treatments-xml/data/C1/0A/55/C10A5544FF8CFFB61553A66FFD6FFDD2.xml

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<document id="941D78B775BBB9BF535D4C4B300DC64C" ID-CLB-Dataset="7100" ID-DOI="10.5852/ejt.2021.748.1335" ID-GBIF-Dataset="04bf840b-8ba9-4059-8190-f2454630a714" ID-ISSN="2118-9773" ID-Zenodo-Dep="4736129" ID-ZooBank="AB54CD6A-B757-4E5A-8EB3-B9B466B74639" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1620111095485" checkinUser="plazi" docAuthor="Sousa, Rafael &amp; Seidel, Matthias" docDate="2021" docId="C10A5544FF8CFFB61553A66FFD6FFDD2" docLanguage="en" docName="ejt-748_15-35.pdf" docOrigin="European Journal of Taxonomy 748 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.748.1335" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Inca LePeletier &amp; Serville 1828" docType="treatment" docVersion="9" lastPageNumber="21" masterDocId="3D332D3CFF8FFFB0177CA434F970FFE8" masterDocTitle="Review of the Inca irroratus species group with description of two new species of Inca LePeletier &amp; Serville, 1828 (Coleoptera, Scarabaeidae, Cetoniinae)" masterLastPageNumber="35" masterPageNumber="15" pageNumber="18" updateTime="1698938226785" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2A850F76DAAB5732C83DB26AA7DA69AF">Review of the Inca irroratus species group with description of two new species of Inca LePeletier &amp; Serville, 1828 (Coleoptera, Scarabaeidae, Cetoniinae)</mods:title>
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<mods:namePart id="3E41BF703D62FF4CA0BBCBC363CCF12F">Sousa, Rafael</mods:namePart>
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<mods:affiliation id="5241D5608BC81D20DFBDE3EE2DBEDD43">Museu de Zoologia da Universidade de São Paulo, Avenida Nazaré 481, Ipiranga, 04263 - 000 São Paulo, São Paulo, Brazil.</mods:affiliation>
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<mods:namePart id="6F0CA20CB55712F996F53D1C856D71A0">Seidel, Matthias</mods:namePart>
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<mods:affiliation id="BBC0D194406A7BC2A1991956CF2C97D1">Centrum für Naturkunde, University of Hamburg, Martin-Luther-King-Platz 3, 20146, Hamburg, Germany.</mods:affiliation>
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<heading id="1254533EFF8CFFB31553A66FFD74FD9D" box="[559,1028,602,629]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="18" reason="2">
Genus
<taxonomicName id="8EA39FD1FF8CFFB315FEA66FFD74FD9D" authority="LePeletier &amp; Serville, 1828" authorityName="LePeletier &amp; Serville" authorityYear="1828" box="[642,1028,602,629]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="7BD73840FF8CFFB315FEA66FFBCAFD9D" bold="true" box="[642,698,603,629]" italics="true" pageId="3" pageNumber="18">Inca</emphasis>
<bibRefCitation id="2D3299A3FF8CFFB315BDA66EFD74FD9D" author="Lepeletier A. L. M. &amp; Serville J. G. A." box="[705,1028,602,629]" pageId="3" pageNumber="18" pagination="345 - 833" refId="ref10104" refString="Lepeletier A. L. M. &amp; Serville J. G. A. 1828. Entomologie, ou histoire naturelle des crustaces, des arachnides et des insectes. Encyclopedie methodique, Histoire naturelle 10: 345 - 833." type="journal article" year="1828">LePeletier &amp; Serville, 1828</bibRefCitation>
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<paragraph id="491CE452FF8CFFB317C1A6AAFB0FFD5F" blockId="3.[189,639,669,695]" box="[189,639,669,695]" pageId="3" pageNumber="18">
<treatmentCitationGroup id="69B3C37CFF8CFFB317C1A6AAFB0FFD5F" box="[189,639,669,695]" pageId="3" pageNumber="18">
<treatmentCitation id="C802C243FF8CFFB317C1A6AAFB0BFD5F" author="Lepeletier A. L. M. &amp; Serville J. G. A." box="[189,635,669,695]" page="380" pageId="3" pageNumber="18" year="1828">
<taxonomicName id="8EA39FD1FF8CFFB317C1A6AAFB0BFD5F" authority="LePeletier &amp; Serville, 1828: 380" authorityName="LePeletier &amp; Serville" authorityPageNumber="380" authorityYear="1828" box="[189,635,669,695]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="7BD73840FF8CFFB317C1A6AAF982FD5F" box="[189,242,670,695]" italics="true" pageId="3" pageNumber="18">Inca</emphasis>
<bibRefCitation id="2D3299A3FF8CFFB31785A6A9FB0BFD5F" author="Lepeletier A. L. M. &amp; Serville J. G. A." box="[249,635,669,695]" pageId="3" pageNumber="18" pagination="345 - 833" refId="ref10104" refString="Lepeletier A. L. M. &amp; Serville J. G. A. 1828. Entomologie, ou histoire naturelle des crustaces, des arachnides et des insectes. Encyclopedie methodique, Histoire naturelle 10: 345 - 833." type="journal article" year="1828">LePeletier &amp; Serville, 1828: 380</bibRefCitation>
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.
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<paragraph id="491CE452FF8CFFB317C1A6D4FAE9FD12" blockId="3.[189,921,735,762]" box="[189,921,735,762]" pageId="3" pageNumber="18">
<treatmentCitationGroup id="69B3C37CFF8CFFB317C1A6D4FAE5FD12" box="[189,917,735,762]" pageId="3" pageNumber="18">
<taxonomicName id="8EA39FD1FF8CFFB317C1A6D4F988FD11" ID-CoL="89WN" box="[189,248,736,761]" class="Insecta" family="Scarabaeidae" genus="Ynca" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="7BD73840FF8CFFB317C1A6D4F988FD11" box="[189,248,736,761]" italics="true" pageId="3" pageNumber="18">Ynca</emphasis>
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<treatmentCitation id="C802C243FF8CFFB3166AA6EBFAE5FD12" author="Chevrolat " box="[278,917,735,762]" pageId="3" pageNumber="18" year="1833">
<bibRefCitation id="2D3299A3FF8CFFB3166AA6EBF8A0FD11" author="Chevrolat A." box="[278,464,735,761]" pageId="3" pageNumber="18" refId="ref9632" refString="Chevrolat A. 1833. Description d'especes nouvelles. Revue Entomologique - Coleopteres 10: unpaginated. Available from https: // www. biodiversitylibrary. org / item / 44561 # page / 349 / mode / 1 up [accessed 6 Apr. 2021]." type="journal volume" year="1833">Chevrolat 1833</bibRefCitation>
: unpaginated (unjustified emendation)
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.
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<treatmentCitationGroup id="69B3C37CFF8CFFB317C1A716FA22FCD4" box="[189,850,801,828]" pageId="3" pageNumber="18">
<taxonomicName id="8EA39FD1FF8CFFB317C1A716F982FCD3" authorityName="LePeletier &amp; Serville" authorityYear="1828" box="[189,242,802,827]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="7BD73840FF8CFFB317C1A716F982FCD3" box="[189,242,802,827]" italics="true" pageId="3" pageNumber="18">Inca</emphasis>
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<treatmentCitation id="C802C243FF8CFFB31673A715FB69FCD4" ID-DOI="http://doi.org/10.5281/zenodo.5062231" ID-Zenodo-Dep="5062231" author="Seidel M. &amp; Arriaga-Varela E. &amp; Sousa R." box="[271,537,801,828]" httpUri="http://treatment.plazi.org/id/03B787A5487EFC46A56805068F2FD026" page="389" pageId="3" pageNumber="18" year="2018">
<bibRefCitation id="2D3299A3FF8CFFB31673A715FB69FCD4" author="Seidel M. &amp; Arriaga-Varela E. &amp; Sousa R." box="[271,537,801,828]" pageId="3" pageNumber="18" pagination="389 - 405" refId="ref10467" refString="Seidel M., Arriaga-Varela E. &amp; Sousa R. 2018. Catalogue of the Incini with the description of the first Archedinus from Honduras (Coleoptera: Scarabaeidae: Cetoniinae). Acta Entomologica Musei Nationalis Pragae 58 (2): 389 - 405." type="journal article" year="2018">
Seidel
<emphasis id="7BD73840FF8CFFB3161CA716F8EBFCD3" box="[352,411,801,827]" italics="true" pageId="3" pageNumber="18">et al.</emphasis>
2018: 389
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(checklist, nomenclature).
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<heading id="1254533EFF8CFFB317C1A750F829FC96" bold="true" box="[189,345,868,894]" fontSize="11" level="3" pageId="3" pageNumber="18" reason="3">
<emphasis id="7BD73840FF8CFFB317C1A750F829FC96" bold="true" box="[189,345,868,894]" pageId="3" pageNumber="18">
<typeStatus id="96185AF0FF8CFFB317C1A750F98BFC96" box="[189,251,868,894]" pageId="3" pageNumber="18">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="491CE452FF8CFFB317C1A7A0FAEBFC46" blockId="3.[189,923,915,942]" box="[189,923,915,942]" pageId="3" pageNumber="18">
<treatmentCitationGroup id="69B3C37CFF8CFFB317C1A7A0FAEBFC46" box="[189,923,915,942]" pageId="3" pageNumber="18">
<taxonomicName id="8EA39FD1FF8CFFB317C1A7A0F888FC46" authority="Weber, 1801" authorityName="Weber" authorityYear="1801" box="[189,504,915,942]" class="Insecta" family="Cetoniidae" genus="Cetonia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="ynca">
<emphasis id="7BD73840FF8CFFB317C1A7A0F82AFC45" box="[189,346,916,942]" italics="true" pageId="3" pageNumber="18">Cetonia ynca</emphasis>
Weber, 1801
</taxonomicName>
, designated by
<treatmentCitation id="C802C243FF8CFFB315D2A7A7FAE7FC46" author="Howden H. F." box="[686,919,915,942]" page="12" pageId="3" pageNumber="18" year="1968">
<bibRefCitation id="2D3299A3FF8CFFB315D2A7A7FAE7FC46" author="Howden H. F." box="[686,919,915,942]" pageId="3" pageNumber="18" pagination="5 - 77" refId="ref9912" refString="Howden H. F. 1968. A review of the Trichiinae of North and Central America (Coleoptera: Scarabaeidae). The Memoirs of the Entomological Society of Canada 100 (S 54): 5 - 77." type="journal article" year="1968">Howden (1968: 12)</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="01B9B7D9FF8CFFB317C1A7E1FB88FB58" pageId="3" pageNumber="18" type="diagnosis">
<paragraph id="491CE452FF8CFFB317C1A7E1F847FC07" blockId="3.[189,311,981,1007]" box="[189,311,981,1007]" pageId="3" pageNumber="18">
<heading id="1254533EFF8CFFB317C1A7E1F847FC07" bold="true" box="[189,311,981,1007]" fontSize="11" level="3" pageId="3" pageNumber="18" reason="3">
<emphasis id="7BD73840FF8CFFB317C1A7E1F847FC07" bold="true" box="[189,311,981,1007]" pageId="3" pageNumber="18">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="491CE452FF8CFFB317C1A031FB88FB58" blockId="3.[188,1399,1029,1200]" pageId="3" pageNumber="18">
The genus can be distinguished from other genera of
<taxonomicName id="8EA39FD1FF8CFFB31457A031FA1FFBF7" authorityName="Burmeister" authorityYear="1842" box="[811,879,1029,1055]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="tribe" tribe="Incini">Incini</taxonomicName>
by the following combination of characters: dorsal surface of pronotum covered with waxy velvet like secretion and setigerous punctures with waxy macula (waxy secretion absent in
<taxonomicName id="8EA39FD1FF8CFFB31531A079FBA3FB8F" authorityName="Moron &amp; Krikken" authorityYear="1990" box="[589,723,1101,1127]" class="Insecta" family="Cetoniidae" genus="Archedinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="7BD73840FF8CFFB31531A079FBA3FB8F" box="[589,723,1101,1127]" italics="true" pageId="3" pageNumber="18">Archedinus</emphasis>
</taxonomicName>
,
<taxonomicName id="8EA39FD1FF8CFFB3159EA079FA03FB8F" authorityName="Burmeister" authorityYear="1842" box="[738,883,1101,1127]" class="Insecta" family="Cetoniidae" genus="Coelocratus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="7BD73840FF8CFFB3159EA079FA03FB8F" box="[738,883,1101,1127]" italics="true" pageId="3" pageNumber="18">Coelocratus</emphasis>
</taxonomicName>
,
<taxonomicName id="8EA39FD1FF8CFFB314FEA079FA91FB8F" authorityName="Thomson" authorityYear="1878" box="[898,993,1101,1127]" class="Insecta" family="Cetoniidae" genus="Golinca" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="7BD73840FF8CFFB314FEA079FA91FB8F" box="[898,993,1101,1127]" italics="true" pageId="3" pageNumber="18">Golinca</emphasis>
</taxonomicName>
and
<taxonomicName id="8EA39FD1FF8CFFB31360A079FDD3FB8F" authorityName="Burmeister" authorityYear="1847" box="[1052,1187,1101,1127]" class="Insecta" family="Cetoniidae" genus="Pantodinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="7BD73840FF8CFFB31360A079FDD3FB8F" box="[1052,1187,1101,1127]" italics="true" pageId="3" pageNumber="18">Pantodinus</emphasis>
</taxonomicName>
); clypeus of male with divergent lateral horns (parallel horns in
<taxonomicName id="8EA39FD1FF8CFFB315B7A046FA5AFB64" authorityName="Thomson" authorityYear="1878" box="[715,810,1138,1164]" class="Insecta" family="Cetoniidae" genus="Golinca" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="7BD73840FF8CFFB315B7A046FA5AFB64" box="[715,810,1138,1164]" italics="true" pageId="3" pageNumber="18">Golinca</emphasis>
</taxonomicName>
; median horn in
<taxonomicName id="8EA39FD1FF8CFFB31491A046FD04FB64" authorityName="Burmeister" authorityYear="1847" box="[1005,1140,1138,1164]" class="Insecta" family="Cetoniidae" genus="Pantodinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="7BD73840FF8CFFB31491A046FD04FB64" box="[1005,1140,1138,1164]" italics="true" pageId="3" pageNumber="18">Pantodinus</emphasis>
</taxonomicName>
; absent in
<taxonomicName id="8EA39FD1FF8CFFB31393A046FC05FB64" authorityName="Moron &amp; Krikken" authorityYear="1990" box="[1263,1397,1138,1164]" class="Insecta" family="Cetoniidae" genus="Archedinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="7BD73840FF8CFFB31393A046FC05FB64" box="[1263,1397,1138,1164]" italics="true" pageId="3" pageNumber="18">Archedinus</emphasis>
</taxonomicName>
and
<taxonomicName id="8EA39FD1FF8CFFB31793A0A2F8F0FB58" authorityName="Burmeister" authorityYear="1842" box="[239,384,1174,1200]" class="Insecta" family="Cetoniidae" genus="Coelocratus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="7BD73840FF8CFFB31793A0A2F8F0FB58" box="[239,384,1174,1200]" italics="true" pageId="3" pageNumber="18">Coelocratus</emphasis>
</taxonomicName>
) with inner area densely setose.
