treatments-xml/data/FA/65/87/FA6587FFFF81FF8D6B94F99DFE4CD3DA.xml

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<document id="DDAFC87B2918BF69ED5BE1D6091358D6" ID="10.11646/zootaxa.4268.4.6SLASH11068" ID-CLB-Dataset="33022" ID-DOI="10.11646/zootaxa.4268.4.6" ID-GBIF-Dataset="196c385a-b33d-4f66-9140-c08987fe2347" ID-ISSN="1175-5326" ID-Zenodo-Dep="580966" ID-ZooBank="432A29A1-0C34-4728-8AC1-07BC5EF7EED8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1495097136037" checkinUser="plazi" docAuthor="Makarova, Olga L. &amp; Huhta, Veikko" docDate="2017" docId="FA6587FFFF81FF8D6B94F99DFE4CD3DA" docLanguage="en" docName="zootaxa.4268.4.6SLASH11068.pdf" docOrigin="Zootaxa 4268 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Arctoseius ambiguus Makarova &amp; Huhta, sp. nov." docType="treatment" docVersion="8" lastPageNumber="559" masterDocId="065CFF87FF80FF886B03FA4DFFEDD012" masterDocTitle="A new species of Arctoseius Thor, 1930 (Acari: Ascidae) from taiga regions of the Palaearctic, with a key to Arctoseius species of Fennoscandia, NW Europe" masterLastPageNumber="562" masterPageNumber="554" pageNumber="555" updateTime="1698439715192" updateUser="ExternalLinkService">
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<mods:title id="54EEE9A59561F501AA4F568B9666D48B">A new species of Arctoseius Thor, 1930 (Acari: Ascidae) from taiga regions of the Palaearctic, with a key to Arctoseius species of Fennoscandia, NW Europe</mods:title>
</mods:titleInfo>
<mods:name id="351EADCCF10349871C4675C4D5041B2A" type="personal">
<mods:role id="A61EEBA92B2A1FA05735035D74C01C1B">
<mods:roleTerm id="7497C1F0F97A2CA9F670AE82AC742B2D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7A2E11219CDB8DB1B62A338B80CBD28E">Makarova, Olga L.</mods:namePart>
</mods:name>
<mods:name id="585023412F25CDB41E0092A69220DB0B" type="personal">
<mods:role id="A1215F548633078C9DCA211721D9EAC9">
<mods:roleTerm id="E90697155927FE4B8C642240C62B6486">Author</mods:roleTerm>
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<mods:namePart id="577204BE6AD4FFBA182450D07D657F04">Huhta, Veikko</mods:namePart>
</mods:name>
<mods:typeOfResource id="B1F13DED1AA1462FA11DA3CA142A4140">text</mods:typeOfResource>
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<mods:title id="EC06B86792B1B5BCFDF8892F358A256A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="04680313A77595A2C743F2C4956D455B">
<mods:date id="A5AAE72F3463E2880AA66122027C324C">2017</mods:date>
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<mods:identifier id="936F1A252D642AFDF14C06458AC1ADCF" type="CLB-Dataset">33022</mods:identifier>
<mods:identifier id="65FDEE11D4DFF4B9D25AAA7A5E6E75CE" type="DOI">10.11646/zootaxa.4268.4.6</mods:identifier>
<mods:identifier id="7BC5C2E4C43FFBD9A13CB196E404F857" type="GBIF-Dataset">196c385a-b33d-4f66-9140-c08987fe2347</mods:identifier>
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<treatment id="FA6587FFFF81FF8D6B94F99DFE4CD3DA" ID-DOI="http://doi.org/10.5281/zenodo.6024012" ID-GBIF-Taxon="130810313" ID-Zenodo-Dep="6024012" LSID="urn:lsid:plazi:treatment:FA6587FFFF81FF8D6B94F99DFE4CD3DA" httpUri="http://treatment.plazi.org/id/FA6587FFFF81FF8D6B94F99DFE4CD3DA" lastPageId="5" lastPageNumber="559" pageId="1" pageNumber="555">
<subSubSection id="3AD66562FF81FF896B94F99DFEF5D41E" pageId="1" pageNumber="555" type="nomenclature">
<paragraph id="727336E9FF81FF896B94F99DFD14D3F9" blockId="1.[151,761,976,1037]" box="[151,761,976,1003]" pageId="1" pageNumber="555">
<heading id="293B8185FF81FF896B94F99DFD14D3F9" bold="true" box="[151,761,976,1003]" fontSize="11" level="1" pageId="1" pageNumber="555" reason="1">
<emphasis id="40B8EAFBFF81FF896B94F99DFD14D3F9" bold="true" box="[151,761,976,1003]" pageId="1" pageNumber="555">
<taxonomicName id="B5CC4D6AFF81FF896B94F99DFD79D3F9" ID-CoL="9FSKJ" authority="Makarova &amp; Huhta" authorityName="Makarova &amp; Huhta" box="[151,660,976,1003]" class="Arachnida" family="Ascidae" genus="Arctoseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="555" phylum="Arthropoda" rank="species" species="ambiguus" status="sp. nov.">
<emphasis id="40B8EAFBFF81FF896B94F99DFE79D3F8" bold="true" box="[151,404,976,1002]" italics="true" pageId="1" pageNumber="555">Arctoseius ambiguus</emphasis>
Makarova &amp; Huhta
</taxonomicName>
<taxonomicNameLabel id="5B8B5780FF81FF896998F99CFD14D3F9" box="[667,761,977,1003]" pageId="1" pageNumber="555" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="727336E9FF81FF896B94F9B9FEF5D41E" blockId="1.[151,761,976,1037]" box="[151,280,1012,1037]" pageId="1" pageNumber="555">
(
<figureCitation id="EAF72A6CFF81FF896B9CF9B9FEE2D41E" box="[159,271,1012,1037]" captionStart-0="FIGURES 1 5" captionStart-1="FIGURES 6 11" captionStartId-0="3.[151,264,1930,1953]" captionStartId-1="4.[151,264,1925,1948]" captionTargetBox-0="[179,1427,199,1902]" captionTargetBox-1="[162,1430,195,1884]" captionTargetId-0="figure@3.[151,1436,195,1907]" captionTargetId-1="figure@4.[151,1436,193,1904]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 1 5. Arctoseius ambiguus sp. nov., female. 1. tritosternum; 2. ambulacrum and tarsus II distally, ventral aspect; 3. gnathotectum; 4. peritrematal region; 5. idiosomal dorsum. Scales: 1 2 25 µm, 3 5 50 µm." captionText-1="FIGURES 6 11. Arctoseius ambiguus sp. nov., female. 6. acetabula II IV and exopodal platelets; 7. subcapitulum; 8. idiosomal ventrum; 9. anal shield; 10. cheliceral digits, dorsal aspect; 11. chelicera without basal joint, paraxial aspect. Scales: 6 9 50 µm, 10 11 25 µm." httpUri-0="https://zenodo.org/record/580967/files/figure.png" httpUri-1="https://zenodo.org/record/580968/files/figure.png" pageId="1" pageNumber="555">Figs 111</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3AD66562FF81FF896B94FE71FD4ED58A" pageId="1" pageNumber="555" type="materials_examined">
<paragraph id="727336E9FF81FF896B94FE71FD4ED58A" blockId="1.[151,1437,1084,2009]" pageId="1" pageNumber="555">
<materialsCitation id="C2A43CB4FF81FF896B94FE71FD97D48F" ID-GBIF-Occurrence="1503373107" collectingDate="2005-08-04" collectorName="V. Huhta &amp; Zoological Museum &amp; Helsinki University" country="Finland" latitude="67.48333" location="Material" longLatPrecision="990" longitude="24.116667" municipality="Kolari" pageId="1" pageNumber="555" specimenCount="1" typeStatus="holotype">
<emphasis id="40B8EAFBFF81FF896B94FE71FE96D447" bold="true" box="[151,379,1084,1109]" pageId="1" pageNumber="555">
<location id="77136032FF81FF896B94FE71FF12D447" LSID="urn:lsid:plazi:treatment:FA6587FFFF81FF8D6B94F99DFE4CD3DA:77136032FF81FF896B94FE71FF12D447" box="[151,255,1084,1109]" country="Finland" latitude="67.48333" longLatPrecision="990" longitude="24.116667" municipality="Kolari" name="Material" pageId="1" pageNumber="555">Material</location>
examined
</emphasis>
.
<typeStatus id="AD77884BFF81FF896A8FFE71FE1AD447" box="[396,503,1084,1109]" pageId="1" pageNumber="555" type="holotype">Holotype</typeStatus>
, female:
<collectingCountry id="0ADB7679FF81FF896962FE71FD57D447" box="[609,698,1084,1109]" name="Finland" pageId="1" pageNumber="555">Finland</collectingCountry>
,
<collectingMunicipality id="9217AC93FF81FF8969C4FE71FCE2D447" box="[711,783,1084,1109]" pageId="1" pageNumber="555">Kolari</collectingMunicipality>
(
<geoCoordinate id="17F8502EFF81FF896823FE71FC63D446" box="[800,910,1084,1109]" direction="north" orientation="latitude" pageId="1" pageNumber="555" precision="925" value="67.48333">67°29 N</geoCoordinate>
,
<geoCoordinate id="17F8502EFF81FF896898FE71FC1AD446" box="[923,1015,1084,1108]" direction="east" orientation="longitude" pageId="1" pageNumber="555" precision="925" value="24.116667">24°7 E</geoCoordinate>
), wet coniferous forest (
<taxonomicName id="B5CC4D6AFF81FF896E12FE70FA78D447" box="[1297,1429,1084,1109]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="1" pageNumber="555" phylum="Tracheophyta" rank="species" species="abies">
<emphasis id="40B8EAFBFF81FF896E12FE70FA78D447" box="[1297,1429,1084,1109]" italics="true" pageId="1" pageNumber="555">Picea abies</emphasis>
</taxonomicName>
,
<taxonomicName id="B5CC4D6AFF81FF896B94FE2CFEA1D46A" box="[151,332,1119,1144]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="1" pageNumber="555" phylum="Tracheophyta" rank="species" species="sylvestris">
<emphasis id="40B8EAFBFF81FF896B94FE2CFEA1D46A" box="[151,332,1119,1144]" italics="true" pageId="1" pageNumber="555">Pinus sylvestris</emphasis>
</taxonomicName>
,
<taxonomicName id="B5CC4D6AFF81FF896A5CFE2CFE39D46A" box="[351,468,1121,1144]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="1" pageNumber="555" phylum="Tracheophyta" rank="genus">
<emphasis id="40B8EAFBFF81FF896A5CFE2CFE39D46A" box="[351,468,1121,1144]" italics="true" pageId="1" pageNumber="555">Vaccinium</emphasis>
</taxonomicName>
spp., mosses), soil/litter sample,
<date id="06721029FF81FF896864FE2DFBF7D46A" box="[871,1050,1119,1144]" pageId="1" pageNumber="555" value="2005-08-04">
<collectingDate id="1636E9C1FF81FF896864FE2DFBF7D46A" box="[871,1050,1119,1144]" pageId="1" pageNumber="555" value="2005-08-04">4 August 2005</collectingDate>
</date>
, leg.
