423 lines
51 KiB
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423 lines
51 KiB
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<document id="FB2BC4B0DDED5889C64A9FDDA9486F7D" ID-CLB-Dataset="273183" ID-DOI="10.1080/00222933.2023.2223386" ID-GBIF-Dataset="5cd7307d-1cec-4f08-8965-9c8e1ca2f625" ID-ISSN="1464-5262" ID-Zenodo-Dep="10015601" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1696936349817" checkinUser="plazi" docAuthor="Sutcharit, Chirasak, Ngor, Peng Bun, Páll-Gergely, Barna, Jeratthitikul, Ekgachai, Siriwut, Warut, Srisonchai, Ruttapon, Ng, Ting Hui, Jirapatrasilp, Parin & Panha, Somsak" docDate="2023" docId="03E387B0FFFCCB63FEABFB94FD17BBC7" docLanguage="en" docName="JNATHIST.57.25-28.1287-1303.pdf" docOrigin="Journal of Natural History 57 (25 - 28)" docSource="http://dx.doi.org/10.1080/00222933.2023.2223386" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Gyliotrachela khmeriana Sutcharit and Panha 2023, sp. n." docType="treatment" docVersion="6" lastPageNumber="1303" masterDocId="FFDAFFC8FFF5CB6EFFCAFFFDFFC8B914" masterDocTitle="Notes on the hypselostomatid snails (Gastropoda: Heterobranchia) from limestone hills in Western Cambodia with a new record and a new species" masterLastPageNumber="1303" masterPageNumber="1287" pageNumber="1287" updateTime="1699694407893" updateUser="plazi">
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<mods:titleInfo id="327CEDD182B580E437E502F3F875FA8F">
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<mods:title id="F20C57472943FECDF22525F478A3DA80">Notes on the hypselostomatid snails (Gastropoda: Heterobranchia) from limestone hills in Western Cambodia with a new record and a new species</mods:title>
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<mods:name id="03C4088907A1C16F95C20F5F9765E5FD" type="personal">
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<mods:namePart id="F74F5C06BBE4D6A3D8D96CDCFA2172C5">Sutcharit, Chirasak</mods:namePart>
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</mods:role>
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<mods:namePart id="C197D4314DF8219555265A12104F8A38">Ngor, Peng Bun</mods:namePart>
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</mods:name>
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<mods:roleTerm id="4902413C786C951461CD7E0DF28A057F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B50CFB96CB66577DEC0F8DE8678AEFD2">Páll-Gergely, Barna</mods:namePart>
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</mods:name>
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<mods:name id="2E1B5457577D4B6C897C352C3310FADB" type="personal">
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<mods:roleTerm id="B82C2A8D17312F983C53018AFF909B7C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="61161F11E2B311DEA2DDBE0E780D7138">Jeratthitikul, Ekgachai</mods:namePart>
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</mods:role>
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<mods:namePart id="5EF18CBB02E33127F4CF3E603FF0D998">Siriwut, Warut</mods:namePart>
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</mods:name>
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<mods:name id="2F6EAC194AEEECF594190CF8B0B5FD9F" type="personal">
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<mods:roleTerm id="08140C59CC6D145271567C2DEEC67444">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E13AE10FF25E26E689CDB6AA262C6584">Srisonchai, Ruttapon</mods:namePart>
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</mods:name>
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<mods:name id="2DE13084C6AA722F587AB66DFAA79DFD" type="personal">
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<mods:roleTerm id="A4B19020FB4D201238A602A672D53BEA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="748831167ECEE5032E188579869C2BCA">Ng, Ting Hui</mods:namePart>
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</mods:name>
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<mods:name id="27649DE49F219B9C6A3BCEF06384A64B" type="personal">
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</mods:role>
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<mods:namePart id="70FA9193C605F1FE8B014FDF66FFFB4E">Jirapatrasilp, Parin</mods:namePart>
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</mods:name>
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<mods:name id="6625B1DC2FB2EECE7D1846E2D9C8C343" type="personal">
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</mods:role>
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<mods:namePart id="9CC77B2A97DB222520963E3FD4FC2DE5">Panha, Somsak</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="F1B80C86091FE23FC3225731DCC45A8E">text</mods:typeOfResource>
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<mods:titleInfo id="31DF3C303EB9634CB737F04118C31FC8">
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<mods:title id="7FD58B8B711575D00055661B6D20F02E">Journal of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part id="02E92CDBA6BA88C68BAC9A6221D533DE">
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<mods:date id="E5509F049B4EEB75C48CFDE3488B47ED">2023</mods:date>
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<mods:detail id="4E8923448FE19E0DC99D44FF5ACE4668" type="pubDate">
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<mods:number id="E848239D8F429C98C8E0E0D68AE57E6D">2023-09-27</mods:number>
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<mods:detail id="C891EA1A5C7C82184344729802ACB708" type="volume">
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<mods:number id="EAE5234866C0DD48ECC6AFFABBAE73F3">57</mods:number>
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</mods:detail>
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<mods:detail id="0A82C99D9FF185C7CA4C472102037C81" type="issue">
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<mods:number id="201462A69D9F9D9762AD0F85A1B0D665">25 - 28</mods:number>
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<mods:start id="81E062F6A6808DDD676B566E0509F035">1287</mods:start>
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<mods:end id="AAEDC7000E17164838B972FE7FFA237C">1303</mods:end>
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<mods:url id="CECB9542BF9632026A8BD00048A1028D">http://dx.doi.org/10.1080/00222933.2023.2223386</mods:url>
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<mods:classification id="A88F2445C82712E4B6AE86674776091B">journal article</mods:classification>
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<mods:identifier id="C163C5E406E4674AE1CD648615A70C0A" type="CLB-Dataset">273183</mods:identifier>
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<mods:identifier id="A90878989894F466D03C453C3494001E" type="DOI">10.1080/00222933.2023.2223386</mods:identifier>
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<mods:identifier id="8D5EDE929C9547FD32B5D31257A41110" type="ISSN">1464-5262</mods:identifier>
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<mods:identifier id="7C4A625969EB3E6AA3597ACC7114EDB5" type="Zenodo-Dep">10015601</mods:identifier>
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</mods:mods>
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<treatment id="03E387B0FFFCCB63FEABFB94FD17BBC7" ID-DOI="http://doi.org/10.5281/zenodo.8428079" ID-GBIF-Taxon="214036415" ID-Zenodo-Dep="8428079" LSID="urn:lsid:plazi:treatment:03E387B0FFFCCB63FEABFB94FD17BBC7" httpUri="http://treatment.plazi.org/id/03E387B0FFFCCB63FEABFB94FD17BBC7" lastPageId="13" pageId="9">
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<subSubSection id="C350652DFFFCCB67FEABFB94FBBBBDBA" pageId="9" type="nomenclature">
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<paragraph id="8BF536A6FFFCCB67FEABFB94FC0ABD97" blockId="9.[176,1139,1129,1272]" box="[353,962,1129,1155]" pageId="9">
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<heading id="D0BD81CAFFFCCB67FEABFB94FC0ABD97" box="[353,962,1129,1155]" centered="true" fontSize="11" level="2" pageId="9" reason="2">
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<taxonomicName id="4C4A4D25FFFCCB67FEABFB94FCBBBD97" ID-CoL="36925d4f-310b-44b0-9bb1-fb725c3762f1" authority="Sutcharit and Panha" authorityName="Sutcharit and Panha" authorityYear="2023" box="[353,883,1129,1155]" class="Gastropoda" family="Gastrocoptidae" genus="Gyliotrachela" kingdom="Animalia" order="Stylommatophora" pageId="9" phylum="Mollusca" rank="species" species="khmeriana" status="sp. nov.">
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<emphasis id="B93EEAB4FFFCCB67FEABFB94FDB2BD97" box="[353,634,1129,1155]" italics="true" pageId="9">Gyliotrachela khmeriana</emphasis>
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Sutcharit and Panha
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</taxonomicName>
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,
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<emphasis id="B93EEAB4FFFCCB67FCB4FB94FC0ABD97" bold="true" box="[894,962,1129,1155]" pageId="9">