</paragraph>
</subSubSection>
<subSubSection id="01B9B7D9FF8CFFB417C1A0ECF86BFC6F" lastPageId="4" lastPageNumber="19" pageId="3" pageNumber="18" type="description">
<paragraph id="491CE452FF8CFFB317C1A0ECF81DFB1A" blockId="3.[189,365,1240,1266]" box="[189,365,1240,1266]" pageId="3" pageNumber="18">
<heading id="1254533EFF8CFFB317C1A0ECF81DFB1A" bold="true" box="[189,365,1240,1266]" fontSize="11" level="3" pageId="3" pageNumber="18" reason="3">
<emphasis id="7BD73840FF8CFFB317C1A0ECF81DFB1A" bold="true" box="[189,365,1240,1266]" pageId="3" pageNumber="18">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="491CE452FF8CFFB31796A13CF85AFACA" blockId="3.[189,1399,1288,1422]" box="[234,298,1288,1314]" pageId="3" pageNumber="18">
<emphasis id="7BD73840FF8CFFB31796A13CF85AFACA" bold="true" box="[234,298,1288,1314]" pageId="3" pageNumber="18">Male</emphasis>
</paragraph>
<paragraph id="491CE452FF8CFFB317C1A118FB65FA66" blockId="3.[189,1399,1288,1422]" pageId="3" pageNumber="18">
BODY. Length including the clypeal horns
<quantity id="8E5B49B7FF8CFFB315B4A118FA3FFAAE" box="[712,847,1324,1350]" metricMagnitude="-2" metricUnit="m" metricValue="4.15" metricValueMax="6.0" metricValueMin="2.3" pageId="3" pageNumber="18" unit="mm" value="41.5" valueMax="60.0" valueMin="23.0">2360 mm</quantity>
; width across humeri
<quantity id="8E5B49B7FF8CFFB3131EA118FD99FAAE" box="[1122,1257,1324,1350]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="2.1" metricValueMin="1.0" pageId="3" pageNumber="18" unit="mm" value="15.5" valueMax="21.0" valueMin="10.0">1021 mm</quantity>
. Surface of pronotum, scutellar shield, elytra and pygidium covered with waxy, velvet like secretion and setigerous punctures with waxy macula.
</paragraph>
<paragraph id="491CE452FF8CFFB317C1A183FBC4F8FE" blockId="3.[188,1400,1462,1814]" pageId="3" pageNumber="18">HEAD. Surface of frons with waxy secretion on disc and setigerous and heterogeneous punctures with light yellow waxy macula; short yellowish setae near to supraocular area; lateral portion of epistomal suture distinct near to canthus. Canthus angulate. Clypeus densely punctate with posterior dorsal margin projecting forward; distal area with divergent lateral horns slightly projecting forward and upward; inner area of horns densely setose, setae long and yellow. Labrum transverse. Mandibles with molar lobe rounded and striated; right molar lobe convex; left molar lobe concave. Maxillae setose; cardo with proximal area densely punctate with long yellow setae; galea inwards projected and with brushlike setae. Hypopharynx with long setae. Labium transverse, deeply and densely punctate; punctures with long yellow setae; anterior margin of mentum emarginated. Antennae with 10 antennomeres; club fusiform composed of three antennomeres.</paragraph>
<paragraph id="491CE452FF8CFFB317C1A30AFAE4F800" blockId="3.[189,1399,1854,2024]" pageId="3" pageNumber="18">
THORAX. Pronotal surface convex, disc with a longitudinal groove; densely punctate and with short yellow setae; lateral margin sinuated and irregularly crenulated. Hypomeron densely punctate; anterior area with irregular striated punctures. Anterior prosternal process projected (except in
<taxonomicName id="8EA39FD1FF8CFFB313AFA3B3FC31F848" authorityName="Burmeister &amp; Schaum" authorityYear="1840" box="[1235,1345,1926,1953]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="besckii">
<emphasis id="7BD73840FF8CFFB313AFA3B3FDADF848" box="[1235,1245,1927,1952]" italics="true" pageId="3" pageNumber="18">I</emphasis>
.
<emphasis id="7BD73840FF8CFFB31393A3B2FC31F848" box="[1263,1345,1926,1952]" italics="true" pageId="3" pageNumber="18">besckii</emphasis>
</taxonomicName>
and
<taxonomicName id="8EA39FD1FF8CFFB317C1A39FF821F82C" baseAuthorityName="Gyllenhal" baseAuthorityYear="1817" box="[189,337,1962,1989]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="bonplandi">
<emphasis id="7BD73840FF8CFFB317C1A39FF9B7F82C" box="[189,199,1963,1988]" italics="true" pageId="3" pageNumber="18">I</emphasis>
.
<emphasis id="7BD73840FF8CFFB317A5A39EF821F82C" box="[217,337,1962,1988]" italics="true" pageId="3" pageNumber="18">bonplandi</emphasis>
</taxonomicName>
). Metaventrite densely setose except in the discrimen area. Elytra densely punctate and maculate; elytral striae barely distinct; distal angle acuminate.
</paragraph>
<paragraph id="491CE452FF8BFFB417C1A53DFB14FE79" blockId="4.[189,1399,265,401]" pageId="4" pageNumber="19">LEGS. Profemur with an inner distal tooth. Protibia with a posterior distal tooth and an inner proximal emargination; spur short. Protarsomeres IIIV with inner striae. Mesotibia with one short and bifurcate spur; transversal carina I with stout setae. Outer posterior angle of metacoxa rounded. Metatibia with transversal carina I with stout setae.</paragraph>
<paragraph id="491CE452FF8BFFB417C1A58DFB05FDF4" blockId="4.[189,1400,441,540]" pageId="4" pageNumber="19">ABDOMEN. Spiracle VII placed near to anterior margin of sternite. Sternite VIII (last exposed sternite) with a rounded fovea on anteriomedian area. Propygidium totally or partially covered by resting elytra. Pygidium posterior margin parabolic.</paragraph>
<paragraph id="491CE452FF8BFFB417C1A671F8D7FD6C" blockId="4.[189,1399,581,644]" pageId="4" pageNumber="19">TERMINALIA. Aedeagus: apodeme of phallobase larger than phallobase. Parameres glabrous and with an outer distal process.</paragraph>
<paragraph id="491CE452FF8BFFB417C1A698F869FD2E" blockId="4.[189,281,684,710]" box="[189,281,684,710]" pageId="4" pageNumber="19">
<heading id="1254533EFF8BFFB417C1A698F869FD2E" bold="true" box="[189,281,684,710]" fontSize="11" level="3" pageId="4" pageNumber="19" reason="3">
<emphasis id="7BD73840FF8BFFB417C1A698F869FD2E" bold="true" box="[189,281,684,710]" pageId="4" pageNumber="19">Female</emphasis>
</heading>
</paragraph>
<paragraph id="491CE452FF8BFFB417C1A6E8F86BFC6F" blockId="4.[189,1399,731,903]" pageId="4" pageNumber="19">
BODY. Length
<quantity id="8E5B49B7FF8BFFB41614A6EFF89AFD1E" box="[360,490,731,758]" metricMagnitude="-2" metricUnit="m" metricValue="3.85" metricValueMax="5.0" metricValueMin="2.7" pageId="4" pageNumber="19" unit="mm" value="38.5" valueMax="50.0" valueMin="27.0">2750 mm</quantity>
; width across humeri
<quantity id="8E5B49B7FF8BFFB41591A6E8FA1FFD1E" box="[749,879,732,758]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.7" metricValueMin="1.0" pageId="4" pageNumber="19" unit="mm" value="13.5" valueMax="17.0" valueMin="10.0">1017 mm</quantity>
. The morphology of females of
<taxonomicName id="8EA39FD1FF8BFFB41397A6E8FC50FD1D" authorityName="LePeletier &amp; Serville" authorityYear="1828" box="[1259,1312,732,757]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="7BD73840FF8BFFB41397A6E8FC50FD1D" box="[1259,1312,732,757]" italics="true" pageId="4" pageNumber="19">Inca</emphasis>
</taxonomicName>
is very similar to the male, except by: posterior area of frons with waxy secretion. Clypeus without horns; anterior angles rounded; frontal area elongated. Outer teeth of protibia larger than in males. Transversal carinae I of meso- and metatibia well-marked; mesotibia with two long and simple spurs. Paraproct distinct.
</paragraph>
</subSubSection>
<subSubSection id="01B9B7D9FF8BFFB417C1A784FC1BFB8F" pageId="4" pageNumber="19" type="discussion">
<paragraph id="491CE452FF8BFFB417C1A784F840FC22" blockId="4.[189,304,944,970]" box="[189,304,944,970]" pageId="4" pageNumber="19">
<heading id="1254533EFF8BFFB417C1A784F840FC22" bold="true" box="[189,304,944,970]" fontSize="11" level="3" pageId="4" pageNumber="19" reason="3">
<emphasis id="7BD73840FF8BFFB417C1A784F840FC22" bold="true" box="[189,304,944,970]" pageId="4" pageNumber="19">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="491CE452FF8BFFB417C1A7D4FC1BFB8F" blockId="4.[189,1399,992,1128]" pageId="4" pageNumber="19">
As observed in some works (e.g.,
<bibRefCitation id="2D3299A3FF8BFFB4153FA7D4FB8EFC12" author="Ricchiardi E." box="[579,766,992,1018]" pageId="4" pageNumber="19" pagination="3 - 7" refId="ref10356" refString="Ricchiardi E. 2002. Notes on the genus Coelocratus Burmeister, 1842 (Coleoptera: Scarabaeidae). Cetoniimania 2 (1): 3 - 7." type="journal article" year="2002">Ricchiardi 2002</bibRefCitation>
;
<bibRefCitation id="2D3299A3FF8BFFB41472A7D4FD13FC12" author="Moron M. A. &amp; Vaz-de-Mello F. Z." box="[782,1123,992,1018]" pageId="4" pageNumber="19" pagination="110 - 119" refId="ref10291" refString="Moron M. A. &amp; Vaz-de-Mello F. Z. 2007. A new species of Archedinus (Coleoptera: Scarabaeoidea: Trichiinae) from Oaxaca, Mexico. The Pan-Pacific Entomologist 83: 110 - 119." type="journal article" year="2007">Morón &amp; Vaz-de-Mello 2007</bibRefCitation>
;
<bibRefCitation id="2D3299A3FF8BFFB4130DA7D4FC4CFC12" author="Seidel M. &amp; Arriaga-Varela E. &amp; Sousa R." box="[1137,1340,992,1018]" pageId="4" pageNumber="19" pagination="389 - 405" refId="ref10467" refString="Seidel M., Arriaga-Varela E. &amp; Sousa R. 2018. Catalogue of the Incini with the description of the first Archedinus from Honduras (Coleoptera: Scarabaeidae: Cetoniinae). Acta Entomologica Musei Nationalis Pragae 58 (2): 389 - 405." type="journal article" year="2018">
Seidel
<emphasis id="7BD73840FF8BFFB413BCA7D5FD8AFC12" box="[1216,1274,992,1018]" italics="true" pageId="4" pageNumber="19">et al.</emphasis>
2018
</bibRefCitation>
), the species of
<taxonomicName id="8EA39FD1FF8BFFB4164BA031F81CFBF6" authorityName="LePeletier &amp; Serville" authorityYear="1828" box="[311,364,1029,1054]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="7BD73840FF8BFFB4164BA031F81CFBF6" box="[311,364,1029,1054]" italics="true" pageId="4" pageNumber="19">Inca</emphasis>
</taxonomicName>
can be distinguished from the other members of the tribe
<taxonomicName id="8EA39FD1FF8BFFB4136CA030FD24FBF6" authorityName="Burmeister" authorityYear="1842" box="[1040,1108,1028,1054]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="tribe" tribe="Incini">Incini</taxonomicName>
mainly by the male with divergent lateral horns with internal face densely setose with long setae and elytra with yellow waxy maculae. In the brachycerous males the outer angles of clypeal horn can be indistinct in some species.