<collectorName id="DF39533FFF81FF896F61FE2DFB23D46A" box="[1122,1230,1119,1144]" pageId="1" pageNumber="555">V. Huhta</collectorName>
, deposited in the
<collectorName id="DF39533FFF81FF896B94FEC9FE94D48E" box="[151,377,1156,1181]" pageId="1" pageNumber="555">Zoological Museum</collectorName>
of
<collectorName id="DF39533FFF81FF896A9EFEC9FD97D48F" box="[413,634,1156,1181]" pageId="1" pageNumber="555">Helsinki University</collectorName>
</materialsCitation>
,
<materialsCitation id="C2A43CB4FF81FF896986FEC9FB9ED48F" ID-GBIF-Occurrence="1503373111" box="[645,1139,1156,1181]" country="Finland" location="Finland" pageId="1" pageNumber="555" specimenCount="1" typeStatus="paratype">
<collectingCountry id="0ADB7679FF81FF896986FEC9FD37D48F" box="[645,730,1156,1181]" name="Finland" pageId="1" pageNumber="555">Finland</collectingCountry>
(MusZHki).
<typeStatus id="AD77884BFF81FF89686DFEC8FC32D48F" box="[878,991,1157,1181]" pageId="1" pageNumber="555" type="paratype">Paratypes</typeStatus>
: two females
</materialsCitation>
,
<materialsCitation id="C2A43CB4FF81FF896F7DFEC8FE5ED4D2" ID-GBIF-Occurrence="1503373116" country="Finland" location="Finland" pageId="1" pageNumber="555" specimenCount="1" typeStatus="holotype">
same data as the
<typeStatus id="AD77884BFF81FF896E39FEC9FA71D48F" box="[1338,1436,1156,1181]" pageId="1" pageNumber="555" type="holotype">holotype</typeStatus>
(MusZHki), two females
</materialsCitation>
,
<materialsCitation id="C2A43CB4FF81FF896AC3FEE5FF08D4F7" ID-GBIF-Occurrence="1503373112" country="Russia" location="Finland" pageId="1" pageNumber="555" specimenCount="1" stateProvince="Saint Petersburg" typeStatus="holotype">
same data, Zoological Institute of the Russian Academy of Sciences,
<collectingRegion id="B008F80BFF81FF896FD8FEEAFA75D4D2" box="[1243,1432,1191,1216]" country="Russia" name="Saint Petersburg" pageId="1" pageNumber="555">Saint Petersburg</collectingRegion>
,
<collectingCountry id="0ADB7679FF81FF896B94FE81FF08D4F7" box="[151,229,1228,1253]" name="Russia" pageId="1" pageNumber="555">Russia</collectingCountry>
</materialsCitation>
;
<materialsCitation id="C2A43CB4FF81FF896BF3FE81FCF7D542" ID-GBIF-Occurrence="1503373109" collectingDate="2006-06-27" collectorName="South Siberia, NE &amp; N. V. Ananieva &amp; Vladimirova &amp; Siberian Zoological Museum &amp; Institute &amp; Systematics &amp; Ecology &amp; Animals &amp; Russian Academy &amp; Sciences" country="Russia" county="Archa Mt." elevation="1600" latitude="51.516666" location="South Siberia" longLatPrecision="1090" longitude="87.433334" municipality="Range" pageId="1" pageNumber="555" specimenCount="1" stateProvince="Altai" typeStatus="holotype">
four females,
<collectorName id="DF39533FFF81FF896A8FFE81FDBBD4F6" box="[396,598,1228,1253]" pageId="1" pageNumber="555">
<location id="77136032FF81FF896A8FFE81FDCBD4F7" LSID="urn:lsid:plazi:treatment:FA6587FFFF81FF8D6B94F99DFE4CD3DA:77136032FF81FF896A8FFE81FDCBD4F7" box="[396,550,1228,1253]" country="Russia" county="Archa Mt." latitude="51.516666" longLatPrecision="1090" longitude="87.433334" municipality="Range" name="South Siberia" pageId="1" pageNumber="555" stateProvince="Altai">South Siberia</location>
, NE
</collectorName>
<collectingRegion id="B008F80BFF81FF89695DFE81FD74D4F7" box="[606,665,1228,1253]" country="Russia" name="Altay" pageId="1" pageNumber="555">Altai</collectingRegion>
,
<location id="77136032FF81FF8969A7FE81FC8AD4F7" LSID="urn:lsid:plazi:treatment:FA6587FFFF81FF8D6B94F99DFE4CD3DA:77136032FF81FF8969A7FE81FC8AD4F7" box="[676,871,1228,1253]" country="Russia" county="Archa Mt." latitude="51.516666" longLatPrecision="1090" longitude="87.433334" municipality="Range" name="Turochak district" pageId="1" pageNumber="555" stateProvince="Altai">Turochak district</location>
, Altyn-Tu
<collectingMunicipality id="9217AC93FF81FF8968E3FE80FBC7D4F7" box="[992,1066,1229,1253]" pageId="1" pageNumber="555">Range</collectingMunicipality>
,
<collectingCounty id="9B124E65FF81FF896F35FE81FB44D4F6" box="[1078,1193,1228,1253]" pageId="1" pageNumber="555">Archa Mt.</collectingCounty>
(
<geoCoordinate id="17F8502EFF81FF896FBAFE81FACDD4F6" box="[1209,1312,1228,1253]" direction="north" orientation="latitude" pageId="1" pageNumber="555" precision="925" value="51.516666">51°31' N</geoCoordinate>
,
<geoCoordinate id="17F8502EFF81FF896E28FE81FA62D4F6" box="[1323,1423,1228,1253]" direction="east" orientation="longitude" pageId="1" pageNumber="555" precision="925" value="87.433334">87°26' E</geoCoordinate>
),
<elevation id="F9E1D1DAFF81FF896B94FEA2FEC0D51A" box="[151,301,1263,1288]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="555" unit="m" value="1600.0">
<quantity id="B5349B0CFF81FF896B94FEA2FF03D515" box="[151,238,1263,1288]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="555" unit="m" value="1600.0">1600 m</quantity>
a.s.l.
</elevation>
, light forest (
<taxonomicName id="B5CC4D6AFF81FF896ACFFEBCFD87D51A" box="[460,618,1263,1288]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="1" pageNumber="555" phylum="Tracheophyta" rank="species" species="sibirica">
<emphasis id="40B8EAFBFF81FF896ACFFEBCFD87D51A" box="[460,618,1263,1288]" italics="true" pageId="1" pageNumber="555">Pinus sibirica</emphasis>
</taxonomicName>
,
<taxonomicName id="B5CC4D6AFF81FF896978FEA2FCBFD51A" box="[635,850,1263,1288]" class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" order="Fagales" pageId="1" pageNumber="555" phylum="Tracheophyta" rank="species" species="rotundifolia">
<emphasis id="40B8EAFBFF81FF896978FEA2FCBFD51A" box="[635,850,1263,1288]" italics="true" pageId="1" pageNumber="555">Betula rotundifolia</emphasis>
</taxonomicName>
) with screes, from soil sample,
<date id="06721029FF81FF896FCBFEBDFA88D51A" box="[1224,1381,1263,1288]" pageId="1" pageNumber="555" value="2006-06-27">
<collectingDate id="1636E9C1FF81FF896FCBFEBDFA88D51A" box="[1224,1381,1263,1288]" pageId="1" pageNumber="555" value="2006-06-27">27 June 2006</collectingDate>
</date>
, leg.
<collectorName id="DF39533FFF81FF896B94FF58FED3D53F" box="[151,318,1300,1325]" pageId="1" pageNumber="555">N.V. Ananieva</collectorName>
[
<collectorName id="DF39533FFF81FF896A4EFF59FE0DD53F" box="[333,480,1300,1325]" pageId="1" pageNumber="555">Vladimirova</collectorName>
] (
<collectorName id="DF39533FFF81FF896AF4FF59FCABD53E" box="[503,838,1300,1325]" pageId="1" pageNumber="555">Siberian Zoological Museum</collectorName>
,
<collectorName id="DF39533FFF81FF896850FF59FC42D53F" box="[851,943,1300,1325]" pageId="1" pageNumber="555">Institute</collectorName>
of
<collectorName id="DF39533FFF81FF8968D4FF59FBB0D53F" box="[983,1117,1300,1325]" pageId="1" pageNumber="555">Systematics</collectorName>
and
<collectorName id="DF39533FFF81FF896F94FF59FB19D53F" box="[1175,1268,1300,1325]" pageId="1" pageNumber="555">Ecology</collectorName>
of
<collectorName id="DF39533FFF81FF896E1EFF59FA91D53F" box="[1309,1404,1300,1325]" pageId="1" pageNumber="555">Animals</collectorName>
of the
<collectorName id="DF39533FFF81FF896BC1FF7AFE62D542" box="[194,399,1335,1360]" pageId="1" pageNumber="555">Russian Academy</collectorName>
of
<collectorName id="DF39533FFF81FF896ABBFF7AFDF0D542" box="[440,541,1335,1360]" pageId="1" pageNumber="555">Sciences</collectorName>
,
<collectingRegion id="B008F80BFF81FF896929FF7AFD5BD542" box="[554,694,1335,1360]" country="Russia" name="Novosibirsk" pageId="1" pageNumber="555">Novosibirsk</collectingRegion>
,
<collectingCountry id="0ADB7679FF81FF8969C1FF7AFCFFD542" box="[706,786,1335,1360]" name="Russia" pageId="1" pageNumber="555">Russia</collectingCountry>
)
</materialsCitation>
.
