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<document id="9C3D36ACC511F7F7906B3D400BCDE4A8" ID-CLB-Dataset="38713" ID-DOI="10.11646/zootaxa.4066.4.4" ID-GBIF-Dataset="d7e1540e-9a08-41e6-8d31-408b77fdf9c1" ID-ISSN="1175-5326" ID-Zenodo-Dep="258317" ID-ZooBank="03DC485E-A9B6-4E5B-93D8-A7639BB4BF22" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1466871784755" checkinUser="plazi" docAuthor="González-Durán, Gustavo A." docDate="2016" docId="7E1C7771FFDABD04CAAE34D9FA50882C" docLanguage="en" docName="zootaxa.4066.4.4.pdf.imf" docOrigin="Zootaxa 4066 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pristimantis stictus González-Durán, 2016, sp. nov." docType="treatment" docVersion="8" lastPageNumber="435" masterDocId="82250F09FFDBBD0ACA39321FFFCD8B71" masterDocTitle="A new small frog species of the genus Pristimantis (Anura: Craugastoridae) from the northern paramos of Colombia" masterLastPageNumber="437" masterPageNumber="421" pageNumber="422" updateTime="1698663197653" updateUser="ExternalLinkService">
<mods:mods id="BA3F54E653DEE8239A54DB674CE0F854" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="1BE56567819E2C7C8E6A95CA3A249C8D">A new small frog species of the genus Pristimantis (Anura: Craugastoridae) from the northern paramos of Colombia</mods:title>
</mods:titleInfo>
<mods:name id="B0E6B831AC25EB9DBF75C5EB479F9C33" type="personal">
<mods:role id="48BC6A70611A72016B19712E2118F196">
<mods:roleTerm id="7092489092EE9B7EF75813A557E91E0E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A1AFFAAE5ABE65CAF4CCFE3AE52DEDC7">González-Durán, Gustavo A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="9D75625B8341446ABA929E5216823536">text</mods:typeOfResource>
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<mods:title id="980A182CCA4A83EEA4BE8EC50A954C3B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="73BE0A5B7302307316FF8CCB8B146122">
<mods:date id="715A9BA2E4AB49784EFB64147138C0C0">2016</mods:date>
<mods:detail id="0D4F22085EA561125D27DD1DF6065DB4" type="volume">
<mods:number id="B45E59C06C822E68820847FAF00ED0C5">4066</mods:number>
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<mods:number id="D7727D2116EF232062F64978345879B4">4</mods:number>
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<mods:classification id="9B85DA7465C5442786C9094F6C354049">journal article</mods:classification>
<mods:identifier id="7F5087B54216386B73FD98E480E7C631" type="CLB-Dataset">38713</mods:identifier>
<mods:identifier id="C4ECF1F946540EC5F9B6050695E66063" type="DOI">10.11646/zootaxa.4066.4.4</mods:identifier>
<mods:identifier id="BDA4856322C10C468A1A3A998C2911FC" type="GBIF-Dataset">d7e1540e-9a08-41e6-8d31-408b77fdf9c1</mods:identifier>
<mods:identifier id="308585D956EB27B30A7091BD9105FFC9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="CB9F6FA138D0BADE8E19BF406EF91EF8" type="Zenodo-Dep">258317</mods:identifier>
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</mods:mods>
<treatment id="7E1C7771FFDABD04CAAE34D9FA50882C" ID-DOI="http://doi.org/10.5281/zenodo.6058401" ID-GBIF-Taxon="121067087" ID-Zenodo-Dep="6058401" LSID="urn:lsid:plazi:treatment:7E1C7771FFDABD04CAAE34D9FA50882C" httpUri="http://treatment.plazi.org/id/7E1C7771FFDABD04CAAE34D9FA50882C" lastPageId="14" lastPageNumber="435" pageId="1" pageNumber="422">
<subSubSection id="BEAF95ECFFDABD0BCAAE34D9FEE28C73" pageId="1" pageNumber="422" type="nomenclature">
<paragraph id="F60AC667FFDABD0BCAAE34D9FEE28C73" blockId="1.[151,478,1734,1794]" pageId="1" pageNumber="422">
<taxonomicName id="31B5BDE4FFDABD0BCAAE34D9FEB38D91" box="[151,382,1734,1760]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="1" pageNumber="422" phylum="Chordata" rank="species" species="stictus" status="sp. nov.">
<emphasis id="C4C11A75FFDABD0BCAAE34D9FEB38D91" bold="true" box="[151,382,1734,1760]" italics="true" pageId="1" pageNumber="422">Pristimantis stictus</emphasis>
</taxonomicName>
<emphasis id="C4C11A75FFDABD0BCBBD34D8FE138D91" bold="true" box="[388,478,1735,1760]" pageId="1" pageNumber="422">
<taxonomicNameLabel id="DFF2A70EFFDABD0BCBBD34D8FE138D91" box="[388,478,1735,1760]" pageId="1" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="6E8EDAE2FFDABD0BCAA734F6FEE88C73" box="[158,293,1769,1794]" captionStart-0="FIGURE 1. A" captionStart-1="FIGURE 2. A" captionStartId-0="2.[151,250,1748,1770]" captionStartId-1="3.[151,250,1912,1934]" captionTargetBox-0="[264,1322,265,1725]" captionTargetBox-1="[192,1408,607,1858]" captionTargetId-0="figure@2.[264,1323,265,1727]" captionTargetId-1="figure@3.[151,1436,541,1898]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. A. Ventral view of Pristimantis stictus sp. nov., adult female holotype, (ICN- 55689). B. Dorsal view of P. stictus sp. nov., adult female holotype, (ICN- 55689). C. Ventral view of hand of P. stictus sp. nov., adult female holotype, (ICN- 55689). D. Ventral view of foot of P. stictus sp. nov., adult female holotype, (ICN- 55689). E. Lateral view of P. stictus sp. nov., adult female holotype (ICN- 55689). White bars equal 4 mm." captionText-1="FIGURE 2. A. Lateral aspect of Pristimantis stictus sp. nov., adult male (ICN- 55740). B. Lateral view of P. stictus sp. nov., adult female (ICN 55715). C. Lateral view of P. stictus sp. nov., adult female (ICN- 55713). D. Lateral aspect of P. stictus sp. nov., adult female (ICN- 55716). E. Ventral view of P. stictus sp. nov., adult female (ICN- 55713). F. Ventral view of P. stictus sp. nov., adult female (ICN- 55716)." httpUri-0="https://zenodo.org/record/258318/files/figure.png" httpUri-1="https://zenodo.org/record/258319/files/figure.png" pageId="1" pageNumber="422">Figures 1–2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="BEAF95ECFFDABD09CAAE352EFDC88ADD" lastPageId="3" lastPageNumber="424" pageId="1" pageNumber="422" type="materials_examined">
<paragraph id="F60AC667FFDABD0BCAAE352EFA8E8C1F" blockId="1.[151,1436,1841,2010]" pageId="1" pageNumber="422">
<emphasis id="C4C11A75FFDABD0BCAAE352EFEC68C3B" bold="true" box="[151,267,1841,1866]" pageId="1" pageNumber="422">
<typeStatus id="290E78C5FFDABD0BCAAE352EFECB8C3B" box="[151,262,1841,1866]" pageId="1" pageNumber="422" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="90A45EA2FFDABD0BCB2A352DFE8E8C3B" box="[275,323,1842,1866]" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="1" pageNumber="422">ICN</collectionCode>
55689, adult female collected at the vereda el Vergel, Municipality of Marulanda, Department of Caldas,
<collectingCountry id="8EA286F7FFDABD0BCAC8354AFE928C1F" box="[241,351,1877,1902]" name="Colombia" pageId="1" pageNumber="422">Colombia</collectingCountry>
(
<geoCoordinate id="9381A0A0FFDABD0BCB57354AFE168C1C" box="[366,475,1877,1902]" direction="north" orientation="latitude" pageId="1" pageNumber="422" precision="5" value="5.2372">5.2372 N</geoCoordinate>
,
<geoCoordinate id="9381A0A0FFDABD0BCBDF354AFDA88C1C" box="[486,613,1877,1902]" direction="west" orientation="longitude" pageId="1" pageNumber="422" precision="5" value="-75.3512">75.3512 W</geoCoordinate>
;
<quantity id="314D6B82FFDABD0BC8493549FD068C1F" box="[624,715,1877,1902]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="1" pageNumber="422" unit="m" value="3700.0">3700 m</quantity>
.a.s.l), by Gustavo Gonzalez-Duran, on
<date id="820BE0A7FFDABD0BCEBB3549FAF28C1F" box="[1154,1343,1878,1902]" pageId="1" pageNumber="422" value="2010-01-30">30 January 2010</date>
.
</paragraph>
<paragraph id="F60AC667FFDABD08CAFE3566FF288BA5" blockId="1.[151,1436,1841,2010]" lastBlockId="2.[151,1436,151,212]" lastPageId="2" lastPageNumber="423" pageId="1" pageNumber="422">
<emphasis id="C4C11A75FFDABD0BCAFE3566FE8B8CE3" bold="true" box="[199,326,1913,1938]" pageId="1" pageNumber="422">
<typeStatus id="290E78C5FFDABD0BCAFE3566FE8C8CE3" box="[199,321,1913,1938]" pageId="1" pageNumber="422" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Nine adult females ICN-55690-92, 55696-99, 55702, 55704 and eight adult males ICN-55693-95, 55700-01, 55703, 55705-06, collected with the
<typeStatus id="290E78C5FFDABD0BC8823582FCD08CC7" box="[699,797,1949,1974]" pageId="1" pageNumber="422" type="holotype">holotype</typeStatus>
on
<date id="820BE0A7FFDABD0BC9753581FBF58CC7" box="[844,1080,1950,1974]" pageId="1" pageNumber="422" value="2010-01-29" valueMax="2010-01-30" valueMin="2010-01-29">29–30 January 2010</date>
. Six adult females ICN-55710, 55712-16 and four adult males ICN-55707-09, 55711, collected on
<date id="820BE0A7FFDABD0BC9DB35DDFB128CAB" box="[994,1247,1985,2010]" pageId="1" pageNumber="422" value="2012-11-03" valueMax="2012-11-04" valueMin="2012-11-03">3–4 November 2012</date>
, in San Pablo, Municipality of Neira, Department of Caldas,
<collectingCountry id="8EA286F7FFD9BD08C8913288FCDB8BC1" box="[680,790,151,176]" name="Colombia" pageId="2" pageNumber="423">Colombia</collectingCountry>
(
<geoCoordinate id="9381A0A0FFD9BD08C91F3288FC598BDE" box="[806,916,151,176]" direction="north" orientation="latitude" pageId="2" pageNumber="423" precision="5" value="5.1193">5.1193 N</geoCoordinate>
,
<geoCoordinate id="9381A0A0FFD9BD08C9983288FBEF8BDE" box="[929,1058,151,176]" direction="west" orientation="longitude" pageId="2" pageNumber="423" precision="5" value="-75.3234">75.3234 W</geoCoordinate>
;
<quantity id="314D6B82FFD9BD08CE163287FB458BC1" box="[1071,1160,152,176]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="2" pageNumber="423" unit="m" value="3700.0">3700 m</quantity>
), by Gustavo Gonzalez- Durán.
</paragraph>
<caption id="A2CA96EFFFD9BD08CAAE34CBFD338C37" httpUri="https://zenodo.org/record/258318/files/figure.png" pageId="2" pageNumber="423" targetBox="[264,1322,265,1725]" targetPageId="2">
<paragraph id="F60AC667FFD9BD08CAAE34CBFD338C37" blockId="2.[151,1436,1748,1862]" pageId="2" pageNumber="423">
<emphasis id="C4C11A75FFD9BD08CAAE34CBFEDE8D98" bold="true" box="[151,275,1748,1770]" pageId="2" pageNumber="423">FIGURE 1.</emphasis>
A. Ventral view of
<taxonomicName id="31B5BDE4FFD9BD08CBE734CAFD6F8D9B" box="[478,674,1749,1770]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="423" phylum="Chordata" rank="species" species="stictus" status="sp. nov.">
<emphasis id="C4C11A75FFD9BD08CBE734CAFD6F8D9B" box="[478,674,1749,1770]" italics="true" pageId="2" pageNumber="423">Pristimantis stictus</emphasis>
</taxonomicName>
<emphasis id="C4C11A75FFD9BD08C89034CAFD348D9B" bold="true" box="[681,761,1749,1770]" pageId="2" pageNumber="423">
<taxonomicNameLabel id="DFF2A70EFFD9BD08C89034CAFD348D9B" box="[681,761,1749,1770]" pageId="2" pageNumber="423" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult female holotype, (ICN-55689). B. Dorsal view of
<taxonomicName id="31B5BDE4FFD9BD08CF7834CAFA518D9B" box="[1345,1436,1749,1770]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="423" phylum="Chordata" rank="species" species="stictus" status="sp. nov.">
<emphasis id="C4C11A75FFD9BD08CF7834CAFA808D9B" box="[1345,1357,1749,1770]" italics="true" pageId="2" pageNumber="423">P</emphasis>
.
<emphasis id="C4C11A75FFD9BD08CF6234CAFA518D9B" box="[1371,1436,1749,1770]" italics="true" pageId="2" pageNumber="423">stictus</emphasis>
</taxonomicName>
<emphasis id="C4C11A75FFD9BD08CAAE34ECFF278C79" bold="true" box="[151,234,1779,1800]" pageId="2" pageNumber="423">
<taxonomicNameLabel id="DFF2A70EFFD9BD08CAAE34ECFF278C79" box="[151,234,1779,1800]" pageId="2" pageNumber="423" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult female holotype, (ICN-55689). C. Ventral view of hand of
<taxonomicName id="31B5BDE4FFD9BD08C99D34EBFBCC8C78" box="[932,1025,1780,1801]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="423" phylum="Chordata" rank="species" species="stictus" status="sp. nov.">
<emphasis id="C4C11A75FFD9BD08C99D34EBFC7C8C78" box="[932,945,1780,1801]" italics="true" pageId="2" pageNumber="423">P</emphasis>
.
<emphasis id="C4C11A75FFD9BD08C9F934EBFBCC8C78" box="[960,1025,1780,1801]" italics="true" pageId="2" pageNumber="423">stictus</emphasis>
</taxonomicName>
<emphasis id="C4C11A75FFD9BD08CE3334ECFB908C79" bold="true" box="[1034,1117,1779,1800]" pageId="2" pageNumber="423">
<taxonomicNameLabel id="DFF2A70EFFD9BD08CE3334ECFB908C79" box="[1034,1117,1779,1800]" pageId="2" pageNumber="423" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult female holotype, (ICN- 55689). D. Ventral view of foot of
<taxonomicName id="31B5BDE4FFD9BD08CBCD350DFD838C56" box="[500,590,1810,1831]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="423" phylum="Chordata" rank="species" species="stictus" status="sp. nov.">
<emphasis id="C4C11A75FFD9BD08CBCD350DFDCD8C56" box="[500,512,1810,1831]" italics="true" pageId="2" pageNumber="423">P</emphasis>
.
<emphasis id="C4C11A75FFD9BD08C834350DFD838C56" box="[525,590,1810,1831]" italics="true" pageId="2" pageNumber="423">stictus</emphasis>
</taxonomicName>
<emphasis id="C4C11A75FFD9BD08C86D350DFD698C56" bold="true" box="[596,676,1810,1831]" pageId="2" pageNumber="423">
<taxonomicNameLabel id="DFF2A70EFFD9BD08C86D350DFD698C56" box="[596,676,1810,1831]" pageId="2" pageNumber="423" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult female holotype, (ICN-55689). E. Lateral view of
<taxonomicName id="31B5BDE4FFD9BD08CEDC350DFAF28C56" box="[1253,1343,1810,1831]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="2" pageNumber="423" phylum="Chordata" rank="species" species="stictus" status="sp. nov.">
<emphasis id="C4C11A75FFD9BD08CEDC350DFB3C8C56" box="[1253,1265,1810,1831]" italics="true" pageId="2" pageNumber="423">P</emphasis>
.
<emphasis id="C4C11A75FFD9BD08CEC7350DFAF28C56" box="[1278,1343,1810,1831]" italics="true" pageId="2" pageNumber="423">stictus</emphasis>
</taxonomicName>
<emphasis id="C4C11A75FFD9BD08CF7C350DFA588C56" bold="true" box="[1349,1429,1810,1831]" pageId="2" pageNumber="423">
<taxonomicNameLabel id="DFF2A70EFFD9BD08CF7C350DFA588C56" box="[1349,1429,1810,1831]" pageId="2" pageNumber="423" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult female holotype (ICN-55689). White bars equal 4 mm.
</paragraph>
</caption>
<paragraph id="F60AC667FFD9BD09CAFE356CFDC88ADD" blockId="2.[151,1437,1907,2004]" lastBlockId="3.[151,1437,151,501]" lastPageId="3" lastPageNumber="424" pageId="2" pageNumber="423">
<emphasis id="C4C11A75FFD9BD08CAFE356CFE768CFD" bold="true" box="[199,443,1907,1932]" pageId="2" pageNumber="423">Referred specimens.</emphasis>
ICN-14417 one adult male and ICN-14418-23, 14432 eight adult females collected on
<date id="820BE0A7FFD9BD08CAAE3588FEFC8CDE" box="[151,305,1943,1967]" pageId="2" pageNumber="423" value="1984-06-23">
23 June
<quantity id="314D6B82FFD9BD08CAC03588FEFC8CDE" box="[249,305,1943,1967]" metricMagnitude="1" metricUnit="m" metricValue="5.03936" pageId="2" pageNumber="423" unit="in" value="1984.0">1984</quantity>
</date>
in the vicinity of the Hotel Termales del Ruiz, Municipality of Villamaria, Department of Caldas,
<collectingCountry id="8EA286F7FFD9BD08CAAE35A4FEC88CA5" box="[151,261,1979,2004]" name="Colombia" pageId="2" pageNumber="423">Colombia</collectingCountry>
(
<geoCoordinate id="9381A0A0FFD9BD08CB2A35A4FEB28CA2" box="[275,383,1979,2003]" direction="north" orientation="latitude" pageId="2" pageNumber="423" precision="5" value="4.9711">4.9711 N</geoCoordinate>
,
<geoCoordinate id="9381A0A0FFD9BD08CBB335A4FDC48CA2" box="[394,521,1979,2004]" direction="west" orientation="longitude" pageId="2" pageNumber="423" precision="5" value="-75.3844">75.3844 W</geoCoordinate>
;
<quantity id="314D6B82FFD9BD08C82A35A4FDA18CA2" box="[531,620,1979,2003]" metricMagnitude="3" metricUnit="m" metricValue="3.37" pageId="2" pageNumber="423" unit="m" value="3370.0">3370 m</quantity>
). ICN-55591-
<specimenCount id="E0B30DEEFFD9BD08C93135A4FCAF8CA5" box="[776,866,1979,2004]" pageId="2" pageNumber="423" type="adult">92 adult</specimenCount>
female collected on
<date id="820BE0A7FFD9BD08CE7235A4FA868CA2" box="[1099,1355,1979,2004]" pageId="2" pageNumber="423" value="2011-11-28" valueMax="2011-11-29" valueMin="2011-11-28">28–29 November 2011</date>
vereda El Desquite, Municipality of Manizales, Department of Caldas,
<collectingCountry id="8EA286F7FFD8BD09C9573288FC118BC1" box="[878,988,151,176]" name="Colombia" pageId="3" pageNumber="424">Colombia</collectingCountry>
(
<geoCoordinate id="9381A0A0FFD8BD09C9D53288FB978BDE" box="[1004,1114,151,176]" direction="north" orientation="latitude" pageId="3" pageNumber="424" precision="5" value="5.065">5.0650 N</geoCoordinate>
,
<geoCoordinate id="9381A0A0FFD8BD09CE5E3288FB2A8BDE" box="[1127,1255,151,176]" direction="west" orientation="longitude" pageId="3" pageNumber="424" precision="5" value="-75.3716">75.3716 W</geoCoordinate>
;
<quantity id="314D6B82FFD8BD09CECD3288FA838BC1" box="[1268,1358,151,176]" metricMagnitude="3" metricUnit="m" metricValue="3.75" pageId="3" pageNumber="424" unit="m" value="3750.0">3750 m</quantity>
). ICN- 55717 adult female collected on
<date id="820BE0A7FFD8BD09C81932A3FCCE8BA5" box="[544,771,188,213]" pageId="3" pageNumber="424" value="2007-11-25">
25 November
<quantity id="314D6B82FFD8BD09C8F232A3FCCE8BA5" box="[715,771,188,212]" metricMagnitude="1" metricUnit="m" metricValue="5.097779999999999" pageId="3" pageNumber="424" unit="in" value="2007.0">2007</quantity>
</date>
in the vereda Hojas Anchas, Municipality of Salamina, Department of Caldas,
<collectingCountry id="8EA286F7FFD8BD09CB9832C0FDC28B89" box="[417,527,223,248]" name="Colombia" pageId="3" pageNumber="424">Colombia</collectingCountry>
(
<geoCoordinate id="9381A0A0FFD8BD09C81832C0FD5D8B86" box="[545,656,223,248]" direction="north" orientation="latitude" pageId="3" pageNumber="424" precision="5" value="5.2423">5.2423 N</geoCoordinate>
,
<geoCoordinate id="9381A0A0FFD8BD09C8A432C0FCD38B86" box="[669,798,223,248]" direction="west" orientation="longitude" pageId="3" pageNumber="424" precision="5" value="-75.3702">75.3702 W</geoCoordinate>
;
<quantity id="314D6B82FFD8BD09C91532C0FC4A8B89" box="[812,903,223,248]" metricMagnitude="3" metricUnit="m" metricValue="3.67" pageId="3" pageNumber="424" unit="m" value="3670.0">3670 m</quantity>
). ICN-55718-31 collected on
<date id="820BE0A7FFD8BD09CEE632C0FA518B89" box="[1247,1436,223,248]" pageId="3" pageNumber="424" value="2014-03-01" valueMax="2014-03-05" valueMin="2014-03-01">1–5 March 2014</date>
vereda El Desquite, Municipality of Manizales, Department of Caldas,
<collectingCountry id="8EA286F7FFD8BD09C98E331BFBE88A6C" box="[951,1061,260,285]" name="Colombia" pageId="3" pageNumber="424">Colombia</collectingCountry>
(
<geoCoordinate id="9381A0A0FFD8BD09CE0D331BFB6C8A6D" box="[1076,1185,260,285]" direction="north" orientation="latitude" pageId="3" pageNumber="424" precision="5" value="5.0651">5.0651 N</geoCoordinate>
,
<geoCoordinate id="9381A0A0FFD8BD09CE95331BFAE68A6D" box="[1196,1323,260,285]" direction="west" orientation="longitude" pageId="3" pageNumber="424" precision="5" value="-75.3791">75.3791 W</geoCoordinate>
;
<quantity id="314D6B82FFD8BD09CF0E331BFA5D8A6D" box="[1335,1424,260,285]" metricMagnitude="3" metricUnit="m" metricValue="3.62" pageId="3" pageNumber="424" unit="m" value="3620.0">3620 m</quantity>
). ICN-55732-
<specimenCount id="E0B30DEEFFD8BD09CB183338FE638A31" box="[289,430,295,320]" pageId="3" pageNumber="424" type="juvenile">33 juveniles</specimenCount>
collected on
<date id="820BE0A7FFD8BD09C8603337FCD28A31" box="[601,799,296,320]" pageId="3" pageNumber="424" value="2010-01-30">
30 January
<quantity id="314D6B82FFD8BD09C8DE3337FCD28A31" box="[743,799,296,320]" metricMagnitude="1" metricUnit="m" metricValue="5.1053999999999995" pageId="3" pageNumber="424" unit="in" value="2010.0">2010</quantity>
</date>
in the vereda el Vergel, Municipality of Marulanda, Department of Caldas,
<collectingCountry id="8EA286F7FFD8BD09CBA03353FDCA8A14" box="[409,519,332,357]" name="Colombia" pageId="3" pageNumber="424">Colombia</collectingCountry>
(
<geoCoordinate id="9381A0A0FFD8BD09C82C3353FD4F8A15" box="[533,642,332,357]" direction="north" orientation="latitude" pageId="3" pageNumber="424" precision="5" value="5.2372">5.2372 N</geoCoordinate>
,
<geoCoordinate id="9381A0A0FFD8BD09C8B43353FCC68A15" box="[653,779,332,357]" direction="west" orientation="longitude" pageId="3" pageNumber="424" precision="5" value="-75.3512">75.3512 W</geoCoordinate>
;
<quantity id="314D6B82FFD8BD09C92F3353FCA38A15" box="[790,878,332,356]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="3" pageNumber="424" unit="m" value="3700.0">3700 m</quantity>
). ICN-55734-39 six juveniles and ICN-55741 one adult male collected on
<date id="820BE0A7FFD8BD09CB99336FFD458AF9" box="[416,648,367,392]" pageId="3" pageNumber="424" value="2012-11-03" valueMax="2012-11-04" valueMin="2012-11-03">3–4 November 2012</date>
, in San Pablo, Municipality of Neira, Department of Caldas,
<collectingCountry id="8EA286F7FFD8BD09CF173370FA518AF9" box="[1326,1436,367,392]" name="Colombia" pageId="3" pageNumber="424">Colombia</collectingCountry>
(
<geoCoordinate id="9381A0A0FFD8BD09CAA7338BFEC78ADD" box="[158,266,404,429]" direction="north" orientation="latitude" pageId="3" pageNumber="424" precision="5" value="5.1193">5.1193 N</geoCoordinate>
,
<geoCoordinate id="9381A0A0FFD8BD09CB2C338BFE598ADD" box="[277,404,404,429]" direction="west" orientation="longitude" pageId="3" pageNumber="424" precision="5" value="-75.3234">75.3234 W</geoCoordinate>
;
<quantity id="314D6B82FFD8BD09CBA6338BFE348ADD" box="[415,505,404,428]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="3" pageNumber="424" unit="m" value="3700.0">3700 m</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="BEAF95ECFFD8BD0ECAFE33A8FC7D8CA7" lastPageId="4" lastPageNumber="425" pageId="3" pageNumber="424" type="etymology">
<paragraph id="F60AC667FFD8BD09CAFE33A8FD9F8A84" blockId="3.[151,1437,151,501]" pageId="3" pageNumber="424">
<emphasis id="C4C11A75FFD8BD09CAFE33A8FE838AA1" bold="true" box="[199,334,439,464]" pageId="3" pageNumber="424">Etymology.</emphasis>
The specific epithet s
<emphasis id="C4C11A75FFD8BD09C87833A7FDB38ABE" box="[577,638,440,463]" italics="true" pageId="3" pageNumber="424">tictus</emphasis>
is derived from greek
<emphasis id="C4C11A75FFD8BD09C94333A7FC0F8AA1" box="[890,962,440,464]" italics="true" pageId="3" pageNumber="424">stiktos</emphasis>
(spotted) due to the spots on the groin and belly. The name is used as an adjective.
</paragraph>
<caption id="A2CA96EFFFD8BD09CAAE3567FDC98C9B" httpUri="https://zenodo.org/record/258319/files/figure.png" pageId="3" pageNumber="424" targetBox="[192,1408,607,1858]" targetPageId="3">
<paragraph id="F60AC667FFD8BD09CAAE3567FDC98C9B" blockId="3.[151,1436,1912,2026]" pageId="3" pageNumber="424">
<emphasis id="C4C11A75FFD8BD09CAAE3567FED98CFC" bold="true" box="[151,276,1912,1934]" pageId="3" pageNumber="424">FIGURE 2.</emphasis>
A. Lateral aspect of
<taxonomicName id="31B5BDE4FFD8BD09CBD73566FD7F8CFF" box="[494,690,1913,1934]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="424" phylum="Chordata" rank="species" species="stictus" status="sp. nov.">
<emphasis id="C4C11A75FFD8BD09CBD73566FD7F8CFF" box="[494,690,1913,1934]" italics="true" pageId="3" pageNumber="424">Pristimantis stictus</emphasis>
</taxonomicName>
<emphasis id="C4C11A75FFD8BD09C8803567FCC78CFC" bold="true" box="[697,778,1912,1933]" pageId="3" pageNumber="424">
<taxonomicNameLabel id="DFF2A70EFFD8BD09C8803567FCC78CFC" box="[697,778,1912,1933]" pageId="3" pageNumber="424" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult male (ICN-55740). B. Lateral view of
<taxonomicName id="31B5BDE4FFD8BD09CED83566FAF18CFF" box="[1249,1340,1913,1934]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="424" phylum="Chordata" rank="species" species="stictus" status="sp. nov.">
<emphasis id="C4C11A75FFD8BD09CED83566FB208CFF" box="[1249,1261,1913,1934]" italics="true" pageId="3" pageNumber="424">P</emphasis>
.
<emphasis id="C4C11A75FFD8BD09CEC23566FAF18CFF" box="[1275,1340,1913,1934]" italics="true" pageId="3" pageNumber="424">stictus</emphasis>
</taxonomicName>
<emphasis id="C4C11A75FFD8BD09CF7D3567FA588CFC" bold="true" box="[1348,1429,1912,1933]" pageId="3" pageNumber="424">
<taxonomicNameLabel id="DFF2A70EFFD8BD09CF7D3567FA588CFC" box="[1348,1429,1912,1933]" pageId="3" pageNumber="424" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult female (ICN55715). C. Lateral view of
<taxonomicName id="31B5BDE4FFD8BD09C8523587FD0A8CDC" box="[619,711,1943,1965]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="424" phylum="Chordata" rank="species" species="stictus" status="sp. nov.">
<emphasis id="C4C11A75FFD8BD09C8523587FDBA8CDC" box="[619,631,1944,1965]" italics="true" pageId="3" pageNumber="424">P</emphasis>
.
<emphasis id="C4C11A75FFD8BD09C8BF3587FD0A8CDC" box="[646,711,1944,1965]" italics="true" pageId="3" pageNumber="424">stictus</emphasis>
</taxonomicName>
<emphasis id="C4C11A75FFD8BD09C8F73588FCED8CDD" bold="true" box="[718,800,1943,1964]" pageId="3" pageNumber="424">
<taxonomicNameLabel id="DFF2A70EFFD8BD09C8F73588FCED8CDD" box="[718,800,1943,1964]" pageId="3" pageNumber="424" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult female (ICN-55713). D. Lateral aspect of
<taxonomicName id="31B5BDE4FFD8BD09CF223587FABB8CDC" box="[1307,1398,1943,1965]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="424" phylum="Chordata" rank="species" species="stictus" status="sp. nov.">
<emphasis id="C4C11A75FFD8BD09CF223587FAE58CDC" box="[1307,1320,1944,1965]" italics="true" pageId="3" pageNumber="424">P</emphasis>
.
<emphasis id="C4C11A75FFD8BD09CF0C3587FABB8CDC" box="[1333,1398,1944,1965]" italics="true" pageId="3" pageNumber="424">stictus</emphasis>
</taxonomicName>
<emphasis id="C4C11A75FFD8BD09CF443588FF0E8CBA" bold="true" pageId="3" pageNumber="424">
<taxonomicNameLabel id="DFF2A70EFFD8BD09CF443588FF0E8CBA" pageId="3" pageNumber="424" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult female (ICN-55716). E. Ventral view of
<taxonomicName id="31B5BDE4FFD8BD09C89335A9FCC88CBA" box="[682,773,1974,1995]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="424" phylum="Chordata" rank="species" species="stictus" status="sp. nov.">
<emphasis id="C4C11A75FFD8BD09C89335A9FD7B8CBA" box="[682,694,1974,1995]" italics="true" pageId="3" pageNumber="424">P</emphasis>
.
<emphasis id="C4C11A75FFD8BD09C8FD35A9FCC88CBA" box="[708,773,1974,1995]" italics="true" pageId="3" pageNumber="424">stictus</emphasis>
</taxonomicName>
<emphasis id="C4C11A75FFD8BD09C93535A9FC908CBA" bold="true" box="[780,861,1974,1995]" pageId="3" pageNumber="424">
<taxonomicNameLabel id="DFF2A70EFFD8BD09C93535A9FC908CBA" box="[780,861,1974,1995]" pageId="3" pageNumber="424" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult female (ICN-55713). F. Ventral view of
<taxonomicName id="31B5BDE4FFD8BD09CF7835A9FA518CBA" box="[1345,1436,1974,1995]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="424" phylum="Chordata" rank="species" species="stictus" status="sp. nov.">
<emphasis id="C4C11A75FFD8BD09CF7835A9FA838CBA" box="[1345,1358,1974,1995]" italics="true" pageId="3" pageNumber="424">P</emphasis>
.
<emphasis id="C4C11A75FFD8BD09CF6235A9FA518CBA" box="[1371,1436,1974,1995]" italics="true" pageId="3" pageNumber="424">stictus</emphasis>
</taxonomicName>
<emphasis id="C4C11A75FFD8BD09CAAE35CBFF2A8C98" bold="true" box="[151,231,2004,2025]" pageId="3" pageNumber="424">
<taxonomicNameLabel id="DFF2A70EFFD8BD09CAAE35CBFF2A8C98" box="[151,231,2004,2025]" pageId="3" pageNumber="424" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult female (ICN-55716).
</paragraph>
</caption>
<caption id="A2CA96EFFFDFBD0ECAAE3437FBB48DEB" httpUri="https://zenodo.org/record/258320/files/figure.png" pageId="4" pageNumber="425" targetBox="[151,1436,193,1552]" targetPageId="4">
<paragraph id="F60AC667FFDFBD0ECAAE3437FBB48DEB" blockId="4.[151,1436,1576,1690]" pageId="4" pageNumber="425">
<emphasis id="C4C11A75FFDFBD0ECAAE3437FEDB8D4C" bold="true" box="[151,278,1576,1598]" pageId="4" pageNumber="425">FIGURE 3.</emphasis>
Variation in life coloration. A.
