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<document ID-DOI="10.5252/a2015n2a2" ID-ISSN="1639-4798" ID-Zenodo-Dep="5208903" approvalRequired="50" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="44" approvalRequired_for_treatments="5" checkinTime="1629150308770" checkinUser="carolina" docAuthor="Daly, Douglas C., Raharimampionona, Jeannie &amp; Federman, Sarah" docDate="2015" docId="03F5FA39BC49FFECB1D1FACA33AEF98C" docLanguage="en" docName="Adansonia.37.2.277-345.pdf" docOrigin="Adansonia 37 (2)" docSource="https://www.mendeley.com/catalogue/2522e05a-0ab0-3e53-96aa-d45147a2ca6c/" docStyle="DocumentStyle:E7A860AFEA63D44BCC37955D07FCAFC7.2:Adansonia.2015-2017.journal_article" docStyleId="E7A860AFEA63D44BCC37955D07FCAFC7" docStyleName="Adansonia.2015-2017.journal_article" docStyleVersion="2" docTitle="Canarium betamponae Daly, Raharim. &amp; Federman 2015, sp. nov." docType="treatment" docVersion="2" lastPageId="18" lastPageNumber="295" masterDocId="FFCC8241BC58FFFEB157FF99310AFFA1" masterDocTitle="A revision of Canarium L. (Burseraceae) in Madagascar" masterLastPageNumber="345" masterPageNumber="277" pageId="17" pageNumber="294" updateTime="1629150974147" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of Canarium L. (Burseraceae) in Madagascar</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Daly, Douglas C.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Raharimampionona, Jeannie</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Federman, Sarah</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Adansonia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="pubDate">
<mods:number>2015-12-31</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>37</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>277</mods:start>
<mods:end>345</mods:end>
</mods:extent>
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<mods:location>
<mods:url>https://www.mendeley.com/catalogue/2522e05a-0ab0-3e53-96aa-d45147a2ca6c/</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5252/a2015n2a2</mods:identifier>
<mods:identifier type="ISSN">1639-4798</mods:identifier>
<mods:identifier type="Zenodo-Dep">5208903</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03F5FA39BC49FFECB1D1FACA33AEF98C" httpUri="http://treatment.plazi.org/id/03F5FA39BC49FFECB1D1FACA33AEF98C" lastPageId="18" lastPageNumber="295" pageId="17" pageNumber="294">
<subSubSection box="[134,773,1363,1390]" pageId="17" pageNumber="294" type="nomenclature">
<paragraph blockId="17.[134,773,1363,1421]" box="[134,773,1363,1390]" pageId="17" pageNumber="294">
<heading box="[134,773,1363,1390]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="294" reason="3">
<taxonomicName authority="Daly, Raharim. &amp; Federman" authorityName="Daly, Raharim. &amp; Federman" authorityYear="2015" box="[134,683,1363,1390]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="17" pageNumber="294" phylum="Tracheophyta" rank="species" species="betamponae" status="sp. nov.">
<emphasis bold="true" box="[134,375,1363,1390]" italics="true" pageId="17" pageNumber="294">Canarium betamponae</emphasis>
Daly, Raharim. &amp; Federman
</taxonomicName>
,
<taxonomicNameLabel box="[692,773,1364,1390]" pageId="17" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="18" lastPageNumber="295" pageId="17" pageNumber="294" type="description">
<paragraph blockId="17.[134,773,1363,1421]" box="[371,537,1395,1421]" pageId="17" pageNumber="294">
(
<figureCitation box="[378,446,1395,1421]" captionStart="FIG" captionStartId="15.[132,143,1228,1246]" captionTargetBox="[131,775,215,1191]" captionTargetId="figure-739@15.[132,775,216,1191]" captionTargetPageId="15" captionText="FIG. 6. — Distributions of Canarium betamponae Daly, Raharim. &amp; Federman, sp. nov., C. bullatum (Leenh.) Daly, Raharim. &amp; Federman, comb. et stat. nov., C. egregium Daly, Raharim. &amp; Federman, sp. nov., and C. indistinctum Daly, Raharim. &amp; Federman, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5208913" httpUri="https://zenodo.org/record/5208913/files/figure.png" pageId="17" pageNumber="294">Figs 6</figureCitation>
;
<figureCitation box="[458,471,1395,1421]" captionStart="FIG" captionStartId="16.[132,143,1939,1957]" captionTargetBox="[206,1378,214,1866]" captionTargetId="figure-19@16.[205,1383,208,1871]" captionTargetPageId="16" captionText="FIG. 7. — A-D, Canarium betamponae Daly, Raharim. &amp; Federman, sp. nov.: A, fruiting branchlet, with detail of leaflet (A’); B, stipules; C, cross-section of fruit (above) and pyrene; D, detail of infructescence; E, F, Canarium moramangae Daly, Raharim. &amp; Federman, sp. nov.: E, fruiting branchlet, with detail of leaflet (E’); F, stipule and scar; A, C, Daly et al. 13079 (NY sheet and field photos); B, Federman et al. 116 (NY); D, Daly et al. 13073 (field photo by D. Daly); E, F, Service Forestier-21894 no. 1748-R-100. Scale bars: A, C-E, 2 cm; B, F, 0.5 cm; A’, E’, not to scale." figureDoi="http://doi.org/10.5281/zenodo.5208915" httpUri="https://zenodo.org/record/5208915/files/figure.png" pageId="17" pageNumber="294">7</figureCitation>
A-D).
</paragraph>
<paragraph blockId="17.[131,777,1454,2027]" pageId="17" pageNumber="294">
<emphasis italics="true" pageId="17" pageNumber="294">Small to medium-sized trees, leaves 3-4(5)-jugate, fruits broadly obovoid (rarely slightly ovoid), glabrous and glossy, distinguished from</emphasis>
<taxonomicName authority="Daly, Raharim. &amp; Federman" authorityName="Daly, Raharim. &amp; Federman" authorityYear="2015" box="[132,515,1507,1531]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="17" pageNumber="294" phylum="Tracheophyta" rank="species" species="galokense" status="sp. nov.">
C. galokense
<emphasis box="[253,515,1507,1531]" italics="true" pageId="17" pageNumber="294">Daly, Raharim. &amp; Federman</emphasis>
</taxonomicName>
<emphasis box="[515,595,1507,1531]" italics="true" pageId="17" pageNumber="294">
,
<taxonomicNameLabel box="[522,592,1508,1531]" pageId="17" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
<taxonomicName authority="Daly, Raharim. &amp; Federman" authorityName="Daly, Raharim. &amp; Federman" authorityYear="2015" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="17" pageNumber="294" phylum="Tracheophyta" rank="species" species="globosum" status="sp. nov.">
C. globosum
<emphasis italics="true" pageId="17" pageNumber="294">Daly, Raharim. &amp; Federman</emphasis>
</taxonomicName>
<emphasis box="[343,425,1533,1557]" italics="true" pageId="17" pageNumber="294">
,
<taxonomicNameLabel box="[351,422,1534,1557]" pageId="17" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
<taxonomicName authority="Daly, Raharim. &amp; Federman" authorityName="Daly, Raharim. &amp; Federman" authorityYear="2015" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="17" pageNumber="294" phylum="Tracheophyta" rank="species" species="manongarivum" status="sp. nov.">
C. manongarivum
<emphasis italics="true" pageId="17" pageNumber="294">Daly, Raharim. &amp; Federman</emphasis>
</taxonomicName>
<emphasis box="[227,344,1560,1584]" italics="true" pageId="17" pageNumber="294">
,
<taxonomicNameLabel box="[235,304,1561,1584]" pageId="17" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
and
</emphasis>
<taxonomicName authority="Daly, Raharim. &amp; Federman" authorityName="Daly, Raharim. &amp; Federman" authorityYear="2015" box="[350,769,1560,1584]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="17" pageNumber="294" phylum="Tracheophyta" rank="species" species="subsidarium" status="sp. nov.">
C. subsidarium
<emphasis box="[500,769,1560,1584]" italics="true" pageId="17" pageNumber="294">Daly, Raharim. &amp; Federman</emphasis>
</taxonomicName>
<emphasis italics="true" pageId="17" pageNumber="294">
,
<taxonomicNameLabel box="[131,201,1587,1610]" pageId="17" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
by membranaceous stipules and subglabrous inflorescence axes.
