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<document id="182AC7693267AFC02DCA15DDE541A8BD" ID-DOI="10.5281/zenodo.273742" ID-GBIF-Dataset="69846e24-baf5-4d82-b776-64331168c4c7" ID-ISSN="1175-5326" ID-Zenodo-Dep="273742" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459836927901" checkinUser="plazi" docAuthor="Michelsen, Verner" docDate="2007" docId="995687C62B3A510DAEA3FE713369DBE3" docLanguage="en" docName="zt01469p057.pdf" docOrigin="Zootaxa 1469" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Delia cardui" docType="treatment" docVersion="8" lastPageNumber="52" masterDocId="656FFFBE2B3B510CAE34FF8E3636DC57" masterDocTitle="Two new European species of Delia Robineau-Desvoidy (Diptera: Anthomyiidae) with a bipartite male sternite III" masterLastPageNumber="57" masterPageNumber="51" pageNumber="52" updateTime="1698223783679" updateUser="plazi">
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<mods:title id="DA618F97054B6A72579E78AC97F2F2D3">Two new European species of Delia Robineau-Desvoidy (Diptera: Anthomyiidae) with a bipartite male sternite III</mods:title>
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<mods:namePart id="84AFF17650FE60E779A9AB56F40E4EAA">Michelsen, Verner</mods:namePart>
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<mods:date id="5BAA6A3EA5F6FCE57601AAFBF18502AB">2007</mods:date>
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<treatment id="995687C62B3A510DAEA3FE713369DBE3" ID-DOI="http://doi.org/10.5281/zenodo.6248988" ID-GBIF-Taxon="119359202" ID-Zenodo-Dep="6248988" LSID="urn:lsid:plazi:treatment:995687C62B3A510DAEA3FE713369DBE3" httpUri="http://treatment.plazi.org/id/995687C62B3A510DAEA3FE713369DBE3" lastPageNumber="52" pageId="1" pageNumber="52">
<subSubSection id="59E5655B2B3A510DAEA3FE7132BCDFF3" pageId="1" pageNumber="52" type="nomenclature">
<paragraph id="114036D02B3A510DAEA3FE713423DE4E" blockId="1.[151,533,511,537]" box="[151,533,511,537]" pageId="1" pageNumber="52">
<heading id="4A0881BC2B3A510DAEA3FE713423DE4E" bold="true" box="[151,533,511,537]" fontSize="11" level="1" pageId="1" pageNumber="52" reason="1">
<emphasis id="238BEAC22B3A510DAEA3FE713423DE4E" bold="true" box="[151,533,511,537]" pageId="1" pageNumber="52">
The
<taxonomicName id="D6FF4D532B3A510DAEFBFE713753DE4E" ID-CoL="92QZV" box="[207,357,511,537]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="cardui">
<emphasis id="238BEAC22B3A510DAEFBFE713753DE4E" bold="true" box="[207,357,511,537]" italics="true" pageId="1" pageNumber="52">Delia cardui</emphasis>
</taxonomicName>
species group
</emphasis>
</heading>
</paragraph>
<paragraph id="114036D02B3A510DAEA3FDC432BCDFF3" blockId="1.[151,1437,586,1972]" pageId="1" pageNumber="52">
The two new species described below have a ventro-basal thickening on mid tarsomere
<quantity id="D6079B352B3A510DAAACFDC432F2DE33" box="[1176,1220,586,612]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="1" pageNumber="52" unit="in" value="2.0">2 in</quantity>
the male sex (
<figureCitation id="89C42A552B3A510DAB59FDC43692DEDB" captionStart="FIGURES 1 3" captionStartId="3.[151,269,1892,1916]" captionTargetBox="[260,1331,488,1891]" captionTargetId="figure@3.[257,1331,441,1896]" captionTargetPageId="3" captionText="FIGURES 1 3. Delia rimiventris, male. 1. Right mid tarsus. Arrow points at basal thickening of tarsomere 2. 2. Tergites I VI, dorsal. 3. Tergites I VI, left lateral. Different scales for 1 and 2 3." httpUri="https://zenodo.org/record/273743/files/figure.png" pageId="1" pageNumber="52">Fig. 1</figureCitation>
), a derived character used by Hennig (1974a) to define a
<taxonomicName id="D6FF4D532B3A510DAD79FDFC35D6DEDC" box="[845,992,626,651]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="cardui">
<emphasis id="238BEAC22B3A510DAD79FDFC35D6DEDC" box="[845,992,626,651]" italics="true" pageId="1" pageNumber="52">Delia cardui</emphasis>