<taxonomicName id="31B5BDE4FFDFBD0EC85F3436FCAF8D4F" box="[614,866,1576,1598]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="425" phylum="Chordata" rank="species" species="leptolophus">
<emphasis id="C4C11A75FFDFBD0EC85F3436FCAF8D4F" box="[614,866,1576,1598]" italics="true" pageId="4" pageNumber="425">Pristimantis leptolophus</emphasis>
</taxonomicName>
, adult female (ICN-7799) (Photo: John D. Lynch); B.
<taxonomicName id="31B5BDE4FFDFBD0ECAAE3457FE478D2D" box="[151,394,1606,1629]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="425" phylum="Chordata" rank="species" species="uranobates">
<emphasis id="C4C11A75FFDFBD0ECAAE3457FE478D2D" box="[151,394,1606,1629]" italics="true" pageId="4" pageNumber="425">Pristimantis uranobates</emphasis>
</taxonomicName>
, adult female (ICN-55588); C.
<taxonomicName id="31B5BDE4FFDFBD0EC8FE3457FC6A8D2C" box="[711,935,1608,1629]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="425" phylum="Chordata" rank="species" species="peraticus">
<emphasis id="C4C11A75FFDFBD0EC8FE3457FC6A8D2C" box="[711,935,1608,1629]" italics="true" pageId="4" pageNumber="425">Pristimantis peraticus</emphasis>
</taxonomicName>
, adult female (ICN-55742) (Photo: Marco Rada); D.
<taxonomicName id="31B5BDE4FFDFBD0ECA813479FE658D0A" box="[184,424,1637,1659]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="425" phylum="Chordata" rank="species" species="maculosus">
<emphasis id="C4C11A75FFDFBD0ECA813479FE658D0A" box="[184,424,1637,1659]" italics="true" pageId="4" pageNumber="425">Pristimantis maculosus</emphasis>
</taxonomicName>
, adult female (ICN-55578); E.
<taxonomicName id="31B5BDE4FFDFBD0EC8CA3479FC2C8D0A" box="[755,993,1638,1659]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="425" phylum="Chordata" rank="species" species="parectatus">
<emphasis id="C4C11A75FFDFBD0EC8CA3479FC2C8D0A" box="[755,993,1638,1659]" italics="true" pageId="4" pageNumber="425">Pristimantis parectatus</emphasis>
</taxonomicName>
, adult female (ICN-55743) (Photo: Marco Rada); F.
<taxonomicName id="31B5BDE4FFDFBD0ECAC0349AFDDA8DEB" box="[249,535,1668,1690]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="425" phylum="Chordata" rank="species" species="scoloblepharus">
<emphasis id="C4C11A75FFDFBD0ECAC0349AFDDA8DEB" box="[249,535,1668,1690]" italics="true" pageId="4" pageNumber="425">Pristimantis scoloblepharus</emphasis>
</taxonomicName>
, holotype adult female (ICN-8583) (Photo: John D. Lynch).
</paragraph>
</caption>
<paragraph id="F60AC667FFDFBD0ECAFE34DEFC7D8CA7" blockId="4.[151,1436,1729,2006]" pageId="4" pageNumber="425">
<emphasis id="C4C11A75FFDFBD0ECAFE34DEFD878DAB" bold="true" box="[199,586,1729,1754]" pageId="4" pageNumber="425">Generic and familial allocation.</emphasis>
Despite the absence of phenotypic synapomorphies for
<taxonomicName id="31B5BDE4FFDFBD0ECEE234DDFAA98DA8" box="[1243,1380,1730,1753]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="425" phylum="Chordata" rank="genus">
<emphasis id="C4C11A75FFDFBD0ECEE234DDFAA98DA8" box="[1243,1380,1730,1753]" italics="true" pageId="4" pageNumber="425">Pristimantis</emphasis>
</taxonomicName>
(see
<bibRefCitation id="9224BB96FFDFBD0ECAAE34FAFEA18D8F" author="Padial" box="[151,364,1765,1790]" pageId="4" pageNumber="436" refString="Padial, J. M., Grant, T. &amp; Frost, D. R. (2014 a) Molecular systematics of terraranas (Anura: Brachycephaloidea) with an assessment of the effects of alignment and optimality criteria. Zootaxa, 3825 (1), 1 - 132. http: // dx. doi. org / 10.11646 / zootaxa. 3825.1.1" type="journal article" year="2014" yearSuffix="a">
Padial
<emphasis id="C4C11A75FFDFBD0ECADD34F9FED98D8C" box="[228,276,1765,1789]" italics="true" pageId="4" pageNumber="425">et al</emphasis>
. 2014a
</bibRefCitation>
,
<bibRefCitation id="9224BB96FFDFBD0ECB4934FAFEB28D8F" author="Padial" box="[368,383,1765,1790]" pageId="4" pageNumber="436" refString="Padial, J. M., Grant, T. &amp; Frost, D. R. (2014 b) Corrections to &quot; Padial et al. (2014) Molecular systematics of terraranas (Anura: Brachycephaloidea) with an assessment of the effects of alignment and optimality criteria &quot;. Zootaxa, 3827 (4), 599 - 600. http: // dx. doi. org / 10.11646 / zootaxa. 3827.4.10" type="journal article" year="2014" yearSuffix="b">b</bibRefCitation>
), the new species is assigned to this genus based on its similarity to other species in this genus occurring in the area, as well as by characters listed by
<bibRefCitation id="9224BB96FFDFBD0EC9303516FC3F8C50" box="[777,1010,1801,1826]" pageId="4" pageNumber="436" refString="Hedges, S. B., Duellman, W. E. &amp; Heinicke, M. P. (2008) New World direct-developing frogs (Anura, Terrarana), molecular phylogeny, classification, biogeography, and conservation. Zootaxa, 1737, 1 - 182." type="journal article">
Hedges
<emphasis id="C4C11A75FFDFBD0EC95D3515FC508C50" box="[868,925,1801,1825]" italics="true" pageId="4" pageNumber="425">et al.</emphasis>
(2008)
</bibRefCitation>
, such as the absence of cranial crests, expanded terminal discs on digits, well-defined circumferential grooves with T-shaped terminal phalanges, and non-protruding subarticular tubercles. The new species is assigned to the
<taxonomicName id="31B5BDE4FFDFBD0EC9E9354DFBA28C18" box="[976,1135,1873,1897]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="425" phylum="Chordata" rank="species" species="leptolophus">
<emphasis id="C4C11A75FFDFBD0EC9E9354DFBA28C18" box="[976,1135,1873,1897]" italics="true" pageId="4" pageNumber="425">P. leptolophus</emphasis>
</taxonomicName>
group due to females with a maximum SVL of less than
<quantity id="314D6B82FFDFBD0ECBD53569FDF08CFC" box="[492,573,1910,1934]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="425" unit="mm" value="30.0">30 mm</quantity>
, narrow heads, short snout, Finger I shorter than Finger II, and Toe V much longer than Toe III and it extends to the distal edge of the distal subarticular tubercle on Toe IV. The position of the new species should be evaluated through future phylogenetic analyses.
</paragraph>
</subSubSection>
<caption id="A2CA96EFFFDEBD0FCAAE3287FB4A8BDE" ID-Table-UUID="A2CA96EFFFDEBD0FCAAE3287FB4A8BDE" box="[151,1159,151,175]" httpUri="http://table.plazi.org/id/A2CA96EFFFDEBD0FCAAE3287FB4A8BDE" pageId="5" pageNumber="426" targetBox="[151,1427,202,818]" targetIsTable="true" targetPageId="5">
<paragraph id="F60AC667FFDEBD0FCAAE3287FB4A8BDE" blockId="5.[151,1159,151,175]" box="[151,1159,151,175]" pageId="5" pageNumber="426">
<emphasis id="C4C11A75FFDEBD0FCAAE3287FEC78BDE" bold="true" box="[151,266,152,175]" pageId="5" pageNumber="426">TABLE 1.</emphasis>
Diagnostic morphological characters in the
<taxonomicName id="31B5BDE4FFDEBD0FC8DB3287FC2A8BDE" box="[738,999,151,175]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="426" phylum="Chordata" rank="species" species="leptolophus">
<emphasis id="C4C11A75FFDEBD0FC8DB3287FC2A8BDE" box="[738,999,151,175]" italics="true" pageId="5" pageNumber="426">Pristimantis leptolophus</emphasis>
</taxonomicName>
species group.
</paragraph>
</caption>
<paragraph id="F60AC667FFDEBD0FCBBD32D5FEC58843" pageId="5" pageNumber="426">
<table id="84B534C7FFDE42F5CAAE32D5FA5E8843" box="[151,1427,202,818]" gridcols="6" gridrows="9" pageId="5" pageNumber="426">
<tr id="4885C425FFDE42F5CAAE32D5FA5E8B8E" box="[151,1427,202,255]" gridrow="0" pageId="5" pageNumber="426" rowspan-0="1">
<th id="0B54AD59FFDE42F5CBBD32D5FE388B8E" box="[388,501,202,255]" gridcol="1" gridrow="0" pageId="5" pageNumber="426">SVL males</th>
<th id="0B54AD59FFDE42F5C82432D5FD6F8B8E" box="[541,674,202,255]" gridcol="2" gridrow="0" pageId="5" pageNumber="426">SVL females</th>
<th id="0B54AD59FFDE42F5C88E32D5FC678B8E" box="[695,938,202,255]" gridcol="3" gridrow="0" pageId="5" pageNumber="426">Dorsal coloration in life</th>
<th id="0B54AD59FFDE42F5C9F332D5FB408B8E" box="[970,1165,202,255]" gridcol="4" gridrow="0" pageId="5" pageNumber="426">Groin coloration in life</th>
<th id="0B54AD59FFDE42F5CE9C32D5FA5E8B8E" box="[1189,1427,202,255]" gridcol="5" gridrow="0" pageId="5" pageNumber="426">Dorsolaterals folds</th>
</tr>
<tr id="4885C425FFDE42F5CAAE330DFA5E8A59" box="[151,1427,274,296]" gridrow="1" pageId="5" pageNumber="426">
<th id="0B54AD59FFDE42F5CAAE330DFE9E8A59" box="[151,339,274,296]" gridcol="0" gridrow="1" pageId="5" pageNumber="426">
<taxonomicName id="31B5BDE4FFDEBD0FCAA6330CFEFC8A59" box="[159,305,274,296]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="426" phylum="Chordata" rank="species" species="lasalleorum">
<emphasis id="C4C11A75FFDEBD0FCAA6330CFF618A59" box="[159,172,275,296]" italics="true" pageId="5" pageNumber="426">P</emphasis>
.
<emphasis id="C4C11A75FFDEBD0FCA81330DFEFC8A59" box="[184,305,274,296]" italics="true" pageId="5" pageNumber="426">lasalleorum</emphasis>
</taxonomicName>
</th>
<td id="0B54AD59FFDE42F5CBBD330DFE388A59" box="[388,501,274,296]" gridcol="1" gridrow="1" pageId="5" pageNumber="426">15.6</td>
<td id="0B54AD59FFDE42F5C824330DFD6F8A59" box="[541,674,274,296]" gridcol="2" gridrow="1" pageId="5" pageNumber="426">18.4–21,0</td>
<td id="0B54AD59FFDE42F5C88E330DFC678A59" box="[695,938,274,296]" gridcol="3" gridrow="1" pageId="5" pageNumber="426">Dark brown above</td>
<td id="0B54AD59FFDE42F5C9F3330DFB408A59" box="[970,1165,274,296]" gridcol="4" gridrow="1" pageId="5" pageNumber="426">Brown</td>
<td id="0B54AD59FFDE42F5CE9C330DFA5E8A59" box="[1189,1427,274,296]" gridcol="5" gridrow="1" pageId="5" pageNumber="426">Low</td>
</tr>
<tr id="4885C425FFDE42F5CAAE3323FA5E8A01" box="[151,1427,316,368]" gridrow="2" pageId="5" pageNumber="426">
<th id="0B54AD59FFDE42F5CAAE3323FE9E8A01" box="[151,339,316,368]" gridcol="0" gridrow="2" pageId="5" pageNumber="426">
<taxonomicName id="31B5BDE4FFDEBD0FCAA63322FEE38A23" box="[159,302,316,338]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="426" phylum="Chordata" rank="species" species="leptolophus">
<emphasis id="C4C11A75FFDEBD0FCAA63322FF618A23" box="[159,172,317,338]" italics="true" pageId="5" pageNumber="426">P</emphasis>
.
<emphasis id="C4C11A75FFDEBD0FCA813323FEE38A23" box="[184,302,316,338]" italics="true" pageId="5" pageNumber="426">leptolophus</emphasis>
</taxonomicName>
</th>
<td id="0B54AD59FFDE42F5CBBD3323FE388A01" box="[388,501,316,368]" gridcol="1" gridrow="2" pageId="5" pageNumber="426">14.3–18.2</td>
<td id="0B54AD59FFDE42F5C8243323FD6F8A01" box="[541,674,316,368]" gridcol="2" gridrow="2" pageId="5" pageNumber="426">21.2–25.1</td>
<td id="0B54AD59FFDE42F5C88E3323FC678A01" box="[695,938,316,368]" gridcol="3" gridrow="2" pageId="5" pageNumber="426">Brown with darker markings</td>
<td id="0B54AD59FFDE42F5C9F33323FB408A01" box="[970,1165,316,368]" gridcol="4" gridrow="2" pageId="5" pageNumber="426">Brown</td>
<td id="0B54AD59FFDE42F5CE9C3323FA5E8A01" box="[1189,1427,316,368]" gridcol="5" gridrow="2" pageId="5" pageNumber="426">Low and thin</td>
</tr>
<tr id="4885C425FFDE42F5CAAE339BFA5E8AA6" box="[151,1427,388,471]" gridrow="3" pageId="5" pageNumber="426">
<th id="0B54AD59FFDE42F5CAAE339BFE9E8AA6" box="[151,339,388,471]" gridcol="0" gridrow="3" pageId="5" pageNumber="426">
<taxonomicName id="31B5BDE4FFDEBD0FCAA6339AFEEE8AEB" box="[159,291,388,410]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="426" phylum="Chordata" rank="species" species="maculosus">
<emphasis id="C4C11A75FFDEBD0FCAA6339AFF618AEB" box="[159,172,389,410]" italics="true" pageId="5" pageNumber="426">P</emphasis>
.
<emphasis id="C4C11A75FFDEBD0FCA81339BFEEE8AEB" box="[184,291,388,410]" italics="true" pageId="5" pageNumber="426">maculosus</emphasis>
</taxonomicName>
</th>
<td id="0B54AD59FFDE42F5CBBD339BFE388AA6" box="[388,501,388,471]" gridcol="1" gridrow="3" pageId="5" pageNumber="426">17.8–19.2</td>
<td id="0B54AD59FFDE42F5C824339BFD6F8AA6" box="[541,674,388,471]" gridcol="2" gridrow="3" pageId="5" pageNumber="426">23.8–26.3</td>
<td id="0B54AD59FFDE42F5C88E339BFC678AA6" box="[695,938,388,471]" gridcol="3" gridrow="3" pageId="5" pageNumber="426">Dark brown</td>
<td id="0B54AD59FFDE42F5C9F3339BFB408AA6" box="[970,1165,388,471]" gridcol="4" gridrow="3" pageId="5" pageNumber="426">Black with large white spots</td>
<td id="0B54AD59FFDE42F5CE9C339BFA5E8AA6" box="[1189,1427,388,471]" gridcol="5" gridrow="3" pageId="5" pageNumber="426">Inconspicuous, reach anterior to level of sacrum</td>
</tr>
<tr id="4885C425FFDE42F5CAAE33F5FA5E896E" box="[151,1427,490,543]" gridrow="4" pageId="5" pageNumber="426">
<th id="0B54AD59FFDE42F5CAAE33F5FE9E896E" box="[151,339,490,543]" gridcol="0" gridrow="4" pageId="5" pageNumber="426">
<taxonomicName id="31B5BDE4FFDEBD0FCAA633F4FEEF8971" box="[159,290,491,512]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="426" phylum="Chordata" rank="species" species="parectatus">
<emphasis id="C4C11A75FFDEBD0FCAA633F4FF618971" box="[159,172,491,512]" italics="true" pageId="5" pageNumber="426">P</emphasis>
.
<emphasis id="C4C11A75FFDEBD0FCA8133F4FEEF8971" box="[184,290,491,512]" italics="true" pageId="5" pageNumber="426">parectatus</emphasis>
</taxonomicName>
</th>
<td id="0B54AD59FFDE42F5CBBD33F5FE38896E" box="[388,501,490,543]" gridcol="1" gridrow="4" pageId="5" pageNumber="426">14.2–16.0</td>
<td id="0B54AD59FFDE42F5C82433F5FD6F896E" box="[541,674,490,543]" gridcol="2" gridrow="4" pageId="5" pageNumber="426">22.5–27.0</td>
<td id="0B54AD59FFDE42F5C88E33F5FC67896E" box="[695,938,490,543]" gridcol="3" gridrow="4" pageId="5" pageNumber="426">Brown with darker markings</td>
<td id="0B54AD59FFDE42F5C9F333F5FB40896E" box="[970,1165,490,543]" gridcol="4" gridrow="4" pageId="5" pageNumber="426">Pale brown or cream</td>
<td id="0B54AD59FFDE42F5CE9C33F5FA5E896E" box="[1189,1427,490,543]" gridcol="5" gridrow="4" pageId="5" pageNumber="426">Low and thin</td>
</tr>
<tr id="4885C425FFDE42F5CAAE302DFA5E8939" box="[151,1427,562,584]" gridrow="5" pageId="5" pageNumber="426">
<th id="0B54AD59FFDE42F5CAAE302DFE9E8939" box="[151,339,562,584]" gridcol="0" gridrow="5" pageId="5" pageNumber="426">
<taxonomicName id="31B5BDE4FFDEBD0FCAA6302CFEDB8939" box="[159,278,563,584]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="426" phylum="Chordata" rank="species" species="peraticus">
<emphasis id="C4C11A75FFDEBD0FCAA6302CFF618939" box="[159,172,563,584]" italics="true" pageId="5" pageNumber="426">P</emphasis>
.
<emphasis id="C4C11A75FFDEBD0FCA81302CFEDB8939" box="[184,278,563,584]" italics="true" pageId="5" pageNumber="426">peraticus</emphasis>
</taxonomicName>
</th>
<td id="0B54AD59FFDE42F5CBBD302DFE388939" box="[388,501,562,584]" gridcol="1" gridrow="5" pageId="5" pageNumber="426">15.3–18.3</td>
<td id="0B54AD59FFDE42F5C824302DFD6F8939" box="[541,674,562,584]" gridcol="2" gridrow="5" pageId="5" pageNumber="426">20.7–26.0</td>
<td id="0B54AD59FFDE42F5C88E302DFC678939" box="[695,938,562,584]" gridcol="3" gridrow="5" pageId="5" pageNumber="426">Pale Brown</td>
<td id="0B54AD59FFDE42F5C9F3302DFB408939" box="[970,1165,562,584]" gridcol="4" gridrow="5" pageId="5" pageNumber="426">Brown</td>
<td id="0B54AD59FFDE42F5CE9C302DFA5E8939" box="[1189,1427,562,584]" gridcol="5" gridrow="5" pageId="5" pageNumber="426">Low and thin</td>
</tr>
<tr id="4885C425FFDE42F5CAAE3043FA5E89DE" box="[151,1427,604,687]" gridrow="6" pageId="5" pageNumber="426">
<th id="0B54AD59FFDE42F5CAAE3043FE9E89DE" box="[151,339,604,687]" gridcol="0" gridrow="6" pageId="5" pageNumber="426">
<taxonomicName id="31B5BDE4FFDEBD0FCAA63042FE9E8903" box="[159,339,604,626]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="426" phylum="Chordata" rank="species" species="scoloblepharus">
<emphasis id="C4C11A75FFDEBD0FCAA63042FF618903" box="[159,172,605,626]" italics="true" pageId="5" pageNumber="426">P</emphasis>
.
<emphasis id="C4C11A75FFDEBD0FCA813043FE9E8903" box="[184,339,604,626]" italics="true" pageId="5" pageNumber="426">scoloblepharus</emphasis>
</taxonomicName>
</th>
<td id="0B54AD59FFDE42F5CBBD3043FE3889DE" box="[388,501,604,687]" gridcol="1" gridrow="6" pageId="5" pageNumber="426">17.4–20,3</td>
<td id="0B54AD59FFDE42F5C8243043FD6F89DE" box="[541,674,604,687]" gridcol="2" gridrow="6" pageId="5" pageNumber="426">23.7</td>
<td id="0B54AD59FFDE42F5C88E3043FC6789DE" box="[695,938,604,687]" gridcol="3" gridrow="6" pageId="5" pageNumber="426">Brown above with dark brown mottling, flanks paler</td>
<td id="0B54AD59FFDE42F5C9F33043FB4089DE" box="[970,1165,604,687]" gridcol="4" gridrow="6" pageId="5" pageNumber="426">Yellow spots</td>
<td id="0B54AD59FFDE42F5CE9C3043FA5E89DE" box="[1189,1427,604,687]" gridcol="5" gridrow="6" pageId="5" pageNumber="426">Thickened with conical warts</td>
</tr>
<tr id="4885C425FFDE42F5CAAE30DDFA5E8986" box="[151,1427,706,759]" gridrow="7" pageId="5" pageNumber="426">
<th id="0B54AD59FFDE42F5CAAE30DDFE9E8986" box="[151,339,706,759]" gridcol="0" gridrow="7" pageId="5" pageNumber="426">
<taxonomicName id="31B5BDE4FFDEBD0FCAA630DCFEE789A9" box="[159,298,706,728]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="426" phylum="Chordata" rank="species" species="uranobates">
<emphasis id="C4C11A75FFDEBD0FCAA630DCFF6189A9" box="[159,172,707,728]" italics="true" pageId="5" pageNumber="426">P</emphasis>
.
<emphasis id="C4C11A75FFDEBD0FCA8130DDFEE789A9" box="[184,298,706,728]" italics="true" pageId="5" pageNumber="426">uranobates</emphasis>
</taxonomicName>
</th>
<td id="0B54AD59FFDE42F5CBBD30DDFE388986" box="[388,501,706,759]" gridcol="1" gridrow="7" pageId="5" pageNumber="426">15.7–19.8</td>
<td id="0B54AD59FFDE42F5C82430DDFD6F8986" box="[541,674,706,759]" gridcol="2" gridrow="7" pageId="5" pageNumber="426">23.1–27.8</td>
<td id="0B54AD59FFDE42F5C88E30DDFC678986" box="[695,938,706,759]" gridcol="3" gridrow="7" pageId="5" pageNumber="426">Brown with markings (polymorphic pattern)</td>
<td id="0B54AD59FFDE42F5C9F330DDFB408986" box="[970,1165,706,759]" gridcol="4" gridrow="7" pageId="5" pageNumber="426">Brown spots and reticulation</td>
<td id="0B54AD59FFDE42F5CE9C30DDFA5E8986" box="[1189,1427,706,759]" gridcol="5" gridrow="7" pageId="5" pageNumber="426">Thickened, polymorphic coloration</td>
</tr>
<tr id="4885C425FFDE42F5CAAE3104FA5E8843" box="[151,1427,795,818]" gridrow="8" pageId="5" pageNumber="426" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="0B54AD59FFDE42F5CAAE3104FE9E8843" box="[151,339,795,818]" gridcol="0" gridrow="8" pageId="5" pageNumber="426">continued.</th>
</tr>
</table>
</paragraph>
<paragraph id="F60AC667FFDEBD0FCBBB3151FEC58E4A" pageId="5" pageNumber="426">
<table id="84B534C7FFDE42F5CAAE3151FA408E4A" box="[151,1421,846,1339]" gridcols="7" gridrows="9" pageId="5" pageNumber="426">
<tr id="4885C425FFDE42F5CAAE3151FA4088F3" box="[151,1421,846,898]" gridrow="0" pageId="5" pageNumber="426" rowspan-0="1">
<th id="0B54AD59FFDE42F5CBBB3151FDDB88F3" box="[386,534,846,898]" gridcol="1" gridrow="0" pageId="5" pageNumber="426">Lateralfringes on finger</th>
<th id="0B54AD59FFDE42F5C8093151FD7C88F3" box="[560,689,846,898]" gridcol="2" gridrow="0" pageId="5" pageNumber="426">Nuptial pads on thumb</th>
<th id="0B54AD59FFDE42F5C8EA3151FC8E88F3" box="[723,835,846,898]" gridcol="3" gridrow="0" pageId="5" pageNumber="426">Ulnar tubercles</th>
<th id="0B54AD59FFDE42F5C9563151FC2488F3" box="[879,1001,846,898]" gridcol="4" gridrow="0" pageId="5" pageNumber="426">Tarsal tubercles</th>
<th id="0B54AD59FFDE42F5CE2A3151FB0788F3" box="[1043,1226,846,898]" gridcol="5" gridrow="0" pageId="5" pageNumber="426">Heel tubercle</th>
<th id="0B54AD59FFDE42F5CEDA3151FA4088F3" box="[1251,1421,846,898]" gridcol="6" gridrow="0" pageId="5" pageNumber="426">Tubercles on upper eyelid</th>
</tr>
<tr id="4885C425FFDE42F5CAAE3189FA4088BB" box="[151,1421,918,970]" gridrow="1" pageId="5" pageNumber="426">
<th id="0B54AD59FFDE42F5CAAE3189FE9E88BB" box="[151,339,918,970]" gridcol="0" gridrow="1" pageId="5" pageNumber="426">
<taxonomicName id="31B5BDE4FFDEBD0FCAA63188FEFC88DD" authority="Present Large Subconical Small Present, Conical" authorityName="Present Large Subconical Small Present, Conical" box="[159,305,918,940]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="426" phylum="Chordata" rank="species" species="lasalleorum">
<emphasis id="C4C11A75FFDEBD0FCAA63188FF6188DD" box="[159,172,919,940]" italics="true" pageId="5" pageNumber="426">P</emphasis>
.
<emphasis id="C4C11A75FFDEBD0FCA813189FEFC88DD" box="[184,305,918,940]" italics="true" pageId="5" pageNumber="426">lasalleorum</emphasis>
</taxonomicName>
</th>
<td id="0B54AD59FFDE42F5CBBB3189FDDB88BB" box="[386,534,918,970]" gridcol="1" gridrow="1" pageId="5" pageNumber="426">Present</td>
<td id="0B54AD59FFDE42F5C8093189FD7C88BB" box="[560,689,918,970]" gridcol="2" gridrow="1" pageId="5" pageNumber="426">Large</td>
<td id="0B54AD59FFDE42F5C8EA3189FC8E88BB" box="[723,835,918,970]" gridcol="3" gridrow="1" pageId="5" pageNumber="426">Subconical</td>
<td id="0B54AD59FFDE42F5C9563189FC2488BB" box="[879,1001,918,970]" gridcol="4" gridrow="1" pageId="5" pageNumber="426">Small</td>
<td id="0B54AD59FFDE42F5CE2A3189FB0788BB" box="[1043,1226,918,970]" gridcol="5" gridrow="1" pageId="5" pageNumber="426">Present, subconicals</td>
<td id="0B54AD59FFDE42F5CEDA3189FA4088BB" box="[1251,1421,918,970]" gridcol="6" gridrow="1" pageId="5" pageNumber="426">Conical</td>
</tr>
<tr id="4885C425FFDE42F5CAAE31C1FA408885" box="[151,1421,990,1012]" gridrow="2" pageId="5" pageNumber="426">
<th id="0B54AD59FFDE42F5CAAE31C1FE9E8885" box="[151,339,990,1012]" gridcol="0" gridrow="2" pageId="5" pageNumber="426">
<taxonomicName id="31B5BDE4FFDEBD0FCAA631C0FEE38885" box="[159,302,990,1012]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="426" phylum="Chordata" rank="species" species="leptolophus">
<emphasis id="C4C11A75FFDEBD0FCAA631C0FF618885" box="[159,172,991,1012]" italics="true" pageId="5" pageNumber="426">P</emphasis>
.
<emphasis id="C4C11A75FFDEBD0FCA8131C1FEE38885" box="[184,302,990,1012]" italics="true" pageId="5" pageNumber="426">leptolophus</emphasis>
</taxonomicName>
</th>
<td id="0B54AD59FFDE42F5CBBB31C1FDDB8885" box="[386,534,990,1012]" gridcol="1" gridrow="2" pageId="5" pageNumber="426">Narrow</td>
<td id="0B54AD59FFDE42F5C80931C1FD7C8885" box="[560,689,990,1012]" gridcol="2" gridrow="2" pageId="5" pageNumber="426">Absent</td>
<td id="0B54AD59FFDE42F5C8EA31C1FC8E8885" box="[723,835,990,1012]" gridcol="3" gridrow="2" pageId="5" pageNumber="426">Elongate</td>
<td id="0B54AD59FFDE42F5C95631C1FC248885" box="[879,1001,990,1012]" gridcol="4" gridrow="2" pageId="5" pageNumber="426">Conical</td>
<td id="0B54AD59FFDE42F5CE2A31C1FB078885" box="[1043,1226,990,1012]" gridcol="5" gridrow="2" pageId="5" pageNumber="426">Conical</td>
<td id="0B54AD59FFDE42F5CEDA31C1FA408885" box="[1251,1421,990,1012]" gridcol="6" gridrow="2" pageId="5" pageNumber="426">Present, small</td>
</tr>
<tr id="4885C425FFDE42F5CAAE3618FA408F6C" box="[151,1421,1031,1053]" gridrow="3" pageId="5" pageNumber="426">
<th id="0B54AD59FFDE42F5CAAE3618FE9E8F6C" box="[151,339,1031,1053]" gridcol="0" gridrow="3" pageId="5" pageNumber="426">
<taxonomicName id="31B5BDE4FFDEBD0FCAA63617FEEE8F6C" box="[159,291,1031,1053]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="426" phylum="Chordata" rank="species" species="maculosus">
<emphasis id="C4C11A75FFDEBD0FCAA63617FF618F6C" box="[159,172,1032,1053]" italics="true" pageId="5" pageNumber="426">P</emphasis>
.
<emphasis id="C4C11A75FFDEBD0FCA813618FEEE8F6C" box="[184,291,1031,1053]" italics="true" pageId="5" pageNumber="426">maculosus</emphasis>
</taxonomicName>
</th>
<td id="0B54AD59FFDE42F5CBBB3618FDDB8F6C" box="[386,534,1031,1053]" gridcol="1" gridrow="3" pageId="5" pageNumber="426">Lateral keels</td>
<td id="0B54AD59FFDE42F5C8093618FD7C8F6C" box="[560,689,1031,1053]" gridcol="2" gridrow="3" pageId="5" pageNumber="426">Present</td>
<td id="0B54AD59FFDE42F5C8EA3618FC8E8F6C" box="[723,835,1031,1053]" gridcol="3" gridrow="3" pageId="5" pageNumber="426">Small</td>
<td id="0B54AD59FFDE42F5C9563618FC248F6C" box="[879,1001,1031,1053]" gridcol="4" gridrow="3" pageId="5" pageNumber="426">Small</td>
<td id="0B54AD59FFDE42F5CE2A3618FB078F6C" box="[1043,1226,1031,1053]" gridcol="5" gridrow="3" pageId="5" pageNumber="426">Present, small</td>
<td id="0B54AD59FFDE42F5CEDA3618FA408F6C" box="[1251,1421,1031,1053]" gridcol="6" gridrow="3" pageId="5" pageNumber="426">Small warts</td>
</tr>
<tr id="4885C425FFDE42F5CAAE362EFA408F14" box="[151,1421,1073,1125]" gridrow="4" pageId="5" pageNumber="426">
<th id="0B54AD59FFDE42F5CAAE362EFE9E8F14" box="[151,339,1073,1125]" gridcol="0" gridrow="4" pageId="5" pageNumber="426">
<taxonomicName id="31B5BDE4FFDEBD0FCAA6362DFEEF8F36" box="[159,290,1074,1095]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="426" phylum="Chordata" rank="species" species="parectatus">
<emphasis id="C4C11A75FFDEBD0FCAA6362DFF618F36" box="[159,172,1074,1095]" italics="true" pageId="5" pageNumber="426">P</emphasis>
.
<emphasis id="C4C11A75FFDEBD0FCA81362DFEEF8F36" box="[184,290,1074,1095]" italics="true" pageId="5" pageNumber="426">parectatus</emphasis>
</taxonomicName>
</th>
<td id="0B54AD59FFDE42F5CBBB362EFDDB8F14" box="[386,534,1073,1125]" gridcol="1" gridrow="4" pageId="5" pageNumber="426">Absent</td>
<td id="0B54AD59FFDE42F5C809362EFD7C8F14" box="[560,689,1073,1125]" gridcol="2" gridrow="4" pageId="5" pageNumber="426">Absent</td>
<td id="0B54AD59FFDE42F5C8EA362EFC8E8F14" box="[723,835,1073,1125]" gridcol="3" gridrow="4" pageId="5" pageNumber="426">Subconical</td>
<td id="0B54AD59FFDE42F5C956362EFC248F14" box="[879,1001,1073,1125]" gridcol="4" gridrow="4" pageId="5" pageNumber="426">Subconicals</td>
<td id="0B54AD59FFDE42F5CE2A362EFB078F14" box="[1043,1226,1073,1125]" gridcol="5" gridrow="4" pageId="5" pageNumber="426">Present, subconicals</td>
<td id="0B54AD59FFDE42F5CEDA362EFA408F14" box="[1251,1421,1073,1125]" gridcol="6" gridrow="4" pageId="5" pageNumber="426">Short conical</td>
</tr>
<tr id="4885C425FFDE42F5CAAE3666FA408FFE" box="[151,1421,1145,1167]" gridrow="5" pageId="5" pageNumber="426">
<th id="0B54AD59FFDE42F5CAAE3666FE9E8FFE" box="[151,339,1145,1167]" gridcol="0" gridrow="5" pageId="5" pageNumber="426">
<taxonomicName id="31B5BDE4FFDEBD0FCAA63665FEDB8FFE" box="[159,278,1146,1167]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="426" phylum="Chordata" rank="species" species="peraticus">
<emphasis id="C4C11A75FFDEBD0FCAA63665FF618FFE" box="[159,172,1146,1167]" italics="true" pageId="5" pageNumber="426">P</emphasis>
.