</paragraph>
</subSubSection>
<subSubSection id="01B9B7D9FF8BFFB417C1A0A4FACBFB32" box="[189,955,1168,1242]" pageId="4" pageNumber="19" type="distribution">
<paragraph id="491CE452FF8BFFB417C1A0A4F828FB42" blockId="4.[189,344,1168,1194]" box="[189,344,1168,1194]" pageId="4" pageNumber="19">
<heading id="1254533EFF8BFFB417C1A0A4F828FB42" bold="true" box="[189,344,1168,1194]" fontSize="11" level="3" pageId="4" pageNumber="19" reason="3">
<emphasis id="7BD73840FF8BFFB417C1A0A4F828FB42" bold="true" box="[189,344,1168,1194]" pageId="4" pageNumber="19">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="491CE452FF8BFFB417C1A0F4FACBFB32" blockId="4.[189,955,1216,1242]" box="[189,955,1216,1242]" pageId="4" pageNumber="19">
From Tamaulipas in Mexico to
<collectingCountry id="31B4A4C2FF8BFFB41553A0F4FBECFB32" box="[559,668,1216,1242]" name="Paraguay" pageId="4" pageNumber="19">Paraguay</collectingCountry>
and northern
<collectingCountry id="31B4A4C2FF8BFFB41442A0F4FAC7FB32" box="[830,951,1216,1242]" name="Argentina" pageId="4" pageNumber="19">Argentina</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="01B9B7D9FF8BFFB417C1A137F88EFAF5" box="[189,510,1282,1309]" pageId="4" pageNumber="19" type="discussion">
<paragraph id="491CE452FF8BFFB417C1A137F88EFAF5" blockId="4.[189,510,1282,1309]" box="[189,510,1282,1309]" pageId="4" pageNumber="19">
<heading id="1254533EFF8BFFB417C1A137F88EFAF5" bold="true" box="[189,510,1282,1309]" fontSize="11" level="3" pageId="4" pageNumber="19" reason="3">
<emphasis id="7BD73840FF8BFFB417C1A137F88EFAF5" bold="true" box="[189,510,1282,1309]" pageId="4" pageNumber="19">
Key to the species of
<taxonomicName id="8EA39FD1FF8BFFB416BAA137F88EFAF5" authorityName="LePeletier &amp; Serville" authorityYear="1828" box="[454,510,1283,1309]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="7BD73840FF8BFFB416BAA137F88EFAF5" bold="true" box="[454,510,1283,1309]" italics="true" pageId="4" pageNumber="19">Inca</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="01B9B7D9FF8BFFB617C1A107FC06FE82" lastPageId="6" lastPageNumber="21" pageId="4" pageNumber="19" type="key">
<key id="3D32B363FF8BFFB617C1A107FC06FE82" lastPageId="6" lastPageNumber="21" pageId="4" pageNumber="19">
<keyStep id="F257FCF7FF8BFFB417C1A107FC06FA99" pageId="4" pageNumber="19">
<paragraph id="491CE452FF8BFFB417C1A107FC07FAA5" blockId="4.[189,1399,1330,2025]" box="[189,1399,1330,1357]" pageId="4" pageNumber="19">
<keyLead id="F2524767FF8BFFB417C1A107FC07FAA5" box="[189,1399,1330,1357]" pageId="4" pageNumber="19">1. Clypeal horn present (male)............................................................................................................... 2</keyLead>
</paragraph>
<paragraph id="491CE452FF8BFFB417BDA163FC06FA99" blockId="4.[189,1399,1330,2025]" box="[193,1398,1367,1394]" pageId="4" pageNumber="19">
<keyLead id="F2524767FF8BFFB417BDA163FC06FA99" box="[193,1398,1367,1394]" pageId="4" pageNumber="19"> Clypeal horn absent (female)........................................................................................................... 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="F257FCF7FF8BFFB417C1A1AEFC07FA30" pageId="4" pageNumber="19">
<paragraph id="491CE452FF8BFFB417C1A1AEFC07FA5C" blockId="4.[189,1399,1330,2025]" box="[189,1399,1433,1460]" pageId="4" pageNumber="19">
<keyLead id="F2524767FF8BFFB417C1A1AEFC07FA5C" box="[189,1399,1433,1460]" pageId="4" pageNumber="19">2. Dorsolateral margin of clypeal horn with a long tooth...................................................................... 3</keyLead>
</paragraph>
<paragraph id="491CE452FF8BFFB417BDA18BFC07FA30" blockId="4.[189,1399,1330,2025]" box="[193,1399,1470,1497]" pageId="4" pageNumber="19">
<keyLead id="F2524767FF8BFFB417BDA18BFC07FA30" box="[193,1399,1470,1497]" pageId="4" pageNumber="19"> Dorsolateral margin of clypeal horn without a long tooth................................................................. 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="F257FCF7FF8BFFB417C1A235FC07F93A" pageId="4" pageNumber="19">
<paragraph id="491CE452FF8BFFB417C1A235FC07F98C" blockId="4.[189,1399,1330,2025]" pageId="4" pageNumber="19">
<keyLead id="F2524767FF8BFFB417C1A235FC07F98C" pageId="4" pageNumber="19">
3. Inner dorsal carina of clypeal horn abruptly interrupted at apex. Elytral disc with large waxy maculae. Outer distal process of parameres long and acuminate.
<collectingCountry id="31B4A4C2FF8BFFB4141DA211FAD9F9D7" box="[865,937,1573,1599]" name="Brazil" pageId="4" pageNumber="19">Brazil</collectingCountry>
(
<collectingRegion id="8B672AB0FF8BFFB414CBA211FD40F9D7" box="[951,1072,1573,1599]" country="Brazil" name="Sao Paulo" pageId="4" pageNumber="19">São Paulo</collectingRegion>
,
<collectingRegion id="8B672AB0FF8BFFB41346A211FDFAF9D7" box="[1082,1162,1573,1599]" country="Brazil" name="Parana" pageId="4" pageNumber="19">Paraná</collectingRegion>
and
<collectingRegion id="8B672AB0FF8BFFB413BCA211FC1CF9A8" box="[1216,1388,1573,1600]" country="Brazil" name="Santa Catarina" pageId="4" pageNumber="19">
Santa
<taxonomicName id="8EA39FD1FF8BFFB4127BA211FC1CF9A8" ID-CoL="94MGZ" authorityName="Vojnits &amp; de Laever" authorityYear="1973" box="[1287,1388,1573,1600]" class="Insecta" family="Geometridae" genus="Catarina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="22" phylum="Arthropoda" rank="genus">Catarina</taxonomicName>
</collectingRegion>
) (
<figureCitation id="D198F8D7FF8BFFB4178DA27EF8E0F98C" box="[241,400,1610,1636]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[265,1323,265,1849]" captionTargetId="figure-17@8.[265,1323,265,1849]" captionTargetPageId="8" captionText="Fig. 1. Males of Inca spp. AE, P. Inca axeli sp. nov., holotype (MZSP 22036). FJ, Q. Inca neglectus sp. nov., paratype (MZSP). KO, R. Inca irroratus Chevrolat, 1833, neotype (BMNH). A, F, K. Dorsal habitus. B, G, L. Ventral habitus. C, H, M. Lateral habitus. D, I, N. Dorsal view of aedeagus. E, J, O. Lateral view of aedeagus. PR. Dorsal view of head. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4736133" httpUri="https://zenodo.org/record/4736133/files/figure.png" pageId="4" pageNumber="19">Figs 1AE, P</figureCitation>
,
<figureCitation id="D198F8D7FF8BFFB416E7A27EF8B0F98C" box="[411,448,1610,1636]" captionStart="Fig" captionStartId="14.[189,232,1823,1849]" captionTargetBox="[201,1387,266,1779]" captionTargetId="figure-125@14.[200,1387,265,1779]" captionTargetPageId="14" captionText="Fig. 4. Images of liveInca spp.A. Inca axelisp. nov. (photo by R. Barros and R. Koike).B. Inca neglectus sp. nov. (photo by N. Marchito). C. Inca clathratus sommeri Westwood, 1844 (photo by A. González Gallina). D. Inca clathratus clathratus (Olivier, 1792) (photo by R. Barros and R. Koike). E. Inca clathratus quesneli Boos &amp; Ratcliffe, 1985 (photo by M. Hulme). F. Inca pulverulentus (Olivier, 1789) mating (photo by R. Barros and R. Koike). G. Inca bonplandi (Gyllenhal, 1817) (photo by J. Touroult). H. Inca besckii Burmeister &amp; Schaum, 1840 feeding on tree sap (photo by N. Wisnik)." figureDoi="http://doi.org/10.5281/zenodo.4736143" httpUri="https://zenodo.org/record/4736143/files/figure.png" pageId="4" pageNumber="19">4A</figureCitation>
).......................................................................................................
<taxonomicName id="8EA39FD1FF8BFFB413BAA27EFC6CF98C" authorityName="Sousa &amp; Seidel" authorityYear="2021" box="[1222,1308,1609,1636]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="axeli">
<emphasis id="7BD73840FF8BFFB413BAA27EFC6CF98C" bold="true" box="[1222,1308,1609,1636]" pageId="4" pageNumber="19">
<emphasis id="7BD73840FF8BFFB413BAA27EFDA1F98C" bold="true" box="[1222,1233,1610,1636]" italics="true" pageId="4" pageNumber="19">I</emphasis>
.
<emphasis id="7BD73840FF8BFFB4139DA27DFC6CF98C" bold="true" box="[1249,1308,1609,1636]" italics="true" pageId="4" pageNumber="19">axeli</emphasis>
</emphasis>
</taxonomicName>
sp nov.
</keyLead>
</paragraph>
<paragraph id="491CE452FF8BFFB417BDA25BFC07F93A" blockId="4.[189,1399,1330,2025]" pageId="4" pageNumber="19">
<keyLead id="F2524767FF8BFFB417BDA25BFC07F93A" pageId="4" pageNumber="19">
Inner dorsal carina of clypeal horn gradually interrupted at apeX. Elytral disc without large waXy maculae. Outer distal process of parameres long and rounded.
<collectingCountry id="31B4A4C2FF8BFFB414B6A2A7FD62F945" box="[970,1042,1683,1709]" name="Brazil" pageId="4" pageNumber="19">Brazil</collectingCountry>
(
<collectingRegion id="8B672AB0FF8BFFB4135DA2A7FDBDF945" box="[1057,1229,1683,1709]" country="Brazil" name="Espirito Santo" pageId="4" pageNumber="19">Espírito Santo</collectingRegion>
,
<collectingRegion id="8B672AB0FF8BFFB413A5A2A7FC07F945" box="[1241,1399,1683,1709]" country="Brazil" name="Minas Gerais" pageId="4" pageNumber="19">Minas Gerais</collectingRegion>
and
<collectingRegion id="8B672AB0FF8BFFB41660A283F8B8F93A" box="[284,456,1719,1746]" country="Brazil" name="Rio de Janeiro" pageId="4" pageNumber="19">Rio de Janeiro</collectingRegion>
) (
<figureCitation id="D198F8D7FF8BFFB416A3A283FB0AF939" box="[479,634,1719,1745]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[265,1323,265,1849]" captionTargetId="figure-17@8.[265,1323,265,1849]" captionTargetPageId="8" captionText="Fig. 1. Males of Inca spp. AE, P. Inca axeli sp. nov., holotype (MZSP 22036). FJ, Q. Inca neglectus sp. nov., paratype (MZSP). KO, R. Inca irroratus Chevrolat, 1833, neotype (BMNH). A, F, K. Dorsal habitus. B, G, L. Ventral habitus. C, H, M. Lateral habitus. D, I, N. Dorsal view of aedeagus. E, J, O. Lateral view of aedeagus. PR. Dorsal view of head. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4736133" httpUri="https://zenodo.org/record/4736133/files/figure.png" pageId="4" pageNumber="19">Figs 1FJ, Q</figureCitation>
,
<figureCitation id="D198F8D7FF8BFFB415FAA283FBDAF939" box="[646,682,1719,1745]" captionStart="Fig" captionStartId="14.[189,232,1823,1849]" captionTargetBox="[201,1387,266,1779]" captionTargetId="figure-125@14.[200,1387,265,1779]" captionTargetPageId="14" captionText="Fig. 4. Images of liveInca spp.A. Inca axelisp. nov. (photo by R. Barros and R. Koike).B. Inca neglectus sp. nov. (photo by N. Marchito). C. Inca clathratus sommeri Westwood, 1844 (photo by A. González Gallina). D. Inca clathratus clathratus (Olivier, 1792) (photo by R. Barros and R. Koike). E. Inca clathratus quesneli Boos &amp; Ratcliffe, 1985 (photo by M. Hulme). F. Inca pulverulentus (Olivier, 1789) mating (photo by R. Barros and R. Koike). G. Inca bonplandi (Gyllenhal, 1817) (photo by J. Touroult). H. Inca besckii Burmeister &amp; Schaum, 1840 feeding on tree sap (photo by N. Wisnik)." figureDoi="http://doi.org/10.5281/zenodo.4736143" httpUri="https://zenodo.org/record/4736143/files/figure.png" pageId="4" pageNumber="19">4B</figureCitation>
) ...............................................................
