treatments-xml/data/7E/50/4E/7E504E7BA54A5C5FA870A13BF970B161.xml

<document ID-CLB-Dataset="7250" ID-DOI="http://dx.doi.org/10.3897/mycokeys.79.63363" ID-GBIF-Dataset="cec973b7-30eb-4570-afc1-159895cffa7c" ID-PMC="PMC8076164" ID-Pensoft-Pub="1314-4049-79-17" ID-Pensoft-UUID="CEDC3742C169540583072E7995018533" ID-PubMed="33958950" ModsDocID="1314-4049-79-17" checkinTime="1618878569608" checkinUser="pensoft" docAuthor="Eberhardt, Ursula, Beker, Henry J., Borgen, Torbjorn, Knudsen, Henning, Schuetz, Nicole &amp; Elborne, Steen A." docDate="2021" docId="7E504E7BA54A5C5FA870A13BF970B161" docLanguage="en" docName="MycoKeys 79: 17-118" docOrigin="MycoKeys 79" docPubDate="2021-04-19" docSource="http://dx.doi.org/10.3897/mycokeys.79.63363" docTitle="Hebeloma arcticum Beker &amp; U. Eberh. sp. nov." docType="treatment" docVersion="6" id="CEDC3742C169540583072E7995018533" lastPageNumber="17" masterDocId="CEDC3742C169540583072E7995018533" masterDocTitle="A survey of Hebeloma (Hymenogastraceae) in Greenland" masterLastPageNumber="118" masterPageNumber="17" pageNumber="17" updateTime="1720462698628" updateUser="ExternalLinkService">
<mods:mods id="7ED093D1E35E86B77B73EF89032F1D87" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C4228A47FFCBB69DE88C2BFFA70AD976">
<mods:title id="FE2BDBD86702E1F89581549CFAE0956D">A survey of Hebeloma (Hymenogastraceae) in Greenland</mods:title>
</mods:titleInfo>
<mods:name id="C1381E95DBAB0725BAAA233D4F1644FE" type="personal">
<mods:role id="93BB2C0A66E6B1AFA71858D316CB5271">
<mods:roleTerm id="8F3C53EEF01E61622742C6A2C1EC4C27">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D7B83811B361A9356A9E3F50C66C43D0">Eberhardt, Ursula</mods:namePart>
<mods:nameIdentifier id="1DE977D79327F1AA4849694B7A06A354" type="ORCID">https://orcid.org/0000-0003-1221-7074</mods:nameIdentifier>
<mods:affiliation id="D1EF09CB35B1DC9107DC682745763157">Staatliches Museum fuer Naturkunde Stuttgart, Rosenstein 1, D- 70191 Stuttgart, Germany</mods:affiliation>
<mods:nameIdentifier id="054FA22FF972624A80F196FB49177DAF" type="email">ursula.eberhardt@smns-bw.de</mods:nameIdentifier>
</mods:name>
<mods:name id="3C57CD830F21E2E8791519D0C243F43C" type="personal">
<mods:role id="98357669735F8FD3A520E0BA5FF462F4">
<mods:roleTerm id="D47C6670152AC4DA9827D3243C74D619">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8BC8F192317221C337873E6D612C4BDF">Beker, Henry J.</mods:namePart>
<mods:nameIdentifier id="18BB6481CF8577C500C74ED990CAE70A" type="ORCID">https://orcid.org/0000-0001-9620-1701</mods:nameIdentifier>
<mods:affiliation id="31A1FE145542E8988B404E75935EF63B">Rue Pere de Deken 19, B- 1040 Bruxelles, Belgium &amp; Royal Holloway College, University of London, Egham, UK &amp; Plantentuin Meise, Nieuwelaan 38, B- 1860 Meise, Belgium</mods:affiliation>
</mods:name>
<mods:name id="0E51345D5D2DE6E234D0A181672D5BF5" type="personal">
<mods:role id="8127C64453DA2245D609FC57883BA19E">
<mods:roleTerm id="ED41A2B4E559CB4FAFA21B14F7EE45F2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6D36FEADEE85060B800F575CCD3CA5BE">Borgen, Torbjorn</mods:namePart>
<mods:nameIdentifier id="CF0D0BCF796BD3FBD599D4F9DD3833D1" type="ORCID">https://orcid.org/0000-0003-1174-9466</mods:nameIdentifier>
<mods:affiliation id="4325A129472138B63E4EA2447B119371">Sensommervej 142, 8600 Silkeborg, Denmark</mods:affiliation>
</mods:name>
<mods:name id="BD45D0668CD1DE74EC9A158E65B58ED2" type="personal">
<mods:role id="101DD1ADD4CD0A3EC482E4F1B8C0D150">
<mods:roleTerm id="BCE2E8B467DA41998AA9F2421D68C373">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="49C4E5EF3E249DCC1BB3891C877CE4FA">Knudsen, Henning</mods:namePart>
<mods:affiliation id="B380AED93A7E6B3227A5D92481C0876D">Hauchsvej 15, 1825 Frederiksberg, Denmark</mods:affiliation>
</mods:name>
<mods:name id="373BC8BC5DD85D23EA51CB0D6C2ADA02" type="personal">
<mods:role id="43663976B41C516DBF03EAFF15D3CDCC">
<mods:roleTerm id="D770321FF6CF586D3ADE0757CB9D777A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FB61DF95452F7939AEB24EF74BDC3A30">Schuetz, Nicole</mods:namePart>
<mods:affiliation id="41C66D48DCAB736D8E392E940E8B3527">Staatliches Museum fuer Naturkunde Stuttgart, Rosenstein 1, D- 70191 Stuttgart, Germany</mods:affiliation>
</mods:name>
<mods:name id="7776BED16FEB928D4002E8ADF8DD2432" type="personal">
<mods:role id="BE12A53195C881124671E82EE22A2154">
<mods:roleTerm id="4F665CBFCCE0EB8C87A719E047877BEE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="10E6F115BBB6AE74B7322D222A912851">Elborne, Steen A.</mods:namePart>
