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<document id="ABA5E7EA654696EE8C142A532974452F" ID-DOI="10.1111/j.1096-3642.2008.00415.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5446700" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630461491835" checkinUser="felipe" docAuthor="Wu, Shu-Ping, Hwang, Chung-Chi &amp; Lin, Yao-Sung" docDate="2008" docId="E45287EFFFFF4314FCC0FE37819757D0" docLanguage="en" docName="j.1096-3642.2008.00415.x.pdf" docOrigin="Zoological Journal of the Linnean Society 154 (3)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00415.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Satsuma hagiomontis Wu &amp; Hwang &amp; Lin 2008, SP. NOV." docType="treatment" docVersion="4" lastPageNumber="449" masterDocId="186BFF97FFF54318FF92FFF482155506" masterDocTitle="Systematic revision of the arboreal snail Satsuma albida species complex (Mollusca: Camaenidae) with descriptions of 14 new species from Taiwan" masterLastPageNumber="493" masterPageNumber="437" pageNumber="447" updateTime="1699279363994" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="1A55030C6823A4080BB3C2ED157BB659">Systematic revision of the arboreal snail Satsuma albida species complex (Mollusca: Camaenidae) with descriptions of 14 new species from Taiwan</mods:title>
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<mods:namePart id="153D6F6DFBFC911B2BD277DC96452625">Wu, Shu-Ping</mods:namePart>
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<mods:namePart id="41D34F71D631BCB056D78F05FECF71E5">Hwang, Chung-Chi</mods:namePart>
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<mods:namePart id="F325C1D40E480E2AAD310DD98D2AAC96">Lin, Yao-Sung</mods:namePart>
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<mods:title id="CB82E0BA2C482096E98FEAA7CA55DE7B">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="CB3DD0A5A0411C1F9C70639FB6C7E0AF">2008</mods:date>
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<mods:number id="CF76A58B088A3B1A620314175529DA23">2008-10-31</mods:number>
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<treatment id="E45287EFFFFF4314FCC0FE37819757D0" ID-DOI="http://doi.org/10.5281/zenodo.5492602" ID-Zenodo-Dep="5492602" LSID="urn:lsid:plazi:treatment:E45287EFFFFF4314FCC0FE37819757D0" httpUri="http://treatment.plazi.org/id/E45287EFFFFF4314FCC0FE37819757D0" lastPageId="12" lastPageNumber="449" pageId="10" pageNumber="447">
<subSubSection id="24E16572FFFF4312FCC0FE37877454DF" box="[850,1377,451,473]" pageId="10" pageNumber="447" type="nomenclature">
<paragraph id="6C4436F9FFFF4312FCC0FE37877454DF" blockId="10.[850,1377,451,473]" box="[850,1377,451,473]" pageId="10" pageNumber="447">
<heading id="370C8195FFFF4312FCC0FE37877454DF" box="[850,1377,451,473]" centered="true" fontSize="9" level="2" pageId="10" pageNumber="447" reason="2">
<taxonomicName id="ABFB4D7AFFFF4312FCC0FE37867B54DE" ID-CoL="6XQB9" authority="Wu &amp; Hwang &amp; Lin, 2008" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" box="[850,1134,451,472]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="447" phylum="Mollusca" rank="species" species="hagiomontis" status="sp. nov.">SATSUMA HAGIOMONTIS</taxonomicName>
<emphasis id="5E8FEAEBFFFF4312FBEAFE3286C854DE" bold="true" box="[1144,1245,451,472]" pageId="10" pageNumber="447">