<materialsCitation id="C2A43CB4FF81FF896825FF7AFD4ED58A" ID-GBIF-Occurrence="1503373115" collectingDate="1994-08" country="Finland" latitude="62.216667" location="Other" longLatPrecision="1020" longitude="25.483334" municipality="Jyvaskyla" pageId="1" pageNumber="555" specimenCount="1" typeStatus="holotype">
<location id="77136032FF81FF896825FF7AFC8BD542" LSID="urn:lsid:plazi:treatment:FA6587FFFF81FF8D6B94F99DFE4CD3DA:77136032FF81FF896825FF7AFC8BD542" box="[806,870,1335,1360]" country="Finland" latitude="62.216667" longLatPrecision="1020" longitude="25.483334" municipality="Jyvaskyla" name="Other" pageId="1" pageNumber="555">Other</location>
material: one female:
<collectingCountry id="0ADB7679FF81FF896F69FF7AFB2FD542" box="[1130,1218,1335,1360]" name="Finland" pageId="1" pageNumber="555">Finland</collectingCountry>
,
<collectingMunicipality id="9217AC93FF81FF896FCCFF7AFAD0D542" box="[1231,1341,1335,1360]" pageId="1" pageNumber="555">Jyväskylä</collectingMunicipality>
(
<geoCoordinate id="17F8502EFF81FF896E4CFF7AFF43D566" direction="north" orientation="latitude" pageId="1" pageNumber="555" precision="925" value="62.216667">62°13 N</geoCoordinate>
,
<geoCoordinate id="17F8502EFF81FF896BBAFF11FECDD566" box="[185,288,1372,1397]" direction="east" orientation="longitude" pageId="1" pageNumber="555" precision="925" value="25.483334">25°29 E</geoCoordinate>
), old mesic coniferous forest (
<taxonomicName id="B5CC4D6AFF81FF896970FF10FD18D567" box="[627,757,1372,1397]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="1" pageNumber="555" phylum="Tracheophyta" rank="species" species="abies">
<emphasis id="40B8EAFBFF81FF896970FF10FD18D567" box="[627,757,1372,1397]" italics="true" pageId="1" pageNumber="555">Picea abies</emphasis>
</taxonomicName>
,
<taxonomicName id="B5CC4D6AFF81FF896801FF10FC5CD567" box="[770,945,1372,1397]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="1" pageNumber="555" phylum="Tracheophyta" rank="species" species="sylvestris">
<emphasis id="40B8EAFBFF81FF896801FF10FC5CD567" box="[770,945,1372,1397]" italics="true" pageId="1" pageNumber="555">Pinus sylvestris</emphasis>
</taxonomicName>
,
<taxonomicName id="B5CC4D6AFF81FF8968BCFF10FB70D567" box="[959,1181,1372,1397]" class="Magnoliopsida" family="Ericaceae" genus="Vaccinium" kingdom="Plantae" order="Ericales" pageId="1" pageNumber="555" phylum="Tracheophyta" rank="species" species="myrtillus">
<emphasis id="40B8EAFBFF81FF8968BCFF10FB70D567" box="[959,1181,1372,1397]" italics="true" pageId="1" pageNumber="555">Vaccinium myrtillus</emphasis>
</taxonomicName>
, mosses)
<date id="06721029FF81FF896E0FFF11FA71D566" box="[1292,1436,1372,1396]" pageId="1" pageNumber="555" value="1994-08">
<collectingDate id="1636E9C1FF81FF896E0FFF11FA71D566" box="[1292,1436,1372,1396]" pageId="1" pageNumber="555" value="1994-08">August 1994</collectingDate>
</date>
(this specimen was lost during postal transfer).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3AD66562FF81FF896BC4FFE9FDA6D67D" pageId="1" pageNumber="555" type="diagnosis">
<paragraph id="727336E9FF81FF896BC4FFE9FDA6D67D" blockId="1.[151,1437,1084,2009]" pageId="1" pageNumber="555">
<emphasis id="40B8EAFBFF81FF896BC4FFE9FE0BD5AF" bold="true" box="[199,486,1444,1469]" pageId="1" pageNumber="555">Diagnosis (female only).</emphasis>
Dorsal shield with lateral incisions, reticulate, with full set of relatively short setae. Sternal shield fused with endopodal elements III and IIIII, reticulate. Anal shield rather large, longer than sternal shield, broadly oval or roundish-triangular, widest posterior to level of gv3, its lateral margins uneven, with roundish ledges; post-anal seta twice or more longer than para-anal setae. Peritremes long. Gnathotectum with three simple prongs. Corniculus and chela of normal proportions, fixed digit with four teeth. Claw I smaller than claws IIIV. Tibia IV with seven setae.
</paragraph>
</subSubSection>
<subSubSection id="3AD66562FF81FF8A6BC4FC31FE61D787" lastPageId="2" lastPageNumber="556" pageId="1" pageNumber="555" type="description">
<paragraph id="727336E9FF81FF896BC4FC31FDA1D6CF" blockId="1.[151,1437,1084,2009]" pageId="1" pageNumber="555">
<emphasis id="40B8EAFBFF81FF896BC4FC31FE2FD687" bold="true" box="[199,450,1660,1685]" pageId="1" pageNumber="555">Description (female).</emphasis>
Middle sized, short-legged, yellowish or brownish mites. Idiosomal shields moderately sclerotized, mainly reticulate. Most dorsal idiosomal setae and leg setae short, needle-shaped. Some setae of tarsi I and IV elongated and hair-like distally.
</paragraph>
<paragraph id="727336E9FF81FF8A6BC4FCAAFC33D0EA" blockId="1.[151,1437,1084,2009]" lastBlockId="2.[151,1437,151,1941]" lastPageId="2" lastPageNumber="556" pageId="1" pageNumber="555">
<emphasis id="40B8EAFBFF81FF896BC4FCAAFE79D712" box="[199,404,1767,1792]" italics="true" pageId="1" pageNumber="555">Idiosomal dorsum</emphasis>
(
<figureCitation id="EAF72A6CFF81FF896AA6FCAAFDE3D712" box="[421,526,1767,1792]" captionStart="FIGURES 1 5" captionStartId="3.[151,264,1930,1953]" captionTargetBox="[179,1427,199,1902]" captionTargetId="figure@3.[151,1436,195,1907]" captionTargetPageId="3" captionText="FIGURES 1 5. Arctoseius ambiguus sp. nov., female. 1. tritosternum; 2. ambulacrum and tarsus II distally, ventral aspect; 3. gnathotectum; 4. peritrematal region; 5. idiosomal dorsum. Scales: 1 2 25 µm, 3 5 50 µm." httpUri="https://zenodo.org/record/580967/files/figure.png" pageId="1" pageNumber="555">Figs 4, 5</figureCitation>
). Dorsal shield 468512 x 264288,
<emphasis id="40B8EAFBFF81FF8968BAFCAAFBE9D712" box="[953,1028,1767,1792]" italics="true" pageId="1" pageNumber="555">lD/wD</emphasis>
ca 1.721.82; its maximal width at level of opisthonotal setae
<emphasis id="40B8EAFBFF81FF896ACAFD40FE3AD736" box="[457,471,1805,1828]" italics="true" pageId="1" pageNumber="555">S</emphasis>
3 (
<figureCitation id="EAF72A6CFF81FF896AF6FD41FDD6D737" box="[501,571,1804,1829]" captionStart="FIGURES 1 5" captionStartId="3.[151,264,1930,1953]" captionTargetBox="[179,1427,199,1902]" captionTargetId="figure@3.[151,1436,195,1907]" captionTargetPageId="3" captionText="FIGURES 1 5. Arctoseius ambiguus sp. nov., female. 1. tritosternum; 2. ambulacrum and tarsus II distally, ventral aspect; 3. gnathotectum; 4. peritrematal region; 5. idiosomal dorsum. Scales: 1 2 25 µm, 3 5 50 µm." httpUri="https://zenodo.org/record/580967/files/figure.png" pageId="1" pageNumber="555">Fig. 5</figureCitation>
); lateral incisions of moderate length (3044), transverse, relatively narrow, often extended by lyrifissure
<emphasis id="40B8EAFBFF81FF896AEEFD62FDE3D75A" box="[493,526,1839,1864]" italics="true" pageId="1" pageNumber="555">idz</emphasis>
6; shield reticulation more distinct laterally. Podonotal region with 17 pairs of simple setae (
<emphasis id="40B8EAFBFF81FF896A31FD18FED0D77E" box="[306,317,1877,1900]" italics="true" pageId="1" pageNumber="555">z</emphasis>
3 present);
<emphasis id="40B8EAFBFF81FF896ABFFD18FE2AD77E" box="[444,455,1877,1900]" italics="true" pageId="1" pageNumber="555">s</emphasis>
2 usually on soft cuticle, seldom asymmetrically on shield margin. Opisthonotal region with 14 pairs of setae (
<emphasis id="40B8EAFBFF81FF896A9FFD35FE47D79D" box="[412,426,1912,1935]" italics="true" pageId="1" pageNumber="555">S</emphasis>
2 always absent). Among podonotal setae,
<emphasis id="40B8EAFBFF81FF896892FD34FC71D782" box="[913,924,1913,1936]" italics="true" pageId="1" pageNumber="555">z</emphasis>
1 (1218),
<emphasis id="40B8EAFBFF81FF896F19FD34FBC8D782" box="[1050,1061,1913,1936]" italics="true" pageId="1" pageNumber="555">s</emphasis>
1 (1822) and
<emphasis id="40B8EAFBFF81FF896FCFFD34FB3AD782" box="[1228,1239,1913,1936]" italics="true" pageId="1" pageNumber="555">s</emphasis>
2 (1823) shorter than most others (1833). On opisthonotal region, setae
<emphasis id="40B8EAFBFF81FF89680BFDD0FCF9D7A6" box="[776,788,1949,1972]" italics="true" pageId="1" pageNumber="555">J</emphasis>
14 (1321) shorter than others (1930), except
<emphasis id="40B8EAFBFF81FF896E2CFDD0FAD6D7A6" box="[1327,1339,1949,1972]" italics="true" pageId="1" pageNumber="555">J</emphasis>
5 clearly shortest (1217) and Z5 clearly longest (3542);
<emphasis id="40B8EAFBFF81FF8969EDFD8CFD17D7CA" box="[750,762,1985,2008]" italics="true" pageId="1" pageNumber="555">J</emphasis>
(14)
<emphasis id="40B8EAFBFF81FF896834FD8CFC4DD7CA" box="[823,928,1985,2008]" italics="true" pageId="1" pageNumber="555">min/jmax</emphasis>
0.620.76,
<emphasis id="40B8EAFBFF81FF896F30FD8CFBB5D7CA" box="[1075,1112,1985,2008]" italics="true" pageId="1" pageNumber="555">Z5/</emphasis>
(
