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183 lines
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<mods:title id="4AC7B09CEA3D708C88DFD2A093BB3BB3">Revision of Eccoptolonthus Bernhauer (Coleoptera: Staphylininae: Philonthina) with descriptions of four new species from China</mods:title>
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<mods:namePart id="0E002C4E671A6CAA990B4094FAA0DBAD">Fei, Xu-Dong</mods:namePart>
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<mods:affiliation id="E1CE28AB9BF2F4A86127D3BA3D03B349">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, 1 Beichen West Rd., Chaoyang District, Beijing 100101, P. R. China. & University of the Chinese Academy of Sciences, 19 A Yuquan Rd., Shijingshan District, Beijing 100049, P. R. China. & feixudong @ ioz. ac. cn; https: // orcid. org / 0000 - 0003 - 0386 - 5918</mods:affiliation>
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<mods:nameIdentifier id="3BAC788F71A8E19EFBD5E6FC194D1773" type="email">feixudong@ioz.ac.cn</mods:nameIdentifier>
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<mods:namePart id="772A9D604F0EB1BD8825F86B6F9FCA05">Zhou, Hong-Zhang</mods:namePart>
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<mods:affiliation id="97EF34C01B78B6EB0D8886BF6B2C90D1">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, 1 Beichen West Rd., Chaoyang District, Beijing 100101, P. R. China. & University of the Chinese Academy of Sciences, 19 A Yuquan Rd., Shijingshan District, Beijing 100049, P. R. China.</mods:affiliation>
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<treatment id="7E5287DBE3562F7BFF2EF985FF3F069C" ID-DOI="http://doi.org/10.5281/zenodo.4640493" ID-GBIF-Taxon="180780154" ID-Zenodo-Dep="4640493" LSID="urn:lsid:plazi:treatment:7E5287DBE3562F7BFF2EF985FF3F069C" httpUri="http://treatment.plazi.org/id/7E5287DBE3562F7BFF2EF985FF3F069C" lastPageNumber="487" pageId="14" pageNumber="487">
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<subSubSection id="BEE16546E3562F7BFF2EF985FD77070A" box="[151,656,1626,1653]" pageId="14" pageNumber="487" type="nomenclature">
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<paragraph id="F64436CDE3562F7BFF2EF985FD77070A" blockId="14.[151,656,1626,1653]" box="[151,656,1626,1653]" pageId="14" pageNumber="487">
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<heading id="AD0C81A1E3562F7BFF2EF985FD77070A" bold="true" box="[151,656,1626,1653]" fontSize="11" level="1" pageId="14" pageNumber="487" reason="1">
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<emphasis id="C48FEADFE3562F7BFF2EF985FD77070A" bold="true" box="[151,656,1626,1653]" pageId="14" pageNumber="487">
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6.
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<taxonomicName id="31FB4D4EE3562F7BFF0CF985FD77070A" ID-CoL="8XZWL" authority="(Sharp, 1874)" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[181,656,1626,1653]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="487" phylum="Arthropoda" rank="species" species="gastralis">
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<emphasis id="C48FEADFE3562F7BFF0CF985FE07070B" bold="true" box="[181,480,1626,1653]" italics="true" pageId="14" pageNumber="487">Eccoptolonthus gastralis</emphasis>
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(
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<bibRefCitation id="926A4B3CE3562F7BFE56F985FD6F070A" author="Sharp, D." box="[495,648,1626,1652]" pageId="14" pageNumber="487" pagination="47 - 48" refId="ref16531" refString="Sharp, D. (1874) The Staphylinidae of Japan. Transactions of the Entomological Society of London, 1, 47 - 48. https: // doi. org / 10.1111 / j. 1365 - 2311.1874. tb 00159. x" type="journal article" year="1874">Sharp, 1874</bibRefCitation>
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)
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="BEE16546E3562F7BFF2EF940FA60062E" pageId="14" pageNumber="487" type="description">
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<paragraph id="F64436CDE3562F7BFF2EF940FA60062E" blockId="14.[151,1437,1695,1873]" pageId="14" pageNumber="487">
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<bibRefCitation id="926A4B3CE3562F7BFF2EF940FEDC07C9" author="Sharp, D." box="[151,315,1695,1719]" pageId="14" pageNumber="487" pagination="47 - 48" refId="ref16531" refString="Sharp, D. (1874) The Staphylinidae of Japan. Transactions of the Entomological Society of London, 1, 47 - 48. https: // doi. org / 10.1111 / j. 1365 - 2311.1874. tb 00159. x" type="journal article" year="1874">Sharp, 1874: 48</bibRefCitation>