<taxonomicNameLabel id="45BC5790FFFF4312FBEAFE3286C854DE" box="[1144,1245,451,472]" pageId="10" pageNumber="447" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="F4C02A7CFFFF4312FB7DFE37872954DF" box="[1263,1340,451,473]" captionStart="Figure 5" captionStartId="10.[144,223,1824,1843]" captionTargetBox="[369,1199,1076,1794]" captionTargetId="figure-318@10.[367,1199,1073,1794]" captionTargetPageId="10" captionText="Figure 5. Satsuma hagiomontis sp. nov. A–D, holotype (TMMT 0604, shell height = 17.7 mm). E, F, paratype (TMMT 0615, shell height = 14 mm). G, H, paratype (TMMT 0616, shell height = 14 mm). I, J, living specimens. Arrow indicates the pre-apertural constriction." figureDoi="http://doi.org/10.5281/zenodo.5446711" httpUri="https://zenodo.org/record/5446711/files/figure.png" pageId="10" pageNumber="447">FIGS 5</figureCitation>
,
<figureCitation id="F4C02A7CFFFF4312FAD9FE37874C54DF" box="[1355,1369,451,473]" captionStart="Figure 6" captionStartId="11.[164,243,1784,1803]" captionTargetBox="[167,1442,196,1751]" captionTargetId="figure-0@11.[164,1444,193,1754]" captionTargetPageId="11" captionText="Figure 6. Reproductive system of Satsuma hagiomontis sp. nov. (paratype, TMMT 0615). A, whole genitalia; B, interior of genitalia. Scale bar = 5 mm. See text for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5446713" httpUri="https://zenodo.org/record/5446713/files/figure.png" pageId="10" pageNumber="447">6</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="24E16572FFFF4314FCB4FE1E819757D0" lastPageId="12" lastPageNumber="449" pageId="10" pageNumber="447" type="materials_examined">
<paragraph id="6C4436F9FFFF4312FCB4FE1E81EB54F9" blockId="10.[806,1422,490,695]" box="[806,1022,490,511]" pageId="10" pageNumber="447">
<emphasis id="5E8FEAEBFFFF4312FCB4FE1E81EB54F9" box="[806,1022,490,511]" italics="true" pageId="10" pageNumber="447">Material examined</emphasis>
</paragraph>
<paragraph id="6C4436F9FFFF4312FCB4FDFD861557B1" blockId="10.[806,1422,490,695]" pageId="10" pageNumber="447">
<emphasis id="5E8FEAEBFFFF4312FCB4FDFD81CD5718" box="[806,984,521,542]" italics="true" pageId="10" pageNumber="447">Type specimen:</emphasis>
<materialsCitation id="DC933CA4FFFF4312FC7BFDFC861557B1" collectionCode="TMMT, BMNH, ANSP" pageId="10" pageNumber="447" specimenCode="TMMT 0604, TMMT 0615, TMMT 0616, BMNH 20060751, BMNH 20060752, ANSP 413674" specimenCount="2" typeStatus="holotype">
<typeStatus id="B340885BFFFF4312FC7BFDFC86465718" box="[1001,1107,520,542]" pageId="10" pageNumber="447" type="holotype">Holotype</typeStatus>
:
<specimenCode id="3C5D9E82FFFF4312FBF5FDFD86E85718" box="[1127,1277,520,542]" collectionCode="TMMT" pageId="10" pageNumber="447">TMMT 0604</specimenCode>
(from type locality, dry shell, tissue in alcohol). Five
<typeStatus id="B340885BFFFF4312FA87FDD3879C573A" box="[1301,1417,551,572]" pageId="10" pageNumber="447" type="paratype">paratypes</typeStatus>
: all from type locality,
<specimenCode id="3C5D9E82FFFF4312FB8CFDB286BE575D" box="[1054,1195,581,603]" collectionCode="TMMT" pageId="10" pageNumber="447">TMMT 0615</specimenCode>
(dry shell, tissue in alcohol, dissected);
<specimenCode id="3C5D9E82FFFF4312FB98FD90868E577C" box="[1034,1179,612,634]" collectionCode="TMMT" pageId="10" pageNumber="447">TMMT 0616</specimenCode>
(dry shell, tissue in alcohol);
<specimenCode id="3C5D9E82FFFF4312FC02FD778648579E" box="[912,1117,642,664]" collectionCode="BMNH" pageId="10" pageNumber="447">BMNH 20060751</specimenCode>
,
<specimenCode id="3C5D9E82FFFF4312FBFEFD77872C579E" box="[1132,1337,642,664]" collectionCode="BMNH" pageId="10" pageNumber="447">BMNH 20060752</specimenCode>
,
<specimenCode id="3C5D9E82FFFF4312FAD5FD77816D57B1" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="10" pageNumber="447">ANSP 413674</specimenCode>
(dry shell).