<emphasis id="C4C11A75FFDEBD0FCA813665FEDB8FFE" box="[184,278,1146,1167]" italics="true" pageId="5" pageNumber="426">peraticus</emphasis>
</taxonomicName>
</th>
<td id="0B54AD59FFDE42F5CBBB3666FDDB8FFE" box="[386,534,1145,1167]" gridcol="1" gridrow="5" pageId="5" pageNumber="426">Narrow</td>
<td id="0B54AD59FFDE42F5C8093666FD7C8FFE" box="[560,689,1145,1167]" gridcol="2" gridrow="5" pageId="5" pageNumber="426">Absent</td>
<td id="0B54AD59FFDE42F5C8EA3666FC8E8FFE" box="[723,835,1145,1167]" gridcol="3" gridrow="5" pageId="5" pageNumber="426">Elongate</td>
<td id="0B54AD59FFDE42F5C9563666FC248FFE" box="[879,1001,1145,1167]" gridcol="4" gridrow="5" pageId="5" pageNumber="426">Small</td>
<td id="0B54AD59FFDE42F5CE2A3666FB078FFE" box="[1043,1226,1145,1167]" gridcol="5" gridrow="5" pageId="5" pageNumber="426">Small subconicals</td>
<td id="0B54AD59FFDE42F5CEDA3666FA408FFE" box="[1251,1421,1145,1167]" gridcol="6" gridrow="5" pageId="5" pageNumber="426">Present, small</td>
</tr>
<tr id="4885C425FFDE42F5CAAE36BDFA408FC9" box="[151,1421,1186,1208]" gridrow="6" pageId="5" pageNumber="426">
<th id="0B54AD59FFDE42F5CAAE36BDFE9E8FC9" box="[151,339,1186,1208]" gridcol="0" gridrow="6" pageId="5" pageNumber="426">
<taxonomicName id="31B5BDE4FFDEBD0FCAA636BCFE9E8FC9" box="[159,339,1186,1208]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="426" phylum="Chordata" rank="species" species="scoloblepharus">
<emphasis id="C4C11A75FFDEBD0FCAA636BCFF618FC9" box="[159,172,1187,1208]" italics="true" pageId="5" pageNumber="426">P</emphasis>
.
<emphasis id="C4C11A75FFDEBD0FCA8136BDFE9E8FC9" box="[184,339,1186,1208]" italics="true" pageId="5" pageNumber="426">scoloblepharus</emphasis>
</taxonomicName>
</th>
<td id="0B54AD59FFDE42F5CBBB36BDFDDB8FC9" box="[386,534,1186,1208]" gridcol="1" gridrow="6" pageId="5" pageNumber="426">Narrow</td>
<td id="0B54AD59FFDE42F5C80936BDFD7C8FC9" box="[560,689,1186,1208]" gridcol="2" gridrow="6" pageId="5" pageNumber="426">Present</td>
<td id="0B54AD59FFDE42F5C8EA36BDFC8E8FC9" box="[723,835,1186,1208]" gridcol="3" gridrow="6" pageId="5" pageNumber="426">Present</td>
<td id="0B54AD59FFDE42F5C95636BDFC248FC9" box="[879,1001,1186,1208]" gridcol="4" gridrow="6" pageId="5" pageNumber="426">Small</td>
<td id="0B54AD59FFDE42F5CE2A36BDFB078FC9" box="[1043,1226,1186,1208]" gridcol="5" gridrow="6" pageId="5" pageNumber="426">Conicals</td>
<td id="0B54AD59FFDE42F5CEDA36BDFA408FC9" box="[1251,1421,1186,1208]" gridcol="6" gridrow="6" pageId="5" pageNumber="426">Present, conicals</td>
</tr>
<tr id="4885C425FFDE42F5CAAE36D4FA408E71" box="[151,1421,1227,1280]" gridrow="7" pageId="5" pageNumber="426">
<th id="0B54AD59FFDE42F5CAAE36D4FE9E8E71" box="[151,339,1227,1280]" gridcol="0" gridrow="7" pageId="5" pageNumber="426">
<taxonomicName id="31B5BDE4FFDEBD0FCAA636D3FEE78F90" box="[159,298,1227,1249]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="426" phylum="Chordata" rank="species" species="uranobates">
<emphasis id="C4C11A75FFDEBD0FCAA636D3FF618F90" box="[159,172,1228,1249]" italics="true" pageId="5" pageNumber="426">P</emphasis>
.
<emphasis id="C4C11A75FFDEBD0FCA8136D4FEE78F90" box="[184,298,1227,1249]" italics="true" pageId="5" pageNumber="426">uranobates</emphasis>
</taxonomicName>
</th>
<td id="0B54AD59FFDE42F5CBBB36D4FDDB8E71" box="[386,534,1227,1280]" gridcol="1" gridrow="7" pageId="5" pageNumber="426">Narrow</td>
<td id="0B54AD59FFDE42F5C80936D4FD7C8E71" box="[560,689,1227,1280]" gridcol="2" gridrow="7" pageId="5" pageNumber="426">Present</td>
<td id="0B54AD59FFDE42F5C8EA36D4FC8E8E71" box="[723,835,1227,1280]" gridcol="3" gridrow="7" pageId="5" pageNumber="426">Elongate</td>
<td id="0B54AD59FFDE42F5C95636D4FC248E71" box="[879,1001,1227,1280]" gridcol="4" gridrow="7" pageId="5" pageNumber="426">Subconicals</td>
<td id="0B54AD59FFDE42F5CE2A36D4FB078E71" box="[1043,1226,1227,1280]" gridcol="5" gridrow="7" pageId="5" pageNumber="426">Present, subconicals</td>
<td id="0B54AD59FFDE42F5CEDA36D4FA408E71" box="[1251,1421,1227,1280]" gridcol="6" gridrow="7" pageId="5" pageNumber="426">Present, subconicals</td>
</tr>
<tr id="4885C425FFDE42F5CAAE373BFA408E4A" box="[151,1421,1316,1339]" gridrow="8" pageId="5" pageNumber="426" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="0B54AD59FFDE42F5CAAE373BFE9E8E4A" box="[151,339,1316,1339]" gridcol="0" gridrow="8" pageId="5" pageNumber="426">continued.</th>
</tr>
</table>
</paragraph>
<table id="84B534C7FFDE42F5CAA63748FA5C8C14" box="[159,1425,1367,1893]" gridcols="6" gridrows="8" pageId="5" pageNumber="426">
<paragraph id="F60AC667FFDEBD0FCBBA3748FA478C14" pageId="5" pageNumber="426">
<tr id="4885C425FFDE42F5CAA63748FA5C8EFA" box="[159,1425,1367,1419]" gridrow="0" pageId="5" pageNumber="426" rowspan-0="1">
<th id="0B54AD59FFDE42F5CBBA3748FD7C8EFA" box="[387,689,1367,1419]" gridcol="1" gridrow="0" pageId="5" pageNumber="426">Snout</th>
<th id="0B54AD59FFDE42F5C8E43748FC9D8EFA" box="[733,848,1367,1419]" gridcol="2" gridrow="0" pageId="5" pageNumber="426">Tympanum</th>
<th id="0B54AD59FFDE42F5C9423748FC0B8EFA" box="[891,966,1367,1419]" gridcol="3" gridrow="0" pageId="5" pageNumber="426">Cranial crests</th>
<th id="0B54AD59FFDE42F5CE393748FB478EFA" box="[1024,1162,1367,1419]" gridcol="4" gridrow="0" pageId="5" pageNumber="426">Vomerine odontophores</th>
<th id="0B54AD59FFDE42F5CE993748FA5C8EFA" box="[1184,1425,1367,1419]" gridcol="5" gridrow="0" pageId="5" pageNumber="426">Source</th>
</tr>
<tr id="4885C425FFDE42F5CAA63780FA5C8EA2" box="[159,1425,1439,1491]" gridrow="1" pageId="5" pageNumber="426">
<th id="0B54AD59FFDE42F5CAA63780FE9E8EA2" box="[159,339,1439,1491]" gridcol="0" gridrow="1" pageId="5" pageNumber="426">
<taxonomicName id="31B5BDE4FFDEBD0FCAA637BFFEFC8EC4" box="[159,305,1439,1461]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="426" phylum="Chordata" rank="species" species="lasalleorum">
<emphasis id="C4C11A75FFDEBD0FCAA637BFFF618EC4" box="[159,172,1440,1461]" italics="true" pageId="5" pageNumber="426">P</emphasis>
.
<emphasis id="C4C11A75FFDEBD0FCA813780FEFC8EC4" box="[184,305,1439,1461]" italics="true" pageId="5" pageNumber="426">lasalleorum</emphasis>
</taxonomicName>
</th>
<td id="0B54AD59FFDE42F5CBBA3780FD7C8EA2" box="[387,689,1439,1491]" gridcol="1" gridrow="1" pageId="5" pageNumber="426">Short, rounded in dorsal and lateral view</td>
<td id="0B54AD59FFDE42F5C8E43780FC9D8EA2" box="[733,848,1439,1491]" gridcol="2" gridrow="1" pageId="5" pageNumber="426">Present</td>
<td id="0B54AD59FFDE42F5C9423780FC0B8EA2" box="[891,966,1439,1491]" gridcol="3" gridrow="1" pageId="5" pageNumber="426">Absent</td>
<td id="0B54AD59FFDE42F5CE393780FB478EA2" box="[1024,1162,1439,1491]" gridcol="4" gridrow="1" pageId="5" pageNumber="426">Absent</td>
<td id="0B54AD59FFDE42F5CE993780FA5C8EA2" box="[1184,1425,1439,1491]" gridcol="5" gridrow="1" pageId="5" pageNumber="426">Lynch, 1995, this work.</td>
</tr>
<tr id="4885C425FFDE42F5CAA637F8FA5C8D6A" box="[159,1425,1511,1563]" gridrow="2" pageId="5" pageNumber="426">
<th id="0B54AD59FFDE42F5CAA637F8FE9E8D6A" box="[159,339,1511,1563]" gridcol="0" gridrow="2" pageId="5" pageNumber="426">
<taxonomicName id="31B5BDE4FFDEBD0FCAA637F7FEE38E8C" box="[159,302,1511,1533]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="426" phylum="Chordata" rank="species" species="leptolophus">
<emphasis id="C4C11A75FFDEBD0FCAA637F7FF618E8C" box="[159,172,1512,1533]" italics="true" pageId="5" pageNumber="426">P</emphasis>
.
<emphasis id="C4C11A75FFDEBD0FCA8137F8FEE38E8C" box="[184,302,1511,1533]" italics="true" pageId="5" pageNumber="426">leptolophus</emphasis>
</taxonomicName>
</th>
<td id="0B54AD59FFDE42F5CBBA37F8FD7C8D6A" box="[387,689,1511,1563]" gridcol="1" gridrow="2" pageId="5" pageNumber="426">Short, rounded in dorsal and lateral view</td>
<td id="0B54AD59FFDE42F5C8E437F8FC9D8D6A" box="[733,848,1511,1563]" gridcol="2" gridrow="2" pageId="5" pageNumber="426">Present</td>
<td id="0B54AD59FFDE42F5C94237F8FC0B8D6A" box="[891,966,1511,1563]" gridcol="3" gridrow="2" pageId="5" pageNumber="426">Absent</td>
<td id="0B54AD59FFDE42F5CE3937F8FB478D6A" box="[1024,1162,1511,1563]" gridcol="4" gridrow="2" pageId="5" pageNumber="426">Absent</td>
<td id="0B54AD59FFDE42F5CE9937F8FA5C8D6A" box="[1184,1425,1511,1563]" gridcol="5" gridrow="2" pageId="5" pageNumber="426">Lynch, 1980, Lynch, 1991.</td>
</tr>
<tr id="4885C425FFDE42F5CAA63430FA5C8D12" box="[159,1425,1583,1635]" gridrow="3" pageId="5" pageNumber="426">
<th id="0B54AD59FFDE42F5CAA63430FE9E8D12" box="[159,339,1583,1635]" gridcol="0" gridrow="3" pageId="5" pageNumber="426">
<taxonomicName id="31B5BDE4FFDEBD0FCAA6342FFEEE8D34" box="[159,291,1583,1605]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="426" phylum="Chordata" rank="species" species="maculosus">
<emphasis id="C4C11A75FFDEBD0FCAA6342FFF618D34" box="[159,172,1584,1605]" italics="true" pageId="5" pageNumber="426">P</emphasis>
.
<emphasis id="C4C11A75FFDEBD0FCA813430FEEE8D34" box="[184,291,1583,1605]" italics="true" pageId="5" pageNumber="426">maculosus</emphasis>
</taxonomicName>
</th>
<td id="0B54AD59FFDE42F5CBBA3430FD7C8D12" box="[387,689,1583,1635]" gridcol="1" gridrow="3" pageId="5" pageNumber="426">Short, subacuminate in dorsal and rounded in lateral view</td>
<td id="0B54AD59FFDE42F5C8E43430FC9D8D12" box="[733,848,1583,1635]" gridcol="2" gridrow="3" pageId="5" pageNumber="426">Present</td>
<td id="0B54AD59FFDE42F5C9423430FC0B8D12" box="[891,966,1583,1635]" gridcol="3" gridrow="3" pageId="5" pageNumber="426">Absent</td>
<td id="0B54AD59FFDE42F5CE393430FB478D12" box="[1024,1162,1583,1635]" gridcol="4" gridrow="3" pageId="5" pageNumber="426">Small, low</td>
<td id="0B54AD59FFDE42F5CE993430FA5C8D12" box="[1184,1425,1583,1635]" gridcol="5" gridrow="3" pageId="5" pageNumber="426">Lynch, 1991, this work.</td>
</tr>
<tr id="4885C425FFDE42F5CAA63468FA5C8DDA" box="[159,1425,1655,1707]" gridrow="4" pageId="5" pageNumber="426">
<th id="0B54AD59FFDE42F5CAA63468FE9E8DDA" box="[159,339,1655,1707]" gridcol="0" gridrow="4" pageId="5" pageNumber="426">
<taxonomicName id="31B5BDE4FFDEBD0FCAA63467FEEF8DFC" box="[159,290,1656,1677]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="426" phylum="Chordata" rank="species" species="parectatus">
<emphasis id="C4C11A75FFDEBD0FCAA63467FF618DFC" box="[159,172,1656,1677]" italics="true" pageId="5" pageNumber="426">P</emphasis>
.
<emphasis id="C4C11A75FFDEBD0FCA813467FEEF8DFC" box="[184,290,1656,1677]" italics="true" pageId="5" pageNumber="426">parectatus</emphasis>
</taxonomicName>
</th>
<td id="0B54AD59FFDE42F5CBBA3468FD7C8DDA" box="[387,689,1655,1707]" gridcol="1" gridrow="4" pageId="5" pageNumber="426">Short, subacuminate in dorsal and rounded in lateral view</td>
<td id="0B54AD59FFDE42F5C8E43468FC9D8DDA" box="[733,848,1655,1707]" gridcol="2" gridrow="4" pageId="5" pageNumber="426">Present</td>
<td id="0B54AD59FFDE42F5C9423468FC0B8DDA" box="[891,966,1655,1707]" gridcol="3" gridrow="4" pageId="5" pageNumber="426">Absent</td>
<td id="0B54AD59FFDE42F5CE393468FB478DDA" box="[1024,1162,1655,1707]" gridcol="4" gridrow="4" pageId="5" pageNumber="426">Low, oval</td>
<td id="0B54AD59FFDE42F5CE993468FA5C8DDA" box="[1184,1425,1655,1707]" gridcol="5" gridrow="4" pageId="5" pageNumber="426">Lynch &amp; Rueda- Almonacid, 1998.</td>
</tr>
<tr id="4885C425FFDE42F5CAA634A0FA5C8D82" box="[159,1425,1727,1779]" gridrow="5" pageId="5" pageNumber="426">
<th id="0B54AD59FFDE42F5CAA634A0FE9E8D82" box="[159,339,1727,1779]" gridcol="0" gridrow="5" pageId="5" pageNumber="426">
<taxonomicName id="31B5BDE4FFDEBD0FCAA634DFFEDB8DA4" box="[159,278,1728,1749]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="426" phylum="Chordata" rank="species" species="peraticus">
<emphasis id="C4C11A75FFDEBD0FCAA634DFFF618DA4" box="[159,172,1728,1749]" italics="true" pageId="5" pageNumber="426">P</emphasis>
.
<emphasis id="C4C11A75FFDEBD0FCA8134DFFEDB8DA4" box="[184,278,1728,1749]" italics="true" pageId="5" pageNumber="426">peraticus</emphasis>
</taxonomicName>
</th>
<td id="0B54AD59FFDE42F5CBBA34A0FD7C8D82" box="[387,689,1727,1779]" gridcol="1" gridrow="5" pageId="5" pageNumber="426">Short, subacuminate in dorsal and rounded in lateral profile</td>
<td id="0B54AD59FFDE42F5C8E434A0FC9D8D82" box="[733,848,1727,1779]" gridcol="2" gridrow="5" pageId="5" pageNumber="426">Absent</td>
<td id="0B54AD59FFDE42F5C94234A0FC0B8D82" box="[891,966,1727,1779]" gridcol="3" gridrow="5" pageId="5" pageNumber="426">Low</td>
<td id="0B54AD59FFDE42F5CE3934A0FB478D82" box="[1024,1162,1727,1779]" gridcol="4" gridrow="5" pageId="5" pageNumber="426">Low</td>
<td id="0B54AD59FFDE42F5CE9934A0FA5C8D82" box="[1184,1425,1727,1779]" gridcol="5" gridrow="5" pageId="5" pageNumber="426">Lynch, 1980, this work</td>
</tr>
<tr id="4885C425FFDE42F5CAA63518FA5C8C4A" box="[159,1425,1799,1851]" gridrow="6" pageId="5" pageNumber="426">
<th id="0B54AD59FFDE42F5CAA63518FE9E8C4A" box="[159,339,1799,1851]" gridcol="0" gridrow="6" pageId="5" pageNumber="426">
<taxonomicName id="31B5BDE4FFDEBD0FCAA63517FE9E8C6C" box="[159,339,1799,1821]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="426" phylum="Chordata" rank="species" species="scoloblepharus">
<emphasis id="C4C11A75FFDEBD0FCAA63517FF618C6C" box="[159,172,1800,1821]" italics="true" pageId="5" pageNumber="426">P</emphasis>
.
<emphasis id="C4C11A75FFDEBD0FCA813518FE9E8C6C" box="[184,339,1799,1821]" italics="true" pageId="5" pageNumber="426">scoloblepharus</emphasis>
</taxonomicName>
</th>
<td id="0B54AD59FFDE42F5CBBA3518FD7C8C4A" box="[387,689,1799,1851]" gridcol="1" gridrow="6" pageId="5" pageNumber="426">Subacuminate in dorsal and rounded in lateral view</td>
<td id="0B54AD59FFDE42F5C8E43518FC9D8C4A" box="[733,848,1799,1851]" gridcol="2" gridrow="6" pageId="5" pageNumber="426">Present</td>
<td id="0B54AD59FFDE42F5C9423518FC0B8C4A" box="[891,966,1799,1851]" gridcol="3" gridrow="6" pageId="5" pageNumber="426">Absent</td>
<td id="0B54AD59FFDE42F5CE393518FB478C4A" box="[1024,1162,1799,1851]" gridcol="4" gridrow="6" pageId="5" pageNumber="426">Obliques</td>
<td id="0B54AD59FFDE42F5CE993518FA5C8C4A" box="[1184,1425,1799,1851]" gridcol="5" gridrow="6" pageId="5" pageNumber="426">Lynch, 1991.</td>
</tr>
<tr id="4885C425FFDE42F5CAA63550FA5C8C14" box="[159,1425,1871,1893]" gridrow="7" pageId="5" pageNumber="426">
<th id="0B54AD59FFDE42F5CAA63550FE9E8C14" box="[159,339,1871,1893]" gridcol="0" gridrow="7" pageId="5" pageNumber="426">
<taxonomicName id="31B5BDE4FFDEBD0FCAA6354FFEE78C14" authority="Round" authorityName="Round" box="[159,298,1871,1893]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="426" phylum="Chordata" rank="species" species="uranobates">
<emphasis id="C4C11A75FFDEBD0FCAA6354FFF618C14" box="[159,172,1872,1893]" italics="true" pageId="5" pageNumber="426">P</emphasis>
.
<emphasis id="C4C11A75FFDEBD0FCA813550FEE78C14" box="[184,298,1871,1893]" italics="true" pageId="5" pageNumber="426">uranobates</emphasis>
</taxonomicName>
</th>
<td id="0B54AD59FFDE42F5CBBA3550FD7C8C14" box="[387,689,1871,1893]" gridcol="1" gridrow="7" pageId="5" pageNumber="426">Round in dorsal and lateral</td>
<td id="0B54AD59FFDE42F5C8E43550FC9D8C14" box="[733,848,1871,1893]" gridcol="2" gridrow="7" pageId="5" pageNumber="426">Present</td>
<td id="0B54AD59FFDE42F5C9423550FC0B8C14" box="[891,966,1871,1893]" gridcol="3" gridrow="7" pageId="5" pageNumber="426">Absent</td>
<td id="0B54AD59FFDE42F5CE393550FB478C14" box="[1024,1162,1871,1893]" gridcol="4" gridrow="7" pageId="5" pageNumber="426">Obliques</td>
<td id="0B54AD59FFDE42F5CE993550FA5C8C14" box="[1184,1425,1871,1893]" gridcol="5" gridrow="7" pageId="5" pageNumber="426">
<bibRefCitation id="9224BB96FFDEBD0FCE993550FAEC8C14" author="Lynch" box="[1184,1313,1871,1893]" pageId="5" pageNumber="436" refString="Lynch, J. D. (1991) New diminutive Eleutherodactylus from the Cordillera Central of Colombia (Amphibia: Leptodactylidae). Journal of Herpetology, 25 (3), 344 - 352. http: // dx. doi. org / 10.2307 / 1564595" type="journal article" year="1991">Lynch, 1991</bibRefCitation>
, this work
</td>
</tr>
</paragraph>
<paragraph id="F60AC667FFDEBD0FC8A53550FD008C14" box="[668,717,1871,1893]" pageId="5" pageNumber="426">view</paragraph>
</table>
<caption id="A2CA96EFFFDDBD0CCAAE3797FC808E8B" httpUri="https://zenodo.org/record/258321/files/figure.png" pageId="6" pageNumber="427" targetBox="[151,1436,193,1394]" targetPageId="6">
<paragraph id="F60AC667FFDDBD0CCAAE3797FC808E8B" blockId="6.[151,1436,1416,1530]" pageId="6" pageNumber="427">
<emphasis id="C4C11A75FFDDBD0CCAAE3797FEDF8EEC" bold="true" box="[151,274,1416,1438]" pageId="6" pageNumber="427">FIGURE 4.</emphasis>
Variation in lateral view of groin color of
<taxonomicName id="31B5BDE4FFDDBD0CC8843796FC4D8EEF" box="[701,896,1417,1438]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="427" phylum="Chordata" rank="species" species="stictus" status="sp. nov.">
<emphasis id="C4C11A75FFDDBD0CC8843796FC4D8EEF" box="[701,896,1417,1438]" italics="true" pageId="6" pageNumber="427">Pristimantis stictus</emphasis>
</taxonomicName>
<emphasis id="C4C11A75FFDDBD0CC9BF3796FC1B8EEF" bold="true" box="[902,982,1417,1438]" pageId="6" pageNumber="427">
<taxonomicNameLabel id="DFF2A70EFFDDBD0CC9BF3796FC1B8EEF" box="[902,982,1417,1438]" pageId="6" pageNumber="427" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and comparisons with other similar taxa. A.
<taxonomicName id="31B5BDE4FFDDBD0CCAAE37B7FF3C8ECC" box="[151,241,1448,1469]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="427" phylum="Chordata" rank="species" species="stictus" status="sp. nov.">
<emphasis id="C4C11A75FFDDBD0CCAAE37B7FF698ECC" box="[151,164,1448,1469]" italics="true" pageId="6" pageNumber="427">P</emphasis>
.
<emphasis id="C4C11A75FFDDBD0CCA8937B7FF3C8ECC" box="[176,241,1448,1469]" italics="true" pageId="6" pageNumber="427">stictus</emphasis>
</taxonomicName>
<emphasis id="C4C11A75FFDDBD0CCACE37B8FE858ECD" bold="true" box="[247,328,1447,1468]" pageId="6" pageNumber="427">
<taxonomicNameLabel id="DFF2A70EFFDDBD0CCACE37B8FE858ECD" box="[247,328,1447,1468]" pageId="6" pageNumber="427" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult female (ICN-55702); B.
<taxonomicName id="31B5BDE4FFDDBD0CC8B137B7FCC68ECC" box="[648,779,1448,1469]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="427" phylum="Chordata" rank="species" species="parectatus">
<emphasis id="C4C11A75FFDDBD0CC8B137B7FD598ECC" box="[648,660,1448,1469]" italics="true" pageId="6" pageNumber="427">P</emphasis>
.
<emphasis id="C4C11A75FFDDBD0CC89837B7FCC68ECC" box="[673,779,1448,1469]" italics="true" pageId="6" pageNumber="427">parectatus</emphasis>
</taxonomicName>
, adult female (ICN-55740); C.
<taxonomicName id="31B5BDE4FFDDBD0CCE7237B7FB028ECC" box="[1099,1231,1447,1469]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="427" phylum="Chordata" rank="species" species="maculosus">
<emphasis id="C4C11A75FFDDBD0CCE7237B7FB9A8ECC" box="[1099,1111,1448,1469]" italics="true" pageId="6" pageNumber="427">P</emphasis>
.
<emphasis id="C4C11A75FFDDBD0CCE5D37B8FB028ECC" box="[1124,1231,1447,1469]" italics="true" pageId="6" pageNumber="427">maculosus</emphasis>
</taxonomicName>
, adult female (ICN- 55578); D.
<taxonomicName id="31B5BDE4FFDDBD0CCB3437D8FE0F8EAA" box="[269,450,1477,1500]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="427" phylum="Chordata" rank="species" species="scoloblepharus">
<emphasis id="C4C11A75FFDDBD0CCB3437D8FED78EAD" box="[269,282,1479,1500]" italics="true" pageId="6" pageNumber="427">P</emphasis>
.
<emphasis id="C4C11A75FFDDBD0CCB1E37DAFE0F8EAA" box="[295,450,1477,1499]" italics="true" pageId="6" pageNumber="427">scoloblepharus</emphasis>
</taxonomicName>
, adult females (ICN-55744) (Photo: Marco Rada), SVL: 26.5 mm; E.
<taxonomicName id="31B5BDE4FFDDBD0CCEAC37D8FAEF8EAA" box="[1173,1314,1477,1500]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="427" phylum="Chordata" rank="species" species="uranobates">
<emphasis id="C4C11A75FFDDBD0CCEAC37D8FB6C8EAD" box="[1173,1185,1479,1500]" italics="true" pageId="6" pageNumber="427">P</emphasis>
.
<emphasis id="C4C11A75FFDDBD0CCE8937DAFAEF8EAA" box="[1200,1314,1477,1499]" italics="true" pageId="6" pageNumber="427">uranobates</emphasis>
</taxonomicName>
, female not collected; F.
<taxonomicName id="31B5BDE4FFDDBD0CCB2137FAFE5D8E8B" box="[280,400,1509,1530]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="427" phylum="Chordata" rank="species" species="peraticus">
<emphasis id="C4C11A75FFDDBD0CCB2137FAFEE98E8B" box="[280,292,1509,1530]" italics="true" pageId="6" pageNumber="427">P</emphasis>
.
<emphasis id="C4C11A75FFDDBD0CCB0B37FAFE5D8E8B" box="[306,400,1509,1530]" italics="true" pageId="6" pageNumber="427">peraticus</emphasis>
</taxonomicName>
, female not collected (Photo: Marco Rada).
</paragraph>
</caption>
<subSubSection id="BEAF95ECFFDDBD0DCAFE3438FC3F8ADD" lastPageId="7" lastPageNumber="428" pageId="6" pageNumber="427" type="diagnosis">
<paragraph id="F60AC667FFDDBD0CCAFE3438FBA18C9E" blockId="6.[151,1436,1575,2041]" pageId="6" pageNumber="427">
<emphasis id="C4C11A75FFDDBD0CCAFE3438FE8C8D31" bold="true" box="[199,321,1575,1600]" pageId="6" pageNumber="427">Diagnosis.</emphasis>
The new species can be diagnosed by the following combination of characters: (1) Dorsal skin smooth; dorsolateral folds present, continuous from eyelid to the sacrum; ventral skin coarsely areolate; discoidal fold absent. (2) Tympanum superficial, tympanic annulus prominent, round, corresponding to 1/2–1/3 (35%–46%) of eye length, with an obscure stripe on the supratympanic fold. (3) Snout short, subacuminate in dorsal view, rounded in lateral profile; canthus rostralis concave. (4) IOD wider than upper eyelid; craneal crests absent, upper eyelid bearing small subconical tubercle. (5) Vomerine odontophores absent. (6) Males with vocal slits and subgular vocal sac; nuptial pads in males absent. (7) Finger I shorter than II, with large rounded digital disc. (8) Lateral fringes on fingers. (9) Ulnar tubercles small, subconical. (10) Subconical tubercles on heel and outer edge of tarsus, small inner tarsal fold. (11) Two oval inner metatarsal tubercles, representing six times the size of outer tubercle; supernumerary plantar tubercles present. (12) Toes with lateral fringes; toe discs slightly narrower than those on fingers. (13) Dorsum brown, yellowish or copper, with golden tones, in some cases spotted yellow; laterally light brown; groin yellowish with dark spots; venter and legs yellow with dark spots. (14) Adults small, males 17.8–22.2 (
<emphasis id="C4C11A75FFDDBD0CCB5D35C2FEBF8C88" bold="true" box="[356,370,2013,2041]" pageId="6" pageNumber="427">x</emphasis>
= 19.9 ± 1.2; n=15) and females 22.0–28.2 (
<emphasis id="C4C11A75FFDDBD0CC94935C2FCB38C88" bold="true" box="[880,894,2013,2041]" pageId="6" pageNumber="427">x</emphasis>
= 25.4 ± 1.6; n=17).
</paragraph>
<paragraph id="F60AC667FFDCBD0DCAFE3288FC3F8ADD" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="428">
<emphasis id="C4C11A75FFDCBD0DCAFE3288FEA18BC1" bold="true" box="[199,364,151,176]" pageId="7" pageNumber="428">Comparisons.</emphasis>
<taxonomicName id="31B5BDE4FFDCBD0DCB423287FD968BDE" box="[379,603,152,175]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="428" phylum="Chordata" rank="species" species="stictus">
<emphasis id="C4C11A75FFDCBD0DCB423287FD968BDE" box="[379,603,152,175]" italics="true" pageId="7" pageNumber="428">Pristimantis stictus</emphasis>
</taxonomicName>
differs from other species of the cloud forest from the northern of Cordillera Central (
<taxonomicName id="31B5BDE4FFDCBD0DCB4A32A2FDCB8BA5" box="[371,518,188,212]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="428" phylum="Chordata" rank="species" species="maculosus">
<emphasis id="C4C11A75FFDCBD0DCB4A32A2FE4C8BA5" box="[371,385,189,212]" italics="true" pageId="7" pageNumber="428">P</emphasis>
.
<emphasis id="C4C11A75FFDCBD0DCBA932A3FDCB8BA5" box="[400,518,188,212]" italics="true" pageId="7" pageNumber="428">maculosus</emphasis>
</taxonomicName>
,
<taxonomicName id="31B5BDE4FFDCBD0DC82C32A2FD6A8BA5" box="[533,679,189,212]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="428" phylum="Chordata" rank="species" species="parectatus">
<emphasis id="C4C11A75FFDCBD0DC82C32A2FDEE8BA5" box="[533,547,189,212]" italics="true" pageId="7" pageNumber="428">P</emphasis>
.
<emphasis id="C4C11A75FFDCBD0DC80B32A2FD6A8BA5" box="[562,679,189,212]" italics="true" pageId="7" pageNumber="428">parectatus</emphasis>
</taxonomicName>
, and
<taxonomicName id="31B5BDE4FFDCBD0DC8DC32A2FC608BA5" box="[741,941,188,212]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="428" phylum="Chordata" rank="species" species="scoloblepharus">
<emphasis id="C4C11A75FFDCBD0DC8DC32A2FD3E8BA5" box="[741,755,189,212]" italics="true" pageId="7" pageNumber="428">P</emphasis>
.
<emphasis id="C4C11A75FFDCBD0DC93B32A3FC608BA5" box="[770,941,188,212]" italics="true" pageId="7" pageNumber="428">scoloblepharus</emphasis>
</taxonomicName>
) by the absence of vomerine odontophores, low and thin dorsolateral folds extended from the posterior eyelid to beyond the sacrum, small tubercles on the eyelid, and by lacking nuptial pads (
<figureCitation id="6E8EDAE2FFDCBD0DC86C331BFD708A6D" box="[597,701,260,285]" captionStart="FIGURE 3" captionStartId="4.[151,250,1576,1598]" captionTargetBox="[151,1436,193,1552]" captionTargetId="figure@4.[151,1436,193,1554]" captionTargetPageId="4" captionText="FIGURE 3. Variation in life coloration. A. Pristimantis leptolophus, adult female (ICN- 7799) (Photo: John D. Lynch); B. Pristimantis uranobates, adult female (ICN- 55588); C. Pristimantis peraticus, adult female (ICN- 55742) (Photo: Marco Rada); D. Pristimantis maculosus, adult female (ICN- 55578); E. Pristimantis parectatus, adult female (ICN- 55743) (Photo: Marco Rada); F. Pristimantis scoloblepharus, holotype adult female (ICN- 8583) (Photo: John D. Lynch)." httpUri="https://zenodo.org/record/258320/files/figure.png" pageId="7" pageNumber="428">Figure 3</figureCitation>
).