</emphasis>
<typeStatus box="[132,194,1636,1659]" pageId="17" pageNumber="294">TYPUS</typeStatus>
. —
<emphasis bold="true" box="[239,363,1636,1660]" pageId="17" pageNumber="294">
<collectingCountry box="[239,358,1636,1660]" name="Madagascar" pageId="17" pageNumber="294">Madagascar</collectingCountry>
.
</emphasis>
<collectingRegion box="[370,474,1636,1660]" country="Madagascar" name="Toamasina" pageId="17" pageNumber="294">Toamasina</collectingRegion>
<emphasis bold="true" box="[474,483,1637,1661]" pageId="17" pageNumber="294">/</emphasis>
Tamatave, District
<collectingRegion box="[670,775,1636,1660]" country="Madagascar" name="Toamasina" pageId="17" pageNumber="294">Toamasina</collectingRegion>
II, Ambodirina, Réserve Naturelle Intégrale Betampona,
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elev.,
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,
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,
<date box="[516,609,1689,1713]" pageId="17" pageNumber="294" value="2006-02-03">3.II.2006</date>
(fr),
<emphasis italics="true" pageId="17" pageNumber="294">D. C. Daly, R. Ranaivojaona, R. B. Iambana, &amp; C. Razafy 13073</emphasis>
(holo-, NY!; iso-, MO!, TEF!).
</paragraph>
<paragraph blockId="17.[131,777,1454,2027]" lastBlockId="17.[811,1460,215,652]" pageId="17" pageNumber="294">
<materialsCitation collectingDate="1956-02-11" collectingDateMax="2008-05-10" collectingDateMin="1956-02-11" collectionCode="NY, TEF" collectorName="D. C. Daly &amp; R. Ranaivojaona &amp; R. B. Iambana &amp; C. Razafy &amp; S. Federman &amp; T. Treuer &amp; J. Sylvain &amp; Ambohimanga &amp; Ambaro &amp; Ifanadiana &amp; Service Forestier &amp; Atsinanana &amp; Reserve Naturelle Integrale Betampona &amp; To" country="Madagascar" county="Atsinanana" elevation="350" latitude="-17.91" location="Naturelle Integrale de Betampona" longLatPrecision="21" longitude="49.202778" municipality="Reserve Naturelle Integrale Betampona" pageId="17" pageNumber="294" specimenCount="1" stateProvince="Toamasina" typeStatus="paratype">
<typeStatus box="[132,221,1792,1815]" pageId="17" pageNumber="294">PARATYPI</typeStatus>
. —
<emphasis bold="true" box="[267,390,1791,1815]" pageId="17" pageNumber="294">
<collectingCountry box="[267,386,1791,1815]" name="Madagascar" pageId="17" pageNumber="294">Madagascar</collectingCountry>
.
</emphasis>
<collectingRegion box="[396,503,1791,1815]" country="Madagascar" name="Toamasina" pageId="17" pageNumber="294">Toamasina</collectingRegion>
,
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,
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,
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,
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,
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<collectingDate box="[134,229,1844,1868]" pageId="17" pageNumber="294" value="2006-02-04">4.II.2006</collectingDate>
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,
<emphasis italics="true" pageId="17" pageNumber="294">
<collectorName box="[237,341,1844,1868]" pageId="17" pageNumber="294">D. C. Daly</collectorName>
,
<collectorName box="[350,509,1845,1868]" pageId="17" pageNumber="294">R. Ranaivojaona</collectorName>
,
<collectorName box="[517,654,1844,1868]" pageId="17" pageNumber="294">R. B. Iambana</collectorName>
&amp;
<collectorName box="[684,774,1844,1868]" pageId="17" pageNumber="294">C. Razafy</collectorName>
13079
</emphasis>
(
<collectionCode box="[206,242,1871,1895]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="17" pageNumber="294" type="Herbarium">NY</collectionCode>
,
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),
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<elevation box="[313,422,1870,1894]" metricMagnitude="2" metricUnit="m" metricValue="3.5" metricValueMax="3.75" metricValueMin="3.25" pageId="17" pageNumber="294" unit="m" value="350.0" valueMax="375.0" valueMin="325.0">325-375 m</elevation>
</quantity>
,
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,
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,
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<collectingDate box="[677,770,1870,1894]" pageId="17" pageNumber="294" value="2006-02-04">4.II.2006</collectingDate>
</date>
,
<emphasis box="[131,775,1897,1921]" italics="true" pageId="17" pageNumber="294">
<collectorName box="[131,242,1897,1921]" pageId="17" pageNumber="294">D. C. Daly</collectorName>
,
<collectorName box="[253,416,1898,1921]" pageId="17" pageNumber="294">R. Ranaivojaona</collectorName>
,
<collectorName box="[427,571,1897,1921]" pageId="17" pageNumber="294">R. B. Iambana</collectorName>
&amp;
<collectorName box="[608,703,1897,1921]" pageId="17" pageNumber="294">C. Razafy</collectorName>
13080
</emphasis>
(
<collectionCode box="[139,175,1924,1948]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="17" pageNumber="294" type="Herbarium">NY</collectionCode>
,
<collectionCode box="[185,234,1924,1948]" country="Madagascar" lsid="urn:lsid:biocol.org:col:12774" name="Centre National de la Recherche Appliquée au Developement Rural" pageId="17" pageNumber="294" type="Herbarium">TEF</collectionCode>
),
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<elevation box="[252,366,1923,1947]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="17" pageNumber="294" unit="m" value="500.0" valueMax="600.0" valueMin="400.0">400-600 m</elevation>
</quantity>
,
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,
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,
<date box="[632,745,1923,1947]" pageId="17" pageNumber="294" value="2008-04-15">
<collectingDate box="[632,745,1923,1947]" pageId="17" pageNumber="294" value="2008-04-15">15.IV.2008</collectingDate>
</date>
,
<emphasis italics="true" pageId="17" pageNumber="294">
<collectorName pageId="17" pageNumber="294">S. Federman</collectorName>
,
<collectorName box="[237,322,1951,1974]" pageId="17" pageNumber="294">T. Treuer</collectorName>
&amp;
<collectorName box="[354,444,1950,1974]" pageId="17" pageNumber="294">J. Sylvain</collectorName>
9
</emphasis>
(
<collectionCode box="[477,516,1950,1974]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="17" pageNumber="294" type="Herbarium">NY</collectionCode>
),
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<elevation box="[533,646,1950,1974]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="17" pageNumber="294" unit="m" value="500.0" valueMax="600.0" valueMin="400.0">400-600 m</elevation>
</quantity>
,
<geoCoordinate box="[656,770,1950,1974]" degrees="17" direction="south" minutes="54" orientation="latitude" pageId="17" pageNumber="294" precision="15" seconds="33" value="-17.909166">17°54’33”S</geoCoordinate>
,
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,
<date box="[255,357,1976,2000]" pageId="17" pageNumber="294" value="2008-03-07">
<collectingDate box="[255,357,1976,2000]" pageId="17" pageNumber="294" value="2008-03-07">7.III.2008</collectingDate>
</date>
,
<emphasis box="[364,713,1976,2000]" italics="true" pageId="17" pageNumber="294">
<collectorName box="[364,481,1976,2000]" pageId="17" pageNumber="294">S. Federman</collectorName>
,
<collectorName box="[487,568,1977,2000]" pageId="17" pageNumber="294">T.Treuer</collectorName>
&amp;
<collectorName box="[597,683,1976,2000]" pageId="17" pageNumber="294">J. Sylvain</collectorName>
28
</emphasis>
(
<collectionCode box="[724,762,1976,2000]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="17" pageNumber="294" type="Herbarium">NY</collectionCode>
),
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<elevation box="[132,245,2003,2027]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="17" pageNumber="294" unit="m" value="500.0" valueMax="600.0" valueMin="400.0">400-600 m</elevation>
</quantity>
,
<geoCoordinate box="[256,371,2003,2027]" degrees="17" direction="south" minutes="54" orientation="latitude" pageId="17" pageNumber="294" precision="15" seconds="29" value="-17.908054">17°54’29”S</geoCoordinate>
,
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,
<date box="[510,614,2003,2027]" pageId="17" pageNumber="294" value="2008-03-09">