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<taxonomicNameLabel id="A20D57CFFFFCCB67FCB4FB94FC0ABD97" box="[894,962,1129,1155]" pageId="9" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF536A6FFFCCB67FF7AFB6EFBBBBDBA" blockId="9.[176,1139,1129,1272]" box="[176,1139,1171,1198]" pageId="9">http://www.zoobank.org/urn:lsid:zoobank.org:act:58A3E65E-0289-48D2-ABD5-</paragraph>
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|
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</subSubSection>
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|||
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<subSubSection id="C350652DFFFCCB67FDF9FB48FCC7BDEC" pageId="9" type="description">
|
|||
|
|
<paragraph id="8BF536A6FFFCCB67FDF9FB48FD38BDC4" blockId="9.[176,1139,1129,1272]" box="[563,752,1205,1232]" pageId="9">D52762D0E754</paragraph>
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<paragraph id="8BF536A6FFFCCB67FDDEFB23FCC7BDEC" blockId="9.[176,1139,1129,1272]" box="[532,783,1246,1272]" pageId="9">
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(
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<figureCitation id="13712A23FFFCCB67FDD6FB23FD4FBDEC" box="[540,647,1246,1272]" captionStart="Figure 1" captionStartId="3.[160,225,968,991]" captionTargetBox="[162,1154,142,935]" captionTargetId="figure-103@3.[162,1154,142,936]" captionTargetPageId="3" captionText="Figure 1. Distribution map of all hypselostomatids known from Cambodia. The small black dots indicate the approximate type locality for 1. Acinolaemus carcharodon, 2. Acinolaemus pyramidalis, 3. Acinolaemus rectus, 4. Anauchen chaunosalpinx, 5. Anauchen depressus, 6. Aulacospira conica, 7. Aulacospira furtiva, 8. Gyliotrachela khmeriana sp. nov., 9. Gyliotrachela torticollis, 10. Hypselostoma benetuitum, 11. Hypselostoma cambodjense, 12. Hypselostoma discobasis, and 13. Hypselostoma srakeoensis. The approximate location of limestone areas in Cambodia (Sisophon-Battambang, Stung Treng, and Kampot areas) is modified from Mouret (2004). Asterisks (*) indicate new localities recorded in this study." figureDoi="http://doi.org/10.5281/zenodo.10015603" httpUri="https://zenodo.org/record/10015603/files/figure.png" pageId="9">Figures 1</figureCitation>
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,
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<figureCitation id="13712A23FFFCCB67FD5FFB23FD6BBDEC" box="[661,675,1246,1272]" captionStart="Figure 4" captionStartId="10.[160,225,1233,1256]" captionTargetBox="[162,1154,142,1201]" captionTargetId="figure-37@10.[162,1154,142,1201]" captionTargetPageId="10" captionText="Figure 4. Gyliotrachela khmeriana sp. nov. (a, b) Holotype CUMZ 14206/1 from Phnom Sampov, Battambang, (a) shell with apertural dentition arrangement, and (b) protoconch sculpture. (c) Paratype CUMZ 14206/2 from the type locality with apertural dentition (very reduced basal lamella). Abbreviations: a = angular lamella, b = basal lamella, c = columellar lamella, lpl = lower palatal lamella, p = parietal lamella, upl = upper palatal lamella." figureDoi="http://doi.org/10.5281/zenodo.10015609" httpUri="https://zenodo.org/record/10015609/files/figure.png" pageId="9">4</figureCitation>
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;
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<tableCitation id="C6C8031DFFFCCB67FD7BFB23FCCFBDEC" box="[689,775,1246,1272]" captionStart="Table 1" captionStartId="4.[160,216,142,165]" captionTargetBox="[171,1144,237,1008]" captionTargetPageId="4" captionText="Table 1. List of all hypselostomatid species recorded from Cambodia.References: 1 = Benthem Jutting" httpUri="http://table.plazi.org/id/DF35662EFFF1CB6AFF6AFF73FE08B9F4" pageId="9" tableUuid="DF35662EFFF1CB6AFF6AFF73FE08B9F4">Table 1</tableCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C350652DFFFCCB67FF6AFAC1FDA8BF3A" pageId="9" type="materials_examined">
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<paragraph id="8BF536A6FFFCCB67FF6AFAC1FE8CBC42" blockId="9.[160,1156,1340,1436]" box="[160,324,1340,1366]" pageId="9">
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<emphasis id="B93EEAB4FFFCCB67FF6AFAC1FE8CBC42" bold="true" box="[160,324,1340,1366]" italics="true" pageId="9">Type material</emphasis>
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</paragraph>
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<paragraph id="8BF536A6FFFCCB67FF6AFA9DFB4CBC88" blockId="9.[160,1156,1340,1436]" pageId="9">
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<materialsCitation id="3B223CFBFFFCCB67FF6AFA9DFB4CBC6D" ID-GBIF-Occurrence="4426912306" box="[160,1156,1376,1401]" collectionCode="CUMZ" pageId="9" specimenCode="CUMZ 14206" specimenCount="2" typeStatus="holotype">
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<typeStatus id="54F18804FFFCCB67FF6AFA9DFECFBC6D" box="[160,263,1376,1401]" pageId="9" type="holotype">Holotype</typeStatus>
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<specimenCode id="DBEC9EDDFFFCCB67FEDBFA9DFE63BC6D" box="[273,427,1376,1401]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="9">CUMZ 14206</specimenCode>
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/1 (height
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<quantity id="4CB29B43FFFCCB67FDD5FA9DFDB1BC6D" box="[543,633,1376,1401]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="9" unit="mm" value="3.1">3.1 mm</quantity>
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, width
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<quantity id="4CB29B43FFFCCB67FD1AFA9DFCE2BC6D" box="[720,810,1376,1401]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="9" unit="mm" value="3.4">3.4 mm</quantity>
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, and 5 whorls;
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<figureCitation id="13712A23FFFCCB67FC17FA9DFBF5BC6D" box="[989,1085,1376,1401]" captionStart="Figure 4" captionStartId="10.[160,225,1233,1256]" captionTargetBox="[162,1154,142,1201]" captionTargetId="figure-37@10.[162,1154,142,1201]" captionTargetPageId="10" captionText="Figure 4. Gyliotrachela khmeriana sp. nov. (a, b) Holotype CUMZ 14206/1 from Phnom Sampov, Battambang, (a) shell with apertural dentition arrangement, and (b) protoconch sculpture. (c) Paratype CUMZ 14206/2 from the type locality with apertural dentition (very reduced basal lamella). Abbreviations: a = angular lamella, b = basal lamella, c = columellar lamella, lpl = lower palatal lamella, p = parietal lamella, upl = upper palatal lamella." figureDoi="http://doi.org/10.5281/zenodo.10015609" httpUri="https://zenodo.org/record/10015609/files/figure.png" pageId="9">Figure 4</figureCitation>
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(a, b)),
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</materialsCitation>
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<materialsCitation id="3B223CFBFFFCCB67FF6AFA7EFB4CBC88" ID-GBIF-Occurrence="4426912303" box="[160,1156,1411,1436]" collectionCode="CUMZ" pageId="9" specimenCode="CUMZ 14206" specimenCount="2" typeStatus="paratype">
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<typeStatus id="54F18804FFFCCB67FF6AFA7EFECCBC88" box="[160,260,1411,1436]" pageId="9" type="paratype">paratype</typeStatus>
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<specimenCode id="DBEC9EDDFFFCCB67FEC0FA7EFE54BC88" box="[266,412,1411,1436]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="9">CUMZ 14206</specimenCode>
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/2 (1 shell; height
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<quantity id="4CB29B43FFFCCB67FDAAFA7EFD7EBC88" box="[608,694,1411,1436]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="9" unit="mm" value="3.2">3.2 mm</quantity>
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|
|
, width
|
|||
|
|
<quantity id="4CB29B43FFFCCB67FCCCFA7EFC94BC88" box="[774,860,1411,1436]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="9" unit="mm" value="3.4">3.4 mm</quantity>
|
|||
|
|
, and 5 whorls;
|
|||
|
|
<figureCitation id="13712A23FFFCCB67FBCAFA7EFB93BC88" box="[1024,1115,1411,1436]" captionStart="Figure 4" captionStartId="10.[160,225,1233,1256]" captionTargetBox="[162,1154,142,1201]" captionTargetId="figure-37@10.[162,1154,142,1201]" captionTargetPageId="10" captionText="Figure 4. Gyliotrachela khmeriana sp. nov. (a, b) Holotype CUMZ 14206/1 from Phnom Sampov, Battambang, (a) shell with apertural dentition arrangement, and (b) protoconch sculpture. (c) Paratype CUMZ 14206/2 from the type locality with apertural dentition (very reduced basal lamella). Abbreviations: a = angular lamella, b = basal lamella, c = columellar lamella, lpl = lower palatal lamella, p = parietal lamella, upl = upper palatal lamella." figureDoi="http://doi.org/10.5281/zenodo.10015609" httpUri="https://zenodo.org/record/10015609/files/figure.png" pageId="9">Figure 4</figureCitation>
|
|||
|
|
(c)).
