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<document id="02207F79E0A658748AC4B05F499D46D3" ID-CLB-Dataset="29980" ID-DOI="10.11646/zootaxa.4425.3.13" ID-GBIF-Dataset="6d1ae1e9-fd43-4421-9770-7062f3525dd7" ID-ISSN="1175-5326" ID-Zenodo-Dep="1270819" ID-ZooBank="C76B9ACC-9E4C-4AE4-814D-D140A9224753" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1528146147922" checkinUser="plazi" docAuthor="Chen, Zhi-Teng" docDate="2018" docId="FA6587D1FFFA6F71FF28BC82FCAEF802" docLanguage="en" docName="zootaxa.4425.3.13.pdf.imf" docOrigin="Zootaxa 4425 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pinguisoperla yangzhouensis Chen 2018" docType="treatment" docVersion="7" lastPageNumber="598" masterDocId="065CFFA9FFFB6F73FFBFBA34FFFFFFF0" masterDocTitle="Pinguisoperla, a new fossil genus of Perlidae (Insecta: Plecoptera) from mid-Cretaceous Burmese amber" masterLastPageNumber="600" masterPageNumber="596" pageNumber="597" updateTime="1698546672822" updateUser="ExternalLinkService">
<mods:mods id="9474B3051A8A356AF675BC5543E1C408" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C41BCC7133A016410824C8C961700EA2">
<mods:title id="044A40F2E0C2952BD7EF82DA8087B6D4">Pinguisoperla, a new fossil genus of Perlidae (Insecta: Plecoptera) from mid-Cretaceous Burmese amber</mods:title>
</mods:titleInfo>
<mods:name id="9BF5140D59027055BFC7AAC19AC156F5" type="personal">
<mods:role id="F714AB8783104AFC781636E523B6B095">
<mods:roleTerm id="03194E841A74DA28FBFC4A6AEA1FE237">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="49251E18B84FF71D286711DB28D26105">Chen, Zhi-Teng</mods:namePart>
</mods:name>
<mods:typeOfResource id="63FF0002A8A26607688FA68E0642B9FD">text</mods:typeOfResource>
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<mods:title id="A208ECBF584A5D39349CCCCF5DC3717F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8C0A099A99B050A997B0BEB15F8713CC">
<mods:date id="D1A4E142B319277A0DA8F0B0378CA68F">2018</mods:date>
<mods:detail id="B0D99562038F5A57FE83C43FAC4855A9" type="volume">
<mods:number id="B65DA101B0EE3B5BF6B85E2C62720076">4425</mods:number>
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<mods:detail id="1C2F69196A70B473903D59B6C3EFE5CA" type="issue">
<mods:number id="33661253B4CF33385610D28798E2A2C8">3</mods:number>
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<mods:classification id="11118C0D1D17446D82EF3210884BC8B7">journal article</mods:classification>
<mods:identifier id="89035554B3FA7879ED5E5089EC6B671E" type="CLB-Dataset">29980</mods:identifier>
<mods:identifier id="9FC7B303A9214537EC6C5076BD0B06A6" type="DOI">10.11646/zootaxa.4425.3.13</mods:identifier>
<mods:identifier id="0CB0533F46FFEF0E281653D1FDBBC760" type="GBIF-Dataset">6d1ae1e9-fd43-4421-9770-7062f3525dd7</mods:identifier>
<mods:identifier id="CBBD98FC86349FCE6EABAB7C50735465" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="42C04E700C253446458C13B1C376D409" type="Zenodo-Dep">1270819</mods:identifier>
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<treatment id="FA6587D1FFFA6F71FF28BC82FCAEF802" ID-DOI="http://doi.org/10.5281/zenodo.5957429" ID-GBIF-Taxon="144235684" ID-Zenodo-Dep="5957429" LSID="urn:lsid:plazi:treatment:FA6587D1FFFA6F71FF28BC82FCAEF802" httpUri="http://treatment.plazi.org/id/FA6587D1FFFA6F71FF28BC82FCAEF802" lastPageId="2" lastPageNumber="598" pageId="1" pageNumber="597">