<mods:affiliation id="F008BCA3A8803B9389D3789A5F4220A9">Frederik VII's Vej 29, 3450 Allerod, Denmark</mods:affiliation>
</mods:name>
<mods:typeOfResource id="B933999B9F17DE2A68C663CF7BBD6E8C">text</mods:typeOfResource>
<mods:relatedItem id="BC7582645EA48725CF58A2DE347C7AA8" type="host">
<mods:titleInfo id="1C474FD2F403D5155AA845C22E54AE59">
<mods:title id="140AFBC2D9001DC1D8EE98D6371F6921">MycoKeys</mods:title>
</mods:titleInfo>
<mods:part id="DD0E4EE68BD2597B52AB188D651BCA50">
<mods:date id="1A3B28D3D58CF2951D81468CD440D54F">2021</mods:date>
<mods:detail id="DBBAA1A4615FA59785B7029C981C055E" type="pubDate">
<mods:number id="5D3C1CC7D9E2D11F80942623FFFF7BB7">2021-04-19</mods:number>
</mods:detail>
<mods:detail id="9AAE04BB7CC4CE83271885137A819170" type="volume">
<mods:number id="B8804FE8F8922EA66DC123913CA4BC80">79</mods:number>
</mods:detail>
<mods:extent id="05CB1EC17381008AAB6363AB502A65C6" unit="page">
<mods:start id="1006BBE6CD6099A106422AE7D3E283FC">17</mods:start>
<mods:end id="D90706FE8A1DB1DDCBED839607EC8DE9">118</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="C183DB63558607FE0D950A1237589C4D">
<mods:url id="235D53FBBC382CAB97D68A6AAF8A6244">http://dx.doi.org/10.3897/mycokeys.79.63363</mods:url>
</mods:location>
<mods:classification id="E857E5C088DA854982318F24BD46F932">journal article</mods:classification>
<mods:identifier id="A30F2AD53015119F1FAB460716635B63" type="DOI">http://dx.doi.org/10.3897/mycokeys.79.63363</mods:identifier>
<mods:identifier id="D158E036D8F401C00F500C242E6FCEF9" type="Pensoft-Pub">1314-4049-79-17</mods:identifier>
<mods:identifier id="F8410854DA92B3C38896268AB3D270D6" type="Pensoft-UUID">CEDC3742C169540583072E7995018533</mods:identifier>
</mods:mods>
<treatment id="7E504E7BA54A5C5FA870A13BF970B161" ID-GBIF-Taxon="181518925" LSID="urn:lsid:plazi:treatment:7E504E7BA54A5C5FA870A13BF970B161" httpUri="http://treatment.plazi.org/id/7E504E7BA54A5C5FA870A13BF970B161" lastPageNumber="17" pageId="0" pageNumber="17">
<subSubSection id="8AFD3AC76C95A3233A546BE08D08E6FF" pageId="0" pageNumber="17" type="nomenclature">
<paragraph id="57AA471419C2CD3F2CD575608166329E" pageId="0" pageNumber="17">
<taxonomicName id="6046C0304E5BB886BFB4135FD69958B8" ID-CoL="9Z8SK" LSID="7E504E7B-A54A-5C5F-A870-A13BF970B161" authority="Beker &amp; U. Eberh. sp. nov." authorityName="Beker &amp; U. Eberh. sp. nov." class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" higherTaxonomySource="treatment-meta" kingdom="Fungi" lsidName="Hebeloma arcticum" order="Agaricales" pageId="0" pageNumber="17" phylum="Basidiomycota" rank="species" species="arcticum">Hebeloma arcticum Beker &amp; U. Eberh. sp. nov.</taxonomicName>
<figureCitation id="11963DD7F5F3280FA639E0AE9B1C8FB3" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Hebeloma arcticum A HK 16.119 (holotype), photograph H. Knudsen B distribution of cited collections." figureDoi="10.3897/mycokeys.79.63363.figure22" httpUri="https://binary.pensoft.net/fig/533413" pageId="0" pageNumber="17">Figs 22</figureCitation>
<figureCitation id="27B45F81AF857D8DFF65F4537DC18076" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Hebeloma arcticum HK 16.119 (holotype) A spores x 1600 and B spore ornamentation x 1600 in Melzer's reagent C spores and D spore ornamentation x 1600 in KOH E basidia cheilocystidia and F cheilocystidia x 1000 in Melzer's reagent G caulocystidia x 500 in KOH H cutis x 125 in KOH. Scale bars: 5 µm (A-F); 10 µm (G); 50 µm (H); microphotographs H. K. J. Beker." figureDoi="10.3897/mycokeys.79.63363.figure23" httpUri="https://binary.pensoft.net/fig/533414" pageId="0" pageNumber="17">, 23</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="4F70EB8C0CB3958DCB2952D31AE768D3" pageId="0" pageNumber="17" type="materials_examined">
<paragraph id="6F88735BCEC685A0C0ABAA2DEA6F5FF0" pageId="0" pageNumber="17">
<typeStatus id="2C103E21CFA1BDF01A7765427F4ADE43">Type</typeStatus>
.
</paragraph>
<paragraph id="997C17EA9ADC3892FA0851913CA09E6A" pageId="0" pageNumber="17">
<materialsCitation id="3968DC5710EAD4A27DBC29EA9E1F5487" ID-GBIF-Occurrence="3080424301" collectingDate="2016-08-14" collectorName="H. Knudsen" country="Greenland" latitude="66.94432" location="Sisimiut" longLatPrecision="1" longitude="-53.670444" specimenCount="1" typeStatus="holotype">
<collectingCountry id="6C45C51C35B60DF9B7AEDDF175B8E947" name="Greenland">Greenland</collectingCountry>
. 
