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<document id="A00736B3ABCBA0845A54F6C567C741FE" ID-CLB-Dataset="8826" ID-DOI="10.11646/zootaxa.4822.3.6" ID-GBIF-Dataset="a6962404-c25a-4f39-89a3-94ddc02bbe66" ID-ISSN="1175-5326" ID-Zenodo-Dep="4401755" ID-ZooBank="184B0899-2522-4974-8BF4-3902DBBE4BF6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609338286988" checkinUser="plazi" docAuthor="Sasakawa, Kôji, Berlov, Oleg &amp; Okuzaki, Yutaka" docDate="2020" docId="F8316A14FE47FFCAFF54CF1EBFF461B8" docLanguage="en" docName="zootaxa.4822.3.6.pdf" docOrigin="Zootaxa 4822 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pterostichus (Lenapterus) marginatus : Matsumura 1911" docType="treatment" docVersion="7" lastPageNumber="420" masterDocId="0408126CFE43FFCEFFC3CD3CBD666756" masterDocTitle="Taxonomic and nomenclatural changes in three species of Pterostichus Bonelli (Coleoptera: Carabidae) from the Far East" masterLastPageNumber="424" masterPageNumber="416" pageNumber="420" updateTime="1698849747236" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="6F97310C9BA948503D129632E8F51716">Taxonomic and nomenclatural changes in three species of Pterostichus Bonelli (Coleoptera: Carabidae) from the Far East</mods:title>
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<mods:namePart id="A07690F176CB5EEEC809DDA8A4834ACF">Sasakawa, Kôji</mods:namePart>
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<mods:affiliation id="B18329247BFFB1AF2C1C7C0293700683">Laboratory of Zoology, Department of Science Education, Faculty of Education, Chiba University, Yayoi-cho, Inage-ku, Chiba, Japan. ksasa @ chiba-u. jp; https: // orcid. org / 0000 - 0001 - 9246 - 5777</mods:affiliation>
<mods:nameIdentifier id="7F178C256D77D322E521B50397002BFE" type="email">ksasa@chiba-u.jp</mods:nameIdentifier>
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<mods:namePart id="2BD10C35A62812403F3162758D2F6882">Berlov, Oleg</mods:namePart>
<mods:affiliation id="176630F64B810A6140FA9ABA83BAFF4B">Laboratory of Experimental Animals, Anti-Plague Institute of Siberia and the Russian Far East, Irkutsk, Russia.</mods:affiliation>
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<mods:namePart id="A932FA6510A7B6BF09CF1AD45ACCAFE1">Okuzaki, Yutaka</mods:namePart>
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<mods:affiliation id="3B215C983E3DFA1EFBCF15CADC4F063B">Field Science Center for Northern Biosphere, Hokkaido University, Sapporo, Hokkaido, Japan. &amp; Department of General Systems Studies, Graduate School of Arts and Sciences, University of Tokyo, Komaba, Meguro-ku, Tokyo, Japan. yutaka @ g. ecc. u-tokyo. ac. jp; https: // orcid. org / 0000 - 0002 - 9362 - 0078</mods:affiliation>
<mods:nameIdentifier id="4C2ECEA4EE1B576BC728D2E964FAA34C" type="email">yutaka@g.ecc.u-tokyo.ac.jp</mods:nameIdentifier>
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<mods:date id="0FCA01903BFDCA835D6849659D962A38">2020</mods:date>
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<mods:number id="579DB48B0E9910984B83333517406DD7">4822</mods:number>
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<taxonomicName id="B798A081FE47FFCAFF54CF1EBE33656A" ID-CoL="8MMXP" authority="Matsumura, 1911" authorityName=": Matsumura" authorityYear="1911" box="[151,853,546,573]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="420" phylum="Arthropoda" rank="species" species="marginatus" subGenus="Lenapterus">
<emphasis id="42EC0710FE47FFCAFF54CF1EBE33656A" bold="true" box="[151,853,546,573]" pageId="4" pageNumber="420">
<emphasis id="42EC0710FE47FFCAFF54CF1EBC56656B" bold="true" box="[151,304,546,573]" italics="true" pageId="4" pageNumber="420">Pterostichus</emphasis>
(
<emphasis id="42EC0710FE47FFCAFE81CF1FBCA8656B" bold="true" box="[322,462,547,573]" italics="true" pageId="4" pageNumber="420">Lenapterus</emphasis>
)
<emphasis id="42EC0710FE47FFCAFE23CF1FBF0B656B" bold="true" box="[480,621,547,573]" italics="true" pageId="4" pageNumber="420">marginatus</emphasis>
<bibRefCitation id="1409A6F3FE47FFCAFDB6CF1FBE33656A" author="Matsumura, S." box="[629,853,546,573]" pageId="4" pageNumber="420" pagination="1 - 145" refId="ref6655" refString="Matsumura, S. (1911) Erster Beitrag zur Insekten-Fauna von Sachalin. Journal of the College of Agriculture, Tohoku Imperial University, Sapporo, Japan, 4, 1 - 145." type="journal article" year="1911">Matsumura, 1911</bibRefCitation>
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<figureCitation id="E8A3C787FE47FFCAFF54CF7BBDBC6537" box="[151,218,583,609]" captionStart="FIGURE 3" captionStartId="5.[151,250,1249,1273]" captionTargetBox="[151,1436,181,1225]" captionTargetId="figure-26@5.[151,1436,181,1225]" captionTargetPageId="5" captionText="FIGURE 3. Habitus (A–E), male genitalia (F–I), and attached labels (G, H) of Pterostichus marginatus marginatus (A–C, F, G, J, K) and P. m. subrugosus (D, E, H, I). A, dorsal view of syntype male; B, dorsal view of syntype female; C, dorsal view of male from Porechje envir., Sakhalin; D, dorsal view of male from Kumono-daira; E, dorsal view of female from Kumono-daira; F, H, right lateral view of everted endophallus; G, I, left lateral view of everted endophallus; J, label attached to syntype male; K, labels attached to syntype female. dl, dorsal lob; gp, gonopore; ll, left lateral lobe; lv-1, basal sub-lobe of left ventral lobe; lv-2, apical sub-lobe of left ventral lobe; rv-1, basal sub-lobe of right ventral lobe; rv-2, apical sub-lobe of right ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.4401761" httpUri="https://zenodo.org/record/4401761/files/figure.png" pageId="4" pageNumber="420">Fig. 3</figureCitation>