<taxonomicName id="31B5BDE4FFDCBD0DC8E3331AFC748A6D" box="[730,953,261,284]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="428" phylum="Chordata" rank="species" species="stictus">
<emphasis id="C4C11A75FFDCBD0DC8E3331AFC748A6D" box="[730,953,261,284]" italics="true" pageId="7" pageNumber="428">Pristimantis stictus</emphasis>
</taxonomicName>
differs from similar paramo species (
<taxonomicName id="31B5BDE4FFDCBD0DCFBE331AFED08A31" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="428" phylum="Chordata" rank="species" species="lasalleorum">
<emphasis id="C4C11A75FFDCBD0DCFBE331AFA588A6D" box="[1415,1429,261,284]" italics="true" pageId="7" pageNumber="428">P</emphasis>
.
<emphasis id="C4C11A75FFDCBD0DCAAE3337FED08A31" box="[151,285,296,320]" italics="true" pageId="7" pageNumber="428">lasalleorum</emphasis>
</taxonomicName>
,
<taxonomicName id="31B5BDE4FFDCBD0DCB093336FE188A31" box="[304,469,296,320]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="428" phylum="Chordata" rank="species" species="leptolophus">
<emphasis id="C4C11A75FFDCBD0DCB093336FEF38A31" box="[304,318,297,320]" italics="true" pageId="7" pageNumber="428">P</emphasis>
.
<emphasis id="C4C11A75FFDCBD0DCB6B3337FE188A31" box="[338,469,296,320]" italics="true" pageId="7" pageNumber="428">leptolophus</emphasis>
</taxonomicName>
and
<taxonomicName id="31B5BDE4FFDCBD0DC82C3336FD528A4E" box="[533,671,296,320]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="428" phylum="Chordata" rank="species" species="peraticus">
<emphasis id="C4C11A75FFDCBD0DC82C3336FDEE8A31" box="[533,547,297,320]" italics="true" pageId="7" pageNumber="428">P</emphasis>
.
<emphasis id="C4C11A75FFDCBD0DC80E3337FD528A4E" box="[567,671,296,319]" italics="true" pageId="7" pageNumber="428">peraticus</emphasis>
</taxonomicName>
) by presenting tympanic annulus and cavum tympanicum, more enlarged discs, and broad lateral fringes on fingers (
<figureCitation id="6E8EDAE2FFDCBD0DC8D73353FC9F8A15" box="[750,850,332,357]" captionStart="FIGURE 3" captionStartId="4.[151,250,1576,1598]" captionTargetBox="[151,1436,193,1552]" captionTargetId="figure@4.[151,1436,193,1554]" captionTargetPageId="4" captionText="FIGURE 3. Variation in life coloration. A. Pristimantis leptolophus, adult female (ICN- 7799) (Photo: John D. Lynch); B. Pristimantis uranobates, adult female (ICN- 55588); C. Pristimantis peraticus, adult female (ICN- 55742) (Photo: Marco Rada); D. Pristimantis maculosus, adult female (ICN- 55578); E. Pristimantis parectatus, adult female (ICN- 55743) (Photo: Marco Rada); F. Pristimantis scoloblepharus, holotype adult female (ICN- 8583) (Photo: John D. Lynch)." httpUri="https://zenodo.org/record/258320/files/figure.png" pageId="7" pageNumber="428">Figure 3</figureCitation>
). It can be easily distinguished from the sympatric species
<taxonomicName id="31B5BDE4FFDCBD0DCAD6336EFE448AF9" box="[239,393,368,392]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="428" phylum="Chordata" rank="species" species="uranobates">
<emphasis id="C4C11A75FFDCBD0DCAD6336EFF308AF9" box="[239,253,369,392]" italics="true" pageId="7" pageNumber="428">P</emphasis>
.
<emphasis id="C4C11A75FFDCBD0DCB32336FFE448AF9" box="[267,393,368,392]" italics="true" pageId="7" pageNumber="428">uranobates</emphasis>
</taxonomicName>
by the absence of vomerine teeth and nuptial pads, and yellow groin with black reticulations (
<figureCitation id="6E8EDAE2FFDCBD0DCAA7338BFECC8ADD" box="[158,257,404,429]" captionStart="FIGURE 4" captionStartId="6.[151,250,1416,1438]" captionTargetBox="[151,1436,193,1394]" captionTargetId="figure@6.[151,1436,193,1395]" captionTargetPageId="6" captionText="FIGURE 4. Variation in lateral view of groin color of Pristimantis stictus sp. nov., and comparisons with other similar taxa. A. P. stictus sp. nov., adult female (ICN- 55702); B. P. parectatus, adult female (ICN- 55740); C. P. maculosus, adult female (ICN- 55578); D. P. scoloblepharus, adult females (ICN- 55744) (Photo: Marco Rada), SVL: 26.5 mm; E. P. uranobates, female not collected; F. P. peraticus, female not collected (Photo: Marco Rada)." httpUri="https://zenodo.org/record/258321/files/figure.png" pageId="7" pageNumber="428">Figure 4</figureCitation>
). Additional differences among species are summarized in
<tableCitation id="BB37F3DCFFDCBD0DC9A1338BFC218ADD" box="[920,1004,404,429]" captionStart="TABLE 1" captionStartId="5.[151,239,152,175]" captionTargetBox="[151,1427,202,818]" captionTargetPageId="5" captionText="TABLE 1. Diagnostic morphological characters in the Pristimantis leptolophus species group." httpUri="http://table.plazi.org/id/A2CA96EFFFDEBD0FCAAE3287FB4A8BDE" pageId="7" pageNumber="428" tableUuid="A2CA96EFFFDEBD0FCAAE3287FB4A8BDE">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BEAF95ECFFDCBD03CAFE33A8FCA98A84" lastPageId="9" lastPageNumber="430" pageId="7" pageNumber="428" type="description">
<paragraph id="F60AC667FFDCBD0DCAFE33A8FF168E24" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="428">
<emphasis id="C4C11A75FFDCBD0DCAFE33A8FD558AA1" bold="true" box="[199,664,439,464]" pageId="7" pageNumber="428">
Description of the
<typeStatus id="290E78C5FFDCBD0DCB9033A8FDC28AA1" box="[425,527,439,464]" pageId="7" pageNumber="428" type="holotype">holotype</typeStatus>
(
<figureCitation id="6E8EDAE2FFDCBD0DC81933A8FD458AA1" box="[544,648,439,464]" captionStart="FIGURE 1. A" captionStartId="2.[151,250,1748,1770]" captionTargetBox="[264,1322,265,1725]" captionTargetId="figure@2.[264,1323,265,1727]" captionTargetPageId="2" captionText="FIGURE 1. A. Ventral view of Pristimantis stictus sp. nov., adult female holotype, (ICN- 55689). B. Dorsal view of P. stictus sp. nov., adult female holotype, (ICN- 55689). C. Ventral view of hand of P. stictus sp. nov., adult female holotype, (ICN- 55689). D. Ventral view of foot of P. stictus sp. nov., adult female holotype, (ICN- 55689). E. Lateral view of P. stictus sp. nov., adult female holotype (ICN- 55689). White bars equal 4 mm." httpUri="https://zenodo.org/record/258318/files/figure.png" pageId="7" pageNumber="428">Figure 1</figureCitation>
).
</emphasis>
Head wider than body; snout rounded in dorsal view and in lateral profile. Snout short, canthus rostralis concave, loreal region is slightly concave, sloping abruptly to lips; lips not flared. Nostrils protruding, directed laterally. Upper eyelid bearing one small subconical tubercle. No cranial crests. Supratympanic fold thin, obscure on upper and posterior edges of tympanum. Tympanum superficial, round, separated from eye by a distance equal to its size, tympanic annulus prominent. Postrictal tubercles small, subconical. Choanae small, round, not concealed by palatal shelf of maxillary arch. Vomerine odontophores absent, represented by oblique keels posteromedial to choanae, not bearing teeth. Tongue longer than wide, posterior edge not notched, posterior one-half not adherent to floor of mouth. Skin of dorsum finely shagreen with small and low spicules. Dorsolateral folds extending from eye to sacral region, lacking thickening or distinctive coloration. Flanks granular, more distinctive than dorsum, with numerous low warts. Groin smooth. Tympanic folds with superposition of tubercles, especially posterior to tympanum. Ventral surface areolate and throat weakly areolate. Discoidal fold indistinct. Upper surface of limbs smooth, without ridges. No cloacal sheath. Two postcloacal warts. Forearm bearing a row of three subconical ulnar tubercles. A broad lateral fringe along outer edge of palm and continuing along outer edge on finger IV. Palmar tubercle bifid, wider than oval tenar tubercle. Six supernumerary tubercles. Subarticular tubercles rounded and nonconical. Fingers bearing broad lateral fringes, and round disks. Pad on thumb slightly wider than digit proximal to pad. The pads on fingers II–IV wider than digits, those of fingers III and IV have discs as large as tympanum or twice the width of digit; ventral pads, broader than long. First finger shorter than second. Subconical tubercle on heel, two small tubercles on outer edge of tarsus. Inner tarsal fold present, its length equals one third of tarsus. Inner metatarsal tubercule twice as long as wide, six times the size of the rounded subconical outer tubercle. Plantar surfaces bearing numerous low supernumerary tubercles. Subarticular tubercles rounded, nonconical. Toes bearing lateral fringes, narrower than fringes on the fingers; toes lack webbing. Discs on toes III, IV and V wider than digits; discs on toes IV and V wider than those of the inner toes; ventral pad as wide as long. Outer pads on toes narrower than the pads on outer fingers. Tip of toe V reaches proximal third of distal subarticular tubercle of toe IV, tip of toe III reaches the distal border of penultimate subarticular tubercle of toe IV. Heels broadly overlapping when hindlimbs are flexed perpendicular to the sagittal plane.
</paragraph>
<paragraph id="F60AC667FFDCBD0DCAFE3740FB548E94" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="428">
<emphasis id="C4C11A75FFDCBD0DCAFE3740FD658E09" bold="true" box="[199,680,1375,1400]" pageId="7" pageNumber="428">
Measurements of the
<typeStatus id="290E78C5FFDCBD0DCBF63740FDFB8E09" box="[463,566,1375,1400]" pageId="7" pageNumber="428" type="holotype">holotype</typeStatus>
(in mm).
</emphasis>
Snout-vent length =26.7, radio-ulna length=6.3, hand length=8.3, tibia length=12.5, foot length=12.7, head length=10.1, head width=8.9, upper eyelid width=2.3, eye diameter=3.2, eye to nostril distance=2.2, internarial distance=2.4, snout to eye distance=4.3, interorbital distance=3.5, eye to tympanum distance=0.8, tympanum diameter=1.5, finger I length=2.2, finger II length=3.0.
</paragraph>
<paragraph id="F60AC667FFDCBD0DCAFE37F0FD1C8DCD" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="428">
<emphasis id="C4C11A75FFDCBD0DCAFE37F0FDC18D79" bold="true" box="[199,524,1519,1544]" pageId="7" pageNumber="428">
Coloration of the
<typeStatus id="290E78C5FFDCBD0DCBA637F0FDC58D79" box="[415,520,1519,1544]" pageId="7" pageNumber="428" type="holotype">holotype</typeStatus>
.
</emphasis>
In life, copper-colored or brown dorsally; laterally and dorsal surfaces of limbs have a lighter brown color. The groin presentes a yellowish color with elongated black spots. The belly is bright yellow with small black dots. The throat is copper. The undersides of thighs have small black dots, while the spots are elongated on the posterodorsal surface. Cloacal triangle absent. Dorsolateral folds lacking a distinctive color. Iris yellow with a reddish transversal band. Rostral markings inconspicuous. In ethanol, the patterns remain, but the dorsum turns gray and ventral surfaces turn cream.
</paragraph>
<paragraph id="F60AC667FFDCBD02CAFE34D8FDB98A31" blockId="7.[151,1437,151,2013]" lastBlockId="8.[151,1436,151,320]" lastPageId="8" lastPageNumber="429" pageId="7" pageNumber="428">
<emphasis id="C4C11A75FFDCBD0DCAFE34D8FDAF8D91" bold="true" box="[199,610,1735,1760]" pageId="7" pageNumber="428">
Adult cranial osteology (
<figureCitation id="6E8EDAE2FFDCBD0DCBD534D8FD998D91" box="[492,596,1735,1760]" captionStart="FIGURE 5" captionStartId="8.[151,250,1725,1747]" captionTargetBox="[151,1436,410,1704]" captionTargetId="figure@8.[151,1436,373,1704]" captionTargetPageId="8" captionText="FIGURE 5. Cranium of adult female of P. stictus (ICN- 55591; SVL: 25.6 mm) in dorsal (A) and ventral view (B), bars represent measurements of 2.5 mm, black indicates foramina, gray cartilage and stippling bones; C. Photography of oblique keel of vomer slightly concealed by mucosa in P. stictus (ICN- 55697; paratype); D. Vomer in cleared and stained specimen of adult female of P. stictus (ICN- 55692)." httpUri="https://zenodo.org/record/258322/files/figure.png" pageId="7" pageNumber="428">Figure 5</figureCitation>
).
</emphasis>
The maximum width of the skull is at the posterior level of the maxillae. Bones of the skull are well ossified. Both centers of ossification of the sphenethmoid are fused dorso and ventromedially; dorsally, it slightly overlaps the anterior part of frontoparietals and a small portion of the posterolateral border of the nasals. The nasals are medially separated; the maxillary process does not reach the preorbital process of the maxilla. Nasals do not articulate with the frontoparietals. The frontoparietal fenestra is absent. No cranial crest. Posterolaterally frontoparietals extend to the level of the epiotic eminence of the prootic. The frontoparietals are not fused with the prootic. The sphenethmoid ventrally subtends the dentigerous processes of vomers, and the cultriform process of parasphenoid. The cultriform process reaches anterior to the level of palatines. Dorsally, the otoccipital bears a cartilaginous parotic crista that connects with the otic ramus of the squamosal. Stapes present. The vomers distinctly medially separated, prechoanal process indistinct; the postchoanal process is directed laterally and narrow. Dentigerous processes of vomers are elongated and narrow, directed posteromedially, extending anteriorly to palatines; externally, it is represented by oblique keel of vomer (
<figureCitation id="6E8EDAE2FFD3BD02CAA73338FF308A31" box="[158,253,295,320]" captionStart="FIGURE 5" captionStartId="8.[151,250,1725,1747]" captionTargetBox="[151,1436,410,1704]" captionTargetId="figure@8.[151,1436,373,1704]" captionTargetPageId="8" captionText="FIGURE 5. Cranium of adult female of P. stictus (ICN- 55591; SVL: 25.6 mm) in dorsal (A) and ventral view (B), bars represent measurements of 2.5 mm, black indicates foramina, gray cartilage and stippling bones; C. Photography of oblique keel of vomer slightly concealed by mucosa in P. stictus (ICN- 55697; paratype); D. Vomer in cleared and stained specimen of adult female of P. stictus (ICN- 55692)." httpUri="https://zenodo.org/record/258322/files/figure.png" pageId="8" pageNumber="429">Figure 5</figureCitation>
C–D). Vomerine teeth are absent.
</paragraph>
<caption id="A2CA96EFFFD3BD02CAAE34A2FDEF8C41" httpUri="https://zenodo.org/record/258322/files/figure.png" pageId="8" pageNumber="429" targetBox="[151,1436,410,1704]" targetPageId="8">
<paragraph id="F60AC667FFD3BD02CAAE34A2FDEF8C41" blockId="8.[151,1436,1725,1840]" pageId="8" pageNumber="429">
<emphasis id="C4C11A75FFD3BD02CAAE34A2FEDB8DA2" bold="true" box="[151,278,1725,1747]" pageId="8" pageNumber="429">FIGURE 5.</emphasis>
Cranium of adult female of
<taxonomicName id="31B5BDE4FFD3BD02C87434A0FD668DA5" box="[589,683,1726,1748]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="429" phylum="Chordata" rank="species" species="stictus">
<emphasis id="C4C11A75FFD3BD02C87434A0FD948DA5" box="[589,601,1727,1748]" italics="true" pageId="8" pageNumber="429">P</emphasis>
.
<emphasis id="C4C11A75FFD3BD02C85334A0FD668DA5" box="[618,683,1727,1748]" italics="true" pageId="8" pageNumber="429">stictus</emphasis>
</taxonomicName>
(ICN-55591; SVL: 25.6 mm) in dorsal (A) and ventral view (B), bars represent measurements of 2.5 mm, black indicates foramina, gray cartilage and stippling bones; C. Photography of oblique keel of vomer slightly concealed by mucosa in
<taxonomicName id="31B5BDE4FFD3BD02C84034E3FD198C60" box="[633,724,1788,1809]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="429" phylum="Chordata" rank="species" species="stictus">
<emphasis id="C4C11A75FFD3BD02C84034E3FD4B8C60" box="[633,646,1788,1809]" italics="true" pageId="8" pageNumber="429">P</emphasis>
.
<emphasis id="C4C11A75FFD3BD02C8AA34E3FD198C60" box="[659,724,1788,1809]" italics="true" pageId="8" pageNumber="429">stictus</emphasis>
</taxonomicName>
(ICN-55697; paratype); D. Vomer in cleared and stained specimen of adult female of
<taxonomicName id="31B5BDE4FFD3BD02CB0F3504FE5C8C41" box="[310,401,1819,1840]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="429" phylum="Chordata" rank="species" species="stictus">
<emphasis id="C4C11A75FFD3BD02CB0F3504FE8F8C41" box="[310,322,1819,1840]" italics="true" pageId="8" pageNumber="429">P</emphasis>
.
<emphasis id="C4C11A75FFD3BD02CB693504FE5C8C41" box="[336,401,1819,1840]" italics="true" pageId="8" pageNumber="429">stictus</emphasis>
</taxonomicName>
(ICN-55692).
</paragraph>
</caption>
<paragraph id="F60AC667FFD3BD03CAFE3548FCA98A84" blockId="8.[151,1436,1879,2012]" lastBlockId="9.[151,1437,151,501]" lastPageId="9" lastPageNumber="430" pageId="8" pageNumber="429">
<emphasis id="C4C11A75FFD3BD02CAFE3548FEF68C01" bold="true" box="[199,315,1879,1904]" pageId="8" pageNumber="429">Variation.</emphasis>
Measurements and ratios of males and females are shown in
<tableCitation id="BB37F3DCFFD3BD02CE3C3548FB908C1E" box="[1029,1117,1879,1904]" captionStart="TABLE 2" captionStartId="9.[151,239,544,567]" captionTargetBox="[159,1384,626,1912]" captionTargetPageId="9" captionText="TABLE 2. Measurements and morphological proportions (in percentages) of adult females and males of Pristimantis stictus (range, mean, and standard deviation (SD) in mm). Head width (HW); snout-vent length (SVL)." httpUri="http://table.plazi.org/id/A2CA96EFFFD2BD03CAAE303FFB2C8926" pageId="8" pageNumber="429" tableUuid="A2CA96EFFFD2BD03CAAE303FFB2C8926">Table 2</tableCitation>
. The head is wider than the body in males and narrower or equal in females. The snout is round in lateral profile in females and in some it may be slightly protruded (ICN-55701, 55707-08). Males and some females (ICN-55691, 55693-95, 55697, 55701, 55703, 55707-709, 55711, 55714-16) have a tubercle on the tip of the snout. The supratympanic stripe is not evident in some females, but it is present in all males and some females (ICN-55690, 55693, 55695, 55697, 55702, 55713). All females lack vomerine odontophores, however, some (ICN-55692-93, 55696-97, 55699, 55701-02, 55704, 55712) possess the oblique keel that lies posterior to the choanae. Some males (ICN-55705, 55708-09, 55711) do not present the oblique keel while others (ICN-55698, 55707) may have a tooth in the posterior end of the oblique keel. The dorsal skin color varies from dark brown to yellowish or copper. The groin may be yellow in females, with small dark dots or elongated spots (ICN-55690, 55692, 55696, 55713). Males usually possess spots on groin (ICN-55694, 55709, 55711, 55714). The belly is yellowish with black spots in females, while males do not present such pattern (ICN-55694, 55705, 55708-09, 55711). The gular region of males is dark grey. Females present small black dots on the ventral part of thighs and elongated spots on posterodorsal areas. Dorsolateral folds have no distinctive color. The iris varies from reddish to yellow.
</paragraph>
</subSubSection>
<caption id="A2CA96EFFFD2BD03CAAE303FFB2C8926" ID-Table-UUID="A2CA96EFFFD2BD03CAAE303FFB2C8926" httpUri="http://table.plazi.org/id/A2CA96EFFFD2BD03CAAE303FFB2C8926" pageId="9" pageNumber="430" targetBox="[159,1384,626,1912]" targetIsTable="true" targetPageId="9">
<paragraph id="F60AC667FFD2BD03CAAE303FFB2C8926" blockId="9.[151,1436,544,599]" pageId="9" pageNumber="430">
<emphasis id="C4C11A75FFD2BD03CAAE303FFEC18946" bold="true" box="[151,268,544,567]" pageId="9" pageNumber="430">TABLE 2.</emphasis>
Measurements and morphological proportions (in percentages) of adult females and males of
<taxonomicName id="31B5BDE4FFD2BD03CF23303FFF168926" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="430" phylum="Chordata" rank="species" species="stictus">
<emphasis id="C4C11A75FFD2BD03CF23303FFF168926" italics="true" pageId="9" pageNumber="430">Pristimantis stictus</emphasis>
</taxonomicName>
(range, mean, and standard deviation (SD) in mm). Head width (HW); snout-vent length (SVL).
</paragraph>
</caption>
<paragraph id="F60AC667FFD2BD03C858306DFAA58C09" pageId="9" pageNumber="430">
<table id="84B534C7FFD242F5CAA6306DFAA58C09" box="[159,1384,626,1912]" gridcols="7" gridrows="31" pageId="9" pageNumber="430">
<tr id="4885C425FFD242F5CAA6306DFAA589F9" box="[159,1384,626,648]" gridrow="0" pageId="9" pageNumber="430" rowspan-0="1" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1">
<th id="0B54AD59FFD242F5C858306DFD3D89F9" box="[609,752,626,648]" gridcol="1" gridrow="0" pageId="9" pageNumber="430">Females n=17</th>
<th id="0B54AD59FFD242F5CE3E306DFB4B89F9" box="[1031,1158,626,648]" gridcol="4" gridrow="0" pageId="9" pageNumber="430">Males n=15</th>
</tr>
<tr id="4885C425FFD242F5CAA63081FAA589C5" box="[159,1384,670,692]" gridrow="1" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA63081FE3089C5" box="[159,509,670,692]" gridcol="0" gridrow="1" pageId="9" pageNumber="430">Character</th>
<td id="0B54AD59FFD242F5C8583081FD3D89C5" box="[609,752,670,692]" gridcol="1" gridrow="1" pageId="9" pageNumber="430">Range</td>
<td id="0B54AD59FFD242F5C90E3081FCBC89C5" box="[823,881,670,692]" gridcol="2" gridrow="1" pageId="9" pageNumber="430">Mean</td>
<td id="0B54AD59FFD242F5C99B3081FC0F89C5" box="[930,962,670,692]" gridcol="3" gridrow="1" pageId="9" pageNumber="430">SD</td>
<td id="0B54AD59FFD242F5CE3E3081FB4B89C5" box="[1031,1158,670,692]" gridcol="4" gridrow="1" pageId="9" pageNumber="430">Range</td>
<td id="0B54AD59FFD242F5CEE43081FADA89C5" box="[1245,1303,670,692]" gridcol="5" gridrow="1" pageId="9" pageNumber="430">Mean</td>
<td id="0B54AD59FFD242F5CF713081FAA589C5" box="[1352,1384,670,692]" gridcol="6" gridrow="1" pageId="9" pageNumber="430">SD</td>
</tr>
<tr id="4885C425FFD242F5CAA630D5FAA58991" box="[159,1384,714,736]" gridrow="2" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA630D5FE308991" box="[159,509,714,736]" gridcol="0" gridrow="2" pageId="9" pageNumber="430">Snout-vent length</th>
<td id="0B54AD59FFD242F5C85830D5FD3D8991" box="[609,752,714,736]" gridcol="1" gridrow="2" pageId="9" pageNumber="430">22–28.2</td>
<td id="0B54AD59FFD242F5C90E30D5FCBC8991" box="[823,881,714,736]" gridcol="2" gridrow="2" pageId="9" pageNumber="430">25.4</td>
<td id="0B54AD59FFD242F5C99B30D5FC0F8991" box="[930,962,714,736]" gridcol="3" gridrow="2" pageId="9" pageNumber="430">1.6</td>
<td id="0B54AD59FFD242F5CE3E30D5FB4B8991" box="[1031,1158,714,736]" gridcol="4" gridrow="2" pageId="9" pageNumber="430">17.8–22.2</td>
<td id="0B54AD59FFD242F5CEE430D5FADA8991" box="[1245,1303,714,736]" gridcol="5" gridrow="2" pageId="9" pageNumber="430">19.9</td>
<td id="0B54AD59FFD242F5CF7130D5FAA58991" box="[1352,1384,714,736]" gridcol="6" gridrow="2" pageId="9" pageNumber="430">1.2</td>
</tr>
<tr id="4885C425FFD242F5CAA630EBFAA5887B" box="[159,1384,756,778]" gridrow="3" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA630EBFE30887B" box="[159,509,756,778]" gridcol="0" gridrow="3" pageId="9" pageNumber="430">Radioulna length</th>
<td id="0B54AD59FFD242F5C85830EBFD3D887B" box="[609,752,756,778]" gridcol="1" gridrow="3" pageId="9" pageNumber="430">5.8–7.4</td>
<td id="0B54AD59FFD242F5C90E30EBFCBC887B" box="[823,881,756,778]" gridcol="2" gridrow="3" pageId="9" pageNumber="430">6.4</td>
<td id="0B54AD59FFD242F5C99B30EBFC0F887B" box="[930,962,756,778]" gridcol="3" gridrow="3" pageId="9" pageNumber="430">0.4</td>
<td id="0B54AD59FFD242F5CE3E30EBFB4B887B" box="[1031,1158,756,778]" gridcol="4" gridrow="3" pageId="9" pageNumber="430">4.6–5.7</td>
<td id="0B54AD59FFD242F5CEE430EBFADA887B" box="[1245,1303,756,778]" gridcol="5" gridrow="3" pageId="9" pageNumber="430">5.1</td>
<td id="0B54AD59FFD242F5CF7130EBFAA5887B" box="[1352,1384,756,778]" gridcol="6" gridrow="3" pageId="9" pageNumber="430">0.3</td>
</tr>
<tr id="4885C425FFD242F5CAA63101FAA58845" box="[159,1384,798,820]" gridrow="4" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA63101FE308845" box="[159,509,798,820]" gridcol="0" gridrow="4" pageId="9" pageNumber="430">Hand length</th>
<td id="0B54AD59FFD242F5C8583101FD3D8845" box="[609,752,798,820]" gridcol="1" gridrow="4" pageId="9" pageNumber="430">7.4–9.4</td>
<td id="0B54AD59FFD242F5C90E3101FCBC8845" box="[823,881,798,820]" gridcol="2" gridrow="4" pageId="9" pageNumber="430">8.4</td>
<td id="0B54AD59FFD242F5C99B3101FC0F8845" box="[930,962,798,820]" gridcol="3" gridrow="4" pageId="9" pageNumber="430">0.5</td>
<td id="0B54AD59FFD242F5CE3E3101FB4B8845" box="[1031,1158,798,820]" gridcol="4" gridrow="4" pageId="9" pageNumber="430">5.8–7.2</td>
<td id="0B54AD59FFD242F5CEE43101FADA8845" box="[1245,1303,798,820]" gridcol="5" gridrow="4" pageId="9" pageNumber="430">6.5</td>
<td id="0B54AD59FFD242F5CF713101FAA58845" box="[1352,1384,798,820]" gridcol="6" gridrow="4" pageId="9" pageNumber="430">0.4</td>
</tr>
<tr id="4885C425FFD242F5CAA63157FAA5882F" box="[159,1384,840,862]" gridrow="5" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA63157FE30882F" box="[159,509,840,862]" gridcol="0" gridrow="5" pageId="9" pageNumber="430">Tibia length</th>
<td id="0B54AD59FFD242F5C8583157FD3D882F" box="[609,752,840,862]" gridcol="1" gridrow="5" pageId="9" pageNumber="430">11.8–13.8</td>
<td id="0B54AD59FFD242F5C90E3157FCBC882F" box="[823,881,840,862]" gridcol="2" gridrow="5" pageId="9" pageNumber="430">12.9</td>
<td id="0B54AD59FFD242F5C99B3157FC0F882F" box="[930,962,840,862]" gridcol="3" gridrow="5" pageId="9" pageNumber="430">0.6</td>
<td id="0B54AD59FFD242F5CE3E3157FB4B882F" box="[1031,1158,840,862]" gridcol="4" gridrow="5" pageId="9" pageNumber="430">9.3–11.2</td>
<td id="0B54AD59FFD242F5CEE43157FADA882F" box="[1245,1303,840,862]" gridcol="5" gridrow="5" pageId="9" pageNumber="430">10.1</td>
<td id="0B54AD59FFD242F5CF713157FAA5882F" box="[1352,1384,840,862]" gridcol="6" gridrow="5" pageId="9" pageNumber="430">0.5</td>
</tr>
<tr id="4885C425FFD242F5CAA6316DFAA588F9" box="[159,1384,882,904]" gridrow="6" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA6316DFE3088F9" box="[159,509,882,904]" gridcol="0" gridrow="6" pageId="9" pageNumber="430">Foot length</th>
<td id="0B54AD59FFD242F5C858316DFD3D88F9" box="[609,752,882,904]" gridcol="1" gridrow="6" pageId="9" pageNumber="430">11.7–14.9</td>
<td id="0B54AD59FFD242F5C90E316DFCBC88F9" box="[823,881,882,904]" gridcol="2" gridrow="6" pageId="9" pageNumber="430">13.2</td>
<td id="0B54AD59FFD242F5C99B316DFC0F88F9" box="[930,962,882,904]" gridcol="3" gridrow="6" pageId="9" pageNumber="430">0.9</td>
<td id="0B54AD59FFD242F5CE3E316DFB4B88F9" box="[1031,1158,882,904]" gridcol="4" gridrow="6" pageId="9" pageNumber="430">8.8–11.4</td>
<td id="0B54AD59FFD242F5CEE4316DFADA88F9" box="[1245,1303,882,904]" gridcol="5" gridrow="6" pageId="9" pageNumber="430">10.2</td>
<td id="0B54AD59FFD242F5CF71316DFAA588F9" box="[1352,1384,882,904]" gridcol="6" gridrow="6" pageId="9" pageNumber="430">0.8</td>
</tr>
<tr id="4885C425FFD242F5CAA63183FAA588C3" box="[159,1384,924,946]" gridrow="7" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA63183FE3088C3" box="[159,509,924,946]" gridcol="0" gridrow="7" pageId="9" pageNumber="430">Head width</th>