<location id="AF5F65EA8266F21A3DBFD02D00F0C4C1" LSID="urn:lsid:plazi:treatment:7E504E7BA54A5C5FA870A13BF970B161:AF5F65EA8266F21A3DBFD02D00F0C4C1" country="Greenland" latitude="66.94432" longLatPrecision="1" longitude="-53.670444" name="Sisimiut">Sisimiut</location>
: at the bridge on the road to the airport (approx. 
<geoCoordinate id="59B9DF3ECB6EC57D3F939D42A3AD27C3" degrees="66.944317" direction="north" orientation="latitude" precision="1" value="66.94432">66.944317N</geoCoordinate>
, 
<geoCoordinate id="AA47A28EE4B9F1C82F553163C4FDB89B" degrees="53.670444" direction="west" orientation="longitude" precision="1" value="-53.670444">53.670444W</geoCoordinate>
, alt. approx. 0 m a.s.l.), 
<collectingDate id="288504BFBB5B9558A4E3E3770064A17C" value="2016-08-14">14 Aug. 2016</collectingDate>
, 
<collectorName id="BA0EB993BB4EDCDCC8E848F3256E78A0">H. Knudsen</collectorName>
(HK16.119)
</materialsCitation>
<materialsCitation id="EA60F79B35A5D2413F4423758368C7FF" ID-GBIF-Occurrence="3080424302" accessionNumber="HJB17673, MW445587, MW452584, MW452593" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="HK" specimenCount="1" typeStatus="Holotype">
(
<emphasis id="ACA5E3E3B611772563400225B1A8F843" bold="true" italics="true" pageId="0" pageNumber="17">
<typeStatus id="5C2B4B6FA2323A0D6E03B36D34CD69AB">Holotype</typeStatus>
</emphasis>
: C-F-104149; Database Record: 
<accessionNumber id="ECB37EF55FF842C077C941BDE7FA3F2B" httpUri="http://www.ncbi.nlm.nih.gov/protein/HJB17673">HJB17673</accessionNumber>
; Genbank: ITS = 
<accessionNumber id="59E2CF00BBFA67394D66682F539586D9" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/MW445587">MW445587</accessionNumber>
, 
<emphasis id="2B0424DDA3CC996B418DB6F780707663" italics="true" pageId="0" pageNumber="17">Tef1</emphasis>
a = 
<accessionNumber id="8F5ED7D9A7ED73CDDFFAFDE7FBB4F238" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/MW452584">MW452584</accessionNumber>
and 
<emphasis id="82909546FAC3BDC84F306E4F5572D788" italics="true" pageId="0" pageNumber="17">RPB2</emphasis>
= 
<accessionNumber id="DAF5F53BB6B996CE8C95A239EAC249CA" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/MW452593">MW452593</accessionNumber>
)
</materialsCitation>
.
</paragraph>
<caption id="3A0234477D8B0A4C919417581A4324AD" doi="10.3897/mycokeys.79.63363.figure22" httpUri="https://binary.pensoft.net/fig/533413" pageId="0" pageNumber="17" start="Figure 22" startId="F22">
<paragraph id="9AF593725606A884BA65D3DD155E822B" pageId="0" pageNumber="17">
<emphasis id="97D7750BAC2BF653E6448671983AE3AA" bold="true" pageId="0" pageNumber="17">Figure 22.</emphasis>
<taxonomicName id="E5631D4B114E7343C1D97F7BAF14F020" class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" kingdom="Fungi" lsidName="Hebeloma arcticum" order="Ostropales" pageId="0" pageNumber="17" phylum="Basidiomycota" rank="species" species="arcticum">
<emphasis id="7A7E48B6F44C2A708C8D3AA27C5C1D1D" italics="true" pageId="0" pageNumber="17">Hebeloma arcticum</emphasis>
</taxonomicName>
<emphasis id="D23B31851596F3A2D8BB3E5E35A16D35" bold="true" pageId="0" pageNumber="17">A</emphasis>
<collectingCountry id="BB3578D6B33E56EAD5C114F396E6C369" name="Hong Kong">HK</collectingCountry>
16.119 (
<typeStatus id="8B1008AB088BB6E6CDFB3ECDF921633B">holotype</typeStatus>
), photograph H. Knudsen 
<emphasis id="B9E86ED3B938E10B3D469FF7C0BEB08E" bold="true" pageId="0" pageNumber="17">B</emphasis>
distribution of cited collections.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C3493F8CAED9A69CD5D0A226E5906B29" pageId="0" pageNumber="17" type="diagnosis">
<paragraph id="397064F6E9F9B68056A390AD4AC3789A" pageId="0" pageNumber="17">Diagnosis.</paragraph>
<paragraph id="9FD0BB6317C9858DB72E67647181EE81" pageId="0" pageNumber="17">Favoring arctic type habitats with many cheilocystidia clavate-stipitate and rather strongly dextrinoid but indistinctly ornamented spores.</paragraph>
</subSubSection>
<subSubSection id="E3F7F51DDEF1A17FE35E24BE1BABD141" pageId="0" pageNumber="17" type="etymology">
<paragraph id="35CEE16DDDF7CC72BE01C1A163739025" pageId="0" pageNumber="17">Etymology.</paragraph>
<paragraph id="4BFABF7018D677C5941A6DC46C00DBC0" pageId="0" pageNumber="17">
<emphasis id="1087B57C468F53D27B449D0D17173884" italics="true" pageId="0" pageNumber="17">arcticus</emphasis>
(adj. Latin), meaning the arctic, to emphasize the habitat within which this mushroom has been discovered.