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BF536A6FFFCCB67FF6AFA32FEFEBCFD" blockId="9.[160,1156,1487,1582]" box="[160,310,1487,1513]" pageId="9">
|
|||
|
|
<emphasis id="B93EEAB4FFFCCB67FF6AFA32FEFEBCFD" bold="true" box="[160,310,1487,1513]" italics="true" pageId="9">Type locality</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BF536A6FFFCCB67FF6AFA0FFDA8BF3A" blockId="9.[160,1156,1487,1582]" pageId="9">
|
|||
|
|
<materialsCitation id="3B223CFBFFFCCB67FF6AFA0FFD94BF3A" ID-GBIF-Occurrence="4426912312" country="Cambodia" county="Banan District" latitude="13.026" location="The" longLatPrecision="1" longitude="103.10101" municipality="Phnom Sampeov Mountain" pageId="9" specimenCount="1" stateProvince="Battambang">
|
|||
|
|
<location id="8E95607DFFFCCB67FF6AFA0FFF01BF1F" LSID="urn:lsid:plazi:treatment:03E387B0FFFCCB63FEABFB94FD17BBC7:8E95607DFFFCCB67FF6AFA0FFF01BF1F" box="[160,201,1522,1547]" country="Cambodia" county="Banan District" latitude="13.026" longLatPrecision="1" longitude="103.10101" municipality="Phnom Sampeov Mountain" name="The" pageId="9" stateProvince="Battambang">The</location>
|
|||
|
|
limestone hills at
|
|||
|
|
<collectingMunicipality id="6B91ACDCFFFCCB67FE51FA0FFD1CBF1F" box="[411,724,1522,1547]" pageId="9">Phnom Sampeov Mountain</collectingMunicipality>
|
|||
|
|
,
|
|||
|
|
<collectingCounty id="62944E2AFFFCCB67FD15FA0FFC48BF1F" box="[735,896,1522,1547]" pageId="9">Banan District</collectingCounty>
|
|||
|
|
,
|
|||
|
|
<collectingRegion id="498EF844FFFCCB67FC41FA0FFB48BF1F" box="[907,1152,1522,1547]" country="Cambodia" name="Battambang" pageId="9">Battambang Province</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="F35D7636FFFCCB67FF6AF9E8FEDBBF3A" box="[160,275,1557,1582]" name="Cambodia" pageId="9">Cambodia</collectingCountry>
|
|||
|
|
(
|
|||
|
|
<geoCoordinate id="EE7E5061FFFCCB67FEEAF9E8FE79BF3A" box="[288,433,1557,1582]" degrees="13.026000" direction="north" orientation="latitude" pageId="9" precision="1" value="13.026">13.026000°N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE7E5061FFFCCB67FE76F9E8FD9EBF3A" box="[444,598,1557,1582]" degrees="103.101011" direction="east" orientation="longitude" pageId="9" precision="1" value="103.10101">103.101011°E</geoCoordinate>
|
|||
|
|
)
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C350652DFFFCCB67FF6AF99CFE01BFD4" pageId="9" type="etymology">
|
|||
|
|
<paragraph id="8BF536A6FFFCCB67FF6AF99CFED5BF6F" blockId="9.[160,1156,1633,1728]" box="[160,285,1633,1659]" pageId="9">
|
|||
|
|
<emphasis id="B93EEAB4FFFCCB67FF6AF99CFED5BF6F" bold="true" box="[160,285,1633,1659]" italics="true" pageId="9">Etymology</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BF536A6FFFCCB67FF6AF978FE01BFD4" blockId="9.[160,1156,1633,1728]" pageId="9">
|
|||
|
|
The specific name ̍
|
|||
|
|
<taxonomicName id="4C4A4D25FFFCCB67FEB3F978FE26BF8A" authorityName="Sutcharit and Panha" authorityYear="2023" box="[377,494,1669,1694]" class="Gastropoda" family="Gastrocoptidae" genus="Gyliotrachela" kingdom="Animalia" order="Stylommatophora" pageId="9" phylum="Mollusca" rank="species" species="khmeriana">
|
|||
|
|
<emphasis id="B93EEAB4FFFCCB67FEB3F978FE26BF8A" box="[377,494,1669,1694]" italics="true" pageId="9">khmeriana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
′ refers to the most numerous ethnic groups of
|
|||
|
|
<collectingCountry id="F35D7636FFFCCB67FBC1F978FB48BF8A" box="[1035,1152,1669,1694]" name="Cambodia" pageId="9">Cambodia</collectingCountry>
|
|||
|
|
; a Latin feminine adjective.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<caption id="DF35662EFFFFCB64FF6AFB2CFCD7BC49" ID-DOI="http://doi.org/10.5281/zenodo.10015609" ID-Zenodo-Dep="10015609" httpUri="https://zenodo.org/record/10015609/files/figure.png" pageId="10" startId="10.[160,225,1233,1256]" targetBox="[162,1154,142,1201]" targetPageId="10" targetType="figure">
|
|||
|
|
<paragraph id="8BF536A6FFFFCB64FF6AFB2CFCD7BC49" blockId="10.[160,1157,1233,1373]" pageId="10">
|
|||
|
|
<emphasis id="B93EEAB4FFFFCB64FF6AFB2CFF36BDFC" bold="true" box="[160,254,1233,1256]" pageId="10">Figure 4.</emphasis>
|
|||
|
|
<taxonomicName id="4C4A4D25FFFFCB64FEC3FB2CFE3FBDFC" authority="Sutcharit & Ngor & Páll-Gergely & Jeratthitikul & Siriwut & Srisonchai & Ng & Jirapatrasilp & Panha, 2023" authorityName="Sutcharit & Ngor & Páll-Gergely & Jeratthitikul & Siriwut & Srisonchai & Ng & Jirapatrasilp & Panha" authorityYear="2023" box="[265,503,1233,1256]" class="Gastropoda" family="Gastrocoptidae" genus="Gyliotrachela" kingdom="Animalia" order="Stylommatophora" pageId="10" phylum="Mollusca" rank="species" species="khmeriana" status="sp. nov.">
|
|||
|
|
<emphasis id="B93EEAB4FFFFCB64FEC3FB2CFE3FBDFC" box="[265,503,1233,1256]" italics="true" pageId="10">Gyliotrachela khmeriana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A20D57CFFFFFCB64FDC8FB2CFD9CBDFC" box="[514,596,1233,1256]" pageId="10" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
(a, b) Holotype CUMZ 14206/1 from Phnom Sampov, Battambang, (a) shell with apertural dentition arrangement, and (b) protoconch sculpture. (c) Paratype CUMZ 14206/2 from the type locality with apertural dentition (very reduced basal lamella). Abbreviations: a = angular lamella, b = basal lamella, c = columellar lamella, lpl = lower palatal lamella, p = parietal lamella, upl = upper palatal lamella.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<subSubSection id="C350652DFFFFCB64FF6AFA49FEEEBF4C" pageId="10" type="diagnosis">
|
|||
|
|
<paragraph id="8BF536A6FFFFCB64FF6AFA49FEDCBCDA" blockId="10.[160,1156,1460,1624]" box="[160,276,1460,1486]" pageId="10">
|
|||
|
|
<emphasis id="B93EEAB4FFFFCB64FF6AFA49FEDCBCDA" bold="true" box="[160,276,1460,1486]" italics="true" pageId="10">Diagnosis</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BF536A6FFFFCB64FF6AFA2AFEEEBF4C" blockId="10.[160,1156,1460,1624]" pageId="10">Shell minute heliciform, and slightly short and straight tuba. Protoconch sculptures with shallow pits and spiral striations; teleoconch without spiral striation. Apertural dentition with six lamellae (parietal, angular, upper and lower palatals, basal, and columellar).</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<caption id="DF35662EFFFECB65FF6AFA9FFE74BF18" ID-DOI="http://doi.org/10.5281/zenodo.10015611" ID-Zenodo-Dep="10015611" httpUri="https://zenodo.org/record/10015611/files/figure.png" pageId="11" startId="11.[160,225,1378,1401]" targetBox="[162,1154,142,1347]" targetPageId="11" targetType="figure">