</paragraph>
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<paragraph id="7027DB02FE47FFCAFF54CFB7BF5E65B7" blockId="4.[151,1437,651,982]" pageId="4" pageNumber="420">
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<treatmentCitation id="F139FD13FE47FFCAFF54CFB7BFF565F5" author="Matsumura, S." box="[151,659,651,675]" page="111" pageId="4" pageNumber="420" year="1911">
<taxonomicName id="B798A081FE47FFCAFF54CFB7BF0265F5" ID-CoL="8MMXP" authority=": Matsumura (1911)" authorityName=": Matsumura" authorityYear="1911" box="[151,612,651,675]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="420" phylum="Arthropoda" rank="species" species="marginatus">
<emphasis id="42EC0710FE47FFCAFF54CFB7BCF565F5" box="[151,403,651,675]" italics="true" pageId="4" pageNumber="420">Pterostichus marginatus</emphasis>
:
<bibRefCitation id="1409A6F3FE47FFCAFE63CFB0BF0265F5" author="Matsumura, S." box="[416,612,651,675]" pageId="4" pageNumber="420" pagination="1 - 145" refId="ref6655" refString="Matsumura, S. (1911) Erster Beitrag zur Insekten-Fauna von Sachalin. Journal of the College of Agriculture, Tohoku Imperial University, Sapporo, Japan, 4, 1 - 145." type="journal article" year="1911">Matsumura (1911)</bibRefCitation>
</taxonomicName>
: 111
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(original description), type locality: “Tonnaitcha, Ushoro” [written as “Oshiyoro” on the labels of
<typeStatus id="AF2365A0FE47FFCAFE7CCF96BF7A6594" box="[447,540,682,706]" pageId="4" pageNumber="420" type="syntype">syntypes</typeStatus>
, but both corresponding to the same locality, Okhotskoe, southern
<collectingRegion id="B25C15E0FE47FFCAFB09CF96B84D6594" box="[1226,1323,682,706]" country="Russia" name="Sakhalin" pageId="4" pageNumber="420">Sakhalin</collectingRegion>
];
<treatmentCitation id="F139FD13FE47FFCAFAF8CF96BC2265B6" author="Bousquet, Y." page="515" pageId="4" pageNumber="420" year="2003">
<bibRefCitation id="1409A6F3FE47FFCAFAF8CF96BC7665B7" author="Bousquet, Y." pageId="4" pageNumber="420" pagination="1810" refId="ref5615" refString="Bousquet, Y. (2003) Tribe Pterostichini Bonelli, 1810. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata-Myxophaga-Adephaga. Apollo Books, Stenstrup, pp. 469 - 520." type="book chapter" year="2003">Bousquet (2003)</bibRefCitation>
: 515
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;
<treatmentCitation id="F139FD13FE47FFCAFE8CCFF5BF5265B7" author="Bousquet, Y." box="[335,564,713,737]" page="747" pageId="4" pageNumber="420" year="2017">
<bibRefCitation id="1409A6F3FE47FFCAFE8CCFF5BF6665B7" author="Bousquet, Y." box="[335,512,713,737]" pageId="4" pageNumber="420" pagination="1810" refId="ref5666" refString="Bousquet, Y. (2017) Tribe Pterostichini Bonelli, 1810. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata-Myxophaga-Adephaga. Brill, Leiden, Boston, pp. 675 - 755." type="book chapter" year="2017">Bousquet (2017)</bibRefCitation>
: 747
</treatmentCitation>
.
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</paragraph>
<paragraph id="7027DB02FE47FFCAFF54CFDBB90C6448" blockId="4.[151,1437,651,982]" pageId="4" pageNumber="420">
<treatmentCitationGroup id="5088FC2CFE47FFCAFF54CFDBB90C6448" pageId="4" pageNumber="420">
<taxonomicName id="B798A081FE47FFCAFF54CFDBBF8D65A9" authority=": Straneo (1955)" authorityName="Straneo" authorityYear="1955" box="[151,747,743,767]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="420" phylum="Arthropoda" rank="species" species="subrugosus" subGenus="Lyperopherus">
<emphasis id="42EC0710FE47FFCAFF54CFDBBC7E65A9" box="[151,280,743,767]" italics="true" pageId="4" pageNumber="420">Pterostichus</emphasis>
(
<emphasis id="42EC0710FE47FFCAFEEACFDBBCD165A9" box="[297,439,743,767]" italics="true" pageId="4" pageNumber="420">Lyperopherus</emphasis>
)
<emphasis id="42EC0710FE47FFCAFE0BCFDBBF5865A9" box="[456,574,743,767]" italics="true" pageId="4" pageNumber="420">subrugosus</emphasis>
:
<bibRefCitation id="1409A6F3FE47FFCAFD8DCFDBBF8D65A9" author="Straneo, S. L." box="[590,747,743,767]" pageId="4" pageNumber="420" pagination="85 - 101" refId="ref7016" refString="Straneo, S. L. (1955) Nuove specie di Pterostichus del Giappone (Coleopt. Carabis.) Annali del Museo Civico di Storia Naturale &quot; Giacomo Doria &quot;, 68, 85 - 101." type="journal article" year="1955">Straneo (1955)</bibRefCitation>
</taxonomicName>
(original description): 88,
<typeStatus id="AF2365A0FE47FFCAFBCACFD4B9506456" box="[1033,1078,744,768]" pageId="4" pageNumber="420">type</typeStatus>
locality “M. Daisetsu, Hokuchindake” [Mt. Hokuchin-dake, the Daisetsuzan Mountains,
<collectingRegion id="B25C15E0FE47FFCAFCD2CE3ABE1B6448" box="[785,893,774,798]" country="Japan" name="Hokkaido" pageId="4" pageNumber="420">Hokkaidô</collectingRegion>
];
<treatmentCitation id="F139FD13FE47FFCAFC4ECE3AB9006448" author="Jedlicka, A." box="[909,1126,774,798]" page="266" pageId="4" pageNumber="420" year="1962">
<bibRefCitation id="1409A6F3FE47FFCAFC4ECE3AB9546448" author="Jedlicka, A." box="[909,1074,774,798]" pageId="4" pageNumber="420" pagination="177 - 346" refId="ref5989" refString="Jedlicka, A. (1962) Monographie des Tribus Pterostichini aus Ostasien (Pterostichini, Trigonotomi, Myadi) (Coleoptera-Ca- rabidae). Entomologische Anhandlungen und Berichte aus dem Staatliche Museum fur Tierkunde in Dresden, 26 (21), 177 - 346." type="journal article" year="1962">Jedlička (1962)</bibRefCitation>
: 266
</treatmentCitation>
.