</taxonomicName>
species group. The following species described from the western Palaearctic region (see Hennig 1974a, b, c) share this character:
<taxonomicName id="D6FF4D532B3A510DAAD0FD14371DDE8B" authority="Zetterstedt" authorityName="Zetterstedt" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="brunnescens">
<emphasis id="238BEAC22B3A510DAAD0FD1433AADEE4" box="[1252,1436,666,691]" italics="true" pageId="1" pageNumber="52">D. brunnescens</emphasis>
(Zetterstedt)
</taxonomicName>
,
<taxonomicName id="D6FF4D532B3A510DAF01FD4C342DDE8B" authority="Meigen" authorityName="Meigen" box="[309,539,706,732]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="cardui">
<emphasis id="238BEAC22B3A510DAF01FD4C3795DE8C" box="[309,419,706,731]" italics="true" pageId="1" pageNumber="52">D. cardui</emphasis>
(Meigen)
</taxonomicName>
,
<taxonomicName id="D6FF4D532B3A510DAC12FD4C34C1DE8B" authority="Hennig" authorityName="Hennig" box="[550,759,706,732]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="crinita">
<emphasis id="238BEAC22B3A510DAC12FD4C34A1DE8C" box="[550,663,706,731]" italics="true" pageId="1" pageNumber="52">D. crinita</emphasis>
Hennig
</taxonomicName>
,
<taxonomicName id="D6FF4D532B3A510DAD36FD4C3271DE8B" authority="Zetterstedt" authorityName="Zetterstedt" box="[770,1095,706,732]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="criniventris">
<emphasis id="238BEAC22B3A510DAD36FD4C359ADE8C" box="[770,940,706,731]" italics="true" pageId="1" pageNumber="52">D. criniventris</emphasis>
(Zetterstedt)
</taxonomicName>
,
<taxonomicName id="D6FF4D532B3A510DAA66FD4C36D0DF53" authority="Robineau-Desvoidy" authorityName="Robineau-Desvoidy" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="floricola">
<emphasis id="238BEAC22B3A510DAA66FD4C32ECDE8C" box="[1106,1242,706,731]" italics="true" pageId="1" pageNumber="52">D. floricola</emphasis>
(Robineau-Desvoidy)
</taxonomicName>
,
<taxonomicName id="D6FF4D532B3A510DAEC1FD643788DF53" authority="Strobl" authorityName="Strobl" box="[245,446,746,772]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="nuda">
<emphasis id="238BEAC22B3A510DAEC1FD643760DF54" box="[245,342,746,771]" italics="true" pageId="1" pageNumber="52">D. nuda</emphasis>
(Strobl)
</taxonomicName>
,
<taxonomicName id="D6FF4D532B3A510DAFFAFD6434C8DF53" authority="Rondani" authorityName="Rondani" box="[462,766,746,772]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="penicillaris">
<emphasis id="238BEAC22B3A510DAFFAFD64344DDF54" box="[462,635,746,771]" italics="true" pageId="1" pageNumber="52">D. penicillaris</emphasis>
(Rondani)
</taxonomicName>
,
<taxonomicName id="D6FF4D532B3A510DAD3AFD64322FDF53" authority="Hennig" authorityName="Hennig" box="[782,1049,746,772]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="penicillosa">
<emphasis id="238BEAC22B3A510DAD3AFD643580DF54" box="[782,950,746,771]" italics="true" pageId="1" pageNumber="52">D. penicillosa</emphasis>
Hennig
</taxonomicName>
,
<taxonomicName id="D6FF4D532B3A510DAA1CFD64333FDF53" authority="Hennig" authorityName="Hennig" box="[1064,1289,746,772]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="persica">
<emphasis id="238BEAC22B3A510DAA1CFD643293DF54" box="[1064,1189,746,771]" italics="true" pageId="1" pageNumber="52">D. persica</emphasis>
Hennig
</taxonomicName>
,
<taxonomicName id="D6FF4D532B3A510DAB2CFD6436D0DF7B" authority="Stein" authorityName="Stein" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="pilitibia">
<emphasis id="238BEAC22B3A510DAB2CFD6433AADF54" box="[1304,1436,746,771]" italics="true" pageId="1" pageNumber="52">D. pilitibia</emphasis>
(Stein)
</taxonomicName>
and
<taxonomicName id="D6FF4D532B3A510DAF2AFC9C340FDF7B" authority="Strobl" authorityName="Strobl" box="[286,569,786,812]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="pseudofugax">