|
|||
|
|
<paragraph id="8BF536A6FFFECB65FF6AFA9FFE74BF18" blockId="11.[160,1156,1378,1548]" pageId="11">
|
|||
|
|
<emphasis id="B93EEAB4FFFECB65FF6AFA9FFF34BC6D" bold="true" box="[160,252,1378,1401]" pageId="11">Figure 5.</emphasis>
|
|||
|
|
Comparative shell (tuba) and apertural dentition among the three genera based on the respective type species. (a)
|
|||
|
|
<taxonomicName id="4C4A4D25FFFECB65FE66FA7DFD45BC83" authority="Pilsbry, 1917" authorityName="Pilsbry" authorityYear="1917" box="[428,653,1408,1431]" class="Gastropoda" family="Hypselostomatidae" genus="Anauchen" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B93EEAB4FFFECB65FE66FA7DFDC3BC83" box="[428,523,1408,1431]" italics="true" pageId="11">Anauchen</emphasis>
|
|||
|
|
<bibRefCitation id="EFDB4B57FFFECB65FDD8FA7DFD45BC83" author="Pilsbry HA" box="[530,653,1408,1431]" pageId="11" refId="ref7609" refString="Pilsbry HA. 1917. Manual of Conchology, Structure and Systematic, with Illustrations of the Species, 2 nd Series. Vol. 24. Pupillidae (Gastrocoptinae). Philadelphia (PA): Academy of Natural Science; Pupillidae (Gastrocoptinae)." type="book" year="1917">Pilsbry, 1917</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(syntype MNHN-IM-2000-35,161 of
|
|||
|
|
<taxonomicName id="4C4A4D25FFFECB65FC24FA7DFB49BC83" authorityName="Bavay and Dautzenberg" authorityYear="1904" box="[1006,1153,1408,1431]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="11" phylum="Mollusca" rank="species" species="gereti">Boysidia gereti</taxonomicName>
|
|||
|
|
;
|
|||
|
|
<bibRefCitation id="EFDB4B57FFFECB65FF6AFA60FE0DBCA0" author="Bavay A & Dautzenberg P" box="[160,453,1437,1460]" journalOrPublisher="J Conchyl." pageId="11" pagination="201 - 236" part="51" refId="ref6351" refString="Bavay A, Dautzenberg P. 1904. Description de coquilles nouvelles de lIndo-Chine. J Conchyl. 51: 201 - 236. [1903]." title="Description de coquilles nouvelles de lIndo-Chine" type="journal article" year="1904">Bavay and Dautzenberg, 1904</bibRefCitation>
|
|||
|
|
). (b)
|
|||
|
|
<taxonomicName id="4C4A4D25FFFECB65FE33FA60FD33BCA0" authority="Tomlin, 1930" authorityName="Tomlin" authorityYear="1930" box="[505,763,1437,1460]" class="Gastropoda" family="Gastrocoptidae" genus="Gyliotrachela" kingdom="Animalia" order="Stylommatophora" pageId="11" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B93EEAB4FFFECB65FE33FA60FDBEBCA0" box="[505,630,1437,1460]" italics="true" pageId="11">Gyliotrachela</emphasis>
|
|||
|
|
<bibRefCitation id="EFDB4B57FFFECB65FDB6FA60FD33BCA0" author="Tomlin JRLB" box="[636,763,1437,1460]" pageId="11" pagination="22 - 24" refId="ref8029" refString="Tomlin JRLB. 1930. Some preoccupied generic names. Proc Malacol Soc London. 19: 22 - 24." type="journal article" year="1930">Tomlin, 1930</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(lectotype SMF 4587/1 of
|
|||
|
|
<taxonomicName id="4C4A4D25FFFECB65FC34FA60FE33BCC5" authority="Mollendorff, 1891" authorityName="Mollendorff" authorityYear="1891" class="Gastropoda" family="Gastrocoptidae" genus="Hypselostoma" kingdom="Animalia" order="Stylommatophora" pageId="11" phylum="Mollusca" rank="species" species="hungerfordianum">
|
|||
|
|
<emphasis id="B93EEAB4FFFECB65FC34FA60FE8FBCC5" italics="true" pageId="11">Hypselostoma hungerfordianum</emphasis>
|
|||
|
|
<bibRefCitation id="EFDB4B57FFFECB65FE84FA47FE33BCC5" author="Mollendorff OF" box="[334,507,1466,1489]" pageId="11" pagination="330 - 348" refId="ref7010" refString="Mollendorff OF von. 1891. On the land and freshwater shells of Perak. Proc Zool Soc London. 59: 330 - 348. doi: 10.1111 / j. 1096 - 3642.1891. tb 01757. x." type="journal article" year="1891">Möllendorff, 1891</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
). (c)
|
|||
|
|
<taxonomicName id="4C4A4D25FFFECB65FDE6FA47FCF3BCC5" authority="Benson, 1856" authorityName="Benson" authorityYear="1856" box="[556,827,1466,1489]" class="Gastropoda" family="Gastrocoptidae" genus="Hypselostoma" kingdom="Animalia" order="Stylommatophora" pageId="11" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B93EEAB4FFFECB65FDE6FA47FD79BCC5" box="[556,689,1466,1489]" italics="true" pageId="11">Hypselostoma</emphasis>
|
|||
|
|
<bibRefCitation id="EFDB4B57FFFECB65FD7DFA47FCF3BCC5" author="Benson WH" box="[695,827,1466,1489]" pageId="11" pagination="129 - 131" refId="ref6379" refString="Benson WH. 1856 a. Description of Tanystoma tubiferum, a Burmese form related to the genus Anostoma of Lamarck. Ann Mag Nat Hist. 17: 129 - 131. Ser 2. doi: 10.1080 / 00222935608697483." type="journal article" year="1856">Benson, 1856</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(syntype NHMUK 1954.6.2.568 of
|
|||
|
|
<taxonomicName id="4C4A4D25FFFECB65FF6AFA25FDCABCFB" authority="Benson, 1856" authorityName="Benson" authorityYear="1856" box="[160,514,1496,1519]" class="Gastropoda" family="Gastrocoptidae" genus="Tanystoma" kingdom="Animalia" order="Stylommatophora" pageId="11" phylum="Mollusca" rank="species" species="tubiferum">
|
|||
|
|
<emphasis id="B93EEAB4FFFECB65FF6AFA25FEB8BCFB" box="[160,368,1496,1519]" italics="true" pageId="11">Tanystoma tubiferum</emphasis>
|
|||
|
|
<bibRefCitation id="EFDB4B57FFFECB65FEB0FA25FDCABCFB" author="Benson WH" box="[378,514,1496,1519]" pageId="11" pagination="129 - 131" refId="ref6379" refString="Benson WH. 1856 a. Description of Tanystoma tubiferum, a Burmese form related to the genus Anostoma of Lamarck. Ann Mag Nat Hist. 17: 129 - 131. Ser 2. doi: 10.1080 / 00222935608697483." type="journal article" year="1856">Benson, 1856</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; after
|
|||
|
|
<bibRefCitation id="EFDB4B57FFFECB65FD82FA25FCC8BCFB" author="Preece RC & White TS & Raheem DC & Ketchum H & Ablett J & Taylor H & Webb K & Naggs F" box="[584,768,1496,1519]" pageId="11" pagination="1 - 434" refId="ref7687" refString="Preece RC, White TS, Raheem DC, Ketchum H, Ablett J, Taylor H, Webb K, Naggs F. 2022. William Benson and the origins of the golden age of malacology in British India: biography, illustrated catalogue and evaluation of his molluscan types. Trop Nat Hist. Suppl. 6: 1 - 434." type="journal article" year="2022">
