treatments-xml/data/74/12/87/741287F4FFE78C21FF25F99C96CF2799.xml

<document id="739FFDB9751DBC6A3863E5AC2B15E57A" ID-DOI="10.11646/zootaxa.4044.2.1" ID-GBIF-Dataset="4be691ce-6649-4fc2-9c81-b54c72c2179d" ID-ISSN="1175-5326" ID-Zenodo-Dep="235030" ID-ZooBank="4ABB85F0-D989-4702-9962-BA6E4D26BDE2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461338415922" checkinUser="plazi" docAuthor="Kupriyanova, Elena K. &amp; Ippolitov, Alexei P." docDate="2015" docId="741287F4FFE78C21FF25F99C96CF2799" docLanguage="en" docName="zt04044p200.pdf" docOrigin="Zootaxa 4044 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hyalopomatus dieteri Kupriyanova &amp; Ippolitov, 2015, sp. nov." docType="treatment" docVersion="10" lastPageNumber="180" masterDocId="882BFF8CFFFD8C3CFFB2FFBD972B2071" masterDocTitle="Deep-sea serpulids (Annelida: Polychaeta) in tetragonal tubes: on a tube convergence path from the Mesozoic to Recent" masterLastPageNumber="200" masterPageNumber="151" pageNumber="177" updateTime="1698619758002" updateUser="plazi">
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<mods:title id="95FE4DF0962EFAE67C6151E4D8BEDAD5">Deep-sea serpulids (Annelida: Polychaeta) in tetragonal tubes: on a tube convergence path from the Mesozoic to Recent</mods:title>
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<mods:namePart id="A7881762B8E71AC0C40E94E7EF0DCFC1">Kupriyanova, Elena K.</mods:namePart>
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<mods:namePart id="3045BD1BF3795A6717129E380667FC00">Ippolitov, Alexei P.</mods:namePart>
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<treatment id="741287F4FFE78C21FF25F99C96CF2799" ID-DOI="http://doi.org/10.5281/zenodo.5324188" ID-GBIF-Taxon="119644917" ID-Zenodo-Dep="5324188" LSID="urn:lsid:plazi:treatment:741287F4FFE78C21FF25F99C96CF2799" httpUri="http://treatment.plazi.org/id/741287F4FFE78C21FF25F99C96CF2799" lastPageId="29" lastPageNumber="180" pageId="26" pageNumber="177">
<subSubSection id="B4A16569FFE78C26FF25F99C964B262C" pageId="26" pageNumber="177" type="nomenclature">
<paragraph id="FC0436E2FFE78C26FF25F99C952B264A" blockId="26.[151,512,1569,1629]" box="[151,512,1569,1595]" pageId="26" pageNumber="177">
<heading id="A74C818EFFE78C26FF25F99C952B264A" bold="true" box="[151,512,1569,1595]" fontSize="11" level="1" pageId="26" pageNumber="177" reason="1">
<emphasis id="CECFEAF0FFE78C26FF25F99C952B264A" bold="true" box="[151,512,1569,1595]" pageId="26" pageNumber="177">
<taxonomicName id="3BBB4D61FFE78C26FF25F99C96B0264A" ID-CoL="3MWYS" box="[151,411,1569,1595]" class="Polychaeta" family="Serpulidae" genus="Hyalopomatus" kingdom="Animalia" order="Sabellida" pageId="26" pageNumber="177" phylum="Annelida" rank="species" species="dieteri" status="sp. nov.">
<emphasis id="CECFEAF0FFE78C26FF25F99C96B0264A" bold="true" box="[151,411,1569,1595]" italics="true" pageId="26" pageNumber="177">Hyalopomatus dieteri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D5FC578BFFE78C26FE10F99C952B264A" box="[418,512,1569,1595]" pageId="26" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="FC0436E2FFE78C26FF25F9F9964B262C" blockId="26.[151,512,1569,1629]" box="[151,352,1604,1629]" pageId="26" pageNumber="177">
<figureCitation id="64802A67FFE78C26FF25F9F997D7262C" box="[151,252,1604,1629]" captionStart="FIGURE 1" captionStartId="3.[151,250,1766,1788]" captionTargetBox="[244,1343,195,1750]" captionTargetId="figure@3.[244,1343,195,1751]" captionTargetPageId="3" captionText="FIGURE 1. External morphology of polygonal serpulid tubes. Recent species: A— Spirodiscus grimaldii, R / V VITYAZ- 2, St. 78. B, C— Bathyditrupa hovei, R / V VITYAZ- 2, St. 79. D— Spirodiscus groenlandicus comb. nov., R / V VITYAZ- 2, St. 79. E— Bathyvermilia gregrousei sp. nov., R / V VITYAZ, St. 5074. F— Hyalopomatus dieteri sp. nov., R / V JEAN CHARCOT, cruise BIOCAL, St. CP 27. G— Zibrovermilia zibrowii gen. et sp. nov., R / V ALIS, cruise MUSORSTOM 6, St. CP 438. Fossil tubes: H, I— Nogrobs cf. vermicularis, PIN 5485 / 23. J – K— Glandifera cf. vertebralis, spirally coiled juvenile tube with median keel running along the lateral tube side, PIN 5485 / 02. L, M— Glandifera cf. vertebralis, a fragment of straight adult tube with regular peristomes, PIN 5485 / 01. N, O— Tetraserpula barremica, PIN 5485 / 06. P, Q— Tetraditrupa canteriata, PIN 5485 / 04. R – T— Tubulostium discoideum, ARI Od 224 / 70 (the specimen figured by Chiplonkar &amp; Tapaswi, 1973, pl. 11, fig. 15)." httpUri="https://zenodo.org/record/235031/files/figure.png" pageId="26" pageNumber="177">Figures 1</figureCitation>
F, 11, 12
</paragraph>
</subSubSection>
<subSubSection id="B4A16569FFE78C26FF25F93192B72744" pageId="26" pageNumber="177" type="materials_examined">
<paragraph id="FC0436E2FFE78C26FF25F93195BB269C" blockId="26.[151,1437,1676,1989]" pageId="26" pageNumber="177">
<emphasis id="CECFEAF0FFE78C26FF25F93196AA26D4" bold="true" box="[151,385,1676,1701]" pageId="26" pageNumber="177">Material examined.</emphasis>
R/V CORIOLIS, cruise BIOGEOCAL, 1987, East of 
<collectingCountry id="84AC7672FFE78C26FC55F93093B226D4" box="[999,1177,1676,1701]" name="New Caledonia" pageId="26" pageNumber="177">New Caledonia</collectingCountry>
, St. CP260: 
<date id="88051022FFE78C26FA97F93092BC26D4" box="[1317,1431,1677,1701]" pageId="26" pageNumber="177" value="1987-04-17">17.4.1987</date>
, 
<geoCoordinate id="998F5025FFE78C26FF25F90D963E26B9" box="[151,277,1712,1736]" direction="south" orientation="latitude" pageId="26" pageNumber="177" precision="9" value="-21.0">21°00.00'S</geoCoordinate>
, 
<geoCoordinate id="998F5025FFE78C26FE92F90D968626B8" box="[288,429,1712,1737]" direction="east" orientation="longitude" pageId="26" pageNumber="177" precision="9" value="167.97234">167°58.34'E</geoCoordinate>
, 
<quantity id="3B439B07FFE78C26FE0BF90D957E26B9" box="[441,597,1712,1736]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="1.98" metricValueMin="1.8199999999999998" pageId="26" pageNumber="177" unit="m" value="1900.0" valueMax="1980.0" valueMin="1820.0">1820–1980 m</quantity>
(
<emphasis id="CECFEAF0FFE78C26FDD7F90D940626B8" bold="true" box="[613,813,1712,1737]" pageId="26" pageNumber="177">
<typeStatus id="23008840FFE78C26FDD7F90D95E726B8" box="[613,716,1712,1737]" pageId="26" pageNumber="177" type="holotype">holotype</typeStatus>
<collectionCode id="9AAAAE27FFE78C26FD66F90D940626B8" LSID="urn:lsid:biocol.org:col:34988" box="[724,813,1712,1737]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="26" pageNumber="177">MNHN</collectionCode>
</emphasis>
<collectionCode id="9AAAAE27FFE78C26FC86F90D945126B9" box="[820,890,1712,1736]" pageId="26" pageNumber="177">POLY</collectionCode>
<typeStatus id="23008840FFE78C26FC30F90C94E326B9" box="[898,968,1713,1736]" pageId="26" pageNumber="177">
<collectionCode id="9AAAAE27FFE78C26FC30F90C94E326B9" box="[898,968,1713,1736]" pageId="26" pageNumber="177">TYPE</collectionCode>
</typeStatus>
1564, 1 tube fragment prepared for 
<collectionCode id="9AAAAE27FFE78C26FAD0F90D92B726B9" box="[1378,1436,1712,1736]" pageId="26" pageNumber="177">SEM</collectionCode>
and X-ray diffraction analysis 
<emphasis id="CECFEAF0FFE78C26FE5EF9699537269C" bold="true" box="[492,540,1748,1773]" pageId="26" pageNumber="177">
<collectionCode id="9AAAAE27FFE78C26FE5EF9699537269C" box="[492,540,1748,1773]" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="26" pageNumber="177">PIN</collectionCode>
</emphasis>
5485/16).