<emphasis id="238BEAC22B3A510DAF2AFC9C37E3DF7C" box="[286,469,786,811]" italics="true" pageId="1" pageNumber="52">D. pseudofugax</emphasis>
(Strobl)
</taxonomicName>
. A few species in this list are further recorded from
<collectingCountry id="69E876402B3A510DAAA1FC9C32EADF7B" box="[1173,1244,786,812]" name="Japan" pageId="1" pageNumber="52">Japan</collectingCountry>
:
<taxonomicName id="D6FF4D532B3A510DAAD2FC9C3704DF03" authority="Suwa 1999" authorityName="Suwa" authorityYear="1999" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="brunnescens">
<emphasis id="238BEAC22B3A510DAAD2FC9C33AADF7C" box="[1254,1436,786,811]" italics="true" pageId="1" pageNumber="52">D. brunnescens</emphasis>
(Suwa 1999)
</taxonomicName>
,
<collectingCountry id="69E876402B3A510DAF74FCB437BDDF03" box="[320,395,826,852]" name="China" pageId="1" pageNumber="52">China</collectingCountry>
:
<taxonomicName id="D6FF4D532B3A510DAFACFCB4343FDF04" box="[408,521,826,851]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="cardui">
<emphasis id="238BEAC22B3A510DAFACFCB4343FDF04" box="[408,521,826,851]" italics="true" pageId="1" pageNumber="52">D. cardui</emphasis>
</taxonomicName>
,
<taxonomicName id="D6FF4D532B3A510DAC2EFCB434F3DF04" box="[538,709,826,851]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="penicillaris">
<emphasis id="238BEAC22B3A510DAC2EFCB434F3DF04" box="[538,709,826,851]" italics="true" pageId="1" pageNumber="52">D. penicillaris</emphasis>
</taxonomicName>
and
<taxonomicName id="D6FF4D532B3A510DAD36FCB43248DF03" authority="Wei et al. 1999" authorityName="Wei et al." authorityYear="1999" box="[770,1150,826,852]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="penicillosa">
<emphasis id="238BEAC22B3A510DAD36FCB4359FDF04" box="[770,937,826,851]" italics="true" pageId="1" pageNumber="52">D. penicillosa</emphasis>
(Wei
<emphasis id="238BEAC22B3A510DADC0FCB43207DF04" box="[1012,1073,826,851]" italics="true" pageId="1" pageNumber="52">et al.</emphasis>
1999)
</taxonomicName>
and North
<collectingCountry id="69E876402B3A510DAB3DFCB43345DF03" box="[1289,1395,826,852]" name="United States of America" pageId="1" pageNumber="52">America</collectingCountry>
:
<taxonomicName id="D6FF4D532B3A510DAB4BFCB436D4DF2C" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="cardui">
<emphasis id="238BEAC22B3A510DAB4BFCB436D4DF2C" italics="true" pageId="1" pageNumber="52">D. cardui</emphasis>
</taxonomicName>
and
<taxonomicName id="D6FF4D532B3A510DAF28FCEC3478DF2B" authority="(Griffiths 1991a)" baseAuthorityName="Griffiths" baseAuthorityYear="1991" box="[284,590,866,892]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="nuda">
<emphasis id="238BEAC22B3A510DAF28FCEC374ADF2C" box="[284,380,866,891]" italics="true" pageId="1" pageNumber="52">D. nuda</emphasis>
(Griffiths 1991a)
</taxonomicName>
. Species apparently endemic to
<collectingCountry id="69E876402B3A510DADFDFCEC320FDF2B" box="[969,1081,866,892]" name="China" pageId="1" pageNumber="52">East Asia</collectingCountry>
are
<taxonomicName id="D6FF4D532B3A510DAA58FCEC3376DF2B" authority="Suwa" authorityName="Suwa" box="[1132,1344,866,892]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="bipartita">
<emphasis id="238BEAC22B3A510DAA58FCEC32C1DF2C" box="[1132,1271,866,891]" italics="true" pageId="1" pageNumber="52">D. bipartita</emphasis>
Suwa
</taxonomicName>
(
<collectingCountry id="69E876402B3A510DAB64FCEC33A1DF2B" box="[1360,1431,866,892]" name="Japan" pageId="1" pageNumber="52">Japan</collectingCountry>
,
<collectingCountry id="69E876402B3A510DAEA3FC0436E9DFF3" box="[151,223,906,932]" name="China" pageId="1" pageNumber="52">China</collectingCountry>
) and
<taxonomicName id="D6FF4D532B3A510DAF2BFC043460DFF3" authority="Fan" authorityName="Fan" box="[287,598,906,932]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="penicilella">