<taxonomicName id="8EA39FD1FF8BFFB413FBA283FC64F93A" authority="Sousa &amp; Seidel, 2021" authorityName="Sousa &amp; Seidel" authorityYear="2021" box="[1159,1300,1719,1746]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="neglectus" status="sp. nov.">
<emphasis id="7BD73840FF8BFFB413FBA283FC64F93A" bold="true" box="[1159,1300,1719,1746]" italics="true" pageId="4" pageNumber="19">I. neglectus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60E4853BFF8BFFB41260A28CFC07F93A" box="[1308,1399,1720,1746]" pageId="4" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="F257FCF7FF8BFFB417C1A2CEFC07F86A" pageId="4" pageNumber="19">
<paragraph id="491CE452FF8BFFB417C1A2CEFC07F8D0" blockId="4.[189,1399,1330,2025]" pageId="4" pageNumber="19">
<keyLead id="F2524767FF8BFFB417C1A2CEFC07F8D0" pageId="4" pageNumber="19">
4. Posterior tooth of protibia short and rounded. Posterior angles of pronotum rounded.
<collectingCountry id="31B4A4C2FF8BFFB413ACA2CEFC68F8FC" box="[1232,1304,1786,1812]" name="Brazil" pageId="4" pageNumber="19">Brazil</collectingCountry>
(
<collectingRegion id="8B672AB0FF8BFFB4125BA2CEF831F8D1" country="Brazil" name="Rio de Janeiro" pageId="4" pageNumber="19">Rio de Janeiro</collectingRegion>
) (
<figureCitation id="D198F8D7FF8BFFB41625A32AF88BF8D1" box="[345,507,1822,1849]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[265,1323,265,1849]" captionTargetId="figure-17@8.[265,1323,265,1849]" captionTargetPageId="8" captionText="Fig. 1. Males of Inca spp. AE, P. Inca axeli sp. nov., holotype (MZSP 22036). FJ, Q. Inca neglectus sp. nov., paratype (MZSP). KO, R. Inca irroratus Chevrolat, 1833, neotype (BMNH). A, F, K. Dorsal habitus. B, G, L. Ventral habitus. C, H, M. Lateral habitus. D, I, N. Dorsal view of aedeagus. E, J, O. Lateral view of aedeagus. PR. Dorsal view of head. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4736133" httpUri="https://zenodo.org/record/4736133/files/figure.png" pageId="4" pageNumber="19">Fig. 1KO, R</figureCitation>
) ..........................................................................
<taxonomicName id="8EA39FD1FF8BFFB41357A32BFC07F8D0" authority="Chevrolat, 1833" authorityName="Chevrolat" authorityYear="1833" box="[1067,1399,1822,1849]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="irroratus">
<emphasis id="7BD73840FF8BFFB41357A32BFDC2F8D0" bold="true" box="[1067,1202,1822,1849]" pageId="4" pageNumber="19">
<emphasis id="7BD73840FF8BFFB41357A32BFD46F8D1" bold="true" box="[1067,1078,1823,1849]" italics="true" pageId="4" pageNumber="19">I</emphasis>
.
<emphasis id="7BD73840FF8BFFB41339A32AFDC2F8D0" bold="true" box="[1093,1202,1822,1848]" italics="true" pageId="4" pageNumber="19">irroratus</emphasis>
</emphasis>
<bibRefCitation id="2D3299A3FF8BFFB413C5A32AFC07F8D0" author="Chevrolat A." box="[1209,1399,1822,1848]" pageId="4" pageNumber="19" refId="ref9632" refString="Chevrolat A. 1833. Description d'especes nouvelles. Revue Entomologique - Coleopteres 10: unpaginated. Available from https: // www. biodiversitylibrary. org / item / 44561 # page / 349 / mode / 1 up [accessed 6 Apr. 2021]." type="journal volume" year="1833">Chevrolat, 1833</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="491CE452FF8BFFB417BDA377FC07F86A" blockId="4.[189,1399,1330,2025]" pageId="4" pageNumber="19">
<keyLead id="F2524767FF8BFFB417BDA377FC07F86A" pageId="4" pageNumber="19"> Posterior tooth of protibia long and acute or short and rounded. Posterior angles of pronotum acuminate........................................................................................................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="F257FCF7FF8BFFB417C1A39EFC07F801" pageId="4" pageNumber="19">
<paragraph id="491CE452FF8BFFB417C1A39EFC07F82C" blockId="4.[189,1399,1330,2025]" box="[189,1399,1962,1989]" pageId="4" pageNumber="19">
<keyLead id="F2524767FF8BFFB417C1A39EFC07F82C" box="[189,1399,1962,1989]" pageId="4" pageNumber="19">5. Clypeal horn with proximal emargination. Posterior tooth of protibia short and rounded................ 6</keyLead>
</paragraph>
<paragraph id="491CE452FF8BFFB417BDA3FBFC07F801" blockId="4.[189,1399,1330,2025]" box="[193,1399,1998,2025]" pageId="4" pageNumber="19">
<keyLead id="F2524767FF8BFFB417BDA3FBFC07F801" box="[193,1399,1998,2025]" pageId="4" pageNumber="19"> Clypeal horn without proXimal emargination. Posterior tooth of protibial long and acute............... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="F257FCF7FF8AFFB517C1A53DFC06FE5A" pageId="5" pageNumber="20">
<paragraph id="491CE452FF8AFFB517C1A53DFC07FE83" blockId="5.[189,1399,265,2025]" pageId="5" pageNumber="20">
<keyLead id="F2524767FF8AFFB517C1A53DFC07FE83" pageId="5" pageNumber="20">
6. Inner dorsal carina of clypeal horn distinct. Legs without red reflections.
<collectingCountry id="31B4A4C2FF8AFFB5133AA53DFDFEFECB" box="[1094,1166,265,291]" name="Brazil" pageId="5" pageNumber="20">Brazil</collectingCountry>
(
<collectingRegion id="8B672AB0FF8AFFB513E2A53DFC4FFECB" box="[1182,1343,265,291]" country="Brazil" name="Minas Gerais" pageId="5" pageNumber="20">Minas Gerais</collectingRegion>
,
<collectingRegion id="8B672AB0FF8AFFB51230A53DF815FEAF" country="Brazil" name="Rio de Janeiro" pageId="5" pageNumber="20">Rio de Janeiro</collectingRegion>
,
<collectingRegion id="8B672AB0FF8AFFB5160DA519F8B4FEAF" box="[369,452,301,327]" country="Brazil" name="Parana" pageId="5" pageNumber="20">Paraná</collectingRegion>
,
<collectingRegion id="8B672AB0FF8AFFB516B3A519FB0BFEAF" box="[463,635,301,327]" country="Brazil" name="Santa Catarina" pageId="5" pageNumber="20">
Santa
<taxonomicName id="8EA39FD1FF8AFFB51564A519FB0BFEAF" ID-CoL="94MGZ" authorityName="Vojnits &amp; de Laever" authorityYear="1973" box="[536,635,301,327]" class="Insecta" family="Geometridae" genus="Catarina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="genus">Catarina</taxonomicName>
</collectingRegion>
and
<collectingRegion id="8B672AB0FF8AFFB515C8A519FAE1FEA0" box="[692,913,301,328]" country="Brazil" name="Rio Grande do Sul" pageId="5" pageNumber="20">Rio Grande do Sul</collectingRegion>
) (
<figureCitation id="D198F8D7FF8AFFB514D5A519FD75FEAF" box="[937,1029,301,327]" captionStart="Fig" captionStartId="14.[189,232,1823,1849]" captionTargetBox="[201,1387,266,1779]" captionTargetId="figure-125@14.[200,1387,265,1779]" captionTargetPageId="14" captionText="Fig. 4. Images of liveInca spp.A. Inca axelisp. nov. (photo by R. Barros and R. Koike).B. Inca neglectus sp. nov. (photo by N. Marchito). C. Inca clathratus sommeri Westwood, 1844 (photo by A. González Gallina). D. Inca clathratus clathratus (Olivier, 1792) (photo by R. Barros and R. Koike). E. Inca clathratus quesneli Boos &amp; Ratcliffe, 1985 (photo by M. Hulme). F. Inca pulverulentus (Olivier, 1789) mating (photo by R. Barros and R. Koike). G. Inca bonplandi (Gyllenhal, 1817) (photo by J. Touroult). H. Inca besckii Burmeister &amp; Schaum, 1840 feeding on tree sap (photo by N. Wisnik)." figureDoi="http://doi.org/10.5281/zenodo.4736143" httpUri="https://zenodo.org/record/4736143/files/figure.png" pageId="5" pageNumber="20">Fig. 4H</figureCitation>
)................................................ ................................................................................................
<taxonomicName id="8EA39FD1FF8AFFB514CDA565FC07FE83" authority="Burmeister &amp; Schaum, 1840" authorityName="Burmeister &amp; Schaum" authorityYear="1840" box="[945,1399,336,363]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="besckii">
<emphasis id="7BD73840FF8AFFB514CDA565FD6FFE83" bold="true" box="[945,1055,336,363]" italics="true" pageId="5" pageNumber="20">I. besckii</emphasis>
<bibRefCitation id="2D3299A3FF8AFFB5135BA564FC07FE83" author="Burmeister H. &amp; Schaum H." box="[1063,1399,336,363]" pageId="5" pageNumber="20" pagination="353 - 420" refId="ref9569" refString="Burmeister H. &amp; Schaum H. 1840. Kritische Revision der Lamellicornia melitophila. Erstes Stuck (Trichiadae). Zeitschrift fur die Entomologie 2: 353 - 420." type="journal article" year="1840">Burmeister &amp; Schaum, 1840</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="491CE452FF8AFFB517BDA541FC06FE5A" blockId="5.[189,1399,265,2025]" pageId="5" pageNumber="20">
<keyLead id="F2524767FF8AFFB517BDA541FC06FE5A" pageId="5" pageNumber="20">
Inner dorsal carina of clypeal horn indistinct. Legs with red reflections.
<collectingCountry id="31B4A4C2FF8AFFB5133FA540FDCBFE66" box="[1091,1211,372,398]" name="Argentina" pageId="5" pageNumber="20">Argentina</collectingCountry>
,
<collectingCountry id="31B4A4C2FF8AFFB513B4A540FC51FE66" box="[1224,1313,372,398]" name="Bolivia" pageId="5" pageNumber="20">Bolivia</collectingCountry>
,
<collectingCountry id="31B4A4C2FF8AFFB51253A540FC07FE66" box="[1327,1399,372,398]" name="Brazil" pageId="5" pageNumber="20">Brazil</collectingCountry>
and
<collectingCountry id="31B4A4C2FF8AFFB51660A5ADF8F9FE5B" box="[284,393,409,435]" name="Paraguay" pageId="5" pageNumber="20">Paraguay</collectingCountry>
(
<figureCitation id="D198F8D7FF8AFFB516E5A5ACF885FE5A" box="[409,501,408,434]" captionStart="Fig" captionStartId="14.[189,232,1823,1849]" captionTargetBox="[201,1387,266,1779]" captionTargetId="figure-125@14.[200,1387,265,1779]" captionTargetPageId="14" captionText="Fig. 4. Images of liveInca spp.A. Inca axelisp. nov. (photo by R. Barros and R. Koike).B. Inca neglectus sp. nov. (photo by N. Marchito). C. Inca clathratus sommeri Westwood, 1844 (photo by A. González Gallina). D. Inca clathratus clathratus (Olivier, 1792) (photo by R. Barros and R. Koike). E. Inca clathratus quesneli Boos &amp; Ratcliffe, 1985 (photo by M. Hulme). F. Inca pulverulentus (Olivier, 1789) mating (photo by R. Barros and R. Koike). G. Inca bonplandi (Gyllenhal, 1817) (photo by J. Touroult). H. Inca besckii Burmeister &amp; Schaum, 1840 feeding on tree sap (photo by N. Wisnik)." figureDoi="http://doi.org/10.5281/zenodo.4736143" httpUri="https://zenodo.org/record/4736143/files/figure.png" pageId="5" pageNumber="20">Fig. 4G</figureCitation>
).......................................................................
