treatments-xml/data/72/42/47/7242476B8847F050FE2A7373F5339412.xml

<document ID-DOI="10.24349/z72f-jdlc" ID-ISSN="2107-7207" ID-Zenodo-Dep="7159892" approvalRequired="14" approvalRequired_for_textStreams="14" checkinTime="1665194714052" checkinUser="felipe" docAuthor="Ermilov, Sergey G. &amp; Martens, Jochen" docDate="2021" docId="7242476B8847F050FE2A7373F5339412" docLanguage="en" docName="Acarologia.61.4.995-1014.pdf.pdf" docOrigin="Acarologia 61 (4)" docSource="https://www1.montpellier.inrae.fr/CBGP/acarologia/article.php?id=4480" docStyle="DocumentStyle:0387F17C4AF9F8D6336E46CC5E5664D0.4:Acarologia.2018-.journal_article" docStyleId="0387F17C4AF9F8D6336E46CC5E5664D0" docStyleName="Acarologia.2018-.journal_article" docStyleVersion="4" docTitle="Leobodes schawalleri Ermilov &amp; Martens 2021, n. sp." docType="treatment" docVersion="3" lastPageNumber="1012" masterDocId="8E7B3F138849F042FFC17710F17B911B" masterDocTitle="Contribution to the knowledge of the oribatid mite genus Leobodes (Acari, Oribatida, Nippobodidae)" masterLastPageNumber="1014" masterPageNumber="995" pageNumber="1008" updateTime="1665196467129" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo>
<mods:title>Contribution to the knowledge of the oribatid mite genus Leobodes (Acari, Oribatida, Nippobodidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ermilov, Sergey G.</mods:namePart>
<mods:affiliation>Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Tyumen, Russia.</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Martens, Jochen</mods:namePart>
<mods:affiliation>Johannes Gutenberg-Universität, Institut für Organismische und Molekulare Evolutionsbiologie (iomE), Mainz, Germany ; Senckenberg Research Institute, Arachnology, Frankfurt am Main, Germany. &amp; Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Tyumen, Russia.</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Acarologia</mods:title>
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<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-12-10</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>61</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
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<mods:start>995</mods:start>
<mods:end>1014</mods:end>
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<mods:url>https://www1.montpellier.inrae.fr/CBGP/acarologia/article.php?id=4480</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.24349/z72f-jdlc</mods:identifier>
<mods:identifier type="ISSN">2107-7207</mods:identifier>
<mods:identifier type="Zenodo-Dep">7159892</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7160161" ID-Zenodo-Dep="7160161" LSID="urn:lsid:plazi:treatment:7242476B8847F050FE2A7373F5339412" httpUri="http://treatment.plazi.org/id/7242476B8847F050FE2A7373F5339412" lastPageId="18" lastPageNumber="1012" pageId="14" pageNumber="1008">
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<paragraph blockId="14.[491,841,1123,1149]" box="[491,841,1123,1149]" pageId="14" pageNumber="1008">
<heading bold="true" box="[491,841,1123,1149]" fontSize="11" level="2" pageId="14" pageNumber="1008" reason="3">
<emphasis bold="true" box="[491,841,1123,1149]" pageId="14" pageNumber="1008">
<taxonomicName authority="Ermilov &amp; Martens, 2021" authorityName="Ermilov &amp; Martens" authorityYear="2021" box="[491,766,1123,1149]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="1008" phylum="Arthropoda" rank="species" species="schawalleri" status="n. sp.">
<emphasis bold="true" box="[491,766,1123,1149]" italics="true" pageId="14" pageNumber="1008">Leobodes schawalleri</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[773,841,1123,1149]" pageId="14" pageNumber="1008" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="14.[491,1539,1179,2028]" box="[491,778,1179,1198]" pageId="14" pageNumber="1008">Zoobank: xxxxxxxxxxxxxxxxxxxxxx</paragraph>
</subSubSection>
<subSubSection box="[531,1013,1210,1234]" pageId="14" pageNumber="1008" type="description">
