BD036F4FFF1EE04F09B53EEDEEB1B2C" blockId="3.[151,1437,1302,2013]" pageId="3" pageNumber="518">
<emphasis id="B91BEAE6FFF1EE04F09B53EEDB471B09" bold="true" box="[199,353,1518,1544]" pageId="3" pageNumber="518">Distribution.</emphasis>
Northeast Atlantic Ocean, Mediterranean Sea: Blanes submarine canyon, Balearic slope and
<collectingCountry id="F3787664FFF1EE04F53A53EEDA8D1B2C" name="Malta" pageId="3" pageNumber="518">Malta</collectingCountry>
escarpment, (
<figureCitation id="13542A71FFF1EE04F1145012DBA81B2C" box="[328,398,1554,1581]" captionStart="FIGURE 4" captionStartId="7.[151,250,1296,1320]" captionTargetBox="[155,1430,422,1270]" captionTargetId="figure-128@7.[151,1436,417,1273]" captionTargetPageId="7" captionText="FIGURE 4. Distributional map of the Mediterranean deep-sea Laonice species: circles, Laonice (Sarsiana) alberti sp. nov.; triangles, Laonice (Appelloefia) barcinensis sp. nov.; star, Laonice (Laonice) sp.; T, type localities." figureDoi="http://doi.org/10.5281/zenodo.4447226" httpUri="https://zenodo.org/record/4447226/files/figure.png" pageId="3" pageNumber="518">Fig. 4</figureCitation>
).
<quantity id="4C979B11FFF1EE04F1FE5012D8661B2C" box="[418,576,1554,1581]" metricMagnitude="3" metricUnit="m" metricValue="1.773" metricValueMax="2.346" metricValueMin="1.2" pageId="3" pageNumber="518" unit="m" value="1773.0" valueMax="2346.0" valueMin="1200.0">1200–2346 m</quantity>
. Probably: Gulf of Cadiz (Mercator mud volcano),
<specimenCount id="9D69FD7DFFF1EE04F4235012DEEB1B2C" box="[1151,1229,1554,1581]" pageId="3" pageNumber="518" type="generic">
<quantity id="4C979B11FFF1EE04F4235012DEEF1B2C" box="[1151,1225,1554,1581]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="518" unit="m" value="350.0">350 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="C375657FFFF1EE03F09B5037DB6A1DD4" lastPageId="4" lastPageNumber="519" pageId="3" pageNumber="518" type="discussion">
<paragraph id="8BD036F4FFF1EE03F09B5037DB6A1DD4" blockId="3.[151,1437,1302,2013]" lastBlockId="4.[151,1436,150,213]" lastPageId="4" lastPageNumber="519" pageId="3" pageNumber="518">
<emphasis id="B91BEAE6FFF1EE04F09B5037DB1D1B50" bold="true" box="[199,315,1591,1617]" pageId="3" pageNumber="518">Remarks.</emphasis>
<taxonomicName id="4C6F4D77FFF1EE04F11F5037D8561B51" authority="Sikorski &amp; Pavlova &amp; Sardá &amp; Langeneck &amp; Gil &amp; Ravara, 2021" authorityName="Sikorski &amp; Pavlova &amp; Sardá &amp; Langeneck &amp; Gil &amp; Ravara" authorityYear="2021" box="[323,624,1590,1617]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="3" pageNumber="518" phylum="Annelida" rank="species" species="alberti" status="sp. nov." subGenus="Sarsiana">
<emphasis id="B91BEAE6FFF1EE04F11F5037D8561B51" box="[323,624,1590,1617]" italics="true" pageId="3" pageNumber="518">Laonice (Sarsiana) alberti</emphasis>
</taxonomicName>
<emphasis id="B91BEAE6FFF1EE04F2245037D8F51B50" bold="true" box="[632,723,1591,1617]" pageId="3" pageNumber="518">
<taxonomicNameLabel id="A228579DFFF1EE04F2245037D8F51B50" box="[632,723,1591,1617]" pageId="3" pageNumber="518" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
belongs to the subgenus
<taxonomicName id="4C6F4D77FFF1EE04F3A85037DE7E1B50" authorityName="Sikorski, Gunton &amp; Pavlova" authorityYear="2017" box="[1012,1112,1591,1617]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="3" pageNumber="518" phylum="Annelida" rank="subGenus" subGenus="Sarsiana">
<emphasis id="B91BEAE6FFF1EE04F3A85037DE7E1B50" box="[1012,1112,1591,1617]" italics="true" pageId="3" pageNumber="518">Sarsiana</emphasis>