|
|||
|
|
Preece
|
|||
|
|
<emphasis id="B93EEAB4FFFECB65FD5EFA25FD0ABCFB" box="[660,706,1496,1519]" italics="true" pageId="11">et al</emphasis>
|
|||
|
|
. 2022
|
|||
|
|
</bibRefCitation>
|
|||
|
|
: fig. 70d). Abbreviations: a = angular lamella, p = parietal lamella.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<subSubSection id="C350652DFFF9CB62FF6AFF73FC56BBC7" pageId="12" type="description">
|
|||
|
|
<paragraph id="8BF536A6FFF9CB62FF6AFF73FEEEB9BC" blockId="12.[160,1156,142,723]" box="[160,294,142,168]" pageId="12">
|
|||
|
|
<emphasis id="B93EEAB4FFF9CB62FF6AFF73FEEEB9BC" bold="true" box="[160,294,142,168]" italics="true" pageId="12">Description</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BF536A6FFF9CB62FF6AFF4FFC56BBC7" blockId="12.[160,1156,142,723]" pageId="12">
|
|||
|
|
Shell heliciform, last whorl large, straight tuba, and spire conical. Shell colour brownish,
|
|||
|
|
<quantity id="4CB29B43FFF9CB62FF6AFF29FF3DB9F9" box="[160,245,212,237]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="12" unit="mm" value="3.1">3.1 mm</quantity>
|
|||
|
|
in length,
|
|||
|
|
<quantity id="4CB29B43FFF9CB62FEBBFF29FE0DB9F9" box="[369,453,212,237]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="12" unit="mm" value="3.4">3.4 mm</quantity>
|
|||
|
|
in width, and with five whorls. Protoconch about two whorls, sculptured with roundish shallow pits, and thin spiral striations present. Teleoconch with irregularly or wrinkled arranged on strong or weak growth lines. Spire raised, whorls slightly angular, resulting in a scalariform shape of the upper whorls; suture wide and impressed. Last whorl angular and short tuba projecting anteriorly. Only wide and shallow columellar furrow present and extended to onset of tuba. Outer outline of peristome rounded, aperture triangular due to the well-developed sinulus and the angled parietocolumellar and columellar-basal junctions; peristome free with broadly expanded lip margins. Apertural dentition with six prominent lamellae: angular, parietal, upper and lower palatals, columellar (sometime very tiny), and basal. Parietal (p) lamella very tall and situated deep inside on parietal wall. Angular (a) lamella relatively small and situated anteriorly close to expanded lip. Palatal wall with tall and strong upper (upl) and lower palatal (lpl) lamellae. Basal (b) lamella small, knob shaped and located between lower palatal plica and the columellar lamella. Columella lamella strong and prominent. All apertural dentition with smooth surfaces. Umbilicus wide and deep.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C350652DFFF9CB62FF6AFCF3FBDABAA7" pageId="12" type="distribution">
|
|||
|
|
<paragraph id="8BF536A6FFF9CB62FF6AFCF3FEE4BA3C" blockId="12.[160,1155,782,947]" box="[160,300,782,808]" pageId="12">
|
|||
|
|
<emphasis id="B93EEAB4FFF9CB62FF6AFCF3FEE4BA3C" bold="true" box="[160,300,782,808]" italics="true" pageId="12">Distribution</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BF536A6FFF9CB62FF6AFCCFFBDABAA7" blockId="12.[160,1155,782,947]" pageId="12">
|
|||
|
|
<taxonomicName id="4C4A4D25FFF9CB62FF6AFCCFFE62BA5F" authority="Sutcharit & Ngor & Páll-Gergely & Jeratthitikul & Siriwut & Srisonchai & Ng & Jirapatrasilp & Panha, 2023" authorityName="Sutcharit & Ngor & Páll-Gergely & Jeratthitikul & Siriwut & Srisonchai & Ng & Jirapatrasilp & Panha" authorityYear="2023" box="[160,426,818,843]" class="Gastropoda" family="Gastrocoptidae" genus="Gyliotrachela" kingdom="Animalia" order="Stylommatophora" pageId="12" phylum="Mollusca" rank="species" species="khmeriana" status="sp. nov.">
|
|||
|
|
<emphasis id="B93EEAB4FFF9CB62FF6AFCCFFE62BA5F" box="[160,426,818,843]" italics="true" pageId="12">Gyliotrachela khmeriana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A20D57CFFFF9CB62FE7AFCCFFDCEBA5F" box="[432,518,818,843]" pageId="12" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
is known only from the
|
|||
|
|
<typeStatus id="54F18804FFF9CB62FCD8FCCFFC8CBA5F" box="[786,836,818,843]" pageId="12">type</typeStatus>
|
|||
|
|
locality (
|
|||
|
|
<figureCitation id="13712A23FFF9CB62FC62FCCFFBCBBA5F" box="[936,1027,818,843]" captionStart="Figure 1" captionStartId="3.[160,225,968,991]" captionTargetBox="[162,1154,142,935]" captionTargetId="figure-103@3.[162,1154,142,936]" captionTargetPageId="3" captionText="Figure 1. Distribution map of all hypselostomatids known from Cambodia. The small black dots indicate the approximate type locality for 1. Acinolaemus carcharodon, 2. Acinolaemus pyramidalis, 3. Acinolaemus rectus, 4. Anauchen chaunosalpinx, 5. Anauchen depressus, 6. Aulacospira conica, 7. Aulacospira furtiva, 8. Gyliotrachela khmeriana sp. nov., 9. Gyliotrachela torticollis, 10. Hypselostoma benetuitum, 11. Hypselostoma cambodjense, 12. Hypselostoma discobasis, and 13. Hypselostoma srakeoensis. The approximate location of limestone areas in Cambodia (Sisophon-Battambang, Stung Treng, and Kampot areas) is modified from Mouret (2004). Asterisks (*) indicate new localities recorded in this study." figureDoi="http://doi.org/10.5281/zenodo.10015603" httpUri="https://zenodo.org/record/10015603/files/figure.png" pageId="12">Figure 1</figureCitation>
|
|||
|
|
). The snails were found on a limestone wall outside the cave, occurring sympatrically also perhaps syntopically with
|
|||
|
|
<taxonomicName id="4C4A4D25FFF9CB62FEAFFC8AFE2DBA84" authorityName="Benthem Jutting" authorityYear="1962" box="[357,485,887,912]" class="Gastropoda" family="Gastrocoptidae" genus="Gyliotrachela" kingdom="Animalia" order="Stylommatophora" pageId="12" phylum="Mollusca" rank="species" species="torticollis">
|
|||
|
|
<emphasis id="B93EEAB4FFF9CB62FEAFFC8AFE2DBA84" box="[357,485,887,912]" italics="true" pageId="12">G. torticollis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C4A4D25FFF9CB62FDD4FC8AFD68BA84" box="[542,672,887,912]" class="Gastropoda" family="Hydrocenidae" genus="Georissa" kingdom="Animalia" order="Cycloneritida" pageId="12" phylum="Mollusca" rank="species" species="undetermined">
|
|||
|
|
<emphasis id="B93EEAB4FFF9CB62FDD4FC8AFDB1BA84" box="[542,633,887,912]" italics="true" pageId="12">Georissa</emphasis>
|
|||
|
|
sp.