<collectingDate box="[510,614,2003,2027]" pageId="17" pageNumber="294" value="2008-03-09">9.III.2008</collectingDate>
</date>
,
<emphasis box="[625,1021,215,2027]" italics="true" pageId="17" pageNumber="294">
<collectorName box="[625,747,2003,2027]" pageId="17" pageNumber="294">S. Federman</collectorName>
,
<collectorName box="[756,872,216,2027]" pageId="17" pageNumber="294">T. Treuer</collectorName>
&amp;
<collectorName box="[902,991,215,239]" pageId="17" pageNumber="294">J. Sylvain</collectorName>
53
</emphasis>
(
<collectionCode box="[1033,1072,215,239]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="17" pageNumber="294" type="Herbarium">NY</collectionCode>
),
<quantity box="[1089,1200,215,239]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="17" pageNumber="294" unit="m" value="500.0" valueMax="600.0" valueMin="400.0">
<elevation box="[1089,1200,215,239]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="17" pageNumber="294" unit="m" value="500.0" valueMax="600.0" valueMin="400.0">400-600 m</elevation>
</quantity>
,
<geoCoordinate box="[1209,1324,215,239]" degrees="17" direction="south" minutes="54" orientation="latitude" pageId="17" pageNumber="294" precision="15" seconds="30" value="-17.908333">17°54’30”S</geoCoordinate>
,
<geoCoordinate box="[1333,1450,215,239]" degrees="49" direction="east" minutes="12" orientation="longitude" pageId="17" pageNumber="294" precision="15" seconds="14" value="49.203888">49°12’14”E</geoCoordinate>
,
<date box="[811,925,241,265]" pageId="17" pageNumber="294" value="2008-04-23">
<collectingDate box="[811,925,241,265]" pageId="17" pageNumber="294" value="2008-04-23">23.IV.2008</collectingDate>
</date>
,
<emphasis box="[934,1301,241,265]" italics="true" pageId="17" pageNumber="294">
<collectorName box="[934,1053,241,265]" pageId="17" pageNumber="294">S. Federman</collectorName>
,
<collectorName box="[1060,1142,242,265]" pageId="17" pageNumber="294">T.Treuer</collectorName>
&amp;
<collectorName box="[1171,1258,241,265]" pageId="17" pageNumber="294">J. Sylvain</collectorName>
116
</emphasis>
(
<collectionCode box="[1311,1350,241,265]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="17" pageNumber="294" type="Herbarium">NY</collectionCode>
);
<collectorName pageId="17" pageNumber="294">Ambohimanga</collectorName>
,
<collectorName box="[889,971,268,292]" pageId="17" pageNumber="294">Ambaro</collectorName>
,
<collectorName box="[979,1084,268,292]" pageId="17" pageNumber="294">Ifanadiana</collectorName>
,
<date box="[1093,1200,268,292]" pageId="17" pageNumber="294" value="1956-02-11">
<collectingDate box="[1093,1200,268,292]" pageId="17" pageNumber="294" value="1956-02-11">11.II.1956</collectingDate>
</date>
,
<emphasis box="[1207,1455,268,292]" italics="true" pageId="17" pageNumber="294">
<collectorName box="[1207,1355,268,291]" pageId="17" pageNumber="294">Service Forestier</collectorName>
20-R-332
</emphasis>
[sic] (
<collectionCode box="[865,901,294,318]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="17" pageNumber="294" type="Herbarium">NY</collectionCode>
,
<collectionCode box="[909,954,295,318]" country="Madagascar" lsid="urn:lsid:biocol.org:col:12774" name="Centre National de la Recherche Appliquée au Developement Rural" pageId="17" pageNumber="294" type="Herbarium">TEF</collectionCode>
[not seen]);
<collectorName box="[1075,1187,294,318]" pageId="17" pageNumber="294">Atsinanana</collectorName>
,
<collectorName pageId="17" pageNumber="294">Réserve Naturelle Intégrale Betampona</collectorName>
,
<quantity box="[939,1056,321,345]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="17" pageNumber="294" unit="m" value="500.0" valueMax="600.0" valueMin="400.0">
<elevation box="[939,1056,321,345]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="17" pageNumber="294" unit="m" value="500.0" valueMax="600.0" valueMin="400.0">400-600 m</elevation>
</quantity>
,
<geoCoordinate box="[1069,1185,321,345]" degrees="17" direction="south" minutes="53" orientation="latitude" pageId="17" pageNumber="294" precision="15" seconds="48" value="-17.896667">17°53’48”S</geoCoordinate>
,
<geoCoordinate box="[1198,1317,321,345]" degrees="49" direction="east" minutes="13" orientation="longitude" pageId="17" pageNumber="294" precision="15" seconds="11" value="49.219723">49°13’11”E</geoCoordinate>
,
<date box="[1329,1423,321,345]" pageId="17" pageNumber="294" value="2008-05-04">
<collectingDate box="[1329,1423,321,345]" pageId="17" pageNumber="294" value="2008-05-04">4.V.2008</collectingDate>
</date>
,
<emphasis italics="true" pageId="17" pageNumber="294">
<collectorName pageId="17" pageNumber="294">T. Treuer</collectorName>
,
<collectorName box="[883,999,347,371]" pageId="17" pageNumber="294">S. Federman</collectorName>
&amp;
<collectorName box="[1029,1118,347,371]" pageId="17" pageNumber="294">J. Sylvain</collectorName>
70
</emphasis>
(
<collectionCode box="[1160,1199,347,371]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="17" pageNumber="294" type="Herbarium">NY</collectionCode>
),
<quantity box="[1215,1326,347,371]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="17" pageNumber="294" unit="m" value="500.0" valueMax="600.0" valueMin="400.0">
<elevation box="[1215,1326,347,371]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="17" pageNumber="294" unit="m" value="500.0" valueMax="600.0" valueMin="400.0">400-600 m</elevation>
</quantity>
,
<geoCoordinate box="[1335,1450,347,371]" degrees="17" direction="south" minutes="54" orientation="latitude" pageId="17" pageNumber="294" precision="15" seconds="36" value="-17.91">17°54’36”S</geoCoordinate>
,
<geoCoordinate box="[811,924,374,398]" degrees="49" direction="east" minutes="12" orientation="longitude" pageId="17" pageNumber="294" precision="15" seconds="10" value="49.202778">49°12’10”E</geoCoordinate>
,
<date box="[932,1034,374,398]" pageId="17" pageNumber="294" value="2008-05-10">
<collectingDate box="[932,1034,374,398]" pageId="17" pageNumber="294" value="2008-05-10">10.V.2008</collectingDate>
</date>
,
<emphasis box="[1041,1394,374,398]" italics="true" pageId="17" pageNumber="294">
<collectorName box="[1041,1121,374,397]" pageId="17" pageNumber="294">T.Treuer</collectorName>
,
<collectorName box="[1129,1241,374,398]" pageId="17" pageNumber="294">S. Federman</collectorName>
&amp;
<collectorName box="[1268,1353,374,398]" pageId="17" pageNumber="294">J. Sylvain</collectorName>
110
</emphasis>
(
<collectionCode box="[1405,1443,374,398]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="17" pageNumber="294" type="Herbarium">NY</collectionCode>
). DISTRIBUTION AND ECOLOGY. —
<collectorName box="[1145,1171,422,445]" pageId="17" pageNumber="294">To</collectorName>
date,
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[1232,1449,421,445]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="17" pageNumber="294" phylum="Tracheophyta" rank="species" species="betamponae" status="sp. nov.">
<emphasis box="[1232,1449,421,445]" italics="true" pageId="17" pageNumber="294">Canarium betamponae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[813,885,448,471]" pageId="17" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
appears to be restricted to theRéserve
<location LSID="urn:lsid:plazi:treatment:03F5FA39BC49FFECB1D1FACA33AEF98C:8E831DF4BC49FFEFB588FE593297FE53" country="Madagascar" county="Atsinanana" latitude="-17.91" longLatPrecision="21" longitude="49.202778" municipality="Reserve Naturelle Integrale Betampona" name="Naturelle Integrale de Betampona" pageId="17" pageNumber="294" stateProvince="Toamasina">Naturelle Intégrale de Betampona</location>
in
</materialsCitation>
<collectingRegion box="[955,1062,474,498]" country="Madagascar" name="Toamasina" pageId="17" pageNumber="294">Toamasina</collectingRegion>
, where it is usually a large tree occurring on sandy clay soils in primary humid forest with open canopy and many poles and lianas; it is found between
<quantity box="[1244,1356,527,551]" metricMagnitude="2" metricUnit="m" metricValue="4.625" metricValueMax="6.0" metricValueMin="3.25" pageId="17" pageNumber="294" unit="m" value="462.5" valueMax="600.0" valueMin="325.0">325-600 m</quantity>
on slopes usually not exposed to cyclones. Fruiting Feb.-Apr.