<emphasis id="238BEAC22B3A510DAF2BFC043789DFF4" box="[287,447,906,931]" italics="true" pageId="1" pageNumber="52">D. penicilella</emphasis>
Fan (
<collectingCountry id="69E876402B3A510DAC36FC043478DFF3" box="[514,590,906,932]" name="China" pageId="1" pageNumber="52">China</collectingCountry>
)
</taxonomicName>
, while
<taxonomicName id="D6FF4D532B3A510DAC9EFC0435BADFF3" authority="Griffiths" authorityName="Griffiths" box="[682,908,906,932]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="polaris">
<emphasis id="238BEAC22B3A510DAC9EFC043517DFF4" box="[682,801,906,931]" italics="true" pageId="1" pageNumber="52">D. polaris</emphasis>
Griffiths
</taxonomicName>
is a Nearctic species.
</paragraph>
</subSubSection>
<subSubSection id="59E5655B2B3A510DAEF3FC3C3369DBE3" pageId="1" pageNumber="52" type="description">
<paragraph id="114036D02B3A510DAEF3FC3C34B7D8B3" blockId="1.[151,1437,586,1972]" pageId="1" pageNumber="52">
In addition to the tarsal character of the
<taxonomicName id="D6FF4D532B3A510DAC83FC3C3578DF9C" box="[695,846,946,971]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="cardui">
<emphasis id="238BEAC22B3A510DAC83FC3C3578DF9C" box="[695,846,946,971]" italics="true" pageId="1" pageNumber="52">Delia cardui</emphasis>
</taxonomicName>
species group, both new species have strikingly short, posteriorly excavated male tergites IIIIV separated from each other and from tergites II and V by relatively wide areas of flexible inter-segmental cuticle (
<figureCitation id="89C42A552B3A510DAD3BFB8C356FD84B" box="[783,857,1026,1052]" captionStart="FIGURES 1 3" captionStartId="3.[151,269,1892,1916]" captionTargetBox="[260,1331,488,1891]" captionTargetId="figure@3.[257,1331,441,1896]" captionTargetPageId="3" captionText="FIGURES 1 3. Delia rimiventris, male. 1. Right mid tarsus. Arrow points at basal thickening of tarsomere 2. 2. Tergites I VI, dorsal. 3. Tergites I VI, left lateral. Different scales for 1 and 2 3." httpUri="https://zenodo.org/record/273743/files/figure.png" pageId="1" pageNumber="52">Fig. 2</figureCitation>
, x). The exposed male abdominal intersegments are characteristic of nearly all species of the
<taxonomicName id="D6FF4D532B3A510DAC86FBA43571D814" box="[690,839,1066,1091]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="cardui">
<emphasis id="238BEAC22B3A510DAC86FBA43571D814" box="[690,839,1066,1091]" italics="true" pageId="1" pageNumber="52">Delia cardui</emphasis>
</taxonomicName>
species group, but are also present in some obviously closely related species with simple male mid tarsi,
<taxonomicName id="D6FF4D532B3A510DAD7CFBDD3263D83B" authority="Hennig" authorityName="Hennig" box="[840,1109,1106,1132]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="alatavensis">
<emphasis id="238BEAC22B3A510DAD7CFBDD35C5D83B" box="[840,1011,1107,1132]" italics="true" pageId="1" pageNumber="52">D. alatavensis</emphasis>
Hennig
</taxonomicName>
,
<taxonomicName id="D6FF4D532B3A510DAA50FBDD33AAD83B" authority="Schnabl" authorityName="Schnabl" box="[1124,1436,1106,1132]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="carduiformis">
<emphasis id="238BEAC22B3A510DAA50FBDD3316D83B" box="[1124,1312,1107,1132]" italics="true" pageId="1" pageNumber="52">D. carduiformis</emphasis>
(Schnabl)
</taxonomicName>
and
<taxonomicName id="D6FF4D532B3A510DAEFEFBF537DFD8C3" authority="Pokorny" authorityName="Pokorny" box="[202,489,1146,1172]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="piliventris">
<emphasis id="238BEAC22B3A510DAEFEFBF53750D8C3" box="[202,358,1147,1172]" italics="true" pageId="1" pageNumber="52">D. piliventris</emphasis>
(Pokorny)
</taxonomicName>
. The condition allows an upward flexure of the caudal part of the abdomen and resulting exposure of the variously specialized sternites IIIIV (see Hennig 1974a for further details and a suggestion on its functional significance).