</paragraph>
</subSubSection>
<subSubSection id="6FFA0743F7D62D237780C7A6F7FB26F1" pageId="0" pageNumber="17" type="description">
<paragraph id="4B6D0E00E01FFB0FAF217BCECFF1D9CC" pageId="0" pageNumber="17">Macroscopic description.</paragraph>
<paragraph id="5C594750A2A2D6EA65A4858E3D153A3F" pageId="0" pageNumber="17">
Cap (1.7-) 2.4-2.9 (-3.8) cm, plano-convex, with or without suggestion of obtuse umbo, with decurved margin then applanate, only weakly viscid even after heavy rainfall, dull, almost glabrous, marginal zone whitish pale due to a fine adpressed tomentum, often eroded, usually bicolored, center ochre, paler towards the margin becoming very pale brownish, hardly hygrophanous but a few hygrophanous dots. Lamellae emarginate with tooth, ventricose, number of lamellae {L} 36-44, up to 4 mm wide, initially pale, darker (clay buff) at maturity with whitish fimbriate edge and usually some watery droplets or droplets visible with a x10 lens. Stem (2.8-) 4.1-4.4 (-6.0) 
<normalizedToken id="1BC08247D17A4DF6128F08DAE566F61E" originalValue="×">x</normalizedToken>
0.5-0.7 (-0.9) cm, to 0.6-0.9 cm at base, whitish fibrillose to pruinose, cylindrical, narrowly fistulose, downwards pale yellowish to pale brownish when rubbed. Context firm, thick particularly in center, whitish pale, watery when moist, in age slightly brownish downwards in stipe. Smell weakly herbaceous-raphanoid. Taste mild. Spore print at most clay buff (Munsell 10YR6/4; in TB 90.071).
</paragraph>
<caption id="37D785C212CBA54C5C5907F16D0D01CB" doi="10.3897/mycokeys.79.63363.figure23" httpUri="https://binary.pensoft.net/fig/533414" pageId="0" pageNumber="17" start="Figure 23" startId="F23">
<paragraph id="2F2D088C652E8E2EF0A87DE0E88968DA" pageId="0" pageNumber="17">
<emphasis id="69DDE4A6A895EC0FFEF76085639729D4" bold="true" pageId="0" pageNumber="17">Figure 23.</emphasis>
<taxonomicName id="F35481FC84947222DCB78250683AA583" class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" kingdom="Fungi" lsidName="Hebeloma arcticum" order="Ostropales" pageId="0" pageNumber="17" phylum="Basidiomycota" rank="species" species="arcticum">
<emphasis id="ABB205151F5137B2CD61A8AD7F7ED0C4" italics="true" pageId="0" pageNumber="17">Hebeloma arcticum</emphasis>
</taxonomicName>
HK16.119 (holotype) 
<emphasis id="4848D506F83530ECE39096FDCE7EE69C" bold="true" pageId="0" pageNumber="17">A</emphasis>
spores 
<normalizedToken id="7371E68AA0E9E77B667CA8227EC41982" originalValue="×">x</normalizedToken>
1600 and 
<emphasis id="199534004A68B99FD8DB04743F008C2F" bold="true" pageId="0" pageNumber="17">B</emphasis>
spore ornamentation 
<normalizedToken id="425E3367421FF58D718EC871D55D70F6" originalValue="×">x</normalizedToken>
1600 in 
<normalizedToken id="B6CAB2399E5CB601979FC993856A0405" originalValue="Melzer’s">Melzer's</normalizedToken>
reagent 
<emphasis id="902B27BBC0F096FC7E30BD8614DE0732" bold="true" pageId="0" pageNumber="17">C</emphasis>
spores and 
<emphasis id="9BAF0ACAD0786829AB8570E2C15E9DB1" bold="true" pageId="0" pageNumber="17">D</emphasis>
spore ornamentation 
<normalizedToken id="039B03A4038ADBCD076740BEF711856A" originalValue="×">x</normalizedToken>
1600 in KOH 
<emphasis id="907EFB0D37D1B4DE47B6B7165A18D2C4" bold="true" pageId="0" pageNumber="17">E</emphasis>
basidia cheilocystidia and 
<emphasis id="44D1EAA82924A6AE05F4ACC0A34E0E87" bold="true" pageId="0" pageNumber="17">F</emphasis>
cheilocystidia 
<normalizedToken id="E27C7B662BA5CC7FB6B7AE58A87535E4" originalValue="×">x</normalizedToken>
1000 in 
<normalizedToken id="31183D4B2D3C617B8C9423A648B72B99" originalValue="Melzer’s">Melzer's</normalizedToken>
reagent 
<emphasis id="866A3B40BF5CD46962BE251BDC9E4A3C" bold="true" pageId="0" pageNumber="17">G</emphasis>
caulocystidia 
<normalizedToken id="A16D801521CB0E6DEF2F70E12BD34C24" originalValue="×">x</normalizedToken>
500 in KOH 
<emphasis id="8D64BAF24D17F7810B1427E22076ED2A" bold="true" pageId="0" pageNumber="17">H</emphasis>
cutis 
<normalizedToken id="B3715C85570C519BCE3D495DEB933E82" originalValue="×">x</normalizedToken>
125 in KOH. Scale bars: 5 
<normalizedToken id="F49677C5B08C7DFA2C3908CA9E0D397D" originalValue="µm">µm</normalizedToken>
(
<emphasis id="5EDAAB90AAADB6AECABA12BF02FD4DF2" bold="true" pageId="0" pageNumber="17">A-F</emphasis>
); 10 
<normalizedToken id="CFF1A9EC7718423A1B8369C0E655DA54" originalValue="µm">µm</normalizedToken>
(
<emphasis id="E8763603091B2C1E18B6C3C2416E3B73" bold="true" pageId="0" pageNumber="17">G</emphasis>
); 50 
<normalizedToken id="0FEE7A5533CECB2C1E6D8B1848581EA5" originalValue="µm">µm</normalizedToken>