<taxonomicName id="8EA39FD1FF8AFFB51374A5ACFC06FE5A" authority="(Gyllenhal, 1817)" baseAuthorityName="Gyllenhal" baseAuthorityYear="1817" box="[1032,1398,408,435]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="bonplandi">
<emphasis id="7BD73840FF8AFFB51374A5ACFDEEFE5B" bold="true" box="[1032,1182,408,435]" italics="true" pageId="5" pageNumber="20">I. bonplandi</emphasis>
(Gyllenhal, 1817)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="F257FCF7FF8AFFB517C1A5EFFC06FDD4" pageId="5" pageNumber="20">
<paragraph id="491CE452FF8AFFB517C1A5EFFC06FDF0" blockId="5.[189,1399,265,2025]" pageId="5" pageNumber="20">
<keyLead id="F2524767FF8AFFB517C1A5EFFC06FDF0" pageId="5" pageNumber="20">
7. Medial and lateral area of pronotum and pygidium without yellow longitudinal bands.
<collectingCountry id="31B4A4C2FF8AFFB5125BA5EEFC02FE1C" box="[1319,1394,474,500]" name="Brazil" pageId="5" pageNumber="20">Brazil</collectingCountry>
,
<collectingCountry id="31B4A4C2FF8AFFB51796A5CAF812FDF0" box="[234,354,510,536]" name="Argentina" pageId="5" pageNumber="20">Argentina</collectingCountry>
,
<collectingCountry id="31B4A4C2FF8AFFB51612A5CAF8ABFDF0" box="[366,475,510,536]" name="Paraguay" pageId="5" pageNumber="20">Paraguay</collectingCountry>
(
<figureCitation id="D198F8D7FF8AFFB51697A5CAFB32FDF0" box="[491,578,510,536]" captionStart="Fig" captionStartId="14.[189,232,1823,1849]" captionTargetBox="[201,1387,266,1779]" captionTargetId="figure-125@14.[200,1387,265,1779]" captionTargetPageId="14" captionText="Fig. 4. Images of liveInca spp.A. Inca axelisp. nov. (photo by R. Barros and R. Koike).B. Inca neglectus sp. nov. (photo by N. Marchito). C. Inca clathratus sommeri Westwood, 1844 (photo by A. González Gallina). D. Inca clathratus clathratus (Olivier, 1792) (photo by R. Barros and R. Koike). E. Inca clathratus quesneli Boos &amp; Ratcliffe, 1985 (photo by M. Hulme). F. Inca pulverulentus (Olivier, 1789) mating (photo by R. Barros and R. Koike). G. Inca bonplandi (Gyllenhal, 1817) (photo by J. Touroult). H. Inca besckii Burmeister &amp; Schaum, 1840 feeding on tree sap (photo by N. Wisnik)." figureDoi="http://doi.org/10.5281/zenodo.4736143" httpUri="https://zenodo.org/record/4736143/files/figure.png" pageId="5" pageNumber="20">Fig. 4F</figureCitation>
) ..........................................................
<taxonomicName id="8EA39FD1FF8AFFB51482A5CAFC06FDF0" authority="(Olivier, 1789)" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[1022,1398,509,536]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="pulverulentus">
<emphasis id="7BD73840FF8AFFB51482A5CAFDB0FDF0" bold="true" box="[1022,1216,509,536]" italics="true" pageId="5" pageNumber="20">I. pulverulentus</emphasis>
(Olivier, 1789)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="491CE452FF8AFFB517BDA616FC06FDD4" blockId="5.[189,1399,265,2025]" box="[193,1398,546,572]" pageId="5" pageNumber="20">
<keyLead id="F2524767FF8AFFB517BDA616FC06FDD4" box="[193,1398,546,572]" pageId="5" pageNumber="20">
Medial and lateral area of pronotum and pygidium with yellow longitudinal bands (
<taxonomicName id="8EA39FD1FF8AFFB513C4A616FC38FDD4" baseAuthorityName="Olivier" baseAuthorityYear="1792" box="[1208,1352,546,572]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="clathratus">
<emphasis id="7BD73840FF8AFFB513C4A616FDB2FDD3" box="[1208,1218,546,571]" italics="true" pageId="5" pageNumber="20">I</emphasis>
.
<emphasis id="7BD73840FF8AFFB513ACA616FC38FDD4" box="[1232,1352,546,572]" italics="true" pageId="5" pageNumber="20">clathratus</emphasis>
</taxonomicName>
)...8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="F257FCF7FF8AFFB517C1A650FC07FD2D" pageId="5" pageNumber="20">
<paragraph id="491CE452FF8AFFB517C1A650FC07FD4A" blockId="5.[189,1399,265,2025]" pageId="5" pageNumber="20">
<keyLead id="F2524767FF8AFFB517C1A650FC07FD4A" pageId="5" pageNumber="20">
8. Inner dorsal carina of clypeal horn abruptly interrupted at apex.
<collectingCountry id="31B4A4C2FF8AFFB514A6A650FD4EFD96" box="[986,1086,612,638]" name="Trinidad and Tobago" pageId="5" pageNumber="20">Trinidad</collectingCountry>
(
<figureCitation id="D198F8D7FF8AFFB51332A650FDD6FD96" box="[1102,1190,612,638]" captionStart="Fig" captionStartId="14.[189,232,1823,1849]" captionTargetBox="[201,1387,266,1779]" captionTargetId="figure-125@14.[200,1387,265,1779]" captionTargetPageId="14" captionText="Fig. 4. Images of liveInca spp.A. Inca axelisp. nov. (photo by R. Barros and R. Koike).B. Inca neglectus sp. nov. (photo by N. Marchito). C. Inca clathratus sommeri Westwood, 1844 (photo by A. González Gallina). D. Inca clathratus clathratus (Olivier, 1792) (photo by R. Barros and R. Koike). E. Inca clathratus quesneli Boos &amp; Ratcliffe, 1985 (photo by M. Hulme). F. Inca pulverulentus (Olivier, 1789) mating (photo by R. Barros and R. Koike). G. Inca bonplandi (Gyllenhal, 1817) (photo by J. Touroult). H. Inca besckii Burmeister &amp; Schaum, 1840 feeding on tree sap (photo by N. Wisnik)." figureDoi="http://doi.org/10.5281/zenodo.4736143" httpUri="https://zenodo.org/record/4736143/files/figure.png" pageId="5" pageNumber="20">Fig. 4E</figureCitation>
).......................... ....................................................................................
<taxonomicName id="8EA39FD1FF8AFFB51427A6BCFC07FD4A" authority="Boos &amp; Ratcliffe, 1985" authorityName="Boos &amp; Ratcliffe" authorityYear="1985" box="[859,1399,647,674]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="subSpecies" species="clathratus" subSpecies="quesneli">
<emphasis id="7BD73840FF8AFFB51427A6BCFD2FFD4A" bold="true" box="[859,1119,647,674]" italics="true" pageId="5" pageNumber="20">I. clathratus quesneli</emphasis>
<bibRefCitation id="2D3299A3FF8AFFB5131BA6BCFC07FD4A" author="Boos J. &amp; Ratcliffe B. C." box="[1127,1399,647,674]" pageId="5" pageNumber="20" pagination="381 - 389" refId="ref9385" refString="Boos J. &amp; Ratcliffe B. C. 1985. A new subspecies of Inca clathrata from Trinidad, West Indies, and range extensions for Inca clathrata. The Coleopterists Bulletin 39: 381 - 389." type="journal article" year="1985">Boos &amp; Ratcliffe, 1985</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="491CE452FF8AFFB517BDA698FC07FD2D" blockId="5.[189,1399,265,2025]" box="[193,1399,683,710]" pageId="5" pageNumber="20">
<keyLead id="F2524767FF8AFFB517BDA698FC07FD2D" box="[193,1399,683,710]" pageId="5" pageNumber="20"> Inner dorsal carina of clypeal horns gradually interrupted at apeX.................................................... 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="F257FCF7FF8AFFB517C1A6D9FC06FC52" pageId="5" pageNumber="20">
<paragraph id="491CE452FF8AFFB517C1A6D9FC07FCA7" blockId="5.[189,1399,265,2025]" pageId="5" pageNumber="20">
<keyLead id="F2524767FF8AFFB517C1A6D9FC07FCA7" pageId="5" pageNumber="20">
9. Inner dorsal carina convergent to the inner apical angle.
<collectingCountry id="31B4A4C2FF8AFFB514DEA6D9FA8FFCEF" box="[930,1023,749,775]" name="Mexico" pageId="5" pageNumber="20">Mexico</collectingCountry>
,
<collectingCountry id="31B4A4C2FF8AFFB51372A6D9FDE0FCEF" box="[1038,1168,749,775]" name="Guatemala" pageId="5" pageNumber="20">Guatemala</collectingCountry>
,
<collectingCountry id="31B4A4C2FF8AFFB513DCA6D9FD9EFCEF" box="[1184,1262,749,775]" name="Belize" pageId="5" pageNumber="20">Belize</collectingCountry>
,
<collectingCountry id="31B4A4C2FF8AFFB51381A6D9FC02FCEF" box="[1277,1394,749,775]" name="Honduras" pageId="5" pageNumber="20">Honduras</collectingCountry>
,
<collectingCountry id="31B4A4C2FF8AFFB51796A725F816FCC3" box="[234,358,785,811]" name="Nicaragua" pageId="5" pageNumber="20">Nicaragua</collectingCountry>
,
<collectingCountry id="31B4A4C2FF8AFFB5160EA725F885FCC3" box="[370,501,785,811]" name="Costa Rica" pageId="5" pageNumber="20">Costa Rica</collectingCountry>
,
<collectingCountry id="31B4A4C2FF8AFFB5157CA725FB10FCC3" box="[512,608,785,811]" name="Panama" pageId="5" pageNumber="20">Panama</collectingCountry>
,
<collectingCountry id="31B4A4C2FF8AFFB51510A725FB90FCC3" box="[620,736,785,811]" name="Colombia" pageId="5" pageNumber="20">Colombia</collectingCountry>
and
<collectingCountry id="31B4A4C2FF8AFFB51464A725FA0AFCC3" box="[792,890,785,811]" name="Ecuador" pageId="5" pageNumber="20">Ecuador</collectingCountry>
(
<figureCitation id="D198F8D7FF8AFFB514F6A725FAADFCC4" box="[906,989,785,812]" captionStart="Fig" captionStartId="14.[189,232,1823,1849]" captionTargetBox="[201,1387,266,1779]" captionTargetId="figure-125@14.[200,1387,265,1779]" captionTargetPageId="14" captionText="Fig. 4. Images of liveInca spp.A. Inca axelisp. nov. (photo by R. Barros and R. Koike).B. Inca neglectus sp. nov. (photo by N. Marchito). C. Inca clathratus sommeri Westwood, 1844 (photo by A. González Gallina). D. Inca clathratus clathratus (Olivier, 1792) (photo by R. Barros and R. Koike). E. Inca clathratus quesneli Boos &amp; Ratcliffe, 1985 (photo by M. Hulme). F. Inca pulverulentus (Olivier, 1789) mating (photo by R. Barros and R. Koike). G. Inca bonplandi (Gyllenhal, 1817) (photo by J. Touroult). H. Inca besckii Burmeister &amp; Schaum, 1840 feeding on tree sap (photo by N. Wisnik)." figureDoi="http://doi.org/10.5281/zenodo.4736143" httpUri="https://zenodo.org/record/4736143/files/figure.png" pageId="5" pageNumber="20">Fig.4C</figureCitation>
) ..................................................... ..............................................................................................
<taxonomicName id="8EA39FD1FF8AFFB514D8A701FC07FCA7" authority="Westwood, 1844" authorityName="Westwood" authorityYear="1844" box="[932,1399,820,847]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="subSpecies" species="clathratus" subSpecies="sommeri">
<emphasis id="7BD73840FF8AFFB514D8A701FDDBFCA7" bold="true" box="[932,1195,820,847]" italics="true" pageId="5" pageNumber="20">I. clathratus sommeri</emphasis>
Westwood, 1844
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="491CE452FF8AFFB517BDA76DFC06FC52" blockId="5.[189,1399,265,2025]" pageId="5" pageNumber="20">
<keyLead id="F2524767FF8AFFB517BDA76DFC06FC52" pageId="5" pageNumber="20">
Inner dorsal carina convergent to the outer apical angle.