</paragraph>
<paragraph id="FC0436E2FFE78C26FF75F94592B72744" blockId="26.[151,1437,1676,1989]" pageId="26" pageNumber="177">
R/V 
<collectionCode id="9AAAAE27FFE78C26FEB2F94496FE2761" box="[256,469,1784,1809]" pageId="26" pageNumber="177">JEAN CHARCOT</collectionCode>
, cruise BIOCAL, 1985, approximately 
<quantity id="3B439B07FFE78C26FC2CF94594DD2760" box="[926,1014,1784,1809]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="26" pageNumber="177" unit="km" value="100.0">100 km</quantity>
South of 
<collectingCountry id="84AC7672FFE78C26FBDFF944920B2760" box="[1133,1312,1784,1809]" name="New Caledonia" pageId="26" pageNumber="177">New Caledonia</collectingCountry>
, St. CP27, 
<date id="88051022FFE78C26FF25F8A196222744" box="[151,265,1820,1845]" pageId="26" pageNumber="177" value="1985-08-29">29.8.1985</date>
, 
<geoCoordinate id="998F5025FFE78C26FEA7F8A196B92744" box="[277,402,1820,1845]" direction="south" orientation="latitude" pageId="26" pageNumber="177" precision="9" value="-23.092">23°05.52'S</geoCoordinate>
, 
<geoCoordinate id="998F5025FFE78C26FE2FF8A195012744" box="[413,554,1820,1845]" direction="east" orientation="longitude" pageId="26" pageNumber="177" precision="9" value="166.44017">166°26.41'E</geoCoordinate>
, 
<quantity id="3B439B07FFE78C26FD87F8A095A12745" box="[565,650,1821,1845]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="26" pageNumber="177" unit="m" value="1900.0">1900 m</quantity>
(1 
<typeStatus id="23008840FFE78C26FD1DF8A094242744" box="[687,783,1821,1845]" pageId="26" pageNumber="177" type="paratype">paratype</typeStatus>
without radioles 
<emphasis id="CECFEAF0FFE78C26FC60F8A1932B2744" bold="true" box="[978,1024,1820,1845]" pageId="26" pageNumber="177">AM</emphasis>
W.46386, partly prepared for 
<collectionCode id="9AAAAE27FFE78C26FAE0F8A092BB2744" box="[1362,1424,1821,1845]" pageId="26" pageNumber="177">SEM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="B4A16569FFE78C21FF75F8FD937A25F5" lastPageId="29" lastPageNumber="180" pageId="26" pageNumber="177" type="description">
<paragraph id="FC0436E2FFE78C26FF75F8FD940127B5" blockId="26.[151,1437,1676,1989]" pageId="26" pageNumber="177">
<emphasis id="CECFEAF0FFE78C26FF75F8FD96722728" bold="true" box="[199,345,1856,1881]" pageId="26" pageNumber="177">Description.</emphasis>
<emphasis id="CECFEAF0FFE78C26FED9F8FD968A2728" box="[363,417,1856,1881]" italics="true" pageId="26" pageNumber="177">Tube</emphasis>
: white opaque, ostensibly free, with shiny surface, straight, thick-walled, mostly quadrangular in cross-section, edges rounded and never denticulate. Juvenile tubes with sharper keels, becoming rounded in adults. Tubes slightly twisted, growth stops accompanied by sudden turns of tube around growth axis. Sides usually slightly convex, sometimes slightly concave.
</paragraph>
<caption id="A8C4666AFFE68C27FF25F94194E12799" httpUri="https://zenodo.org/record/235040/files/figure.png" pageId="27" pageNumber="178" targetBox="[237,1354,193,1765]" targetPageId="27">
<paragraph id="FC0436E2FFE68C27FF25F94194E12799" blockId="27.[151,1436,1788,2024]" pageId="27" pageNumber="178">
<emphasis id="CECFEAF0FFE68C27FF25F941960A2760" bold="true" box="[151,289,1788,1810]" pageId="27" pageNumber="178">FIGURE 11.</emphasis>
Tube ultrastructures of 
<taxonomicName id="3BBB4D61FFE68C27FDADF94195D52763" box="[543,766,1788,1810]" class="Polychaeta" family="Serpulidae" genus="Hyalopomatus" kingdom="Animalia" order="Sabellida" pageId="27" pageNumber="178" phylum="Annelida" rank="species" species="dieteri" status="sp. nov.">
<emphasis id="CECFEAF0FFE68C27FDADF94195D52763" box="[543,766,1788,1810]" italics="true" pageId="27" pageNumber="178">Hyalopomatus dieteri</emphasis>
</taxonomicName>
<emphasis id="CECFEAF0FFE68C27FCB5F94194712760" bold="true" box="[775,858,1788,1809]" pageId="27" pageNumber="178">
<taxonomicNameLabel id="D5FC578BFFE68C27FCB5F94194712760" box="[775,858,1788,1809]" pageId="27" pageNumber="178" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A–E: 
<emphasis id="CECFEAF0FFE68C27FC11F94194E42760" bold="true" box="[931,975,1788,1809]" pageId="27" pageNumber="178">PIN</emphasis>
5485/16, A—longitudinal wall section with parabolic texture in middle wall part, B—enlarged outer part consisting of larger slightly elongated and isometric crystals, Cenlarged middle wall part, parabolic texture produced by orientation of elongated crystals along growth lines, D—enlarged inner wall part showing slight transition to isometric smaller crystals, E—oblique longitudinal section through inner wall part showing orientation of crystals transversely to growth axis, F—tube cross-section, G—enlarged cross-section across tube edge, H—enlarged outer wall part, I—details of middle wall part consisting of isometric to slightly elongated crystals in tube edge area, J—details of inner wall part showing layer with ordered cigar-shaped crystals. Arrows indicate direction of tube growth (where 
<taxonomicName id="3BBB4D61FFE68C27FF51F86F96072799" box="[227,300,2002,2024]" class="Polychaeta" family="Serpulidae" genus="Ditrypa" higherTaxonomySource="CoL" kingdom="Animalia" order="Sabellida" pageId="27" pageNumber="178" phylum="Annelida" rank="species" species="known">known</taxonomicName>
). Abbreviations: g. l.—growth lamellae, o. s.—outer tube surface.