</taxonomicName>
due to the presence of short nuchal organs with a narrow range of length variation, and due to a prostomium and peristomium fused only by a membrane, inconspicuous when observed in dorsal view (although the membrane may also be hidden in the groove between prostomium and peristomium). While the material examined only showed neuropodial hooks, notopodial hooks might be present in the posterior-most parapodia of complete specimens, as they are known to occur in other species within the subgenus
<taxonomicName id="4C6F4D77FFF1EE04F26650EBD8B81A04" authorityName="Sikorski, Gunton &amp; Pavlova" authorityYear="2017" box="[570,670,1771,1797]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="3" pageNumber="518" phylum="Annelida" rank="subGenus" subGenus="Sarsiana">
<emphasis id="B91BEAE6FFF1EE04F26650EBD8B81A04" box="[570,670,1771,1797]" italics="true" pageId="3" pageNumber="518">Sarsiana</emphasis>
</taxonomicName>
, and especially in those having prostomium and peristomium fused by a membrane. The new species has relatively short nuchal organs. Two species of the subgenus
<taxonomicName id="4C6F4D77FFF1EE04F4B8510FDF6E1A28" authorityName="Sikorski, Gunton &amp; Pavlova" authorityYear="2017" box="[1252,1352,1807,1833]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="3" pageNumber="518" phylum="Annelida" rank="subGenus" subGenus="Sarsiana">
<emphasis id="B91BEAE6FFF1EE04F4B8510FDF6E1A28" box="[1252,1352,1807,1833]" italics="true" pageId="3" pageNumber="518">Sarsiana</emphasis>
</taxonomicName>
:
<taxonomicName id="4C6F4D77FFF1EE04F50B510FDBFA1A4C" authority="Ward, 1981" authorityName="Ward" authorityYear="1981" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="3" pageNumber="518" phylum="Annelida" rank="species" species="papillibranchiae" subGenus="Sarsiana">
<emphasis id="B91BEAE6FFF1EE04F50B510FDB741A4D" italics="true" pageId="3" pageNumber="518">L. (S.) papillibranchiae</emphasis>
<bibRefCitation id="EFFE4B05FFF1EE04F1055132DBFA1A4C" author="Ward, L. A." box="[345,476,1842,1869]" pageId="3" pageNumber="518" pagination="713 - 730" refId="ref9052" refString="Ward, L. A. (1981) Spionidae (Polychaeta: Annelida) from Hawaii, with descriptions of five new species. Proceedings of the Biological Society of Washington, 94 (3), 713 - 730." type="journal article" year="1981">Ward, 1981</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C6F4D77FFF1EE04F24F5133DE0C1A4C" authority="Aguirrezabalaga &amp; Ceberio, 2005" authorityName="Aguirrezabalaga &amp; Ceberio" authorityYear="2005" box="[531,1066,1842,1869]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="3" pageNumber="518" phylum="Annelida" rank="species" species="junoyi" subGenus="Sarsiana">
<emphasis id="B91BEAE6FFF1EE04F24F5133D8801A4D" box="[531,678,1843,1869]" italics="true" pageId="3" pageNumber="518">L. (S.) junoyi</emphasis>
<bibRefCitation id="EFFE4B05FFF1EE04F2F25132DE0C1A4C" author="Aguirrezabalaga, F. &amp; Ceberio, A." box="[686,1066,1842,1869]" pageId="3" pageNumber="518" pagination="267 - 280" refId="ref6577" refString="Aguirrezabalaga, F. &amp; Ceberio, A. (2005) Spionidae (Annelida: Polychaeta) from the Capbreton Canyon (Bay of Biscay, NE Atlantic) with descriptions of a new genus and three new species. Marine Biology Research, 1 (4), 267 - 280. https: // doi. org / 10.1080 / 17451000500262066" type="journal article" year="2005">Aguirrezabalaga &amp; Ceberio, 2005</bibRefCitation>