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</paragraph>
<paragraph id="7027DB02FE47FFCAFF54CE18B9E6640D" blockId="4.[151,1437,651,982]" pageId="4" pageNumber="420">
<treatmentCitationGroup id="5088FC2CFE47FFCAFF54CE18B9E6640D" pageId="4" pageNumber="420">
<treatmentCitation id="F139FD13FE47FFCAFF54CE18BE1E646B" author="Kirschenhofer, E." box="[151,888,804,829]" page="224" pageId="4" pageNumber="420" year="1985">
<taxonomicName id="B798A081FE47FFCAFF54CE18BE23646A" ID-CoL="8MMJR" authority=": Kirschenhofer (1985)" authorityName=": Kirschenhofer" authorityYear="1985" box="[151,837,804,829]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="420" phylum="Arthropoda" rank="species" species="wellschmiedi" subGenus="Lyperopherus">
<emphasis id="42EC0710FE47FFCAFF54CE18BC7E646A" box="[151,280,804,828]" italics="true" pageId="4" pageNumber="420">Pterostichus</emphasis>
(
<emphasis id="42EC0710FE47FFCAFEEACE18BCD1646A" box="[297,439,804,828]" italics="true" pageId="4" pageNumber="420">Lyperopherus</emphasis>
)
<emphasis id="42EC0710FE47FFCAFE0BCE18BF29646A" box="[456,591,804,828]" italics="true" pageId="4" pageNumber="420">wellschmiedi</emphasis>
:
<materialsCitation id="C0F0D15FFE47FFCAFD9CCE18BE23646A" ID-GBIF-Occurrence="3019437309" box="[607,837,804,828]" location="Mt. Chekhov" pageId="4" pageNumber="420" specimenCount="1" stateProvince="Sakhalin">
<bibRefCitation id="1409A6F3FE47FFCAFD9CCE18BE23646A" author="Kirschenhofer, E." box="[607,837,804,828]" pageId="4" pageNumber="420" pagination="221 - 231" refId="ref6265" refString="Kirschenhofer, E. (1985) Zweineue Pterostichus - Arten aus Sachalin: Pterostichus (Steropus) sachalinensis sp. n. und Pterostichus (Lyperopherus) wellschmiedi sp. n. Entomofauna, 6, 221 - 231." type="journal article" year="1985">Kirschenhofer (1985)</bibRefCitation>
</materialsCitation>
</taxonomicName>
: 224
</treatmentCitation>
(original description),
<typeStatus id="AF2365A0FE47FFCAFBAECE19B9FC646B" box="[1133,1178,805,829]" pageId="4" pageNumber="420">type</typeStatus>
locality: “Süd-Sachalin, Umgeb. Nowoalexandrowsk, Berg Tschkowa” [
<location id="75478DD9FE47FFCAFD76CE78BE26640D" LSID="urn:lsid:plazi:treatment:F8316A14FE47FFCAFF54CF1EBFF461B8:75478DD9FE47FFCAFD76CE78BE26640D" box="[693,832,835,859]" name="Mt. Chekhov" pageId="4" pageNumber="420" stateProvince="Sakhalin">Mt. Chekhov</location>
, southern
<collectingRegion id="B25C15E0FE47FFCAFC68CE7FB96A640D" box="[939,1036,835,859]" country="Russia" name="Sakhalin" pageId="4" pageNumber="420">Sakhalin</collectingRegion>
].
<emphasis id="42EC0710FE47FFCAFBDFCE7FB9E6640D" bold="true" box="[1052,1152,835,859]" pageId="4" pageNumber="420">Syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="7027DB02FE47FFCAFF54CE5EBEB5642C" blockId="4.[151,1437,651,982]" box="[151,979,866,890]" pageId="4" pageNumber="420">
<treatmentCitationGroup id="5088FC2CFE47FFCAFF54CE5EBEB5642C" box="[151,979,866,890]" pageId="4" pageNumber="420">
<treatmentCitation id="F139FD13FE47FFCAFF54CE5EBFB9642C" author="Morita, S." box="[151,735,866,890]" page="4" pageId="4" pageNumber="420" year="1995">
<taxonomicName id="B798A081FE47FFCAFF54CE5EBFA3642C" authority=": Morita (1995)" authorityName=": Morita" authorityYear="1995" box="[151,709,866,890]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="420" phylum="Arthropoda" rank="species" species="subrugosus" subGenus="Lenapterus">
<emphasis id="42EC0710FE47FFCAFF54CE5EBC7E642C" box="[151,280,866,890]" italics="true" pageId="4" pageNumber="420">Pterostichus</emphasis>
(
<emphasis id="42EC0710FE47FFCAFEE4CE5FBCFD642C" box="[295,411,867,890]" italics="true" pageId="4" pageNumber="420">Lenapterus</emphasis>
)
<emphasis id="42EC0710FE47FFCAFE68CE5EBF47642C" box="[427,545,866,890]" italics="true" pageId="4" pageNumber="420">subrugosus</emphasis>
:
<bibRefCitation id="1409A6F3FE47FFCAFDECCE5EBFA3642C" author="Morita, S." box="[559,709,866,890]" pageId="4" pageNumber="420" pagination="1 - 7" refId="ref6725" refString="Morita, S. (1995) Carabid beetles of the Island of Rishiri-to off northern Hokkaido, northeast Japan. Rishiri Studies, 15, 1 - 7." type="journal article" year="1995">Morita (1995)</bibRefCitation>
</taxonomicName>
: 4
</treatmentCitation>
;
<treatmentCitation id="F139FD13FE47FFCAFD29CE5EBEA9642C" author="Bousquet, Y." box="[746,975,866,890]" page="497" pageId="4" pageNumber="420" year="2003">
<bibRefCitation id="1409A6F3FE47FFCAFD29CE5EBEFD642C" author="Bousquet, Y." box="[746,923,866,890]" pageId="4" pageNumber="420" pagination="1810" refId="ref5615" refString="Bousquet, Y. (2003) Tribe Pterostichini Bonelli, 1810. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata-Myxophaga-Adephaga. Apollo Books, Stenstrup, pp. 469 - 520." type="book chapter" year="2003">Bousquet (2003)</bibRefCitation>
: 497
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="7027DB02FE47FFCAFF54CEBCBEF664CE" blockId="4.[151,1437,651,982]" box="[151,912,896,921]" pageId="4" pageNumber="420">
<treatmentCitationGroup id="5088FC2CFE47FFCAFF54CEBCBEF664CE" box="[151,912,896,921]" pageId="4" pageNumber="420">
<treatmentCitation id="F139FD13FE47FFCAFF54CEBCBEEA64CE" author="Kryzhanovskij, O. L. &amp; Belousov, I. A. &amp; Kabak, I. I. &amp; Kataev, B. M. &amp; Makarov, K. V. &amp; Shilenkov, V. G." box="[151,908,896,921]" page="105" pageId="4" pageNumber="420" year="1995">