</paragraph>
</caption>
<caption id="A8C4666AFFE18C20FF25F8E8934D2794" httpUri="https://zenodo.org/record/235041/files/figure.png" pageId="28" pageNumber="179" targetBox="[234,1364,180,1846]" targetPageId="28">
<paragraph id="FC0436E2FFE18C20FF25F8E8934D2794" blockId="28.[151,1436,1877,2021]" pageId="28" pageNumber="179">
<emphasis id="CECFEAF0FFE18C20FF25F8E8960B271B" bold="true" box="[151,288,1877,1899]" pageId="28" pageNumber="179">FIGURE 12.</emphasis>
Photos and SEM images of 
<taxonomicName id="3BBB4D61FFE18C20FDF5F8E8940F271A" box="[583,804,1877,1899]" class="Polychaeta" family="Serpulidae" genus="Hyalopomatus" kingdom="Animalia" order="Sabellida" pageId="28" pageNumber="179" phylum="Annelida" rank="species" species="dieteri" status="sp. nov.">
<emphasis id="CECFEAF0FFE18C20FDF5F8E8940F271A" box="[583,804,1877,1899]" italics="true" pageId="28" pageNumber="179">Hyalopomatus dieteri</emphasis>
</taxonomicName>
<emphasis id="CECFEAF0FFE18C20FC99F8E89456271B" bold="true" box="[811,893,1877,1898]" pageId="28" pageNumber="179">
<taxonomicNameLabel id="D5FC578BFFE18C20FC99F8E89456271B" box="[811,893,1877,1898]" pageId="28" pageNumber="179" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A–B: 
<emphasis id="CECFEAF0FFE18C20FC71F8E89338271B" bold="true" box="[963,1043,1877,1898]" pageId="28" pageNumber="179">MNHN</emphasis>
POLY TYPE 1564, photos, A—distal view of radioles and operculum on thin smooth peduncle, B—lateral view of thorax and basal part of radiolar lobes. C–J: paratype 
<emphasis id="CECFEAF0FFE18C20FF44F82F960B27D6" bold="true" box="[246,288,1938,1959]" pageId="28" pageNumber="179">AM</emphasis>
W.46386, SEM images, C—bundle of collar chaetae with fin-and-blade chaetae, D—close-up view of fin-andblade collar chaetae, E—thoracic chaetae, F—anterior thoracic uncini, G—close-up view of anterior pegs of thoracic uncini, H—distal part of abdominal chaeta, I—abdominal uncini, J ‒ anterior pegs of abdominal uncini.
</paragraph>
</caption>
<paragraph id="FC0436E2FFE08C21FF75FF2A94FB21BE" blockId="29.[151,1437,151,2024]" pageId="29" pageNumber="180">
<emphasis id="CECFEAF0FFE08C21FF75FF2A968820C1" box="[199,419,151,176]" italics="true" pageId="29" pageNumber="180">Tube ultrastructure</emphasis>
: wall unilayered with irregularly oriented prismatic (IOP) structure, in inner wall part transforming into semi-ordered irregularly oriented prismatic (SOIOP). Tube wall not clearly subdivided into layers, in longitudinal section revealing only larger isometric to slightly elongated crystals with size up to 5 µm long (
<figureCitation id="64802A67FFE08C21FF68FEB99601216D" box="[218,298,260,285]" captionStart="FIGURE 11" captionStartId="27.[151,250,1788,1810]" captionTargetBox="[237,1354,193,1765]" captionTargetId="figure@27.[237,1354,193,1766]" captionTargetPageId="27" captionText="FIGURE 11. Tube ultrastructures of Hyalopomatus dieteri sp. nov. A – E: PIN 5485 / 16, A—longitudinal wall section with parabolic texture in middle wall part, B—enlarged outer part consisting of larger slightly elongated and isometric crystals, Cenlarged middle wall part, parabolic texture produced by orientation of elongated crystals along growth lines, D—enlarged inner wall part showing slight transition to isometric smaller crystals, E—oblique longitudinal section through inner wall part showing orientation of crystals transversely to growth axis, F—tube cross-section, G—enlarged cross-section across tube edge, H—enlarged outer wall part, I—details of middle wall part consisting of isometric to slightly elongated crystals in tube edge area, J—details of inner wall part showing layer with ordered cigar-shaped crystals. Arrows indicate direction of tube growth (where known). Abbreviations: g. l. — growth lamellae, o. s. — outer tube surface." httpUri="https://zenodo.org/record/235040/files/figure.png" pageId="29" pageNumber="180">Fig. 11</figureCitation>
A–D). Crystal size smallest near lumen (
<figureCitation id="64802A67FFE08C21FCB0FEB9947F216D" box="[770,852,260,285]" captionStart="FIGURE 11" captionStartId="27.[151,250,1788,1810]" captionTargetBox="[237,1354,193,1765]" captionTargetId="figure@27.[237,1354,193,1766]" captionTargetPageId="27" captionText="FIGURE 11. Tube ultrastructures of Hyalopomatus dieteri sp. nov. A – E: PIN 5485 / 16, A—longitudinal wall section with parabolic texture in middle wall part, B—enlarged outer part consisting of larger slightly elongated and isometric crystals, Cenlarged middle wall part, parabolic texture produced by orientation of elongated crystals along growth lines, D—enlarged inner wall part showing slight transition to isometric smaller crystals, E—oblique longitudinal section through inner wall part showing orientation of crystals transversely to growth axis, F—tube cross-section, G—enlarged cross-section across tube edge, H—enlarged outer wall part, I—details of middle wall part consisting of isometric to slightly elongated crystals in tube edge area, J—details of inner wall part showing layer with ordered cigar-shaped crystals. Arrows indicate direction of tube growth (where known). Abbreviations: g. l. — growth lamellae, o. s. — outer tube surface." httpUri="https://zenodo.org/record/235040/files/figure.png" pageId="29" pageNumber="180">Fig. 11</figureCitation>
D), increasing in middle (
<figureCitation id="64802A67FFE08C21FBCCFEB993E4216D" box="[1150,1231,260,285]" captionStart="FIGURE 11" captionStartId="27.[151,250,1788,1810]" captionTargetBox="[237,1354,193,1765]" captionTargetId="figure@27.[237,1354,193,1766]" captionTargetPageId="27" captionText="FIGURE 11. Tube ultrastructures of Hyalopomatus dieteri sp. nov. A – E: PIN 5485 / 16, A—longitudinal wall section with parabolic texture in middle wall part, B—enlarged outer part consisting of larger slightly elongated and isometric crystals, Cenlarged middle wall part, parabolic texture produced by orientation of elongated crystals along growth lines, D—enlarged inner wall part showing slight transition to isometric smaller crystals, E—oblique longitudinal section through inner wall part showing orientation of crystals transversely to growth axis, F—tube cross-section, G—enlarged cross-section across tube edge, H—enlarged outer wall part, I—details of middle wall part consisting of isometric to slightly elongated crystals in tube edge area, J—details of inner wall part showing layer with ordered cigar-shaped crystals. Arrows indicate direction of tube growth (where known). Abbreviations: g. l. — growth lamellae, o. s. — outer tube surface." httpUri="https://zenodo.org/record/235040/files/figure.png" pageId="29" pageNumber="180">Fig. 11</figureCitation>
C), and becoming largest in outer part of the wall (
<figureCitation id="64802A67FFE08C21FDA0FE9A954D2131" box="[530,614,295,320]" captionStart="FIGURE 11" captionStartId="27.[151,250,1788,1810]" captionTargetBox="[237,1354,193,1765]" captionTargetId="figure@27.[237,1354,193,1766]" captionTargetPageId="27" captionText="FIGURE 11. Tube ultrastructures of Hyalopomatus dieteri sp. nov. A – E: PIN 5485 / 16, A—longitudinal wall section with parabolic texture in middle wall part, B—enlarged outer part consisting of larger slightly elongated and isometric crystals, Cenlarged middle wall part, parabolic texture produced by orientation of elongated crystals along growth lines, D—enlarged inner wall part showing slight transition to isometric smaller crystals, E—oblique longitudinal section through inner wall part showing orientation of crystals transversely to growth axis, F—tube cross-section, G—enlarged cross-section across tube edge, H—enlarged outer wall part, I—details of middle wall part consisting of isometric to slightly elongated crystals in tube edge area, J—details of inner wall part showing layer with ordered cigar-shaped crystals. Arrows indicate direction of tube growth (where known). Abbreviations: g. l. — growth lamellae, o. s. — outer tube surface." httpUri="https://zenodo.org/record/235040/files/figure.png" pageId="29" pageNumber="180">Fig. 11</figureCitation>
B). Cross-section showing distinct zone in inner wall part consisting of highly ordered cigar-shaped crystals (5–6 µm in length, 1–1.5 µm in diameter, 