</materialsCitation>
</paragraph>
<paragraph id="6C4436F9FFFF4312FCB4FD0C81FE566F" blockId="10.[806,1421,760,874]" pageId="10" pageNumber="447">
<emphasis id="5E8FEAEBFFFF4312FCB4FD0C860D560B" box="[806,1048,760,781]" italics="true" pageId="10" pageNumber="447">Additional material:</emphasis>
<materialsCitation id="DC933CA4FFFF4312FBBAFD0C81F2566F" collectionCode="NTUM" country="Taiwan" location="southern Taiwan" municipality="Museum of Zoology" pageId="10" pageNumber="447" specimenCount="1" stateProvince="Pingtung">
<collectingMunicipality id="8C20AC83FFFF4312FBBAFD0C87035608" box="[1064,1302,760,782]" pageId="10" pageNumber="447">Museum of Zoology</collectingMunicipality>
, National
<collectingCountry id="14EC7669FFFF4312FCB4FCE3816F562B" box="[806,890,791,813]" name="Taiwan" pageId="10" pageNumber="447">Taiwan</collectingCountry>
University (
<collectionCode id="0AEAAE3CFFFF4312FB9CFCE38676562B" box="[1038,1123,791,813]" pageId="10" pageNumber="447">NTUM</collectionCode>
): two dry shells, collected from Kuwarusu (= Taiwu,
<collectingRegion id="AE3FF81BFFFF4312FBC4FCC1870E564C" box="[1110,1307,821,843]" country="Taiwan" name="Pingtung" pageId="10" pageNumber="447">Pingtung County</collectingRegion>
,
<location id="69246022FFFF4312FAB5FCC1816E566C" LSID="urn:lsid:plazi:treatment:E45287EFFFFF4314FCC0FE37819757D0:69246022FFFF4312FAB5FCC1816E566C" country="Taiwan" municipality="Museum of Zoology" name="southern Taiwan" pageId="10" pageNumber="447" stateProvince="Pingtung">southern Taiwan</location>
) in 1918
</materialsCitation>
.
</paragraph>
<paragraph id="6C4436F9FFFF4312FCB4FC5F81A256C6" blockId="10.[806,1421,939,1022]" box="[806,951,939,960]" pageId="10" pageNumber="447">
<emphasis id="5E8FEAEBFFFF4312FCB4FC5F81A256C6" box="[806,951,939,960]" italics="true" pageId="10" pageNumber="447">
<typeStatus id="B340885BFFFF4312FCB4FC5F814F56C6" box="[806,858,939,960]" pageId="10" pageNumber="447">Type</typeStatus>
locality
</emphasis>
</paragraph>
<paragraph id="6C4436F9FFFF4312FCB4FC3E81F756F8" blockId="10.[806,1421,939,1022]" pageId="10" pageNumber="447">
<materialsCitation id="DC933CA4FFFF4312FCB4FC3E81C856F8" collectorName="M. Beidawushan" country="Taiwan" location="Pingtung County" pageId="10" pageNumber="447" specimenCount="1" stateProvince="Pingtung" typeStatus="holotype">
<collectorName id="C10E532FFFFF4312FCB4FC3E81FE56D9" box="[806,1003,969,991]" pageId="10" pageNumber="447">M. Beidawushan</collectorName>
,
<collectingRegion id="AE3FF81BFFFF4312FC6AFC3D86A856D8" box="[1016,1213,969,991]" country="Taiwan" name="Pingtung" pageId="10" pageNumber="447">Pingtung County</collectingRegion>
, southern
<collectingCountry id="14EC7669FFFF4312FAABFC3D879856D9" box="[1337,1421,969,991]" name="Taiwan" pageId="10" pageNumber="447">Taiwan</collectingCountry>
(
<figureCitation id="F4C02A7CFFFF4312FCBCFC1C817A56FB" box="[814,879,1000,1022]" captionStart="Figure 1" captionStartId="1.[164,243,1882,1901]" captionTargetBox="[387,1219,874,1853]" captionTargetId="graphics-312@1.[386,1220,874,1853]" captionTargetPageId="1" captionText="Figure 1. Sampling localities of all taxa in Taiwan. Bold line, Central Range; thin line, administrative boundary." figureDoi="http://doi.org/10.5281/zenodo.5446702" httpUri="https://zenodo.org/record/5446702/files/figure.png" pageId="10" pageNumber="447">Fig. 1</figureCitation>
,
<tableCitation id="21790342FFFF4312FC12FC1C81C756F8" box="[896,978,1000,1022]" captionStart="Table 1" captionStartId="2.[144,208,200,219]" captionText="Table 1. List of the sampling localities showing altitude (metres) and WGS84 coordinate" pageId="10" pageNumber="447">Table 1</tableCitation>
)
</materialsCitation>
.