<taxonomicName id="B798A081FE47FFCAFF54CEBCBE3E64CE" ID-CoL="8MMJR" authority=": Kryzhanovskij et al. (1995)" authorityName="Kryzhanovskij" authorityYear="1995" box="[151,856,896,921]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="420" phylum="Arthropoda" rank="species" species="wellschmiedi" subGenus="Steroperis">
<emphasis id="42EC0710FE47FFCAFF54CEBCBC7E64CE" box="[151,280,896,920]" italics="true" pageId="4" pageNumber="420">Pterostichus</emphasis>
(
<emphasis id="42EC0710FE47FFCAFEE4CEBDBCF664CE" box="[295,400,897,920]" italics="true" pageId="4" pageNumber="420">Steroperis</emphasis>
)
<emphasis id="42EC0710FE47FFCAFE63CEBCBF4164CE" box="[416,551,896,920]" italics="true" pageId="4" pageNumber="420">wellschmiedi</emphasis>
:
<bibRefCitation id="1409A6F3FE47FFCAFDF6CEBCBE3E64CE" author="Kryzhanovskij, O. L. &amp; Belousov, I. A. &amp; Kabak, I. I. &amp; Kataev, B. M. &amp; Makarov, K. V. &amp; Shilenkov, V. G." box="[565,856,896,920]" pageId="4" pageNumber="420" refId="ref6365" refString="Kryzhanovskij, O. L., Belousov, I. A., Kabak, I. I., Kataev, B. M., Makarov, K. V. &amp; Shilenkov, V. G. (1995) A Checklist of the Ground-Beetles of Russia and Adjacent Lands (Insecta, Coleoptera, Carabidae). Pensoft Publishers, Sofia, 271 pp." type="book" year="1995">
Kryzhanovskij
<emphasis id="42EC0710FE47FFCAFD17CEBDBE6F64CE" box="[724,777,896,920]" italics="true" pageId="4" pageNumber="420">et al.</emphasis>
(1995)
</bibRefCitation>
</taxonomicName>
: 105
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="7027DB02FE47FFCAFF54CEA3BCD66480" blockId="4.[151,1437,651,982]" pageId="4" pageNumber="420">
<treatmentCitationGroup id="5088FC2CFE47FFCAFF54CEA3BCD66480" pageId="4" pageNumber="420">
<treatmentCitation id="F139FD13FE47FFCAFF54CEA3BEE164E1" author="Berlov, E. &amp; Berlov, O." box="[151,903,927,951]" page="39" pageId="4" pageNumber="420" year="1997">
<taxonomicName id="B798A081FE47FFCAFF54CEA3BE3B64E1" ID-CoL="8MMJR" authority=": Berlov &amp; Berlov (1997 d)" authorityName=": Berlov &amp; Berlov" authorityYear="1997" box="[151,861,927,951]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="420" phylum="Arthropoda" rank="species" species="wellschmiedi" subGenus="Lenapterus">
<emphasis id="42EC0710FE47FFCAFF54CEA3BC7E64E1" box="[151,280,927,951]" italics="true" pageId="4" pageNumber="420">Pterostichus</emphasis>
(
<emphasis id="42EC0710FE47FFCAFEE9CE9CBCF864E1" box="[298,414,928,951]" italics="true" pageId="4" pageNumber="420">Lenapterus</emphasis>
)
<emphasis id="42EC0710FE47FFCAFE72CEA3BF5E64E1" box="[433,568,927,951]" italics="true" pageId="4" pageNumber="420">wellschmiedi</emphasis>
:
<bibRefCitation id="1409A6F3FE47FFCAFD8ACEA3BE3B64E1" author="Berlov, E. &amp; Berlov, O." box="[585,861,927,951]" pageId="4" pageNumber="420" pagination="36 - 46" refId="ref5521" refString="Berlov, E. &amp; Berlov, O. (1997 d) A key to species of the genus Pterostichus (Coleoptera, Carabidae) from Sakhalin Island. Vestnik of the Irkutsk State Agricultural Academy, 5, 36 - 46." type="journal article" year="1997">Berlov &amp; Berlov (1997d)</bibRefCitation>
</taxonomicName>
: 39
</treatmentCitation>
;
<treatmentCitation id="F139FD13FE47FFCAFC56CE9CB91B64E1" author="Bousquet, Y." box="[917,1149,927,951]" page="497" pageId="4" pageNumber="420" year="2003">
<bibRefCitation id="1409A6F3FE47FFCAFC56CE9CB92F64E1" author="Bousquet, Y." box="[917,1097,927,951]" pageId="4" pageNumber="420" pagination="1810" refId="ref5615" refString="Bousquet, Y. (2003) Tribe Pterostichini Bonelli, 1810. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata-Myxophaga-Adephaga. Apollo Books, Stenstrup, pp. 469 - 520." type="book chapter" year="2003">Bousquet (2003)</bibRefCitation>
: 497
</treatmentCitation>
(synonym of
<taxonomicName id="B798A081FE47FFCAFAD4CEA3B8EB64E1" authorityName=": Morita" authorityYear="1995" box="[1303,1421,927,951]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="420" phylum="Arthropoda" rank="species" species="subrugosus">
<emphasis id="42EC0710FE47FFCAFAD4CEA3B8EB64E1" box="[1303,1421,927,951]" italics="true" pageId="4" pageNumber="420">subrugosus</emphasis>
</taxonomicName>
);
<treatmentCitation id="F139FD13FE47FFCAFF04CE82BCCA6480" author="Bousquet, Y." box="[199,428,958,982]" page="718" pageId="4" pageNumber="420" year="2017">
<bibRefCitation id="1409A6F3FE47FFCAFF04CE82BC1F6480" author="Bousquet, Y." box="[199,377,958,982]" pageId="4" pageNumber="420" pagination="1810" refId="ref5666" refString="Bousquet, Y. (2017) Tribe Pterostichini Bonelli, 1810. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata-Myxophaga-Adephaga. Brill, Leiden, Boston, pp. 675 - 755." type="book chapter" year="2017">Bousquet (2017)</bibRefCitation>
: 718
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="38828889FE47FFCAFF54C938BFF461B8" pageId="4" pageNumber="420" type="materials_examined">
<paragraph id="7027DB02FE47FFCAFF54C938B9C163A0" blockId="4.[151,1437,1028,1775]" pageId="4" pageNumber="420">
<emphasis id="42EC0710FE47FFCAFF54C938BCF56348" bold="true" box="[151,403,1028,1054]" pageId="4" pageNumber="420">Materials examined.</emphasis>
<taxonomicName id="B798A081FE47FFCAFE5DC938BE256348" authority="marginatus" box="[414,835,1028,1054]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="420" phylum="Arthropoda" rank="subSpecies" species="marginatus" subSpecies="marginatus">