<figureCitation id="64802A67FFE08C21FB90FEF193592115" box="[1058,1138,332,357]" captionStart="FIGURE 11" captionStartId="27.[151,250,1788,1810]" captionTargetBox="[237,1354,193,1765]" captionTargetId="figure@27.[237,1354,193,1766]" captionTargetPageId="27" captionText="FIGURE 11. Tube ultrastructures of Hyalopomatus dieteri sp. nov. A – E: PIN 5485 / 16, A—longitudinal wall section with parabolic texture in middle wall part, B—enlarged outer part consisting of larger slightly elongated and isometric crystals, Cenlarged middle wall part, parabolic texture produced by orientation of elongated crystals along growth lines, D—enlarged inner wall part showing slight transition to isometric smaller crystals, E—oblique longitudinal section through inner wall part showing orientation of crystals transversely to growth axis, F—tube cross-section, G—enlarged cross-section across tube edge, H—enlarged outer wall part, I—details of middle wall part consisting of isometric to slightly elongated crystals in tube edge area, J—details of inner wall part showing layer with ordered cigar-shaped crystals. Arrows indicate direction of tube growth (where known). Abbreviations: g. l. — growth lamellae, o. s. — outer tube surface." httpUri="https://zenodo.org/record/235040/files/figure.png" pageId="29" pageNumber="180">Fig. 11</figureCitation>
J) with long axes oriented parallel to tube surface (
<figureCitation id="64802A67FFE08C21FE01FED2952E21F9" box="[435,517,367,392]" captionStart="FIGURE 11" captionStartId="27.[151,250,1788,1810]" captionTargetBox="[237,1354,193,1765]" captionTargetId="figure@27.[237,1354,193,1766]" captionTargetPageId="27" captionText="FIGURE 11. Tube ultrastructures of Hyalopomatus dieteri sp. nov. A – E: PIN 5485 / 16, A—longitudinal wall section with parabolic texture in middle wall part, B—enlarged outer part consisting of larger slightly elongated and isometric crystals, Cenlarged middle wall part, parabolic texture produced by orientation of elongated crystals along growth lines, D—enlarged inner wall part showing slight transition to isometric smaller crystals, E—oblique longitudinal section through inner wall part showing orientation of crystals transversely to growth axis, F—tube cross-section, G—enlarged cross-section across tube edge, H—enlarged outer wall part, I—details of middle wall part consisting of isometric to slightly elongated crystals in tube edge area, J—details of inner wall part showing layer with ordered cigar-shaped crystals. Arrows indicate direction of tube growth (where known). Abbreviations: g. l. — growth lamellae, o. s. — outer tube surface." httpUri="https://zenodo.org/record/235040/files/figure.png" pageId="29" pageNumber="180">Fig. 11</figureCitation>
E, J) resulting in SOIOP ultrastructure; in longitudinal sections this structure is unrecognizable. Transition from IOP to SOIOP structure gradual (
<figureCitation id="64802A67FFE08C21FC36FE2994FD21DD" box="[900,982,404,429]" captionStart="FIGURE 11" captionStartId="27.[151,250,1788,1810]" captionTargetBox="[237,1354,193,1765]" captionTargetId="figure@27.[237,1354,193,1766]" captionTargetPageId="27" captionText="FIGURE 11. Tube ultrastructures of Hyalopomatus dieteri sp. nov. A – E: PIN 5485 / 16, A—longitudinal wall section with parabolic texture in middle wall part, B—enlarged outer part consisting of larger slightly elongated and isometric crystals, Cenlarged middle wall part, parabolic texture produced by orientation of elongated crystals along growth lines, D—enlarged inner wall part showing slight transition to isometric smaller crystals, E—oblique longitudinal section through inner wall part showing orientation of crystals transversely to growth axis, F—tube cross-section, G—enlarged cross-section across tube edge, H—enlarged outer wall part, I—details of middle wall part consisting of isometric to slightly elongated crystals in tube edge area, J—details of inner wall part showing layer with ordered cigar-shaped crystals. Arrows indicate direction of tube growth (where known). Abbreviations: g. l. — growth lamellae, o. s. — outer tube surface." httpUri="https://zenodo.org/record/235040/files/figure.png" pageId="29" pageNumber="180">Fig. 11</figureCitation>
J). Parabolic growth lamellae (
<figureCitation id="64802A67FFE08C21FA83FE2992AB21DD" box="[1329,1408,404,429]" captionStart="FIGURE 11" captionStartId="27.[151,250,1788,1810]" captionTargetBox="[237,1354,193,1765]" captionTargetId="figure@27.[237,1354,193,1766]" captionTargetPageId="27" captionText="FIGURE 11. Tube ultrastructures of Hyalopomatus dieteri sp. nov. A – E: PIN 5485 / 16, A—longitudinal wall section with parabolic texture in middle wall part, B—enlarged outer part consisting of larger slightly elongated and isometric crystals, Cenlarged middle wall part, parabolic texture produced by orientation of elongated crystals along growth lines, D—enlarged inner wall part showing slight transition to isometric smaller crystals, E—oblique longitudinal section through inner wall part showing orientation of crystals transversely to growth axis, F—tube cross-section, G—enlarged cross-section across tube edge, H—enlarged outer wall part, I—details of middle wall part consisting of isometric to slightly elongated crystals in tube edge area, J—details of inner wall part showing layer with ordered cigar-shaped crystals. Arrows indicate direction of tube growth (where known). Abbreviations: g. l. — growth lamellae, o. s. — outer tube surface." httpUri="https://zenodo.org/record/235040/files/figure.png" pageId="29" pageNumber="180">Fig. 11</figureCitation>
A, C) present in all studied sections, axis of parabolae located in wall centre.
</paragraph>
<paragraph id="FC0436E2FFE08C21FF75FE6194DD2185" blockId="29.[151,1437,151,2024]" box="[199,1014,476,504]" pageId="29" pageNumber="180">
<emphasis id="CECFEAF0FFE08C21FF75FE6196A62184" box="[199,397,476,501]" italics="true" pageId="29" pageNumber="180">Tube mineralogy:</emphasis>
97–100% aragonite (I arag=69), doubtful calcite content.
</paragraph>
<paragraph id="FC0436E2FFE08C21FF75FDBF93442212" blockId="29.[151,1437,151,2024]" pageId="29" pageNumber="180">
<emphasis id="CECFEAF0FFE08C21FF75FDBF9654226A" box="[199,383,514,539]" italics="true" pageId="29" pageNumber="180">Radiolar crown</emphasis>
: with 7 pairs of radioles in 
<typeStatus id="23008840FFE08C21FD5DFDBF947A226A" box="[751,849,514,539]" pageId="29" pageNumber="180" type="holotype">holotype</typeStatus>
(radioles mostly missing in 
<typeStatus id="23008840FFE08C21FB08FDBE9234226A" box="[1210,1311,515,539]" pageId="29" pageNumber="180" type="paratype">paratype</typeStatus>
), arranged pectinately, easily detachable from short radiolar lobes. Inter-radiolar membrane and stylodes absent. Terminal filaments of radioles thin, spirally twisted. Radiolar eyes and mouth palps not observed.
</paragraph>
<paragraph id="FC0436E2FFE08C21FF75FDD3938B22DF" blockId="29.[151,1437,151,2024]" pageId="29" pageNumber="180">
<emphasis id="CECFEAF0FFE08C21FF75FDD3960422F6" box="[199,303,622,647]" italics="true" pageId="29" pageNumber="180">Peduncle</emphasis>
: smooth, cylindrical, thin (approximately same thickness as radioles), distal wings absent; inserted conspicuously outside radiolar crown proper, between base of 1st and 2nd radioles (
<figureCitation id="64802A67FFE08C21FB9FFD28935422DF" box="[1069,1151,661,686]" captionStart="FIGURE 12" captionStartId="28.[151,250,1877,1899]" captionTargetBox="[234,1364,180,1846]" captionTargetId="figure@28.[229,1368,177,1850]" captionTargetPageId="28" captionText="FIGURE 12. Photos and SEM images of Hyalopomatus dieteri sp. nov. A – B: MNHN POLY TYPE 1564, photos, A—distal view of radioles and operculum on thin smooth peduncle, B—lateral view of thorax and basal part of radiolar lobes. C – J: paratype AM W. 46386, SEM images, C—bundle of collar chaetae with fin-and-blade chaetae, D—close-up view of fin-andblade collar chaetae, E—thoracic chaetae, F—anterior thoracic uncini, G—close-up view of anterior pegs of thoracic uncini, H—distal part of abdominal chaeta, I—abdominal uncini, J ‒ anterior pegs of abdominal uncini." httpUri="https://zenodo.org/record/235041/files/figure.png" pageId="29" pageNumber="180">Fig. 12</figureCitation>
B).