</paragraph>
<caption id="38846671FFFF4312FF02F8D4802F5268" ID-DOI="http://doi.org/10.5281/zenodo.5446711" ID-Zenodo-Dep="5446711" httpUri="https://zenodo.org/record/5446711/files/figure.png" pageId="10" pageNumber="447" startId="10.[144,223,1824,1843]" targetBox="[369,1199,1076,1794]" targetPageId="10">
<paragraph id="6C4436F9FFFF4312FF02F8D4802F5268" blockId="10.[144,1421,1823,1902]" pageId="10" pageNumber="447">
<emphasis id="5E8FEAEBFFFF4312FF02F8D4806D5235" bold="true" box="[144,632,1824,1843]" pageId="10" pageNumber="447">
Figure 5.
<taxonomicName id="ABFB4D7AFFFF4312FE96F8D4801E5235" authority="Wu &amp; Hwang &amp; Lin, 2008" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" box="[260,523,1824,1843]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="447" phylum="Mollusca" rank="species" species="hagiomontis" status="sp. nov.">
<emphasis id="5E8FEAEBFFFF4312FE96F8D4801E5235" bold="true" box="[260,523,1824,1843]" italics="true" pageId="10" pageNumber="447">Satsuma hagiomontis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45BC5790FFFF4312FD8BF8D4806D5235" box="[537,632,1824,1843]" pageId="10" pageNumber="447" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A–D, holotype (TMMT 0604, shell height = 17.7 mm). E, F, paratype (TMMT 0615, shell height = 14 mm). G, H, paratype (TMMT 0616, shell height = 14 mm). I, J, living specimens. Arrow indicates the pre-apertural constriction.
</paragraph>
</caption>
<caption id="38846671FFFE4313FF36F90C8193522E" ID-DOI="http://doi.org/10.5281/zenodo.5446713" ID-Zenodo-Dep="5446713" httpUri="https://zenodo.org/record/5446713/files/figure.png" pageId="11" pageNumber="448" startId="11.[164,243,1784,1803]" targetBox="[167,1442,196,1751]" targetPageId="11">
<paragraph id="6C4436F9FFFE4313FF36F90C8193522E" blockId="11.[164,1441,1784,1833]" pageId="11" pageNumber="448">
<emphasis id="5E8FEAEBFFFE4313FF36F90C8319520D" bold="true" box="[164,268,1784,1803]" pageId="11" pageNumber="448">Figure 6.</emphasis>
Reproductive system of
<emphasis id="5E8FEAEBFFFE4313FDB9F90C818A520D" bold="true" box="[555,927,1784,1803]" pageId="11" pageNumber="448">
<taxonomicName id="ABFB4D7AFFFE4313FDB9F90C8127520D" authority="Wu &amp; Hwang &amp; Lin, 2008" authorityName="Wu &amp; Hwang &amp; Lin" authorityYear="2008" box="[555,818,1784,1803]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="448" phylum="Mollusca" rank="species" species="hagiomontis" status="sp. nov.">
<emphasis id="5E8FEAEBFFFE4313FDB9F90C8127520D" bold="true" box="[555,818,1784,1803]" italics="true" pageId="11" pageNumber="448">Satsuma hagiomontis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45BC5790FFFE4313FCD2F90C818A520D" box="[832,927,1784,1803]" pageId="11" pageNumber="448" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(paratype, TMMT 0615). A, whole genitalia; B, interior of genitalia. Scale bar = 5 mm. See text for abbreviations.