<emphasis id="42EC0710FE47FFCAFE5DC938BE256348" box="[414,835,1028,1054]" italics="true" pageId="4" pageNumber="420">Pterostichus marginatus marginatus</emphasis>
</taxonomicName>
,
<materialsCitation id="C0F0D15FFE47FFCAFC96C938B9AF6384" ID-GBIF-Occurrence="3019437308" collectingDate="1906-08-11" collectingDateMax="1991-07-28" collectingDateMin="1906-08-11" collectionCode="HUM" collectorName="E. Berlov" county="Mt. Chekhov" elevation="500" location="Sakhalin" municipality="Novoaleksandrovsk" pageId="4" pageNumber="420" specimenCount="1934" specimenCount-female="6" specimenCount-male="17" typeStatus="syntype">
<specimenCount id="669E108BFE47FFCAFC96C938BE186348" box="[853,894,1028,1054]" pageId="4" pageNumber="420" type="male">9♂</specimenCount>
<specimenCount id="669E108BFE47FFCAFCBDC938BEF86348" box="[894,926,1028,1054]" pageId="4" pageNumber="420" type="female">3♀</specimenCount>
,
<location id="75478DD9FE47FFCAFC6DC938B9746348" LSID="urn:lsid:plazi:treatment:F8316A14FE47FFCAFF54CF1EBFF461B8:75478DD9FE47FFCAFC6DC938B9746348" box="[942,1042,1028,1054]" county="Mt. Chekhov" municipality="Novoaleksandrovsk" name="Sakhalin" pageId="4" pageNumber="420">Sakhalin</location>
(
<taxonomicName id="B798A081FE47FFCAFBE5C939B8FA6348" authority="Matsumura, 1911" authorityName=": Matsumura" authorityYear="1911" box="[1062,1436,1028,1055]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="420" phylum="Arthropoda" rank="species" species="marginatus">
<emphasis id="42EC0710FE47FFCAFBE5C939B9A06348" box="[1062,1222,1029,1054]" italics="true" pageId="4" pageNumber="420">P. marginatus</emphasis>
<bibRefCitation id="1409A6F3FE47FFCAFB12C939B8FA6348" author="Matsumura, S." box="[1233,1436,1028,1055]" pageId="4" pageNumber="420" pagination="1 - 145" refId="ref6655" refString="Matsumura, S. (1911) Erster Beitrag zur Insekten-Fauna von Sachalin. Journal of the College of Agriculture, Tohoku Imperial University, Sapporo, Japan, 4, 1 - 145." type="journal article" year="1911">
Matsumura,
<specimenCount id="669E108BFE47FFCAFAA7C938B8FA6348" box="[1380,1436,1028,1054]" hasTypeStatus="true" pageId="4" pageNumber="420">1911</specimenCount>
</bibRefCitation>
</taxonomicName>
<typeStatus id="AF2365A0FE47FFCAFF54C915BD9D6315" box="[151,251,1065,1091]" pageId="4" pageNumber="420" type="syntype">syntypes</typeStatus>
<specimenCount id="669E108BFE47FFCAFEC7C914BC4C6314" box="[260,298,1064,1090]" pageId="4" pageNumber="420" type="male">1♂</specimenCount>
, “/
<date id="0426FDC2FE47FFCAFE88C914BC856314" box="[331,483,1064,1090]" pageId="4" pageNumber="420" value="1906-08-11">
<collectingDate id="1462042AFE47FFCAFE88C914BC856314" box="[331,483,1064,1090]" pageId="4" pageNumber="420" value="1906-08-11">11/VIII 1906</collectingDate>
</date>
Oshiyoro” [partly in Japanese] and
<specimenCount id="669E108BFE47FFCAFC48C914BED66314" box="[907,944,1064,1090]" pageId="4" pageNumber="420" type="female">1♀</specimenCount>
, “
<date id="0426FDC2FE47FFCAFC0AC914B9056314" box="[969,1123,1064,1090]" pageId="4" pageNumber="420" value="1906-08-11">
<collectingDate id="1462042AFE47FFCAFC0AC914B9056314" box="[969,1123,1064,1090]" pageId="4" pageNumber="420" value="1906-08-11">11/VIII 1906</collectingDate>
</date>
Oshiyoro” [partly in Japanese], “P. margina?? n sp.” in
<collectionCode id="168943C7FE47FFCAFE3CC971BF236331" box="[511,581,1101,1127]" country="Germany" httpUri="http://grbio.org/cool/kch5-hv72" name="Humboldt University Zoologisches Museum" pageId="4" pageNumber="420">HUM</collectionCode>
;
<specimenCount id="669E108BFE47FFCAFD90C970BF1B6330" box="[595,637,1100,1126]" pageId="4" pageNumber="420" type="male">3♂</specimenCount>
<specimenCount id="669E108BFE47FFCAFDBEC970BFFD6330" box="[637,667,1100,1126]" pageId="4" pageNumber="420" type="female">1♀</specimenCount>
,
<location id="75478DD9FE47FFCAFD68C970BE3E6330" LSID="urn:lsid:plazi:treatment:F8316A14FE47FFCAFF54CF1EBFF461B8:75478DD9FE47FFCAFD68C970BE3E6330" box="[683,856,1100,1126]" county="Mt. Chekhov" municipality="Novoaleksandrovsk" name="Makarov distr." pageId="4" pageNumber="420">Makarov distr.</location>
,
<location id="75478DD9FE47FFCAFCA5C970BEAC6330" LSID="urn:lsid:plazi:treatment:F8316A14FE47FFCAFF54CF1EBFF461B8:75478DD9FE47FFCAFCA5C970BEAC6330" box="[870,970,1100,1126]" county="Mt. Chekhov" municipality="Novoaleksandrovsk" name="Porechje" pageId="4" pageNumber="420">Porechje</location>
envir.,
<quantity id="B76076E7FE47FFCAFBE6C970B9FC6330" box="[1061,1178,1100,1127]" metricMagnitude="4" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="4" pageNumber="420" unit="km" value="12.5" valueMax="15.0" valueMin="10.0">10–15 km</quantity>
from sea shore, larch forest,
<date id="0426FDC2FE47FFCAFF2BC94CBC0863DC" box="[232,366,1136,1162]" pageId="4" pageNumber="420" value="1991-07-28">
<collectingDate id="1462042AFE47FFCAFF2BC94CBC0863DC" box="[232,366,1136,1162]" pageId="4" pageNumber="420" value="1991-07-28">28.vii.1991</collectingDate>
</date>
,
<collectorName id="DD6DBED4FE47FFCAFEB9C94DBC8F63DC" box="[378,489,1136,1163]" pageId="4" pageNumber="420">E. Berlov</collectorName>
leg. in BO coll.,
<specimenCount id="669E108BFE47FFCAFD77C94CBFBA63DC" box="[692,732,1136,1162]" pageId="4" pageNumber="420" type="male">1♂</specimenCount>
<specimenCount id="669E108BFE47FFCAFD1FC94CBF9A63DC" box="[732,764,1136,1162]" pageId="4" pageNumber="420" type="female">1♀</specimenCount>
,