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(
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<taxonomicName id="31FB4D4EE3562F7BFEF2F940FE5E07C9" authorityName="Stephens" authorityYear="1829" box="[331,441,1695,1719]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="487" phylum="Arthropoda" rank="genus">
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<emphasis id="C48FEADFE3562F7BFEF2F940FE5E07C9" box="[331,441,1695,1719]" italics="true" pageId="14" pageNumber="487">Philonthus</emphasis>
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</taxonomicName>
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,
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<typeStatus id="2940886FE3562F7BFE7EF97FFE1407C6" box="[455,499,1696,1720]" pageId="14" pageNumber="487">type</typeStatus>
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locality:
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<collectingCountry id="8EEC765DE3562F7BFDECF97FFD7607C9" box="[597,657,1696,1719]" name="Japan" pageId="14" pageNumber="487">Japan</collectingCountry>
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,
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<collectingRegion id="343FF82FE3562F7BFD24F940FCE707C9" box="[669,768,1695,1719]" country="Japan" name="Nagasaki" pageId="14" pageNumber="487">Nagasaki</collectingRegion>
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);
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<bibRefCitation id="926A4B3CE3562F7BFCABF940FC3F07C9" author="Thaxter, R." box="[786,984,1695,1719]" pageId="14" pageNumber="487" pagination="217 - 469" refId="ref16626" refString="Thaxter, R. (1908) Contribution toward a monograph of the Laboulbeniaceae. Part II. Memoirs of the American Academy of Arts and Sciences, 13, 217 - 469. https: // doi. org / 10.2307 / 25058090" type="journal article" year="1908">Thaxter, 1908: 424</bibRefCitation>
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;
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<bibRefCitation id="926A4B3CE3562F7BFC5DF940FAA707C9" author="Bernhauer, M. & Schubert, K." box="[996,1344,1695,1719]" pageId="14" pageNumber="487" pagination="289 - 408" refId="ref14553" refString="Bernhauer, M. & Schubert, K. (1914) Staphylinidae IV. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus. Vol. 5. Pars 57. Junk, Berlin, pp. 289 - 408." type="book chapter" year="1914">Bernhauer & Schubert, 1914: 340</bibRefCitation>
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(
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<taxonomicName id="31FB4D4EE3562F7BFAE9F940FF1607A8" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="487" phylum="Arthropoda" rank="genus">
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<emphasis id="C48FEADFE3562F7BFAE9F940FF1607A8" italics="true" pageId="14" pageNumber="487">Philonthus</emphasis>
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</taxonomicName>
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, catalog);
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<bibRefCitation id="926A4B3CE3562F7BFEDBF961FDBB07A8" author="Scheerpeltz, O." box="[354,604,1726,1750]" pageId="14" pageNumber="487" pagination="989 - 1500" refId="ref16140" refString="Scheerpeltz, O. (1933) Staphylinidae VII. In: Schenkling, S. (Eds.), Coleopterorum Catalogus. 6 (129). Junk, Berlin, pp. 989 - 1500." type="book chapter" year="1933">Scheerpeltz, 1933: 1344</bibRefCitation>
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(
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<taxonomicName id="31FB4D4EE3562F7BFDD4F961FD3C07A8" authorityName="Stephens" authorityYear="1829" box="[621,731,1726,1750]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="487" phylum="Arthropoda" rank="genus">
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<emphasis id="C48FEADFE3562F7BFDD4F961FD3C07A8" box="[621,731,1726,1750]" italics="true" pageId="14" pageNumber="487">Philonthus</emphasis>
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</taxonomicName>
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, catalog);
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<bibRefCitation id="926A4B3CE3562F7BFCF2F961FBF307A8" author="Lepesme, P." box="[843,1044,1726,1750]" pageId="14" pageNumber="487" pagination="57 - 80" refId="ref15733" refString="Lepesme, P. (1942) Revision des Rhachomyces palearctiques (Laboulbeniaceae). Bulletin de la Societe mycologique de France, 58, 57 - 80." type="journal article" year="1942">Lepesme, 1942: 79</bibRefCitation>
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;