<emphasis id="40B8EAFBFF81FF896F61FD8CFB9CD7CA" box="[1122,1137,1985,2008]" italics="true" pageId="1" pageNumber="555">Z</emphasis>
5-
<emphasis id="40B8EAFBFF81FF896F8BFD8CFB7AD7CA" box="[1160,1175,1985,2008]" italics="true" pageId="1" pageNumber="555">Z</emphasis>
5) 0.350.46; (
<emphasis id="40B8EAFBFF81FF896E4FFD8CFAB5D7CA" box="[1356,1368,1985,2008]" italics="true" pageId="1" pageNumber="555">J</emphasis>
4-
<emphasis id="40B8EAFBFF81FF896E73FD8CFA91D7CA" box="[1392,1404,1985,2008]" italics="true" pageId="1" pageNumber="555">J</emphasis>
4)
<emphasis id="40B8EAFBFF81FF896E97FD8CFA71D7CA" box="[1428,1436,1985,2008]" italics="true" pageId="1" pageNumber="555">/</emphasis>
(
<emphasis id="40B8EAFBFF82FF8A6BA3FAD4FF41D0A2" box="[160,172,153,176]" italics="true" pageId="2" pageNumber="556">J</emphasis>
3-
<emphasis id="40B8EAFBFF82FF8A6BC7FAD4FF3DD0A2" box="[196,208,153,176]" italics="true" pageId="2" pageNumber="556">J</emphasis>
3)1.331.72. Dorsal shield with five pairs of gland pores: poorly visible
<emphasis id="40B8EAFBFF82FF8A6F0AFADAFBC0D0A2" box="[1033,1069,151,176]" italics="true" pageId="2" pageNumber="556">gdj</emphasis>
2,
<emphasis id="40B8EAFBFF82FF8A6F49FADAFB80D0A2" box="[1098,1133,151,176]" italics="true" pageId="2" pageNumber="556">gdj</emphasis>
4,
<emphasis id="40B8EAFBFF82FF8A6F8FFADAFB5ED0A2" box="[1164,1203,151,176]" italics="true" pageId="2" pageNumber="556">gdz</emphasis>
6, and distinct
<emphasis id="40B8EAFBFF82FF8A6E58FADAFA6AD0A2" box="[1371,1415,151,176]" italics="true" pageId="2" pageNumber="556">gdZ</emphasis>
3,
<emphasis id="40B8EAFBFF82FF8A6B94FAF1FF2ED0C7" box="[151,195,188,213]" italics="true" pageId="2" pageNumber="556">gdZ</emphasis>
4. All marginal setae on soft cuticle; four setae in series
<emphasis id="40B8EAFBFF82FF8A6841FAF0FCA0D0C6" box="[834,845,189,212]" italics="true" pageId="2" pageNumber="556">r</emphasis>
(
<emphasis id="40B8EAFBFF82FF8A685DFAF0FC84D0C6" box="[862,873,189,212]" italics="true" pageId="2" pageNumber="556">r</emphasis>
25; their length 1927) and 45 setae in series
<emphasis id="40B8EAFBFF82FF8A6E88FAF0FA71D0C6" box="[1419,1436,189,212]" italics="true" pageId="2" pageNumber="556">R</emphasis>
(
<emphasis id="40B8EAFBFF82FF8A6BA3FAACFF5CD0EA" box="[160,177,225,248]" italics="true" pageId="2" pageNumber="556">R</emphasis>
14 or
<emphasis id="40B8EAFBFF82FF8A6A03FAACFEFCD0EA" box="[256,273,225,248]" italics="true" pageId="2" pageNumber="556">R</emphasis>
1-5; 1522); setae of series
<emphasis id="40B8EAFBFF82FF8A694BFAACFDB4D0EA" box="[584,601,225,248]" italics="true" pageId="2" pageNumber="556">R</emphasis>
usually inserted ventrally (
<figureCitation id="EAF72A6CFF82FF8A6889FA92FC22D0EA" box="[906,975,223,248]" captionStart="FIGURES 6 11" captionStartId="4.[151,264,1925,1948]" captionTargetBox="[162,1430,195,1884]" captionTargetId="figure@4.[151,1436,193,1904]" captionTargetPageId="4" captionText="FIGURES 6 11. Arctoseius ambiguus sp. nov., female. 6. acetabula II IV and exopodal platelets; 7. subcapitulum; 8. idiosomal ventrum; 9. anal shield; 10. cheliceral digits, dorsal aspect; 11. chelicera without basal joint, paraxial aspect. Scales: 6 9 50 µm, 10 11 25 µm." httpUri="https://zenodo.org/record/580968/files/figure.png" pageId="2" pageNumber="556">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="727336E9FF82FF8A6BC4FB49FD11D44A" blockId="2.[151,1437,151,1941]" pageId="2" pageNumber="556">
<emphasis id="40B8EAFBFF82FF8A6BC4FB49FE6AD10E" box="[199,391,260,285]" italics="true" pageId="2" pageNumber="556">Idiosomal venter</emphasis>
(
<figureCitation id="EAF72A6CFF82FF8A6A9BFB49FE0FD10E" box="[408,482,260,285]" captionStart="FIGURES 1 5" captionStartId="3.[151,264,1930,1953]" captionTargetBox="[179,1427,199,1902]" captionTargetId="figure@3.[151,1436,195,1907]" captionTargetPageId="3" captionText="FIGURES 1 5. Arctoseius ambiguus sp. nov., female. 1. tritosternum; 2. ambulacrum and tarsus II distally, ventral aspect; 3. gnathotectum; 4. peritrematal region; 5. idiosomal dorsum. Scales: 1 2 25 µm, 3 5 50 µm." httpUri="https://zenodo.org/record/580967/files/figure.png" pageId="2" pageNumber="556">Figs 1</figureCitation>
,
<figureCitation id="EAF72A6CFF82FF8A6AF0FB49FDACD10E" box="[499,577,260,285]" captionStart="FIGURES 6 11" captionStartId="4.[151,264,1925,1948]" captionTargetBox="[162,1430,195,1884]" captionTargetId="figure@4.[151,1436,193,1904]" captionTargetPageId="4" captionText="FIGURES 6 11. Arctoseius ambiguus sp. nov., female. 6. acetabula II IV and exopodal platelets; 7. subcapitulum; 8. idiosomal ventrum; 9. anal shield; 10. cheliceral digits, dorsal aspect; 11. chelicera without basal joint, paraxial aspect. Scales: 6 9 50 µm, 10 11 25 µm." httpUri="https://zenodo.org/record/580968/files/figure.png" pageId="2" pageNumber="556">6, 8, 9</figureCitation>
). Base of tritosternum trapezoidal (
<date id="06721029FF82FF8A68DFFB49FB8ED10F" box="[988,1123,260,285]" pageId="2" pageNumber="556" value="2016-10-21" valueMax="2016-10-23" valueMin="2016-10-21">2123 x 16</date>
18); laciniae (6880) with sparse large barbs, free for
<emphasis id="40B8EAFBFF82FF8A6ADFFB64FE1AD152" box="[476,503,297,320]" italics="true" pageId="2" pageNumber="556">ca</emphasis>
0.80.9 of lengths, their fused basal area fringed anteriorly (
<figureCitation id="EAF72A6CFF82FF8A6FC9FB6AFAE2D152" box="[1226,1295,295,320]" captionStart="FIGURES 1 5" captionStartId="3.[151,264,1930,1953]" captionTargetBox="[179,1427,199,1902]" captionTargetId="figure@3.[151,1436,195,1907]" captionTargetPageId="3" captionText="FIGURES 1 5. Arctoseius ambiguus sp. nov., female. 1. tritosternum; 2. ambulacrum and tarsus II distally, ventral aspect; 3. gnathotectum; 4. peritrematal region; 5. idiosomal dorsum. Scales: 1 2 25 µm, 3 5 50 µm." httpUri="https://zenodo.org/record/580967/files/figure.png" pageId="2" pageNumber="556">Fig. 1</figureCitation>
). Presternal platelets small, weakly sclerotized, lineate, adjacent to sternal shield (
<figureCitation id="EAF72A6CFF82FF8A68B3FB01FC18D176" box="[944,1013,332,357]" captionStart="FIGURES 6 11" captionStartId="4.[151,264,1925,1948]" captionTargetBox="[162,1430,195,1884]" captionTargetId="figure@4.[151,1436,193,1904]" captionTargetPageId="4" captionText="FIGURES 6 11. Arctoseius ambiguus sp. nov., female. 6. acetabula II IV and exopodal platelets; 7. subcapitulum; 8. idiosomal ventrum; 9. anal shield; 10. cheliceral digits, dorsal aspect; 11. chelicera without basal joint, paraxial aspect. Scales: 6 9 50 µm, 10 11 25 µm." httpUri="https://zenodo.org/record/580968/files/figure.png" pageId="2" pageNumber="556">Fig. 8</figureCitation>
). Sternal shield as long (108115) as wide (102116),
<emphasis id="40B8EAFBFF82FF8A6A57FB22FE4DD19A" box="[340,416,367,392]" italics="true" pageId="2" pageNumber="556">lSt/wSt</emphasis>
0.971.06, fused with endopodal platelets between coxae III and IIIII. Narrow distal part of endopodal projection between coxae III nearly bacilliform, sometimes separated, their lateral part encompassing opening of gland
<emphasis id="40B8EAFBFF82FF8A6906FBFAFDC0D1C2" box="[517,557,439,464]" italics="true" pageId="2" pageNumber="556">gvb</emphasis>
. Sternal shield entirely reticulated with ornamentation more distinct in lateral parts; anterior margin straight or slightly concave; posterior margin more or less straight. Sternal shield with typical setae
<emphasis id="40B8EAFBFF82FF8A6BDBF84CFF06D20A" box="[216,235,513,536]" italics="true" pageId="2" pageNumber="556">st</emphasis>
13 (2635) and lyrifissures
<date id="06721029FF82FF8A6935F84CFD96D20A" box="[566,635,512,536]" pageId="2" pageNumber="556">
<emphasis id="40B8EAFBFF82FF8A6935F84CFDA7D20A" box="[566,586,513,536]" italics="true" pageId="2" pageNumber="556">iv</emphasis>
13.
</date>
Setae
<emphasis id="40B8EAFBFF82FF8A69C9F84CFD30D20A" box="[714,733,513,536]" italics="true" pageId="2" pageNumber="556">st</emphasis>
4 (2428) on soft cuticle. Endopodal strips between coxae III and IV free, rather narrow. Epigynal shield narrowly flask-shaped (118136 х 4752), reticulated posteriorly, fully punctate with dots much denser in anterior part (
<figureCitation id="EAF72A6CFF82FF8A69DBF80AFCCCD272" box="[728,801,583,608]" captionStart="FIGURES 6 11" captionStartId="4.[151,264,1925,1948]" captionTargetBox="[162,1430,195,1884]" captionTargetId="figure@4.[151,1436,193,1904]" captionTargetPageId="4" captionText="FIGURES 6 11. Arctoseius ambiguus sp. nov., female. 6. acetabula II IV and exopodal platelets; 7. subcapitulum; 8. idiosomal ventrum; 9. anal shield; 10. cheliceral digits, dorsal aspect; 11. chelicera without basal joint, paraxial aspect. Scales: 6 9 50 µm, 10 11 25 µm." httpUri="https://zenodo.org/record/580968/files/figure.png" pageId="2" pageNumber="556">Fig. 8</figureCitation>