<collectingCountry id="31B4A4C2FF8AFFB514C5A76CFD40FC9A" box="[953,1072,856,882]" name="Colombia" pageId="5" pageNumber="20">Colombia</collectingCountry>
,
<collectingCountry id="31B4A4C2FF8AFFB5133EA76CFDFDFC9A" box="[1090,1165,856,882]" name="Brazil" pageId="5" pageNumber="20">Brazil</collectingCountry>
,
<collectingCountry id="31B4A4C2FF8AFFB513E2A76CFC72FC9A" box="[1182,1282,856,882]" name="Ecuador" pageId="5" pageNumber="20">Ecuador</collectingCountry>
,
<collectingCountry id="31B4A4C2FF8AFFB51268A76DFC02FC9B" box="[1300,1394,857,883]" name="Guyana" pageId="5" pageNumber="20">Guyana</collectingCountry>
,
<collectingCountry id="31B4A4C2FF8AFFB51796A748F82DFC7E" box="[234,349,892,918]" name="Suriname" pageId="5" pageNumber="20">Suriname</collectingCountry>
, French
<collectingCountry id="31B4A4C2FF8AFFB516BEA748FB50FC7E" box="[450,544,892,918]" name="Guyana" pageId="5" pageNumber="20">Guyana</collectingCountry>
,
<collectingCountry id="31B4A4C2FF8AFFB51550A749FB15FC7F" box="[556,613,893,919]" name="Peru" pageId="5" pageNumber="20">Peru</collectingCountry>
,
<collectingCountry id="31B4A4C2FF8AFFB5150DA748FBB7FC7E" box="[625,711,892,918]" name="Bolivia" pageId="5" pageNumber="20">Bolivia</collectingCountry>
and
<collectingCountry id="31B4A4C2FF8AFFB51582A748FA03FC7E" box="[766,883,892,918]" name="Argentina" pageId="5" pageNumber="20">Argentina</collectingCountry>
(
<figureCitation id="D198F8D7FF8AFFB514FFA748FAAFFC7E" box="[899,991,892,918]" captionStart="Fig" captionStartId="14.[189,232,1823,1849]" captionTargetBox="[201,1387,266,1779]" captionTargetId="figure-125@14.[200,1387,265,1779]" captionTargetPageId="14" captionText="Fig. 4. Images of liveInca spp.A. Inca axelisp. nov. (photo by R. Barros and R. Koike).B. Inca neglectus sp. nov. (photo by N. Marchito). C. Inca clathratus sommeri Westwood, 1844 (photo by A. González Gallina). D. Inca clathratus clathratus (Olivier, 1792) (photo by R. Barros and R. Koike). E. Inca clathratus quesneli Boos &amp; Ratcliffe, 1985 (photo by M. Hulme). F. Inca pulverulentus (Olivier, 1789) mating (photo by R. Barros and R. Koike). G. Inca bonplandi (Gyllenhal, 1817) (photo by J. Touroult). H. Inca besckii Burmeister &amp; Schaum, 1840 feeding on tree sap (photo by N. Wisnik)." figureDoi="http://doi.org/10.5281/zenodo.4736143" httpUri="https://zenodo.org/record/4736143/files/figure.png" pageId="5" pageNumber="20">Fig. 4D</figureCitation>
) ..................................................... ..............................................................................................
<taxonomicName id="8EA39FD1FF8AFFB514DBA794FC06FC52" authority="clathratus (Olivier, 1792)" authorityName="clathratus (Olivier" authorityYear="1792" baseAuthorityName="Olivier" baseAuthorityYear="1792" box="[935,1398,928,955]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="subSpecies" species="clathratus" subSpecies="clathratus">
<emphasis id="7BD73840FF8AFFB514DBA794FDB1FC53" bold="true" box="[935,1217,928,955]" italics="true" pageId="5" pageNumber="20">I. clathratus clathratus</emphasis>
(Olivier, 1792)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="F257FCF7FF8AFFB517C1A7D6FC06FBC8" pageId="5" pageNumber="20">
<paragraph id="491CE452FF8AFFB517C1A7D6FC06FC14" blockId="5.[189,1399,265,2025]" box="[189,1398,994,1021]" pageId="5" pageNumber="20">
<keyLead id="F2524767FF8AFFB517C1A7D6FC06FC14" box="[189,1398,994,1021]" pageId="5" pageNumber="20">10. Anterior margin of clypeus with a medial tooth...............................................................................11</keyLead>
</paragraph>
<paragraph id="491CE452FF8AFFB517BDA032FC06FBC8" blockId="5.[189,1399,265,2025]" box="[193,1398,1030,1056]" pageId="5" pageNumber="20">
<keyLead id="F2524767FF8AFFB517BDA032FC06FBC8" box="[193,1398,1030,1056]" pageId="5" pageNumber="20"> Anterior margin of clypeus without a medial tooth......................................................................... 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="F257FCF7FF8AFFB517C1A07CFC06FB6E" pageId="5" pageNumber="20">
<paragraph id="491CE452FF8AFFB517C1A07CFC06FB8A" blockId="5.[189,1399,265,2025]" box="[189,1398,1096,1122]" pageId="5" pageNumber="20">
<keyLead id="F2524767FF8AFFB517C1A07CFC06FB8A" box="[189,1398,1096,1122]" pageId="5" pageNumber="20">11. Medial tooth of the anterior margin rounded................................................................................... 12</keyLead>
</paragraph>
<paragraph id="491CE452FF8AFFB517BDA058FC06FB6E" blockId="5.[189,1399,265,2025]" box="[193,1398,1132,1158]" pageId="5" pageNumber="20">
<keyLead id="F2524767FF8AFFB517BDA058FC06FB6E" box="[193,1398,1132,1158]" pageId="5" pageNumber="20"> Medial tooth of the anterior margin acute ....................................................................................... 17</keyLead>
</paragraph>
</keyStep>
<keyStep id="F257FCF7FF8AFFB517C1A09AFC06FABF" pageId="5" pageNumber="20">
<paragraph id="491CE452FF8AFFB517C1A09AFC07FAE7" blockId="5.[189,1399,265,2025]" pageId="5" pageNumber="20">
<keyLead id="F2524767FF8AFFB517C1A09AFC07FAE7" pageId="5" pageNumber="20">
12. Disc of clypeus with a large fovea. Lateral margin of pronotum with deep crenulation. Marks of medial area of elytral epipleuron evident.
<collectingCountry id="31B4A4C2FF8AFFB515AAA0E5FA6EFB03" box="[726,798,1233,1259]" name="Brazil" pageId="5" pageNumber="20">Brazil</collectingCountry>
(
<collectingRegion id="8B672AB0FF8AFFB51451A0E5FAAAFB04" box="[813,986,1233,1260]" country="Brazil" name="Rio de Janeiro" pageId="5" pageNumber="20">Rio de Janeiro</collectingRegion>
) (
<figureCitation id="D198F8D7FF8AFFB5148DA0E5FDFFFB04" box="[1009,1167,1233,1260]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[253,1334,267,1885]" captionTargetId="figure-17@10.[253,1334,267,1885]" captionTargetPageId="10" captionText="Fig. 2. Females of Inca spp. and type labels. AD, M. Inca axeli sp. nov. EH, N. Inca neglectus sp. nov. IL, O. Inca irroratus Chevrolat, 1833. A, E, I. Dorsal habitus. B, F, J. Ventral habitus. C, G, K. Lateral habitus. D, H, L. Type labels. MO. Frontal view of head. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4736137" httpUri="https://zenodo.org/record/4736137/files/figure.png" pageId="5" pageNumber="20">Figs 2IK, O</figureCitation>
,
<figureCitation id="D198F8D7FF8AFFB513E7A0E6FDAEFB04" box="[1179,1246,1234,1260]" captionStart="Fig" captionStartId="13.[189,232,1892,1918]" captionTargetBox="[265,1323,265,1848]" captionTargetId="figure-12@13.[265,1323,265,1848]" captionTargetPageId="13" captionText="Fig. 3. Morphological details of Inca spp. A, D, GN. I. axeli sp. nov. (A, D: ♀, GN ♂). B, E. I. neglectus sp. nov. C, F. I. irroratus Chevrolat, 1833. AC. Protibia, lateral view. DF. Details of elytral epipleuron. GN. Mouthparts. GI. Right mandibles (dorsal, inner and ventral view). JK. Labrum (dorsal and frontal view). L. Right maxilla (dorsal view). M. Epipharynx. N. Labium." figureDoi="http://doi.org/10.5281/zenodo.4736141" httpUri="https://zenodo.org/record/4736141/files/figure.png" pageId="5" pageNumber="20">3C, F</figureCitation>
) .................. ................................................................................................................
<taxonomicName id="8EA39FD1FF8AFFB51357A0C2FC07FAE7" authority="Chevrolat, 1833" authorityName="Chevrolat" authorityYear="1833" box="[1067,1399,1269,1296]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="irroratus">
<emphasis id="7BD73840FF8AFFB51357A0C2FDC2FAE7" bold="true" box="[1067,1202,1269,1296]" pageId="5" pageNumber="20">
<emphasis id="7BD73840FF8AFFB51357A0C2FD46FAF8" bold="true" box="[1067,1078,1270,1296]" italics="true" pageId="5" pageNumber="20">I</emphasis>
.
<emphasis id="7BD73840FF8AFFB51339A0C1FDC2FAE7" bold="true" box="[1093,1202,1269,1295]" italics="true" pageId="5" pageNumber="20">irroratus</emphasis>
</emphasis>
<bibRefCitation id="2D3299A3FF8AFFB513C5A0C1FC07FAE7" author="Chevrolat A." box="[1209,1399,1269,1295]" pageId="5" pageNumber="20" refId="ref9632" refString="Chevrolat A. 1833. Description d'especes nouvelles. Revue Entomologique - Coleopteres 10: unpaginated. Available from https: // www. biodiversitylibrary. org / item / 44561 # page / 349 / mode / 1 up [accessed 6 Apr. 2021]." type="journal volume" year="1833">Chevrolat, 1833</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="491CE452FF8AFFB517BDA12EFC06FABF" blockId="5.[189,1399,265,2025]" pageId="5" pageNumber="20">
<keyLead id="F2524767FF8AFFB517BDA12EFC06FABF" pageId="5" pageNumber="20"> Clypeus without fovea. Lateral margin of pronotum with shallow crenulation. Marks of medial area of elytral epipleuron indistinct......................................................................................................... 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="F257FCF7FF8AFFB517C1A14BFC07F9C0" pageId="5" pageNumber="20">
<paragraph id="491CE452FF8AFFB517C1A14BFC07FA55" blockId="5.[189,1399,265,2025]" pageId="5" pageNumber="20">
<keyLead id="F2524767FF8AFFB517C1A14BFC07FA55" pageId="5" pageNumber="20">
13. Medial tooth of anterior margin of clypeus evident. Elytral disc with large waxy maculae.
<collectingCountry id="31B4A4C2FF8AFFB51253A14BFC07FA71" box="[1327,1399,1407,1433]" name="Brazil" pageId="5" pageNumber="20">Brazil</collectingCountry>
(
<collectingRegion id="8B672AB0FF8AFFB5178FA197F81DFA54" box="[243,365,1442,1469]" country="Brazil" name="Sao Paulo" pageId="5" pageNumber="20">São Paulo</collectingRegion>
,
<collectingRegion id="8B672AB0FF8AFFB51605A197F8B9FA55" box="[377,457,1443,1469]" country="Brazil" name="Parana" pageId="5" pageNumber="20">Paraná</collectingRegion>
and
<collectingRegion id="8B672AB0FF8AFFB5157EA197FBDFFA55" box="[514,687,1442,1469]" country="Brazil" name="Santa Catarina" pageId="5" pageNumber="20">
Santa
<taxonomicName id="8EA39FD1FF8AFFB51536A196FBDFFA55" ID-CoL="94MGZ" authorityName="Vojnits &amp; de Laever" authorityYear="1973" box="[586,687,1442,1469]" class="Insecta" family="Geometridae" genus="Catarina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="22" phylum="Arthropoda" rank="genus">Catarina</taxonomicName>
</collectingRegion>
) (
<figureCitation id="D198F8D7FF8AFFB515BBA196FA04FA55" box="[711,884,1442,1469]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[253,1334,267,1885]" captionTargetId="figure-17@10.[253,1334,267,1885]" captionTargetPageId="10" captionText="Fig. 2. Females of Inca spp. and type labels. AD, M. Inca axeli sp. nov. EH, N. Inca neglectus sp. nov. IL, O. Inca irroratus Chevrolat, 1833. A, E, I. Dorsal habitus. B, F, J. Ventral habitus. C, G, K. Lateral habitus. D, H, L. Type labels. MO. Frontal view of head. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4736137" httpUri="https://zenodo.org/record/4736137/files/figure.png" pageId="5" pageNumber="20">Figs 2AC, M</figureCitation>
,
<figureCitation id="D198F8D7FF8AFFB514FCA197FAB8FA55" box="[896,968,1443,1469]" captionStart="Fig" captionStartId="13.[189,232,1892,1918]" captionTargetBox="[265,1323,265,1848]" captionTargetId="figure-12@13.[265,1323,265,1848]" captionTargetPageId="13" captionText="Fig. 3. Morphological details of Inca spp. A, D, GN. I. axeli sp. nov. (A, D: ♀, GN ♂). B, E. I. neglectus sp. nov. C, F. I. irroratus Chevrolat, 1833. AC. Protibia, lateral view. DF. Details of elytral epipleuron. GN. Mouthparts. GI. Right mandibles (dorsal, inner and ventral view). JK. Labrum (dorsal and frontal view). L. Right maxilla (dorsal view). M. Epipharynx. N. Labium." figureDoi="http://doi.org/10.5281/zenodo.4736141" httpUri="https://zenodo.org/record/4736141/files/figure.png" pageId="5" pageNumber="20">3A, D</figureCitation>
)................................