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<bibRefCitation id="926A4B3CE3562F7BFB98F961FB3007AB" author="Ishikawa, M." box="[1057,1239,1726,1750]" pageId="14" pageNumber="487" pagination="2 - 7" refId="ref15593" refString="Ishikawa, M. (1952) An essay on Laboulbeniales. Shin Konchu, 5 (10), 2 - 7. [in Japanese]" type="journal article" year="1952">
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<collectingRegion id="343FF82FE3562F7BFB98F961FB9B07A8" box="[1057,1148,1726,1750]" country="Japan" name="Ishikawa" pageId="14" pageNumber="487">Ishikawa</collectingRegion>
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1952: 7
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</bibRefCitation>
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;
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<bibRefCitation id="926A4B3CE3562F7BFB5DF961FA7B07A8" author="Nakane, T." box="[1252,1436,1726,1750]" pageId="14" pageNumber="487" refId="ref15937" refString="Nakane, T. (1963) Staphylinidae. In: Nakane, T., Ohbayashi, K., Nomura, S., & Kurosawa, Y. (Eds.), Iconographia Insectorum Japonicorum, Colore naturali edita. Vol. 2. Hokuryukan, Tokyo, pp. 81 - 100. [in Japanese]" type="book" year="1963">Nakane, 1963: 91</bibRefCitation>
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(
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<taxonomicName id="31FB4D4EE3562F7BFF69F903FED9078A" authorityName="Stephens" authorityYear="1829" box="[208,318,1756,1780]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="487" phylum="Arthropoda" rank="genus">
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<emphasis id="C48FEADFE3562F7BFF69F903FED9078A" box="[208,318,1756,1780]" italics="true" pageId="14" pageNumber="487">Philonthus</emphasis>
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</taxonomicName>
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, characters, habitus image,
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<collectingCountry id="8EEC765DE3562F7BFDE5F902FD7D078B" box="[604,666,1757,1781]" name="Japan" pageId="14" pageNumber="487">Japan</collectingCountry>
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);
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<bibRefCitation id="926A4B3CE3562F7BFD15F903FC14078B" author="Watanabe, Y. & Shibata, Y." box="[684,1011,1756,1781]" pageId="14" pageNumber="487" pagination="321" refId="ref16866" refString="Watanabe, Y. & Shibata, Y. (1965) The staphylinid-beetles from Rishiri and Rebun Isls., Hokkaido, Japan, with descriptions of three new species. Kontyu, 33 (3), 321." type="journal article" year="1965">Watanabe & Shibata, 1965: 321</bibRefCitation>
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(subgenus
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<taxonomicName id="31FB4D4EE3562F7BFBD0F902FB53078A" authorityName="Stephens" authorityYear="1829" box="[1129,1204,1757,1780]" class="Insecta" family="Staphylinidae" genus="Bisnius" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="487" phylum="Arthropoda" rank="genus">
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<emphasis id="C48FEADFE3562F7BFBD0F902FB53078A" box="[1129,1204,1757,1780]" italics="true" pageId="14" pageNumber="487">Bisnius</emphasis>
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</taxonomicName>
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of
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<taxonomicName id="31FB4D4EE3562F7BFB6EF903FAA2078A" authorityName="Stephens" authorityYear="1829" box="[1239,1349,1756,1780]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="487" phylum="Arthropoda" rank="genus">
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<emphasis id="C48FEADFE3562F7BFB6EF903FAA2078A" box="[1239,1349,1756,1780]" italics="true" pageId="14" pageNumber="487">Philonthus</emphasis>
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</taxonomicName>
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,
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<collectingCountry id="8EEC765DE3562F7BFAEAF902FA76078B" box="[1363,1425,1757,1781]" name="Japan" pageId="14" pageNumber="487">Japan</collectingCountry>
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);
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<bibRefCitation id="926A4B3CE3562F7BFF7EF924FE78066D" author="Scheloske, H. W." box="[199,415,1787,1811]" pageId="14" pageNumber="487" pagination="1 - 176" refId="ref16338" refString="Scheloske, H. W. (1969) Beitrage zur Bilogie, Okologie und Systematik der Laboulbeniales (Ascomycetes) unter besonderer Berucksichtigung des Parasit-Wirt-Verhaltnisser. Parasitologische Schriftenreihe, 19, 1 - 176." type="journal article" year="1969">Scheloske, 1969: 142</bibRefCitation>