<subSubSection id="3AD6654CFFFA6F72FF28BC82FEF9F902" pageId="1" pageNumber="597" type="nomenclature">
<paragraph id="727336C7FFFA6F72FF28BC82FD50F920" blockId="1.[151,687,1718,1778]" box="[151,687,1718,1744]" pageId="1" pageNumber="597">
<heading id="293B81ABFFFA6F72FF28BC82FD50F920" bold="true" box="[151,687,1718,1744]" fontSize="11" level="1" pageId="1" pageNumber="597" reason="1">
<emphasis id="40B8EAD5FFFA6F72FF28BC82FD50F920" bold="true" box="[151,687,1718,1744]" pageId="1" pageNumber="597">
<taxonomicName id="B5CC4D44FFFA6F72FF28BC82FE06F920" ID-CoL="8NGQ5" authority="Chen, 2018" authorityName="Chen" authorityYear="2018" box="[151,505,1718,1744]" class="Insecta" family="Perlidae" genus="Pinguisoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="600" phylum="Arthropoda" rank="species" species="yangzhouensis" status="sp. nov.">
<emphasis id="40B8EAD5FFFA6F72FF28BC82FE06F920" bold="true" box="[151,505,1718,1744]" italics="true" pageId="1" pageNumber="597">Pinguisoperla yangzhouensis</emphasis>
</taxonomicName>
Chen, sp. nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="727336C7FFFA6F72FF28BCEDFEF9F902" blockId="1.[151,687,1718,1778]" box="[151,262,1753,1778]" pageId="1" pageNumber="597">
<figureCitation id="EAF72A42FFFA6F72FF28BCEDFEFDF902" box="[151,258,1753,1778]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1341,1363]" captionStartId-1="3.[151,250,1208,1230]" captionStartId-2="3.[151,250,1781,1803]" captionStartId-3="4.[151,250,762,784]" captionTargetBox-0="[313,1257,598,1287]" captionTargetBox-1="[313,1273,228,1157]" captionTargetBox-2="[325,1253,1316,1723]" captionTargetId-0="figure@2.[313,1273,598,1287]" captionTargetId-1="figure@3.[313,1273,228,1157]" captionTargetId-2="figure@3.[313,1273,1295,1740]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="3" captionText-0="FIGURE 1. Pinguisoperla yangzhouensis, sp. nov. (A) Adult habitus, ventral view; (B) Reconstruction of adult habitus, dorsal view." captionText-1="FIGURE 2. Pinguisoperla yangzhouensis, sp. nov. (A) Head and pronotum, ventral view; (B) Antennae and palpi." captionText-2="FIGURE 3. Pinguisoperla yangzhouensis, sp. nov. Reconstruction of left forewing, dorsal view." captionText-3="FIGURE 4. Pinguisoperla yangzhouensis, sp. nov. Terminalia, ventral view;" httpUri-0="https://zenodo.org/record/1270821/files/figure.png" httpUri-1="https://zenodo.org/record/1270823/files/figure.png" httpUri-2="https://zenodo.org/record/1270827/files/figure.png" pageId="1" pageNumber="597">Figs. 1–4</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3AD6654CFFFA6F72FF28BD15FE27F8CA" box="[151,472,1825,1850]" pageId="1" pageNumber="597" type="diagnosis">
<paragraph id="727336C7FFFA6F72FF28BD15FE27F8CA" blockId="1.[151,1437,1825,2030]" box="[151,472,1825,1850]" pageId="1" pageNumber="597">
<emphasis id="40B8EAD5FFFA6F72FF28BD15FEEEF8CA" bold="true" box="[151,273,1825,1850]" pageId="1" pageNumber="597">Diagnosis.</emphasis>
As for the genus.
