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<document id= "179DE74C04B70E5EEC6BA8DAEDB55792" ID-CLB-Dataset= "34195" ID-DOI= "http://dx.doi.org/10.3897/zookeys.228.3401" ID-GBIF-Dataset= "14392d6c-4f18-4c00-863b-4d0c709409a5" ID-PMC= "PMC11321467" ID-Pensoft-Pub= "1313-2970-228-77" ID-PubMed= "39139796" ModsDocAuthor= "" ModsDocDate= "2012" ModsDocID= "1313-2970-228-77" ModsDocOrigin= "ZooKeys 228" ModsDocTitle= "Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines" checkinTime= "1451248671826" checkinUser= "pensoft" docAuthor= "Zhao, Danyang & Tian, Mingyi" docDate= "2012" docId= "7E0E3BFCD8260C9B7D3852414329D2DC" docLanguage= "en" docName= "ZooKeys 228: 77-100" docOrigin= "ZooKeys 228" docSource= "http://dx.doi.org/10.3897/zookeys.228.3401" docTitle= "Macrocheilus niger Andrewes 1920" docType= "treatment" docVersion= "4" lastPageNumber= "84" masterDocId= "FFD6FFE3FF8DFFD04741FF923F7CFFAC" masterDocTitle= "Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines" masterLastPageNumber= "100" masterPageNumber= "77" pageNumber= "84" updateTime= "1732929750356" updateUser= "ExternalLinkService" >
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<mods:title id= "29D682D8232B623F419D82469D586B14" > Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines</mods:title>
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<mods:namePart id= "EEE4E4C06D3B111F036651A429699CD3" > Zhao, Danyang</mods:namePart>
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<mods:namePart id= "8863EEBD6C43AA98114EBEFD94738A00" > Tian, Mingyi</mods:namePart>
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<mods:typeOfResource id= "0836BC97245BD8BFDA87DA669EF44E3F" > text</mods:typeOfResource>
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<mods:title id= "E603A34ECA5D359059A2624D09B8B9B9" > ZooKeys</mods:title>
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<mods:date id= "3BE56EB807A1D52934FB2F41BBC8E68C" > 2012</mods:date>
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<treatment id= "7E0E3BFCD8260C9B7D3852414329D2DC" ID-GBIF-Taxon= "152037331" LSID= "urn:lsid:plazi:treatment:7E0E3BFCD8260C9B7D3852414329D2DC" httpUri= "http://treatment.plazi.org/id/7E0E3BFCD8260C9B7D3852414329D2DC" lastPageId= "8" lastPageNumber= "84" pageId= "7" pageNumber= "84" scope_family= "Carabidae" scope_order= "Coleoptera" scope_tribe= "Helluonini" >
<subSubSection id= "3B81123D941A5F9DB4D4BC620A6723CC" pageId= "7" pageNumber= "84" type= "nomenclature" >
<paragraph id= "F9D7FA352C3F7DDEDAF9A78C902A4077" pageId= "7" pageNumber= "84" >
<taxonomicName id= "4A59EB8C041B8667421A702D38714D2D" ID-CoL= "3X3VM" LSID= "http://species-id.net/wiki/Macrocheilus_niger" authority= "Andrewes, 1920" authorityName= "Andrewes" authorityYear= "1920" class= "Insecta" family= "Carabidae" genus= "Macrocheilus" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Macrocheilus niger" order= "Coleoptera" pageId= "7" pageNumber= "84" phylum= "Arthropoda" rank= "species" species= "niger" > Macrocheilus niger Andrewes, 1920</taxonomicName>
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Figs 4, 12, 20, 29, 30, 39, 46
</paragraph>
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<subSubSection id= "1FE9B3DA42E5E8EA31E194A8C913E0E0" pageId= "7" pageNumber= "84" type= "reference_group" >
<paragraph id= "144853F05C5AD92902B94B50C1DD3FB0" pageId= "7" pageNumber= "84" >
<taxonomicName id= "DBEFC2119F51F298BE6263FC03BDA2F2" class= "Insecta" family= "Carabidae" genus= "Macrocheilus" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Macrocheilus niger" order= "Coleoptera" pageId= "7" pageNumber= "84" phylum= "Arthropoda" rank= "species" species= "niger" > Macrocheilus niger</taxonomicName>
<bibRefCitation id= "62363D96A64ACAFBC1D8063ED93BCDE8" author= "Andrewes, HE" journalOrPublisher= "The Annals and Magazine of Natural History," pageId= "14" pageNumber= "91" pagination= "493 - 506" title= "Papers on Oriental Carabidae. - V." url= "10.1080/00222932008632476" volume= "9" year= "1920" > Andrewes 1920</bibRefCitation>
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: 499.
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<bibRefCitation id= "A5F955E22649C0A6D4D5A14928F82C1F" pageId= "7" pageNumber= "84" > Csiki 1932</bibRefCitation>
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: 1574;
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<bibRefCitation id= "F7E47D3282A9D2E3C672B5AFB3772556" pageId= "7" pageNumber= "84" > Lorenz 2005</bibRefCitation>
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: 512. Type locality: India (Nilgiri Hills), deposited in NHML.