); its hyaline flap well distant from sternal shield, and posterior margin broadly convex; setae
<emphasis id="40B8EAFBFF82FF8A6958F820FD83D296" box="[603,622,621,644]" italics="true" pageId="2" pageNumber="556">st</emphasis>
5 (2426) and paragenital poroids
<date id="06721029FF82FF8A68FBF820FBF7D297" box="[1016,1050,620,645]" pageId="2" pageNumber="556">
<emphasis id="40B8EAFBFF82FF8A68FBF820FBE1D296" box="[1016,1036,621,644]" italics="true" pageId="2" pageNumber="556">iv</emphasis>
5
</date>
on soft cuticle. Opening of gland
<emphasis id="40B8EAFBFF82FF8A6B94F8DCFF5FD2BA" box="[151,178,657,680]" italics="true" pageId="2" pageNumber="556">gv</emphasis>
2 on soft cuticle between
<emphasis id="40B8EAFBFF82FF8A6ACCF8DCFE0FD2BA" box="[463,482,657,680]" italics="true" pageId="2" pageNumber="556">st</emphasis>
5 and end of peritrematal-exopodal strip. Postgenital platelets not visible. Anal shield relatively large (116124 x 101120; [
<emphasis id="40B8EAFBFF82FF8A6951F8F9FD94D2DF" box="[594,633,692,717]" italics="true" pageId="2" pageNumber="556">lAn</emphasis>
x
<emphasis id="40B8EAFBFF82FF8A699AF8F9FD26D2D9" box="[665,715,692,715]" italics="true" pageId="2" pageNumber="556">wAn</emphasis>
] / [
<emphasis id="40B8EAFBFF82FF8A69FBF8F9FCF9D2DF" box="[760,788,692,717]" italics="true" pageId="2" pageNumber="556">lD</emphasis>
x
<emphasis id="40B8EAFBFF82FF8A6836F8F8FCB1D2DE" box="[821,860,693,716]" italics="true" pageId="2" pageNumber="556">wD</emphasis>
] 8.410.6 %), slightly longer than wide (
<emphasis id="40B8EAFBFF82FF8A6E3FF8F9FA71D2DF" box="[1340,1436,692,717]" italics="true" pageId="2" pageNumber="556">lAn/wAn</emphasis>
1.031.19), broadly oval or roundish-triangular, widest posterior to level of gland
<emphasis id="40B8EAFBFF82FF8A6F2BF894FBAED2E2" box="[1064,1091,729,752]" italics="true" pageId="2" pageNumber="556">gv</emphasis>
3; its lateral margins bent, with roundish ledges of different size (
<figureCitation id="EAF72A6CFF82FF8A6912F8B1FD9AD306" box="[529,631,764,789]" captionStart="FIGURES 6 11" captionStartId="4.[151,264,1925,1948]" captionTargetBox="[162,1430,195,1884]" captionTargetId="figure@4.[151,1436,193,1904]" captionTargetPageId="4" captionText="FIGURES 6 11. Arctoseius ambiguus sp. nov., female. 6. acetabula II IV and exopodal platelets; 7. subcapitulum; 8. idiosomal ventrum; 9. anal shield; 10. cheliceral digits, dorsal aspect; 11. chelicera without basal joint, paraxial aspect. Scales: 6 9 50 µm, 10 11 25 µm." httpUri="https://zenodo.org/record/580968/files/figure.png" pageId="2" pageNumber="556">Figs 8, 9</figureCitation>
); shield weakly reticulate anteriorly and coarsely punctate laterally and especially posteriorly; post-anal seta (4449) twice or more longer than para-anal setae (2022); papillae of cribrum forming rows and arcs. Opisthogastral setae (810 pairs,
<emphasis id="40B8EAFBFF82FF8A6884F908FC49D34E" box="[903,932,837,860]" italics="true" pageId="2" pageNumber="556">JV</emphasis>
15,
<emphasis id="40B8EAFBFF82FF8A68DDF908FC12D34E" box="[990,1023,837,860]" italics="true" pageId="2" pageNumber="556">ZV</emphasis>
15;
<emphasis id="40B8EAFBFF82FF8A6F39F908FBB6D34E" box="[1082,1115,837,860]" italics="true" pageId="2" pageNumber="556">ZV</emphasis>
3 and
<emphasis id="40B8EAFBFF82FF8A6FA0F908FB29D34E" box="[1187,1220,837,860]" italics="true" pageId="2" pageNumber="556">ZV</emphasis>
5 often omitted) of varying length,
<emphasis id="40B8EAFBFF82FF8A6A49F924FE8AD392" box="[330,359,873,896]" italics="true" pageId="2" pageNumber="556">JV</emphasis>
5 usually longest (3236),
<emphasis id="40B8EAFBFF82FF8A699AF924FD54D392" box="[665,697,873,896]" italics="true" pageId="2" pageNumber="556">ZV</emphasis>
1 usually shortest (1416); among others, setae in series
<emphasis id="40B8EAFBFF82FF8A6E40F924FA8DD392" box="[1347,1376,873,896]" italics="true" pageId="2" pageNumber="556">JV</emphasis>
(20 34) usually longer than setae in series
<emphasis id="40B8EAFBFF82FF8A6948F9C0FD81D3B6" box="[587,620,909,932]" italics="true" pageId="2" pageNumber="556">ZV</emphasis>
(1626). Anterior metapodal platelets small, of variable form; posterior metapodal platelets elongate oval (32
<date id="06721029FF82FF8A694BF9FDFD53D3DA" box="[584,702,944,968]" pageId="2" pageNumber="556" value="1940-10-07" valueMax="1940-10-10" valueMin="1940-10-07">40 x 710</date>
). Exopodal platelets between coxae II and III, as well as between coxae III and IV, small rudimentary (
<figureCitation id="EAF72A6CFF82FF8A693FF999FD6FD3FF" box="[572,642,980,1005]" captionStart="FIGURES 6 11" captionStartId="4.[151,264,1925,1948]" captionTargetBox="[162,1430,195,1884]" captionTargetId="figure@4.[151,1436,193,1904]" captionTargetPageId="4" captionText="FIGURES 6 11. Arctoseius ambiguus sp. nov., female. 6. acetabula II IV and exopodal platelets; 7. subcapitulum; 8. idiosomal ventrum; 9. anal shield; 10. cheliceral digits, dorsal aspect; 11. chelicera without basal joint, paraxial aspect. Scales: 6 9 50 µm, 10 11 25 µm." httpUri="https://zenodo.org/record/580968/files/figure.png" pageId="2" pageNumber="556">Fig. 6</figureCitation>
), not visible in ventral aspect. Peritrematal shield normally developed (
<figureCitation id="EAF72A6CFF82FF8A6B9CF9BAFF04D402" box="[159,233,1015,1040]" captionStart="FIGURES 1 5" captionStartId="3.[151,264,1930,1953]" captionTargetBox="[179,1427,199,1902]" captionTargetId="figure@3.[151,1436,195,1907]" captionTargetPageId="3" captionText="FIGURES 1 5. Arctoseius ambiguus sp. nov., female. 1. tritosternum; 2. ambulacrum and tarsus II distally, ventral aspect; 3. gnathotectum; 4. peritrematal region; 5. idiosomal dorsum. Scales: 1 2 25 µm, 3 5 50 µm." httpUri="https://zenodo.org/record/580967/files/figure.png" pageId="2" pageNumber="556">Figs 4</figureCitation>
,
<figureCitation id="EAF72A6CFF82FF8A6BF5F9B5FEE8D402" box="[246,261,1016,1040]" captionStart="FIGURES 6 11" captionStartId="4.[151,264,1925,1948]" captionTargetBox="[162,1430,195,1884]" captionTargetId="figure@4.[151,1436,193,1904]" captionTargetPageId="4" captionText="FIGURES 6 11. Arctoseius ambiguus sp. nov., female. 6. acetabula II IV and exopodal platelets; 7. subcapitulum; 8. idiosomal ventrum; 9. anal shield; 10. cheliceral digits, dorsal aspect; 11. chelicera without basal joint, paraxial aspect. Scales: 6 9 50 µm, 10 11 25 µm." httpUri="https://zenodo.org/record/580968/files/figure.png" pageId="2" pageNumber="556">8</figureCitation>
); its posterior edge fused with exopodal strip surrounding coxa IV posteriorly (
<figureCitation id="EAF72A6CFF82FF8A6F8FF9BAFB3CD402" box="[1164,1233,1015,1040]" captionStart="FIGURES 6 11" captionStartId="4.[151,264,1925,1948]" captionTargetBox="[162,1430,195,1884]" captionTargetId="figure@4.[151,1436,193,1904]" captionTargetPageId="4" captionText="FIGURES 6 11. Arctoseius ambiguus sp. nov., female. 6. acetabula II IV and exopodal platelets; 7. subcapitulum; 8. idiosomal ventrum; 9. anal shield; 10. cheliceral digits, dorsal aspect; 11. chelicera without basal joint, paraxial aspect. Scales: 6 9 50 µm, 10 11 25 µm." httpUri="https://zenodo.org/record/580968/files/figure.png" pageId="2" pageNumber="556">Fig. 8</figureCitation>
); poroids
<emphasis id="40B8EAFBFF82FF8A6E46F9B4FAB6D402" box="[1349,1371,1017,1040]" italics="true" pageId="2" pageNumber="556">ip</emphasis>
1,
<emphasis id="40B8EAFBFF82FF8A6E7BF9B4FA63D402" box="[1400,1422,1017,1040]" italics="true" pageId="2" pageNumber="556">ip</emphasis>
2 and gland pores
<emphasis id="40B8EAFBFF82FF8A6A50FE50FE82D426" box="[339,367,1053,1076]" italics="true" pageId="2" pageNumber="556">gp</emphasis>
1,
<emphasis id="40B8EAFBFF82FF8A6A8FFE50FE45D426" box="[396,424,1053,1076]" italics="true" pageId="2" pageNumber="556">gp</emphasis>
2 present. Peritreme not shortened (226240 х 911), extending to frontal level of seta
<emphasis id="40B8EAFBFF82FF8A6E85FE50FA63D426" box="[1414,1422,1053,1076]" italics="true" pageId="2" pageNumber="556">j</emphasis>
2 anteriorly (
<figureCitation id="EAF72A6CFF82FF8A6A10FE72FEB5D44A" box="[275,344,1087,1112]" captionStart="FIGURES 1 5" captionStartId="3.[151,264,1930,1953]" captionTargetBox="[179,1427,199,1902]" captionTargetId="figure@3.[151,1436,195,1907]" captionTargetPageId="3" captionText="FIGURES 1 5. Arctoseius ambiguus sp. nov., female. 1. tritosternum; 2. ambulacrum and tarsus II distally, ventral aspect; 3. gnathotectum; 4. peritrematal region; 5. idiosomal dorsum. Scales: 1 2 25 µm, 3 5 50 µm." httpUri="https://zenodo.org/record/580967/files/figure.png" pageId="2" pageNumber="556">Fig. 4</figureCitation>
). Spermathecal apparatus not visible.