|
|||
|
|
</taxonomicName>
|
|||
|
|
However,
|
|||
|
|
<taxonomicName id="4C4A4D25FFF9CB62FCDDFC8AFC62BA84" authority="Sutcharit & Ngor & Páll-Gergely & Jeratthitikul & Siriwut & Srisonchai & Ng & Jirapatrasilp & Panha, 2023" authorityName="Sutcharit & Ngor & Páll-Gergely & Jeratthitikul & Siriwut & Srisonchai & Ng & Jirapatrasilp & Panha" authorityYear="2023" box="[791,938,887,912]" class="Gastropoda" family="Gastrocoptidae" genus="Gyliotrachela" kingdom="Animalia" order="Stylommatophora" pageId="12" phylum="Mollusca" rank="species" species="khmeriana" status="sp. nov.">
|
|||
|
|
<emphasis id="B93EEAB4FFF9CB62FCDDFC8AFC62BA84" box="[791,938,887,912]" italics="true" pageId="12">G. khmeriana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A20D57CFFFF9CB62FC7BFC8AFBC0BA84" box="[945,1032,887,912]" pageId="12" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
was found in lower abundances than the others, and only
|
|||
|
|
<specimenCount id="9D4CFD2FFFF9CB62FD7DFC67FCA8BAA7" box="[695,864,922,947]" pageId="12" type="generic">two specimens</specimenCount>
|
|||
|
|
were collected.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C350652DFFF9CB62FF6AFC13FDEEBF8F" pageId="12" type="diagnosis">
|
|||
|
|
<paragraph id="8BF536A6FFF9CB62FF6AFC13FE55BD1C" blockId="12.[160,1156,1006,1691]" box="[160,413,1006,1032]" pageId="12">
|
|||
|
|
<emphasis id="B93EEAB4FFF9CB62FF6AFC13FE55BD1C" bold="true" box="[160,413,1006,1032]" italics="true" pageId="12">Differential diagnosis</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BF536A6FFF9CB62FF6AFBEFFF2CBCBC" blockId="12.[160,1156,1006,1691]" pageId="12">
|
|||
|
|
This new species differs from all other known hypselostomatids from
|
|||
|
|
<collectingCountry id="F35D7636FFF9CB62FC69FBEFFBDEBD3F" box="[931,1046,1042,1067]" name="Cambodia" pageId="12">Cambodia</collectingCountry>
|
|||
|
|
in having slightly angular whorl, short and straight tuba, strong angular lamella, last whorl angular, protoconch with rounded pits, and spiral striations. In contrast,
|
|||
|
|
<taxonomicName id="4C4A4D25FFF9CB62FCB5FBAAFBE6BD64" authorityName="Benthem Jutting" authorityYear="1962" box="[895,1070,1111,1136]" class="Gastropoda" family="Gastrocoptidae" genus="Hypselostoma" kingdom="Animalia" order="Stylommatophora" pageId="12" phylum="Mollusca" rank="species" species="cambodjense">
|
|||
|
|
<emphasis id="B93EEAB4FFF9CB62FCB5FBAAFBE6BD64" box="[895,1070,1111,1136]" italics="true" pageId="12">H. cambodjense</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
has an elevated spire and a shouldered last whorl, is without angular lamella, and has an infraparietal plica;
|
|||
|
|
<taxonomicName id="4C4A4D25FFF9CB62FEBBFB61FD24BDA1" authority="Vermeulen et al., 2007" authorityName="Vermeulen" authorityYear="2007" box="[369,748,1180,1205]" class="Gastropoda" family="Hypselostomatidae" genus="Anauchen" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" phylum="Mollusca" rank="species" species="informis">
|
|||
|
|
<emphasis id="B93EEAB4FFF9CB62FEBBFB61FE2FBDA1" box="[369,487,1180,1205]" italics="true" pageId="12">A. informis</emphasis>
|
|||
|
|
<bibRefCitation id="EFDB4B57FFF9CB62FE25FB61FD24BDA1" author="Vermeulen JJ & Phung CL & Truong QT" box="[495,748,1180,1205]" pageId="12" pagination="81 - 92" refId="ref8340" refString="Vermeulen JJ, Phung CL, Truong QT. 2007. New species of terrestrial molluscs (Caenogastropoda, Pupinidae & Pulmonata: Vertiginidae) of the Hon Chong - ha Tien limestone hills, Southern Vietnam. Basteria. 71: 81 - 92." type="journal article" year="2007">
|
|||
|
|
Vermeulen
|
|||
|
|
<emphasis id="B93EEAB4FFF9CB62FDBBFB61FD6ABDA1" box="[625,674,1180,1205]" italics="true" pageId="12">et al</emphasis>
|
|||
|
|
., 2007
|
|||
|
|
</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
from
|
|||
|
|
<collectingCountry id="F35D7636FFF9CB62FCF8FB61FC47BDA1" box="[818,911,1180,1205]" name="Vietnam" pageId="12">Vietnam</collectingCountry>
|
|||
|
|
possesses a rounded last whorl, spiral striations on teleoconch, and adnate peristome, is without angular lamella, and has upper palatal and basal lamellae that are sometimes absent (
|
|||
|
|
<bibRefCitation id="EFDB4B57FFF9CB62FBD6FB1FFEFCBC09" author="Benthem Jutting WSS" pageId="12" pagination="3 - 15" refId="ref6458" refString="Benthem Jutting WSS van. 1962. Coquilles terrestres nouvelles de quelques collines calcaires du Cambodge et du Sud Vietnam. J Conchyl. 102: 3 - 15." type="journal article" year="1962">Benthem Jutting 1962</bibRefCitation>
|
|||
|
|
;
|
|||
|
|
<bibRefCitation id="EFDB4B57FFF9CB62FE8FFAF9FD84BC09" author="Vermeulen JJ & Phung CL & Truong QT" box="[325,588,1284,1309]" pageId="12" pagination="81 - 92" refId="ref8340" refString="Vermeulen JJ, Phung CL, Truong QT. 2007. New species of terrestrial molluscs (Caenogastropoda, Pupinidae & Pulmonata: Vertiginidae) of the Hon Chong - ha Tien limestone hills, Southern Vietnam. Basteria. 71: 81 - 92." type="journal article" year="2007">
|
|||
|
|
Vermeulen
|
|||
|
|
<emphasis id="B93EEAB4FFF9CB62FE07FAF9FDCABC09" box="[461,514,1284,1309]" italics="true" pageId="12">et al</emphasis>
|
|||
|
|
. 2007
|
|||
|
|
</bibRefCitation>
|
|||
|
|
). This new species also differs from
|
|||
|
|
<taxonomicName id="4C4A4D25FFF9CB62FBCDFAF9FE6CBC54" authority="Benthem Jutting, 1962" authorityName="Benthem Jutting" authorityYear="1962" class="Gastropoda" family="Gastrocoptidae" genus="Hypselostoma" kingdom="Animalia" order="Stylommatophora" pageId="12" phylum="Mollusca" rank="species" species="rupestre">
|
|||
|
|
<emphasis id="B93EEAB4FFF9CB62FBCDFAF9FB4BBC09" box="[1031,1155,1284,1309]" italics="true" pageId="12">H. rupestre</emphasis>
|
|||
|
|
<bibRefCitation id="EFDB4B57FFF9CB62FF6AFADAFE6CBC54" author="Benthem Jutting WSS" box="[160,420,1319,1344]" pageId="12" pagination="3 - 15" refId="ref6458" refString="Benthem Jutting WSS van. 1962. Coquilles terrestres nouvelles de quelques collines calcaires du Cambodge et du Sud Vietnam. J Conchyl. 102: 3 - 15." type="journal article" year="1962">Benthem Jutting, 1962</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C4A4D25FFF9CB62FE2BFADAFCB1BC54" authority="Benthem Jutting, 1962" authorityName="Benthem Jutting" authorityYear="1962" box="[481,889,1319,1344]" class="Gastropoda" family="Gastrocoptidae" genus="Hypselostoma" kingdom="Animalia" order="Stylommatophora" pageId="12" phylum="Mollusca" rank="species" species="dilatatum">
|
|||
|
|