</paragraph>
<paragraph blockId="17.[811,1460,215,652]" pageId="17" pageNumber="294">
COMMON NAMES AND USES. — Ramy. Resin used for caulking canoes, also for illumination, and for religious (
<emphasis box="[1255,1322,628,652]" italics="true" pageId="17" pageNumber="294">tromba</emphasis>
) ceremonies.
</paragraph>
<paragraph blockId="17.[810,1457,694,2028]" box="[813,957,694,720]" pageId="17" pageNumber="294">DESCRIPTION</paragraph>
<paragraph blockId="17.[810,1457,694,2028]" lastBlockId="18.[132,775,1364,1581]" lastPageId="18" lastPageNumber="295" pageId="17" pageNumber="294">
Trees, reproductive size
<quantity box="[1057,1148,726,752]" metricMagnitude="1" metricUnit="m" metricValue="1.65" metricValueMax="2.2" metricValueMin="1.1" pageId="17" pageNumber="294" unit="m" value="16.5" valueMax="22.0" valueMin="11.0">11-22 m</quantity>
<emphasis box="[1154,1170,726,751]" italics="true" pageId="17" pageNumber="294">×</emphasis>
<quantity box="[1175,1277,725,752]" metricMagnitude="-1" metricUnit="m" metricValue="4.25" metricValueMax="6.5" metricValueMin="2.0" pageId="17" pageNumber="294" unit="cm" value="42.5" valueMax="65.0" valueMin="20.0">20-65 cm</quantity>
diam, with thick low plank buttresses.Outer bark dark gray(-brown), thick, with deep short fissures, shed in elongate irregular plates; inner bark pale red or orange to tan, thick.Leaves
<quantity box="[1194,1292,821,848]" metricMagnitude="-1" metricUnit="m" metricValue="2.1" metricValueMax="2.7" metricValueMin="1.5" pageId="17" pageNumber="294" unit="cm" value="21.0" valueMax="27.0" valueMin="15.0">15-27 cm</quantity>
long, 3-4(5)-jugate; petiole
<quantity box="[943,1059,853,880]" metricMagnitude="-2" metricUnit="m" metricValue="4.6" metricValueMax="5.7" metricValueMin="3.5" pageId="17" pageNumber="294" unit="cm" value="4.6" valueMax="5.7" valueMin="3.5">3.5-5.7 cm</quantity>
long, petiole and rachis with scattered capitate glands and a few scattered ascending to appressed pale golden hairs to
<quantity box="[970,1066,917,944]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="17" pageNumber="294" unit="mm" value="0.25">0.25 mm</quantity>
long, also with scattered elongate lenticels; stipules
<quantity box="[959,1071,949,975]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="3.8" metricValueMin="1.1" pageId="17" pageNumber="294" unit="mm" value="24.5" valueMax="38.0" valueMin="11.0">11-38 mm</quantity>
from petiole insertion,
<quantity box="[1312,1397,949,975]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="17" pageNumber="294" unit="mm" value="5.0" valueMax="7.0" valueMin="3.0">3-7 mm</quantity>
long, orbicular, membranaceous, with sparse appressed pale golden hairs to
<quantity box="[892,985,1012,1039]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="294" unit="mm" value="0.15">0.15mm</quantity>
long, the scar
<quantity box="[1124,1207,1013,1039]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="17" pageNumber="294" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1-2 mm</quantity>
long; all lateral petiolules
<quantity box="[810,892,1044,1071]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="9.0" metricValueMin="4.0" pageId="17" pageNumber="294" unit="mm" value="6.5" valueMax="9.0" valueMin="4.0">4-9 mm</quantity>
long, terminal one
<quantity box="[1079,1174,1044,1071]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="17" pageNumber="294" unit="mm" value="8.5" valueMax="10.0" valueMin="7.0">7-10 mm</quantity>
long,petiolules canaliculate, lateral pulvinuli inconspicuous; basal leaflets 3.2-4.9 ×
<quantity box="[1354,1448,1076,1103]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" metricValueMax="3.0" metricValueMin="2.2" pageId="17" pageNumber="294" unit="cm" value="2.6" valueMax="3.0" valueMin="2.2">2.2-3cm</quantity>
, broadly ovate to suborbicular or oblong, other laterals 3.7-6.5
<emphasis box="[1439,1455,1109,1134]" italics="true" pageId="17" pageNumber="294">×</emphasis>
<quantity box="[810,927,1140,1167]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" metricValueMax="3.2" metricValueMin="2.2" pageId="17" pageNumber="294" unit="cm" value="2.7" valueMax="3.2" valueMin="2.2">2.2-3.2 cm</quantity>
, broadly elliptic to slightly ovate; terminal one 3.5- 6.2
<emphasis box="[851,867,1173,1198]" italics="true" pageId="17" pageNumber="294">×</emphasis>
<quantity box="[870,985,1172,1198]" metricMagnitude="-2" metricUnit="m" metricValue="2.8499999999999996" metricValueMax="3.3" metricValueMin="2.4" pageId="17" pageNumber="294" unit="cm" value="2.8499999999999996" valueMax="3.3" valueMin="2.4">2.4-3.3 cm</quantity>
, broadly elliptic, rarely slightly obovate; leaflet apex usually abruptly and broadly short-acuminate, the acumen
<quantity box="[810,897,1236,1262]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="7.0" metricValueMin="2.0" pageId="17" pageNumber="294" unit="mm" value="4.5" valueMax="7.0" valueMin="2.0">2-7 mm</quantity>
long; lateral leaflet base symmetrical to slightly asymmetrical, obtuse or rounded to truncate or sometimes slightly cordate;leaflet margin flat to slightly revolute; leaflets coriaceous, drying brown; secondary vein framework festooned brochidodromous, usually looping far from the margin, secondaries in 6-9(11) pairs, spreading, spacing decreasing toward apex and base, the angle increasing toward the apex and sometimes toward the base, perpendicular epimedial tertiaries usually present, intercostal tertiaries irregular-reticulate,frequently some admedial tertiaries branching toward secondary vein axils from adjacent secondaries, quaternaries regular-polygonal; on abaxial side all veins narrowly prominent, the surface glabrous; on adaxial side all veins narrowly prominulous except the midvein sunk in a groove, glabrous. Flowers unknown. Infructescences
<quantity box="[1365,1455,1650,1677]" metricMagnitude="-1" metricUnit="m" metricValue="1.45" metricValueMax="2.0" metricValueMin="0.9" pageId="17" pageNumber="294" unit="cm" value="14.5" valueMax="20.0" valueMin="9.0">9-20 cm</quantity>
long, the secondary axes
<quantity box="[1080,1157,1682,1709]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="17" pageNumber="294" unit="cm" value="5.0" valueMax="7.0" valueMin="3.0">3-7 cm</quantity>