<paragraph blockId="14.[491,1539,1179,2028]" box="[531,1013,1210,1234]" pageId="14" pageNumber="1008">
(
<figureCitation box="[539,702,1210,1234]" captionStart="Figure 3" captionStartId="6.[119,186,1985,2006]" captionTargetBox="[220,1366,247,1866]" captionTargetId="figure-2@6.[193,1402,221,1931]" captionTargetPageId="6" captionText="Figure 3 Legs ofLeobodes spp., dissected adults: a – leg I ofL. becki n. sp., right, antiaxial view; b – leg II ofL. becki n. sp., left, antiaxial view; c – leg III ofL. becki n. sp., right, paraxial view; d – leg IV ofL. becki n. sp., left, antiaxial view; e – leg trochanter III ofL. schalleri n. sp., left, antiaxial view; f – leg trochanter IV ofL. schallerin. sp., left, antiaxial view; g – leg trochanter III ofL. schawallerin. sp., left, antiaxial view; h – leg trochanter IV ofL. schawallerin. sp., left, antiaxial view. Scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.7159900" httpUri="https://zenodo.org/record/7159900/files/figure.png" pageId="14" pageNumber="1008">
Figures 
<quantity box="[631,661,1210,1234]" metricMagnitude="-3" metricUnit="kg" metricValue="3.0" pageId="14" pageNumber="1008" unit="g" value="3.0">3g</quantity>
, 3h
</figureCitation>
, 
<figureCitation box="[714,780,1210,1234]" captionStart="Figure 6" captionStartId="9.[119,186,1704,1725]" captionTargetBox="[103,1494,221,1647]" captionTargetId="figure-2@9.[103,1494,221,1650]" captionTargetPageId="9" captionText="Figure 6 Leobodes spp., adult, microscope images: a –L. becki n. sp., dorsal view; b –L. becki n. sp., lateral view; c – L. schalleri n. sp., dorsal view; d –L. schallerin. sp., lateral view; e – L. schawallerin. sp., dorsal view; f –L. schawallerin. sp., lateral view. Magnification:" figureDoi="http://doi.org/10.5281/zenodo.7159912" httpUri="https://zenodo.org/record/7159912/files/figure.png" pageId="14" pageNumber="1008">6e, 6f</figureCitation>
, 
<figureCitation box="[791,817,1210,1234]" captionStart="Figure 10" captionStartId="15.[119,186,2041,2062]" captionTargetBox="[135,1476,227,1966]" captionTargetId="figure-2@15.[118,1479,221,1987]" captionTargetPageId="15" captionText="Figure 10 Leobodes schawallerin. sp., adult: a – dorsal view (legs omitted); b – ventral view (gnathosoma and legs omitted); c – lateral view (gnathosoma and legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7159926" httpUri="https://zenodo.org/record/7159926/files/figure.png" pageId="14" pageNumber="1008">10</figureCitation>
a-c, 11a-d, 12a-d)
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="1008" type="diagnosis">
<paragraph blockId="14.[491,1539,1179,2028]" pageId="14" pageNumber="1008">
<emphasis bold="true" box="[531,640,1245,1269]" pageId="14" pageNumber="1008">Diagnosis</emphasis>
— Adult. Body size: 597–664× 365–431. Body surface with dense granulate microsculpturing; notogaster and partially anogenital region foveolate. Prodorsal processes connected medially at a relatively large distance; their anterior parts forming rounded prodorsal aperture. Rostral, lamellar and interlamellar setae slightly thickened; 
<emphasis box="[1262,1283,1349,1372]" italics="true" pageId="14" pageNumber="1008">in</emphasis>
shortest, erect. Bothridial seta with distinctly lanceolate head. Notogastral tubercle present. Distance between medial notogastral condyles comparatively short. Thirteen pairs of notogastral setae present: seta 
<emphasis box="[540,552,1453,1476]" italics="true" pageId="14" pageNumber="1008">c</emphasis>
long, with slightly phylliform basal part and attenuate mediodistal part, straight; other setae of medium length, thickened. Subcapitular seta 
<emphasis box="[1059,1072,1486,1510]" italics="true" pageId="14" pageNumber="1008">h</emphasis>
setiform. Seta 
<emphasis box="[1234,1247,1486,1510]" italics="true" pageId="14" pageNumber="1008">l′</emphasis>
on trochanter III and 
<emphasis box="[1479,1497,1486,1510]" italics="true" pageId="14" pageNumber="1008">v′</emphasis>
on leg trochanters III, IV long, thick.