</paragraph>
<paragraph id="114036D02B3A510DAEF3FB7C34FFD9FB" blockId="1.[151,1437,586,1972]" pageId="1" pageNumber="52">
Unfortunately, as pointed out by Hennig (1974a) and Griffiths (1991a), the male tarsal character is somewhat unstable and liable to secondary reduction. The same applies to the modifications of the male tergites IIIIV as described above. Firstly this character complex is barely apparent in
<taxonomicName id="D6FF4D532B3A510DAA07FACD3282D90B" box="[1075,1204,1347,1372]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="pilitibia">
<emphasis id="238BEAC22B3A510DAA07FACD3282D90B" box="[1075,1204,1347,1372]" italics="true" pageId="1" pageNumber="52">D. pilitibia</emphasis>
</taxonomicName>
of the
<taxonomicName id="D6FF4D532B3A510DAB3CFACD33AAD90B" box="[1288,1436,1347,1372]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="cardui">
<emphasis id="238BEAC22B3A510DAB3CFACD33AAD90B" box="[1288,1436,1347,1372]" italics="true" pageId="1" pageNumber="52">Delia cardui</emphasis>
</taxonomicName>
species group, and secondly the same character complex is seen in a moderated version in some more distantly related species, e.g.,
<taxonomicName id="D6FF4D532B3A510DAFE0FA1D34F3D9FB" authority="Seguy" authorityName="Seguy" box="[468,709,1426,1452]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="echinata">
<emphasis id="238BEAC22B3A510DAFE0FA1D346BD9FB" box="[468,605,1427,1452]" italics="true" pageId="1" pageNumber="52">D. echinata</emphasis>
(Séguy)
</taxonomicName>
.
</paragraph>
<paragraph id="114036D02B3A510DAEF3FA343361DABB" blockId="1.[151,1437,586,1972]" pageId="1" pageNumber="52">
Griffiths (1991a) abandoned the male mid tarsal character as the only criterion for inclusion in his
<taxonomicName id="D6FF4D532B3A510DAB6FFA3536D5D9AB" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="cardui">
<emphasis id="238BEAC22B3A510DAB6FFA3536D5D9AB" italics="true" pageId="1" pageNumber="52">Delia cardui</emphasis>
</taxonomicName>
superspecies by adding two species (
<taxonomicName id="D6FF4D532B3A510DAC80FA6D3569D9AB" box="[692,863,1507,1532]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="alatavensis">
<emphasis id="238BEAC22B3A510DAC80FA6D3569D9AB" box="[692,863,1507,1532]" italics="true" pageId="1" pageNumber="52">D. alatavensis</emphasis>
</taxonomicName>
,
<taxonomicName id="D6FF4D532B3A510DAD44FA6D3238D9AB" box="[880,1038,1507,1532]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="piliventris">
<emphasis id="238BEAC22B3A510DAD44FA6D3238D9AB" box="[880,1038,1507,1532]" italics="true" pageId="1" pageNumber="52">D. piliventris</emphasis>
</taxonomicName>
) without this character. Unfortunately, the integrity of the resulting superspecies against his
<taxonomicName id="D6FF4D532B3A510DAD44F9853214DA73" box="[880,1058,1547,1572]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="pruinosa">
<emphasis id="238BEAC22B3A510DAD44F9853214DA73" box="[880,1058,1547,1572]" italics="true" pageId="1" pageNumber="52">Delia pruinosa</emphasis>
</taxonomicName>
superspecies and
<taxonomicName id="D6FF4D532B3A510DAB3BF98533ADDA73" box="[1295,1435,1547,1572]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="fallax">
<emphasis id="238BEAC22B3A510DAB3BF98533ADDA73" box="[1295,1435,1547,1572]" italics="true" pageId="1" pageNumber="52">Delia fallax</emphasis>
</taxonomicName>
superspecies is rather ambiguous and needs some adjustment. I suggest the following diagnosis for the