<td id="0B54AD59FFD242F5C8583183FD3D88C3" box="[609,752,924,946]" gridcol="1" gridrow="7" pageId="9" pageNumber="430">8.6–10.2</td>
<td id="0B54AD59FFD242F5C90E3183FCBC88C3" box="[823,881,924,946]" gridcol="2" gridrow="7" pageId="9" pageNumber="430">9.6</td>
<td id="0B54AD59FFD242F5C99B3183FC0F88C3" box="[930,962,924,946]" gridcol="3" gridrow="7" pageId="9" pageNumber="430">0.5</td>
<td id="0B54AD59FFD242F5CE3E3183FB4B88C3" box="[1031,1158,924,946]" gridcol="4" gridrow="7" pageId="9" pageNumber="430">6.8–8.6</td>
<td id="0B54AD59FFD242F5CEE43183FADA88C3" box="[1245,1303,924,946]" gridcol="5" gridrow="7" pageId="9" pageNumber="430">7.6</td>
<td id="0B54AD59FFD242F5CF713183FAA588C3" box="[1352,1384,924,946]" gridcol="6" gridrow="7" pageId="9" pageNumber="430">0.5</td>
</tr>
<tr id="4885C425FFD242F5CAA631D9FAA588AD" box="[159,1384,966,988]" gridrow="8" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA631D9FE3088AD" box="[159,509,966,988]" gridcol="0" gridrow="8" pageId="9" pageNumber="430">Head length</th>
<td id="0B54AD59FFD242F5C85831D9FD3D88AD" box="[609,752,966,988]" gridcol="1" gridrow="8" pageId="9" pageNumber="430">7.7–9.6</td>
<td id="0B54AD59FFD242F5C90E31D9FCBC88AD" box="[823,881,966,988]" gridcol="2" gridrow="8" pageId="9" pageNumber="430">8.4</td>
<td id="0B54AD59FFD242F5C99B31D9FC0F88AD" box="[930,962,966,988]" gridcol="3" gridrow="8" pageId="9" pageNumber="430">0.5</td>
<td id="0B54AD59FFD242F5CE3E31D9FB4B88AD" box="[1031,1158,966,988]" gridcol="4" gridrow="8" pageId="9" pageNumber="430">6.2–7.4</td>
<td id="0B54AD59FFD242F5CEE431D9FADA88AD" box="[1245,1303,966,988]" gridcol="5" gridrow="8" pageId="9" pageNumber="430">6.9</td>
<td id="0B54AD59FFD242F5CF7131D9FAA588AD" box="[1352,1384,966,988]" gridcol="6" gridrow="8" pageId="9" pageNumber="430">0.4</td>
</tr>
<tr id="4885C425FFD242F5CAA631EFFAA58F77" box="[159,1384,1008,1030]" gridrow="9" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA631EFFE308F77" box="[159,509,1008,1030]" gridcol="0" gridrow="9" pageId="9" pageNumber="430">Upper eyelid width</th>
<td id="0B54AD59FFD242F5C85831EFFD3D8F77" box="[609,752,1008,1030]" gridcol="1" gridrow="9" pageId="9" pageNumber="430">1.8–2.5</td>
<td id="0B54AD59FFD242F5C90E31EFFCBC8F77" box="[823,881,1008,1030]" gridcol="2" gridrow="9" pageId="9" pageNumber="430">2.1</td>
<td id="0B54AD59FFD242F5C99B31EFFC0F8F77" box="[930,962,1008,1030]" gridcol="3" gridrow="9" pageId="9" pageNumber="430">0.2</td>
<td id="0B54AD59FFD242F5CE3E31EFFB4B8F77" box="[1031,1158,1008,1030]" gridcol="4" gridrow="9" pageId="9" pageNumber="430">1.6–2.2</td>
<td id="0B54AD59FFD242F5CEE431EFFADA8F77" box="[1245,1303,1008,1030]" gridcol="5" gridrow="9" pageId="9" pageNumber="430">1.8</td>
<td id="0B54AD59FFD242F5CF7131EFFAA58F77" box="[1352,1384,1008,1030]" gridcol="6" gridrow="9" pageId="9" pageNumber="430">0.2</td>
</tr>
<tr id="4885C425FFD242F5CAA63605FAA58F41" box="[159,1384,1050,1072]" gridrow="10" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA63605FE308F41" box="[159,509,1050,1072]" gridcol="0" gridrow="10" pageId="9" pageNumber="430">Eye diameter</th>
<td id="0B54AD59FFD242F5C8583605FD3D8F41" box="[609,752,1050,1072]" gridcol="1" gridrow="10" pageId="9" pageNumber="430">2.7–3.7</td>
<td id="0B54AD59FFD242F5C90E3605FCBC8F41" box="[823,881,1050,1072]" gridcol="2" gridrow="10" pageId="9" pageNumber="430">3.2</td>
<td id="0B54AD59FFD242F5C99B3605FC0F8F41" box="[930,962,1050,1072]" gridcol="3" gridrow="10" pageId="9" pageNumber="430">0.3</td>
<td id="0B54AD59FFD242F5CE3E3605FB4B8F41" box="[1031,1158,1050,1072]" gridcol="4" gridrow="10" pageId="9" pageNumber="430">2.2–2.9</td>
<td id="0B54AD59FFD242F5CEE43605FADA8F41" box="[1245,1303,1050,1072]" gridcol="5" gridrow="10" pageId="9" pageNumber="430">2.2</td>
<td id="0B54AD59FFD242F5CF713605FAA58F41" box="[1352,1384,1050,1072]" gridcol="6" gridrow="10" pageId="9" pageNumber="430">0.2</td>
</tr>
<tr id="4885C425FFD242F5CAA6365BFAA58F2B" box="[159,1384,1092,1114]" gridrow="11" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA6365BFE308F2B" box="[159,509,1092,1114]" gridcol="0" gridrow="11" pageId="9" pageNumber="430">Eye to nostril distance</th>
<td id="0B54AD59FFD242F5C858365BFD3D8F2B" box="[609,752,1092,1114]" gridcol="1" gridrow="11" pageId="9" pageNumber="430">2–2.6</td>
<td id="0B54AD59FFD242F5C90E365BFCBC8F2B" box="[823,881,1092,1114]" gridcol="2" gridrow="11" pageId="9" pageNumber="430">2.3</td>
<td id="0B54AD59FFD242F5C99B365BFC0F8F2B" box="[930,962,1092,1114]" gridcol="3" gridrow="11" pageId="9" pageNumber="430">0.1</td>
<td id="0B54AD59FFD242F5CE3E365BFB4B8F2B" box="[1031,1158,1092,1114]" gridcol="4" gridrow="11" pageId="9" pageNumber="430">1.3–2.1</td>
<td id="0B54AD59FFD242F5CEE4365BFADA8F2B" box="[1245,1303,1092,1114]" gridcol="5" gridrow="11" pageId="9" pageNumber="430">1.2</td>
<td id="0B54AD59FFD242F5CF71365BFAA58F2B" box="[1352,1384,1092,1114]" gridcol="6" gridrow="11" pageId="9" pageNumber="430">0.2</td>
</tr>
<tr id="4885C425FFD242F5CAA63671FAA58FF5" box="[159,1384,1134,1156]" gridrow="12" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA63671FE308FF5" box="[159,509,1134,1156]" gridcol="0" gridrow="12" pageId="9" pageNumber="430">Internarial distance</th>
<td id="0B54AD59FFD242F5C8583671FD3D8FF5" box="[609,752,1134,1156]" gridcol="1" gridrow="12" pageId="9" pageNumber="430">2.2–2.8</td>
<td id="0B54AD59FFD242F5C90E3671FCBC8FF5" box="[823,881,1134,1156]" gridcol="2" gridrow="12" pageId="9" pageNumber="430">2.6</td>
<td id="0B54AD59FFD242F5C99B3671FC0F8FF5" box="[930,962,1134,1156]" gridcol="3" gridrow="12" pageId="9" pageNumber="430">0.2</td>
<td id="0B54AD59FFD242F5CE3E3671FB4B8FF5" box="[1031,1158,1134,1156]" gridcol="4" gridrow="12" pageId="9" pageNumber="430">1.8–2.3</td>
<td id="0B54AD59FFD242F5CEE43671FADA8FF5" box="[1245,1303,1134,1156]" gridcol="5" gridrow="12" pageId="9" pageNumber="430">2.1</td>
<td id="0B54AD59FFD242F5CF713671FAA58FF5" box="[1352,1384,1134,1156]" gridcol="6" gridrow="12" pageId="9" pageNumber="430">0.2</td>
</tr>
<tr id="4885C425FFD242F5CAA63687FAA58FDF" box="[159,1384,1176,1198]" gridrow="13" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA63687FE308FDF" box="[159,509,1176,1198]" gridcol="0" gridrow="13" pageId="9" pageNumber="430">Snout to eye distance</th>
<td id="0B54AD59FFD242F5C8583687FD3D8FDF" box="[609,752,1176,1198]" gridcol="1" gridrow="13" pageId="9" pageNumber="430">3.5–4.3</td>
<td id="0B54AD59FFD242F5C90E3687FCBC8FDF" box="[823,881,1176,1198]" gridcol="2" gridrow="13" pageId="9" pageNumber="430">4.0</td>
<td id="0B54AD59FFD242F5C99B3687FC0F8FDF" box="[930,962,1176,1198]" gridcol="3" gridrow="13" pageId="9" pageNumber="430">0.2</td>
<td id="0B54AD59FFD242F5CE3E3687FB4B8FDF" box="[1031,1158,1176,1198]" gridcol="4" gridrow="13" pageId="9" pageNumber="430">2.9–3.5</td>
<td id="0B54AD59FFD242F5CEE43687FADA8FDF" box="[1245,1303,1176,1198]" gridcol="5" gridrow="13" pageId="9" pageNumber="430">3.2</td>
<td id="0B54AD59FFD242F5CF713687FAA58FDF" box="[1352,1384,1176,1198]" gridcol="6" gridrow="13" pageId="9" pageNumber="430">0.2</td>
</tr>
<tr id="4885C425FFD242F5CAA636DDFAA58FA9" box="[159,1384,1218,1240]" gridrow="14" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA636DDFE308FA9" box="[159,509,1218,1240]" gridcol="0" gridrow="14" pageId="9" pageNumber="430">Interorbital distance</th>
<td id="0B54AD59FFD242F5C85836DDFD3D8FA9" box="[609,752,1218,1240]" gridcol="1" gridrow="14" pageId="9" pageNumber="430">3–3.9</td>
<td id="0B54AD59FFD242F5C90E36DDFCBC8FA9" box="[823,881,1218,1240]" gridcol="2" gridrow="14" pageId="9" pageNumber="430">3.3</td>
<td id="0B54AD59FFD242F5C99B36DDFC0F8FA9" box="[930,962,1218,1240]" gridcol="3" gridrow="14" pageId="9" pageNumber="430">0.2</td>
<td id="0B54AD59FFD242F5CE3E36DDFB4B8FA9" box="[1031,1158,1218,1240]" gridcol="4" gridrow="14" pageId="9" pageNumber="430">2.1–3</td>
<td id="0B54AD59FFD242F5CEE436DDFADA8FA9" box="[1245,1303,1218,1240]" gridcol="5" gridrow="14" pageId="9" pageNumber="430">2.6</td>
<td id="0B54AD59FFD242F5CF7136DDFAA58FA9" box="[1352,1384,1218,1240]" gridcol="6" gridrow="14" pageId="9" pageNumber="430">0.2</td>
</tr>
<tr id="4885C425FFD242F5CAA636F3FAA58E73" box="[159,1384,1260,1282]" gridrow="15" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA636F3FE308E73" box="[159,509,1260,1282]" gridcol="0" gridrow="15" pageId="9" pageNumber="430">Narinal snout length</th>
<td id="0B54AD59FFD242F5C85836F3FD3D8E73" box="[609,752,1260,1282]" gridcol="1" gridrow="15" pageId="9" pageNumber="430">1.3–1.9</td>
<td id="0B54AD59FFD242F5C90E36F3FCBC8E73" box="[823,881,1260,1282]" gridcol="2" gridrow="15" pageId="9" pageNumber="430">1.5</td>
<td id="0B54AD59FFD242F5C99B36F3FC0F8E73" box="[930,962,1260,1282]" gridcol="3" gridrow="15" pageId="9" pageNumber="430">0.1</td>
<td id="0B54AD59FFD242F5CE3E36F3FB4B8E73" box="[1031,1158,1260,1282]" gridcol="4" gridrow="15" pageId="9" pageNumber="430">1.1–1.4</td>
<td id="0B54AD59FFD242F5CEE436F3FADA8E73" box="[1245,1303,1260,1282]" gridcol="5" gridrow="15" pageId="9" pageNumber="430">1.3</td>
<td id="0B54AD59FFD242F5CF7136F3FAA58E73" box="[1352,1384,1260,1282]" gridcol="6" gridrow="15" pageId="9" pageNumber="430">0.1</td>
</tr>
<tr id="4885C425FFD242F5CAA63709FAA58E5D" box="[159,1384,1302,1324]" gridrow="16" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA63709FE308E5D" box="[159,509,1302,1324]" gridcol="0" gridrow="16" pageId="9" pageNumber="430">Eye to tympanum distance</th>
<td id="0B54AD59FFD242F5C8583709FD3D8E5D" box="[609,752,1302,1324]" gridcol="1" gridrow="16" pageId="9" pageNumber="430">0.6–1.2</td>
<td id="0B54AD59FFD242F5C90E3709FCBC8E5D" box="[823,881,1302,1324]" gridcol="2" gridrow="16" pageId="9" pageNumber="430">0.8</td>
<td id="0B54AD59FFD242F5C99B3709FC0F8E5D" box="[930,962,1302,1324]" gridcol="3" gridrow="16" pageId="9" pageNumber="430">0.2</td>
<td id="0B54AD59FFD242F5CE3E3709FB4B8E5D" box="[1031,1158,1302,1324]" gridcol="4" gridrow="16" pageId="9" pageNumber="430">0.4–0.8</td>
<td id="0B54AD59FFD242F5CEE43709FADA8E5D" box="[1245,1303,1302,1324]" gridcol="5" gridrow="16" pageId="9" pageNumber="430">0.7</td>
<td id="0B54AD59FFD242F5CF713709FAA58E5D" box="[1352,1384,1302,1324]" gridcol="6" gridrow="16" pageId="9" pageNumber="430">0.1</td>
</tr>
<tr id="4885C425FFD242F5CAA6375FFAA58E27" box="[159,1384,1344,1366]" gridrow="17" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA6375FFE308E27" box="[159,509,1344,1366]" gridcol="0" gridrow="17" pageId="9" pageNumber="430">Tympanum diameter</th>
<td id="0B54AD59FFD242F5C858375FFD3D8E27" box="[609,752,1344,1366]" gridcol="1" gridrow="17" pageId="9" pageNumber="430">1.1–1.6</td>
<td id="0B54AD59FFD242F5C90E375FFCBC8E27" box="[823,881,1344,1366]" gridcol="2" gridrow="17" pageId="9" pageNumber="430">1.4</td>
<td id="0B54AD59FFD242F5C99B375FFC0F8E27" box="[930,962,1344,1366]" gridcol="3" gridrow="17" pageId="9" pageNumber="430">0.1</td>
<td id="0B54AD59FFD242F5CE3E375FFB4B8E27" box="[1031,1158,1344,1366]" gridcol="4" gridrow="17" pageId="9" pageNumber="430">0.9–1.3</td>
<td id="0B54AD59FFD242F5CEE4375FFADA8E27" box="[1245,1303,1344,1366]" gridcol="5" gridrow="17" pageId="9" pageNumber="430">1.1</td>
<td id="0B54AD59FFD242F5CF71375FFAA58E27" box="[1352,1384,1344,1366]" gridcol="6" gridrow="17" pageId="9" pageNumber="430">0.1</td>
</tr>
<tr id="4885C425FFD242F5CAA63775FAA58EF1" box="[159,1384,1386,1408]" gridrow="18" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA63775FE308EF1" box="[159,509,1386,1408]" gridcol="0" gridrow="18" pageId="9" pageNumber="430">Finger I length</th>
<td id="0B54AD59FFD242F5C8583775FD3D8EF1" box="[609,752,1386,1408]" gridcol="1" gridrow="18" pageId="9" pageNumber="430">1.8–2.6</td>
<td id="0B54AD59FFD242F5C90E3775FCBC8EF1" box="[823,881,1386,1408]" gridcol="2" gridrow="18" pageId="9" pageNumber="430">2.2</td>
<td id="0B54AD59FFD242F5C99B3775FC0F8EF1" box="[930,962,1386,1408]" gridcol="3" gridrow="18" pageId="9" pageNumber="430">0.2</td>
<td id="0B54AD59FFD242F5CE3E3775FB4B8EF1" box="[1031,1158,1386,1408]" gridcol="4" gridrow="18" pageId="9" pageNumber="430">1.3–1.9</td>
<td id="0B54AD59FFD242F5CEE43775FADA8EF1" box="[1245,1303,1386,1408]" gridcol="5" gridrow="18" pageId="9" pageNumber="430">1.6</td>
<td id="0B54AD59FFD242F5CF713775FAA58EF1" box="[1352,1384,1386,1408]" gridcol="6" gridrow="18" pageId="9" pageNumber="430">0.2</td>
</tr>
<tr id="4885C425FFD242F5CAA6378BFAA58EDB" box="[159,1384,1428,1450]" gridrow="19" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA6378BFE308EDB" box="[159,509,1428,1450]" gridcol="0" gridrow="19" pageId="9" pageNumber="430">Finger II length</th>
<td id="0B54AD59FFD242F5C858378BFD3D8EDB" box="[609,752,1428,1450]" gridcol="1" gridrow="19" pageId="9" pageNumber="430">2.5–3.5</td>
<td id="0B54AD59FFD242F5C90E378BFCBC8EDB" box="[823,881,1428,1450]" gridcol="2" gridrow="19" pageId="9" pageNumber="430">2.9</td>
<td id="0B54AD59FFD242F5C99B378BFC0F8EDB" box="[930,962,1428,1450]" gridcol="3" gridrow="19" pageId="9" pageNumber="430">0.3</td>
<td id="0B54AD59FFD242F5CE3E378BFB4B8EDB" box="[1031,1158,1428,1450]" gridcol="4" gridrow="19" pageId="9" pageNumber="430">1.9–2.6</td>
<td id="0B54AD59FFD242F5CEE4378BFADA8EDB" box="[1245,1303,1428,1450]" gridcol="5" gridrow="19" pageId="9" pageNumber="430">2.3</td>
<td id="0B54AD59FFD242F5CF71378BFAA58EDB" box="[1352,1384,1428,1450]" gridcol="6" gridrow="19" pageId="9" pageNumber="430">0.2</td>
</tr>
<tr id="4885C425FFD242F5CAA637A1FAA58EA5" box="[159,1384,1470,1492]" gridrow="20" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA637A1FE308EA5" box="[159,509,1470,1492]" gridcol="0" gridrow="20" pageId="9" pageNumber="430">Toe III length</th>
<td id="0B54AD59FFD242F5C85837A1FD3D8EA5" box="[609,752,1470,1492]" gridcol="1" gridrow="20" pageId="9" pageNumber="430">3.9–5.1</td>
<td id="0B54AD59FFD242F5C90E37A1FCBC8EA5" box="[823,881,1470,1492]" gridcol="2" gridrow="20" pageId="9" pageNumber="430">4.4</td>
<td id="0B54AD59FFD242F5C99B37A1FC0F8EA5" box="[930,962,1470,1492]" gridcol="3" gridrow="20" pageId="9" pageNumber="430">0.3</td>
<td id="0B54AD59FFD242F5CE3E37A1FB4B8EA5" box="[1031,1158,1470,1492]" gridcol="4" gridrow="20" pageId="9" pageNumber="430">–</td>
<td id="0B54AD59FFD242F5CEE437A1FADA8EA5" box="[1245,1303,1470,1492]" gridcol="5" gridrow="20" pageId="9" pageNumber="430">–</td>
<td id="0B54AD59FFD242F5CF7137A1FAA58EA5" box="[1352,1384,1470,1492]" gridcol="6" gridrow="20" pageId="9" pageNumber="430">–</td>
</tr>
<tr id="4885C425FFD242F5CAA637F7FAA58E8F" box="[159,1384,1512,1534]" gridrow="21" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA637F7FE308E8F" box="[159,509,1512,1534]" gridcol="0" gridrow="21" pageId="9" pageNumber="430">Toe IV length</th>
<td id="0B54AD59FFD242F5C85837F7FD3D8E8F" box="[609,752,1512,1534]" gridcol="1" gridrow="21" pageId="9" pageNumber="430">6.5–8</td>
<td id="0B54AD59FFD242F5C90E37F7FCBC8E8F" box="[823,881,1512,1534]" gridcol="2" gridrow="21" pageId="9" pageNumber="430">7.3</td>
<td id="0B54AD59FFD242F5C99B37F7FC0F8E8F" box="[930,962,1512,1534]" gridcol="3" gridrow="21" pageId="9" pageNumber="430">0.6</td>
<td id="0B54AD59FFD242F5CE3E37F7FB4B8E8F" box="[1031,1158,1512,1534]" gridcol="4" gridrow="21" pageId="9" pageNumber="430">–</td>
<td id="0B54AD59FFD242F5CEE437F7FADA8E8F" box="[1245,1303,1512,1534]" gridcol="5" gridrow="21" pageId="9" pageNumber="430">–</td>
<td id="0B54AD59FFD242F5CF7137F7FAA58E8F" box="[1352,1384,1512,1534]" gridcol="6" gridrow="21" pageId="9" pageNumber="430">–</td>
</tr>
<tr id="4885C425FFD242F5CAA6340DFAA58D59" box="[159,1384,1554,1576]" gridrow="22" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA6340DFE308D59" box="[159,509,1554,1576]" gridcol="0" gridrow="22" pageId="9" pageNumber="430">Tibia length/SVL</th>
<td id="0B54AD59FFD242F5C858340DFD3D8D59" box="[609,752,1554,1576]" gridcol="1" gridrow="22" pageId="9" pageNumber="430">47.6–55.5</td>
<td id="0B54AD59FFD242F5C90E340DFCBC8D59" box="[823,881,1554,1576]" gridcol="2" gridrow="22" pageId="9" pageNumber="430">50.8</td>
<td id="0B54AD59FFD242F5C99B340DFC0F8D59" box="[930,962,1554,1576]" gridcol="3" gridrow="22" pageId="9" pageNumber="430">2.1</td>
<td id="0B54AD59FFD242F5CE3E340DFB4B8D59" box="[1031,1158,1554,1576]" gridcol="4" gridrow="22" pageId="9" pageNumber="430">46.9–54.5</td>
<td id="0B54AD59FFD242F5CEE4340DFADA8D59" box="[1245,1303,1554,1576]" gridcol="5" gridrow="22" pageId="9" pageNumber="430">51.1</td>
<td id="0B54AD59FFD242F5CF71340DFAA58D59" box="[1352,1384,1554,1576]" gridcol="6" gridrow="22" pageId="9" pageNumber="430">2.7</td>
</tr>
<tr id="4885C425FFD242F5CAA63423FAA58D23" box="[159,1384,1596,1618]" gridrow="23" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA63423FE308D23" box="[159,509,1596,1618]" gridcol="0" gridrow="23" pageId="9" pageNumber="430">Head width/SVL</th>
<td id="0B54AD59FFD242F5C8583423FD3D8D23" box="[609,752,1596,1618]" gridcol="1" gridrow="23" pageId="9" pageNumber="430">35.1–41,9</td>
<td id="0B54AD59FFD242F5C90E3423FCBC8D23" box="[823,881,1596,1618]" gridcol="2" gridrow="23" pageId="9" pageNumber="430">37.9</td>
<td id="0B54AD59FFD242F5C99B3423FC0F8D23" box="[930,962,1596,1618]" gridcol="3" gridrow="23" pageId="9" pageNumber="430">1.4</td>
<td id="0B54AD59FFD242F5CE3E3423FB4B8D23" box="[1031,1158,1596,1618]" gridcol="4" gridrow="23" pageId="9" pageNumber="430">36.1–40.7</td>
<td id="0B54AD59FFD242F5CEE43423FADA8D23" box="[1245,1303,1596,1618]" gridcol="5" gridrow="23" pageId="9" pageNumber="430">38.5</td>
<td id="0B54AD59FFD242F5CF713423FAA58D23" box="[1352,1384,1596,1618]" gridcol="6" gridrow="23" pageId="9" pageNumber="430">2.1</td>
</tr>
<tr id="4885C425FFD242F5CAA63479FAA58D0D" box="[159,1384,1638,1660]" gridrow="24" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA63479FE308D0D" box="[159,509,1638,1660]" gridcol="0" gridrow="24" pageId="9" pageNumber="430">Tympanum diameter/Eye diameter</th>
<td id="0B54AD59FFD242F5C8583479FD3D8D0D" box="[609,752,1638,1660]" gridcol="1" gridrow="24" pageId="9" pageNumber="430">31.0–47.7</td>
<td id="0B54AD59FFD242F5C90E3479FCBC8D0D" box="[823,881,1638,1660]" gridcol="2" gridrow="24" pageId="9" pageNumber="430">46.7</td>
<td id="0B54AD59FFD242F5C99B3479FC0F8D0D" box="[930,962,1638,1660]" gridcol="3" gridrow="24" pageId="9" pageNumber="430">5.4</td>
<td id="0B54AD59FFD242F5CE3E3479FB4B8D0D" box="[1031,1158,1638,1660]" gridcol="4" gridrow="24" pageId="9" pageNumber="430">35.9–56.2</td>
<td id="0B54AD59FFD242F5CEE43479FADA8D0D" box="[1245,1303,1638,1660]" gridcol="5" gridrow="24" pageId="9" pageNumber="430">41.6</td>
<td id="0B54AD59FFD242F5CF713479FAA58D0D" box="[1352,1384,1638,1660]" gridcol="6" gridrow="24" pageId="9" pageNumber="430">5.2</td>
</tr>
<tr id="4885C425FFD242F5CAA6348FFAA58DD7" box="[159,1384,1680,1702]" gridrow="25" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA6348FFE308DD7" box="[159,509,1680,1702]" gridcol="0" gridrow="25" pageId="9" pageNumber="430">Foot length/SVL</th>
<td id="0B54AD59FFD242F5C858348FFD3D8DD7" box="[609,752,1680,1702]" gridcol="1" gridrow="25" pageId="9" pageNumber="430">46.0–57.5</td>
<td id="0B54AD59FFD242F5C90E348FFCBC8DD7" box="[823,881,1680,1702]" gridcol="2" gridrow="25" pageId="9" pageNumber="430">51.1</td>
<td id="0B54AD59FFD242F5C99B348FFC0F8DD7" box="[930,962,1680,1702]" gridcol="3" gridrow="25" pageId="9" pageNumber="430">2.2</td>
<td id="0B54AD59FFD242F5CE3E348FFB4B8DD7" box="[1031,1158,1680,1702]" gridcol="4" gridrow="25" pageId="9" pageNumber="430">47.4–53.4</td>
<td id="0B54AD59FFD242F5CEE4348FFADA8DD7" box="[1245,1303,1680,1702]" gridcol="5" gridrow="25" pageId="9" pageNumber="430">51.2</td>
<td id="0B54AD59FFD242F5CF71348FFAA58DD7" box="[1352,1384,1680,1702]" gridcol="6" gridrow="25" pageId="9" pageNumber="430">3.1</td>
</tr>
<tr id="4885C425FFD242F5CAA634A5FAA58DA1" box="[159,1384,1722,1744]" gridrow="26" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA634A5FE308DA1" box="[159,509,1722,1744]" gridcol="0" gridrow="26" pageId="9" pageNumber="430">Head width/head length</th>
<td id="0B54AD59FFD242F5C85834A5FD3D8DA1" box="[609,752,1722,1744]" gridcol="1" gridrow="26" pageId="9" pageNumber="430">102.7–123.4</td>
<td id="0B54AD59FFD242F5C90E34A5FCBC8DA1" box="[823,881,1722,1744]" gridcol="2" gridrow="26" pageId="9" pageNumber="430">114.3</td>
<td id="0B54AD59FFD242F5C99B34A5FC0F8DA1" box="[930,962,1722,1744]" gridcol="3" gridrow="26" pageId="9" pageNumber="430">4.4</td>
<td id="0B54AD59FFD242F5CE3E34A5FB4B8DA1" box="[1031,1158,1722,1744]" gridcol="4" gridrow="26" pageId="9" pageNumber="430">105.6–124.2</td>
<td id="0B54AD59FFD242F5CEE434A5FADA8DA1" box="[1245,1303,1722,1744]" gridcol="5" gridrow="26" pageId="9" pageNumber="430">111.1</td>
<td id="0B54AD59FFD242F5CF7134A5FAA58DA1" box="[1352,1384,1722,1744]" gridcol="6" gridrow="26" pageId="9" pageNumber="430">4.9</td>
</tr>
<tr id="4885C425FFD242F5CAA634FBFAA58D8B" box="[159,1384,1764,1786]" gridrow="27" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA634FBFE308D8B" box="[159,509,1764,1786]" gridcol="0" gridrow="27" pageId="9" pageNumber="430">Eye to nostril distance/HW</th>
<td id="0B54AD59FFD242F5C85834FBFD3D8D8B" box="[609,752,1764,1786]" gridcol="1" gridrow="27" pageId="9" pageNumber="430">19.7–25.3</td>
<td id="0B54AD59FFD242F5C90E34FBFCBC8D8B" box="[823,881,1764,1786]" gridcol="2" gridrow="27" pageId="9" pageNumber="430">24.0</td>
<td id="0B54AD59FFD242F5C99B34FBFC0F8D8B" box="[930,962,1764,1786]" gridcol="3" gridrow="27" pageId="9" pageNumber="430">1.8</td>
<td id="0B54AD59FFD242F5CE3E34FBFB4B8D8B" box="[1031,1158,1764,1786]" gridcol="4" gridrow="27" pageId="9" pageNumber="430">21.7–26.9</td>
<td id="0B54AD59FFD242F5CEE434FBFADA8D8B" box="[1245,1303,1764,1786]" gridcol="5" gridrow="27" pageId="9" pageNumber="430">23.1</td>
<td id="0B54AD59FFD242F5CF7134FBFAA58D8B" box="[1352,1384,1764,1786]" gridcol="6" gridrow="27" pageId="9" pageNumber="430">2.0</td>
</tr>
<tr id="4885C425FFD242F5CAA63511FAA58C55" box="[159,1384,1806,1828]" gridrow="28" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA63511FE308C55" box="[159,509,1806,1828]" gridcol="0" gridrow="28" pageId="9" pageNumber="430">Radioulna length/SVL</th>
<td id="0B54AD59FFD242F5C8583511FD3D8C55" box="[609,752,1806,1828]" gridcol="1" gridrow="28" pageId="9" pageNumber="430">23.6–28.7</td>
<td id="0B54AD59FFD242F5C90E3511FCBC8C55" box="[823,881,1806,1828]" gridcol="2" gridrow="28" pageId="9" pageNumber="430">25.9</td>
<td id="0B54AD59FFD242F5C99B3511FC0F8C55" box="[930,962,1806,1828]" gridcol="3" gridrow="28" pageId="9" pageNumber="430">1.2</td>
<td id="0B54AD59FFD242F5CE3E3511FB4B8C55" box="[1031,1158,1806,1828]" gridcol="4" gridrow="28" pageId="9" pageNumber="430">23.5–27.8</td>
<td id="0B54AD59FFD242F5CEE43511FADA8C55" box="[1245,1303,1806,1828]" gridcol="5" gridrow="28" pageId="9" pageNumber="430">25.7</td>
<td id="0B54AD59FFD242F5CF713511FAA58C55" box="[1352,1384,1806,1828]" gridcol="6" gridrow="28" pageId="9" pageNumber="430">1.4</td>
</tr>
<tr id="4885C425FFD242F5CAA63527FAA58C3F" box="[159,1384,1848,1870]" gridrow="29" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA63527FE308C3F" box="[159,509,1848,1870]" gridcol="0" gridrow="29" pageId="9" pageNumber="430">Interorbital distance /HW</th>
<td id="0B54AD59FFD242F5C8583527FD3D8C3F" box="[609,752,1848,1870]" gridcol="1" gridrow="29" pageId="9" pageNumber="430">25.7–38.9</td>
<td id="0B54AD59FFD242F5C90E3527FCBC8C3F" box="[823,881,1848,1870]" gridcol="2" gridrow="29" pageId="9" pageNumber="430">34.7</td>
<td id="0B54AD59FFD242F5C99B3527FC0F8C3F" box="[930,962,1848,1870]" gridcol="3" gridrow="29" pageId="9" pageNumber="430">2.1</td>
<td id="0B54AD59FFD242F5CE3E3527FB4B8C3F" box="[1031,1158,1848,1870]" gridcol="4" gridrow="29" pageId="9" pageNumber="430">30.7–37.6</td>
<td id="0B54AD59FFD242F5CEE43527FADA8C3F" box="[1245,1303,1848,1870]" gridcol="5" gridrow="29" pageId="9" pageNumber="430">33.8</td>
<td id="0B54AD59FFD242F5CF713527FAA58C3F" box="[1352,1384,1848,1870]" gridcol="6" gridrow="29" pageId="9" pageNumber="430">2.9</td>
</tr>
<tr id="4885C425FFD242F5CAA6357DFAA58C09" box="[159,1384,1890,1912]" gridrow="30" pageId="9" pageNumber="430">
<th id="0B54AD59FFD242F5CAA6357DFE308C09" box="[159,509,1890,1912]" gridcol="0" gridrow="30" pageId="9" pageNumber="430">Finger I length/Finger II length</th>
<td id="0B54AD59FFD242F5C858357DFD3D8C09" box="[609,752,1890,1912]" gridcol="1" gridrow="30" pageId="9" pageNumber="430">60.8–82.6</td>
<td id="0B54AD59FFD242F5C90E357DFCBC8C09" box="[823,881,1890,1912]" gridcol="2" gridrow="30" pageId="9" pageNumber="430">74.5</td>
<td id="0B54AD59FFD242F5C99B357DFC0F8C09" box="[930,962,1890,1912]" gridcol="3" gridrow="30" pageId="9" pageNumber="430">5.2</td>
<td id="0B54AD59FFD242F5CE3E357DFB4B8C09" box="[1031,1158,1890,1912]" gridcol="4" gridrow="30" pageId="9" pageNumber="430">64.3–82.1</td>
<td id="0B54AD59FFD242F5CEE4357DFADA8C09" box="[1245,1303,1890,1912]" gridcol="5" gridrow="30" pageId="9" pageNumber="430">70.7</td>
<td id="0B54AD59FFD242F5CF71357DFAA58C09" box="[1352,1384,1890,1912]" gridcol="6" gridrow="30" pageId="9" pageNumber="430">5.4</td>
</tr>
</table>
</paragraph>
<caption id="A2CA96EFFFD1BD00CAAE3419FF3F8DE7" httpUri="https://zenodo.org/record/258323/files/figure.png" pageId="10" pageNumber="431" targetBox="[319,1268,216,1504]" targetPageId="10">
<paragraph id="F60AC667FFD1BD00CAAE3419FF3F8DE7" blockId="10.[151,1436,1542,1686]" pageId="10" pageNumber="431">
<emphasis id="C4C11A75FFD1BD00CAAE3419FEDF8D6A" bold="true" box="[151,274,1542,1564]" pageId="10" pageNumber="431">FIGURE 6.</emphasis>
Map of Colombia showing the distribution of
<taxonomicName id="31B5BDE4FFD1BD00C8D03418FC668D6D" box="[745,939,1543,1564]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="431" phylum="Chordata" rank="species" species="stictus" status="sp. nov.">
<emphasis id="C4C11A75FFD1BD00C8D03418FC668D6D" box="[745,939,1543,1564]" italics="true" pageId="10" pageNumber="431">Pristimantis stictus</emphasis>
</taxonomicName>
<emphasis id="C4C11A75FFD1BD00C9883419FBCC8D6A" bold="true" box="[945,1025,1542,1563]" pageId="10" pageNumber="431">
<taxonomicNameLabel id="DFF2A70EFFD1BD00C9883419FBCC8D6A" box="[945,1025,1542,1563]" pageId="10" pageNumber="431" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and other species. Black circles indicate localities of
<taxonomicName id="31B5BDE4FFD1BD00CB2E3439FEBE8D4A" box="[279,371,1573,1595]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="431" phylum="Chordata" rank="species" species="stictus" status="sp. nov.">
<emphasis id="C4C11A75FFD1BD00CB2E3439FEE98D4A" box="[279,292,1574,1595]" italics="true" pageId="10" pageNumber="431">P</emphasis>
.