</paragraph>
</subSubSection>
<subSubSection lastPageId="18" lastPageNumber="1012" pageId="14" pageNumber="1008" type="description">
<paragraph blockId="14.[491,1539,1179,2028]" box="[531,1468,1555,1580]" pageId="14" pageNumber="1008">
<emphasis bold="true" box="[531,761,1556,1580]" pageId="14" pageNumber="1008">Description of adult</emphasis>
— Measurements – Body length: 664 (
<typeStatus box="[1216,1316,1555,1579]" pageId="14" pageNumber="1008">holotype</typeStatus>
: female), 597–
</paragraph>
<paragraph blockId="14.[491,1539,1179,2028]" pageId="14" pageNumber="1008">
664 (
<specimenCount box="[552,690,1590,1614]" pageId="14" pageNumber="1008" type="generic" typeStatus="paratypes">16 paratypes</specimenCount>
: 
<specimenCount box="[705,819,1590,1614]" pageId="14" pageNumber="1008" type="male">four males</specimenCount>
and 
<specimenCount box="[879,998,1590,1614]" pageId="14" pageNumber="1008" type="female">12 females</specimenCount>
); body width: 431 (
<typeStatus box="[1225,1325,1590,1614]" pageId="14" pageNumber="1008">holotype</typeStatus>
), 365–431 (
<specimenCount pageId="14" pageNumber="1008" type="generic" typeStatus="paratypes">16 paratypes</specimenCount>
).
</paragraph>
<paragraph blockId="14.[491,1539,1179,2028]" pageId="14" pageNumber="1008">Integument – Body color brown. Body surface with dense granulate microsculpturing. Tutorium and pedotectum I foveate. Region between bothridium and acetabula I–III densely tuberculate (diameter of tubercle up to 4). Notogaster with sparse foveolae (diameter of foveola up to 10). Anogenital region (except ventrolateral parts and region between genital and anal apertures) with more dense foveolae (diameter of foveola up to 10).</paragraph>
<paragraph blockId="14.[491,1539,1179,2028]" lastBlockId="18.[491,1530,229,1290]" lastPageId="18" lastPageNumber="1012" pageId="14" pageNumber="1008">
Prodorsum – Rostrum rounded. Lateral carinae fused medially before rostral margin. Lamella slightly developed. Prolamella present, but inconspicuous. Tutorium and prodorsal hollow of prodorsum well developed. Prodorsal processes connected medially at a relatively large distance. Anterior part of prodorsal processes have rounded prodorsal aperture (fused with prodorsal hollow) anteriorly to connecting region. Rostral (192–205), lamellar (180–192) and interlamellar (151–155) setae slightly thickened, slightly barbed; 
<emphasis box="[1231,1254,2005,2028]" italics="true" pageId="14" pageNumber="1008">ro</emphasis>
with slightly attenuate tip; 
<emphasis box="[536,555,229,253]" italics="true" pageId="18" pageNumber="1012">le</emphasis>
inserted on distinct tubercle; 
<emphasis box="[879,900,230,253]" italics="true" pageId="18" pageNumber="1012">in</emphasis>
erect. Distance 
<emphasis box="[1081,1134,229,253]" italics="true" pageId="18" pageNumber="1012">in–le</emphasis>
equally to 
<emphasis box="[1258,1313,229,253]" italics="true" pageId="18" pageNumber="1012">le–ro</emphasis>
. Slight carina developed between insertions of rostral setae. Bothridial seta (127–135) with long stalk and short, distinctly lanceolate head, roughened.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7159926" ID-Zenodo-Dep="7159926" httpUri="https://zenodo.org/record/7159926/files/figure.png" pageId="15" pageNumber="1009" startId="15.[119,186,2041,2062]" targetBox="[135,1476,227,1966]" targetPageId="15">