<emphasis id="B93EEAB4FFF9CB62FE2BFADAFDA3BC54" box="[481,619,1319,1344]" italics="true" pageId="12">H. dilatatum</emphasis>
|
|||
|
|
<bibRefCitation id="EFDB4B57FFF9CB62FDBFFADAFCB1BC54" author="Benthem Jutting WSS" box="[629,889,1319,1344]" pageId="12" pagination="3 - 15" refId="ref6458" refString="Benthem Jutting WSS van. 1962. Coquilles terrestres nouvelles de quelques collines calcaires du Cambodge et du Sud Vietnam. J Conchyl. 102: 3 - 15." type="journal article" year="1962">Benthem Jutting, 1962</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
from
|
|||
|
|
<collectingCountry id="F35D7636FFF9CB62FC0BFADAFBE9BC54" box="[961,1057,1319,1344]" name="Vietnam" pageId="12">Vietnam</collectingCountry>
|
|||
|
|
, both of which possess a rounded last whorl, elongate conical spire, interpalatal and infrapalatal plicae, and a robust supracolumella, and are without angular lamella (
|
|||
|
|
<bibRefCitation id="EFDB4B57FFF9CB62FC0EFA91FF1FBCBC" author="Benthem Jutting WSS" pageId="12" pagination="3 - 15" refId="ref6458" refString="Benthem Jutting WSS van. 1962. Coquilles terrestres nouvelles de quelques collines calcaires du Cambodge et du Sud Vietnam. J Conchyl. 102: 3 - 15." type="journal article" year="1962">Benthem Jutting 1962</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BF536A6FFF9CB62FF0AFA4FFDEEBF8F" blockId="12.[160,1156,1006,1691]" pageId="12">
|
|||
|
|
In addition, two congeners,
|
|||
|
|
<taxonomicName id="4C4A4D25FFF9CB62FDCDFA4FFC56BCDF" authority="(Panha and Burch, 2002)" baseAuthorityName="Panha and Burch" baseAuthorityYear="2002" box="[519,926,1458,1483]" class="Gastropoda" family="Hydrocenidae" genus="Georissa" kingdom="Animalia" order="Cycloneritida" pageId="12" phylum="Mollusca" rank="species" species="tarutao">
|
|||
|
|
<emphasis id="B93EEAB4FFF9CB62FDCDFA4FFDB1BCDF" box="[519,633,1458,1483]" italics="true" pageId="12">G. tarutao</emphasis>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation id="EFDB4B57FFF9CB62FD42FA4FFC5EBCDF" author="Panha S & Burch JB" box="[648,918,1458,1483]" pageId="12" pagination="77 - 94" refId="ref7480" refString="Panha S, Burch JB. 2002. The Pupillidae genus Paraboysidia in Thailand (Pulmonata: Stylommatophora). Walkerana. 12 (28): 77 - 94. [2001]." type="journal article" year="2002">Panha and Burch, 2002</bibRefCitation>
|
|||
|
|
)
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C4A4D25FFF9CB62FC17FA4FFE97BCF9" authority="Panha and Burch" authorityName="Panha and Burch" class="Gastropoda" family="Hydrocenidae" genus="Georissa" kingdom="Animalia" order="Cycloneritida" pageId="12" phylum="Mollusca" rank="species" species="khaochakan">
|
|||
|
|
<emphasis id="B93EEAB4FFF9CB62FC17FA4FFB4BBCDF" box="[989,1155,1458,1483]" italics="true" pageId="12">G. khaochakan</emphasis>
|
|||
|
|
Panha and Burch
|
|||
|
|
</taxonomicName>
|
|||
|
|
in
|
|||
|
|
<bibRefCitation id="EFDB4B57FFF9CB62FE4BFA29FD82BCF9" author="Burch JB & Panha S & Tongkerd P" box="[385,586,1492,1517]" pageId="12" pagination="129 - 187" refId="ref6488" refString="Burch JB, Panha S, Tongkerd P. 2002 - 2003. New taxa of Pupillidae (Pulmonata: Stylommatophora) from Thailand. Walkerana. 13 (29 / 30): 129 - 187." type="journal article" year="2003">
|
|||
|
|
Burch
|
|||
|
|
<emphasis id="B93EEAB4FFF9CB62FE03FA29FE30BCF9" box="[457,504,1492,1517]" italics="true" pageId="12">et al</emphasis>
|
|||
|
|
. (2003)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
from
|
|||
|
|
<collectingCountry id="F35D7636FFF9CB62FD47FA29FD38BCF9" box="[653,752,1492,1517]" name="Thailand" pageId="12">Thailand</collectingCountry>
|
|||
|
|
, also differ from this new species. In comparison,
|
|||
|
|
<taxonomicName id="4C4A4D25FFF9CB62FEF9FA0AFE6BBF04" baseAuthorityName="Panha and Burch" baseAuthorityYear="2002" box="[307,419,1527,1552]" class="Gastropoda" family="Hydrocenidae" genus="Georissa" kingdom="Animalia" order="Cycloneritida" pageId="12" phylum="Mollusca" rank="species" species="tarutao">
|
|||
|
|
<emphasis id="B93EEAB4FFF9CB62FEF9FA0AFE6BBF04" box="[307,419,1527,1552]" italics="true" pageId="12">G. tarutao</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
has a very short tuba, elongated conical spire, umbilical furrow extended to an expanded lip, and infraparietal and subcolumellar plicae present, while
|
|||
|
|
<taxonomicName id="4C4A4D25FFF9CB62FF6AF9C1FE81BF41" box="[160,329,1596,1621]" class="Gastropoda" family="Hydrocenidae" genus="Georissa" kingdom="Animalia" order="Cycloneritida" pageId="12" phylum="Mollusca" rank="species" species="khaochakan">
|
|||
|
|
<emphasis id="B93EEAB4FFF9CB62FF6AF9C1FE81BF41" box="[160,329,1596,1621]" italics="true" pageId="12">G. khaochakan</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
possesses an elongated conical spire, spiral striations present on the entirety of the shell, the last whorl shouldered, and an infrapalatal plica present (
|
|||
|
|
<bibRefCitation id="EFDB4B57FFF9CB62FBF6F9A2FE99BF8F" author="Panha S & Burch JB" pageId="12" pagination="77 - 94" refId="ref7480" refString="Panha S, Burch JB. 2002. The Pupillidae genus Paraboysidia in Thailand (Pulmonata: Stylommatophora). Walkerana. 12 (28): 77 - 94. [2001]." type="journal article" year="2002">Panha and Burch 2002</bibRefCitation>
|
|||
|
|
;
|
|||
|
|
<bibRefCitation id="EFDB4B57FFF9CB62FE94F97FFDD2BF8F" author="Burch JB & Panha S & Tongkerd P" box="[350,538,1666,1691]" pageId="12" pagination="129 - 187" refId="ref6488" refString="Burch JB, Panha S, Tongkerd P. 2002 - 2003. New taxa of Pupillidae (Pulmonata: Stylommatophora) from Thailand. Walkerana. 13 (29 / 30): 129 - 187." type="journal article" year="2003">
|
|||
|
|
Burch
|
|||
|
|
<emphasis id="B93EEAB4FFF9CB62FE6CF97FFE1EBF8F" box="[422,470,1666,1691]" italics="true" pageId="12">et al</emphasis>
|
|||
|
|
. 2003
|
|||
|
|
</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C350652DFFF8CB63FF6AFF73FD17BBC7" pageId="13" type="discussion">
|
|||
|
|
<paragraph id="8BF536A6FFF8CB63FF6AFF73FECEB9BC" blockId="13.[160,1156,142,723]" box="[160,262,142,168]" pageId="13">
|
|||
|
|
<emphasis id="B93EEAB4FFF8CB63FF6AFF73FECEB9BC" bold="true" box="[160,262,142,168]" italics="true" pageId="13">Remarks</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BF536A6FFF8CB63FF6AFF4FFB97B86C" blockId="13.[160,1156,142,723]" pageId="13">
|
|||
|
|
The type locality, Phnom Sampeau, is located about
|
|||
|
|
<quantity id="4CB29B43FFF8CB63FD24FF4FFCFCB9DF" box="[750,820,178,203]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="13" unit="km" value="13.0">13 km</quantity>