</paragraph>
<paragraph id="FC0436E2FFE08C21FF75FD04921A2288" blockId="29.[151,1437,151,2024]" pageId="29" pageNumber="180">
<emphasis id="CECFEAF0FFE08C21FF75FD04950522A3" box="[199,558,697,722]" italics="true" pageId="29" pageNumber="180">Collar and thoracic membranes</emphasis>
: collar covering radiolar lobes, thin; trilobed, with ventral lobe slightly higher than the lateral ones (
<figureCitation id="64802A67FFE08C21FE35FD5D96F22288" box="[391,473,736,761]" captionStart="FIGURE 12" captionStartId="28.[151,250,1877,1899]" captionTargetBox="[234,1364,180,1846]" captionTargetId="figure@28.[229,1368,177,1850]" captionTargetPageId="28" captionText="FIGURE 12. Photos and SEM images of Hyalopomatus dieteri sp. nov. A – B: MNHN POLY TYPE 1564, photos, A—distal view of radioles and operculum on thin smooth peduncle, B—lateral view of thorax and basal part of radiolar lobes. C – J: paratype AM W. 46386, SEM images, C—bundle of collar chaetae with fin-and-blade chaetae, D—close-up view of fin-andblade collar chaetae, E—thoracic chaetae, F—anterior thoracic uncini, G—close-up view of anterior pegs of thoracic uncini, H—distal part of abdominal chaeta, I—abdominal uncini, J ‒ anterior pegs of abdominal uncini." httpUri="https://zenodo.org/record/235041/files/figure.png" pageId="29" pageNumber="180">Fig. 12</figureCitation>
B). Collar continuous with short thoracic membranes ending at 2nd chaetiger.
</paragraph>
<paragraph id="FC0436E2FFE08C21FF75FCB995B32314" blockId="29.[151,1437,151,2024]" pageId="29" pageNumber="180">
<emphasis id="CECFEAF0FFE08C21FF75FCB9966F236C" box="[199,324,772,797]" italics="true" pageId="29" pageNumber="180">Operculum</emphasis>
: soft membranous, semi-transparent, mostly globular, but with flattened top, slightly differentiated from the basal part; conspicuous constriction and additional small vesicular ampulla between operculum and peduncle (
<figureCitation id="64802A67FFE08C21FEB9FCF196772314" box="[267,348,844,869]" captionStart="FIGURE 12" captionStartId="28.[151,250,1877,1899]" captionTargetBox="[234,1364,180,1846]" captionTargetId="figure@28.[229,1368,177,1850]" captionTargetPageId="28" captionText="FIGURE 12. Photos and SEM images of Hyalopomatus dieteri sp. nov. A – B: MNHN POLY TYPE 1564, photos, A—distal view of radioles and operculum on thin smooth peduncle, B—lateral view of thorax and basal part of radiolar lobes. C – J: paratype AM W. 46386, SEM images, C—bundle of collar chaetae with fin-and-blade chaetae, D—close-up view of fin-andblade collar chaetae, E—thoracic chaetae, F—anterior thoracic uncini, G—close-up view of anterior pegs of thoracic uncini, H—distal part of abdominal chaeta, I—abdominal uncini, J ‒ anterior pegs of abdominal uncini." httpUri="https://zenodo.org/record/235041/files/figure.png" pageId="29" pageNumber="180">Fig. 12</figureCitation>
A). Pseudoperculum absent.
</paragraph>
<paragraph id="FC0436E2FFE08C21FF75FCCD93BA244C" blockId="29.[151,1437,151,2024]" pageId="29" pageNumber="180">
<emphasis id="CECFEAF0FFE08C21FF75FCCD963323F8" box="[199,280,880,905]" italics="true" pageId="29" pageNumber="180">Thorax</emphasis>
: with 6 chaetigerous segments, 5 of which uncinigerous. Small bundle of collar chaetae, of two 
<typeStatus id="23008840FFE08C21FAEBFCCC92B323F8" box="[1369,1432,881,905]" pageId="29" pageNumber="180">types</typeStatus>
: simple limbate and fin-and-blade with the distal blade separated from the basal fin by a short gap (
<figureCitation id="64802A67FFE08C21FAB1FC29927D23DC" box="[1283,1366,916,941]" captionStart="FIGURE 12" captionStartId="28.[151,250,1877,1899]" captionTargetBox="[234,1364,180,1846]" captionTargetId="figure@28.[229,1368,177,1850]" captionTargetPageId="28" captionText="FIGURE 12. Photos and SEM images of Hyalopomatus dieteri sp. nov. A – B: MNHN POLY TYPE 1564, photos, A—distal view of radioles and operculum on thin smooth peduncle, B—lateral view of thorax and basal part of radiolar lobes. C – J: paratype AM W. 46386, SEM images, C—bundle of collar chaetae with fin-and-blade chaetae, D—close-up view of fin-andblade collar chaetae, E—thoracic chaetae, F—anterior thoracic uncini, G—close-up view of anterior pegs of thoracic uncini, H—distal part of abdominal chaeta, I—abdominal uncini, J ‒ anterior pegs of abdominal uncini." httpUri="https://zenodo.org/record/235041/files/figure.png" pageId="29" pageNumber="180">Fig. 12</figureCitation>
C, D). Subsequent chaetae limbate, of two sizes, 
<taxonomicName id="3BBB4D61FFE08C21FDCCFC0495C523A1" box="[638,750,953,976]" class="Polychaeta" family="Serpulidae" genus="Apomatus" kingdom="Animalia" order="Sabellida" pageId="29" pageNumber="180" phylum="Annelida" rank="genus">
<emphasis id="CECFEAF0FFE08C21FDCCFC0495C523A1" box="[638,750,953,976]" italics="true" pageId="29" pageNumber="180">Apomatus</emphasis>
</taxonomicName>
-chaetae absent (
<figureCitation id="64802A67FFE08C21FC1DFC05932823A0" box="[943,1027,952,977]" captionStart="FIGURE 12" captionStartId="28.[151,250,1877,1899]" captionTargetBox="[234,1364,180,1846]" captionTargetId="figure@28.[229,1368,177,1850]" captionTargetPageId="28" captionText="FIGURE 12. Photos and SEM images of Hyalopomatus dieteri sp. nov. A – B: MNHN POLY TYPE 1564, photos, A—distal view of radioles and operculum on thin smooth peduncle, B—lateral view of thorax and basal part of radiolar lobes. C – J: paratype AM W. 46386, SEM images, C—bundle of collar chaetae with fin-and-blade chaetae, D—close-up view of fin-andblade collar chaetae, E—thoracic chaetae, F—anterior thoracic uncini, G—close-up view of anterior pegs of thoracic uncini, H—distal part of abdominal chaeta, I—abdominal uncini, J ‒ anterior pegs of abdominal uncini." httpUri="https://zenodo.org/record/235041/files/figure.png" pageId="29" pageNumber="180">Fig. 12</figureCitation>
E). Uncini along entire thorax raspshaped, with 20–25 small teeth in profile view, with 6–8 teeth in the row above flat anterior peg made of 3–4 rounded lobes, dental formula P:8:7:7:7:6:6:6:6:6:6:6:6:?:?:?:?:?:?:? (
<figureCitation id="64802A67FFE08C21FC17FBBD94DE2468" box="[933,1013,1024,1049]" captionStart="FIGURE 12" captionStartId="28.[151,250,1877,1899]" captionTargetBox="[234,1364,180,1846]" captionTargetId="figure@28.[229,1368,177,1850]" captionTargetPageId="28" captionText="FIGURE 12. Photos and SEM images of Hyalopomatus dieteri sp. nov. A – B: MNHN POLY TYPE 1564, photos, A—distal view of radioles and operculum on thin smooth peduncle, B—lateral view of thorax and basal part of radiolar lobes. C – J: paratype AM W. 46386, SEM images, C—bundle of collar chaetae with fin-and-blade chaetae, D—close-up view of fin-andblade collar chaetae, E—thoracic chaetae, F—anterior thoracic uncini, G—close-up view of anterior pegs of thoracic uncini, H—distal part of abdominal chaeta, I—abdominal uncini, J ‒ anterior pegs of abdominal uncini." httpUri="https://zenodo.org/record/235041/files/figure.png" pageId="29" pageNumber="180">Fig. 12</figureCitation>
F, G). Pair of prostomial eyes absent. Triangular depression absent, thoracic tori almost parallel to mid-ventral line of the thorax.