<taxonomicName id="D6FF4D532B3A510DAB6FF9BD36D5DA23" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="cardui">
<emphasis id="238BEAC22B3A510DAB6FF9BD36D5DA23" italics="true" pageId="1" pageNumber="52">Delia cardui</emphasis>
</taxonomicName>
species group: male mid tarsomere 2 with a ventral swelling and/or male abdominal tergites III and IV distinctly shorter than tergite V, posteriorly excavated and separated by widely exposed intersegmental cuticle. By this diagnosis the limits of the
<taxonomicName id="D6FF4D532B3A510DAC68F92534C7DA93" box="[604,753,1707,1732]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="cardui">
<emphasis id="238BEAC22B3A510DAC68F92534C7DA93" box="[604,753,1707,1732]" italics="true" pageId="1" pageNumber="52">Delia cardui</emphasis>
</taxonomicName>
species group are slightly expanded to further include the Holarctic
<taxonomicName id="D6FF4D532B3A510DAF3FF95D37F3DABB" box="[267,453,1747,1772]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="carduiformis">
<emphasis id="238BEAC22B3A510DAF3FF95D37F3DABB" box="[267,453,1747,1772]" italics="true" pageId="1" pageNumber="52">D. carduiformis</emphasis>
</taxonomicName>
, a species that Griffiths (1991a) referred to his
<taxonomicName id="D6FF4D532B3A510DADCFF95D329ADABB" box="[1019,1196,1747,1772]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="pruinosa">
<emphasis id="238BEAC22B3A510DADCFF95D329ADABB" box="[1019,1196,1747,1772]" italics="true" pageId="1" pageNumber="52">Delia pruinosa</emphasis>
</taxonomicName>
superspecies.
</paragraph>
<paragraph id="114036D02B3A510DAEF3F9743369DBE3" blockId="1.[151,1437,586,1972]" pageId="1" pageNumber="52">
Available biological data suggest that the phytophagous larvae of the
<taxonomicName id="D6FF4D532B3A510DADCAF97532A5DB43" box="[1022,1171,1787,1812]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="cardui">
<emphasis id="238BEAC22B3A510DADCAF97532A5DB43" box="[1022,1171,1787,1812]" italics="true" pageId="1" pageNumber="52">Delia cardui</emphasis>
</taxonomicName>
species group all feed on leaves, shoots or floral parts of species of
<taxonomicName id="D6FF4D532B3A510DAC99F8AC3540DB6B" box="[685,886,1826,1852]" class="Magnoliopsida" family="Caryophyllaceae" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="52" phylum="Tracheophyta" rank="family">Caryophyllaceae</taxonomicName>
. However, as pointed out by Griffiths (1991a), a larval association with that plant family is characteristic of a more inclusive species group within
<taxonomicName id="D6FF4D532B3A510DAB6FF8C533AADB33" box="[1371,1436,1867,1892]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="238BEAC22B3A510DAB6FF8C533AADB33" box="[1371,1436,1867,1892]" italics="true" pageId="1" pageNumber="52">Delia</emphasis>
</taxonomicName>
denoted by Griffiths as the
<taxonomicName id="D6FF4D532B3A510DAFC4F8FD34BEDBDB" box="[496,648,1907,1932]" class="Insecta" family="Anthomyiidae" genus="Delia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="cardui">
<emphasis id="238BEAC22B3A510DAFC4F8FD34BEDBDB" box="[496,648,1907,1932]" italics="true" pageId="1" pageNumber="52">Delia cardui</emphasis>
</taxonomicName>
subsection. Morphologically, this subsection is characterized by more or less pronounced modifications in shape and/or vestiture of the male abdominal segments IIIIV.
</paragraph>
</subSubSection>
</treatment>
</document>