</paragraph>
</caption>
<paragraph id="6C4436F9FFF94314FF02FF32831555DD" blockId="12.[144,759,198,373]" box="[144,256,198,219]" pageId="12" pageNumber="449">
<emphasis id="5E8FEAEBFFF94314FF02FF32831555DD" box="[144,256,198,219]" italics="true" pageId="12" pageNumber="449">Diagnosis</emphasis>
</paragraph>
<paragraph id="6C4436F9FFF94314FF02FF1082E35473" blockId="12.[144,759,198,373]" pageId="12" pageNumber="449">Shell and soft body white with grey pedal stripe; periphery keeled; shell height/diameter ratio close to 1.0; distal vagina short; penis corrugated on surface; penial caecum conical; genitalia with or without spinules.</paragraph>
<paragraph id="6C4436F9FFF94314FF02FE4F831254D6" blockId="12.[144,759,443,556]" box="[144,263,443,464]" pageId="12" pageNumber="449">
<emphasis id="5E8FEAEBFFF94314FF02FE4F831254D6" box="[144,263,443,464]" italics="true" pageId="12" pageNumber="449">Etymology</emphasis>
</paragraph>
<paragraph id="6C4436F9FFF94314FF02FE2E800A572A" blockId="12.[144,759,443,556]" pageId="12" pageNumber="449">
L.
<emphasis id="5E8FEAEBFFF94314FF3DFE2D82FB54E8" box="[175,238,473,494]" italics="true" pageId="12" pageNumber="449">hagio</emphasis>
: holy, L.
<emphasis id="5E8FEAEBFFF94314FECBFE2E83B354E9" box="[345,422,474,495]" italics="true" pageId="12" pageNumber="449">montis</emphasis>
: mountain. The
<typeStatus id="B340885BFFF94314FDF4FE2E808254E9" box="[614,663,474,495]" pageId="12" pageNumber="449">type</typeStatus>
locality, Mt Beidawushan, is a holy mountain of two aboriginal tribes, the Paiwan and Rukai.
</paragraph>
<paragraph id="6C4436F9FFF94314FF02FD878304578E" blockId="12.[144,759,627,985]" box="[144,273,627,648]" pageId="12" pageNumber="449">
<emphasis id="5E8FEAEBFFF94314FF02FD878304578E" box="[144,273,627,648]" italics="true" pageId="12" pageNumber="449">Description</emphasis>
</paragraph>
<paragraph id="6C4436F9FFF94314FF02FD6580A056DF" blockId="12.[144,759,627,985]" pageId="12" pageNumber="449">
<emphasis id="5E8FEAEBFFF94314FF02FD6582DE57A0" box="[144,203,657,678]" italics="true" pageId="12" pageNumber="449">Shell</emphasis>
(
<figureCitation id="F4C02A7CFFF94314FF71FD65833657A1" box="[227,291,657,679]" captionStart="Figure 5" captionStartId="10.[144,223,1824,1843]" captionTargetBox="[369,1199,1076,1794]" captionTargetId="figure-318@10.[367,1199,1073,1794]" captionTargetPageId="10" captionText="Figure 5. Satsuma hagiomontis sp. nov. A–D, holotype (TMMT 0604, shell height = 17.7 mm). E, F, paratype (TMMT 0615, shell height = 14 mm). G, H, paratype (TMMT 0616, shell height = 14 mm). I, J, living specimens. Arrow indicates the pre-apertural constriction." figureDoi="http://doi.org/10.5281/zenodo.5446711" httpUri="https://zenodo.org/record/5446711/files/figure.png" pageId="12" pageNumber="449">Fig. 5</figureCitation>
): dextral, conical, thin, fragile, semitranslucent, medium sized. Apex obtuse. Whorls inflated. Peripheries bluntly angulated with keel extended to peristome. Base inflated. Pre-apertural constriction behind outer and inferior lips present. Shell surface smooth, glossy, with spiral striae. Shell colour white milky with thin periostracum. Aperture diagonal, ovate-lunate. Peristome thin, expanded, reflected at curved inferior lip. Superior columellar lip vertical, reflected, covering most of umbilicus. Inferior columellar lip oblique, continual to inferior lip.