</paragraph>
<paragraph id="FC0436E2FFE08C21FF75FBF597FB2480" blockId="29.[151,1437,151,2024]" pageId="29" pageNumber="180">
<emphasis id="CECFEAF0FFE08C21FF75FBF5961A2410" box="[199,305,1096,1121]" italics="true" pageId="29" pageNumber="180">Abdomen</emphasis>
: with up to 60 segments. Chaetae long, nearly capillary with only narrow geniculate tip clearly made of two rows of pointed teeth (
<figureCitation id="64802A67FFE08C21FE5AFBD1951024F4" box="[488,571,1132,1157]" captionStart="FIGURE 12" captionStartId="28.[151,250,1877,1899]" captionTargetBox="[234,1364,180,1846]" captionTargetId="figure@28.[229,1368,177,1850]" captionTargetPageId="28" captionText="FIGURE 12. Photos and SEM images of Hyalopomatus dieteri sp. nov. A – B: MNHN POLY TYPE 1564, photos, A—distal view of radioles and operculum on thin smooth peduncle, B—lateral view of thorax and basal part of radiolar lobes. C – J: paratype AM W. 46386, SEM images, C—bundle of collar chaetae with fin-and-blade chaetae, D—close-up view of fin-andblade collar chaetae, E—thoracic chaetae, F—anterior thoracic uncini, G—close-up view of anterior pegs of thoracic uncini, H—distal part of abdominal chaeta, I—abdominal uncini, J ‒ anterior pegs of abdominal uncini." httpUri="https://zenodo.org/record/235041/files/figure.png" pageId="29" pageNumber="180">
Fig. 
<date id="88051022FFE08C21FDAFFBD0951024F4" box="[541,571,1133,1157]" pageId="29" pageNumber="180" value="2012-01">12</date>
</figureCitation>
I). Capillary chaetae present in posterior chaetigers. Uncini rasp-shaped with over 20 teeth in profile and up to 9 rows of teeth (
<figureCitation id="64802A67FFE08C21FD6EFB2D941B24D8" box="[732,816,1168,1193]" captionStart="FIGURE 12" captionStartId="28.[151,250,1877,1899]" captionTargetBox="[234,1364,180,1846]" captionTargetId="figure@28.[229,1368,177,1850]" captionTargetPageId="28" captionText="FIGURE 12. Photos and SEM images of Hyalopomatus dieteri sp. nov. A – B: MNHN POLY TYPE 1564, photos, A—distal view of radioles and operculum on thin smooth peduncle, B—lateral view of thorax and basal part of radiolar lobes. C – J: paratype AM W. 46386, SEM images, C—bundle of collar chaetae with fin-and-blade chaetae, D—close-up view of fin-andblade collar chaetae, E—thoracic chaetae, F—anterior thoracic uncini, G—close-up view of anterior pegs of thoracic uncini, H—distal part of abdominal chaeta, I—abdominal uncini, J ‒ anterior pegs of abdominal uncini." httpUri="https://zenodo.org/record/235041/files/figure.png" pageId="29" pageNumber="180">Fig. 12</figureCitation>
J, K) above anterior peg flat divided into 3–5 rounded lobes (crenulated). Dental formula P:9:7:5:4:3:4:3:2:?:? or P:8:5:5:5:4:4:4:?:?. Achaetous anterior abdominal zone long.
</paragraph>
<paragraph id="FC0436E2FFE08C21FF75FB40937A25F5" blockId="29.[151,1437,151,2024]" pageId="29" pageNumber="180">
<emphasis id="CECFEAF0FFE08C21FF75FB4097DF2565" box="[199,244,1277,1300]" italics="true" pageId="29" pageNumber="180">Size</emphasis>
: total body length up to 
<quantity id="3B439B07FFE08C21FDBBFB4195712565" box="[521,602,1276,1301]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="29" pageNumber="180" unit="mm" value="15.0">15 mm</quantity>
, including up to 
<quantity id="3B439B07FFE08C21FCAAFB4094722565" box="[792,857,1277,1301]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="29" pageNumber="180" unit="mm" value="9.0">9 mm</quantity>
long radioles, width of thorax up to 
<quantity id="3B439B07FFE08C21FB46FB4192652565" box="[1268,1358,1276,1301]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="29" pageNumber="180" unit="mm" value="0.5">0.5 mm</quantity>
. Tubes incomplete and broken into fragments, maximum total length of fragments (in 
<typeStatus id="23008840FFE08C21FB9EFA9C93A52548" box="[1068,1166,1313,1337]" pageId="29" pageNumber="180" type="paratype">paratype</typeStatus>
) 
<quantity id="3B439B07FFE08C21FB12FA9C93DE2549" box="[1184,1269,1313,1337]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="29" pageNumber="180" unit="mm" value="40.0">40 mm</quantity>
. External tube diameter up to 
<quantity id="3B439B07FFE08C21FEFBFAF8968F252D" box="[329,420,1349,1373]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="29" pageNumber="180" unit="mm" value="1.3">1.3 mm</quantity>
, corresponding lumen diameter 
<quantity id="3B439B07FFE08C21FCABFAF8945E252D" box="[793,885,1349,1373]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="29" pageNumber="180" unit="mm" value="0.9">0.9 mm</quantity>
. Thickness of tube wall in between the angular margins about 1/3–1/4th of outer diameter, and up to 1/2 when measured across keels.
</paragraph>
</subSubSection>
<subSubSection id="B4A16569FFE08C21FF75FA32968725BD" pageId="29" pageNumber="180" type="etymology">
<paragraph id="FC0436E2FFE08C21FF75FA32968725BD" blockId="29.[151,1437,151,2024]" pageId="29" pageNumber="180">
<emphasis id="CECFEAF0FFE08C21FF75FA32966D25D9" bold="true" box="[199,326,1423,1448]" pageId="29" pageNumber="180">Etymology</emphasis>
. The species is named after Dr. Dieter Fiege (
<emphasis id="CECFEAF0FFE08C21FCF4FA3294AA25D9" bold="true" box="[838,897,1423,1448]" pageId="29" pageNumber="180">SMF</emphasis>
) in recognition of his important contributions to taxonomy of 
<taxonomicName id="3BBB4D61FFE08C21FE9EFA0E968C25BD" box="[300,423,1459,1484]" class="Polychaeta" family="Serpulidae" kingdom="Animalia" order="Sabellida" pageId="29" pageNumber="180" phylum="Annelida" rank="family">Serpulidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="B4A16569FFE08C21FF75FA6A944B259E" box="[199,864,1495,1520]" pageId="29" pageNumber="180" type="distribution">
<paragraph id="FC0436E2FFE08C21FF75FA6A944B259E" blockId="29.[151,1437,151,2024]" box="[199,864,1495,1520]" pageId="29" pageNumber="180">
<emphasis id="CECFEAF0FFE08C21FF75FA6A964B2581" bold="true" box="[199,352,1495,1520]" pageId="29" pageNumber="180">Distribution.</emphasis>
Coral Sea off 
<collectingCountry id="84AC7672FFE08C21FDB1FA6595982581" box="[515,691,1495,1520]" name="New Caledonia" pageId="29" pageNumber="180">New Caledonia</collectingCountry>
, 
<quantity id="3B439B07FFE08C21FD0CFA6A9477259E" box="[702,860,1495,1519]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="1.98" metricValueMin="1.8199999999999998" pageId="29" pageNumber="180" unit="m" value="1900.0" valueMax="1980.0" valueMin="1820.0">1820–1980 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="B4A16569FFE08C21FF75FA4696CF2799" pageId="29" pageNumber="180" type="discussion">
<paragraph id="FC0436E2FFE08C21FF75FA469697262D" blockId="29.[151,1437,151,2024]" pageId="29" pageNumber="180">
<emphasis id="CECFEAF0FFE08C21FF75FA4696102665" bold="true" box="[199,315,1531,1556]" pageId="29" pageNumber="180">Remarks.</emphasis>
The new species from 
<collectingCountry id="84AC7672FFE08C21FD8FFA4195C12665" box="[573,746,1531,1556]" name="New Caledonia" pageId="29" pageNumber="180">New Caledonia</collectingCountry>
resembles 
<taxonomicName id="3BBB4D61FFE08C21FCD5FA41939E2662" authority="Gravier, 1911" authorityName="Gravier" authorityYear="1911" box="[871,1205,1531,1556]" class="Polychaeta" family="Serpulidae" genus="Hyalopomatus" kingdom="Animalia" order="Sabellida" pageId="29" pageNumber="180" phylum="Annelida" rank="species" species="macintoshi">
<emphasis id="CECFEAF0FFE08C21FCD5FA4193282665" box="[871,1027,1531,1556]" italics="true" pageId="29" pageNumber="180">H. macintoshi</emphasis>
(
<bibRefCitation id="982A4B13FFE08C21FBA0FA4693862662" author="Gravier" box="[1042,1197,1531,1556]" pageId="29" pageNumber="180" refString="Gravier, C. (1911) Especes nouvelles d'annelides polychetes. Expedition antarctiques francaise du &quot; Pourquoi-Pas? &quot; dirigee par le Dr. J. B. Charcot (1908 - 1910). Bulletin du Museum National d'Histoire Naturelle, Paris, 17, 310 - 316." type="journal article" year="1911">Gravier, 1911</bibRefCitation>
)
</taxonomicName>
in having a globular transparent, only slightly differentiated operculum. However, the tube of 
<taxonomicName id="3BBB4D61FFE08C21FC54F99D937F2649" box="[998,1108,1567,1592]" class="Polychaeta" family="Serpulidae" genus="Hyalopomatus" kingdom="Animalia" order="Sabellida" pageId="29" pageNumber="180" phylum="Annelida" rank="species" species="dieteri" status="sp. nov.">
<emphasis id="CECFEAF0FFE08C21FC54F99D937F2649" box="[998,1108,1567,1592]" italics="true" pageId="29" pageNumber="180">H. dieteri</emphasis>
</taxonomicName>
<emphasis id="CECFEAF0FFE08C21FBECF9A293902649" bold="true" box="[1118,1211,1567,1592]" pageId="29" pageNumber="180">
<taxonomicNameLabel id="D5FC578BFFE08C21FBECF9A293902649" box="[1118,1211,1567,1592]" pageId="29" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinct in being polygonal (quadrangular).