</paragraph>
</subSubSection>
<subSubSection id="3AD6654CFFFA6F71FF78BD71FF29F8E9" lastPageId="2" lastPageNumber="598" pageId="1" pageNumber="597" type="description">
<paragraph id="727336C7FFFA6F72FF78BD71FF10F83A" blockId="1.[151,1437,1825,2030]" pageId="1" pageNumber="597">
<emphasis id="40B8EAD5FFFA6F72FF78BD71FEA6F8AE" bold="true" box="[199,345,1861,1886]" pageId="1" pageNumber="597">Description.</emphasis>
Macropterous; body length (excluding antennae and cerci) ca.
<quantity id="B5349B22FFFA6F72FB8EBD72FB7BF8AD" box="[1073,1156,1862,1886]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="597" unit="mm" value="10.0">10 mm</quantity>
, generally dark (
<figureCitation id="EAF72A42FFFA6F72FAF8BD71FA75F8AE" box="[1351,1418,1861,1886]" captionStart="FIGURE 1" captionStartId="2.[151,250,1341,1363]" captionTargetBox="[313,1257,598,1287]" captionTargetId="figure@2.[313,1273,598,1287]" captionTargetPageId="2" captionText="FIGURE 1. Pinguisoperla yangzhouensis, sp. nov. (A) Adult habitus, ventral view; (B) Reconstruction of adult habitus, dorsal view." httpUri="https://zenodo.org/record/1270821/files/figure.png" pageId="1" pageNumber="597">Fig. 1</figureCitation>
). Dorsal head and thorax of the specimen are covered by cracks of the amber; apices of antennae and wings are not preserved; dorsal abdomen is covered by wings; ventral abdominal tip slightly decayed, whose detailed structure is unclear.
</paragraph>
<paragraph id="727336C7FFFA6F71FF78BDE1FAE6FEED" blockId="1.[151,1437,1825,2030]" lastBlockId="2.[151,1436,151,501]" lastPageId="2" lastPageNumber="598" pageId="1" pageNumber="597">
Head (
<figureCitation id="EAF72A42FFFA6F72FEA9BDE1FE97F81E" box="[278,360,2005,2030]" captionStart="FIGURE 1" captionStartId="2.[151,250,1341,1363]" captionTargetBox="[313,1257,598,1287]" captionTargetId="figure@2.[313,1273,598,1287]" captionTargetPageId="2" captionText="FIGURE 1. Pinguisoperla yangzhouensis, sp. nov. (A) Adult habitus, ventral view; (B) Reconstruction of adult habitus, dorsal view." httpUri="https://zenodo.org/record/1270821/files/figure.png" pageId="1" pageNumber="597">Figs. 1</figureCitation>
,
<figureCitation id="EAF72A42FFFA6F72FEC5BDE2FE74F81E" box="[378,395,2006,2030]" captionStart="FIGURE 2" captionStartId="3.[151,250,1208,1230]" captionTargetBox="[313,1273,228,1157]" captionTargetId="figure@3.[313,1273,228,1157]" captionTargetPageId="3" captionText="FIGURE 2. Pinguisoperla yangzhouensis, sp. nov. (A) Head and pronotum, ventral view; (B) Antennae and palpi." httpUri="https://zenodo.org/record/1270823/files/figure.png" pageId="1" pageNumber="597">2</figureCitation>
): Head inserted in prothorax, dark brown and sub-quadrate except for the dark, bulbous compound eyes; ocelli invisible. Antenna filiform, at least 24-segmented, each segment almost dark except for the paler apices. Maxillary palp four-segmented, two basal segments longest, third segment slightly thinner and shorter, apical segment shortest and thinnest, about half length of the third segment. Labial palp three-segmented, median segment longest, apical segment shortest and thinnest, about 2/3 length of the median segment.