</paragraph>
<paragraph id="6C4436F9FFF94314FF02FBEB80DA51ED" blockId="12.[144,759,1055,1260]" pageId="12" pageNumber="449">
<emphasis id="5E8FEAEBFFF94314FF02FBEB83545132" box="[144,321,1055,1076]" italics="true" pageId="12" pageNumber="449">Band or stain:</emphasis>
Polymorphism exists in this taxon. Some individuals exhibit a red-brown to black-purple stain around the umbilicus, columellar, columellar lip, inferior lip and apex, others have spiral colour band around the whorls or at the base. The outer lip exhibits no coloration. Band formula = 1 0 0 0 0 0 7 8; 1 0 0 0 5 0 7 0; 1 0 0 4 0 0 7 0; 1 0 0 4 0 0 7 8.
</paragraph>
<paragraph id="6C4436F9FFF94314FF02FAC683F25269" blockId="12.[144,759,1330,1903]" pageId="12" pageNumber="449">
<emphasis id="5E8FEAEBFFF94314FF02FAC6836E504E" box="[144,379,1330,1352]" italics="true" pageId="12" pageNumber="449">Reproductive system</emphasis>
(
<figureCitation id="F4C02A7CFFF94314FE1DFAC683C5504E" box="[399,464,1330,1352]" captionStart="Figure 6" captionStartId="11.[164,243,1784,1803]" captionTargetBox="[167,1442,196,1751]" captionTargetId="figure-0@11.[164,1444,193,1754]" captionTargetPageId="11" captionText="Figure 6. Reproductive system of Satsuma hagiomontis sp. nov. (paratype, TMMT 0615). A, whole genitalia; B, interior of genitalia. Scale bar = 5 mm. See text for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.5446713" httpUri="https://zenodo.org/record/5446713/files/figure.png" pageId="12" pageNumber="449">Fig. 6</figureCitation>
): Bursa stalk long, with apparently expanded and conical base, regularly tapering towards oval bursa copulatrix. Proximal vagina muscular, swollen, smooth externally, with 14–17 strong, smooth folds internally. Middle vagina less muscular, distinctly constrictive, twisted. Distal vagina short, one-quarter of vagina in length. Flagellum long, tapering, not swollen at base. Pilaster in epiphallus wide, weak, three in number. Penial caecum long, depressed-conical. Cecal pilaster two in number, moderately weak, not strongly corrugated on surfaces. Principal pilaster and proximal penis absent. Middle penis long, twisted, depressed, with seven to nine moderately strong, wiggly, corrugated pilasters inside. Distal penis short, constrictive, with three wide and low pilasters vanishing towards atrium internally. Of the two dissected specimens one (
<collectionCode id="0AEAAE3CFFF94314FF05F8CE82FD5249" box="[151,232,1850,1871]" pageId="12" pageNumber="449">TMMT</collectionCode>
0615) has many spinules on the surface of the pilasters in the middle penis.
</paragraph>
<paragraph id="6C4436F9FFF94314FCB4FF3281A455DD" blockId="12.[806,1421,198,342]" box="[806,945,198,219]" pageId="12" pageNumber="449">
<emphasis id="5E8FEAEBFFF94314FCB4FF3281A455DD" box="[806,945,198,219]" italics="true" pageId="12" pageNumber="449">Distribution</emphasis>
</paragraph>
<paragraph id="6C4436F9FFF94314FCB4FF10819D5450" blockId="12.[806,1421,198,342]" pageId="12" pageNumber="449">
<materialsCitation id="DC933CA4FFF94314FCB4FF1081965450" country="Taiwan" elevation="1200" location="Mt Beidawushan" pageId="12" pageNumber="449" specimenCount="1" stateProvince="Pingtung">
The species was found only in a narrow area of mid-altitude (
<quantity id="AB039B1CFFF94314FC51FEF78649541E" box="[963,1116,259,280]" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.4" metricValueMin="1.0" pageId="12" pageNumber="449" unit="m" value="1200.0" valueMax="1400.0" valueMin="1000.0">
<elevation id="E7D6D1CAFFF94314FC51FEF78649541E" box="[963,1116,259,280]" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.4" metricValueMin="1.0" pageId="12" pageNumber="449" unit="m" value="1200.0" valueMax="1400.0" valueMin="1000.0">1000–1400 m</elevation>
</quantity>
) forest near