<location id="75478DD9FE47FFCAFCCAC94CB94A63DC" LSID="urn:lsid:plazi:treatment:F8316A14FE47FFCAFF54CF1EBFF461B8:75478DD9FE47FFCAFCCAC94CB94A63DC" box="[777,1068,1136,1162]" county="Mt. Chekhov" municipality="Novoaleksandrovsk" name="Yuzhno-Sakhalinsk distr." pageId="4" pageNumber="420">Yuzhno-Sakhalinsk distr.</location>
,
<location id="75478DD9FE47FFCAFBFAC94CB87A63DC" LSID="urn:lsid:plazi:treatment:F8316A14FE47FFCAFF54CF1EBFF461B8:75478DD9FE47FFCAFBFAC94CB87A63DC" box="[1081,1308,1136,1162]" county="Mt. Chekhov" municipality="Novoaleksandrovsk" name="Novoaleksandrovsk" pageId="4" pageNumber="420">Novoaleksandrovsk</location>
envir.,
<location id="75478DD9FE47FFCAFAB7C94DBD9963F8" LSID="urn:lsid:plazi:treatment:F8316A14FE47FFCAFF54CF1EBFF461B8:75478DD9FE47FFCAFAB7C94DBD9963F8" county="Mt. Chekhov" municipality="Novoaleksandrovsk" name="Mt. Chekhov" pageId="4" pageNumber="420">Mt. Chekhov</location>
,
<date id="0426FDC2FE47FFCAFECFC9A8BCEC63F8" box="[268,394,1172,1198]" pageId="4" pageNumber="420" value="1990-06-19">
<collectingDate id="1462042AFE47FFCAFECFC9A8BCEC63F8" box="[268,394,1172,1198]" pageId="4" pageNumber="420" value="1990-06-19">19.vi.1990</collectingDate>
</date>
,
<collectorName id="DD6DBED4FE47FFCAFE54C9A9BF6063F8" box="[407,518,1172,1199]" pageId="4" pageNumber="420">E. Berlov</collectorName>
leg. in BO coll.;
<specimenCount id="669E108BFE47FFCAFD16C9A8BF9C63F8" box="[725,762,1172,1198]" pageId="4" pageNumber="420" type="male">3♂</specimenCount>
,
<location id="75478DD9FE47FFCAFCC4C9A8B94D63F8" LSID="urn:lsid:plazi:treatment:F8316A14FE47FFCAFF54CF1EBFF461B8:75478DD9FE47FFCAFCC4C9A8B94D63F8" box="[775,1067,1172,1198]" county="Mt. Chekhov" municipality="Novoaleksandrovsk" name="Yuzhno-Sakhalinsk distr." pageId="4" pageNumber="420">Yuzhno-Sakhalinsk distr.</location>
,
<collectingMunicipality id="90434178FE47FFCAFBF4C9A8B87C63F8" box="[1079,1306,1172,1198]" pageId="4" pageNumber="420">Novoaleksandrovsk</collectingMunicipality>
envir.,
<collectingCounty id="9946A38EFE47FFCAFAB7C9A9BD996384" pageId="4" pageNumber="420">Mt. Chekhov</collectingCounty>
,
<quantity id="B76076E7FE47FFCAFECFC984BCF76385" box="[268,401,1208,1235]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="4" pageNumber="420" unit="m" value="500.0" valueMax="600.0" valueMin="400.0">
<elevation id="FBB53C31FE47FFCAFECFC984BCF76385" box="[268,401,1208,1235]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="4" pageNumber="420" unit="m" value="500.0" valueMax="600.0" valueMin="400.0">400–600 m</elevation>
</quantity>
,
<date id="0426FDC2FE47FFCAFE5DC984BF7D6384" box="[414,539,1208,1234]" pageId="4" pageNumber="420" value="1988-06-10">
<collectingDate id="1462042AFE47FFCAFE5DC984BF7D6384" box="[414,539,1208,1234]" pageId="4" pageNumber="420" value="1988-06-10">10.vi.1988</collectingDate>
</date>
, leg.
<collectorName id="DD6DBED4FE47FFCAFD98C985BFAC6384" box="[603,714,1208,1235]" pageId="4" pageNumber="420">E. Berlov</collectorName>
leg. in BO coll.);
<taxonomicName id="B798A081FE47FFCAFC5CC985B9AF6384" authorityName="Sasakawa &amp; Berlov &amp; Okuzaki" authorityYear="2020" baseAuthorityName=": Straneo" baseAuthorityYear="1955" box="[927,1225,1208,1234]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="420" phylum="Arthropoda" rank="subSpecies" species="marginatus" subSpecies="subrugosus">
<emphasis id="42EC0710FE47FFCAFC5CC985B9AF6384" box="[927,1225,1208,1234]" italics="true" pageId="4" pageNumber="420">P. marginatus subrugosus</emphasis>
</taxonomicName>
</materialsCitation>
,
<materialsCitation id="C0F0D15FFE47FFCAFB1BC984B9C563A0" ID-GBIF-Occurrence="3019437302" collectingDate="2005-08-05" collectorName="N. Yasuda" country="Japan" location="Kumonodaira" municipality="Daisetsuzan Mountains" pageId="4" pageNumber="420" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Hokkaido" typeStatus="syntype">
<specimenCount id="669E108BFE47FFCAFB1BC984B8666384" box="[1240,1280,1208,1234]" pageId="4" pageNumber="420" type="male">2♂</specimenCount>
<specimenCount id="669E108BFE47FFCAFAC2C984B8476384" box="[1281,1313,1208,1234]" pageId="4" pageNumber="420" type="female">2♀</specimenCount>
,
<location id="75478DD9FE47FFCAFAECC985BDB563A0" LSID="urn:lsid:plazi:treatment:F8316A14FE47FFCAFF54CF1EBFF461B8:75478DD9FE47FFCAFAECC985BDB563A0" country="Japan" municipality="Daisetsuzan Mountains" name="Kumonodaira" pageId="4" pageNumber="420" stateProvince="Hokkaido">Kumonodaira</location>
, the
<collectingMunicipality id="90434178FE47FFCAFECAC9E0BF7E63A0" box="[265,536,1244,1270]" pageId="4" pageNumber="420">Daisetsuzan Mountains</collectingMunicipality>
,
<collectingRegion id="B25C15E0FE47FFCAFDE7C9E0BFFE63A0" box="[548,664,1244,1270]" country="Japan" name="Hokkaido" pageId="4" pageNumber="420">Hokkaidô</collectingRegion>
,
<collectingCountry id="088F9B92FE47FFCAFD60C9E1BF8163A1" box="[675,743,1245,1271]" name="Japan" pageId="4" pageNumber="420">Japan</collectingCountry>
,
<date id="0426FDC2FE47FFCAFD30C9E0BE1463A1" box="[755,882,1244,1271]" pageId="4" pageNumber="420" value="2005-08-05">
<collectingDate id="1462042AFE47FFCAFD30C9E0BE1463A1" box="[755,882,1244,1271]" pageId="4" pageNumber="420" value="2005-08-05">5.viii.2005</collectingDate>
</date>
,
<collectorName id="DD6DBED4FE47FFCAFCBDC9E1BE9563A0" box="[894,1011,1244,1271]" pageId="4" pageNumber="420">N. Yasuda</collectorName>
leg. in KS coll
</materialsCitation>
.