long, in fruit the axes (sub) glabrous, sometimes with some bristles to
<quantity box="[1246,1330,1714,1741]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="294" unit="mm" value="0.1">0.1 mm</quantity>
long; bracts deciduous, not semi-clasping; fruiting pedicel 4-7
<emphasis box="[1339,1355,1747,1772]" italics="true" pageId="17" pageNumber="294">×</emphasis>
<quantity box="[1362,1452,1746,1772]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="17" pageNumber="294" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2-3 mm</quantity>
, slightly clavate; fruiting calyx
<quantity box="[1124,1211,1778,1804]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="17" pageNumber="294" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2-3 mm</quantity>
long, spreading to patent, the lobes distinct, with a few short thick ascending hairs to
<quantity box="[811,907,1841,1868]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="294" unit="mm" value="0.15">0.15 mm</quantity>
long. Fruits 3-3.2
<emphasis box="[1094,1110,1842,1867]" italics="true" pageId="17" pageNumber="294">×</emphasis>
<quantity box="[1111,1224,1841,1868]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" metricValueMax="3.2" metricValueMin="2.6" pageId="17" pageNumber="294" unit="cm" value="2.9" valueMax="3.2" valueMin="2.6">2.6-3.2 cm</quantity>
(to
<quantity box="[1263,1334,1842,1868]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="17" pageNumber="294" unit="cm" value="3.4">3.4 cm</quantity>
long fresh), broadly obovoid, rarely slightly ovoid or depressed-globose, sometimes slightly 3-lobed (
<emphasis box="[1102,1279,1905,1931]" italics="true" pageId="17" pageNumber="294">Daly et al. 13073</emphasis>
), the apex round and base obtuse to truncate, surface glaucous to dull turquoise, smooth, glabrous, usually glossy; mesocarp to
<quantity box="[1286,1350,1969,1996]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="294" unit="mm" value="5.0">5 mm</quantity>
thick; the pyrene 2-2.8
<emphasis box="[951,967,2002,2027]" italics="true" pageId="17" pageNumber="294">×</emphasis>
<quantity box="[972,1089,2001,2028]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.4" metricValueMin="2.1" pageId="17" pageNumber="294" unit="cm" value="2.25" valueMax="2.4" valueMin="2.1">2.1-2.4 cm</quantity>
, operculum runs along the pyrene. NOTES
</paragraph>
<caption box="[392,1198,219,237]" pageId="18" pageNumber="295" startId="18.[392,402,219,237]">
<paragraph blockId="18.[392,1198,219,237]" box="[392,1198,219,237]" pageId="18" pageNumber="295">
TABLE 1. — Comparison of the
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[647,836,219,237]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="18" pageNumber="295" phylum="Tracheophyta" rank="species" species="betamponae" status="sp. nov.">
<emphasis box="[647,836,219,237]" italics="true" pageId="18" pageNumber="295">Canarium betamponae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[846,913,219,237]" pageId="18" pageNumber="295" rank="species">sp. nov.</taxonomicNameLabel>
complex (salient features in bold).
</paragraph>
</caption>
<paragraph pageId="18" pageNumber="295">
<table box="[134,1454,294,1305]" gridcols="3" gridrows="40" pageId="18" pageNumber="295">
<tr box="[134,1454,294,314]" gridrow="0" pageId="18" pageNumber="295">
<td box="[134,577,294,314]" gridcol="0" gridrow="0" pageId="18" pageNumber="295">
<emphasis bold="true" box="[357,518,294,314]" pageId="18" pageNumber="295">
<taxonomicName authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[357,512,294,314]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="18" pageNumber="295" phylum="Tracheophyta" rank="species" species="betamponae">
<emphasis bold="true" box="[357,512,294,314]" italics="true" pageId="18" pageNumber="295">C. betamponae</emphasis>
</taxonomicName>
,
</emphasis>
</td>
<td box="[588,1244,294,314]" gridcol="1" gridrow="0" pageId="18" pageNumber="295">
<emphasis bold="true" box="[588,1237,294,314]" pageId="18" pageNumber="295">
<taxonomicName authorityName="Daly, Raharim. &amp; Federman" authorityYear="2015" box="[588,720,294,314]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="18" pageNumber="295" phylum="Tracheophyta" rank="species" species="galokense">
<emphasis bold="true" box="[588,720,294,314]" italics="true" pageId="18" pageNumber="295">C. galokense</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Daly, Raharim. &amp; Federman" authorityYear="2015" box="[809,937,294,314]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="18" pageNumber="295" phylum="Tracheophyta" rank="species" species="globosum">
<emphasis bold="true" box="[809,937,294,314]" italics="true" pageId="18" pageNumber="295">C. globosum</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Daly, Raharim. &amp; Federman" authorityYear="2015" box="[1051,1231,294,314]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="18" pageNumber="295" phylum="Tracheophyta" rank="species" species="manongarivum">
<emphasis bold="true" box="[1051,1231,294,314]" italics="true" pageId="18" pageNumber="295">C. manongarivum</emphasis>
</taxonomicName>
,
</emphasis>
</td>
<td box="[1256,1454,294,314]" gridcol="2" gridrow="0" pageId="18" pageNumber="295">
<emphasis bold="true" box="[1256,1415,294,314]" pageId="18" pageNumber="295">
<taxonomicName authorityName="Daly, Raharim. &amp; Federman" authorityYear="2015" box="[1256,1408,294,314]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="18" pageNumber="295" phylum="Tracheophyta" rank="species" species="subsidarium">
<emphasis bold="true" box="[1256,1408,294,314]" italics="true" pageId="18" pageNumber="295">C. subsidarium</emphasis>
</taxonomicName>
,
</emphasis>
</td>
</tr>
<tr box="[134,1454,320,340]" gridrow="1" pageId="18" pageNumber="295">
<td box="[134,577,320,340]" gridcol="0" gridrow="1" pageId="18" pageNumber="295">
<emphasis bold="true" box="[134,458,320,340]" italics="true" pageId="18" pageNumber="295">
<taxonomicName box="[134,270,320,340]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="18" pageNumber="295" phylum="Tracheophyta" rank="species" species="undetermined">Canarium sp.</taxonomicName>
sp. nov.