(
<emphasis id="FC154CE4F7F370FA5B64A79ABF06877B" bold="true" pageId="0" pageNumber="17">H</emphasis>
); microphotographs H.K.J. Beker.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="39158AB73C8D430C07CB6390C10ACCDB" pageId="0" pageNumber="17" type="description">
<paragraph id="5F05094CD124CCCCD20B186C5781919C" pageId="0" pageNumber="17">Microscopic description.</paragraph>
<paragraph id="7BBD8932E297B3293F0DE8DA24AF756B" pageId="0" pageNumber="17">
Spores amygdaloid, ellipsoid or ovoid, on ave. (across eight collections) 10.8-12.0 
<normalizedToken id="90F8ECFC5CF6032FDBAE16BE951533F6" originalValue="×">x</normalizedToken>
6.4-6.9 
<normalizedToken id="8A77A5368AD63B8522C73A0EF8A20E4F" originalValue="µm">µm</normalizedToken>
, ave. Q = 1.65-1.83, (for the holotype, measuring 102 spores, 5% to 95% percentile range 10.4-13.2 
<normalizedToken id="41F8D34EA3337FD14E4564D52FB69EC2" originalValue="×">x</normalizedToken>
5.9-7.2 
<normalizedToken id="F278BBFAEA40EF0B2C0E7195AEA211E1" originalValue="µm">µm</normalizedToken>
with median 11.9 
<normalizedToken id="301CE9B0300BB8A4B5FFEEA40FE9B570" originalValue="×">x</normalizedToken>
6.5 
<normalizedToken id="16D1413561C4C7CD0F9DBD39F4F803DB" originalValue="µm">µm</normalizedToken>
and ave. 11.9 
<normalizedToken id="2095440F9E69B9EF6FA120919F9DDB3B" originalValue="×">x</normalizedToken>
6.5 
<normalizedToken id="9678C4A78EAF78DDD690A4B4EF3F14F4" originalValue="µm">µm</normalizedToken>
and S.D. for length 0.84 
<normalizedToken id="2D833133282EB453B5BCF98218FF8A37" originalValue="µm">µm</normalizedToken>
and for width 0.39 
<normalizedToken id="871963CE343381B8F453FC17663510ED" originalValue="µm">µm</normalizedToken>
, and Q value 5% to 95% percentile range 1.57-2.04, with median 1.83 and ave. 1.83 with S. D. 0.14), yellow brown, guttulate, not papillate, almost smooth (O1), perispore not loosening (P0), distinctly to strongly dextrinoid (D2D3). Basidia (20) 22-37 
<normalizedToken id="FA8A974EEABA9C4D1254ADAE810023DC" originalValue="×">x</normalizedToken>
6-10 
<normalizedToken id="43DAB9B6B26D1B28839CC4835AD8B80F" originalValue="μm">μm</normalizedToken>
; ave. Q 3.0-3.7, (for the holotype 25-37 
<normalizedToken id="7208D6619EB5F5B4BF60D879855438C1" originalValue="×">x</normalizedToken>
7-9, ave. Q 3.6), mostly four-spored. Cheilocystidia mainly clavate-stipitate, sometimes clavate-ventricose, more rarely capitate-stipitate, occasional apical or median wall thickening, sometimes septate (occasionally clamped), on ave. 36-57 
<normalizedToken id="B545BB3295325A96EABDEA6CE992C268" originalValue="×">x</normalizedToken>
6-9 (apex) 
<normalizedToken id="7BF04D791C24B021773B2A7127007C93" originalValue="×">x</normalizedToken>
4-5 (middle) 
<normalizedToken id="A34F3C55B3A510CE88686E83866261B8" originalValue="×">x</normalizedToken>
4-6.5 (base) 
<normalizedToken id="5F45DFB643B9395275F0C0515A65DB37" originalValue="µm">µm</normalizedToken>
, ratios A/M = 1.5-2.3, A/B = 1.2-2.0, B/M = 1.0-1.4, (for the holotype, width near apex, 5% to 95% percentile range 6.5-8.3 
<normalizedToken id="E28E1C51998C294349CBC624DB0D570F" originalValue="µm">µm</normalizedToken>
, with median 7.4 
<normalizedToken id="BF3E162F26702A80AC6F902305D46427" originalValue="µm">µm</normalizedToken>
and ave. 7.4 
<normalizedToken id="6D89549E336B8EB54B1A8060D04FF11F" originalValue="µm">µm</normalizedToken>
with S.D. 0.63 
<normalizedToken id="650FBBA9846935AB66A3997207A10100" originalValue="µm">µm</normalizedToken>
and over all 45 
<normalizedToken id="BB346877F662DA59E3717C968B18884B" originalValue="×">x</normalizedToken>
7.4 (apex) 
<normalizedToken id="474B0F1AFC0837CFE56DC58C2CBF2C43" originalValue="×">x</normalizedToken>
5 (middle) 
<normalizedToken id="E386F1B3FCCF5270BD283FA866C24D5E" originalValue="×">x</normalizedToken>
6.5 (base) 
<normalizedToken id="7FFBEA09825D9520AD82A4E040DD1D4D" originalValue="µm">µm</normalizedToken>
). Pileipellis an ixocutis, up to 90 
<normalizedToken id="633E47E3D0B8FD54DB8134BA35527FFD" originalValue="µm">µm</normalizedToken>
thick (measured from exsiccata), hyphae up to 6 
<normalizedToken id="224B77C1FDC744288C317669A1041E80" originalValue="µm">µm</normalizedToken>
broad, none encrusted; trama elements beneath subcutis isodiametric to ellipsoid or thick sausage-shaped, up to 17.5 
<normalizedToken id="8FC87ED6DCEFCDF0AEBCE68A4EBD61CB" originalValue="µm">µm</normalizedToken>
wide. Caulocystidia similar to cheilocystidia, up to 90 
<normalizedToken id="20E15A531BE62CE193082633F9C202A3" originalValue="µm">µm</normalizedToken>
long.