</paragraph>
<paragraph id="7027DB02FE47FFCAFF04C83CB8376269" blockId="4.[151,1437,1028,1775]" pageId="4" pageNumber="420">
<emphasis id="42EC0710FE47FFCAFF04C83CBFC4624C" bold="true" box="[199,674,1280,1306]" pageId="4" pageNumber="420">Other consubgeneric species examined.</emphasis>
The following species were investigated through literature:
<taxonomicName id="B798A081FE47FFCAFA8DC83DBCF96269" authority="(Motschulsky)" baseAuthorityName="Motschulsky" baseAuthorityYear="1860" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="420" phylum="Arthropoda" rank="species" species="cancellatus">
<emphasis id="42EC0710FE47FFCAFA8DC83DBD966268" italics="true" pageId="4" pageNumber="420">P. cancellatus</emphasis>
(Motschulsky)
</taxonomicName>
by
<bibRefCitation id="1409A6F3FE47FFCAFE09C818BFF26269" author="Sundukov, Y. N." box="[458,660,1316,1343]" pageId="4" pageNumber="420" pagination="480 - 503" refId="ref7125" refString="Sundukov, Y. N. (2005) A review of species of the subgenus Lenapterus (Coleoptera, Carabidae, Pterostichus) with description of new species and subspecies from the Sikhote-Alin Mountains. Entomological Review, 85 (5), 480 - 503." type="journal article" year="2005">Sundukov (2005)</bibRefCitation>
;
<taxonomicName id="B798A081FE47FFCAFD62C819BE876269" authority="(Menetries)" baseAuthorityName="Menetries" baseAuthorityYear="1851" box="[673,993,1316,1343]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="420" phylum="Arthropoda" rank="species" species="vermiculosus">
<emphasis id="42EC0710FE47FFCAFD62C819BE326268" box="[673,852,1316,1342]" italics="true" pageId="4" pageNumber="420">P. vermiculosus</emphasis>
(Ménétriés)
</taxonomicName>
by
<bibRefCitation id="1409A6F3FE47FFCAFBCFC818B82A6269" author="Sasakawa, K. &amp; Kubota, K." box="[1036,1356,1316,1343]" pageId="4" pageNumber="420" pagination="389 - 404" refId="ref6803" refString="Sasakawa, K. &amp; Kubota, K. (2005) Cryptic species of the subgenus Morphnosoma Lutshnik (Coleoptera: Carabidae; genus Pterostichus) from Japan. Entomological Science, 8, 389 - 404. https: // doi. org / 10.1111 / j. 1479 - 8298.2005.00139. x" type="journal article" year="2005">Sasakawa &amp; Kubota (2005)</bibRefCitation>
.
</paragraph>
<paragraph id="7027DB02FE47FFCAFF04C874BFF461B8" blockId="4.[151,1437,1028,1775]" pageId="4" pageNumber="420">
<emphasis id="42EC0710FE47FFCAFF04C874BC756234" bold="true" box="[199,275,1352,1378]" pageId="4" pageNumber="420">Notes.</emphasis>
<taxonomicName id="B798A081FE47FFCAFEDFC874BF526234" authorityName=": Matsumura" authorityYear="1911" box="[284,564,1352,1378]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="420" phylum="Arthropoda" rank="species" species="marginatus">
<emphasis id="42EC0710FE47FFCAFEDFC874BF526234" box="[284,564,1352,1378]" italics="true" pageId="4" pageNumber="420">Pterostichus marginatus</emphasis>
</taxonomicName>
was described from southern
<collectingRegion id="B25C15E0FE47FFCAFC56C874BE9F6234" box="[917,1017,1352,1378]" country="Russia" name="Sakhalin" pageId="4" pageNumber="420">Sakhalin</collectingRegion>
but has been forgotten since its description and treated as
<emphasis id="42EC0710FE47FFCAFE60C851BF2762D0" box="[419,577,1388,1414]" italics="true" pageId="4" pageNumber="420">incertae sedis</emphasis>
in the Palaearctic Catalogue (
<bibRefCitation id="1409A6F3FE47FFCAFC50C850B92562D0" author="Bousquet, Y." box="[915,1091,1388,1414]" pageId="4" pageNumber="420" pagination="1810" refId="ref5615" refString="Bousquet, Y. (2003) Tribe Pterostichini Bonelli, 1810. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata-Myxophaga-Adephaga. Apollo Books, Stenstrup, pp. 469 - 520." type="book chapter" year="2003">Bousquet 2003</bibRefCitation>
,
<bibRefCitation id="1409A6F3FE47FFCAFB8EC850B9ED62D0" author="Bousquet, Y." box="[1101,1163,1388,1414]" pageId="4" pageNumber="420" pagination="1810" refId="ref5666" refString="Bousquet, Y. (2017) Tribe Pterostichini Bonelli, 1810. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata-Myxophaga-Adephaga. Brill, Leiden, Boston, pp. 675 - 755." type="book chapter" year="2017">2017</bibRefCitation>
). Our comparison of the
<typeStatus id="AF2365A0FE47FFCAFF54C8ADBD9D62FD" box="[151,251,1425,1451]" pageId="4" pageNumber="420" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="B798A081FE47FFCAFEE1C8ADBCD962FC" authorityName=": Matsumura" authorityYear="1911" box="[290,447,1425,1450]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="420" phylum="Arthropoda" rank="species" species="marginatus">
<emphasis id="42EC0710FE47FFCAFEE1C8ADBCD962FC" box="[290,447,1425,1450]" italics="true" pageId="4" pageNumber="420">P. marginatus</emphasis>
</taxonomicName>
with a photo of a female
<typeStatus id="AF2365A0FE47FFCAFD25C8ADBE2F62FD" box="[742,841,1425,1451]" pageId="4" pageNumber="420" type="paratype">paratype</typeStatus>
of
<taxonomicName id="B798A081FE47FFCAFCB3C8ADB94762FC" authorityName=": Berlov &amp; Berlov" authorityYear="1997" box="[880,1057,1424,1450]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="420" phylum="Arthropoda" rank="species" species="wellschmiedi">
<emphasis id="42EC0710FE47FFCAFCB3C8ADB94762FC" box="[880,1057,1424,1450]" italics="true" pageId="4" pageNumber="420">P. wellschmiedi</emphasis>
</taxonomicName>
(
<bibRefCitation id="1409A6F3FE47FFCAFBF3C8ACB87C62FD" author="Kirschenhofer, E." box="[1072,1306,1424,1451]" pageId="4" pageNumber="420" pagination="221 - 231" refId="ref6265" refString="Kirschenhofer, E. (1985) Zweineue Pterostichus - Arten aus Sachalin: Pterostichus (Steropus) sachalinensis sp. n. und Pterostichus (Lyperopherus) wellschmiedi sp. n. Entomofauna, 6, 221 - 231." type="journal article" year="1985">Kirschenhofer 1985</bibRefCitation>