<emphasis id="C4C11A75FFD1BD00CB0B3439FEBE8D4A" box="[306,371,1574,1595]" italics="true" pageId="10" pageNumber="431">stictus</emphasis>
</taxonomicName>
<emphasis id="C4C11A75FFD1BD00CB42343AFE008D4B" bold="true" box="[379,461,1573,1594]" pageId="10" pageNumber="431">
<taxonomicNameLabel id="DFF2A70EFFD1BD00CB42343AFE008D4B" box="[379,461,1573,1594]" pageId="10" pageNumber="431" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and a white circle signals the type locality. Distribution of other species in the
<taxonomicName id="31B5BDE4FFD1BD00CF323439FA518D4B" box="[1291,1436,1572,1595]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="431" phylum="Chordata" rank="species" species="leptolophus">
<emphasis id="C4C11A75FFD1BD00CF323439FADA8D4A" box="[1291,1303,1574,1595]" italics="true" pageId="10" pageNumber="431">P</emphasis>
.
<emphasis id="C4C11A75FFD1BD00CF1F343BFA518D4B" box="[1318,1436,1572,1594]" italics="true" pageId="10" pageNumber="431">leptolophus</emphasis>
</taxonomicName>
group:
<taxonomicName id="31B5BDE4FFD1BD00CAD9345BFEA88D28" box="[224,357,1603,1625]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="431" phylum="Chordata" rank="species" species="maculosus">
<emphasis id="C4C11A75FFD1BD00CAD9345BFF218D28" box="[224,236,1604,1625]" italics="true" pageId="10" pageNumber="431">P</emphasis>
.
<emphasis id="C4C11A75FFD1BD00CAC3345CFEA88D28" box="[250,357,1603,1625]" italics="true" pageId="10" pageNumber="431">maculosus</emphasis>
</taxonomicName>
(black five pointed star),
<taxonomicName id="31B5BDE4FFD1BD00C857345BFD3E8D28" box="[622,755,1604,1625]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="431" phylum="Chordata" rank="species" species="parectatus">
<emphasis id="C4C11A75FFD1BD00C857345BFDB68D28" box="[622,635,1604,1625]" italics="true" pageId="10" pageNumber="431">P</emphasis>
.
<emphasis id="C4C11A75FFD1BD00C8B0345BFD3E8D28" box="[649,755,1604,1625]" italics="true" pageId="10" pageNumber="431">parectatus</emphasis>
</taxonomicName>
(white diamond),
<taxonomicName id="31B5BDE4FFD1BD00C989345BFB8E8D28" box="[944,1091,1603,1625]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="431" phylum="Chordata" rank="species" species="lasalleorum">
<emphasis id="C4C11A75FFD1BD00C989345BFC718D28" box="[944,956,1604,1625]" italics="true" pageId="10" pageNumber="431">P</emphasis>
.
<emphasis id="C4C11A75FFD1BD00C9F3345CFB8E8D28" box="[970,1091,1603,1625]" italics="true" pageId="10" pageNumber="431">lasalleorum</emphasis>
</taxonomicName>
(inverted triangle),
<taxonomicName id="31B5BDE4FFD1BD00CF29345BFA518D28" box="[1296,1436,1603,1625]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="431" phylum="Chordata" rank="species" species="uranobates">
<emphasis id="C4C11A75FFD1BD00CF29345BFAD08D28" box="[1296,1309,1604,1625]" italics="true" pageId="10" pageNumber="431">P</emphasis>
.
<emphasis id="C4C11A75FFD1BD00CF13345CFA518D28" box="[1322,1436,1603,1625]" italics="true" pageId="10" pageNumber="431">uranobates</emphasis>
</taxonomicName>
(white four pointed star),
<taxonomicName id="31B5BDE4FFD1BD00CB9D347CFDD38D09" box="[420,542,1635,1656]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="431" phylum="Chordata" rank="species" species="peraticus">
<emphasis id="C4C11A75FFD1BD00CB9D347CFE7C8D09" box="[420,433,1635,1656]" italics="true" pageId="10" pageNumber="431">P</emphasis>
.
<emphasis id="C4C11A75FFD1BD00CBF9347CFDD38D09" box="[448,542,1635,1656]" italics="true" pageId="10" pageNumber="431">peraticus</emphasis>
</taxonomicName>
(black six pointed star),
<taxonomicName id="31B5BDE4FFD1BD00C91F347CFC108D09" box="[806,989,1634,1656]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="431" phylum="Chordata" rank="species" species="scoloblepharus">
<emphasis id="C4C11A75FFD1BD00C91F347CFCFE8D09" box="[806,819,1635,1656]" italics="true" pageId="10" pageNumber="431">P</emphasis>
.
<emphasis id="C4C11A75FFD1BD00C97B347DFC108D09" box="[834,989,1634,1656]" italics="true" pageId="10" pageNumber="431">scoloblepharus</emphasis>
</taxonomicName>
(black triangle), and
<taxonomicName id="31B5BDE4FFD1BD00CEF9347CFA9F8D09" box="[1216,1362,1634,1656]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="431" phylum="Chordata" rank="species" species="leptolophus">
<emphasis id="C4C11A75FFD1BD00CEF9347CFB008D09" box="[1216,1229,1635,1656]" italics="true" pageId="10" pageNumber="431">P</emphasis>
.
<emphasis id="C4C11A75FFD1BD00CEE5347DFA9F8D09" box="[1244,1362,1634,1656]" italics="true" pageId="10" pageNumber="431">leptolophus</emphasis>
</taxonomicName>
(white squares).
</paragraph>
</caption>
<subSubSection id="BEAF95ECFFD1BD01CAFE34A2FAED888A" lastPageId="11" lastPageNumber="432" pageId="10" pageNumber="431" type="distribution">
<paragraph id="F60AC667FFD1BD00CAFE34A2FB7D8CDF" blockId="10.[151,1436,1725,1967]" pageId="10" pageNumber="431">
<emphasis id="C4C11A75FFD1BD00CAFE34A2FD9D8DA7" bold="true" box="[199,592,1725,1750]" pageId="10" pageNumber="431">Distribution and natural history.</emphasis>
The new species is found at altitudes above
<quantity id="314D6B82FFD1BD00CE7334A2FB6F8DA7" box="[1098,1186,1725,1750]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="10" pageNumber="431" unit="m" value="3500.0">3500 m</quantity>
, reaching a maximum altitude of 3700 meters. At altitudes of
<quantity id="314D6B82FFD1BD00C87634FDFCCF8D8B" box="[591,770,1762,1787]" metricMagnitude="3" metricUnit="m" metricValue="3.55" metricValueMax="3.6" metricValueMin="3.5" pageId="10" pageNumber="431" unit="m" value="3550.0" valueMax="3600.0" valueMin="3500.0">3500 to 3600 m</quantity>
,
<taxonomicName id="31B5BDE4FFD1BD00C93434FCFCBC8D8B" box="[781,881,1763,1786]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="431" phylum="Chordata" rank="species" species="stictus">
<emphasis id="C4C11A75FFD1BD00C93434FCFCBC8D8B" box="[781,881,1763,1786]" italics="true" pageId="10" pageNumber="431">P. stictus</emphasis>
</taxonomicName>
is parapatric with
<taxonomicName id="31B5BDE4FFD1BD00CE7834FCFB178D8B" box="[1089,1242,1762,1786]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="431" phylum="Chordata" rank="species" species="uranobates">
<emphasis id="C4C11A75FFD1BD00CE7834FCFB178D8B" box="[1089,1242,1762,1786]" italics="true" pageId="10" pageNumber="431">P. uranobates</emphasis>
</taxonomicName>
. So far,
<taxonomicName id="31B5BDE4FFD1BD00CF0034FCFA518D8B" box="[1337,1436,1763,1786]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="431" phylum="Chordata" rank="species" species="stictus">
<emphasis id="C4C11A75FFD1BD00CF0034FCFA518D8B" box="[1337,1436,1763,1786]" italics="true" pageId="10" pageNumber="431">P. stictus</emphasis>
</taxonomicName>
is endemic to the department of Caldas, in the northern region of Páramo Los Nevados (
<figureCitation id="6E8EDAE2FFD1BD00CE9D351AFAC48C6F" box="[1188,1289,1797,1822]" captionStart="FIGURE 6" captionStartId="10.[151,250,1542,1564]" captionTargetBox="[319,1268,216,1504]" captionTargetId="figure@10.[302,1284,193,1520]" captionTargetPageId="10" captionText="FIGURE 6. Map of Colombia showing the distribution of Pristimantis stictus sp. nov. and other species. Black circles indicate localities of P. stictus sp. nov., and a white circle signals the type locality. Distribution of other species in the P. leptolophus group: P. maculosus (black five pointed star), P. parectatus (white diamond), P. lasalleorum (inverted triangle), P. uranobates (white four pointed star), P. peraticus (black six pointed star), P. scoloblepharus (black triangle), and P. leptolophus (white squares)." httpUri="https://zenodo.org/record/258323/files/figure.png" pageId="10" pageNumber="431">Figure 6</figureCitation>
), being very abundant throughout its distribution range. It inhabits high altitude vegetation of the high Andean forest, and Subparamo (
<bibRefCitation id="9224BB96FFD1BD00CB123552FDFA8C17" author="Rodriguez" box="[299,567,1869,1894]" pageId="10" pageNumber="437" refString="Rodriguez, N., Armenteras, D., Morales, M &amp; Romero M. (2006) Ecosistemas de los Andes Colombianos. Segunda edicion. Instituto de Investigacion de Recursos Biologicos, Alexander von Humboldt, Bogota, Colombia. 154 pp." type="book" year="2006">
Rodríguez
<emphasis id="C4C11A75FFD1BD00CB973550FE288C17" box="[430,485,1870,1894]" italics="true" pageId="10" pageNumber="431">et al</emphasis>
. 2006
</bibRefCitation>
).
<taxonomicName id="31B5BDE4FFD1BD00C8763550FCE18C17" box="[591,812,1871,1894]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="431" phylum="Chordata" rank="species" species="stictus">
<emphasis id="C4C11A75FFD1BD00C8763550FCE18C17" box="[591,812,1871,1894]" italics="true" pageId="10" pageNumber="431">Pristimantis stictus</emphasis>
</taxonomicName>
was found actively at night mainly associated with terrestrial bromeliads. Males called at heights lower than
<quantity id="314D6B82FFD1BD00C919356DFCA78CFB" box="[800,874,1906,1930]" metricMagnitude="-1" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="431" unit="cm" value="70.0">70 cm</quantity>
. Amplectant pairs and gravid females were found on
<date id="820BE0A7FFD1BD00CA833588FE9D8CDF" box="[186,336,1942,1967]" pageId="10" pageNumber="431" value="2010-01">January 2010</date>
and
<date id="820BE0A7FFD1BD00CBBF358AFDC98CDF" box="[390,516,1941,1966]" pageId="10" pageNumber="431" value="2014-04">April 2014</date>
. Juveniles were found on
<date id="820BE0A7FFD1BD00C91E3588FC708CDF" box="[807,957,1942,1967]" pageId="10" pageNumber="431" value="2009-01">January 2009</date>
and
<date id="820BE0A7FFD1BD00C9CA358AFB618CDF" box="[1011,1196,1941,1966]" pageId="10" pageNumber="431" value="2012-11">November 2012</date>
.
</paragraph>
<paragraph id="F60AC667FFD0BD01CAFE3288FE5A8AFA" blockId="11.[151,1436,151,395]" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CAFE3288FD588BC1" bold="true" box="[199,661,151,176]" pageId="11" pageNumber="432">Morphometry between similar species.</emphasis>
Discriminant analysis allowed to differentiate species. Variables that showed significant differences between means are listed in
<tableCitation id="BB37F3DCFFD0BD01C90932A3FC498BA5" box="[816,900,188,213]" captionStart="TABLE 3" captionStartId="11.[151,239,438,461]" captionTargetBox="[159,1378,553,937]" captionTargetPageId="11" captionText="TABLE 3. Variables in the model for the Discriminant Function Analysis summary and standardized coefficients for the two roots. Wilks’ Lambda: 0.02578, F (44,72) = 8.5544, P &lt;0.0000. Abbreviations: Eigenvalue (Eig), cumulative proportion (Cp)." httpUri="http://table.plazi.org/id/A2CA96EFFFD0BD01CAAE33A9FE8B897C" pageId="11" pageNumber="432" tableUuid="A2CA96EFFFD0BD01CAAE33A9FE8B897C">Table 3</tableCitation>
. The variables indicating a higher proportion of variance were toe III length, snout-eye distance, snout-vent length, Toe IV length, and eye-to-nostril distance (
<tableCitation id="BB37F3DCFFD0BD01CAA7331BFF3B8A6D" box="[158,246,260,285]" captionStart="TABLE 3" captionStartId="11.[151,239,438,461]" captionTargetBox="[159,1378,553,937]" captionTargetPageId="11" captionText="TABLE 3. Variables in the model for the Discriminant Function Analysis summary and standardized coefficients for the two roots. Wilks’ Lambda: 0.02578, F (44,72) = 8.5544, P &lt;0.0000. Abbreviations: Eigenvalue (Eig), cumulative proportion (Cp)." httpUri="http://table.plazi.org/id/A2CA96EFFFD0BD01CAAE33A9FE8B897C" pageId="11" pageNumber="432" tableUuid="A2CA96EFFFD0BD01CAAE33A9FE8B897C">Table 3</tableCitation>
). The Chi-square Test with successive roots removal showed significant association between the species and canonical functions for the first (R= 0.95;
<date id="820BE0A7FFD0BD01C89E3334FD0E8A44" box="[679,707,295,322]" pageId="11" pageNumber="432">X2</date>
= 170.09; df= 44 p &lt;0.00) and second root (R= 0.85;
<date id="820BE0A7FFD0BD01CF1B3334FAF38A44" box="[1314,1342,295,322]" pageId="11" pageNumber="432">X2</date>
= 57.87; df= 21 p &lt;0.00). Such variation allowed the recognition of three distinct groups corresponding to the target species (
<tableCitation id="BB37F3DCFFD0BD01CAA7336DFF3E8AFA" box="[158,243,370,395]" captionStart="TABLE 3" captionStartId="11.[151,239,438,461]" captionTargetBox="[159,1378,553,937]" captionTargetPageId="11" captionText="TABLE 3. Variables in the model for the Discriminant Function Analysis summary and standardized coefficients for the two roots. Wilks’ Lambda: 0.02578, F (44,72) = 8.5544, P &lt;0.0000. Abbreviations: Eigenvalue (Eig), cumulative proportion (Cp)." httpUri="http://table.plazi.org/id/A2CA96EFFFD0BD01CAAE33A9FE8B897C" pageId="11" pageNumber="432" tableUuid="A2CA96EFFFD0BD01CAAE33A9FE8B897C">Table 3</tableCitation>
and
<figureCitation id="6E8EDAE2FFD0BD01CB10336DFE448AFA" box="[297,393,370,395]" captionStart="FIGURE 7" captionStartId="12.[151,250,1899,1921]" captionTargetBox="[248,1297,679,1860]" captionTargetId="figure@12.[226,1360,662,1877]" captionTargetPageId="12" captionText="FIGURE 7. Graphic representation of the two roots of the discriminant analysis for morphometrics." httpUri="https://zenodo.org/record/258324/files/figure.png" pageId="11" pageNumber="432">Figure 7</figureCitation>
).
</paragraph>
<caption id="A2CA96EFFFD0BD01CAAE33A9FE8B897C" ID-Table-UUID="A2CA96EFFFD0BD01CAAE33A9FE8B897C" httpUri="http://table.plazi.org/id/A2CA96EFFFD0BD01CAAE33A9FE8B897C" pageId="11" pageNumber="432" targetBox="[159,1378,553,937]" targetIsTable="true" targetPageId="11">
<paragraph id="F60AC667FFD0BD01CAAE33A9FE8B897C" blockId="11.[151,1436,438,525]" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CAAE33A9FEC48ABC" bold="true" box="[151,265,438,461]" pageId="11" pageNumber="432">TABLE 3.</emphasis>
Variables in the model for the Discriminant Function Analysis summary and standardized coefficients for the two roots. Wilks’ Lambda: 0.02578, F(44,72)=8.5544, P&lt;0.0000. Abbreviations: Eigenvalue (Eig), cumulative proportion (Cp).
</paragraph>
</caption>
<paragraph id="F60AC667FFD0BD01CA9C3036FA9488D8" pageId="11" pageNumber="432">
<table id="84B534C7FFD042F5CAA63036FAAF88D8" box="[159,1378,553,937]" gridcols="7" gridrows="9" pageId="11" pageNumber="432">
<tr id="4885C425FFD042F5CAA63036FAAF892F" box="[159,1378,553,606]" gridrow="0" pageId="11" pageNumber="432">
<th id="0B54AD59FFD042F5CAA63036FE4D892F" box="[159,384,553,606]" gridcol="0" gridrow="0" pageId="11" pageNumber="432">Variables</th>
<th id="0B54AD59FFD042F5C81D3036FDBA892F" box="[548,631,553,606]" gridcol="1" gridrow="0" pageId="11" pageNumber="432">Wilks' Lambda</th>
<th id="0B54AD59FFD042F5C8803036FCC1892F" box="[697,780,553,606]" gridcol="2" gridrow="0" pageId="11" pageNumber="432">Partial Lambda</th>
<th id="0B54AD59FFD042F5C9743036FC39892F" box="[845,1012,553,606]" gridcol="3" gridrow="0" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C9743035FC91894E" box="[845,860,554,575]" italics="true" pageId="11" pageNumber="432">F</emphasis>
-remove (2.39)
</th>
<th id="0B54AD59FFD042F5CE1D3036FB9C892F" box="[1060,1105,553,606]" gridcol="4" gridrow="0" pageId="11" pageNumber="432">P</th>
<th id="0B54AD59FFD042F5CE473036FB01892F" box="[1150,1228,553,606]" gridcol="5" gridrow="0" pageId="11" pageNumber="432">Root 1</th>
<th id="0B54AD59FFD042F5CF2A3036FAAF892F" box="[1299,1378,553,606]" gridcol="6" gridrow="0" pageId="11" pageNumber="432">Root 2</th>
</tr>
<tr id="4885C425FFD042F5CAA6306EFAAF89F6" box="[159,1378,625,647]" gridrow="1" pageId="11" pageNumber="432">
<th id="0B54AD59FFD042F5CAA6306EFE4D89F6" box="[159,384,625,647]" gridcol="0" gridrow="1" pageId="11" pageNumber="432">Snout vent length</th>
<td id="0B54AD59FFD042F5C81D306EFDBA89F6" box="[548,631,625,647]" gridcol="1" gridrow="1" pageId="11" pageNumber="432">0.033</td>
<td id="0B54AD59FFD042F5C880306EFCC189F6" box="[697,780,625,647]" gridcol="2" gridrow="1" pageId="11" pageNumber="432">0.773</td>
<td id="0B54AD59FFD042F5C974306EFC3989F6" box="[845,1012,625,647]" gridcol="3" gridrow="1" pageId="11" pageNumber="432">5.278</td>
<td id="0B54AD59FFD042F5CE1D306EFB9C89F6" box="[1060,1105,625,647]" gridcol="4" gridrow="1" pageId="11" pageNumber="432">0.00</td>
<td id="0B54AD59FFD042F5CE47306EFB0189F6" box="[1150,1228,625,647]" gridcol="5" gridrow="1" pageId="11" pageNumber="432">1.2387</td>
<td id="0B54AD59FFD042F5CF2A306EFAAF89F6" box="[1299,1378,625,647]" gridcol="6" gridrow="1" pageId="11" pageNumber="432">-0.0678</td>
</tr>
<tr id="4885C425FFD042F5CAA63084FAAF89C0" box="[159,1378,667,689]" gridrow="2" pageId="11" pageNumber="432">
<th id="0B54AD59FFD042F5CAA63084FE4D89C0" box="[159,384,667,689]" gridcol="0" gridrow="2" pageId="11" pageNumber="432">Tibia length</th>
<td id="0B54AD59FFD042F5C81D3084FDBA89C0" box="[548,631,667,689]" gridcol="1" gridrow="2" pageId="11" pageNumber="432">0.034</td>
<td id="0B54AD59FFD042F5C8803084FCC189C0" box="[697,780,667,689]" gridcol="2" gridrow="2" pageId="11" pageNumber="432">0.752</td>
<td id="0B54AD59FFD042F5C9743084FC3989C0" box="[845,1012,667,689]" gridcol="3" gridrow="2" pageId="11" pageNumber="432">5.929</td>
<td id="0B54AD59FFD042F5CE1D3084FB9C89C0" box="[1060,1105,667,689]" gridcol="4" gridrow="2" pageId="11" pageNumber="432">0.00</td>
<td id="0B54AD59FFD042F5CE473084FB0189C0" box="[1150,1228,667,689]" gridcol="5" gridrow="2" pageId="11" pageNumber="432">0.8635</td>
<td id="0B54AD59FFD042F5CF2A3084FAAF89C0" box="[1299,1378,667,689]" gridcol="6" gridrow="2" pageId="11" pageNumber="432">-0.3502</td>
</tr>
<tr id="4885C425FFD042F5CAA630DBFAAF89AB" box="[159,1378,708,730]" gridrow="3" pageId="11" pageNumber="432">
<th id="0B54AD59FFD042F5CAA630DBFE4D89AB" box="[159,384,708,730]" gridcol="0" gridrow="3" pageId="11" pageNumber="432">Eye diameter</th>
<td id="0B54AD59FFD042F5C81D30DBFDBA89AB" box="[548,631,708,730]" gridcol="1" gridrow="3" pageId="11" pageNumber="432">0.029</td>
<td id="0B54AD59FFD042F5C88030DBFCC189AB" box="[697,780,708,730]" gridcol="2" gridrow="3" pageId="11" pageNumber="432">0.882</td>
<td id="0B54AD59FFD042F5C97430DBFC3989AB" box="[845,1012,708,730]" gridcol="3" gridrow="3" pageId="11" pageNumber="432">2.403</td>
<td id="0B54AD59FFD042F5CE1D30DBFB9C89AB" box="[1060,1105,708,730]" gridcol="4" gridrow="3" pageId="11" pageNumber="432">0.10</td>
<td id="0B54AD59FFD042F5CE4730DBFB0189AB" box="[1150,1228,708,730]" gridcol="5" gridrow="3" pageId="11" pageNumber="432">0.3053</td>
<td id="0B54AD59FFD042F5CF2A30DBFAAF89AB" box="[1299,1378,708,730]" gridcol="6" gridrow="3" pageId="11" pageNumber="432">-0.4290</td>
</tr>
<tr id="4885C425FFD042F5CAA630F2FAAF8872" box="[159,1378,749,771]" gridrow="4" pageId="11" pageNumber="432">
<th id="0B54AD59FFD042F5CAA630F2FE4D8872" box="[159,384,749,771]" gridcol="0" gridrow="4" pageId="11" pageNumber="432">Eye to nostril distance</th>
<td id="0B54AD59FFD042F5C81D30F2FDBA8872" box="[548,631,749,771]" gridcol="1" gridrow="4" pageId="11" pageNumber="432">0.035</td>
<td id="0B54AD59FFD042F5C88030F2FCC18872" box="[697,780,749,771]" gridcol="2" gridrow="4" pageId="11" pageNumber="432">0.718</td>
<td id="0B54AD59FFD042F5C97430F2FC398872" box="[845,1012,749,771]" gridcol="3" gridrow="4" pageId="11" pageNumber="432">7.034</td>
<td id="0B54AD59FFD042F5CE1D30F2FB9C8872" box="[1060,1105,749,771]" gridcol="4" gridrow="4" pageId="11" pageNumber="432">0.00</td>
<td id="0B54AD59FFD042F5CE4730F2FB018872" box="[1150,1228,749,771]" gridcol="5" gridrow="4" pageId="11" pageNumber="432">1.0696</td>
<td id="0B54AD59FFD042F5CF2A30F2FAAF8872" box="[1299,1378,749,771]" gridcol="6" gridrow="4" pageId="11" pageNumber="432">0.0121</td>
</tr>
<tr id="4885C425FFD042F5CAA63109FAAF885D" box="[159,1378,790,812]" gridrow="5" pageId="11" pageNumber="432">
<th id="0B54AD59FFD042F5CAA63109FE4D885D" box="[159,384,790,812]" gridcol="0" gridrow="5" pageId="11" pageNumber="432">Snout eye distance</th>
<td id="0B54AD59FFD042F5C81D3109FDBA885D" box="[548,631,790,812]" gridcol="1" gridrow="5" pageId="11" pageNumber="432">0.040</td>
<td id="0B54AD59FFD042F5C8803109FCC1885D" box="[697,780,790,812]" gridcol="2" gridrow="5" pageId="11" pageNumber="432">0.635</td>
<td id="0B54AD59FFD042F5C9743109FC39885D" box="[845,1012,790,812]" gridcol="3" gridrow="5" pageId="11" pageNumber="432">10.309</td>
<td id="0B54AD59FFD042F5CE1D3109FB9C885D" box="[1060,1105,790,812]" gridcol="4" gridrow="5" pageId="11" pageNumber="432">0.00</td>
<td id="0B54AD59FFD042F5CE473109FB01885D" box="[1150,1228,790,812]" gridcol="5" gridrow="5" pageId="11" pageNumber="432">-1.3239</td>
<td id="0B54AD59FFD042F5CF2A3109FAAF885D" box="[1299,1378,790,812]" gridcol="6" gridrow="5" pageId="11" pageNumber="432">0.2792</td>
</tr>
<tr id="4885C425FFD042F5CAA6315FFAAF8827" box="[159,1378,832,854]" gridrow="6" pageId="11" pageNumber="432">
<th id="0B54AD59FFD042F5CAA6315FFE4D8827" box="[159,384,832,854]" gridcol="0" gridrow="6" pageId="11" pageNumber="432">Tympanum diameter</th>
<td id="0B54AD59FFD042F5C81D315FFDBA8827" box="[548,631,832,854]" gridcol="1" gridrow="6" pageId="11" pageNumber="432">0.036</td>
<td id="0B54AD59FFD042F5C880315FFCC18827" box="[697,780,832,854]" gridcol="2" gridrow="6" pageId="11" pageNumber="432">0.705</td>
<td id="0B54AD59FFD042F5C974315FFC398827" box="[845,1012,832,854]" gridcol="3" gridrow="6" pageId="11" pageNumber="432">7.496</td>
<td id="0B54AD59FFD042F5CE1D315FFB9C8827" box="[1060,1105,832,854]" gridcol="4" gridrow="6" pageId="11" pageNumber="432">0.00</td>
<td id="0B54AD59FFD042F5CE47315FFB018827" box="[1150,1228,832,854]" gridcol="5" gridrow="6" pageId="11" pageNumber="432">-0.9808</td>
<td id="0B54AD59FFD042F5CF2A315FFAAF8827" box="[1299,1378,832,854]" gridcol="6" gridrow="6" pageId="11" pageNumber="432">0.0280</td>
</tr>
<tr id="4885C425FFD042F5CAA63176FAAF880E" box="[159,1378,873,895]" gridrow="7" pageId="11" pageNumber="432">
<th id="0B54AD59FFD042F5CAA63176FE4D880E" box="[159,384,873,895]" gridcol="0" gridrow="7" pageId="11" pageNumber="432">Toe III length</th>
<td id="0B54AD59FFD042F5C81D3176FDBA880E" box="[548,631,873,895]" gridcol="1" gridrow="7" pageId="11" pageNumber="432">0.035</td>
<td id="0B54AD59FFD042F5C8803176FCC1880E" box="[697,780,873,895]" gridcol="2" gridrow="7" pageId="11" pageNumber="432">0.723</td>
<td id="0B54AD59FFD042F5C9743176FC39880E" box="[845,1012,873,895]" gridcol="3" gridrow="7" pageId="11" pageNumber="432">6.867</td>
<td id="0B54AD59FFD042F5CE1D3176FB9C880E" box="[1060,1105,873,895]" gridcol="4" gridrow="7" pageId="11" pageNumber="432">0.00</td>
<td id="0B54AD59FFD042F5CE473176FB01880E" box="[1150,1228,873,895]" gridcol="5" gridrow="7" pageId="11" pageNumber="432">-1.2393</td>
<td id="0B54AD59FFD042F5CF2A3176FAAF880E" box="[1299,1378,873,895]" gridcol="6" gridrow="7" pageId="11" pageNumber="432">-0.9580</td>
</tr>
<tr id="4885C425FFD042F5CAA6318CFAAF88D8" box="[159,1378,915,937]" gridrow="8" pageId="11" pageNumber="432">
<th id="0B54AD59FFD042F5CAA6318CFE4D88D8" box="[159,384,915,937]" gridcol="0" gridrow="8" pageId="11" pageNumber="432">Toe IV length</th>
<td id="0B54AD59FFD042F5C81D318CFDBA88D8" box="[548,631,915,937]" gridcol="1" gridrow="8" pageId="11" pageNumber="432">0.030</td>
<td id="0B54AD59FFD042F5C880318CFCC188D8" box="[697,780,915,937]" gridcol="2" gridrow="8" pageId="11" pageNumber="432">0.840</td>
<td id="0B54AD59FFD042F5C974318CFC3988D8" box="[845,1012,915,937]" gridcol="3" gridrow="8" pageId="11" pageNumber="432">3.424</td>
<td id="0B54AD59FFD042F5CE1D318CFB9C88D8" box="[1060,1105,915,937]" gridcol="4" gridrow="8" pageId="11" pageNumber="432">0.04</td>
<td id="0B54AD59FFD042F5CE47318CFB0188D8" box="[1150,1228,915,937]" gridcol="5" gridrow="8" pageId="11" pageNumber="432">1.2585</td>
<td id="0B54AD59FFD042F5CF2A318CFAAF88D8" box="[1299,1378,915,937]" gridcol="6" gridrow="8" pageId="11" pageNumber="432">0.7482</td>
</tr>
</table>
</paragraph>
<paragraph id="F60AC667FFD0BD01CE1D31A3FA9488A3" blockId="11.[1060,1369,956,1019]" box="[1060,1369,956,978]" pageId="11" pageNumber="432">Eig 10.1724 2.4713</paragraph>
<paragraph id="F60AC667FFD0BD01CE1D31FAFAED888A" blockId="11.[1060,1369,956,1019]" box="[1060,1312,997,1019]" pageId="11" pageNumber="432">Cp 0.8045 1</paragraph>
</subSubSection>
<subSubSection id="BEAF95ECFFD0BD04CAFE3622FA50882C" lastPageId="14" lastPageNumber="435" pageId="11" pageNumber="432" type="diagnosis">
<paragraph id="F60AC667FFD0BD01CAFE3622FBE08FB1" blockId="11.[151,1436,1084,1217]" pageId="11" pageNumber="432">
<taxonomicName id="31B5BDE4FFD0BD01CAFE3622FE538F25" box="[199,414,1085,1108]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="432" phylum="Chordata" rank="species" species="stictus">
<emphasis id="C4C11A75FFD0BD01CAFE3622FE538F25" box="[199,414,1085,1108]" italics="true" pageId="11" pageNumber="432">Pristimantis stictus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="31B5BDE4FFD0BD01C8093622FD028F25" box="[560,719,1084,1108]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="432" phylum="Chordata" rank="species" species="leptolophus">
<emphasis id="C4C11A75FFD0BD01C8093622FDF38F25" box="[560,574,1085,1108]" italics="true" pageId="11" pageNumber="432">P</emphasis>
.
<emphasis id="C4C11A75FFD0BD01C8753623FD028F25" box="[588,719,1084,1108]" italics="true" pageId="11" pageNumber="432">leptolophus</emphasis>
</taxonomicName>
in a higher snout-vent length, tibia length, eye-diameter, eye-tonostril distance, Toe III length, and Toe IV length (
<tableCitation id="BB37F3DCFFD0BD01C8F2367FFCEF8F09" box="[715,802,1120,1145]" captionStart="TABLE 4" captionStartId="11.[151,239,1260,1283]" captionTargetBox="[159,1353,1310,1710]" captionTargetPageId="11" captionText="TABLE 4. Mean and standard deviation of measurements showing with significant differences in Tukey Test HSD." httpUri="http://table.plazi.org/id/A2CA96EFFFD0BD01CAAE36F3FAAA8E72" pageId="11" pageNumber="432" tableUuid="A2CA96EFFFD0BD01CAAE36F3FAAA8E72">Table 4</tableCitation>
). The average eye-diameter, tympanum diameter, Toe III length, Toe IV length was significantly higher in
<taxonomicName id="31B5BDE4FFD0BD01C8EA369AFCF48FED" box="[723,825,1157,1180]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="432" phylum="Chordata" rank="species" species="stictus">
<emphasis id="C4C11A75FFD0BD01C8EA369AFD2C8FED" box="[723,737,1157,1180]" italics="true" pageId="11" pageNumber="432">P</emphasis>
.
<emphasis id="C4C11A75FFD0BD01C8CB369AFCF48FED" box="[754,825,1157,1180]" italics="true" pageId="11" pageNumber="432">stictus</emphasis>
</taxonomicName>
in comparison with
<taxonomicName id="31B5BDE4FFD0BD01CE13369AFB0A8FED" box="[1066,1223,1156,1180]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="432" phylum="Chordata" rank="species" species="uranobates">
<emphasis id="C4C11A75FFD0BD01CE13369AFBF58FED" box="[1066,1080,1157,1180]" italics="true" pageId="11" pageNumber="432">P</emphasis>
.
<emphasis id="C4C11A75FFD0BD01CE70369BFB0A8FED" box="[1097,1223,1156,1180]" italics="true" pageId="11" pageNumber="432">uranobates</emphasis>
</taxonomicName>
, but eye-to-nostril distance was significantly lower in
<taxonomicName id="31B5BDE4FFD0BD01C81A36B6FD488FB1" box="[547,645,1193,1216]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="432" phylum="Chordata" rank="species" species="stictus">
<emphasis id="C4C11A75FFD0BD01C81A36B6FDFC8FB1" box="[547,561,1193,1216]" italics="true" pageId="11" pageNumber="432">P</emphasis>
.