<taxonomicName id="8EA39FD1FF8AFFB513BAA197FC6CFA55" authorityName="Sousa &amp; Seidel" authorityYear="2021" box="[1222,1308,1442,1469]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="axeli">
<emphasis id="7BD73840FF8AFFB513BAA197FC6CFA55" bold="true" box="[1222,1308,1442,1469]" pageId="5" pageNumber="20">
<emphasis id="7BD73840FF8AFFB513BAA197FDA1FA55" bold="true" box="[1222,1233,1443,1469]" italics="true" pageId="5" pageNumber="20">I</emphasis>
.
<emphasis id="7BD73840FF8AFFB5139DA196FC6CFA55" bold="true" box="[1249,1308,1442,1469]" italics="true" pageId="5" pageNumber="20">axeli</emphasis>
</emphasis>
</taxonomicName>
sp nov.
</keyLead>
</paragraph>
<paragraph id="491CE452FF8AFFB517BDA1F3FC07F9C0" blockId="5.[189,1399,265,2025]" pageId="5" pageNumber="20">
<keyLead id="F2524767FF8AFFB517BDA1F3FC07F9C0" pageId="5" pageNumber="20">
Medial tooth of anterior margin of clypeus almost indistinct. Elytral disc without large waXy maculae.
<collectingCountry id="31B4A4C2FF8AFFB51796A1DEF842F9EC" box="[234,306,1514,1540]" name="Brazil" pageId="5" pageNumber="20">Brazil</collectingCountry>
(
<collectingRegion id="8B672AB0FF8AFFB5163DA1DEF89DF9EC" box="[321,493,1514,1540]" country="Brazil" name="Espirito Santo" pageId="5" pageNumber="20">Espírito Santo</collectingRegion>
,
<collectingRegion id="8B672AB0FF8AFFB51685A1DEFBE7F9EC" box="[505,663,1514,1540]" country="Brazil" name="Minas Gerais" pageId="5" pageNumber="20">Minas Gerais</collectingRegion>
and
<collectingRegion id="8B672AB0FF8AFFB515B3A1DEFA0BF9ED" box="[719,891,1514,1541]" country="Brazil" name="Rio de Janeiro" pageId="5" pageNumber="20">Rio de Janeiro</collectingRegion>
) (
<figureCitation id="D198F8D7FF8AFFB514EEA1DEFD48F9ED" box="[914,1080,1514,1541]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[253,1334,267,1885]" captionTargetId="figure-17@10.[253,1334,267,1885]" captionTargetPageId="10" captionText="Fig. 2. Females of Inca spp. and type labels. AD, M. Inca axeli sp. nov. EH, N. Inca neglectus sp. nov. IL, O. Inca irroratus Chevrolat, 1833. A, E, I. Dorsal habitus. B, F, J. Ventral habitus. C, G, K. Lateral habitus. D, H, L. Type labels. MO. Frontal view of head. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4736137" httpUri="https://zenodo.org/record/4736137/files/figure.png" pageId="5" pageNumber="20">Figs 2EG, N</figureCitation>
,
<figureCitation id="D198F8D7FF8AFFB51338A1DEFDF8F9ED" box="[1092,1160,1514,1541]" captionStart="Fig" captionStartId="13.[189,232,1892,1918]" captionTargetBox="[265,1323,265,1848]" captionTargetId="figure-12@13.[265,1323,265,1848]" captionTargetPageId="13" captionText="Fig. 3. Morphological details of Inca spp. A, D, GN. I. axeli sp. nov. (A, D: ♀, GN ♂). B, E. I. neglectus sp. nov. C, F. I. irroratus Chevrolat, 1833. AC. Protibia, lateral view. DF. Details of elytral epipleuron. GN. Mouthparts. GI. Right mandibles (dorsal, inner and ventral view). JK. Labrum (dorsal and frontal view). L. Right maxilla (dorsal view). M. Epipharynx. N. Labium." figureDoi="http://doi.org/10.5281/zenodo.4736141" httpUri="https://zenodo.org/record/4736141/files/figure.png" pageId="5" pageNumber="20">3B, E</figureCitation>
) .............................. .............................................................................................................................
<taxonomicName id="8EA39FD1FF8AFFB513FBA23AFC64F9C0" authority="Sousa &amp; Seidel, 2021" authorityName="Sousa &amp; Seidel" authorityYear="2021" box="[1159,1300,1549,1576]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="neglectus" status="sp. nov.">
<emphasis id="7BD73840FF8AFFB513FBA23AFC64F9C0" bold="true" box="[1159,1300,1549,1576]" italics="true" pageId="5" pageNumber="20">I. neglectus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60E4853BFF8AFFB51260A23AFC07F9C0" box="[1308,1399,1550,1576]" pageId="5" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="F257FCF7FF8AFFB517C1A264FC06F93E" pageId="5" pageNumber="20">
<paragraph id="491CE452FF8AFFB517C1A264FC06F966" blockId="5.[189,1399,265,2025]" pageId="5" pageNumber="20">
<keyLead id="F2524767FF8AFFB517C1A264FC06F966" pageId="5" pageNumber="20">
14. Pronotum and pygidium densely maculate without yellow longitudinal bands.
<collectingCountry id="31B4A4C2FF8AFFB513DEA264FD9DF982" box="[1186,1261,1616,1642]" name="Brazil" pageId="5" pageNumber="20">Brazil</collectingCountry>
,
<collectingCountry id="31B4A4C2FF8AFFB51386A264FC02F982" box="[1274,1394,1616,1642]" name="Argentina" pageId="5" pageNumber="20">Argentina</collectingCountry>
,
<collectingCountry id="31B4A4C2FF8AFFB51796A240F827F966" box="[234,343,1652,1678]" name="Paraguay" pageId="5" pageNumber="20">Paraguay</collectingCountry>
(
<figureCitation id="D198F8D7FF8AFFB51614A240F8CEF966" box="[360,446,1652,1678]" captionStart="Fig" captionStartId="14.[189,232,1823,1849]" captionTargetBox="[201,1387,266,1779]" captionTargetId="figure-125@14.[200,1387,265,1779]" captionTargetPageId="14" captionText="Fig. 4. Images of liveInca spp.A. Inca axelisp. nov. (photo by R. Barros and R. Koike).B. Inca neglectus sp. nov. (photo by N. Marchito). C. Inca clathratus sommeri Westwood, 1844 (photo by A. González Gallina). D. Inca clathratus clathratus (Olivier, 1792) (photo by R. Barros and R. Koike). E. Inca clathratus quesneli Boos &amp; Ratcliffe, 1985 (photo by M. Hulme). F. Inca pulverulentus (Olivier, 1789) mating (photo by R. Barros and R. Koike). G. Inca bonplandi (Gyllenhal, 1817) (photo by J. Touroult). H. Inca besckii Burmeister &amp; Schaum, 1840 feeding on tree sap (photo by N. Wisnik)." figureDoi="http://doi.org/10.5281/zenodo.4736143" httpUri="https://zenodo.org/record/4736143/files/figure.png" pageId="5" pageNumber="20">Fig. 4F</figureCitation>
) ............................................................................
<taxonomicName id="8EA39FD1FF8AFFB51482A240FC06F966" authority="(Olivier, 1789)" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[1022,1398,1651,1678]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="pulverulentus">
<emphasis id="7BD73840FF8AFFB51482A240FDB0F966" bold="true" box="[1022,1216,1651,1678]" italics="true" pageId="5" pageNumber="20">I. pulverulentus</emphasis>
(Olivier, 1789)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="491CE452FF8AFFB517BDA2ACFC06F93E" blockId="5.[189,1399,265,2025]" pageId="5" pageNumber="20">
<keyLead id="F2524767FF8AFFB517BDA2ACFC06F93E" pageId="5" pageNumber="20">
Pronotum and pygidium weakly maculate with medial and lateral areas with yellow longitudinal bands (
<taxonomicName id="8EA39FD1FF8AFFB5163CA288F8A0F93D" baseAuthorityName="Olivier" baseAuthorityYear="1792" box="[320,464,1723,1750]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="clathratus">
<emphasis id="7BD73840FF8AFFB5163CA288F83AF93D" box="[320,330,1724,1749]" italics="true" pageId="5" pageNumber="20">I</emphasis>
.
<emphasis id="7BD73840FF8AFFB51624A28FF8A0F93D" box="[344,464,1723,1749]" italics="true" pageId="5" pageNumber="20">clathratus</emphasis>
</taxonomicName>
)......................................................................................................................... 15
</keyLead>
</paragraph>
</keyStep>
<keyStep id="F257FCF7FF8AFFB517C1A2C9FC07F8BA" pageId="5" pageNumber="20">
<paragraph id="491CE452FF8AFFB517C1A2C9FC07F8D3" blockId="5.[189,1399,265,2025]" pageId="5" pageNumber="20">
<keyLead id="F2524767FF8AFFB517C1A2C9FC07F8D3" pageId="5" pageNumber="20">
15. Widely distributed in
<collectingCountry id="31B4A4C2FF8AFFB51696A2C9FB34F8FF" box="[490,580,1789,1815]" name="Mexico" pageId="5" pageNumber="20">Mexico</collectingCountry>
and Central America and restricted to South America on the east side of the Andes Mountains in
<collectingCountry id="31B4A4C2FF8AFFB51555A315FBEDF8D3" box="[553,669,1825,1851]" name="Colombia" pageId="5" pageNumber="20">Colombia</collectingCountry>
and
<collectingCountry id="31B4A4C2FF8AFFB515AAA315FA48F8D3" box="[726,824,1825,1851]" name="Ecuador" pageId="5" pageNumber="20">Ecuador</collectingCountry>
..............
<taxonomicName id="8EA39FD1FF8AFFB514D8A315FC07F8D3" authority="Westwood, 1844" authorityName="Westwood" authorityYear="1844" box="[932,1399,1824,1851]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="subSpecies" species="clathratus" subSpecies="sommeri">
<emphasis id="7BD73840FF8AFFB514D8A315FDDBF8D3" bold="true" box="[932,1195,1824,1851]" italics="true" pageId="5" pageNumber="20">I. clathratus sommeri</emphasis>
Westwood, 1844
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="491CE452FF8AFFB517BDA371FC07F8BA" blockId="5.[189,1399,265,2025]" box="[193,1399,1860,1887]" pageId="5" pageNumber="20">
<keyLead id="F2524767FF8AFFB517BDA371FC07F8BA" box="[193,1399,1860,1887]" pageId="5" pageNumber="20">
Distributed in South America, on the west side of the Andes Mountains ...................................... 16
<superScript id="BED6491AFF8AFFB51212A370FC07F8BA" attach="none" box="[1390,1399,1860,1874]" fontSize="6" pageId="5" pageNumber="20">*</superScript>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="F257FCF7FF8AFFB517C1A3B3FC06F800" pageId="5" pageNumber="20">
<paragraph id="491CE452FF8AFFB517C1A3B3FC07F84A" blockId="5.[189,1399,265,2025]" box="[189,1399,1926,1954]" pageId="5" pageNumber="20">
<keyLead id="F2524767FF8AFFB517C1A3B3FC07F84A" box="[189,1399,1926,1954]" pageId="5" pageNumber="20">
16. Restricted to
<collectingCountry id="31B4A4C2FF8AFFB516FAA3B3F89AF849" box="[390,490,1927,1953]" name="Trinidad and Tobago" pageId="5" pageNumber="20">Trinidad</collectingCountry>
.................................................
<taxonomicName id="8EA39FD1FF8AFFB51427A3B3FC07F84A" authority="Boos &amp; Ratcliffe, 1985" authorityName="Boos &amp; Ratcliffe" authorityYear="1985" box="[859,1399,1926,1954]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="subSpecies" species="clathratus" subSpecies="quesneli">
<emphasis id="7BD73840FF8AFFB51427A3B3FD2FF849" bold="true" box="[859,1119,1926,1953]" pageId="5" pageNumber="20">
<emphasis id="7BD73840FF8AFFB51427A3B3FA16F849" bold="true" box="[859,870,1927,1953]" italics="true" pageId="5" pageNumber="20">I</emphasis>
.