</paragraph>
<paragraph id="727336E9FF82FF8A6BC4FE29FD80D5F7" blockId="2.[151,1437,151,1941]" pageId="2" pageNumber="556">
<taxonomicName id="B5CC4D6AFF82FF8A6BC4FE29FEBBD46F" box="[199,342,1124,1149]" class="Arachnida" family="Syringophilidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="556" phylum="Arthropoda" rank="genus">
<emphasis id="40B8EAFBFF82FF8A6BC4FE29FEBBD46F" box="[199,342,1124,1149]" italics="true" pageId="2" pageNumber="556">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="EAF72A6CFF82FF8A6A66FE29FE41D46E" box="[357,428,1124,1149]" captionStart="FIGURES 1 5" captionStartId="3.[151,264,1930,1953]" captionTargetBox="[179,1427,199,1902]" captionTargetId="figure@3.[151,1436,195,1907]" captionTargetPageId="3" captionText="FIGURES 1 5. Arctoseius ambiguus sp. nov., female. 1. tritosternum; 2. ambulacrum and tarsus II distally, ventral aspect; 3. gnathotectum; 4. peritrematal region; 5. idiosomal dorsum. Scales: 1 2 25 µm, 3 5 50 µm." httpUri="https://zenodo.org/record/580967/files/figure.png" pageId="2" pageNumber="556">Figs 3</figureCitation>
,
<figureCitation id="EAF72A6CFF82FF8A6ABAFE28FDF1D46E" box="[441,540,1124,1148]" captionStart="FIGURES 6 11" captionStartId="4.[151,264,1925,1948]" captionTargetBox="[162,1430,195,1884]" captionTargetId="figure@4.[151,1436,193,1904]" captionTargetPageId="4" captionText="FIGURES 6 11. Arctoseius ambiguus sp. nov., female. 6. acetabula II IV and exopodal platelets; 7. subcapitulum; 8. idiosomal ventrum; 9. anal shield; 10. cheliceral digits, dorsal aspect; 11. chelicera without basal joint, paraxial aspect. Scales: 6 9 50 µm, 10 11 25 µm." httpUri="https://zenodo.org/record/580968/files/figure.png" pageId="2" pageNumber="556">7, 10, 11</figureCitation>
). Gnathotectum (
<date id="06721029FF82FF8A69DCFE29FCB3D46E" box="[735,862,1124,1149]" pageId="2" pageNumber="556" value="1930-10-16" valueMax="1930-10-22" valueMin="1930-10-16">1622 x 30</date>
40) triramous, with simple prongs of almost equal size (
<figureCitation id="EAF72A6CFF82FF8A6BD2FECAFEF9D4B2" box="[209,276,1159,1184]" captionStart="FIGURES 1 5" captionStartId="3.[151,264,1930,1953]" captionTargetBox="[179,1427,199,1902]" captionTargetId="figure@3.[151,1436,195,1907]" captionTargetPageId="3" captionText="FIGURES 1 5. Arctoseius ambiguus sp. nov., female. 1. tritosternum; 2. ambulacrum and tarsus II distally, ventral aspect; 3. gnathotectum; 4. peritrematal region; 5. idiosomal dorsum. Scales: 1 2 25 µm, 3 5 50 µm." httpUri="https://zenodo.org/record/580967/files/figure.png" pageId="2" pageNumber="556">Fig. 3</figureCitation>
). Subcapitulum almost as long (108112) as wide (100104) (
<figureCitation id="EAF72A6CFF82FF8A68C8FECAFBE2D4B2" box="[971,1039,1159,1184]" captionStart="FIGURES 6 11" captionStartId="4.[151,264,1925,1948]" captionTargetBox="[162,1430,195,1884]" captionTargetId="figure@4.[151,1436,193,1904]" captionTargetPageId="4" captionText="FIGURES 6 11. Arctoseius ambiguus sp. nov., female. 6. acetabula II IV and exopodal platelets; 7. subcapitulum; 8. idiosomal ventrum; 9. anal shield; 10. cheliceral digits, dorsal aspect; 11. chelicera without basal joint, paraxial aspect. Scales: 6 9 50 µm, 10 11 25 µm." httpUri="https://zenodo.org/record/580968/files/figure.png" pageId="2" pageNumber="556">Fig. 7</figureCitation>
). Deutosternum with seven rows of denticles (1220 denticles in each row); groove delineated laterally, slightly wider in the middle (up to 19). Hypostomatic setae
<emphasis id="40B8EAFBFF82FF8A6A7EFE82FE74D4FA" box="[381,409,1231,1256]" italics="true" pageId="2" pageNumber="556">hp</emphasis>
1 (3336) and
<emphasis id="40B8EAFBFF82FF8A6940FE82FDB2D4FA" box="[579,607,1231,1256]" italics="true" pageId="2" pageNumber="556">hp</emphasis>
3 (3638) longer than
<emphasis id="40B8EAFBFF82FF8A6862FE82FC90D4FA" box="[865,893,1231,1256]" italics="true" pageId="2" pageNumber="556">hp</emphasis>
2 (1921) and
<emphasis id="40B8EAFBFF82FF8A6F25FE9CFBACD4FA" box="[1062,1089,1233,1256]" italics="true" pageId="2" pageNumber="556">pc</emphasis>
(2226); all setae needle-like, attenuate. Corniculi small, of normal proportions, 2934 х 1012;
<emphasis id="40B8EAFBFF82FF8A689CFEB9FC06D51F" box="[927,1003,1268,1293]" italics="true" pageId="2" pageNumber="556">lCo/lD</emphasis>
6.106.64 %. Internal malae slightly shorter than corniculi, with lateral margins fimbriated basally. Chelicera medium-sized (
<figureCitation id="EAF72A6CFF82FF8A6F90FF5AFAF4D522" box="[1171,1305,1303,1328]" captionStart="FIGURES 6 11" captionStartId="4.[151,264,1925,1948]" captionTargetBox="[162,1430,195,1884]" captionTargetId="figure@4.[151,1436,193,1904]" captionTargetPageId="4" captionText="FIGURES 6 11. Arctoseius ambiguus sp. nov., female. 6. acetabula II IV and exopodal platelets; 7. subcapitulum; 8. idiosomal ventrum; 9. anal shield; 10. cheliceral digits, dorsal aspect; 11. chelicera without basal joint, paraxial aspect. Scales: 6 9 50 µm, 10 11 25 µm." httpUri="https://zenodo.org/record/580968/files/figure.png" pageId="2" pageNumber="556">Figs 10, 11</figureCitation>
), its length without basal segment 108116; cheliceral digits of moderate size (4750,
<emphasis id="40B8EAFBFF82FF8A6F02FF71FBA0D547" box="[1025,1101,1340,1365]" italics="true" pageId="2" pageNumber="556">lCh/lD</emphasis>
9.2210.09 %), half as long again as corniculus (
<emphasis id="40B8EAFBFF82FF8A6A7FFF12FE38D56A" box="[380,469,1375,1400]" italics="true" pageId="2" pageNumber="556">lCh/lCo</emphasis>
1.441.64). Fixed digit with a row of three teeth in paraxial position, and pilus dentilis and one tooth in antaxial position. Movable digit bidentate. Palp length 148162; internal seta of trochanter (30 35) one third longer than external seta (2226); palpi with typically specialised setae on femur (
<emphasis id="40B8EAFBFF82FF8A6FC7FFEAFB37D5D2" box="[1220,1242,1447,1472]" italics="true" pageId="2" pageNumber="556">al</emphasis>
2021) and genu (
<emphasis id="40B8EAFBFF82FF8A6BA3FF81FF5BD5F7" box="[160,182,1484,1509]" italics="true" pageId="2" pageNumber="556">al</emphasis>
<date id="06721029FF82FF8A6BB5FF81FEFED5F7" box="[182,275,1484,1509]" pageId="2" pageNumber="556" value="2015-01-16">1 1516</date>
,
<emphasis id="40B8EAFBFF82FF8A6A1CFF81FED8D5F7" box="[287,309,1484,1509]" italics="true" pageId="2" pageNumber="556">al</emphasis>
<date id="06721029FF82FF8A6A37FF81FE7CD5F6" box="[308,401,1484,1508]" pageId="2" pageNumber="556" value="1920-02-21">2 2021</date>
) strong, cultriform.
</paragraph>
<paragraph id="727336E9FF82FF8A6BC4FFBCFAA6D762" blockId="2.[151,1437,151,1941]" pageId="2" pageNumber="556">
<emphasis id="40B8EAFBFF82FF8A6BC4FFBCFF11D61A" box="[199,252,1521,1544]" italics="true" pageId="2" pageNumber="556">Legs</emphasis>
(
<figureCitation id="EAF72A6CFF82FF8A6A09FFA2FEA3D61A" box="[266,334,1519,1544]" captionStart="FIGURES 1 5" captionStartId="3.[151,264,1930,1953]" captionTargetBox="[179,1427,199,1902]" captionTargetId="figure@3.[151,1436,195,1907]" captionTargetPageId="3" captionText="FIGURES 1 5. Arctoseius ambiguus sp. nov., female. 1. tritosternum; 2. ambulacrum and tarsus II distally, ventral aspect; 3. gnathotectum; 4. peritrematal region; 5. idiosomal dorsum. Scales: 1 2 25 µm, 3 5 50 µm." httpUri="https://zenodo.org/record/580967/files/figure.png" pageId="2" pageNumber="556">Fig. 2</figureCitation>
). Legs relatively short (I 408464, II 312364, III 278330, IV 364404); leg I clearly shorter than dorsal shield,
<emphasis id="40B8EAFBFF82FF8A6A3AFC59FE60D63F" box="[313,397,1556,1581]" italics="true" pageId="2" pageNumber="556">lExI/lD</emphasis>
0.850.92. Length of tarsi I 108128,
<date id="06721029FF82FF8A6850FC58FC61D63E" box="[851,908,1556,1580]" pageId="2" pageNumber="556" value="1988-02">II 88</date>
102,
<date id="06721029FF82FF8A68D6FC58FBFAD63E" box="[981,1047,1556,1580]" pageId="2" pageNumber="556" value="1980-03">III 80</date>
92, IV 112127; tarsi shortened:
<emphasis id="40B8EAFBFF82FF8A6B94FC7AFF43D642" box="[151,174,1591,1616]" italics="true" pageId="2" pageNumber="556">lT</emphasis>
IV
<emphasis id="40B8EAFBFF82FF8A6BCFFC7AFF02D642" box="[204,239,1591,1616]" italics="true" pageId="2" pageNumber="556">/lD</emphasis>
0.230.25,
<emphasis id="40B8EAFBFF82FF8A6A74FC7AFE63D642" box="[375,398,1591,1616]" italics="true" pageId="2" pageNumber="556">lT</emphasis>
III
<emphasis id="40B8EAFBFF82FF8A6AA9FC75FE39D65D" box="[426,468,1592,1615]" italics="true" pageId="2" pageNumber="556">/wT</emphasis>
III 2.943.29; tarsus I dilated distally. Legs II, IV slightly thicker than legs I, III. Leg chaetome as described for the genus (
<bibRefCitation id="165D4B18FF82FF8A694AFC11FC9DD667" author="Lindquist" box="[585,880,1628,1653]" pageId="2" pageNumber="556" refString="Lindquist, E. E. &amp; Evans, G. O. (1965) Taxonomic concepts in the Ascidae, with a modified setal nomenclature for the idiosoma of the Gamasina (Acarina: Mesostigmata). Memoirs of the Entomological Society of Canada, 47, 1 - 64. https: // doi. org / 10.4039 / entm 9747 fv" type="journal article" year="1965">Lindquist &amp; Evans, 1965</bibRefCitation>
); tibia IV with seven setae,
<emphasis id="40B8EAFBFF82FF8A6FB5FC11FB26D667" box="[1206,1227,1628,1653]" italics="true" pageId="2" pageNumber="556">pl</emphasis>
present. Most leg setae simple, short or of moderate lengths; length of
<emphasis id="40B8EAFBFF82FF8A69E9FC32FCEBD68A" box="[746,774,1663,1688]" italics="true" pageId="2" pageNumber="556">pd</emphasis>
2 on genu II (1621) clearly shorter than width of segment (4448). Tarsi IIIV each with some rather long setae (up to 64 on tarsus IV), many of them with piliform tips; with subapical setae
<emphasis id="40B8EAFBFF82FF8A6A4BFC8AFEB3D6F2" box="[328,350,1735,1760]" italics="true" pageId="2" pageNumber="556">al</emphasis>
1,
<emphasis id="40B8EAFBFF82FF8A6A78FC8AFE7CD6F2" box="[379,401,1735,1760]" italics="true" pageId="2" pageNumber="556">pl</emphasis>
1 (1417) and
<emphasis id="40B8EAFBFF82FF8A6934FC84FDBFD6F2" box="[567,594,1737,1760]" italics="true" pageId="2" pageNumber="556">av</emphasis>
1,
<emphasis id="40B8EAFBFF82FF8A696CFC84FD67D6F2" box="[623,650,1737,1760]" italics="true" pageId="2" pageNumber="556">pv</emphasis>
1 (1316) about equally distant from tarsus apex (
<figureCitation id="EAF72A6CFF82FF8A6FBCFC8AFAE8D6F2" box="[1215,1285,1735,1760]" captionStart="FIGURES 1 5" captionStartId="3.[151,264,1930,1953]" captionTargetBox="[179,1427,199,1902]" captionTargetId="figure@3.[151,1436,195,1907]" captionTargetPageId="3" captionText="FIGURES 1 5. Arctoseius ambiguus sp. nov., female. 1. tritosternum; 2. ambulacrum and tarsus II distally, ventral aspect; 3. gnathotectum; 4. peritrematal region; 5. idiosomal dorsum. Scales: 1 2 25 µm, 3 5 50 µm." httpUri="https://zenodo.org/record/580967/files/figure.png" pageId="2" pageNumber="556">Fig. 2</figureCitation>
),
<emphasis id="40B8EAFBFF82FF8A6E18FC84FADBD6F2" box="[1307,1334,1737,1760]" italics="true" pageId="2" pageNumber="556">av</emphasis>
1and
<emphasis id="40B8EAFBFF82FF8A6E77FC84FA63D6F2" box="[1396,1422,1737,1760]" italics="true" pageId="2" pageNumber="556">pv</emphasis>
1 noticeably thicker. Ambulacrum I on pedicellate base, claws I (12) smaller than claws IIIV (1518). Tarsus I distally with seven rod-like solenidia (five apical, two subapical); length of sensillum with lanceolate apex 3032. Ambulacra of legs IIIV (length 3238) with rather long paradactyli (2224) extending clearly beyond apices of claws. Tarsi IIIV with apical setae
<emphasis id="40B8EAFBFF82FF8A6929FD1AFDABD762" box="[554,582,1879,1904]" italics="true" pageId="2" pageNumber="556">ad</emphasis>
1 and
<emphasis id="40B8EAFBFF82FF8A6989FD1AFD4BD762" box="[650,678,1879,1904]" italics="true" pageId="2" pageNumber="556">pd</emphasis>
1 threadlike, approximately as long (2630) as ambulacrum.