</paragraph>
</subSubSection>
<subSubSection id="E5198153EBFACFBE09D182EC4164537B" pageId="0" pageNumber="17" type="other collections examined">
<paragraph id="F4EDD8011F31696BF906E9CAD0AD1E6C" pageId="0" pageNumber="17">Other collections examined.</paragraph>
<paragraph id="0BD1182D22385060F3C0A1B6F18AC404" pageId="0" pageNumber="17">
<emphasis id="75EDEE88C8E3FE1F5E7DADF3219895BC" bold="true" pageId="0" pageNumber="17">S-Greenland</emphasis>
: Paamiut, 
<geoCoordinate id="DB659B67957444924C32D2A5FAAD09AE" degrees="62.01" direction="north" orientation="latitude" precision="555" value="62.01">62.01°N</geoCoordinate>
, 
<geoCoordinate id="6F50ED6D62A3ABE0A32C0FB2276612A3" degrees="49.4" direction="west" orientation="longitude" precision="5555" value="-49.4">49.4°W</geoCoordinate>
, 31 Aug 1986, T. Borgen (TB86.277A, C-F-103571), 10 m, with 
<taxonomicName id="D636252D1F6679AD4BBABED6B5823143" class="Dicotyledoneae" family="Salicaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="S. arctophila" order="Salicales" pageId="0" pageNumber="17" phylum="Angiospermae" rank="species" species="arctophila">
<emphasis id="88A3D1D12848E68B564A3EB9D3A72CE3" italics="true" pageId="0" pageNumber="17">S. arctophila</emphasis>
</taxonomicName>
, 
<geoCoordinate id="4F6BDBD0403150D48D6B5F2957172C9E" degrees="62.01" direction="north" orientation="latitude" precision="555" value="62.01">62.01°N</geoCoordinate>
, 
<geoCoordinate id="480FC66578CDC33A149835ED8AF3F3A3" degrees="49.4" direction="west" orientation="longitude" precision="5555" value="-49.4">49.4°W</geoCoordinate>
, 4 Sep 1990, T. Borgen (TB90.083, C-F-103555), 10 m, with 
<taxonomicName id="E5035875E992C1859A0605F6ACBF8F8A" class="Dicotyledoneae" family="Salicaceae" genus="Salix" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Salix glauca" order="Salicales" pageId="0" pageNumber="17" phylum="Angiospermae" rank="species" species="glauca">
<emphasis id="F5572327B974F98109C9C721724DD151" italics="true" pageId="0" pageNumber="17">Salix glauca</emphasis>
</taxonomicName>
and 
<taxonomicName id="2516E08563410A06269878FF9BE3A16C" class="Eudicots" family="Polygonaceae" genus="Bistorta" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Bistorta vivipara" order="Caryophyllales" pageId="0" pageNumber="17" phylum="Magnoliophyta" rank="species" species="vivipara">
<emphasis id="5F847CB786468F9CB8C92E7C62369B11" italics="true" pageId="0" pageNumber="17">Bistorta vivipara</emphasis>
</taxonomicName>
in fenland. Paamiut, 
<geoCoordinate id="94363790F353767D38AFE276D4CACE8E" degrees="62.01" direction="north" orientation="latitude" precision="555" value="62.01">62.01°N</geoCoordinate>
, 
<geoCoordinate id="FCCF68AB7457403AD533125A77586679" degrees="49.4" direction="west" orientation="longitude" precision="5555" value="-49.4">49.4°W</geoCoordinate>
, 1 Sep 1990, T. Borgen (TB90.071, C-F-103483), 10 m, with 
<taxonomicName id="3143BA8AF8B46ED5B9EE036F07DD686C" class="Dicotyledoneae" family="Salicaceae" genus="Salix" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Salix glauca" order="Salicales" pageId="0" pageNumber="17" phylum="Angiospermae" rank="species" species="glauca">
<emphasis id="D9F2FCE0957746E978F8C6BCC1DFCC03" italics="true" pageId="0" pageNumber="17">Salix glauca</emphasis>
</taxonomicName>
and 
<taxonomicName id="E3F909B56AF3D7103E3FE0A30E4482D2" class="Eudicots" family="Polygonaceae" genus="Bistorta" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Bistorta vivipara" order="Caryophyllales" pageId="0" pageNumber="17" phylum="Magnoliophyta" rank="species" species="vivipara">
<emphasis id="BE07984933A036EADF69ACBFBD40B324" italics="true" pageId="0" pageNumber="17">Bistorta vivipara</emphasis>
</taxonomicName>
. Paamiut, near cemetery, 
<geoCoordinate id="05B8FBABFA99A3D0FDDD9483058AE589" degrees="61.9941" direction="north" orientation="latitude" precision="5" value="61.9941">61.9941°N</geoCoordinate>
, 
<geoCoordinate id="209CEA369D87F9AACD32FFD106E80C82" degrees="49.6666" direction="west" orientation="longitude" precision="5" value="-49.6666">49.6666°W</geoCoordinate>
, 21 Aug 2008, T. Borgen (TB08.153, C-F-106751), 15 m, with 
<taxonomicName id="064F4DD2FF1D75BDF133C90E0AEA1F38" class="Dicotyledoneae" family="Salicaceae" genus="Salix" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Salix glauca" order="Salicales" pageId="0" pageNumber="17" phylum="Angiospermae" rank="species" species="glauca">
<emphasis id="CAC153BB4DB54C21E85C7BFC828B0753" italics="true" pageId="0" pageNumber="17">Salix glauca</emphasis>
</taxonomicName>
. 