</emphasis>
</td>
<td box="[588,1244,320,340]" gridcol="1" gridrow="1" pageId="18" pageNumber="295">
<emphasis bold="true" box="[611,1152,320,340]" pageId="18" pageNumber="295">sp. nov. sp. nov. sp. nov.</emphasis>
</td>
<td box="[1256,1454,320,340]" gridcol="2" gridrow="1" pageId="18" pageNumber="295">
<emphasis bold="true" box="[1278,1357,320,340]" pageId="18" pageNumber="295">sp. nov.</emphasis>
</td>
</tr>
<tr box="[134,1454,355,376]" gridrow="2" pageId="18" pageNumber="295">
<td box="[134,577,355,376]" gridcol="0" gridrow="2" pageId="18" pageNumber="295">Provinces of Toamasina</td>
<td box="[588,1244,355,376]" gridcol="1" gridrow="2" pageId="18" pageNumber="295">Antsiranana Antsiranana, Toamasina, Antsiranana</td>
<td box="[1256,1454,355,376]" gridcol="2" gridrow="2" pageId="18" pageNumber="295">Antsiranana</td>
</tr>
<tr box="[134,1454,381,401]" gridrow="3" pageId="18" pageNumber="295" rowspan-2="1">
<td box="[134,577,381,401]" gridcol="0" gridrow="3" pageId="18" pageNumber="295">distribution</td>
<td box="[588,1244,381,401]" gridcol="1" gridrow="3" pageId="18" pageNumber="295">Toliara</td>
</tr>
<tr box="[134,1454,405,426]" gridrow="4" pageId="18" pageNumber="295">
<td box="[134,577,405,426]" gridcol="0" gridrow="4" pageId="18" pageNumber="295">Elevation range (m) 325-600</td>
<td box="[588,1244,405,426]" gridcol="1" gridrow="4" pageId="18" pageNumber="295">
300-700 10-700
<emphasis box="[1051,1069,405,425]" italics="true" pageId="18" pageNumber="295">c.</emphasis>
875
</td>
<td box="[1256,1454,405,426]" gridcol="2" gridrow="4" pageId="18" pageNumber="295">
<emphasis bold="true" box="[1256,1360,406,426]" pageId="18" pageNumber="295">1000-1600</emphasis>
</td>
</tr>
<tr box="[134,1454,430,450]" gridrow="5" pageId="18" pageNumber="295">
<td box="[134,577,430,450]" gridcol="0" gridrow="5" pageId="18" pageNumber="295">No. juga 3-4(5)</td>
<td box="[588,1244,430,450]" gridcol="1" gridrow="5" pageId="18" pageNumber="295">
3-5 3-5
<emphasis bold="true" box="[1051,1063,430,450]" pageId="18" pageNumber="295">3</emphasis>
</td>
<td box="[1256,1454,430,450]" gridcol="2" gridrow="5" pageId="18" pageNumber="295">3-4</td>
</tr>
<tr box="[134,1454,454,474]" gridrow="6" pageId="18" pageNumber="295">
<td box="[134,577,454,474]" gridcol="0" gridrow="6" pageId="18" pageNumber="295">Stipule insertion 11-38</td>
<td box="[588,1244,454,474]" gridcol="1" gridrow="6" pageId="18" pageNumber="295">3-10 (4)8-23(35) 10-16</td>
<td box="[1256,1454,454,474]" gridcol="2" gridrow="6" pageId="18" pageNumber="295">4-9</td>
</tr>
<tr box="[134,1454,480,500]" gridrow="7" pageId="18" pageNumber="295" rowspan-1="1" rowspan-2="1">
<td box="[134,577,480,500]" gridcol="0" gridrow="7" pageId="18" pageNumber="295">from base (mm)</td>
</tr>
<tr box="[134,1454,504,525]" gridrow="8" pageId="18" pageNumber="295">
<td box="[134,577,504,525]" gridcol="0" gridrow="8" pageId="18" pageNumber="295">Stipular 1-2</td>
<td box="[588,1244,504,525]" gridcol="1" gridrow="8" pageId="18" pageNumber="295">
2-4 1-2
<emphasis bold="true" box="[1051,1084,505,525]" pageId="18" pageNumber="295">7-9</emphasis>
</td>
<td box="[1256,1454,504,525]" gridcol="2" gridrow="8" pageId="18" pageNumber="295">2-3 (raised)</td>
</tr>
<tr box="[134,1454,530,550]" gridrow="9" pageId="18" pageNumber="295" rowspan-1="1" rowspan-2="1">
<td box="[134,577,530,550]" gridcol="0" gridrow="9" pageId="18" pageNumber="295">scar length (mm)</td>
</tr>
<tr box="[134,1454,553,574]" gridrow="10" pageId="18" pageNumber="295">
<td box="[134,577,553,574]" gridcol="0" gridrow="10" pageId="18" pageNumber="295">Stipule length (mm) 3-7, orbicular</td>
<td box="[588,1244,553,574]" gridcol="1" gridrow="10" pageId="18" pageNumber="295">4.5-10, orbicular, 3-8, orbicular [Unknown]</td>
<td box="[1256,1454,553,574]" gridcol="2" gridrow="10" pageId="18" pageNumber="295">[Unknown]</td>
</tr>
<tr box="[134,1454,579,599]" gridrow="11" pageId="18" pageNumber="295" rowspan-2="1">
<td box="[134,577,579,599]" gridcol="0" gridrow="11" pageId="18" pageNumber="295">and shape</td>
<td box="[588,1244,579,599]" gridcol="1" gridrow="11" pageId="18" pageNumber="295">substipitate</td>
</tr>
<tr box="[134,1454,603,624]" gridrow="12" pageId="18" pageNumber="295">
<td box="[134,577,603,624]" gridcol="0" gridrow="12" pageId="18" pageNumber="295">
Stipule texture
<emphasis bold="true" box="[357,528,604,624]" pageId="18" pageNumber="295">Membranaceous</emphasis>
</td>
<td box="[588,1244,603,624]" gridcol="1" gridrow="12" pageId="18" pageNumber="295">Coriaceous Becoming fleshy [Unknown]</td>
<td box="[1256,1454,603,624]" gridcol="2" gridrow="12" pageId="18" pageNumber="295">[Unknown]</td>
</tr>
<tr box="[134,1454,628,648]" gridrow="13" pageId="18" pageNumber="295">
<td box="[134,577,628,648]" gridcol="0" gridrow="13" pageId="18" pageNumber="295">Leaf rachis Few scattered</td>
<td box="[588,1244,628,648]" gridcol="1" gridrow="13" pageId="18" pageNumber="295">Scattered thick short Sparse to scattered, fine,Sparse thick erect</td>
<td box="[1256,1454,628,648]" gridcol="2" gridrow="13" pageId="18" pageNumber="295">
<emphasis bold="true" box="[1256,1449,628,648]" pageId="18" pageNumber="295">Dense to scattered</emphasis>
</td>
</tr>
<tr box="[134,1454,654,674]" gridrow="14" pageId="18" pageNumber="295">
<td box="[134,577,654,674]" gridcol="0" gridrow="14" pageId="18" pageNumber="295">pubescence ascending to</td>
<td box="[588,1244,654,674]" gridcol="1" gridrow="14" pageId="18" pageNumber="295">hairs to 0.1 appressed hairs to 0.2 hairs to 0.1 mm</td>
<td box="[1256,1454,654,674]" gridcol="2" gridrow="14" pageId="18" pageNumber="295">
<emphasis bold="true" box="[1278,1430,654,674]" pageId="18" pageNumber="295">fine erect hairs</emphasis>
</td>
</tr>
<tr box="[134,1454,679,700]" gridrow="15" pageId="18" pageNumber="295">
<td box="[134,577,679,700]" gridcol="0" gridrow="15" pageId="18" pageNumber="295">(discounting capitate appressed hairs to</td>
<td box="[588,1244,679,700]" gridcol="1" gridrow="15" pageId="18" pageNumber="295">long</td>
<td box="[1256,1454,679,700]" gridcol="2" gridrow="15" pageId="18" pageNumber="295">
<emphasis bold="true" box="[1278,1388,680,700]" pageId="18" pageNumber="295">to 0.5; also</emphasis>
</td>
</tr>
<tr box="[134,1454,705,725]" gridrow="16" pageId="18" pageNumber="295" rowspan-1="1">
<td box="[134,577,705,725]" gridcol="0" gridrow="16" pageId="18" pageNumber="295">glands) (mm) 0.25</td>
<td box="[1256,1454,705,725]" gridcol="2" gridrow="16" pageId="18" pageNumber="295">