</taxonomicName>
are characterized by having also short nuchal organs extending up to chaetigers 4–6 and 6–7 respectively (
<quantity id="4C979B11FFF1EE04F3975156DE341A71" box="[971,1042,1878,1905]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" metricValueMax="1.778" metricValueMin="1.27" pageId="3" pageNumber="518" unit="in" value="6.0" valueMax="7.0" valueMin="5.0">5–7 in</quantity>
<taxonomicName id="4C6F4D77FFF1EE04F4455157DE891A71" authority="Sikorski &amp; Pavlova &amp; Sardá &amp; Langeneck &amp; Gil &amp; Ravara, 2021" authorityName="Sikorski &amp; Pavlova &amp; Sardá &amp; Langeneck &amp; Gil &amp; Ravara" authorityYear="2021" box="[1049,1199,1878,1905]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="3" pageNumber="518" phylum="Annelida" rank="species" species="alberti" status="sp. nov." subGenus="Sarsiana">
<emphasis id="B91BEAE6FFF1EE04F4455157DE891A71" box="[1049,1199,1878,1905]" italics="true" pageId="3" pageNumber="518">L. (S.) alberti</emphasis>
</taxonomicName>
<emphasis id="B91BEAE6FFF1EE04F4EA5157DF291A70" bold="true" box="[1206,1295,1879,1905]" pageId="3" pageNumber="518">
<taxonomicNameLabel id="A228579DFFF1EE04F4EA5157DF291A70" box="[1206,1295,1879,1905]" pageId="3" pageNumber="518" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), though the prostomium is not fused with the peristomium in any way. In addition, inter-parapodial pouches in those species appear on chaetiger 12 and 8–9, respectively, against
<quantity id="4C979B11FFF1EE04F291519FD9091AB9" box="[717,815,1950,1977]" metricMagnitude="-1" metricUnit="m" metricValue="6.984999999999999" metricValueMax="8.636" metricValueMin="5.334" pageId="3" pageNumber="518" unit="in" value="27.5" valueMax="34.0" valueMin="21.0">21–34 in</quantity>
the new species. Two other deep-sea species, namely
<taxonomicName id="4C6F4D77FFF1EE04F5DA519FD89B1ADC" authority="Sigvaldadottir &amp; Desbruyres, 2003" authorityName="Sigvaldadottir &amp; Desbruyres" authorityYear="2003" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="3" pageNumber="518" phylum="Annelida" rank="species" species="asaccata" subGenus="Sarsiana">
<emphasis id="B91BEAE6FFF1EE04F5DA519FDB0C1ADD" italics="true" pageId="3" pageNumber="518">L. (S.) asaccata</emphasis>
Sigvaldadóttir &amp; Desbruyres, 2003
</taxonomicName>
, and
<taxonomicName id="4C6F4D77FFF1EE04F2A451C3DEB01ADC" authority="Sikorski, 2003" authorityName="Sikorski" authorityYear="2003" box="[760,1174,1986,2013]" class="Polychaeta" family="Spionidae" genus="Laonice" kingdom="Animalia" order="Spionida" pageId="3" pageNumber="518" phylum="Annelida" rank="species" species="shamrockensis" subGenus="Sarsiana">
<emphasis id="B91BEAE6FFF1EE04F2A451C3D9CF1ADD" box="[760,1001,1986,2013]" italics="true" pageId="3" pageNumber="518">L. (S.) shamrockensis</emphasis>
<bibRefCitation id="EFFE4B05FFF1EE04F3AC51C2DEB01ADC" author="Sikorski, A. V." box="[1008,1174,1986,2013]" pageId="3" pageNumber="518" pagination="316 - 345" refId="ref8665" refString="Sikorski, A. V. (2003) Laonice (Polychaeta, Spionidae) in the Arctic and the North Atlantic. Sarsia, 88 (5), 316 - 345. https: // doi. org / 10.1080 / 00364820310002551" type="journal article" year="2003">Sikorski, 2003</bibRefCitation>
</taxonomicName>
, also have short nuchal organs (up to chaetigers 4 and 5, respectively), but their prostomium is clearly fused with the peristomium in the anterior margin.
</paragraph>
</subSubSection>
</treatment>
</document>
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