</paragraph>
<paragraph id="FC0436E2FFE08C21FF75F9D596CF2799" blockId="29.[151,1437,151,2024]" pageId="29" pageNumber="180">
Four other 
<taxonomicName id="3BBB4D61FFE08C21FEF6F9DA96CC26F1" box="[324,487,1639,1664]" class="Polychaeta" family="Serpulidae" genus="Hyalopomatus" kingdom="Animalia" order="Sabellida" pageId="29" pageNumber="180" phylum="Annelida" rank="genus">
<emphasis id="CECFEAF0FFE08C21FEF6F9DA96CC26F1" box="[324,487,1639,1664]" italics="true" pageId="29" pageNumber="180">Hyalopomatus</emphasis>
</taxonomicName>
spp. with 
<taxonomicName id="3BBB4D61FFE08C21FDEEF9DA958326F1" box="[604,680,1639,1664]" class="Polychaeta" family="Serpulidae" genus="Ditrypa" higherTaxonomySource="CoL" kingdom="Animalia" order="Sabellida" pageId="29" pageNumber="180" phylum="Annelida" rank="species" species="known">known</taxonomicName>
tube ultrastructures are 
<taxonomicName id="3BBB4D61FFE08C21FC05F9D592B726F1" authority="Ben-Eliahu &amp; Fiege, 1996" authorityName="Ben-Eliahu &amp; Fiege" authorityYear="1996" box="[951,1436,1639,1664]" class="Polychaeta" family="Serpulidae" genus="Hyalopomatus" kingdom="Animalia" order="Sabellida" pageId="29" pageNumber="180" phylum="Annelida" rank="species" species="variorugosus">
<emphasis id="CECFEAF0FFE08C21FC05F9D59346260E" box="[951,1133,1640,1663]" italics="true" pageId="29" pageNumber="180">H. variorugosus</emphasis>
<bibRefCitation id="982A4B13FFE08C21FBC6F9DA92B726F1" author="Ben-Eliahu" box="[1140,1436,1639,1664]" pageId="29" pageNumber="180" refString="Ben-Eliahu, M. N. &amp; Fiege, D. (1996) Serpulid tube-worms (Annelida, Polychaeta) of the central and eastern Mediterranean with particular attention to the Levant Basin. Senckenbergiana Maritima, 28, 1 - 51. http: // dx. doi. org / 10.1007 / BF 03042821" type="journal article" year="1996">Ben-Eliahu &amp; Fiege, 1996</bibRefCitation>
</taxonomicName>
(see 
<bibRefCitation id="982A4B13FFE08C21FF7EF93696BA26D2" author="Sanfilippo" box="[204,401,1675,1700]" pageId="29" pageNumber="180" refString="Sanfilippo, R. (1998 a) Tube morphology and structure of the bathyal Mediterranean serpulid Hyalopomatus variorugosus Ben- Eliahu &amp; Fiege, 1996 (Annelida, Polychaeta). Rivista Italiana di Paleontologia e Stratigrafia, 104, 131 - 138." type="journal article" year="1998" yearSuffix="a">Sanfilippo 1998a</bibRefCitation>
; Vinn 
<emphasis id="CECFEAF0FFE08C21FE6EF931953D26D5" box="[476,534,1675,1700]" italics="true" pageId="29" pageNumber="180">et al.</emphasis>
2008), 
<taxonomicName id="3BBB4D61FFE08C21FDDCF93194F626D2" authority="Marenzeller, 1878" authorityName="Marenzeller" authorityYear="1878" box="[622,989,1675,1700]" class="Polychaeta" family="Serpulidae" genus="Hyalopomatus" kingdom="Animalia" order="Sabellida" pageId="29" pageNumber="180" phylum="Annelida" rank="species" species="claparedii">
<emphasis id="CECFEAF0FFE08C21FDDCF931942F26D5" box="[622,772,1675,1700]" italics="true" pageId="29" pageNumber="180">H. claparedii</emphasis>
<bibRefCitation id="982A4B13FFE08C21FCBEF93694F626D2" author="Marenzeller" box="[780,989,1675,1700]" pageId="29" pageNumber="180" refString="Marenzeller, E., von. (1878) Die Coelenteraten, Echinodermen und Wurmer der K. K. Osterreichisch - Ungarischen Nordpol - Expedition. Denkschriften Kaiserlichen Akademie der Wissenschaften, Wien (mathematisch-naturwissenschaftliche Classe), 35, 357 - 398." type="journal article" year="1878">Marenzeller, 1878</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="3BBB4D61FFE08C21FC5BF931924026D2" authority="Langerhans, 1884" authorityName="Langerhans" authorityYear="1884" box="[1001,1387,1675,1700]" class="Polychaeta" family="Serpulidae" genus="Hyalopomatus" kingdom="Animalia" order="Sabellida" pageId="29" pageNumber="180" phylum="Annelida" rank="species" species="marenzelleri">
<emphasis id="CECFEAF0FFE08C21FC5BF93193B226D5" box="[1001,1177,1675,1700]" italics="true" pageId="29" pageNumber="180">H. marenzelleri</emphasis>
<bibRefCitation id="982A4B13FFE08C21FB13F936924026D2" author="Langerhans" box="[1185,1387,1675,1700]" pageId="29" pageNumber="180" refString="Langerhans, P. (1884) Die Wurmfauna von Madeira. 4. Zeitschrift fur Wissenschaftliche Zoologie, 40, 247 - 285." type="journal article" year="1884">Langerhans, 1884</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="3BBB4D61FFE08C21FF25F90D953426B6" authority="Sanfilippo, 2009" authorityName="Sanfilippo" authorityYear="2009" box="[151,543,1711,1736]" class="Polychaeta" family="Serpulidae" genus="Hyalopomatus" kingdom="Animalia" order="Sabellida" pageId="29" pageNumber="180" phylum="Annelida" rank="species" species="madreporae">
<emphasis id="CECFEAF0FFE08C21FF25F90D966026B9" box="[151,331,1711,1736]" italics="true" pageId="29" pageNumber="180">H. madreporae</emphasis>
<bibRefCitation id="982A4B13FFE08C21FEE9F912953426B6" author="Sanfilippo" box="[347,543,1711,1736]" pageId="29" pageNumber="180" refString="Sanfilippo, R. (2009) New species of Hyalopomatus Marenzeller, 1878 (Polychaeta, Serpulidae) from Recent Mediterranean deep-water coral mounds and comments on some congeners. Zoosystema, 31, 147 - 161. http: // dx. doi. org / 10.5252 / z 2009 n 1 a 8" type="journal article" year="2009">Sanfilippo, 2009</bibRefCitation>
</taxonomicName>
(see 
<bibRefCitation id="982A4B13FFE08C21FDDCF912941B26B6" author="Sanfilippo" box="[622,816,1711,1736]" pageId="29" pageNumber="180" refString="Sanfilippo, R. (2009) New species of Hyalopomatus Marenzeller, 1878 (Polychaeta, Serpulidae) from Recent Mediterranean deep-water coral mounds and comments on some congeners. Zoosystema, 31, 147 - 161. http: // dx. doi. org / 10.5252 / z 2009 n 1 a 8" type="journal article" year="2009">Sanfilippo 2009</bibRefCitation>
). Ultrastructural 
<typeStatus id="23008840FFE08C21FBB2F90D931B26B9" box="[1024,1072,1712,1736]" pageId="29" pageNumber="180">type</typeStatus>
and shape of crystals of 
<taxonomicName id="3BBB4D61FFE08C21FA33F90D9600269A" class="Polychaeta" family="Serpulidae" genus="Hyalopomatus" kingdom="Animalia" order="Sabellida" pageId="29" pageNumber="180" phylum="Annelida" rank="species" species="variorugosus">
<emphasis id="CECFEAF0FFE08C21FA33F90D9600269A" italics="true" pageId="29" pageNumber="180">H. variorugosus</emphasis>
</taxonomicName>
are very similar to those of 
<taxonomicName id="3BBB4D61FFE08C21FDD4F96995F9269D" box="[614,722,1747,1772]" class="Polychaeta" family="Serpulidae" genus="Hyalopomatus" kingdom="Animalia" order="Sabellida" pageId="29" pageNumber="180" phylum="Annelida" rank="species" species="dieteri" status="sp. nov.">
<emphasis id="CECFEAF0FFE08C21FDD4F96995F9269D" box="[614,722,1747,1772]" italics="true" pageId="29" pageNumber="180">H. dieteri</emphasis>
</taxonomicName>
<emphasis id="CECFEAF0FFE08C21FD6BF96E9418269D" bold="true" box="[729,819,1747,1772]" pageId="29" pageNumber="180">
<taxonomicNameLabel id="D5FC578BFFE08C21FD6BF96E9418269D" box="[729,819,1747,1772]" pageId="29" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, despite chaotic orientation of crystals without defined SOIOP ultrastructure in the former. 