<emphasis bold="true" box="[1278,1403,705,725]" pageId="18" pageNumber="295">bristles (fine</emphasis>
</td>
</tr>
<tr box="[134,1454,731,751]" gridrow="17" pageId="18" pageNumber="295" rowspan-0="1" rowspan-1="1">
<td box="[1256,1454,731,751]" gridcol="2" gridrow="17" pageId="18" pageNumber="295">
<emphasis bold="true" box="[1278,1414,731,751]" pageId="18" pageNumber="295">erect hairs to</emphasis>
</td>
</tr>
<tr box="[134,1454,756,776]" gridrow="18" pageId="18" pageNumber="295" rowspan-0="1" rowspan-1="1">
<td box="[1256,1454,756,776]" gridcol="2" gridrow="18" pageId="18" pageNumber="295">
<emphasis bold="true" box="[1278,1326,756,776]" pageId="18" pageNumber="295">0.05)</emphasis>
</td>
</tr>
<tr box="[134,1454,780,801]" gridrow="19" pageId="18" pageNumber="295">
<td box="[134,577,780,801]" gridcol="0" gridrow="19" pageId="18" pageNumber="295">Lateral leaflet shape Broadly elliptic to</td>
<td box="[588,1244,780,801]" gridcol="1" gridrow="19" pageId="18" pageNumber="295">(Oblong-) lanceolate Oblong or less often Elliptic to slightly</td>
<td box="[1256,1454,780,801]" gridcol="2" gridrow="19" pageId="18" pageNumber="295">Oblong(-lanceolate)</td>
</tr>
<tr box="[134,1454,806,826]" gridrow="20" pageId="18" pageNumber="295">
<td box="[134,577,806,826]" gridcol="0" gridrow="20" pageId="18" pageNumber="295">slightly ovate</td>
<td box="[588,1244,806,826]" gridcol="1" gridrow="20" pageId="18" pageNumber="295">to ovate oblong-elliptic or oblong-</td>
<td box="[1256,1454,806,826]" gridcol="2" gridrow="20" pageId="18" pageNumber="295">to elliptic</td>
</tr>
<tr box="[134,1454,832,852]" gridrow="21" pageId="18" pageNumber="295" rowspan-0="1" rowspan-2="1">
<td box="[588,1244,832,852]" gridcol="1" gridrow="21" pageId="18" pageNumber="295">ovate oblanceolate</td>
</tr>
<tr box="[134,1454,856,876]" gridrow="22" pageId="18" pageNumber="295">
<td box="[134,577,856,876]" gridcol="0" gridrow="22" pageId="18" pageNumber="295">Leaflet apex, Usually abruptly and</td>
<td box="[588,1244,856,876]" gridcol="1" gridrow="22" pageId="18" pageNumber="295">
Gradually and usually
<emphasis bold="true" box="[809,1005,856,876]" pageId="18" pageNumber="295">Rounded to slightly</emphasis>
Broadly short-
</td>
<td box="[1256,1454,856,876]" gridcol="2" gridrow="22" pageId="18" pageNumber="295">Gradually and</td>
</tr>
<tr box="[134,1454,881,902]" gridrow="23" pageId="18" pageNumber="295">
<td box="[134,577,881,902]" gridcol="0" gridrow="23" pageId="18" pageNumber="295">acumen (mm) broadly short-</td>
<td box="[588,1244,881,902]" gridcol="1" gridrow="23" pageId="18" pageNumber="295">
narrowly acuminate,
<emphasis bold="true" box="[832,1033,882,902]" pageId="18" pageNumber="295">emarginate, seldom</emphasis>
acuminate or
</td>
<td box="[1256,1454,881,902]" gridcol="2" gridrow="23" pageId="18" pageNumber="295">
broadly
<tr box="[134,1454,907,928]" gridrow="24" pageId="18" pageNumber="295">
<td box="[134,577,907,928]" gridcol="0" gridrow="24" pageId="18" pageNumber="295">shortacuminate, 2-7</td>
<td box="[588,1244,907,928]" gridcol="1" gridrow="24" pageId="18" pageNumber="295">
(2) 5-10
<emphasis bold="true" box="[832,1041,908,928]" pageId="18" pageNumber="295">abruptly and broadly</emphasis>
rarely obtuse, 0-4
</td>
<td box="[1256,1454,907,928]" gridcol="2" gridrow="24" pageId="18" pageNumber="295">acuminate, 2-6</td>
</tr>
</td>
</tr>
<tr box="[134,1454,933,953]" gridrow="25" pageId="18" pageNumber="295" rowspan-0="1" rowspan-2="1">
<td box="[588,1244,933,953]" gridcol="1" gridrow="25" pageId="18" pageNumber="295">
<emphasis bold="true" box="[832,1007,933,953]" pageId="18" pageNumber="295">short-acuminate,</emphasis>
</td>
</tr>
<tr box="[134,1454,959,979]" gridrow="26" pageId="18" pageNumber="295" rowspan-0="1" rowspan-2="1">
<td box="[588,1244,959,979]" gridcol="1" gridrow="26" pageId="18" pageNumber="295">
<emphasis bold="true" box="[832,894,959,979]" pageId="18" pageNumber="295">0-2 (4)</emphasis>
</td>
</tr>
<tr box="[134,1454,983,1003]" gridrow="27" pageId="18" pageNumber="295">
<td box="[134,577,983,1003]" gridcol="0" gridrow="27" pageId="18" pageNumber="295">Leaflet adaxial surface Glabrous</td>
<td box="[588,1244,983,1003]" gridcol="1" gridrow="27" pageId="18" pageNumber="295">
Few scattered capitate Glabrous
<emphasis bold="true" box="[1051,1207,983,1003]" pageId="18" pageNumber="295">Black-punctate</emphasis>
</td>
<td box="[1256,1454,983,1003]" gridcol="2" gridrow="27" pageId="18" pageNumber="295">Glabrous</td>
</tr>
<tr box="[134,1454,1008,1028]" gridrow="28" pageId="18" pageNumber="295" rowspan-0="1" rowspan-2="1">
<td box="[588,1244,1008,1028]" gridcol="1" gridrow="28" pageId="18" pageNumber="295">glands or glabrous</td>
</tr>
<tr box="[134,1454,1032,1053]" gridrow="29" pageId="18" pageNumber="295">
<td box="[134,577,1032,1053]" gridcol="0" gridrow="29" pageId="18" pageNumber="295">
Inflorescence
<emphasis bold="true" box="[357,498,1033,1053]" pageId="18" pageNumber="295">(Sub)glabrous</emphasis>
,
</td>
<td box="[588,1244,1032,1053]" gridcol="1" gridrow="29" pageId="18" pageNumber="295">Dense to sparse Sparse to dense Sparse to scattered</td>
<td box="[1256,1454,1032,1053]" gridcol="2" gridrow="29" pageId="18" pageNumber="295">Dense to sparse</td>
</tr>
<tr box="[134,1454,1058,1079]" gridrow="30" pageId="18" pageNumber="295">
<td box="[134,577,1058,1079]" gridcol="0" gridrow="30" pageId="18" pageNumber="295">trichomes sometimes with</td>
<td box="[588,1244,1058,1079]" gridcol="1" gridrow="30" pageId="18" pageNumber="295">flexuous hairs flexuous hairs to 0.2 flexuous to erect</td>
<td box="[1256,1454,1058,1079]" gridcol="2" gridrow="30" pageId="18" pageNumber="295">flexuous hairs to</td>
</tr>
<tr box="[134,1454,1084,1104]" gridrow="31" pageId="18" pageNumber="295">
<td box="[134,577,1084,1104]" gridcol="0" gridrow="31" pageId="18" pageNumber="295">(discounting capitate some bristles to 0.1</td>
<td box="[588,1244,1084,1104]" gridcol="1" gridrow="31" pageId="18" pageNumber="295">to 0.25 thick hairs to 0.25</td>
<td box="[1256,1454,1084,1104]" gridcol="2" gridrow="31" pageId="18" pageNumber="295">0.3, also sparse to</td>
</tr>
<tr box="[134,1454,1109,1129]" gridrow="32" pageId="18" pageNumber="295" rowspan-1="1">
<td box="[134,577,1109,1129]" gridcol="0" gridrow="32" pageId="18" pageNumber="295">glands) (mm)</td>
<td box="[1256,1454,1109,1129]" gridcol="2" gridrow="32" pageId="18" pageNumber="295">dense bristles</td>
</tr>
<tr box="[134,1454,1133,1154]" gridrow="33" pageId="18" pageNumber="295">
<td box="[134,577,1133,1154]" gridcol="0" gridrow="33" pageId="18" pageNumber="295">