, fig. 9) and with specimens treated as
<taxonomicName id="B798A081FE47FFCAFE7CC889BF166298" authorityName=": Berlov &amp; Berlov" authorityYear="1997" box="[447,624,1460,1486]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="420" phylum="Arthropoda" rank="species" species="wellschmiedi">
<emphasis id="42EC0710FE47FFCAFE7CC889BF166298" box="[447,624,1460,1486]" italics="true" pageId="4" pageNumber="420">P. wellschmiedi</emphasis>
</taxonomicName>
revealed that
<taxonomicName id="B798A081FE47FFCAFCCEC889BEDB6298" authorityName=": Berlov &amp; Berlov" authorityYear="1997" box="[781,957,1460,1486]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="420" phylum="Arthropoda" rank="species" species="wellschmiedi">
<emphasis id="42EC0710FE47FFCAFCCEC889BEDB6298" box="[781,957,1460,1486]" italics="true" pageId="4" pageNumber="420">P. wellschmiedi</emphasis>
</taxonomicName>
is a synonym of
<taxonomicName id="B798A081FE47FFCAFBBFC889B87E6298" authorityName=": Matsumura" authorityYear="1911" box="[1148,1304,1461,1486]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="420" phylum="Arthropoda" rank="species" species="marginatus">
<emphasis id="42EC0710FE47FFCAFBBFC889B87E6298" box="[1148,1304,1461,1486]" italics="true" pageId="4" pageNumber="420">P. marginatus</emphasis>
</taxonomicName>
. Moreover, comparison of this
<collectingRegion id="B25C15E0FE47FFCAFEB3C8E4BCB262A4" box="[368,468,1496,1522]" country="Russia" name="Sakhalin" pageId="4" pageNumber="420">Sakhalin</collectingRegion>
species with
<taxonomicName id="B798A081FE47FFCAFDAEC8E5BE6C62A4" authorityName=": Morita" authorityYear="1995" box="[621,778,1496,1522]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="420" phylum="Arthropoda" rank="species" species="subrugosus">
<emphasis id="42EC0710FE47FFCAFDAEC8E5BE6C62A4" box="[621,778,1496,1522]" italics="true" pageId="4" pageNumber="420">P. subrugosus</emphasis>
</taxonomicName>
from
<collectingRegion id="B25C15E0FE47FFCAFC8DC8E4BED962A4" box="[846,959,1496,1522]" country="Japan" name="Hokkaido" pageId="4" pageNumber="420">Hokkaidô</collectingRegion>
revealed the relationship between the two taxa. Although the previously reported differences in external characters between the two (e.g., flatter elytra with less frequently interrupted intervals, and the absence of metallic luster of dorsal surface in
<taxonomicName id="B798A081FE47FFCAFB5ECB1DB85D616C" authorityName=": Morita" authorityYear="1995" box="[1181,1339,1568,1594]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="420" phylum="Arthropoda" rank="species" species="subrugosus">
<emphasis id="42EC0710FE47FFCAFB5ECB1DB85D616C" box="[1181,1339,1568,1594]" italics="true" pageId="4" pageNumber="420">P. subrugosus</emphasis>
</taxonomicName>
;
<bibRefCitation id="1409A6F3FE47FFCAFA89CB1CBC636109" author="Sundukov, Y. N." pageId="4" pageNumber="420" pagination="480 - 503" refId="ref7125" refString="Sundukov, Y. N. (2005) A review of species of the subgenus Lenapterus (Coleoptera, Carabidae, Pterostichus) with description of new species and subspecies from the Sikhote-Alin Mountains. Entomological Review, 85 (5), 480 - 503." type="journal article" year="2005">Sundukov 2005</bibRefCitation>
) were confirmed, no conspicuous differences were found in the structure of the endophallus. Thus,
<taxonomicName id="B798A081FE47FFCAFA44CB79BC7F61D4" authorityName=": Morita" authorityYear="1995" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="420" phylum="Arthropoda" rank="species" species="subrugosus">
<emphasis id="42EC0710FE47FFCAFA44CB79BC7F61D4" italics="true" pageId="4" pageNumber="420">P. subrugosus</emphasis>
</taxonomicName>
is downgraded from a distinct species to subspecies
<taxonomicName id="B798A081FE47FFCAFCB2CB55B9F061D4" authorityName="Sasakawa &amp; Berlov &amp; Okuzaki" authorityYear="2020" baseAuthorityName=": Straneo" baseAuthorityYear="1955" box="[881,1174,1640,1666]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="420" phylum="Arthropoda" rank="subSpecies" species="marginatus" status="stat. nov." subSpecies="subrugosus">
<emphasis id="42EC0710FE47FFCAFCB2CB55B9F061D4" box="[881,1174,1640,1666]" italics="true" pageId="4" pageNumber="420">P. marginatus subrugosus</emphasis>
</taxonomicName>
<emphasis id="42EC0710FE47FFCAFB5ECB54B86C61D4" bold="true" box="[1181,1290,1640,1666]" pageId="4" pageNumber="420">
<taxonomicNameLabel id="59DFBA6BFE47FFCAFB5ECB54B86C61D4" box="[1181,1290,1640,1666]" pageId="4" pageNumber="420">stat. nov.</taxonomicNameLabel>
</emphasis>
This species was recorded from the summit of Mt. Rishiri-dake based on two female specimens (
<bibRefCitation id="1409A6F3FE47FFCAFB98CBB0B98861F1" author="Morita, S." box="[1115,1262,1676,1703]" pageId="4" pageNumber="420" pagination="1 - 7" refId="ref6725" refString="Morita, S. (1995) Carabid beetles of the Island of Rishiri-to off northern Hokkaido, northeast Japan. Rishiri Studies, 15, 1 - 7." type="journal article" year="1995">Morita 1995</bibRefCitation>
, as
<taxonomicName id="B798A081FE47FFCAFADACBB1BDDD619C" authorityName=": Morita" authorityYear="1995" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="420" phylum="Arthropoda" rank="species" species="subrugosus">
<emphasis id="42EC0710FE47FFCAFADACBB1BDDD619C" italics="true" pageId="4" pageNumber="420">P. subrugosus</emphasis>
</taxonomicName>
). However, the subspecies identity of the Rishiri population remains undetermined because specimens from the locality were not examined in this study.
</paragraph>
</subSubSection>
</treatment>
</document>
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