</paragraph>
<paragraph id="727336E9FF82FF8A6BC4FD31FE61D787" blockId="2.[151,1437,151,1941]" box="[199,396,1916,1941]" pageId="2" pageNumber="556">
<emphasis id="40B8EAFBFF82FF8A6BC4FD31FEE6D787" bold="true" box="[199,267,1916,1941]" pageId="2" pageNumber="556">Male.</emphasis>
Not found.
</paragraph>
</subSubSection>
<caption id="26B36661FF83FF8B6B94FDC7FBC3D7D2" httpUri="https://zenodo.org/record/580967/files/figure.png" pageId="3" pageNumber="557" targetBox="[179,1427,199,1902]" targetPageId="3">
<paragraph id="727336E9FF83FF8B6B94FDC7FBC3D7D2" blockId="3.[151,1435,1930,1984]" pageId="3" pageNumber="557">
<emphasis id="40B8EAFBFF83FF8B6B94FDC7FED6D7B2" bold="true" box="[151,315,1930,1953]" pageId="3" pageNumber="557">FIGURES 15.</emphasis>
<taxonomicName id="B5CC4D6AFF83FF8B6A41FDC6FDFBD7B2" box="[322,534,1931,1952]" class="Arachnida" family="Ascidae" genus="Arctoseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="557" phylum="Arthropoda" rank="species" species="ambiguus" status="sp. nov.">
<emphasis id="40B8EAFBFF83FF8B6A41FDC6FDFBD7B2" box="[322,534,1931,1952]" italics="true" pageId="3" pageNumber="557">Arctoseius ambiguus</emphasis>
</taxonomicName>
<emphasis id="40B8EAFBFF83FF8B691FFDC6FD83D7B2" bold="true" box="[540,622,1931,1952]" pageId="3" pageNumber="557">
<taxonomicNameLabel id="5B8B5780FF83FF8B691FFDC6FD83D7B2" box="[540,622,1931,1952]" pageId="3" pageNumber="557" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female. 1. tritosternum; 2. ambulacrum and tarsus II distally, ventral aspect; 3. gnathotectum; 4. peritrematal region; 5. idiosomal dorsum. Scales: 12 25 µm, 35 50 µm.
</paragraph>
</caption>
<caption id="26B36661FF84FF8C6B94FDC8FE70D7CB" httpUri="https://zenodo.org/record/580968/files/figure.png" pageId="4" pageNumber="558" targetBox="[162,1430,195,1884]" targetPageId="4">
<paragraph id="727336E9FF84FF8C6B94FDC8FE70D7CB" blockId="4.[151,1437,1925,2009]" pageId="4" pageNumber="558">
<emphasis id="40B8EAFBFF84FF8C6B94FDC8FEA6D789" bold="true" box="[151,331,1925,1948]" pageId="4" pageNumber="558">FIGURES 611.</emphasis>
<taxonomicName id="B5CC4D6AFF84FF8C6A54FDCBFDDDD789" box="[343,560,1926,1947]" class="Arachnida" family="Ascidae" genus="Arctoseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="558" phylum="Arthropoda" rank="species" species="ambiguus" status="sp. nov.">
<emphasis id="40B8EAFBFF84FF8C6A54FDCBFDDDD789" box="[343,560,1926,1947]" italics="true" pageId="4" pageNumber="558">Arctoseius ambiguus</emphasis>
</taxonomicName>
<emphasis id="40B8EAFBFF84FF8C693FFDCBFD7FD789" bold="true" box="[572,658,1926,1947]" pageId="4" pageNumber="558">
<taxonomicNameLabel id="5B8B5780FF84FF8C693FFDCBFD7FD789" box="[572,658,1926,1947]" pageId="4" pageNumber="558" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female. 6. acetabula IIIV and exopodal platelets; 7. subcapitulum; 8. idiosomal ventrum; 9. anal shield; 10. cheliceral digits, dorsal aspect; 11. chelicera without basal joint, paraxial aspect. Scales: 69 50 µm, 1011 25 µm.
</paragraph>
</caption>
<subSubSection id="3AD66562FF85FF8D6BC4FADAFB6BD2BA" pageId="5" pageNumber="559" type="discussion">
<paragraph id="727336E9FF85FF8D6BC4FADAFA78D22E" blockId="5.[151,1436,151,968]" pageId="5" pageNumber="559">
<emphasis id="40B8EAFBFF85FF8D6BC4FADAFED6D0A2" bold="true" box="[199,315,151,176]" pageId="5" pageNumber="559">Remarks.</emphasis>
Females of
<taxonomicName id="B5CC4D6AFF85FF8D6AC8FAD5FDB4D0A2" box="[459,601,151,176]" class="Arachnida" family="Ascidae" genus="Arctoseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="559" phylum="Arthropoda" rank="species" species="ambiguus">
<emphasis id="40B8EAFBFF85FF8D6AC8FAD5FE31D0BD" box="[459,476,152,175]" italics="true" pageId="5" pageNumber="559">A</emphasis>
.
<emphasis id="40B8EAFBFF85FF8D6AEFFADAFDB4D0A2" box="[492,601,151,176]" italics="true" pageId="5" pageNumber="559">ambiguus</emphasis>
</taxonomicName>
are similar to those of
<taxonomicName id="B5CC4D6AFF85FF8D6869FAD5FB50D0A2" authority="Berlese, 1916" authorityName="Berlese" authorityYear="1916" box="[874,1213,151,176]" class="Arachnida" family="Ascidae" genus="Arctoseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="559" phylum="Arthropoda" rank="species" species="venustulus">
<emphasis id="40B8EAFBFF85FF8D6869FAD5FC96D0BD" box="[874,891,152,175]" italics="true" pageId="5" pageNumber="559">A</emphasis>
.
<emphasis id="40B8EAFBFF85FF8D6888FADAFBECD0A2" box="[907,1025,151,176]" italics="true" pageId="5" pageNumber="559">venustulus</emphasis>
(
<bibRefCitation id="165D4B18FF85FF8D6F11FADAFB59D0A2" author="Berlese" box="[1042,1204,151,176]" pageId="5" pageNumber="559" refString="Berlese, A. (1916) Centuria prima di Acari nuovi. Redia, 12, 19 - 67." type="journal article" year="1916">Berlese, 1916</bibRefCitation>
)
</taxonomicName>
. They both possess fully reticulated sternal and dorsal shields (in the former species, reticulation is more pronounced laterally) and relatively large, posteriorly dilated anal shield. However, the idiosoma in
<taxonomicName id="B5CC4D6AFF85FF8D68F9FAADFB67D0EA" box="[1018,1162,223,248]" class="Arachnida" family="Ascidae" genus="Arctoseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="559" phylum="Arthropoda" rank="species" species="ambiguus">
<emphasis id="40B8EAFBFF85FF8D68F9FAADFBE6D0E5" box="[1018,1035,224,247]" italics="true" pageId="5" pageNumber="559">A</emphasis>
.
<emphasis id="40B8EAFBFF85FF8D6F1EFA92FB67D0EA" box="[1053,1162,223,248]" italics="true" pageId="5" pageNumber="559">ambiguus</emphasis>
</taxonomicName>
is larger (dorsal shield length 468512 instead of 365445) and wider (
<emphasis id="40B8EAFBFF85FF8D69B7FB49FD12D10F" box="[692,767,260,285]" italics="true" pageId="5" pageNumber="559">lD/wD</emphasis>
1.721.82 instead of 1.952.12). Dorsal setae in the female of the new species are in general longer and more heterogeneous, setae
<emphasis id="40B8EAFBFF85FF8D68BFFB64FC29D152" box="[956,964,297,320]" italics="true" pageId="5" pageNumber="559">j</emphasis>
2-6 [2028] usually longer than
<emphasis id="40B8EAFBFF85FF8D6E2CFB64FAD6D152" box="[1327,1339,297,320]" italics="true" pageId="5" pageNumber="559">J</emphasis>
1-4 [12 21] (weakly distinguished in
<taxonomicName id="B5CC4D6AFF85FF8D6AE3FB01FD98D177" box="[480,629,332,357]" class="Arachnida" family="Ascidae" genus="Arctoseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="559" phylum="Arthropoda" rank="species" species="venustulus">
<emphasis id="40B8EAFBFF85FF8D6AE3FB01FE1CD171" box="[480,497,332,355]" italics="true" pageId="5" pageNumber="559">A</emphasis>
.
<emphasis id="40B8EAFBFF85FF8D6AFCFB01FD98D177" box="[511,629,332,357]" italics="true" pageId="5" pageNumber="559">venustulus</emphasis>
</taxonomicName>
,
<emphasis id="40B8EAFBFF85FF8D6987FB00FD61D176" box="[644,652,333,356]" italics="true" pageId="5" pageNumber="559">j</emphasis>
5=
<emphasis id="40B8EAFBFF85FF8D69A9FB00FD5BD176" box="[682,694,333,356]" italics="true" pageId="5" pageNumber="559">J</emphasis>
1=10 µm), seta
<emphasis id="40B8EAFBFF85FF8D6864FB00FC9BD176" box="[871,886,333,356]" italics="true" pageId="5" pageNumber="559">Z</emphasis>
5 are significantly longer than other opisthonotal setae, particularly in the series
<emphasis id="40B8EAFBFF85FF8D6AF8FB3CFDEAD19A" box="[507,519,369,392]" italics="true" pageId="5" pageNumber="559">J</emphasis>
(length ratio
<emphasis id="40B8EAFBFF85FF8D69A7FB3CFD5ED19A" box="[676,691,369,392]" italics="true" pageId="5" pageNumber="559">Z</emphasis>
5/
<emphasis id="40B8EAFBFF85FF8D69CAFB3CFD38D19A" box="[713,725,369,392]" italics="true" pageId="5" pageNumber="559">J</emphasis>
1 1.92.4 instead of
<quantity id="B5349B0CFF85FF8D68C1FB3DFBEED19A" box="[962,1027,367,392]" metricMagnitude="-2" metricUnit="m" metricValue="3.302" pageId="5" pageNumber="559" unit="in" value="1.3">1.3 in</quantity>
<taxonomicName id="B5CC4D6AFF85FF8D6F0FFB3DFB4ED19A" box="[1036,1187,367,392]" class="Arachnida" family="Ascidae" genus="Arctoseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="559" phylum="Arthropoda" rank="species" species="venustulus">
<emphasis id="40B8EAFBFF85FF8D6F0FFB3DFBF0D195" box="[1036,1053,368,391]" italics="true" pageId="5" pageNumber="559">A</emphasis>
.
<emphasis id="40B8EAFBFF85FF8D6F2EFB22FB4ED19A" box="[1069,1187,367,392]" italics="true" pageId="5" pageNumber="559">venustulus</emphasis>
</taxonomicName>
). In
<taxonomicName id="B5CC4D6AFF85FF8D6FDFFB3DFA87D19A" box="[1244,1386,367,392]" class="Arachnida" family="Ascidae" genus="Arctoseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="559" phylum="Arthropoda" rank="species" species="ambiguus">
<emphasis id="40B8EAFBFF85FF8D6FDFFB3DFB00D195" box="[1244,1261,368,391]" italics="true" pageId="5" pageNumber="559">A</emphasis>
.