<emphasis id="C4C11A75FFD0BD01C80736B6FD488FB1" box="[574,645,1193,1216]" italics="true" pageId="11" pageNumber="432">stictus</emphasis>
</taxonomicName>
compared to
<taxonomicName id="31B5BDE4FFD0BD01C92636B6FC748FB1" box="[799,953,1192,1216]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="432" phylum="Chordata" rank="species" species="uranobates">
<emphasis id="C4C11A75FFD0BD01C92636B6FCE08FB1" box="[799,813,1193,1216]" italics="true" pageId="11" pageNumber="432">P</emphasis>
.
<emphasis id="C4C11A75FFD0BD01C90236B7FC748FB1" box="[827,953,1192,1216]" italics="true" pageId="11" pageNumber="432">uranobates</emphasis>
</taxonomicName>
(
<tableCitation id="BB37F3DCFFD0BD01C9F136B7FBD28FB1" box="[968,1055,1192,1217]" captionStart="TABLE 4" captionStartId="11.[151,239,1260,1283]" captionTargetBox="[159,1353,1310,1710]" captionTargetPageId="11" captionText="TABLE 4. Mean and standard deviation of measurements showing with significant differences in Tukey Test HSD." httpUri="http://table.plazi.org/id/A2CA96EFFFD0BD01CAAE36F3FAAA8E72" pageId="11" pageNumber="432" tableUuid="A2CA96EFFFD0BD01CAAE36F3FAAA8E72">Table 4</tableCitation>
).
</paragraph>
<caption id="A2CA96EFFFD0BD01CAAE36F3FAAA8E72" ID-Table-UUID="A2CA96EFFFD0BD01CAAE36F3FAAA8E72" box="[151,1383,1260,1283]" httpUri="http://table.plazi.org/id/A2CA96EFFFD0BD01CAAE36F3FAAA8E72" pageId="11" pageNumber="432" targetBox="[159,1353,1310,1710]" targetIsTable="true" targetPageId="11">
<paragraph id="F60AC667FFD0BD01CAAE36F3FAAA8E72" blockId="11.[151,1383,1260,1283]" box="[151,1383,1260,1283]" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CAAE36F3FEC48E72" bold="true" box="[151,265,1260,1283]" pageId="11" pageNumber="432">TABLE 4.</emphasis>
Mean and standard deviation of measurements showing with significant differences in Tukey Test HSD.
</paragraph>
</caption>
<table id="84B534C7FFD042F5CAA63701FA848DDF" box="[159,1353,1310,1710]" gridcols="7" gridrows="10" pageId="11" pageNumber="432">
<paragraph id="F60AC667FFD0BD01CA9C3701FA848DDF" pageId="11" pageNumber="432">
<tr id="4885C425FFD042F5CAA63701FA848E44" box="[159,1353,1310,1333]" gridrow="0" pageId="11" pageNumber="432" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1">
<th id="0B54AD59FFD042F5CAA63701FE4D8E44" box="[159,384,1310,1333]" gridcol="0" gridrow="0" pageId="11" pageNumber="432">Variables</th>
<th id="0B54AD59FFD042F5C8573701FD658E44" box="[622,680,1310,1333]" gridcol="1" gridrow="0" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C857373FFD4F8E44" box="[622,642,1312,1333]" italics="true" pageId="11" pageNumber="432">P.</emphasis>
</th>
<th id="0B54AD59FFD042F5CE303701FB8E8E44" box="[1033,1091,1310,1333]" gridcol="4" gridrow="0" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CE30373FFBD18E44" box="[1033,1052,1312,1333]" italics="true" pageId="11" pageNumber="432">P.</emphasis>
</th>
</tr>
<tr id="4885C425FFD042F5CAA63757FA848E2F" box="[159,1353,1352,1374]" gridrow="1" pageId="11" pageNumber="432" rowspan-0="1">
<td id="0B54AD59FFD042F5C8573757FD658E2F" box="[622,680,1352,1374]" gridcol="1" gridrow="1" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C8573756FD658E2F" box="[622,680,1353,1374]" italics="true" pageId="11" pageNumber="432">Mean</emphasis>
</td>
<td id="0B54AD59FFD042F5C8C63757FCD28E2F" box="[767,799,1352,1374]" gridcol="2" gridrow="1" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C8C63757FCD28E2F" box="[767,799,1352,1374]" italics="true" pageId="11" pageNumber="432">SD</emphasis>
</td>
<td id="0B54AD59FFD042F5C9BA3757FC628E2F" box="[899,943,1352,1374]" gridcol="3" gridrow="1" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C9BA3756FC5C8E2F" box="[899,913,1353,1374]" italics="true" pageId="11" pageNumber="432">P</emphasis>
</td>
<td id="0B54AD59FFD042F5CE303757FB8E8E2F" box="[1033,1091,1352,1374]" gridcol="4" gridrow="1" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CE303756FB8E8E2F" box="[1033,1091,1353,1374]" italics="true" pageId="11" pageNumber="432">Mean</emphasis>
</td>
<td id="0B54AD59FFD042F5CEA03757FB748E2F" box="[1177,1209,1352,1374]" gridcol="5" gridrow="1" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CEA03757FB748E2F" box="[1177,1209,1352,1374]" italics="true" pageId="11" pageNumber="432">SD</emphasis>
</td>
<td id="0B54AD59FFD042F5CF243757FA848E2F" box="[1309,1353,1352,1374]" gridcol="6" gridrow="1" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CF243756FAE68E2F" box="[1309,1323,1353,1374]" italics="true" pageId="11" pageNumber="432">P</emphasis>
</td>
</tr>
<tr id="4885C425FFD042F5CAA6376CFA848EF8" box="[159,1353,1395,1417]" gridrow="2" pageId="11" pageNumber="432">
<th id="0B54AD59FFD042F5CAA6376CFE4D8EF8" box="[159,384,1395,1417]" gridcol="0" gridrow="2" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CAA6376CFE9C8EF8" box="[159,337,1395,1417]" italics="true" pageId="11" pageNumber="432">Snout vent length</emphasis>
</th>
<td id="0B54AD59FFD042F5C857376CFD658EF8" box="[622,680,1395,1417]" gridcol="1" gridrow="2" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C857376CFD578EF8" box="[622,666,1395,1417]" italics="true" pageId="11" pageNumber="432">24.9</emphasis>
</td>
<td id="0B54AD59FFD042F5C8C6376CFCD28EF8" box="[767,799,1395,1417]" gridcol="2" gridrow="2" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C8C6376CFCD28EF8" box="[767,799,1395,1417]" italics="true" pageId="11" pageNumber="432">1.9</emphasis>
</td>
<td id="0B54AD59FFD042F5C9BA376CFC628EF8" box="[899,943,1395,1417]" gridcol="3" gridrow="2" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C9BA376CFC628EF8" box="[899,943,1395,1417]" italics="true" pageId="11" pageNumber="432">0.65</emphasis>
</td>
<td id="0B54AD59FFD042F5CE30376CFB8E8EF8" box="[1033,1091,1395,1417]" gridcol="4" gridrow="2" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CE30376CFBF88EF8" box="[1033,1077,1395,1417]" italics="true" pageId="11" pageNumber="432">23.4</emphasis>
</td>
<td id="0B54AD59FFD042F5CEA0376CFB748EF8" box="[1177,1209,1395,1417]" gridcol="5" gridrow="2" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CEA0376CFB748EF8" box="[1177,1209,1395,1417]" italics="true" pageId="11" pageNumber="432">1.4</emphasis>
</td>
<td id="0B54AD59FFD042F5CF24376CFA848EF8" box="[1309,1353,1395,1417]" gridcol="6" gridrow="2" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CF24376CFA848EF8" box="[1309,1353,1395,1417]" italics="true" pageId="11" pageNumber="432">0.00</emphasis>
</td>
</tr>
<tr id="4885C425FFD042F5CAA63783FA848EC3" box="[159,1353,1436,1458]" gridrow="3" pageId="11" pageNumber="432">
<th id="0B54AD59FFD042F5CAA63783FE4D8EC3" box="[159,384,1436,1458]" gridcol="0" gridrow="3" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CAA63783FED48EC3" box="[159,281,1436,1458]" italics="true" pageId="11" pageNumber="432">Tibia length</emphasis>
</th>
<td id="0B54AD59FFD042F5C8573783FD658EC3" box="[622,680,1436,1458]" gridcol="1" gridrow="3" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C8573783FD578EC3" box="[622,666,1436,1458]" italics="true" pageId="11" pageNumber="432">12.5</emphasis>
</td>
<td id="0B54AD59FFD042F5C8C63783FCD28EC3" box="[767,799,1436,1458]" gridcol="2" gridrow="3" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C8C63783FCD28EC3" box="[767,799,1436,1458]" italics="true" pageId="11" pageNumber="432">0.8</emphasis>
</td>
<td id="0B54AD59FFD042F5C9BA3783FC628EC3" box="[899,943,1436,1458]" gridcol="3" gridrow="3" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C9BA3783FC628EC3" box="[899,943,1436,1458]" italics="true" pageId="11" pageNumber="432">0.26</emphasis>
</td>
<td id="0B54AD59FFD042F5CE303783FB8E8EC3" box="[1033,1091,1436,1458]" gridcol="4" gridrow="3" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CE303783FBF88EC3" box="[1033,1077,1436,1458]" italics="true" pageId="11" pageNumber="432">11.3</emphasis>
</td>
<td id="0B54AD59FFD042F5CEA03783FB748EC3" box="[1177,1209,1436,1458]" gridcol="5" gridrow="3" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CEA03783FB748EC3" box="[1177,1209,1436,1458]" italics="true" pageId="11" pageNumber="432">0.5</emphasis>
</td>
<td id="0B54AD59FFD042F5CF243783FA848EC3" box="[1309,1353,1436,1458]" gridcol="6" gridrow="3" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CF243783FA848EC3" box="[1309,1353,1436,1458]" italics="true" pageId="11" pageNumber="432">0.00</emphasis>
</td>
</tr>
<tr id="4885C425FFD042F5CAA637D9FA848EAD" box="[159,1353,1478,1500]" gridrow="4" pageId="11" pageNumber="432">
<th id="0B54AD59FFD042F5CAA637D9FE4D8EAD" box="[159,384,1478,1500]" gridcol="0" gridrow="4" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CAA637D8FEE98EAD" box="[159,292,1478,1500]" italics="true" pageId="11" pageNumber="432">Eye diameter</emphasis>
</th>
<td id="0B54AD59FFD042F5C85737D9FD658EAD" box="[622,680,1478,1500]" gridcol="1" gridrow="4" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C85737D9FD438EAD" box="[622,654,1478,1500]" italics="true" pageId="11" pageNumber="432">2.9</emphasis>
</td>
<td id="0B54AD59FFD042F5C8C637D9FCD28EAD" box="[767,799,1478,1500]" gridcol="2" gridrow="4" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C8C637D9FCD28EAD" box="[767,799,1478,1500]" italics="true" pageId="11" pageNumber="432">0.2</emphasis>
</td>
<td id="0B54AD59FFD042F5C9BA37D9FC628EAD" box="[899,943,1478,1500]" gridcol="3" gridrow="4" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C9BA37D9FC628EAD" box="[899,943,1478,1500]" italics="true" pageId="11" pageNumber="432">0.00</emphasis>
</td>
<td id="0B54AD59FFD042F5CE3037D9FB8E8EAD" box="[1033,1091,1478,1500]" gridcol="4" gridrow="4" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CE3037D9FBE48EAD" box="[1033,1065,1478,1500]" italics="true" pageId="11" pageNumber="432">2.7</emphasis>
</td>
<td id="0B54AD59FFD042F5CEA037D9FB748EAD" box="[1177,1209,1478,1500]" gridcol="5" gridrow="4" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CEA037D9FB748EAD" box="[1177,1209,1478,1500]" italics="true" pageId="11" pageNumber="432">0.1</emphasis>
</td>
<td id="0B54AD59FFD042F5CF2437D9FA848EAD" box="[1309,1353,1478,1500]" gridcol="6" gridrow="4" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CF2437D9FA848EAD" box="[1309,1353,1478,1500]" italics="true" pageId="11" pageNumber="432">0.00</emphasis>
</td>
</tr>
<tr id="4885C425FFD042F5CAA637EFFA848D77" box="[159,1353,1520,1542]" gridrow="5" pageId="11" pageNumber="432">
<th id="0B54AD59FFD042F5CAA637EFFE4D8D77" box="[159,384,1520,1542]" gridcol="0" gridrow="5" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CAA637EEFE4D8D77" box="[159,384,1520,1542]" italics="true" pageId="11" pageNumber="432">Eye to nostril distance</emphasis>
</th>
<td id="0B54AD59FFD042F5C85737EFFD658D77" box="[622,680,1520,1542]" gridcol="1" gridrow="5" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C85737EFFD438D77" box="[622,654,1520,1542]" italics="true" pageId="11" pageNumber="432">2.5</emphasis>
</td>
<td id="0B54AD59FFD042F5C8C637EFFCD28D77" box="[767,799,1520,1542]" gridcol="2" gridrow="5" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C8C637EFFCD28D77" box="[767,799,1520,1542]" italics="true" pageId="11" pageNumber="432">0.2</emphasis>
</td>
<td id="0B54AD59FFD042F5C9BA37EFFC628D77" box="[899,943,1520,1542]" gridcol="3" gridrow="5" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C9BA37EFFC628D77" box="[899,943,1520,1542]" italics="true" pageId="11" pageNumber="432">0.00</emphasis>
</td>
<td id="0B54AD59FFD042F5CE3037EFFB8E8D77" box="[1033,1091,1520,1542]" gridcol="4" gridrow="5" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CE3037EFFBE48D77" box="[1033,1065,1520,1542]" italics="true" pageId="11" pageNumber="432">2.1</emphasis>
</td>
<td id="0B54AD59FFD042F5CEA037EFFB748D77" box="[1177,1209,1520,1542]" gridcol="5" gridrow="5" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CEA037EFFB748D77" box="[1177,1209,1520,1542]" italics="true" pageId="11" pageNumber="432">0.1</emphasis>
</td>
<td id="0B54AD59FFD042F5CF2437EFFA848D77" box="[1309,1353,1520,1542]" gridcol="6" gridrow="5" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CF2437EFFA848D77" box="[1309,1353,1520,1542]" italics="true" pageId="11" pageNumber="432">0.00</emphasis>
</td>
</tr>
<tr id="4885C425FFD042F5CAA63405FA848D41" box="[159,1353,1562,1584]" gridrow="6" pageId="11" pageNumber="432">
<th id="0B54AD59FFD042F5CAA63405FE4D8D41" box="[159,384,1562,1584]" gridcol="0" gridrow="6" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CAA63405FE908D41" box="[159,349,1562,1584]" italics="true" pageId="11" pageNumber="432">Snout eye distance</emphasis>
</th>
<td id="0B54AD59FFD042F5C8573405FD658D41" box="[622,680,1562,1584]" gridcol="1" gridrow="6" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C8573405FD438D41" box="[622,654,1562,1584]" italics="true" pageId="11" pageNumber="432">4.0</emphasis>
</td>
<td id="0B54AD59FFD042F5C8C63405FCD28D41" box="[767,799,1562,1584]" gridcol="2" gridrow="6" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C8C63405FCD28D41" box="[767,799,1562,1584]" italics="true" pageId="11" pageNumber="432">0.3</emphasis>
</td>
<td id="0B54AD59FFD042F5C9BA3405FC628D41" box="[899,943,1562,1584]" gridcol="3" gridrow="6" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C9BA3405FC628D41" box="[899,943,1562,1584]" italics="true" pageId="11" pageNumber="432">0.99</emphasis>
</td>
<td id="0B54AD59FFD042F5CE303405FB8E8D41" box="[1033,1091,1562,1584]" gridcol="4" gridrow="6" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CE303405FBE48D41" box="[1033,1065,1562,1584]" italics="true" pageId="11" pageNumber="432">4.0</emphasis>
</td>
<td id="0B54AD59FFD042F5CEA03405FB748D41" box="[1177,1209,1562,1584]" gridcol="5" gridrow="6" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CEA03405FB748D41" box="[1177,1209,1562,1584]" italics="true" pageId="11" pageNumber="432">0.2</emphasis>
</td>
<td id="0B54AD59FFD042F5CF243405FA848D41" box="[1309,1353,1562,1584]" gridcol="6" gridrow="6" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CF243405FA848D41" box="[1309,1353,1562,1584]" italics="true" pageId="11" pageNumber="432">0.95</emphasis>
</td>
</tr>
<tr id="4885C425FFD042F5CAA6345BFA848D2B" box="[159,1353,1604,1626]" gridrow="7" pageId="11" pageNumber="432">
<th id="0B54AD59FFD042F5CAA6345BFE4D8D2B" box="[159,384,1604,1626]" gridcol="0" gridrow="7" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CAA6345AFEBC8D2B" box="[159,369,1604,1626]" italics="true" pageId="11" pageNumber="432">Tympanum diameter</emphasis>
</th>
<td id="0B54AD59FFD042F5C857345BFD658D2B" box="[622,680,1604,1626]" gridcol="1" gridrow="7" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C857345BFD438D2B" box="[622,654,1604,1626]" italics="true" pageId="11" pageNumber="432">1.2</emphasis>
</td>
<td id="0B54AD59FFD042F5C8C6345BFCD28D2B" box="[767,799,1604,1626]" gridcol="2" gridrow="7" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C8C6345BFCD28D2B" box="[767,799,1604,1626]" italics="true" pageId="11" pageNumber="432">0.1</emphasis>
</td>
<td id="0B54AD59FFD042F5C9BA345BFC628D2B" box="[899,943,1604,1626]" gridcol="3" gridrow="7" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C9BA345BFC628D2B" box="[899,943,1604,1626]" italics="true" pageId="11" pageNumber="432">0.03</emphasis>
</td>
<td id="0B54AD59FFD042F5CE30345BFB8E8D2B" box="[1033,1091,1604,1626]" gridcol="4" gridrow="7" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CE30345BFBE48D2B" box="[1033,1065,1604,1626]" italics="true" pageId="11" pageNumber="432">1.3</emphasis>
</td>
<td id="0B54AD59FFD042F5CEA0345BFB748D2B" box="[1177,1209,1604,1626]" gridcol="5" gridrow="7" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CEA0345BFB748D2B" box="[1177,1209,1604,1626]" italics="true" pageId="11" pageNumber="432">0.2</emphasis>
</td>
<td id="0B54AD59FFD042F5CF24345BFA848D2B" box="[1309,1353,1604,1626]" gridcol="6" gridrow="7" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CF24345BFA848D2B" box="[1309,1353,1604,1626]" italics="true" pageId="11" pageNumber="432">0.50</emphasis>
</td>
</tr>
<tr id="4885C425FFD042F5CAA63471FA848DF5" box="[159,1353,1646,1668]" gridrow="8" pageId="11" pageNumber="432">
<th id="0B54AD59FFD042F5CAA63471FE4D8DF5" box="[159,384,1646,1668]" gridcol="0" gridrow="8" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CAA63470FEE48DF5" box="[159,297,1646,1668]" italics="true" pageId="11" pageNumber="432">Toe III length</emphasis>
</th>
<td id="0B54AD59FFD042F5C8573471FD658DF5" box="[622,680,1646,1668]" gridcol="1" gridrow="8" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C8573471FD438DF5" box="[622,654,1646,1668]" italics="true" pageId="11" pageNumber="432">3.9</emphasis>
</td>
<td id="0B54AD59FFD042F5C8C63471FCD28DF5" box="[767,799,1646,1668]" gridcol="2" gridrow="8" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C8C63471FCD28DF5" box="[767,799,1646,1668]" italics="true" pageId="11" pageNumber="432">0.3</emphasis>
</td>
<td id="0B54AD59FFD042F5C9BA3471FC628DF5" box="[899,943,1646,1668]" gridcol="3" gridrow="8" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C9BA3471FC628DF5" box="[899,943,1646,1668]" italics="true" pageId="11" pageNumber="432">0.00</emphasis>
</td>
<td id="0B54AD59FFD042F5CE303471FB8E8DF5" box="[1033,1091,1646,1668]" gridcol="4" gridrow="8" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CE303471FBE48DF5" box="[1033,1065,1646,1668]" italics="true" pageId="11" pageNumber="432">3.8</emphasis>
</td>
<td id="0B54AD59FFD042F5CEA03471FB748DF5" box="[1177,1209,1646,1668]" gridcol="5" gridrow="8" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CEA03471FB748DF5" box="[1177,1209,1646,1668]" italics="true" pageId="11" pageNumber="432">0.3</emphasis>
</td>
<td id="0B54AD59FFD042F5CF243471FA848DF5" box="[1309,1353,1646,1668]" gridcol="6" gridrow="8" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CF243471FA848DF5" box="[1309,1353,1646,1668]" italics="true" pageId="11" pageNumber="432">0.00</emphasis>
</td>
</tr>
<tr id="4885C425FFD042F5CAA63487FA848DDF" box="[159,1353,1688,1710]" gridrow="9" pageId="11" pageNumber="432">
<th id="0B54AD59FFD042F5CAA63487FE4D8DDF" box="[159,384,1688,1710]" gridcol="0" gridrow="9" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CAA63486FEE68DDF" box="[159,299,1688,1710]" italics="true" pageId="11" pageNumber="432">Toe IV length</emphasis>
</th>
<td id="0B54AD59FFD042F5C8573487FD658DDF" box="[622,680,1688,1710]" gridcol="1" gridrow="9" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C8573487FD438DDF" box="[622,654,1688,1710]" italics="true" pageId="11" pageNumber="432">6.5</emphasis>
</td>
<td id="0B54AD59FFD042F5C8C63487FCD28DDF" box="[767,799,1688,1710]" gridcol="2" gridrow="9" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C8C63487FCD28DDF" box="[767,799,1688,1710]" italics="true" pageId="11" pageNumber="432">0.5</emphasis>
</td>
<td id="0B54AD59FFD042F5C9BA3487FC628DDF" box="[899,943,1688,1710]" gridcol="3" gridrow="9" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01C9BA3487FC628DDF" box="[899,943,1688,1710]" italics="true" pageId="11" pageNumber="432">0.00</emphasis>
</td>
<td id="0B54AD59FFD042F5CE303487FB8E8DDF" box="[1033,1091,1688,1710]" gridcol="4" gridrow="9" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CE303487FBE48DDF" box="[1033,1065,1688,1710]" italics="true" pageId="11" pageNumber="432">6.3</emphasis>
</td>
<td id="0B54AD59FFD042F5CEA03487FB748DDF" box="[1177,1209,1688,1710]" gridcol="5" gridrow="9" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CEA03487FB748DDF" box="[1177,1209,1688,1710]" italics="true" pageId="11" pageNumber="432">0.4</emphasis>
</td>
<td id="0B54AD59FFD042F5CF243487FA848DDF" box="[1309,1353,1688,1710]" gridcol="6" gridrow="9" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CF243487FA848DDF" box="[1309,1353,1688,1710]" italics="true" pageId="11" pageNumber="432">0.00</emphasis>
</td>
</tr>
</paragraph>
<paragraph id="F60AC667FFD0BD01C8BE3701FB5A8E45" box="[647,1175,1310,1332]" pageId="11" pageNumber="432">uranobates leptolophus</paragraph>
</table>
<paragraph id="F60AC667FFD0BD06CAFE34EDFCE68911" blockId="11.[151,1436,1778,2019]" lastBlockId="12.[151,1437,151,608]" lastPageId="12" lastPageNumber="433" pageId="11" pageNumber="432">
<emphasis id="C4C11A75FFD0BD01CAFE34EDFEF68C7A" bold="true" box="[199,315,1778,1803]" pageId="11" pageNumber="432">Remarks.</emphasis>
Variation in vomerine odontophores has received some attention in the literature (e.g.,
<bibRefCitation id="9224BB96FFD0BD01CF3034EDFA558C7B" author="Lynch" box="[1289,1432,1778,1803]" pageId="11" pageNumber="436" refString="Lynch, J. D. (1980) New species of Eleutherodactylus of Colombia (Amphibia: Leptodactylidae). I: Five new species from the paramos of the Cordillera Central. Caldasia, 13 (61), 165 - 188." type="journal article" year="1980">Lynch, 1980</bibRefCitation>
;
<bibRefCitation id="9224BB96FFD0BD01CAAE3509FE1D8C5E" author="Lynch" box="[151,464,1814,1839]" pageId="11" pageNumber="436" refString="Lynch, J. D. &amp; Duellman, W. E. (1997) Frogs of the genus Eleutherodactylus (Leptodactylidae) in western Ecuador, systematics, ecology, and biogeography. University of Kansas Natural History Museum Special Publications, 23, 1 - 236. http: // dx. doi. org / 10.5962 / bhl. title. 7951" type="journal article" year="1997">Lynch and Duellman, 1997</bibRefCitation>
;
<bibRefCitation id="9224BB96FFD0BD01CBE43509FDA28C5E" author="Lynch" box="[477,623,1814,1839]" pageId="11" pageNumber="436" refString="Lynch, J. D. (2001) Four osteological synapomorphies within Eleutherodactylus: (Amphibia: Leptodactylidae) and their bearing on subgeneric classifications. Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales, 25 (94), 127 - 136." type="journal article" year="2001">Lynch, 2001</bibRefCitation>
;
<bibRefCitation id="9224BB96FFD0BD01C8453509FC698C5E" author="Duellman" box="[636,932,1814,1839]" pageId="11" pageNumber="435" refString="Duellman, W. E. &amp; Lehr, E. (2009) Terrestrial-breeding frogs (Strabomantidae) in Peru. Nature und Tier Verlag, Munster, 384 pp." type="book" year="2009">Duellman and Lehr, 2009</bibRefCitation>
).
<bibRefCitation id="9224BB96FFD0BD01C9803509FB378C5E" author="Lynch" box="[953,1274,1814,1839]" pageId="11" pageNumber="436" refString="Lynch, J. D. &amp; Duellman, W. E. (1997) Frogs of the genus Eleutherodactylus (Leptodactylidae) in western Ecuador, systematics, ecology, and biogeography. University of Kansas Natural History Museum Special Publications, 23, 1 - 236. http: // dx. doi. org / 10.5962 / bhl. title. 7951" type="journal article" year="1997">Lynch and Duellman (1997)</bibRefCitation>
compared the vomerine odontophores among several genera of Brachycephaloidea and discussed the loss of this character in some taxa, as in
<taxonomicName id="31B5BDE4FFD0BD01CB763541FE218C07" box="[335,492,1886,1910]" class="Amphibia" family="Strabomantidae" genus="Atopophrynus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="432" phylum="Chordata" rank="genus">
<emphasis id="C4C11A75FFD0BD01CB763541FE218C07" box="[335,492,1886,1910]" italics="true" pageId="11" pageNumber="432">Atopophrynus</emphasis>
</taxonomicName>
,
<taxonomicName id="31B5BDE4FFD0BD01CBC33541FD548C07" box="[506,665,1886,1910]" class="Amphibia" family="Strabomantidae" genus="Geobatrachus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="432" phylum="Chordata" rank="genus">
<emphasis id="C4C11A75FFD0BD01CBC33541FD548C07" box="[506,665,1886,1910]" italics="true" pageId="11" pageNumber="432">Geobatrachus</emphasis>
</taxonomicName>
,
<taxonomicName id="31B5BDE4FFD0BD01C89F3541FCCA8C07" box="[678,775,1886,1910]" class="Amphibia" family="Strabomantidae" genus="Noblella" kingdom="Animalia" order="Anura" pageId="11" pageNumber="432" phylum="Chordata" rank="genus">
<emphasis id="C4C11A75FFD0BD01C89F3541FCCA8C07" box="[678,775,1886,1910]" italics="true" pageId="11" pageNumber="432">Noblella</emphasis>
</taxonomicName>
,
<taxonomicName id="31B5BDE4FFD0BD01C92D3541FC558C07" box="[788,920,1886,1910]" class="Amphibia" family="Strabomantidae" genus="Bryophryne" kingdom="Animalia" order="Anura" pageId="11" pageNumber="432" phylum="Chordata" rank="genus">
<emphasis id="C4C11A75FFD0BD01C92D3541FC558C07" box="[788,920,1886,1910]" italics="true" pageId="11" pageNumber="432">Bryophryne</emphasis>
</taxonomicName>
, and a few species in the genus of
<taxonomicName id="31B5BDE4FFD0BD01CF253541FA588C07" box="[1308,1429,1886,1910]" class="Amphibia" family="Strabomantidae" genus="Phrynopus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="432" phylum="Chordata" rank="genus">
<emphasis id="C4C11A75FFD0BD01CF253541FA588C07" box="[1308,1429,1886,1910]" italics="true" pageId="11" pageNumber="432">Phrynopus</emphasis>
</taxonomicName>
,
<taxonomicName id="31B5BDE4FFD0BD01CAAE359DFE908CEB" box="[151,349,1922,1946]" class="Amphibia" family="Strabomantidae" genus="Psychrophrynella" kingdom="Animalia" order="Anura" pageId="11" pageNumber="432" phylum="Chordata" rank="genus">
<emphasis id="C4C11A75FFD0BD01CAAE359DFE908CEB" box="[151,349,1922,1946]" italics="true" pageId="11" pageNumber="432">Psychrophrynella</emphasis>
</taxonomicName>
,
<taxonomicName id="31B5BDE4FFD0BD01CB57359DFDF48CEB" box="[366,569,1922,1946]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="432" phylum="Chordata" rank="genus">
<emphasis id="C4C11A75FFD0BD01CB57359DFDF48CEB" box="[366,569,1922,1946]" italics="true" pageId="11" pageNumber="432">Eleutherodactylus</emphasis>
</taxonomicName>
and
<taxonomicName id="31B5BDE4FFD0BD01C84F359DFD3A8CE8" box="[630,759,1922,1945]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="11" pageNumber="432" phylum="Chordata" rank="genus">
<emphasis id="C4C11A75FFD0BD01C84F359DFD3A8CE8" box="[630,759,1922,1945]" italics="true" pageId="11" pageNumber="432">Craugastor</emphasis>
</taxonomicName>
(
<figureCitation id="6E8EDAE2FFD0BD01C930359DFCA68CEB" box="[777,875,1922,1947]" captionStart="FIGURE 8" captionStartId="13.[151,250,1622,1644]" captionTargetBox="[228,1345,240,1520]" captionTargetId="figure@13.[188,1398,193,1600]" captionTargetPageId="13" captionText="FIGURE 8. Vomers variation of some brachyphaloids frogs. A. Eleutherodactylus nitidus (KU- 102657), lacks the dentigerous process; B. Eleutherodactylus leprus (KU- 55963), dentigerous process represented by long sheet, the odontophores are absent; C. Craugastor pygmaeus (KU- 86876), dentigerous process represented by sharp and elongated structure, the odontophores are absent; D. Craugastor escoses, adult female (KU- 117355), dentigerous process consists mainly of a large odontophore; E. Pristimantis danae, adult male (KU- 164062; SVL: 36 mm), elongate dentigerous process with small odontophore; F. Pristimantis orpacobates, adult female (KU- 170134; SVL: 43 mm), dentigerous process with midsize odontophore; G. Pristimantis orestes, adult female (KU- 141995; SVL: 23.5 mm), elongate dentigerous process with low odontophore; H. Pristimantis vicarius, adult male (KU- 170155; SVL: 34 mm), very elongated dentigerous process with odontophore in the distal end. Bars represent measurements of 2 mm; gray denotes cartilage and stippling denotes bones. (All drawings based on original illustrations by Lynch)." httpUri="https://zenodo.org/record/258325/files/figure.png" pageId="11" pageNumber="432">Figure 8</figureCitation>
A–C) (
<bibRefCitation id="9224BB96FFD0BD01C984359DFB9C8CEA" author="Lynch" box="[957,1105,1922,1947]" pageId="11" pageNumber="436" refString="Lynch, J. D. (1975) A review of the Andean leptodactylid frog genus Phrynopus. Occasional Papers of the Museum of Natural History University of Kansas, 35, 1 - 51." type="journal article" year="1975">Lynch, 1975</bibRefCitation>
;
<bibRefCitation id="9224BB96FFD0BD01CE59359DFB398CEB" author="Lynch" box="[1120,1268,1922,1947]" pageId="11" pageNumber="436" refString="Lynch, J. D. (2001) Four osteological synapomorphies within Eleutherodactylus: (Amphibia: Leptodactylidae) and their bearing on subgeneric classifications. Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales, 25 (94), 127 - 136." type="journal article" year="2001">Lynch, 2001</bibRefCitation>
;
<bibRefCitation id="9224BB96FFD0BD01CF3B359DFF1E8CCE" author="Hedges" pageId="11" pageNumber="436" refString="Hedges, S. B., Duellman, W. E. &amp; Heinicke, M. P. (2008) New World direct-developing frogs (Anura, Terrarana), molecular phylogeny, classification, biogeography, and conservation. Zootaxa, 1737, 1 - 182." type="journal article" year="2008">
Hedges
<emphasis id="C4C11A75FFD0BD01CF59359CFA588CEB" box="[1376,1429,1922,1946]" italics="true" pageId="11" pageNumber="432">et al</emphasis>
. 2008
</bibRefCitation>
;
<bibRefCitation id="9224BB96FFD0BD01CAE435B9FE1F8CCE" author="Trueb" box="[221,466,1958,1983]" pageId="11" pageNumber="437" refString="Trueb, L. &amp; Lehr, E. (2008) A new species of Phrynopus (Anura, Strabomantidae) from Peru, with comments on the osteology of the genus. Phyllomedusa: Journal of Herpetology, 7 (1), 11 - 24. http: // dx. doi. org / 10.11606 / issn. 2316 - 9079. v 7 i 1 p 11 - 24" type="journal article" year="2008">Trueb and Lehr, 2008</bibRefCitation>
;
<bibRefCitation id="9224BB96FFD0BD01CBE535B9FD368CCE" author="Lehr" box="[476,763,1958,1983]" pageId="11" pageNumber="436" refString="Lehr, E. &amp; Catenazzi, A. (2008) A new species of Bryophryne (Anura: Strabomantidae) from southern Peru. Zootaxa, 1784, 1 - 10." type="journal article" year="2008">
Lehr and Catenazzi,
<date id="820BE0A7FFD0BD01C8FA35B8FD368CCE" box="[707,763,1959,1983]" pageId="11" pageNumber="432" value="2008" valueMax="2009">2008</date>
</bibRefCitation>
and
<bibRefCitation id="9224BB96FFD0BD01C90935B8FCA18CCE" author="Catenazzi" box="[816,876,1959,1983]" pageId="11" pageNumber="436" refString="Lehr, E. &amp; Catenazzi, A. (2009) A new species of minute Noblella (Anura: Strabomantidae) from southern Peru: the smallest frog of the Andes. Copeia, 2009 (1), 148 - 156. http: // dx. doi. org / 10.1643 / ch- 07 - 270" type="journal article" year="2009">2009</bibRefCitation>
;
<bibRefCitation id="9224BB96FFD0BD01C94E35B9FB848CCE" author="Padial" box="[887,1097,1958,1983]" pageId="11" pageNumber="436" refString="Padial, J. M., Grant, T. &amp; Frost, D. R. (2014 a) Molecular systematics of terraranas (Anura: Brachycephaloidea) with an assessment of the effects of alignment and optimality criteria. Zootaxa, 3825 (1), 1 - 132. http: // dx. doi. org / 10.11646 / zootaxa. 3825.1.1" type="journal article" year="2014" yearSuffix="a">
Padial
<emphasis id="C4C11A75FFD0BD01C9FA35B8FC398CCF" box="[963,1012,1958,1982]" italics="true" pageId="11" pageNumber="432">et al</emphasis>
. 2014a
</bibRefCitation>
;
<bibRefCitation id="9224BB96FFD0BD01CE6D35B9FA898CCE" author="Catenazzi" box="[1108,1348,1958,1983]" pageId="11" pageNumber="435" refString="Catenazzi, A., Uscapi, V. &amp; von May, R. (2015) A new species of Noblella (Amphibia, Anura, Craugastoridae) from the humid montane forests of Cusco, Peru. ZooKeys, (516), 71." type="journal article" year="2015">
Catenazzi
<emphasis id="C4C11A75FFD0BD01CEF035B8FB378CCF" box="[1225,1274,1958,1982]" italics="true" pageId="11" pageNumber="432">et al</emphasis>
. 2015
</bibRefCitation>
).