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(
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<collectingCountry id="8EEC765DE3562F7BFE12F924FE0D066D" box="[427,490,1787,1811]" name="Japan" pageId="14" pageNumber="487">Japan</collectingCountry>
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);
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<bibRefCitation id="926A4B3CE3562F7BFE42F924FD7E066D" author="Frank, J. H." box="[507,665,1787,1811]" pageId="14" pageNumber="487" pagination="41" refId="ref15147" refString="Frank, J. H. (1982) The parasites of the Staphylinidae (Coleoptera). A contribution towards an encyclopedia of the Staphylinidae. University of Florida Agricultural Experiment Station Bulletin, 824, 41." type="journal article" year="1982">Frank, 1982: 41</bibRefCitation>
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(
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<taxonomicName id="31FB4D4EE3562F7BFD1EF924FCF2066D" authorityName="Stephens" authorityYear="1829" box="[679,789,1787,1811]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="487" phylum="Arthropoda" rank="genus">
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<emphasis id="C48FEADFE3562F7BFD1EF924FCF2066D" box="[679,789,1787,1811]" italics="true" pageId="14" pageNumber="487">Philonthus</emphasis>
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</taxonomicName>
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, list of parasites);
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<bibRefCitation id="926A4B3CE3562F7BFC75F924FB47066D" author="Watanabe, Y." box="[972,1184,1787,1811]" pageId="14" pageNumber="487" pagination="256" refId="ref16835" refString="Watanabe, Y. (1995) Additional records of staphylinid species from Okushiripp. to Island, off Southwestern Hokkaido. Elytra, 23 (2), 256." type="journal article" year="1995">Watanabe, 1995: 256</bibRefCitation>
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(
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<taxonomicName id="31FB4D4EE3562F7BFB17F924FAFB066D" authorityName="Stephens" authorityYear="1829" box="[1198,1308,1787,1811]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="487" phylum="Arthropoda" rank="genus">
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<emphasis id="C48FEADFE3562F7BFB17F924FAFB066D" box="[1198,1308,1787,1811]" italics="true" pageId="14" pageNumber="487">Philonthus</emphasis>
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</taxonomicName>
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, Okushiri-tô Island);
|
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<bibRefCitation id="926A4B3CE3562F7BFEA3F8C5FE12064C" author="Herman, L. H." box="[282,501,1818,1842]" pageId="14" pageNumber="487" pagination="2819 - 2949" refId="ref15354" refString="Herman, L. H. (2001 a) Catalog of the Staphylinidae (Insecta: Coleoptera): 1758 to the end of the second millennium. V. Staphylinine group (Part 2). Staphylininae: Diochini, Maorothiini, Othiini, Platyprosopini, Staphylinini (Amblyopinina, Anisolinina, Hyptiomina, Philonthina). Bulletin of the American Museum of Natural History, 265, 2819 - 2949. https: // doi. org / 10.1206 / 0003 - 0090.265.1.5" type="journal article" year="2001">Herman, 2001a: 2831</bibRefCitation>
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(
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<taxonomicName id="31FB4D4EE3562F7BFDBAF8C5FD96064C" authorityName="Stephens" authorityYear="1829" box="[515,625,1818,1842]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="487" phylum="Arthropoda" rank="genus">
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<emphasis id="C48FEADFE3562F7BFDBAF8C5FD96064C" box="[515,625,1818,1842]" italics="true" pageId="14" pageNumber="487">Philonthus</emphasis>
|
||
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</taxonomicName>
|
||
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|
, catalog);
|
||
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|
<bibRefCitation id="926A4B3CE3562F7BFD62F8C5FC43064C" author="Smetana, A." box="[731,932,1818,1842]" pageId="14" pageNumber="487" refId="ref16576" refString="Smetana, A. (2004) Staphylinidae, subtribe Philonthina. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea-Histeroidea-Staphylinoidea. Apollo Books, Stenstrup, pp. 644 - 649." type="book" year="2004">Smetana, 2004: 645</bibRefCitation>
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(