<emphasis id="40B8EAFBFF85FF8D6FFEFB22FA87D19A" box="[1277,1386,367,392]" italics="true" pageId="5" pageNumber="559">ambiguus</emphasis>
</taxonomicName>
, the lateral margins of the anal shield often have broad ledges as in
<figureCitation id="EAF72A6CFF85FF8D6877FBD9FC55D1BE" box="[884,952,404,429]" captionStart="FIGURES 6 11" captionStartId="4.[151,264,1925,1948]" captionTargetBox="[162,1430,195,1884]" captionTargetId="figure@4.[151,1436,193,1904]" captionTargetPageId="4" captionText="FIGURES 6 11. Arctoseius ambiguus sp. nov., female. 6. acetabula II IV and exopodal platelets; 7. subcapitulum; 8. idiosomal ventrum; 9. anal shield; 10. cheliceral digits, dorsal aspect; 11. chelicera without basal joint, paraxial aspect. Scales: 6 9 50 µm, 10 11 25 µm." httpUri="https://zenodo.org/record/580968/files/figure.png" pageId="5" pageNumber="559">Fig. 9</figureCitation>
(usually almost smooth in
<taxonomicName id="B5CC4D6AFF85FF8D6FF0FBD9FA67D1BF" box="[1267,1418,404,429]" class="Arachnida" family="Ascidae" genus="Arctoseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="559" phylum="Arthropoda" rank="species" species="venustulus">
<emphasis id="40B8EAFBFF85FF8D6FF0FBD9FAE9D1B9" box="[1267,1284,404,427]" italics="true" pageId="5" pageNumber="559">A</emphasis>
.
<emphasis id="40B8EAFBFF85FF8D6E17FBD9FA67D1BF" box="[1300,1418,404,429]" italics="true" pageId="5" pageNumber="559">venustulus</emphasis>
</taxonomicName>
); posterior and lateral parts of the shield are coarsely punctate (alternatively reticulate to different degree and finepunctate). The gnathotectum of
<taxonomicName id="B5CC4D6AFF85FF8D690EFB91FD73D1E7" box="[525,670,476,501]" class="Arachnida" family="Ascidae" genus="Arctoseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="559" phylum="Arthropoda" rank="species" species="ambiguus">
<emphasis id="40B8EAFBFF85FF8D690EFB91FDF3D1E1" box="[525,542,476,499]" italics="true" pageId="5" pageNumber="559">A</emphasis>
.
<emphasis id="40B8EAFBFF85FF8D6932FB91FD73D1E7" box="[561,670,476,501]" italics="true" pageId="5" pageNumber="559">ambiguus</emphasis>
</taxonomicName>
is wider (width/length ratio 1.82.6 instead 1.11.7). The sternal shield in
<taxonomicName id="B5CC4D6AFF85FF8D6BFCF84DFE61D20A" box="[255,396,511,536]" class="Arachnida" family="Ascidae" genus="Arctoseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="559" phylum="Arthropoda" rank="species" species="ambiguus">
<emphasis id="40B8EAFBFF85FF8D6BFCF84DFEFDD205" box="[255,272,512,535]" italics="true" pageId="5" pageNumber="559">A</emphasis>
.
<emphasis id="40B8EAFBFF85FF8D6A1CFBB2FE61D20A" box="[287,396,511,536]" italics="true" pageId="5" pageNumber="559">ambiguus</emphasis>
</taxonomicName>
is clearly reticulate only in the lateral parts (totally and evenly so in
<taxonomicName id="B5CC4D6AFF85FF8D6F98F84DFADCD20A" box="[1179,1329,511,536]" class="Arachnida" family="Ascidae" genus="Arctoseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="559" phylum="Arthropoda" rank="species" species="venustulus">
<emphasis id="40B8EAFBFF85FF8D6F98F84DFB41D205" box="[1179,1196,512,535]" italics="true" pageId="5" pageNumber="559">A</emphasis>
.
<emphasis id="40B8EAFBFF85FF8D6FB8FBB2FADCD20A" box="[1211,1329,511,536]" italics="true" pageId="5" pageNumber="559">venustulus</emphasis>
</taxonomicName>
), and the presternal platelets are weakly sclerotized, abutting the sternal shield (well sclerotized, separated in
<taxonomicName id="B5CC4D6AFF85FF8D6FF3F869FA69D22F" box="[1264,1412,548,573]" class="Arachnida" family="Ascidae" genus="Arctoseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="559" phylum="Arthropoda" rank="species" species="venustulus">
<emphasis id="40B8EAFBFF85FF8D6FF3F869FAECD229" box="[1264,1281,548,571]" italics="true" pageId="5" pageNumber="559">A</emphasis>
.
<emphasis id="40B8EAFBFF85FF8D6E0DF869FA69D22F" box="[1294,1412,548,573]" italics="true" pageId="5" pageNumber="559">venustulus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="727336E9FF85FF8D6BC4F80AFB6BD2BA" blockId="5.[151,1436,151,968]" pageId="5" pageNumber="559">
<emphasis id="40B8EAFBFF85FF8D6BC4F80AFEA8D272" bold="true" box="[199,325,583,608]" pageId="5" pageNumber="559">Variability</emphasis>
. The morphological variation in
<taxonomicName id="B5CC4D6AFF85FF8D69D2F805FC8FD272" box="[721,866,583,608]" class="Arachnida" family="Ascidae" genus="Arctoseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="559" phylum="Arthropoda" rank="species" species="ambiquus">
<emphasis id="40B8EAFBFF85FF8D69D2F805FD0FD24D" box="[721,738,584,607]" italics="true" pageId="5" pageNumber="559">A</emphasis>
.
<emphasis id="40B8EAFBFF85FF8D69F6F80AFC8FD272" box="[757,866,583,608]" italics="true" pageId="5" pageNumber="559">ambiquus</emphasis>
</taxonomicName>
is minimal and concerns mainly the number of opisthogastral and posterior marginal setae of the
<emphasis id="40B8EAFBFF85FF8D69D9F820FD06D296" box="[730,747,621,644]" italics="true" pageId="5" pageNumber="559">R</emphasis>
series (
<figureCitation id="EAF72A6CFF85FF8D6844F821FC60D296" box="[839,909,620,645]" captionStart="FIGURES 6 11" captionStartId="4.[151,264,1925,1948]" captionTargetBox="[162,1430,195,1884]" captionTargetId="figure@4.[151,1436,193,1904]" captionTargetPageId="4" captionText="FIGURES 6 11. Arctoseius ambiguus sp. nov., female. 6. acetabula II IV and exopodal platelets; 7. subcapitulum; 8. idiosomal ventrum; 9. anal shield; 10. cheliceral digits, dorsal aspect; 11. chelicera without basal joint, paraxial aspect. Scales: 6 9 50 µm, 10 11 25 µm." httpUri="https://zenodo.org/record/580968/files/figure.png" pageId="5" pageNumber="559">Fig. 8</figureCitation>
), the position of seta
<emphasis id="40B8EAFBFF85FF8D6F8DF820FB74D296" box="[1166,1177,621,644]" italics="true" pageId="5" pageNumber="559">s</emphasis>
2 (rarely on the dorsal shield margin) and the development of lateral protuberances on the anal shield (
<figureCitation id="EAF72A6CFF85FF8D6F10F8C2FB95D2BA" box="[1043,1144,655,680]" captionStart="FIGURES 6 11" captionStartId="4.[151,264,1925,1948]" captionTargetBox="[162,1430,195,1884]" captionTargetId="figure@4.[151,1436,193,1904]" captionTargetPageId="4" captionText="FIGURES 6 11. Arctoseius ambiguus sp. nov., female. 6. acetabula II IV and exopodal platelets; 7. subcapitulum; 8. idiosomal ventrum; 9. anal shield; 10. cheliceral digits, dorsal aspect; 11. chelicera without basal joint, paraxial aspect. Scales: 6 9 50 µm, 10 11 25 µm." httpUri="https://zenodo.org/record/580968/files/figure.png" pageId="5" pageNumber="559">Figs 8, 9</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3AD66562FF85FF8D6BC4F8F9FDB9D2E2" pageId="5" pageNumber="559" type="etymology">
<paragraph id="727336E9FF85FF8D6BC4F8F9FDB9D2E2" blockId="5.[151,1436,151,968]" pageId="5" pageNumber="559">
<emphasis id="40B8EAFBFF85FF8D6BC4F8F9FEA3D2DF" bold="true" box="[199,334,692,717]" pageId="5" pageNumber="559">Etymology.</emphasis>
The specific name
<taxonomicName id="B5CC4D6AFF85FF8D6957F8F9FD2CD2DF" box="[596,705,692,717]" class="Arachnida" family="Ascidae" genus="Arctoseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="559" phylum="Arthropoda" rank="species" species="ambiguus">
<emphasis id="40B8EAFBFF85FF8D6957F8F9FD2CD2DF" box="[596,705,692,717]" italics="true" pageId="5" pageNumber="559">ambiguus</emphasis>
</taxonomicName>
highlights the absence of pronounced, easily recognisable differential features in this new species.
</paragraph>
</subSubSection>
<subSubSection id="3AD66562FF85FF8D6BC4F8B1FE4CD3DA" pageId="5" pageNumber="559" type="distribution">
<paragraph id="727336E9FF85FF8D6BC4F8B1FE4CD3DA" blockId="5.[151,1436,151,968]" pageId="5" pageNumber="559">
<emphasis id="40B8EAFBFF85FF8D6BC4F8B1FE17D307" bold="true" box="[199,506,764,789]" pageId="5" pageNumber="559">Distribution and ecology.</emphasis>
At present the new species is known only from
<collectingCountry id="0ADB7679FF85FF8D6F3CF8B1FB79D307" box="[1087,1172,764,789]" name="Finland" pageId="5" pageNumber="559">Finland</collectingCountry>
and
<collectingRegion id="B008F80BFF85FF8D6FD0F8B1FAE6D307" box="[1235,1291,764,789]" country="Russia" name="Altay" pageId="5" pageNumber="559">Altai</collectingRegion>
Mts., South Siberia. Such distant records possibly reflect either an insufficient level of our knowledge, or peculiarities of
<taxonomicName id="B5CC4D6AFF85FF8D6B94F909FEE1D349" box="[151,268,836,859]" class="Arachnida" family="Ascidae" genus="Arctoseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="559" phylum="Arthropoda" rank="genus">
<emphasis id="40B8EAFBFF85FF8D6B94F909FEE1D349" box="[151,268,836,859]" italics="true" pageId="5" pageNumber="559">Arctoseius</emphasis>
</taxonomicName>
bionomy. All specimens were collected from soil and litter in forests of different humidity. Forest soil is principally a stable environment, but may include unstable and scattered microhabitats. Thus the inclination of the genus to eutrophic and temporary substrata as well as their phoretic associations with dipterans (see
<bibRefCitation id="165D4B18FF85FF8D6E29F9C1FF3DD3DA" author="Makarova" pageId="5" pageNumber="559" refString="Makarova, O. L. (1999) Mesostigmatic mites (Parasitiformes, Mesostigmata) of polar deserts in Severnaya Zemlya. Entomological Review, 79, 982 - 990." type="journal article" year="1999">Makarova 1999</bibRefCitation>
) may be involved.
</paragraph>
</subSubSection>
</treatment>
</document>