<emphasis id="7BD73840FF8AFFB5140AA3B2FD2FF849" bold="true" box="[886,1119,1926,1953]" italics="true" pageId="5" pageNumber="20">clathratus quesneli</emphasis>
</emphasis>
<bibRefCitation id="2D3299A3FF8AFFB5131BA3B3FC07F84A" author="Boos J. &amp; Ratcliffe B. C." box="[1127,1399,1927,1954]" pageId="5" pageNumber="20" pagination="381 - 389" refId="ref9385" refString="Boos J. &amp; Ratcliffe B. C. 1985. A new subspecies of Inca clathrata from Trinidad, West Indies, and range extensions for Inca clathrata. The Coleopterists Bulletin 39: 381 - 389." type="journal article" year="1985">Boos &amp; Ratcliffe, 1985</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="491CE452FF8AFFB517BDA39FFC06F800" blockId="5.[189,1399,265,2025]" pageId="5" pageNumber="20">
<keyLead id="F2524767FF8AFFB517BDA39FFC06F800" pageId="5" pageNumber="20">
Distributed in
<collectingCountry id="31B4A4C2FF8AFFB516C8A39EFB5BF82C" box="[436,555,1962,1988]" name="Colombia" pageId="5" pageNumber="20">Colombia</collectingCountry>
,
<collectingCountry id="31B4A4C2FF8AFFB5153BA39EFBE2F82C" box="[583,658,1962,1988]" name="Brazil" pageId="5" pageNumber="20">Brazil</collectingCountry>
,
<collectingCountry id="31B4A4C2FF8AFFB515D1A39EFA61F82C" box="[685,785,1962,1988]" name="Ecuador" pageId="5" pageNumber="20">Ecuador</collectingCountry>
,
<collectingCountry id="31B4A4C2FF8AFFB51451A39FFAFBF82D" box="[813,907,1963,1989]" name="Guyana" pageId="5" pageNumber="20">Guyana</collectingCountry>
, French
<collectingCountry id="31B4A4C2FF8AFFB5136CA39FFD1EF82D" box="[1040,1134,1963,1989]" name="Guyana" pageId="5" pageNumber="20">Guyana</collectingCountry>
,
<collectingCountry id="31B4A4C2FF8AFFB513F6A39FFDB3F82D" box="[1162,1219,1963,1989]" name="Peru" pageId="5" pageNumber="20">Peru</collectingCountry>
,
<collectingCountry id="31B4A4C2FF8AFFB513A2A39EFC44F82C" box="[1246,1332,1962,1988]" name="Bolivia" pageId="5" pageNumber="20">Bolivia</collectingCountry>
and
<collectingCountry id="31B4A4C2FF8AFFB51796A3FAF82FF800" box="[234,351,1998,2024]" name="Argentina" pageId="5" pageNumber="20">Argentina</collectingCountry>
..............................................................................
<taxonomicName id="8EA39FD1FF8AFFB514DBA3FBFC06F800" authority="clathratus (Olivier, 1792)" authorityName="clathratus (Olivier" authorityYear="1792" baseAuthorityName="Olivier" baseAuthorityYear="1792" box="[935,1398,1998,2025]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="subSpecies" species="clathratus" subSpecies="clathratus">
<emphasis id="7BD73840FF8AFFB514DBA3FBFDB1F801" bold="true" box="[935,1217,1998,2025]" italics="true" pageId="5" pageNumber="20">I. clathratus clathratus</emphasis>
(Olivier, 1792)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="F257FCF7FF89FFB617C1A53DFC06FE82" pageId="6" pageNumber="21">
<paragraph id="491CE452FF89FFB617C1A53DFC07FEAF" blockId="6.[189,1399,265,363]" pageId="6" pageNumber="21">
<keyLead id="F2524767FF89FFB617C1A53DFC07FEAF" pageId="6" pageNumber="21">
17. Legs without red reflections.
<collectingCountry id="31B4A4C2FF89FFB61538A53DFBFCFECB" box="[580,652,265,291]" name="Brazil" pageId="6" pageNumber="21">Brazil</collectingCountry>
(
<collectingRegion id="8B672AB0FF89FFB615E0A53DFA4EFECB" box="[668,830,265,291]" country="Brazil" name="Minas Gerais" pageId="6" pageNumber="21">Minas Gerais</collectingRegion>
,
<collectingRegion id="8B672AB0FF89FFB61437A53DFA8AFECB" box="[843,1018,265,291]" country="Brazil" name="Rio de Janeiro" pageId="6" pageNumber="21">Rio de Janeiro</collectingRegion>
,
<collectingRegion id="8B672AB0FF89FFB6137AA53DFD29FECB" box="[1030,1113,265,291]" country="Brazil" name="Parana" pageId="6" pageNumber="21">Paraná</collectingRegion>
,
<collectingRegion id="8B672AB0FF89FFB61319A53DFC61FECB" box="[1125,1297,265,291]" country="Brazil" name="Santa Catarina" pageId="6" pageNumber="21">
Santa
<taxonomicName id="8EA39FD1FF89FFB613D2A53DFC61FECB" ID-CoL="94MGZ" authorityName="Vojnits &amp; de Laever" authorityYear="1973" box="[1198,1297,265,291]" class="Insecta" family="Geometridae" genus="Catarina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="22" phylum="Arthropoda" rank="genus">Catarina</taxonomicName>
</collectingRegion>
and
<collectingRegion id="8B672AB0FF89FFB61230A53DF8E6FEAF" country="Brazil" name="Rio Grande do Sul" pageId="6" pageNumber="21">Rio Grande do Sul</collectingRegion>
) (
<figureCitation id="D198F8D7FF89FFB616D2A518FB79FEAF" box="[430,521,300,327]" captionStart="Fig" captionStartId="14.[189,232,1823,1849]" captionTargetBox="[201,1387,266,1779]" captionTargetId="figure-125@14.[200,1387,265,1779]" captionTargetPageId="14" captionText="Fig. 4. Images of liveInca spp.A. Inca axelisp. nov. (photo by R. Barros and R. Koike).B. Inca neglectus sp. nov. (photo by N. Marchito). C. Inca clathratus sommeri Westwood, 1844 (photo by A. González Gallina). D. Inca clathratus clathratus (Olivier, 1792) (photo by R. Barros and R. Koike). E. Inca clathratus quesneli Boos &amp; Ratcliffe, 1985 (photo by M. Hulme). F. Inca pulverulentus (Olivier, 1789) mating (photo by R. Barros and R. Koike). G. Inca bonplandi (Gyllenhal, 1817) (photo by J. Touroult). H. Inca besckii Burmeister &amp; Schaum, 1840 feeding on tree sap (photo by N. Wisnik)." figureDoi="http://doi.org/10.5281/zenodo.4736143" httpUri="https://zenodo.org/record/4736143/files/figure.png" pageId="6" pageNumber="21">Fig. 4H</figureCitation>
) ........................................................
<taxonomicName id="8EA39FD1FF89FFB614CDA519FC07FEAF" authority="Burmeister &amp; Schaum, 1840" authorityName="Burmeister &amp; Schaum" authorityYear="1840" box="[945,1399,300,327]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="besckii">
<emphasis id="7BD73840FF89FFB614CDA519FD6FFEAF" bold="true" box="[945,1055,300,327]" italics="true" pageId="6" pageNumber="21">I. besckii</emphasis>
<bibRefCitation id="2D3299A3FF89FFB6135BA518FC07FEAF" author="Burmeister H. &amp; Schaum H." box="[1063,1399,300,327]" pageId="6" pageNumber="21" pagination="353 - 420" refId="ref9569" refString="Burmeister H. &amp; Schaum H. 1840. Kritische Revision der Lamellicornia melitophila. Erstes Stuck (Trichiadae). Zeitschrift fur die Entomologie 2: 353 - 420." type="journal article" year="1840">Burmeister &amp; Schaum, 1840</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="491CE452FF89FFB617BDA565FC06FE82" blockId="6.[189,1399,265,363]" box="[193,1398,336,363]" pageId="6" pageNumber="21">
<keyLead id="F2524767FF89FFB617BDA565FC06FE82" box="[193,1398,336,363]" pageId="6" pageNumber="21">
Legs with red reflections.
<collectingCountry id="31B4A4C2FF89FFB61566A564FBE2FE82" box="[538,658,336,362]" name="Argentina" pageId="6" pageNumber="21">Argentina</collectingCountry>
,
<collectingCountry id="31B4A4C2FF89FFB615E2A564FB87FE82" box="[670,759,336,362]" name="Bolivia" pageId="6" pageNumber="21">Bolivia</collectingCountry>
,
<collectingCountry id="31B4A4C2FF89FFB6147FA564FA3BFE82" box="[771,843,336,362]" name="Brazil" pageId="6" pageNumber="21">Brazil</collectingCountry>
and
<collectingCountry id="31B4A4C2FF89FFB614FFA565FA80FE83" box="[899,1008,337,363]" name="Paraguay" pageId="6" pageNumber="21">Paraguay</collectingCountry>
...
<taxonomicName id="8EA39FD1FF89FFB61374A564FC06FE82" authority="(Gyllenhal, 1817)" baseAuthorityName="Gyllenhal" baseAuthorityYear="1817" box="[1032,1398,336,363]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="bonplandi">
<emphasis id="7BD73840FF89FFB61374A564FDEEFE83" bold="true" box="[1032,1182,336,363]" italics="true" pageId="6" pageNumber="21">I. bonplandi</emphasis>
(Gyllenhal, 1817)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
<subSubSection id="01B9B7D9FF89FFB617C1A5A5FD6FFDD2" pageId="6" pageNumber="21" type="discussion">
<paragraph id="491CE452FF89FFB617C1A5A5FD6FFDD2" blockId="6.[188,1399,401,571]" pageId="6" pageNumber="21">
<superScript id="BED6491AFF89FFB617C1A5A5F9B6FE77" attach="right" box="[189,198,401,415]" fontSize="6" pageId="6" pageNumber="21">*</superScript>
Step 16 separates the females of
<taxonomicName id="8EA39FD1FF89FFB6153EA5A7FBA3FE44" baseAuthorityName="Olivier" baseAuthorityYear="1792" box="[578,723,402,428]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" sensu="stricto" species="clathratus">
<emphasis id="7BD73840FF89FFB6153EA5A7FB3CFE44" box="[578,588,403,428]" italics="true" pageId="6" pageNumber="21">I</emphasis>
.
<emphasis id="7BD73840FF89FFB61527A5A6FBA3FE44" box="[603,723,402,428]" italics="true" pageId="6" pageNumber="21">clathratus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60E4853BFF89FFB615A6A5A6FA67FE44" box="[730,791,402,428]" pageId="6" pageNumber="21" sensu="stricto">s. str.</taxonomicNameLabel>
to
<taxonomicName id="8EA39FD1FF89FFB61447A5A7FD46FE44" authorityName="Boos &amp; Ratcliffe" authorityYear="1985" box="[827,1078,402,428]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="subSpecies" species="clathratus" subSpecies="quesneli">
<emphasis id="7BD73840FF89FFB61447A5A7FA35FE44" box="[827,837,403,428]" italics="true" pageId="6" pageNumber="21">I</emphasis>
.
<emphasis id="7BD73840FF89FFB6142FA5A6FD46FE44" box="[851,1078,402,428]" italics="true" pageId="6" pageNumber="21">clathratus quesneli</emphasis>
</taxonomicName>
. However, the morphology of these species is not informative for the distinction of these taxa. Beyond geographic distribution the females of
<taxonomicName id="8EA39FD1FF89FFB6163FA5EEFB30FE1B" authorityName="Boos &amp; Ratcliffe" authorityYear="1985" box="[323,576,473,500]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="subSpecies" species="clathratus" subSpecies="quesneli">
<emphasis id="7BD73840FF89FFB6163FA5EEF83CFE1B" box="[323,332,474,499]" italics="true" pageId="6" pageNumber="21">I</emphasis>
.
<emphasis id="7BD73840FF89FFB61620A5EDFB30FE1B" box="[348,576,473,499]" italics="true" pageId="6" pageNumber="21">clathratus quesneli</emphasis>
</taxonomicName>
are usually bigger than
<taxonomicName id="8EA39FD1FF89FFB6141EA5EEFA82FE1B" baseAuthorityName="Olivier" baseAuthorityYear="1792" box="[866,1010,473,500]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" sensu="stricto" species="clathratus">
<emphasis id="7BD73840FF89FFB6141EA5EEFA1BFE1B" box="[866,875,474,499]" italics="true" pageId="6" pageNumber="21">I</emphasis>
.
<emphasis id="7BD73840FF89FFB61406A5EDFA82FE1B" box="[890,1010,473,499]" italics="true" pageId="6" pageNumber="21">clathratus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60E4853BFF89FFB61487A5EEFD4AFE1C" box="[1019,1082,474,500]" pageId="6" pageNumber="21" sensu="stricto">s. str.</taxonomicNameLabel>
and can have bluish black tegument coloration beyond the dark green, black or brownish black coloration of
<taxonomicName id="8EA39FD1FF89FFB613E1A5C9FC5EFDFF" baseAuthorityName="Olivier" baseAuthorityYear="1792" box="[1181,1326,509,535]" class="Insecta" family="Cetoniidae" genus="Inca" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" sensu="stricto" species="clathratus">
<emphasis id="7BD73840FF89FFB613E1A5C9FDD5FDFE" box="[1181,1189,509,534]" italics="true" pageId="6" pageNumber="21">I</emphasis>
.
<emphasis id="7BD73840FF89FFB613CAA5C9FC5EFDFF" box="[1206,1326,509,535]" italics="true" pageId="6" pageNumber="21">clathratus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60E4853BFF89FFB6124BA5C9FC07FDFF" box="[1335,1399,509,535]" pageId="6" pageNumber="21" sensu="stricto">s. str.</taxonomicNameLabel>
Further studies are needed to better clarify the taxonomy of these species.
</paragraph>
</subSubSection>
</treatment>
</document>