<bibRefCitation id="9224BB96FFD0BD01CF6C35B9FE468C93" author="Lynch" pageId="11" pageNumber="436" refString="Lynch, J. D. &amp; Duellman, W. E. (1997) Frogs of the genus Eleutherodactylus (Leptodactylidae) in western Ecuador, systematics, ecology, and biogeography. University of Kansas Natural History Museum Special Publications, 23, 1 - 236. http: // dx. doi. org / 10.5962 / bhl. title. 7951" type="journal article" year="1997">Lynch and Duellman (1997)</bibRefCitation>
also noted that species of
<taxonomicName id="31B5BDE4FFD0BD01C8F935D4FC848C93" box="[704,841,1995,2018]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="432" phylum="Chordata" rank="genus">
<emphasis id="C4C11A75FFD0BD01C8F935D4FC848C93" box="[704,841,1995,2018]" italics="true" pageId="11" pageNumber="432">Pristimantis</emphasis>
</taxonomicName>
from western
<collectingCountry id="8EA286F7FFD0BD01C9CB35D5FB828C92" box="[1010,1103,1994,2019]" name="Ecuador" pageId="11" pageNumber="432">Ecuador</collectingCountry>
possess small odontophores, which are concealed beneath the palate tissue—a very distinctive condition for the genus. But despite the variation and potential importance of these features in the taxonomy of brachycephaloids (and in other taxa as well), differences in terminology have obscured the interpretation of observed variation. For example,
<bibRefCitation id="9224BB96FFD7BD06CEFD32C0FF1F8A6D" author="Lynch" pageId="12" pageNumber="436" refString="Lynch, J. D. (1979) Leptodactylid frogs of the genus Eleutherodactylus from the Andes of southern Ecuador. Miscellaneous publication University of Kansas, 66, 1 - 62." type="journal article" year="1979">
Lynch (
<emphasis id="C4C11A75FFD7BD06CF2032FEFA8C8B89" box="[1305,1345,225,248]" italics="true" pageId="12" pageNumber="433">e.g.</emphasis>
, Lynch, 1979
</bibRefCitation>
,
<bibRefCitation id="9224BB96FFD7BD06CAE4331BFEA08A6D" author="Lynch" box="[221,365,260,285]" pageId="12" pageNumber="436" refString="Lynch, J. D. (1980) New species of Eleutherodactylus of Colombia (Amphibia: Leptodactylidae). I: Five new species from the paramos of the Cordillera Central. Caldasia, 13 (61), 165 - 188." type="journal article" year="1980">Lynch, 1980</bibRefCitation>
;
<bibRefCitation id="9224BB96FFD7BD06CB41331BFDC88A6D" author="Lynch" box="[376,517,260,285]" pageId="12" pageNumber="436" refString="Lynch, J. D. (1981) Leptodactylid frogs of the genus Eleutherodactylus in the Andes of northern Ecuador and adjacent Colombia. Miscellaneous publication University of Kansas, 72, 1 - 46. http: // dx. doi. org / 10.5962 / bhl. title. 16289" type="journal article" year="1981">Lynch, 1981</bibRefCitation>
) equates the term dentigerous process of the vomers with vomerine odontophores in many of the descriptions of brachycephaloids frogs.
<bibRefCitation id="9224BB96FFD7BD06C9023338FB408A31" author="Lynch" box="[827,1165,295,320]" pageId="12" pageNumber="436" refString="Lynch, J. D. &amp; Duellman, W. E. (1997) Frogs of the genus Eleutherodactylus (Leptodactylidae) in western Ecuador, systematics, ecology, and biogeography. University of Kansas Natural History Museum Special Publications, 23, 1 - 236. http: // dx. doi. org / 10.5962 / bhl. title. 7951" type="journal article" year="1997">Lynch and Duellman (1997)</bibRefCitation>
also treated vomerine odontophores as a synonym of the dentigerous process of the vomer, although, they recognized that when poorly discernible, the odontophore could be present but concealed beneath the mucosa.
<bibRefCitation id="9224BB96FFD7BD06CE5D3370FACC8AF9" author="Lynch" box="[1124,1281,367,392]" pageId="12" pageNumber="436" refString="Lynch, J. D. (2001) Four osteological synapomorphies within Eleutherodactylus: (Amphibia: Leptodactylidae) and their bearing on subgeneric classifications. Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales, 25 (94), 127 - 136." type="journal article" year="2001">Lynch (2001)</bibRefCitation>
makes some distinctions between odontophore and dentigerous process. Nevertheless, none of them provided accurate definitions. Only
<bibRefCitation id="9224BB96FFD7BD06CB6433A7FDCD8AA1" author="Savage" box="[349,512,440,464]" pageId="12" pageNumber="437" refString="Savage, J. M. (2002) The amphibians and reptiles of Costa Rica: a herpetofauna between two continents, between two seas. University of Chicago Press, Chicago, 934 pp." type="book" year="2002">Savage (2002)</bibRefCitation>
defined the odontophore as tooth-bearing processes of the vomer.
<bibRefCitation id="9224BB96FFD7BD06CED733A8FF2C8A84" pageId="12" pageNumber="435" refString="Duellman, W. E., Lehr, E. &amp; Venegas, P. J. (2006) Two new species of Eleutherodactylus (Anura: Leptodactylidae) from the Andes of northern Peru. Zootaxa, 1285, 51 - 64." type="journal article">
Duellman
<emphasis id="C4C11A75FFD7BD06CF5D33A6FA588AA1" box="[1380,1429,440,464]" italics="true" pageId="12" pageNumber="433">et al</emphasis>
. (2006)
</bibRefCitation>
and
<bibRefCitation id="9224BB96FFD7BD06CB2033C3FDF98A85" author="Duellman" box="[281,564,476,501]" pageId="12" pageNumber="435" refString="Duellman, W. E. &amp; Lehr, E. (2009) Terrestrial-breeding frogs (Strabomantidae) in Peru. Nature und Tier Verlag, Munster, 384 pp." type="book" year="2009">Duellman &amp; Lehr (2009)</bibRefCitation>
reviewed the terminology and proposed using dentigerous process to replace vomerine odontophores.
<collectingCountry id="8EA286F7FFD7BD06CB80301FFE158969" box="[441,472,512,536]" name="American Samoa" pageId="12" pageNumber="433">As</collectingCountry>
a result of the inconsistence in the use of terminology, some species have been described as lacking dentigerous process when the process was in fact concealed underneath the mucosa (
<figureCitation id="6E8EDAE2FFD7BD06CF68303BFF6E8911" captionStart="FIGURE 8" captionStartId="13.[151,250,1622,1644]" captionTargetBox="[228,1345,240,1520]" captionTargetId="figure@13.[188,1398,193,1600]" captionTargetPageId="13" captionText="FIGURE 8. Vomers variation of some brachyphaloids frogs. A. Eleutherodactylus nitidus (KU- 102657), lacks the dentigerous process; B. Eleutherodactylus leprus (KU- 55963), dentigerous process represented by long sheet, the odontophores are absent; C. Craugastor pygmaeus (KU- 86876), dentigerous process represented by sharp and elongated structure, the odontophores are absent; D. Craugastor escoses, adult female (KU- 117355), dentigerous process consists mainly of a large odontophore; E. Pristimantis danae, adult male (KU- 164062; SVL: 36 mm), elongate dentigerous process with small odontophore; F. Pristimantis orpacobates, adult female (KU- 170134; SVL: 43 mm), dentigerous process with midsize odontophore; G. Pristimantis orestes, adult female (KU- 141995; SVL: 23.5 mm), elongate dentigerous process with low odontophore; H. Pristimantis vicarius, adult male (KU- 170155; SVL: 34 mm), very elongated dentigerous process with odontophore in the distal end. Bars represent measurements of 2 mm; gray denotes cartilage and stippling denotes bones. (All drawings based on original illustrations by Lynch)." httpUri="https://zenodo.org/record/258325/files/figure.png" pageId="12" pageNumber="433">Figure 8</figureCitation>
A–C) (see also Duellman and Hedges, 2007;
<bibRefCitation id="9224BB96FFD7BD06C8993058FCD28911" author="Lehr" box="[672,799,583,608]" pageId="12" pageNumber="436" refString="Lehr, E. (2007) New eleutherodactyline frogs (Leptodactylidae: Pristimantis, Phrynopus) from Peru. Bulletin of the Museum of Comparative Zoology, 159 (2), 145 - 178. http: // dx. doi. org / 10.3099 / 0027 - 4100 (2007) 159 [145: NEFLPP] 2.0. CO; 2" type="journal article" year="2007">Lehr, 2007</bibRefCitation>
).
</paragraph>
<caption id="A2CA96EFFFD7BD06CAAE3574FB438CF0" box="[151,1166,1899,1921]" httpUri="https://zenodo.org/record/258324/files/figure.png" pageId="12" pageNumber="433" targetBox="[248,1297,679,1860]" targetPageId="12">
<paragraph id="F60AC667FFD7BD06CAAE3574FB438CF0" blockId="12.[151,1166,1899,1921]" box="[151,1166,1899,1921]" pageId="12" pageNumber="433">
<emphasis id="C4C11A75FFD7BD06CAAE3574FEDF8CF1" bold="true" box="[151,274,1899,1921]" pageId="12" pageNumber="433">FIGURE 7.</emphasis>
Graphic representation of the two roots of the discriminant analysis for morphometrics.
</paragraph>
</caption>
<caption id="A2CA96EFFFD6BD07CAAE3449FE158CF1" httpUri="https://zenodo.org/record/258325/files/figure.png" pageId="13" pageNumber="434" targetBox="[228,1345,240,1520]" targetPageId="13">
<paragraph id="F60AC667FFD6BD07CAAE3449FE158CF1" blockId="13.[151,1437,1622,1920]" pageId="13" pageNumber="434">
<emphasis id="C4C11A75FFD6BD07CAAE3449FEDE8D1A" bold="true" box="[151,275,1622,1644]" pageId="13" pageNumber="434">FIGURE 8.</emphasis>
Vomers variation of some brachyphaloids frogs. A.
<taxonomicName id="31B5BDE4FFD6BD07C91F3449FBE48D1D" box="[806,1065,1622,1644]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="434" phylum="Chordata" rank="species" species="nitidus">
<emphasis id="C4C11A75FFD6BD07C91F3449FBE48D1D" box="[806,1065,1622,1644]" italics="true" pageId="13" pageNumber="434">Eleutherodactylus nitidus</emphasis>
</taxonomicName>
(KU-102657), lacks the dentigerous process; B.
<taxonomicName id="31B5BDE4FFD6BD07CB37346BFDC68DFB" box="[270,523,1652,1674]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="434" phylum="Chordata" rank="species" species="leprus">
<emphasis id="C4C11A75FFD6BD07CB37346BFDC68DFB" box="[270,523,1652,1674]" italics="true" pageId="13" pageNumber="434">Eleutherodactylus leprus</emphasis>
</taxonomicName>
(KU-55963), dentigerous process represented by long sheet, the odontophores are absent; C.
<taxonomicName id="31B5BDE4FFD6BD07CA8D348BFE588DD8" box="[180,405,1684,1705]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="13" pageNumber="434" phylum="Chordata" rank="species" species="pygmaeus">
<emphasis id="C4C11A75FFD6BD07CA8D348BFE588DD8" box="[180,405,1684,1705]" italics="true" pageId="13" pageNumber="434">Craugastor pygmaeus</emphasis>
</taxonomicName>
(KU-86876), dentigerous process represented by sharp and elongated structure, the odontophores are absent; D.
<taxonomicName id="31B5BDE4FFD6BD07CB3334ADFE188DB9" box="[266,469,1714,1736]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="13" pageNumber="434" phylum="Chordata" rank="species" species="escoses">
<emphasis id="C4C11A75FFD6BD07CB3334ADFE188DB9" box="[266,469,1714,1736]" italics="true" pageId="13" pageNumber="434">Craugastor escoses</emphasis>
</taxonomicName>
, adult female (KU-117355), dentigerous process consists mainly of a large odontophore; E.
<taxonomicName id="31B5BDE4FFD6BD07CAAE34CEFE938D97" box="[151,350,1744,1766]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="13" pageNumber="434" phylum="Chordata" rank="species" species="danae">
<emphasis id="C4C11A75FFD6BD07CAAE34CEFE938D97" box="[151,350,1744,1766]" italics="true" pageId="13" pageNumber="434">Pristimantis danae</emphasis>
</taxonomicName>
, adult male (KU-164062; SVL: 36 mm), elongate dentigerous process with small odontophore; F.
<taxonomicName id="31B5BDE4FFD6BD07CAAE34EFFE518C74" box="[151,412,1775,1797]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="13" pageNumber="434" phylum="Chordata" rank="species" species="orpacobates">
<emphasis id="C4C11A75FFD6BD07CAAE34EFFE518C74" box="[151,412,1775,1797]" italics="true" pageId="13" pageNumber="434">Pristimantis orpacobates</emphasis>
</taxonomicName>
, adult female (KU-170134; SVL: 43 mm), dentigerous process with midsize odontophore; G.
<taxonomicName id="31B5BDE4FFD6BD07CAAE3510FEA98C55" box="[151,356,1807,1828]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="13" pageNumber="434" phylum="Chordata" rank="species" species="orestes">
<emphasis id="C4C11A75FFD6BD07CAAE3510FEA98C55" box="[151,356,1807,1828]" italics="true" pageId="13" pageNumber="434">Pristimantis orestes</emphasis>
</taxonomicName>
, adult female (KU-141995; SVL: 23.5 mm), elongate dentigerous process with low odontophore; H.
<taxonomicName id="31B5BDE4FFD6BD07CAAE3531FEA08C32" box="[151,365,1838,1859]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="13" pageNumber="434" phylum="Chordata" rank="species" species="vicarius">
<emphasis id="C4C11A75FFD6BD07CAAE3531FEA08C32" box="[151,365,1838,1859]" italics="true" pageId="13" pageNumber="434">Pristimantis vicarius</emphasis>
</taxonomicName>
, adult male (KU-170155; SVL: 34 mm), very elongated dentigerous process with odontophore in the distal end. Bars represent measurements of 2 mm; gray denotes cartilage and stippling denotes bones. (All drawings based on original illustrations by Lynch).
</paragraph>
</caption>
<paragraph id="F60AC667FFD5BD04CAFE3288FD508A15" blockId="14.[151,1437,151,861]" pageId="14" pageNumber="435">
Another important osteological feature in the taxonomy of brachycephaloid frogs are the oblique keels.
<bibRefCitation id="9224BB96FFD5BD04CF6F3288FF2C8BA5" author="Lynch" pageId="14" pageNumber="436" refString="Lynch, J. D. (1980) New species of Eleutherodactylus of Colombia (Amphibia: Leptodactylidae). I: Five new species from the paramos of the Cordillera Central. Caldasia, 13 (61), 165 - 188." type="journal article" year="1980">Lynch (1980)</bibRefCitation>
first mentioned oblique keels but did not define this character. I define the oblique keels as a ridge concealed or not by the buccal mucosa, and formed by the anterior elongated portion of the posterior process of vomers (
<figureCitation id="6E8EDAE2FFD5BD04CB39331BFEA78A6C" box="[256,362,260,285]" captionStart="FIGURE 5" captionStartId="8.[151,250,1725,1747]" captionTargetBox="[151,1436,410,1704]" captionTargetId="figure@8.[151,1436,373,1704]" captionTargetPageId="8" captionText="FIGURE 5. Cranium of adult female of P. stictus (ICN- 55591; SVL: 25.6 mm) in dorsal (A) and ventral view (B), bars represent measurements of 2.5 mm, black indicates foramina, gray cartilage and stippling bones; C. Photography of oblique keel of vomer slightly concealed by mucosa in P. stictus (ICN- 55697; paratype); D. Vomer in cleared and stained specimen of adult female of P. stictus (ICN- 55692)." httpUri="https://zenodo.org/record/258322/files/figure.png" pageId="14" pageNumber="435">Figure 5</figureCitation>
). It is important to note that the oblique keels and the vomerine odontophores are not homologous. Oblique keels are exposed anteriorly to the odontophores in the posterior process of vomers, while odontophores and teeth may be present or not.
</paragraph>
<paragraph id="F60AC667FFD5BD04CAFE3370FA50882C" blockId="14.[151,1437,151,861]" pageId="14" pageNumber="435">
The vomerine odontophore is the ossified and exposed part of the posterior vomer processes. It may bear teeth and its structure is variable. In brachycephaloids, the dentigerous process can be either absent (
<figureCitation id="6E8EDAE2FFD5BD04CEF6338BFAE38ADD" box="[1231,1326,404,429]" captionStart="FIGURE 8" captionStartId="13.[151,250,1622,1644]" captionTargetBox="[228,1345,240,1520]" captionTargetId="figure@13.[188,1398,193,1600]" captionTargetPageId="13" captionText="FIGURE 8. Vomers variation of some brachyphaloids frogs. A. Eleutherodactylus nitidus (KU- 102657), lacks the dentigerous process; B. Eleutherodactylus leprus (KU- 55963), dentigerous process represented by long sheet, the odontophores are absent; C. Craugastor pygmaeus (KU- 86876), dentigerous process represented by sharp and elongated structure, the odontophores are absent; D. Craugastor escoses, adult female (KU- 117355), dentigerous process consists mainly of a large odontophore; E. Pristimantis danae, adult male (KU- 164062; SVL: 36 mm), elongate dentigerous process with small odontophore; F. Pristimantis orpacobates, adult female (KU- 170134; SVL: 43 mm), dentigerous process with midsize odontophore; G. Pristimantis orestes, adult female (KU- 141995; SVL: 23.5 mm), elongate dentigerous process with low odontophore; H. Pristimantis vicarius, adult male (KU- 170155; SVL: 34 mm), very elongated dentigerous process with odontophore in the distal end. Bars represent measurements of 2 mm; gray denotes cartilage and stippling denotes bones. (All drawings based on original illustrations by Lynch)." httpUri="https://zenodo.org/record/258325/files/figure.png" pageId="14" pageNumber="435">Figure 8</figureCitation>
A—some
<taxonomicName id="31B5BDE4FFD5BD04CAAE33A7FEAF8AA1" box="[151,354,440,464]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="435" phylum="Chordata" rank="genus">
<emphasis id="C4C11A75FFD5BD04CAAE33A7FEAF8AA1" box="[151,354,440,464]" italics="true" pageId="14" pageNumber="435">Eleutherodactylus</emphasis>
</taxonomicName>
and
<taxonomicName id="31B5BDE4FFD5BD04CB9933A7FDCD8AA1" box="[416,512,440,464]" class="Amphibia" family="Strabomantidae" genus="Noblella" kingdom="Animalia" order="Anura" pageId="14" pageNumber="435" phylum="Chordata" rank="genus">
<emphasis id="C4C11A75FFD5BD04CB9933A7FDCD8AA1" box="[416,512,440,464]" italics="true" pageId="14" pageNumber="435">Noblella</emphasis>
</taxonomicName>
), short and mainly formed by the odontophores (
<figureCitation id="6E8EDAE2FFD5BD04CE7F33A8FB678AA1" box="[1094,1194,439,464]" captionStart="FIGURE 8" captionStartId="13.[151,250,1622,1644]" captionTargetBox="[228,1345,240,1520]" captionTargetId="figure@13.[188,1398,193,1600]" captionTargetPageId="13" captionText="FIGURE 8. Vomers variation of some brachyphaloids frogs. A. Eleutherodactylus nitidus (KU- 102657), lacks the dentigerous process; B. Eleutherodactylus leprus (KU- 55963), dentigerous process represented by long sheet, the odontophores are absent; C. Craugastor pygmaeus (KU- 86876), dentigerous process represented by sharp and elongated structure, the odontophores are absent; D. Craugastor escoses, adult female (KU- 117355), dentigerous process consists mainly of a large odontophore; E. Pristimantis danae, adult male (KU- 164062; SVL: 36 mm), elongate dentigerous process with small odontophore; F. Pristimantis orpacobates, adult female (KU- 170134; SVL: 43 mm), dentigerous process with midsize odontophore; G. Pristimantis orestes, adult female (KU- 141995; SVL: 23.5 mm), elongate dentigerous process with low odontophore; H. Pristimantis vicarius, adult male (KU- 170155; SVL: 34 mm), very elongated dentigerous process with odontophore in the distal end. Bars represent measurements of 2 mm; gray denotes cartilage and stippling denotes bones. (All drawings based on original illustrations by Lynch)." httpUri="https://zenodo.org/record/258325/files/figure.png" pageId="14" pageNumber="435">Figure 8</figureCitation>
D), elongated and of different size, shape, and position, but bearing the odontophore at the posterior end (
<figureCitation id="6E8EDAE2FFD5BD04CE6A33C3FB7F8A85" box="[1107,1202,476,501]" captionStart="FIGURE 8" captionStartId="13.[151,250,1622,1644]" captionTargetBox="[228,1345,240,1520]" captionTargetId="figure@13.[188,1398,193,1600]" captionTargetPageId="13" captionText="FIGURE 8. Vomers variation of some brachyphaloids frogs. A. Eleutherodactylus nitidus (KU- 102657), lacks the dentigerous process; B. Eleutherodactylus leprus (KU- 55963), dentigerous process represented by long sheet, the odontophores are absent; C. Craugastor pygmaeus (KU- 86876), dentigerous process represented by sharp and elongated structure, the odontophores are absent; D. Craugastor escoses, adult female (KU- 117355), dentigerous process consists mainly of a large odontophore; E. Pristimantis danae, adult male (KU- 164062; SVL: 36 mm), elongate dentigerous process with small odontophore; F. Pristimantis orpacobates, adult female (KU- 170134; SVL: 43 mm), dentigerous process with midsize odontophore; G. Pristimantis orestes, adult female (KU- 141995; SVL: 23.5 mm), elongate dentigerous process with low odontophore; H. Pristimantis vicarius, adult male (KU- 170155; SVL: 34 mm), very elongated dentigerous process with odontophore in the distal end. Bars represent measurements of 2 mm; gray denotes cartilage and stippling denotes bones. (All drawings based on original illustrations by Lynch)." httpUri="https://zenodo.org/record/258325/files/figure.png" pageId="14" pageNumber="435">Figure 8</figureCitation>
E–H), or lacking the odontophore (Figure B–C) (
<emphasis id="C4C11A75FFD5BD04CBE6301EFDCC8969" box="[479,513,513,536]" italics="true" pageId="14" pageNumber="435">e.g</emphasis>
.,
<taxonomicName id="31B5BDE4FFD5BD04C822301FFCCA8969" box="[539,775,512,536]" class="Amphibia" family="Strabomantidae" genus="Bryophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="435" phylum="Chordata" rank="species" species="cophites">
<emphasis id="C4C11A75FFD5BD04C822301FFCCA8969" box="[539,775,512,536]" italics="true" pageId="14" pageNumber="435">Bryophryne cophites</emphasis>
</taxonomicName>
,
<taxonomicName id="31B5BDE4FFD5BD04C923301FFBDA8969" box="[794,1047,512,536]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="14" pageNumber="435" phylum="Chordata" rank="species" species="pygmaeus">
<emphasis id="C4C11A75FFD5BD04C923301FFBDA8969" box="[794,1047,512,536]" italics="true" pageId="14" pageNumber="435">Craugastor pygmaeus</emphasis>
</taxonomicName>
,
<taxonomicName id="31B5BDE4FFD5BD04CE10301FFABC8969" box="[1065,1393,512,536]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="435" phylum="Chordata" rank="species" species="marnockii">
<emphasis id="C4C11A75FFD5BD04CE10301FFABC8969" box="[1065,1393,512,536]" italics="true" pageId="14" pageNumber="435">Eleutherodactylus marnockii</emphasis>
</taxonomicName>
,
<taxonomicName id="31B5BDE4FFD5BD04CFBD301EFECC894D" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="435" phylum="Chordata" rank="species" species="rufescens">
<emphasis id="C4C11A75FFD5BD04CFBD301EFA588969" box="[1412,1429,513,536]" italics="true" pageId="14" pageNumber="435">E</emphasis>
.
<emphasis id="C4C11A75FFD5BD04CAAE303BFECC894D" box="[151,257,548,572]" italics="true" pageId="14" pageNumber="435">rufescens</emphasis>
</taxonomicName>
,
<taxonomicName id="31B5BDE4FFD5BD04CB28303AFE5C894D" box="[273,401,548,572]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="435" phylum="Chordata" rank="species" species="longipes">
<emphasis id="C4C11A75FFD5BD04CB28303AFEEF894D" box="[273,290,549,572]" italics="true" pageId="14" pageNumber="435">E</emphasis>
.
<emphasis id="C4C11A75FFD5BD04CB0B303BFE5C894D" box="[306,401,548,572]" italics="true" pageId="14" pageNumber="435">longipes</emphasis>
</taxonomicName>
,
<taxonomicName id="31B5BDE4FFD5BD04CB9B303AFDC4894D" box="[418,521,548,572]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="435" phylum="Chordata" rank="species" species="dixoni">
<emphasis id="C4C11A75FFD5BD04CB9B303AFE7E894D" box="[418,435,549,572]" italics="true" pageId="14" pageNumber="435">E</emphasis>
.
<emphasis id="C4C11A75FFD5BD04CBFA303BFDC4894D" box="[451,521,548,572]" italics="true" pageId="14" pageNumber="435">dixoni</emphasis>
</taxonomicName>
,
<taxonomicName id="31B5BDE4FFD5BD04C820303AFD4C894D" box="[537,641,548,572]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="435" phylum="Chordata" rank="species" species="leprus">
<emphasis id="C4C11A75FFD5BD04C820303AFDE7894D" box="[537,554,549,572]" italics="true" pageId="14" pageNumber="435">E</emphasis>
.
<emphasis id="C4C11A75FFD5BD04C802303BFD4C894D" box="[571,641,548,572]" italics="true" pageId="14" pageNumber="435">leprus</emphasis>
</taxonomicName>
).
<taxonomicName id="31B5BDE4FFD5BD04C8A2303AFCE9894D" box="[667,804,549,572]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="14" pageNumber="435" phylum="Chordata" rank="genus">
<emphasis id="C4C11A75FFD5BD04C8A2303AFCE9894D" box="[667,804,549,572]" italics="true" pageId="14" pageNumber="435">Pristimantis</emphasis>
</taxonomicName>
of the
<taxonomicName id="31B5BDE4FFD5BD04C940303AFBD7894D" box="[889,1050,548,572]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="14" pageNumber="435" phylum="Chordata" rank="species" species="leptolophus">
<emphasis id="C4C11A75FFD5BD04C940303AFC4A894D" box="[889,903,549,572]" italics="true" pageId="14" pageNumber="435">P</emphasis>
.
<emphasis id="C4C11A75FFD5BD04C9AE303BFBD7894D" box="[919,1050,548,572]" italics="true" pageId="14" pageNumber="435">leptolophus</emphasis>
</taxonomicName>
group of the paramos, such as
<taxonomicName id="31B5BDE4FFD5BD04CFBE303AFF12892E" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="14" pageNumber="435" phylum="Chordata" rank="species" species="stictus">
<emphasis id="C4C11A75FFD5BD04CFBE303AFA58894D" box="[1415,1429,549,572]" italics="true" pageId="14" pageNumber="435">P</emphasis>
.
<emphasis id="C4C11A75FFD5BD04CAAE3057FF12892E" box="[151,223,584,607]" italics="true" pageId="14" pageNumber="435">stictus</emphasis>
</taxonomicName>
,
<taxonomicName id="31B5BDE4FFD5BD04CACA3056FE558911" box="[243,408,584,608]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="14" pageNumber="435" phylum="Chordata" rank="species" species="leptolophus">
<emphasis id="C4C11A75FFD5BD04CACA3056FECC8911" box="[243,257,585,608]" italics="true" pageId="14" pageNumber="435">P</emphasis>
.
<emphasis id="C4C11A75FFD5BD04CB2C3057FE558911" box="[277,408,584,608]" italics="true" pageId="14" pageNumber="435">leptolophus</emphasis>
</taxonomicName>
, and
<taxonomicName id="31B5BDE4FFD5BD04CBDB3056FD478911" box="[482,650,584,608]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="14" pageNumber="435" phylum="Chordata" rank="species" species="lasalleorum">
<emphasis id="C4C11A75FFD5BD04CBDB3056FE3D8911" box="[482,496,585,608]" italics="true" pageId="14" pageNumber="435">P</emphasis>
.
<emphasis id="C4C11A75FFD5BD04C83D3057FD478911" box="[516,650,584,608]" italics="true" pageId="14" pageNumber="435">lasalleorum</emphasis>
</taxonomicName>
, lack odontophores and vomerine teeth. A thorough study of the variation of these characters in brachycephaloid frogs is required, especially in the poorly know genus
<taxonomicName id="31B5BDE4FFD5BD04CF353072FA5889F5" box="[1292,1429,621,644]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="14" pageNumber="435" phylum="Chordata" rank="genus">
<emphasis id="C4C11A75FFD5BD04CF353072FA5889F5" box="[1292,1429,621,644]" italics="true" pageId="14" pageNumber="435">Pristimantis</emphasis>
</taxonomicName>
. On the other hand, not only is the morphology of
<taxonomicName id="31B5BDE4FFD5BD04C8D9308FFCA489D6" box="[736,873,656,679]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="14" pageNumber="435" phylum="Chordata" rank="genus">
<emphasis id="C4C11A75FFD5BD04C8D9308FFCA489D6" box="[736,873,656,679]" italics="true" pageId="14" pageNumber="435">Pristimantis</emphasis>
</taxonomicName>
poorly known, but also its species diversity and geographic distribution. For example, the paramos of
<collectingCountry id="8EA286F7FFD5BD04C8D630ABFC9089BC" box="[751,861,692,717]" name="Colombia" pageId="14" pageNumber="435">Colombia</collectingCountry>
have so far been little explored, and the knowledge on amphibian communities in the different highland areas is rudimentary. The allopatric distributions of species on paramo areas of mountain tops suggest an important role of vicariant speciation in highland species. Indeed, the presence of endemic species (
<figureCitation id="6E8EDAE2FFD5BD04CBDB3100FD8E8849" box="[482,579,799,824]" captionStart="FIGURE 6" captionStartId="10.[151,250,1542,1564]" captionTargetBox="[319,1268,216,1504]" captionTargetId="figure@10.[302,1284,193,1520]" captionTargetPageId="10" captionText="FIGURE 6. Map of Colombia showing the distribution of Pristimantis stictus sp. nov. and other species. Black circles indicate localities of P. stictus sp. nov., and a white circle signals the type locality. Distribution of other species in the P. leptolophus group: P. maculosus (black five pointed star), P. parectatus (white diamond), P. lasalleorum (inverted triangle), P. uranobates (white four pointed star), P. peraticus (black six pointed star), P. scoloblepharus (black triangle), and P. leptolophus (white squares)." httpUri="https://zenodo.org/record/258323/files/figure.png" pageId="14" pageNumber="435">Figure 6</figureCitation>
) (
<bibRefCitation id="9224BB96FFD5BD04C8633100FC9D8849" author="Bernal" box="[602,848,799,824]" pageId="14" pageNumber="435" refString="Bernal, M. H. &amp; Lynch, J. D. (2008) Review and analysis of altitudinal distribution of the Andean anurans in Colombia. Zootaxa, 1826 (1), 1 - 25. http: // dx. doi. org / 10.11646 / zootaxa. 1826.1.1" type="journal article" year="2008">Bernal &amp; Lynch 2008</bibRefCitation>
) in isolated paramos suggests that other undescribed species could await discovery in the many isolated and unexplored paramos of the Cordillera Central of
<collectingCountry id="8EA286F7FFD5BD04CF1E315BFA54882C" box="[1319,1433,836,861]" name="Colombia" pageId="14" pageNumber="435">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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