</paragraph>
<paragraph id="727336C7FFF96F71FF78BB13FC3EFE05" blockId="2.[151,1436,151,501]" pageId="2" pageNumber="598">
Thorax (
<figureCitation id="EAF72A42FFF96F71FE96BB13FE87FEB0" box="[297,376,295,320]" captionStart="FIGURE 1" captionStartId="2.[151,250,1341,1363]" captionTargetBox="[313,1257,598,1287]" captionTargetId="figure@2.[313,1273,598,1287]" captionTargetPageId="2" captionText="FIGURE 1. Pinguisoperla yangzhouensis, sp. nov. (A) Adult habitus, ventral view; (B) Reconstruction of adult habitus, dorsal view." httpUri="https://zenodo.org/record/1270821/files/figure.png" pageId="2" pageNumber="598">Figs. 1</figureCitation>
,
<figureCitation id="EAF72A42FFF96F71FE37BB1CFE66FEB0" box="[392,409,296,320]" captionStart="FIGURE 2" captionStartId="3.[151,250,1208,1230]" captionTargetBox="[313,1273,228,1157]" captionTargetId="figure@3.[313,1273,228,1157]" captionTargetPageId="3" captionText="FIGURE 2. Pinguisoperla yangzhouensis, sp. nov. (A) Head and pronotum, ventral view; (B) Antennae and palpi." httpUri="https://zenodo.org/record/1270823/files/figure.png" pageId="2" pageNumber="598">2</figureCitation>
): Prothorax much narrower than head, but subequal in length; pronotum dark, trapezoidal with acute corners, anterior margin wider than posterior margin. Mesothorax and metathorax unclear, but strongly sclerotized and wider than prothorax. Legs mostly dark; forelegs shortest, hindlegs longest; femur subequal in length with tibia in forelegs and midlegs, but shorter than tibia in hindlegs; tibia ventrally with two subapical spurs; tarsus three-segmented, two basal segments shortest, subequal in size and with conspicuous euplantulae, apical segment slender and with long apical bristles; claws simple and pointed.
</paragraph>
<caption id="26B3664FFFF96F71FF28BF09FF31FA82" httpUri="https://zenodo.org/record/1270821/files/figure.png" pageId="2" pageNumber="598" startId="2.[151,250,1341,1363]" targetBox="[313,1257,598,1287]" targetPageId="2">
<paragraph id="727336C7FFF96F71FF28BF09FF31FA82" blockId="2.[151,1436,1341,1394]" pageId="2" pageNumber="598">
<emphasis id="40B8EAD5FFF96F71FF28BF09FEEDFAA3" bold="true" box="[151,274,1341,1363]" pageId="2" pageNumber="598">FIGURE 1.</emphasis>
<taxonomicName id="B5CC4D44FFF96F71FEA7BF09FDBEFAA2" authority="Chen, 2018" authorityName="Chen" authorityYear="2018" box="[280,577,1341,1362]" class="Insecta" family="Perlidae" genus="Pinguisoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="598" phylum="Arthropoda" rank="species" species="yangzhouensis" status="sp. nov.">
<emphasis id="40B8EAD5FFF96F71FEA7BF09FDBEFAA2" box="[280,577,1341,1362]" italics="true" pageId="2" pageNumber="598">Pinguisoperla yangzhouensis</emphasis>
</taxonomicName>
,
<emphasis id="40B8EAD5FFF96F71FDF1BF09FD61FAA3" bold="true" box="[590,670,1341,1363]" pageId="2" pageNumber="598">
<taxonomicNameLabel id="5B8B57AEFFF96F71FDF1BF09FD61FAA3" box="[590,670,1341,1363]" pageId="2" pageNumber="598" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) Adult habitus, ventral view; (B) Reconstruction of adult habitus, dorsal view.
</paragraph>
</caption>
<paragraph id="727336C7FFF96F71FF78BFADFEB6F9B2" blockId="2.[151,1437,1433,2034]" pageId="2" pageNumber="598">
Wings (
<figureCitation id="EAF72A42FFF96F71FE9BBFADFE89FA41" box="[292,374,1433,1458]" captionStart="FIGURE 1" captionStartId="2.[151,250,1341,1363]" captionTargetBox="[313,1257,598,1287]" captionTargetId="figure@2.[313,1273,598,1287]" captionTargetPageId="2" captionText="FIGURE 1. Pinguisoperla yangzhouensis, sp. nov. (A) Adult habitus, ventral view; (B) Reconstruction of adult habitus, dorsal view." httpUri="https://zenodo.org/record/1270821/files/figure.png" pageId="2" pageNumber="598">Figs. 1</figureCitation>
,
<figureCitation id="EAF72A42FFF96F71FE35BFADFE65FA41" box="[394,410,1433,1457]" captionStart="FIGURE 3" captionStartId="3.[151,250,1781,1803]" captionTargetBox="[325,1253,1316,1723]" captionTargetId="figure@3.[313,1273,1295,1740]" captionTargetPageId="3" captionText="FIGURE 3. Pinguisoperla yangzhouensis, sp. nov. Reconstruction of left forewing, dorsal view." httpUri="https://zenodo.org/record/1270827/files/figure.png" pageId="2" pageNumber="598">3</figureCitation>
): Apex of left forewing lost, estimated length ca.