</paragraph>
</subSubSection>
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<subSubSection id= "A6B23C8E3A2398A6C247CEB79CA28643" pageId= "7" pageNumber= "84" type= "diagnosis" >
<paragraph id= "7E6575C2E209929862F7B84295F8DB2A" pageId= "7" pageNumber= "84" > Diagnosis.</paragraph>
<paragraph id= "BF706071A4B0557AC7363E4A3E7BD93E" pageId= "7" pageNumber= "84" > Length 16.0-16.3 mm, width 6.0-6.2 mm. Labrum (Fig. 4) convex, with a short furrow on each side of base, narrowed and pointed in front, with three pairs of setae away from the margin; ligula thickened, apex truncate towards sides, apical margin slightly emarginate, deeply depressed towards base, with a small median impression near apex; mentum (Fig. 12) glabrous at base; median tooth shorter than lobes, rather narrow and with apex rather pointed, two pairs of setae at basal area; lateral lobes obtuse at apex. Palpi (Fig. 20) not dilated. Elytral spots transverse, close to the middle of interval 5, covering intervals 3-7.</paragraph>
<paragraph id= "72D02F3A781949D12E3E02F08D4A33E6" pageId= "7" pageNumber= "84" > Male genitalia. Median lobe stout, apical lamella short and round at apex (Figs 29-30).</paragraph>
<paragraph id= "402489F681C0FDF2E0BC38205BF07AC8" pageId= "7" pageNumber= "84" > Female genitalia. Gonocoxite subapically dilated, three setae on dorsal surface, sharp at apex (Fig. 39).</paragraph>
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</subSubSection>
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<subSubSection id= "2AAC49FE5CAC475C6B2F49CB656BDDB1" pageId= "7" pageNumber= "84" type= "remarks" >
<paragraph id= "DA45259736ACB0646EBBC5AD4EE2F34E" pageId= "7" pageNumber= "84" > Remarks.</paragraph>
<paragraph id= "5557375DF9C7EF3B8D6F0DB48F13DFB5" pageId= "7" pageNumber= "84" >
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Similar to
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<taxonomicName id= "50B48ED5F6E563453703A1EE497177CB" class= "Insecta" family= "Carabidae" genus= "Macrocheilus" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Macrocheilus asteriscus" order= "Coleoptera" pageId= "7" pageNumber= "84" phylum= "Arthropoda" rank= "species" species= "asteriscus" > Macrocheilus asteriscus</taxonomicName>
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, but differs with the tooth of mentum plurisetose and elytral spot not cruciform.
</paragraph>
</subSubSection>
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<subSubSection id= "7A723706C6A8D5BF5B050F1671FD3E76" pageId= "7" pageNumber= "84" type= "materials examined" >
<paragraph id= "64AE6030557BFAC5457FCEABCEC858A4" pageId= "7" pageNumber= "84" > Materials examined.</paragraph>
<paragraph id= "F54D998B4FD16F9E2B0BA5A3AC34D52E" pageId= "7" pageNumber= "84" >
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1 female, the holotype, labeled " type, 11, H. L. Andrewes, Nilgiri Hills, H. E. Andrewes Coll., B.M. 1945-97.;
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<taxonomicName id= "7FAD482663A33368295F4EAC58E3E581" class= "Insecta" family= "Carabidae" genus= "Macrochilus" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Macrochilus niger" order= "Coleoptera" pageId= "7" pageNumber= "84" phylum= "Arthropoda" rank= "species" species= "niger" > Macrochilus niger</taxonomicName>
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Type-, Andr., H. E. Andrewes det.;
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<taxonomicName id= "ED4BC7D20C12D70FC1507D07BE5BE428" class= "Insecta" family= "Carabidae" genus= "Macrochilus" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Macrochilus niger" order= "Coleoptera" pageId= "7" pageNumber= "84" phylum= "Arthropoda" rank= "species" species= "niger" > Macrochilus niger</taxonomicName>
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Andrewes, 1920" , deposited in NHML. 1 male and 1 female, " Dehra-Dun 1940-45, Kumaon Himalaya, Liesenfeldt leg." , deposited in MNHN.
</paragraph>
</subSubSection>
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<subSubSection id= "9A5AE327DC4AE1A00F4084F3AB5F4623" lastPageId= "8" lastPageNumber= "85" pageId= "7" pageNumber= "84" type= "distribution" >
<paragraph id= "CB538F957E9325E6470A36AAA4F8D416" pageId= "7" pageNumber= "84" > Distribution.</paragraph>
<paragraph id= "E0741A93DC9806663859B8F9533CC846" lastPageId= "8" lastPageNumber= "85" pageId= "7" pageNumber= "84" >
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India (Nilgiri Hills, Malabar, Madras and Bombay) and Sri Lanka
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<pageBreakToken id= "AC3534D825DEE4A9E07A7065DEBDAE71" pageId= "8" pageNumber= "85" start= "start" > .</pageBreakToken>
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</paragraph>
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</treatment>
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