<taxonomicName id="3BBB4D61FFE08C21FDEAF945943B2761" box="[600,784,1783,1808]" class="Polychaeta" family="Serpulidae" genus="Hyalopomatus" kingdom="Animalia" order="Sabellida" pageId="29" pageNumber="180" phylum="Annelida" rank="species" species="marenzelleri">
<emphasis id="CECFEAF0FFE08C21FDEAF945943B2761" box="[600,784,1783,1808]" italics="true" pageId="29" pageNumber="180">H. marenzelleri</emphasis>
</taxonomicName>
demonstrates squat prismatic crystal shapes, but the ultrastructure 
<typeStatus id="23008840FFE08C21FE86F8A1964F2745" box="[308,356,1820,1844]" pageId="29" pageNumber="180">type</typeStatus>
is still the same (IOP). The closest to 
<taxonomicName id="3BBB4D61FFE08C21FCA0F8A194552745" box="[786,894,1819,1844]" class="Polychaeta" family="Serpulidae" genus="Hyalopomatus" kingdom="Animalia" order="Sabellida" pageId="29" pageNumber="180" phylum="Annelida" rank="species" species="dieteri" status="sp. nov.">
<emphasis id="CECFEAF0FFE08C21FCA0F8A194552745" box="[786,894,1819,1844]" italics="true" pageId="29" pageNumber="180">H. dieteri</emphasis>
</taxonomicName>
<emphasis id="CECFEAF0FFE08C21FC37F8A694F42745" bold="true" box="[901,991,1819,1844]" pageId="29" pageNumber="180">
<taxonomicNameLabel id="D5FC578BFFE08C21FC37F8A694F42745" box="[901,991,1819,1844]" pageId="29" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is 
<taxonomicName id="3BBB4D61FFE08C21FBB3F8A193BE2745" box="[1025,1173,1819,1844]" class="Polychaeta" family="Serpulidae" genus="Hyalopomatus" kingdom="Animalia" order="Sabellida" pageId="29" pageNumber="180" phylum="Annelida" rank="species" species="claparedii">
<emphasis id="CECFEAF0FFE08C21FBB3F8A193BE2745" box="[1025,1173,1819,1844]" italics="true" pageId="29" pageNumber="180">H. claparedii</emphasis>
</taxonomicName>
that has a well-defined SOIOP ultrastructure near the lumen (
<bibRefCitation id="982A4B13FFE08C21FD8CF88294662729" author="Sanfilippo" box="[574,845,1855,1880]" pageId="29" pageNumber="180" refString="Sanfilippo, R. (2009) New species of Hyalopomatus Marenzeller, 1878 (Polychaeta, Serpulidae) from Recent Mediterranean deep-water coral mounds and comments on some congeners. Zoosystema, 31, 147 - 161. http: // dx. doi. org / 10.5252 / z 2009 n 1 a 8" type="journal article" year="2009">Sanfilippo 2009, fig. 6E</bibRefCitation>
) with a gradual transition to IOP ultrastructure in the outer part of the tube wall. The tube of 
<taxonomicName id="3BBB4D61FFE08C21FDFEF8D995DE270D" box="[588,757,1891,1916]" class="Polychaeta" family="Serpulidae" genus="Hyalopomatus" kingdom="Animalia" order="Sabellida" pageId="29" pageNumber="180" phylum="Annelida" rank="species" species="madreporae">
<emphasis id="CECFEAF0FFE08C21FDFEF8D995DE270D" box="[588,757,1891,1916]" italics="true" pageId="29" pageNumber="180">H. madreporae</emphasis>
</taxonomicName>
also shows a similar pattern, but its SOIOP zone (
<bibRefCitation id="982A4B13FFE08C21FA97F8DE961027EE" author="Sanfilippo" pageId="29" pageNumber="180" refString="Sanfilippo, R. (2009) New species of Hyalopomatus Marenzeller, 1878 (Polychaeta, Serpulidae) from Recent Mediterranean deep-water coral mounds and comments on some congeners. Zoosystema, 31, 147 - 161. http: // dx. doi. org / 10.5252 / z 2009 n 1 a 8" type="journal article" year="2009">Sanfilippo 2009, fig. 4G</bibRefCitation>
) seems to be relatively wider than those in both 
<taxonomicName id="3BBB4D61FFE08C21FC10F835933F27D1" box="[930,1044,1927,1952]" class="Polychaeta" family="Serpulidae" genus="Hyalopomatus" kingdom="Animalia" order="Sabellida" pageId="29" pageNumber="180" phylum="Annelida" rank="species" species="dieteri" status="sp. nov.">
<emphasis id="CECFEAF0FFE08C21FC10F835933F27D1" box="[930,1044,1927,1952]" italics="true" pageId="29" pageNumber="180">H. dieteri</emphasis>
</taxonomicName>
<emphasis id="CECFEAF0FFE08C21FB90F83A93A827D1" bold="true" box="[1058,1155,1927,1952]" pageId="29" pageNumber="180">
<taxonomicNameLabel id="D5FC578BFFE08C21FB90F83A93A827D1" box="[1058,1155,1927,1952]" pageId="29" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="3BBB4D61FFE08C21FB7AF835924827D1" box="[1224,1379,1927,1952]" class="Polychaeta" family="Serpulidae" genus="Hyalopomatus" kingdom="Animalia" order="Sabellida" pageId="29" pageNumber="180" phylum="Annelida" rank="species" species="claparedii">
<emphasis id="CECFEAF0FFE08C21FB7AF835924827D1" box="[1224,1379,1927,1952]" italics="true" pageId="29" pageNumber="180">H. claparedii</emphasis>
</taxonomicName>
. All 
<taxonomicName id="3BBB4D61FFE08C21FF25F816961127B5" box="[151,314,1963,1988]" class="Polychaeta" family="Serpulidae" genus="Hyalopomatus" kingdom="Animalia" order="Sabellida" pageId="29" pageNumber="180" phylum="Annelida" rank="genus">
<emphasis id="CECFEAF0FFE08C21FF25F816961127B5" box="[151,314,1963,1988]" italics="true" pageId="29" pageNumber="180">Hyalopomatus</emphasis>
</taxonomicName>
spp. studied to date have very similar tube ultrastructures supporting the generic placement of the new species described herein.
</paragraph>
</subSubSection>
</treatment>
</document>