|
|||
|
|
west of
|
|||
|
|
<collectingRegion id="498EF844FFF8CB63FC5EFF4FFBD6B9DF" box="[916,1054,178,203]" country="Cambodia" name="Battambang" pageId="13">Battambang</collectingRegion>
|
|||
|
|
town on road no. 57 leading to
|
|||
|
|
<collectingRegion id="498EF844FFF8CB63FE53FF29FDF7B9F9" box="[409,575,212,237]" country="Cambodia" name="Pailin" pageId="13">Pailin Province</collectingRegion>
|
|||
|
|
. The mountain is of karstic limestone with limestone forest and is surrounded by villages, plantation areas and paddy fields. Conspicuous human disturbance includes the presence of a temple with complex tourist attractions, such as a bat cave and a memorial commemorating the grisly history of the caves during the previous civil war, and small-scale limestone quarries around the foot of the hills.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BF536A6FFF8CB63FF0AFE7FFD17BBC7" blockId="13.[160,1156,142,723]" pageId="13">
|
|||
|
|
The difference between the genera
|
|||
|
|
<taxonomicName id="4C4A4D25FFF8CB63FDA2FE7FFD3FB88F" authorityName="Tomlin" authorityYear="1930" box="[616,759,386,411]" class="Gastropoda" family="Gastrocoptidae" genus="Gyliotrachela" kingdom="Animalia" order="Stylommatophora" pageId="13" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B93EEAB4FFF8CB63FDA2FE7FFD3FB88F" box="[616,759,386,411]" italics="true" pageId="13">Gyliotrachela</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C4A4D25FFF8CB63FCF2FE7FFBA9B88F" authority="Pilsbry, 1917" authorityName="Pilsbry" authorityYear="1917" box="[824,1121,386,411]" class="Gastropoda" family="Gastrocoptidae" genus="Paraboysidia" kingdom="Animalia" order="Stylommatophora" pageId="13" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B93EEAB4FFF8CB63FCF2FE7FFC0CB88F" box="[824,964,386,411]" italics="true" pageId="13">Paraboysidia</emphasis>
|
|||
|
|
<bibRefCitation id="EFDB4B57FFF8CB63FC05FE7FFBA9B88F" author="Pilsbry HA" box="[975,1121,386,411]" pageId="13" refId="ref7609" refString="Pilsbry HA. 1917. Manual of Conchology, Structure and Systematic, with Illustrations of the Species, 2 nd Series. Vol. 24. Pupillidae (Gastrocoptinae). Philadelphia (PA): Academy of Natural Science; Pupillidae (Gastrocoptinae)." type="book" year="1917">Pilsbry, 1917</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
[=
|
|||
|
|
<taxonomicName id="4C4A4D25FFF8CB63FF6AFE59FD9AB8A9" authority="Pilsbry and Vanatta, 1900" authorityName="Pilsbry and Vanatta" authorityYear="1900" box="[160,594,420,445]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="13" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B93EEAB4FFF8CB63FF6AFE59FEDFB8A9" box="[160,279,420,445]" italics="true" pageId="13">Bensonella</emphasis>
|
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|
|
<bibRefCitation id="EFDB4B57FFF8CB63FEE9FE59FD9AB8A9" author="Pilsbry HA & Vanatta EG" box="[291,594,420,445]" pageId="13" pagination="582 - 611" refId="ref7656" refString="Pilsbry HA, Vanatta EG. 1900. A partial revision of the Pupae of the United States. Proc Acad Nat Sci Philadelphia. 52: 582 - 611." type="journal article" year="1900">Pilsbry and Vanatta, 1900</bibRefCitation>
|
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</taxonomicName>
|
|||
|
|
; see
|
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|
|
<bibRefCitation id="EFDB4B57FFF8CB63FD5CFE59FC30B8A9" author="Pall-Gergely B & White TS" box="[662,1016,420,445]" pageId="13" pagination="45 - 48" refId="ref7434" refString="Pall-Gergely B, White TS. 2023. Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). J Nat Hist. 56: 45 - 48. doi: 10. 1080 / 00222933.2022.2152750" type="journal article" year="2023">Páll-Gergely and White (2023)</bibRefCitation>
|
|||
|
|
] lies in the detached aperture in the former and the adnate aperture in the latter. However, as
|
|||
|
|
<bibRefCitation id="EFDB4B57FFF8CB63FF6AFE17FEF6BB17" author="Pilsbry HA" box="[160,318,490,515]" pageId="13" refId="ref7609" refString="Pilsbry HA. 1917. Manual of Conchology, Structure and Systematic, with Illustrations of the Species, 2 nd Series. Vol. 24. Pupillidae (Gastrocoptinae). Philadelphia (PA): Academy of Natural Science; Pupillidae (Gastrocoptinae)." type="book" year="1917">Pilsbry (1917)</bibRefCitation>
|
|||
|
|
already suspected, there is a nearly perfect morphological continuum between the two extremes, and
|
|||
|
|
<taxonomicName id="4C4A4D25FFF8CB63FDC5FDF1FCD3BB31" authority="Sutcharit & Ngor & Páll-Gergely & Jeratthitikul & Siriwut & Srisonchai & Ng & Jirapatrasilp & Panha, 2023" authorityName="Sutcharit & Ngor & Páll-Gergely & Jeratthitikul & Siriwut & Srisonchai & Ng & Jirapatrasilp & Panha" authorityYear="2023" box="[527,795,524,549]" class="Gastropoda" family="Gastrocoptidae" genus="Gyliotrachela" kingdom="Animalia" order="Stylommatophora" pageId="13" phylum="Mollusca" rank="species" species="khmeriana" status="sp. nov.">
|
|||
|
|
<emphasis id="B93EEAB4FFF8CB63FDC5FDF1FCD3BB31" box="[527,795,524,549]" italics="true" pageId="13">Gyliotrachela khmeriana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A20D57CFFFF8CB63FCE8FDF1FCB2BB31" box="[802,890,524,549]" pageId="13" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
can also be considered an intermediate character state. This new species is similar to some
|
|||
|
|
<taxonomicName id="4C4A4D25FFF8CB63FC55FDD2FB4BBB5C" box="[927,1155,559,584]" pageId="13">
|
|||
|
|
<emphasis id="B93EEAB4FFF8CB63FC55FDD2FBE3BB5C" box="[927,1067,559,584]" italics="true" pageId="13">Paraboysidia</emphasis>
|
|||
|
|
species
|
|||
|
|
</taxonomicName>
|
|||
|
|
from the Malaysian Peninsula, mostly in terms of the arrangement of the apertural barriers. The most similar species is probably
|
|||
|
|
<taxonomicName id="4C4A4D25FFF8CB63FD6CFD89FBB6BB99" authority="Benthem Jutting, 1950" authorityName="Benthem Jutting" authorityYear="1950" box="[678,1150,628,653]" class="Gastropoda" family="Gastrocoptidae" genus="Paraboysidia" kingdom="Animalia" order="Stylommatophora" pageId="13" phylum="Mollusca" rank="species" species="serpa">
|
|||
|
|
<emphasis id="B93EEAB4FFF8CB63FD6CFD89FCBCBB99" box="[678,884,628,653]" italics="true" pageId="13">Paraboysidia serpa</emphasis>
|
|||
|
|
Benthem Jutting, 1950
|
|||
|
|
</taxonomicName>
|
|||
|
|
, which also has a keel on its body whorl. However, the new species has a more strongly detached aperture, and more keeled upper whorls.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|