<quantity id="B5349B22FFF96F71FC48BFADFBB1FA41" box="[1015,1102,1433,1457]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="598" unit="mm" value="11.0">11 mm</quantity>
. ScP not clear; at least five crossveins present between ScP and the anterior margin. RP originated from basal 1/3 of RA and with three branches. M simple, not forked. Cu forked to CuA and CuP, CuA with four branches; CuP simple; area between M and CuA with six crossveins; area between CuA and CuP with six crossveins. AA1 simple, AA2 forked. Hind wings invisible.
</paragraph>
<paragraph id="727336C7FFF96F71FF78BC79FF29F8E9" blockId="2.[151,1437,1433,2034]" pageId="2" pageNumber="598">
Abdomen (
<figureCitation id="EAF72A42FFF96F71FEFABC79FE6DF995" box="[325,402,1613,1638]" captionStart="FIGURE 1" captionStartId="2.[151,250,1341,1363]" captionTargetBox="[313,1257,598,1287]" captionTargetId="figure@2.[313,1273,598,1287]" captionTargetPageId="2" captionText="FIGURE 1. Pinguisoperla yangzhouensis, sp. nov. (A) Adult habitus, ventral view; (B) Reconstruction of adult habitus, dorsal view." httpUri="https://zenodo.org/record/1270821/files/figure.png" pageId="2" pageNumber="598">Figs. 1</figureCitation>
,
<figureCitation id="EAF72A42FFF96F71FE1FBC79FE4EF995" box="[416,433,1613,1637]" captionStart="FIGURE 4" captionStartId="4.[151,250,762,784]" captionText="FIGURE 4. Pinguisoperla yangzhouensis, sp. nov. Terminalia, ventral view;" pageId="2" pageNumber="598">4</figureCitation>
): Abdomen dark, near half the length of the body. Sternum 9 darkly sclerotized, posterior margin distinctly extended backwards, median area paler and decayed, unclear, perhaps with specific structures. Paraprocts invisible, not well developed; abdominal tip triangular in shape. Cercus hairy, with about 7 segments; first basal segment strongly sclerotized, enlarged and elongated, curved outward and backward; several subsequent segments still thick but gradually tapering to the hairy apex; each cercal segment subapically fringed with long setae.
</paragraph>
</subSubSection>
<subSubSection id="3AD6654CFFF96F71FF78BD11FBD2F876" pageId="2" pageNumber="598" type="materials_examined">
<paragraph id="727336C7FFF96F71FF78BD11FBD2F876" blockId="2.[151,1437,1433,2034]" pageId="2" pageNumber="598">
<materialsCitation id="C2A43C9AFFF96F71FF78BD11FBD5F876" ID-GBIF-Occurrence="1848255664" country="China" county="Yangzhou" latitude="26.333334" location="Hukawng Valley" longLatPrecision="1242" longitude="96.6" municipality="Chen Amber Collection" pageId="2" pageNumber="598" specimenCount="3" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="40B8EAD5FFF96F71FF78BD11FE88F8CE" bold="true" box="[199,375,1829,1854]" pageId="2" pageNumber="598">
<typeStatus id="AD778865FFF96F71FF78BD11FEFDF8CE" box="[199,258,1829,1854]" pageId="2" pageNumber="598">Type</typeStatus>
material.