<emphasis id="3FFD4B0BA6B704FB6077F4D0C9DB531B" bold="true" pageId="0" pageNumber="17">W-Greenland</emphasis>
: Kangerlussuaq, 
<normalizedToken id="0C3F3962BE082EBE97947E6038307B41" originalValue="Ringsødalen">Ringsodalen</normalizedToken>
, 
<normalizedToken id="B5D75A6DBBC9C0C58B5F2FEC19D1EE71" originalValue="Ringsøen">Ringsoen</normalizedToken>
, 
<geoCoordinate id="E443D371650CEC14C992631C7B940D94" degrees="66.9853" direction="north" orientation="latitude" precision="5" value="66.9853">66.9853°N</geoCoordinate>
, 
<geoCoordinate id="D509E32A6A4A56DA37BD08DEB68EA23D" degrees="50.9464" direction="west" orientation="longitude" precision="5" value="-50.9464">50.9464°W</geoCoordinate>
, 9 Aug 2016, T. Borgen (TB16.095, C-F-103584), 180 m, with 
<taxonomicName id="7DA353FD59EB730F365331FBE42FDA53" class="Dicotyledoneae" family="Salicaceae" genus="Salix" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Salix glauca" order="Salicales" pageId="0" pageNumber="17" phylum="Angiospermae" rank="species" species="glauca">
<emphasis id="FC1B2EE81349AC08E8DE1A4F70CF7B19" italics="true" pageId="0" pageNumber="17">Salix glauca</emphasis>
</taxonomicName>
. Kangerlussuaq, 
<normalizedToken id="D09F0D87EC0770267B15158BEC5A073B" originalValue="Ringsødalen">Ringsodalen</normalizedToken>
, 
<normalizedToken id="FD5AFF0D8561F0CEA2C792920244EB62" originalValue="Ringsøen">Ringsoen</normalizedToken>
, 
<geoCoordinate id="61868634310A5210EBCEA5E9693A0147" degrees="66.9853" direction="north" orientation="latitude" precision="5" value="66.9853">66.9853°N</geoCoordinate>
, 
<geoCoordinate id="54A4011291FA13B06C2FFFFB6749311F" degrees="50.9464" direction="west" orientation="longitude" precision="5" value="-50.9464">50.9464°W</geoCoordinate>
, 9 Aug 2016, H. Knudsen (HK16.044, C-F-104080), 180 m, with 
<taxonomicName id="B6C6B81DF7C42FC75EEBFE99C6189CB8" class="Dicotyledoneae" family="Salicaceae" genus="Salix" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Salix glauca" order="Salicales" pageId="0" pageNumber="17" phylum="Angiospermae" rank="species" species="glauca">
<emphasis id="D9FC6FA27FCFC750542A71BB324C15A3" italics="true" pageId="0" pageNumber="17">Salix glauca</emphasis>
</taxonomicName>
. Kangerlussuaq, Russels Glacier, 
<geoCoordinate id="24ABC74ABE055EF7840B36169E85B222" degrees="67.0977" direction="north" orientation="latitude" precision="5" value="67.0977">67.0977°N</geoCoordinate>
, 
<geoCoordinate id="350122F3DB616BAEB53859E1FA2EA39D" degrees="50.2318" direction="west" orientation="longitude" precision="5" value="-50.2318">50.2318°W</geoCoordinate>
, 12 Aug 2000, S.A. Elborne (SAE-2000.021-GR, C-F-108472), 220 m, with 
<taxonomicName id="8026562FD73F93FF3E3D8490951C636A" class="Dicotyledoneae" family="Salicaceae" genus="Salix" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Salix glauca" order="Salicales" pageId="0" pageNumber="17" phylum="Angiospermae" rank="species" species="glauca">
<emphasis id="6810A3A84E90B749AF03A780FC8BB010" italics="true" pageId="0" pageNumber="17">Salix glauca</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="A94CBBC274C54A899EDDF8BB3AF6CBC8" pageId="0" pageNumber="17" type="distribution">
<paragraph id="5052051353D355D4603A7B449D2240CE" pageId="0" pageNumber="17">Distribution.</paragraph>
<paragraph id="0069F18342BBBCD30AE11BBC2CCFC3B2" pageId="0" pageNumber="17">This is a new species known from a number of records in the Low Arctic zone in southern and western Greenland. The fact that it was collected several times independently may indicate that it is quite common in these areas.</paragraph>
</subSubSection>
<subSubSection id="B3E333EBAF86F4D409B83B3030A67E6E" pageId="0" pageNumber="17" type="habitat">
<paragraph id="B5E37EE296549E9F0AD0BBC03365EA73" pageId="0" pageNumber="17">Habitat and ecology.</paragraph>
<paragraph id="BFB24726FAAEB3735BB7305C3FB2555F" pageId="0" pageNumber="17">
Eight collections, seven with 
<taxonomicName id="5A5DE9B93B530A97837D910AD08C64F1" class="Dicotyledoneae" family="Salicaceae" genus="Salix" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Salix glauca" order="Salicales" pageId="0" pageNumber="17" phylum="Angiospermae" rank="species" species="glauca">
<emphasis id="28E443E46798608C275605292692E109" italics="true" pageId="0" pageNumber="17">Salix glauca</emphasis>
</taxonomicName>
one with 
<taxonomicName id="70317953B9DF7F21B3CCA03E4AB85FDA" class="Dicotyledoneae" family="Salicaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="S. arctophila" order="Salicales" pageId="0" pageNumber="17" phylum="Angiospermae" rank="species" species="arctophila">
<emphasis id="7C86D4339B1901C8FEFDF1FEDD365648" italics="true" pageId="0" pageNumber="17">S. arctophila</emphasis>
</taxonomicName>
. Most collections are from mineral-poor ground.