Fruit size (cm) 3-3.2
<emphasis box="[412,425,1134,1154]" italics="true" pageId="18" pageNumber="295">×</emphasis>
2.6-3.2
</td>
<td box="[588,1244,1133,1154]" gridcol="1" gridrow="33" pageId="18" pageNumber="295">
3.5-4
<emphasis box="[644,657,1134,1154]" italics="true" pageId="18" pageNumber="295">×</emphasis>
2.5-3 2-3.8
<emphasis box="[865,878,1134,1154]" italics="true" pageId="18" pageNumber="295">×</emphasis>
2-2.5
<emphasis box="[1051,1069,1134,1154]" italics="true" pageId="18" pageNumber="295">c.</emphasis>
2.8-2.9
<emphasis box="[1148,1161,1134,1154]" italics="true" pageId="18" pageNumber="295">×</emphasis>
1.8-2.1
</td>
<td box="[1256,1454,1133,1154]" gridcol="2" gridrow="33" pageId="18" pageNumber="295">
3.6-3.8
<emphasis box="[1329,1342,1134,1154]" italics="true" pageId="18" pageNumber="295">×</emphasis>
2.3-2.8
</td>
</tr>
<tr box="[134,1454,1159,1179]" gridrow="34" pageId="18" pageNumber="295" rowspan-0="1" rowspan-2="1">
<td box="[588,1244,1159,1179]" gridcol="1" gridrow="34" pageId="18" pageNumber="295">(immature?)</td>
</tr>
<tr box="[134,1454,1183,1204]" gridrow="35" pageId="18" pageNumber="295">
<td box="[134,577,1183,1204]" gridcol="0" gridrow="35" pageId="18" pageNumber="295">Fruit shape Broadly obovoid, rarely</td>
<td box="[588,1244,1183,1204]" gridcol="1" gridrow="35" pageId="18" pageNumber="295">
<emphasis bold="true" box="[588,769,1184,1204]" pageId="18" pageNumber="295">Broadly oblong or</emphasis>
Subglobose to (ob)ovoid Slightly ovoid
</td>
<td box="[1256,1454,1183,1204]" gridcol="2" gridrow="35" pageId="18" pageNumber="295">Ovoid</td>
</tr>
<tr box="[134,1454,1209,1229]" gridrow="36" pageId="18" pageNumber="295" rowspan-2="1">
<td box="[134,577,1209,1229]" gridcol="0" gridrow="36" pageId="18" pageNumber="295">slightly ovoid</td>
<td box="[588,1244,1209,1229]" gridcol="1" gridrow="36" pageId="18" pageNumber="295">
<emphasis bold="true" box="[611,749,1209,1229]" pageId="18" pageNumber="295">broadly ovoid</emphasis>
</td>
</tr>
<tr box="[134,1454,1233,1254]" gridrow="37" pageId="18" pageNumber="295">
<td box="[134,577,1233,1254]" gridcol="0" gridrow="37" pageId="18" pageNumber="295">Fruit surface Glabrous, glossy</td>
<td box="[588,1244,1233,1254]" gridcol="1" gridrow="37" pageId="18" pageNumber="295">
<emphasis bold="true" box="[588,706,1234,1254]" pageId="18" pageNumber="295">Lenticellate</emphasis>
Glabrous, glossy
<emphasis bold="true" box="[1051,1213,1234,1254]" pageId="18" pageNumber="295">Lenticellate and</emphasis>
</td>
<td box="[1256,1454,1233,1254]" gridcol="2" gridrow="37" pageId="18" pageNumber="295">Glabrous, dull;</td>
</tr>
<tr box="[134,1454,1259,1279]" gridrow="38" pageId="18" pageNumber="295" rowspan-0="1">
<td box="[588,1244,1259,1279]" gridcol="1" gridrow="38" pageId="18" pageNumber="295">
<emphasis bold="true" box="[1073,1180,1259,1279]" pageId="18" pageNumber="295">pubescent</emphasis>
</td>
<td box="[1256,1454,1259,1279]" gridcol="2" gridrow="38" pageId="18" pageNumber="295">becoming finely</td>
</tr>
<tr box="[134,1454,1285,1305]" gridrow="39" pageId="18" pageNumber="295" rowspan-0="1" rowspan-1="1">
<td box="[1256,1454,1285,1305]" gridcol="2" gridrow="39" pageId="18" pageNumber="295">lenticellate</td>
</tr>
</table>
</paragraph>
<paragraph blockId="18.[132,775,1364,1581]" pageId="18" pageNumber="295">
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[132,363,1395,1422]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="18" pageNumber="295" phylum="Tracheophyta" rank="species" species="betamponae" status="sp. nov.">
<emphasis box="[132,363,1395,1422]" italics="true" pageId="18" pageNumber="295">Canarium betamponae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[378,458,1396,1422]" pageId="18" pageNumber="295" rank="species">sp. nov.</taxonomicNameLabel>
is part of a complex of species including
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[235,354,1427,1453]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="18" pageNumber="295" phylum="Tracheophyta" rank="species" species="galokense" status="sp.nov.">
<emphasis box="[235,354,1427,1453]" italics="true" pageId="18" pageNumber="295">C. galokense</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[364,443,1428,1454]" pageId="18" pageNumber="295" rank="species">sp.nov.</taxonomicNameLabel>
,
<emphasis box="[451,570,1427,1453]" italics="true" pageId="18" pageNumber="295">
C,
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[480,570,1427,1453]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="18" pageNumber="295" phylum="Tracheophyta" rank="species" species="globosum" status="sp. nov.">globosum</taxonomicName>
</emphasis>
,
<taxonomicNameLabel box="[581,658,1428,1454]" pageId="18" pageNumber="295" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="18" pageNumber="295" phylum="Tracheophyta" rank="species" species="manongarivum" status="sp. nov.">
<emphasis italics="true" pageId="18" pageNumber="295">C.manongarivum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[228,307,1459,1485]" pageId="18" pageNumber="295" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[354,507,1459,1485]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="18" pageNumber="295" phylum="Tracheophyta" rank="species" species="subsidarium" status="sp. nov.">
<emphasis box="[354,507,1459,1485]" italics="true" pageId="18" pageNumber="295">C. subsidarium</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[519,598,1459,1485]" pageId="18" pageNumber="295" rank="species">sp. nov.</taxonomicNameLabel>
that have 3-5-jugate leaves with short stipular scars (except
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[583,768,1491,1517]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="18" pageNumber="295" phylum="Tracheophyta" rank="species" species="manongarivum" status="sp. nov.">
<emphasis box="[583,768,1491,1517]" italics="true" pageId="18" pageNumber="295">C. manongarivum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[132,219,1523,1549]" pageId="18" pageNumber="295" rank="species">sp. nov.</taxonomicNameLabel>
), relatively small leaflets that are often oblong, and relatively small fruits. They are compared in
<tableCitation box="[592,670,1554,1581]" captionStart="TABLE" captionStartId="18.[392,402,219,237]" captionText="TABLE 1. — Comparison of the Canarium betamponae, sp. nov. complex (salient features in bold)." pageId="18" pageNumber="295">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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