</emphasis>
<typeStatus id="AD778865FFF96F71FE3FBD11FE17F8CE" box="[384,488,1829,1854]" pageId="2" pageNumber="598" type="holotype">Holotype</typeStatus>
male, an almost complete specimen in a piece of mid-Cretaceous Burmese amber from
<collectingRegion id="B008F825FFF96F71FF66BD7DFED2F892" box="[217,301,1865,1890]" country="Myanmar" name="Kachin" pageId="2" pageNumber="598">Kachin</collectingRegion>
,
<location id="7713601CFFF96F71FE83BD7DFE03F892" LSID="urn:lsid:plazi:treatment:FA6587D1FFFA6F71FF28BC82FCAEF802:7713601CFFF96F71FE83BD7DFE03F892" box="[316,508,1865,1890]" country="China" county="Yangzhou" latitude="26.333334" longLatPrecision="1242" longitude="96.6" municipality="Chen Amber Collection" name="Hukawng Valley" pageId="2" pageNumber="598" stateProvince="Kachin">Hukawng Valley</location>
(
<geoCoordinate id="17F85000FFF96F71FDAFBD7DFD8DF892" box="[528,626,1865,1890]" direction="north" orientation="latitude" pageId="2" pageNumber="598" precision="925" value="26.333334">26°20´N</geoCoordinate>
,
<geoCoordinate id="17F85000FFF96F71FD3DBD7DFD21F892" box="[642,734,1865,1890]" direction="east" orientation="longitude" pageId="2" pageNumber="598" precision="925" value="96.6">96°36´E</geoCoordinate>
) of
<location id="7713601CFFF96F71FCAABD7DFC13F892" LSID="urn:lsid:plazi:treatment:FA6587D1FFFA6F71FF28BC82FCAEF802:7713601CFFF96F71FCAABD7DFC13F892" box="[789,1004,1865,1890]" country="China" county="Yangzhou" latitude="26.333334" longLatPrecision="1242" longitude="96.6" municipality="Chen Amber Collection" name="northern Myanmar" pageId="2" pageNumber="598" stateProvince="Kachin">northern Myanmar</location>
(98.79 ± 0.62 Ma). The
<typeStatus id="AD778865FFF96F71FAA3BD7DFA81F892" box="[1308,1406,1865,1890]" pageId="2" pageNumber="598" type="holotype">holotype</typeStatus>
is deposited in the
<collectingMunicipality id="9217ACBDFFF96F71FEF0BD59FDA4F876" box="[335,603,1901,1926]" pageId="2" pageNumber="598">Chen Amber Collection</collectingMunicipality>
(No. CZT-PLE-MA1),
<collectingCounty id="9B124E4BFFF96F71FCDBBD59FC28F876" box="[868,983,1901,1926]" pageId="2" pageNumber="598">Yangzhou</collectingCounty>
,
<collectingCountry id="0ADB7657FFF96F71FC5CBD59FBD5F876" box="[995,1066,1901,1926]" name="China" pageId="2" pageNumber="598">China</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3AD6654CFFF96F71FF78BDA5FCAEF802" pageId="2" pageNumber="598" type="etymology">
<paragraph id="727336C7FFF96F71FF78BDA5FF25F83E" blockId="2.[151,1437,1433,2034]" pageId="2" pageNumber="598">
<emphasis id="40B8EAD5FFF96F71FF78BDA5FEB1F85A" bold="true" box="[199,334,1937,1962]" pageId="2" pageNumber="598">Etymology.</emphasis>
The species is named after Yangzhou City, where the author has studied and worked for nearly ten years.
</paragraph>
<paragraph id="727336C7FFF96F71FF78BDEDFCAEF802" blockId="2.[151,1437,1433,2034]" box="[199,849,2009,2034]" pageId="2" pageNumber="598">
<emphasis id="40B8EAD5FFF96F71FF78BDEDFE8EF802" bold="true" box="[199,369,2009,2034]" pageId="2" pageNumber="598">Syninclusions.</emphasis>
Plant remains, including leaves and twigs.
</paragraph>
</subSubSection>
</treatment>
</document>
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