</paragraph>
</subSubSection>
<subSubSection id="924D7666427923F599C0543A49B15658" pageId="0" pageNumber="17" type="notes">
<paragraph id="9478FCFD2C1994CDE812781FAD6451B7" pageId="0" pageNumber="17">Notes.</paragraph>
<paragraph id="46E38205FC7C8CE89968707A667B75A9" pageId="0" pageNumber="17">
The mainly clavate-stipitate cheilocystidia indicate that this species belongs in 
<taxonomicName id="228DFA8AE8C56D779EAC267796A0188D" class="Dicotyledoneae" family="Salicaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="H. subsp. sect." order="Salicales" pageId="0" pageNumber="17" phylum="Angiospermae" rank="section" section="Denudata">H. sect. Denudata</taxonomicName>
and there in 
<taxonomicName id="1224A379456AFF47D7CE1BCD26452970" class="Dicotyledoneae" family="Salicaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="H. subsp. subsect." order="Salicales" pageId="0" pageNumber="17" phylum="Angiospermae" rank="subSection" subSection="Crustuliniformia">H. subsect. Crustuliniformia</taxonomicName>
.Few species of the subsection have almost smooth spores or spores that are rather strongly dextrinoid. The combination of almost smooth and rather strongly dextrinoid spores is unique in this subsection. Molecular results support its placement within this subsection. Within 
<taxonomicName id="BF2BE7D26FFFC6170C806ADCBA41A0C0" class="Dicotyledoneae" family="Salicaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="H. subsp. sect." order="Salicales" pageId="0" pageNumber="17" phylum="Angiospermae" rank="section" section="Denudata">H. sect. Denudata</taxonomicName>
, based on the spore characters, 
<taxonomicName id="BBB407BAF2DCD8CA264CC858D8B5833F" class="Dicotyledoneae" family="Salicaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="H. arcticum" order="Salicales" pageId="0" pageNumber="17" phylum="Angiospermae" rank="species" species="arcticum">
<emphasis id="33100F6F96EDD83F40350DB3782E1FDE" italics="true" pageId="0" pageNumber="17">H. arcticum</emphasis>
</taxonomicName>
is readily identifiable. To be noted is also the unusual shape of the spores; while many spores are amygdaloid, many others appear more ellipsoid or cylindrical, again uncommon for this section of 
<taxonomicName id="2623A91FD30CD5159152D7635779AFF6" class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" kingdom="Fungi" lsidName="Hebeloma" order="Ostropales" pageId="0" pageNumber="17" phylum="Basidiomycota" rank="genus">
<emphasis id="5535FEB40D63657FA34E10F5D58C5184" italics="true" pageId="0" pageNumber="17">Hebeloma</emphasis>
</taxonomicName>
. As Figs 
<figureCitation id="7A43F09493062188BF71BB98C4722B0C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A Pruned quasi-median network of the Hebeloma alpinum complex, H. arcticum and H. louiseae. Circles shared by two or more taxa or collections from two regions, from Greenland (G) or collected elsewhere, are divided according to the number of representatives for each class B ML result for Hebeloma subsect. Crustuliniforme, based on concatenated ITS, RPB 2 and Tef 1 a data, rooted with H. louiseae. Thick branches indicate bootstrap support of at least 80 % (450 bootstrap replicates). Species are represented by three collections (if available); Hebeloma arcticum by four collections including the holotype." figureDoi="10.3897/mycokeys.79.63363.figure5" httpUri="https://binary.pensoft.net/fig/533396" pageId="0" pageNumber="17">5A-B</figureCitation>
show, 
<taxonomicName id="45C6F516968C0C4B6004CAC358AA1E1A" class="Dicotyledoneae" family="Salicaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="H. arcticum" order="Salicales" pageId="0" pageNumber="17" phylum="Angiospermae" rank="species" species="arcticum">
<emphasis id="2FB5C92B12E3DC2C2202761A0673F78F" italics="true" pageId="0" pageNumber="17">H. arcticum</emphasis>
</taxonomicName>
is well distinct from related taxa and is in a supported clade (Fig. 
<figureCitation id="BD239918CBD5EB590C38BAA4AD48A5C9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A Pruned quasi-median network of the Hebeloma alpinum complex, H. arcticum and H. louiseae. Circles shared by two or more taxa or collections from two regions, from Greenland (G) or collected elsewhere, are divided according to the number of representatives for each class B ML result for Hebeloma subsect. Crustuliniforme, based on concatenated ITS, RPB 2 and Tef 1 a data, rooted with H. louiseae. Thick branches indicate bootstrap support of at least 80 % (450 bootstrap replicates). Species are represented by three collections (if available); Hebeloma arcticum by four collections including the holotype." figureDoi="10.3897/mycokeys.79.63363.figure5" httpUri="https://binary.pensoft.net/fig/533396" pageId="0" pageNumber="17">5B</figureCitation>
) with other members of 
<taxonomicName id="92B4532B5845E6DC4C929FE53500DB55" class="Dicotyledoneae" family="Salicaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="H. subsp. subsect." order="Salicales" pageId="0" pageNumber="17" phylum="Angiospermae" rank="subSection" subSection="Crustuliniformia">H. subsect. Crustuliniformia</taxonomicName>
, but is not a member of the 
<taxonomicName id="A16CFA199F31FC043601AA413BF35FCF" class="Dicotyledoneae" family="Salicaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="H. alpinum" order="Salicales" pageId="0" pageNumber="17" phylum="Angiospermae" rank="species" species="alpinum">
<emphasis id="5A5801B37EA22672BCE0B85055839B05" italics="true" pageId="0" pageNumber="17">H. alpinum</emphasis>
</taxonomicName>
-complex. The clade of the species is supported by 100% bootstrap in the analysis of three loci (Fig. 
<figureCitation id="9575009E9666BD5BB6AD5821A9815A44" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A Pruned quasi-median network of the Hebeloma alpinum complex, H. arcticum and H. louiseae. Circles shared by two or more taxa or collections from two regions, from Greenland (G) or collected elsewhere, are divided according to the number of representatives for each class B ML result for Hebeloma subsect. Crustuliniforme, based on concatenated ITS, RPB 2 and Tef 1 a data, rooted with H. louiseae. Thick branches indicate bootstrap support of at least 80 % (450 bootstrap replicates). Species are represented by three collections (if available); Hebeloma arcticum by four collections including the holotype." figureDoi="10.3897/mycokeys.79.63363.figure5" httpUri="https://binary.pensoft.net/fig/533396" pageId="0" pageNumber="17">5B</figureCitation>
), but based on current knowledge, ITS alone is sufficient to recognize the taxon; in the ITS tree (not shown) calculated prior to concatenation of the data, 
<taxonomicName id="95011408BC776C9AA36572CFB2457BC8" class="Dicotyledoneae" family="Salicaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="H. arcticum" order="Salicales" pageId="0" pageNumber="17" phylum="Angiospermae" rank="species" species="arcticum">
<emphasis id="3E2F773304D45A9C9CEC22856491B3F6" italics="true" pageId="0" pageNumber="17">H. arcticum</emphasis>
</taxonomicName>
received 84% bootstrap support.
</paragraph>
</subSubSection>
</treatment>
</document>