<location id="69246022FFF94314FB7CFEF78141543E" LSID="urn:lsid:plazi:treatment:E45287EFFFFF4314FCC0FE37819757D0:69246022FFF94314FB7CFEF78141543E" country="Taiwan" name="Mt Beidawushan" pageId="12" pageNumber="449" stateProvince="Pingtung">Mt Beidawushan</location>
,
<collectingRegion id="AE3FF81BFFF94314FCFCFED686555431" box="[878,1088,290,312]" country="Taiwan" name="Pingtung" pageId="12" pageNumber="449">Pingtung County</collectingRegion>
, southern
<collectingCountry id="14EC7669FFF94314FB46FED6873D543E" box="[1236,1320,290,312]" name="Taiwan" pageId="12" pageNumber="449">Taiwan</collectingCountry>
(
<figureCitation id="F4C02A7CFFF94314FAD7FED687935431" box="[1349,1414,290,312]" captionStart="Figure 1" captionStartId="1.[164,243,1882,1901]" captionTargetBox="[387,1219,874,1853]" captionTargetId="graphics-312@1.[386,1220,874,1853]" captionTargetPageId="1" captionText="Figure 1. Sampling localities of all taxa in Taiwan. Bold line, Central Range; thin line, administrative boundary." figureDoi="http://doi.org/10.5281/zenodo.5446702" httpUri="https://zenodo.org/record/5446702/files/figure.png" pageId="12" pageNumber="449">Fig. 1</figureCitation>
,
<tableCitation id="21790342FFF94314FCB4FEB4816D5450" box="[806,888,320,342]" captionStart="Table 1" captionStartId="2.[144,208,200,219]" captionText="Table 1. List of the sampling localities showing altitude (metres) and WGS84 coordinate" pageId="12" pageNumber="449">Table 1</tableCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="6C4436F9FFF94314FCB4FE7A819E54A5" blockId="12.[806,1422,398,726]" box="[806,907,398,419]" pageId="12" pageNumber="449">
<emphasis id="5E8FEAEBFFF94314FCB4FE7A819E54A5" box="[806,907,398,419]" italics="true" pageId="12" pageNumber="449">Remarks</emphasis>
</paragraph>
<paragraph id="6C4436F9FFF94314FCB4FE59819757D0" blockId="12.[806,1422,398,726]" pageId="12" pageNumber="449">
These arboreal herbivores are nocturnal, perching under leaves more than
<quantity id="AB039B1CFFF94314FBC4FE38869454E7" box="[1110,1153,460,481]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="449" unit="m" value="2.0">2 m</quantity>
above ground. Adults were found during summer (
<figureCitation id="F4C02A7CFFF94314FBEDFE1E86FF54F9" box="[1151,1258,490,512]" captionStart="Figure 5" captionStartId="10.[144,223,1824,1843]" captionTargetBox="[369,1199,1076,1794]" captionTargetId="figure-318@10.[367,1199,1073,1794]" captionTargetPageId="10" captionText="Figure 5. Satsuma hagiomontis sp. nov. A–D, holotype (TMMT 0604, shell height = 17.7 mm). E, F, paratype (TMMT 0615, shell height = 14 mm). G, H, paratype (TMMT 0616, shell height = 14 mm). I, J, living specimens. Arrow indicates the pre-apertural constriction." figureDoi="http://doi.org/10.5281/zenodo.5446711" httpUri="https://zenodo.org/record/5446711/files/figure.png" pageId="12" pageNumber="449">Fig. 5I, J</figureCitation>
). The species differs from
<taxonomicName id="ABFB4D7AFFF94314FC28FDFD86335718" authorityName=", Ohara &amp; Otani" authorityYear="2002" box="[954,1062,521,542]" class="Gastropoda" family="Camaenidae" genus="Satsuma" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="449" phylum="Mollusca" rank="species" species="albida">
<emphasis id="5E8FEAEBFFF94314FC28FDFD86335718" box="[954,1062,521,542]" italics="true" pageId="12" pageNumber="449">S. albida</emphasis>
</taxonomicName>
in having a keeled periphery, lower spire with shell height/diameter ratio close to 1.0, short distal vagina; corrugated penis externally, conical penial caecum and spinules inside genitalia. The conical penial caecum and strongly corrugated penis are unique among white-shelled species in west
<collectingCountry id="14EC7669FFF94314FCB4FD34816B57D0" box="[806,894,704,726]" name="Taiwan" pageId="12" pageNumber="449">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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