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<taxonomicName id="31FB4D4EE3562F7BFC0BF8C5FBC7064C" authorityName="Stephens" authorityYear="1829" box="[946,1056,1818,1842]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="487" phylum="Arthropoda" rank="genus">
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<emphasis id="C48FEADFE3562F7BFC0BF8C5FBC7064C" box="[946,1056,1818,1842]" italics="true" pageId="14" pageNumber="487">Philonthus</emphasis>
|
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</taxonomicName>
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||
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|
, catalog of Palaearctic region);
|
||
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<bibRefCitation id="926A4B3CE3562F7BFAD9F8C5FE61062F" author="Schillhammer, H. & Staphylinidae, New Synonyms & In & Lobl, I. & Lobl, D." pageId="14" pageNumber="487" pagination="1" refId="ref16408" refString="Schillhammer, H. (2015) Staphylinidae, New Synonyms. In: Lobl, I. & Lobl, D. (2015) Catalogue of Palaearctic Coleoptera. Vol. 2 / 1. Hydrophiloidea-Staphylinoidea Revised and Updated Edition. Vol. 1. Brill, Leiden and Boston, pp. 23 - 24." type="book chapter" year="2015">Schillhammer, 2015: 24</bibRefCitation>
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(transferred to
|
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<taxonomicName id="31FB4D4EE3562F7BFD9AF8E7FD26062E" authorityName="Bernhauer" authorityYear="1912" box="[547,705,1848,1872]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="487" phylum="Arthropoda" rank="genus">
|
||
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|
<emphasis id="C48FEADFE3562F7BFD9AF8E7FD26062E" box="[547,705,1848,1872]" italics="true" pageId="14" pageNumber="487">Eccoptolonthus</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
, catalog);
|
||
|
|
<bibRefCitation id="926A4B3CE3562F7BFC95F8E7FB92062F" author="Schulke, M. & Smetana, A." box="[812,1141,1848,1873]" pageId="14" pageNumber="487" pagination="1" refId="ref16467" refString="Schulke, M. & Smetana, A. (2015) Staphylinidae, subtribe Philonthina. In: Lobl, I. & Lobl, D. (2015) Catalogue of Palaearctic Coleoptera. Vol. 2 / 1. Hydrophiloidea-Staphylinoidea Revised and Updated Edition. Vol. 1. Brill, Leiden and Boston, pp. 1017 - 1054." type="book chapter" year="2015">Schülke & Smetana, 2015: 1021</bibRefCitation>
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(
|
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|
<taxonomicName id="31FB4D4EE3562F7BFB3DF8E7FAC5062E" authorityName="Bernhauer" authorityYear="1912" box="[1156,1314,1848,1872]" class="Insecta" family="Staphylinidae" genus="Eccoptolonthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="487" phylum="Arthropoda" rank="genus">
|
||
|
|
<emphasis id="C48FEADFE3562F7BFB3DF8E7FAC5062E" box="[1156,1314,1848,1872]" italics="true" pageId="14" pageNumber="487">Eccoptolonthus</emphasis>
|
||
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|
</taxonomicName>
|
||
|
|
, catalog).
|
||
|
|
</paragraph>
|
||
|
|
</subSubSection>
|
||
|
|
<subSubSection id="BEE16546E3562F7BFF2EF8A1FE3106E7" box="[151,470,1918,1945]" pageId="14" pageNumber="487" type="materials_examined">
|
||
|
|
<paragraph id="F64436CDE3562F7BFF2EF8A1FE3106E7" blockId="14.[151,1437,1918,2018]" box="[151,470,1918,1945]" pageId="14" pageNumber="487">
|
||
|
|
<emphasis id="C48FEADFE3562F7BFF2EF8A1FE6B06E6" bold="true" box="[151,396,1918,1944]" pageId="14" pageNumber="487">Materials examined.</emphasis>
|
||
|
|
None.
|
||
|
|
</paragraph>
|
||
|
|
</subSubSection>
|
||
|
|
<subSubSection id="BEE16546E3562F7BFF7EF87DFF3F069C" pageId="14" pageNumber="487" type="distribution">
|
||
|
|
<paragraph id="F64436CDE3562F7BFF7EF87DFF3F069C" blockId="14.[151,1437,1918,2018]" pageId="14" pageNumber="487">
|
||
|
|
<emphasis id="C48FEADFE3562F7BFF7EF87DFE8606C2" bold="true" box="[199,353,1954,1980]" pageId="14" pageNumber="487">Distribution.</emphasis>
|
||
|
|
<collectingCountry id="8EEC765DE3562F7BFED0F87CFE4B06C3" box="[361,428,1955,1981]" name="China" pageId="14" pageNumber="487">China</collectingCountry>
|
||
|
|
(exact locality not recorded),
|
||
|
|
<collectingCountry id="8EEC765DE3562F7BFCBAF87CFCA406C3" box="[771,835,1955,1981]" name="Japan" pageId="14" pageNumber="487">Japan</collectingCountry>
|
||
|
|
(
|
||
|
|
<collectingRegion id="343FF82FE3562F7BFCEAF87CFC2106C3" box="[851,966,1955,1981]" country="Japan" name="Hokkaido" pageId="14" pageNumber="487">Hokkaido</collectingRegion>
|
||
|
|
, Honshu, Kyushu, Shikoku, Tsushima Island).
|
||
|
|
</paragraph>
|
||
|
|
</subSubSection>
|
||
|
|
</treatment>
|
||
|
|
</document>
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