<paragraph blockId="15.[118,1525,2040,2085]" pageId="15" pageNumber="1009">
<emphasis bold="true" box="[119,216,2041,2062]" pageId="15" pageNumber="1009">Figure 10</emphasis>
<taxonomicName authority="Ermilov &amp; Martens, 2021" authorityName="Ermilov &amp; Martens" authorityYear="2021" box="[227,442,2040,2062]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="1009" phylum="Arthropoda" rank="species" species="schawalleri" status="n. sp.">
<emphasis box="[227,442,2040,2062]" italics="true" pageId="15" pageNumber="1009">Leobodes schawalleri</emphasis>
</taxonomicName>
<emphasis bold="true" box="[442,496,2041,2062]" pageId="15" pageNumber="1009">
<taxonomicNameLabel box="[442,496,2041,2062]" pageId="15" pageNumber="1009" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, adult: a – dorsal view (legs omitted); b – ventral view (gnathosoma and legs omitted); c – lateral view (gnathosoma and legs omitted). Scale bar 100 μm.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7159930" ID-Zenodo-Dep="7159930" box="[119,1346,1897,1919]" httpUri="https://zenodo.org/record/7159930/files/figure.png" pageId="16" pageNumber="1010" startId="16.[119,186,1898,1919]" targetBox="[103,1494,285,1843]" targetPageId="16">
<paragraph blockId="16.[119,1346,1897,1919]" box="[119,1346,1897,1919]" pageId="16" pageNumber="1010">
<emphasis bold="true" box="[119,215,1898,1919]" pageId="16" pageNumber="1010">Figure 11</emphasis>
<taxonomicName authority="Ermilov &amp; Martens, 2021" authorityName="Ermilov &amp; Martens" authorityYear="2021" box="[227,442,1897,1919]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="16" pageNumber="1010" phylum="Arthropoda" rank="species" species="schawalleri" status="n. sp.">
<emphasis box="[227,442,1897,1919]" italics="true" pageId="16" pageNumber="1010">Leobodes schawalleri</emphasis>
</taxonomicName>
<emphasis bold="true" box="[442,496,1898,1919]" pageId="16" pageNumber="1010">
<taxonomicNameLabel box="[442,496,1898,1919]" pageId="16" pageNumber="1010" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, adult, SEM micrographs: a, b – dorsal view; c – ventral view; d – ventrolateral view.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7159932" ID-Zenodo-Dep="7159932" box="[119,1424,1896,1918]" httpUri="https://zenodo.org/record/7159932/files/figure.png" pageId="17" pageNumber="1011" startId="17.[119,186,1897,1918]" targetBox="[103,1494,286,1841]" targetPageId="17">
<paragraph blockId="17.[119,1424,1896,1918]" box="[119,1424,1896,1918]" pageId="17" pageNumber="1011">
<emphasis bold="true" box="[119,216,1897,1918]" pageId="17" pageNumber="1011">Figure 12</emphasis>
<taxonomicName authority="Ermilov &amp; Martens, 2021" authorityName="Ermilov &amp; Martens" authorityYear="2021" box="[228,444,1896,1918]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="17" pageNumber="1011" phylum="Arthropoda" rank="species" species="schawalleri" status="n. sp.">
<emphasis box="[228,444,1896,1918]" italics="true" pageId="17" pageNumber="1011">Leobodes schawalleri</emphasis>
</taxonomicName>
<emphasis bold="true" box="[443,498,1897,1918]" pageId="17" pageNumber="1011">
<taxonomicNameLabel box="[443,498,1897,1918]" pageId="17" pageNumber="1011" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, adult, SEM micrographs: a – dorsoanterior view; b, c – lateral view; d – ventroanterior view.
</paragraph>
</caption>
<paragraph blockId="18.[491,1530,229,1290]" pageId="18" pageNumber="1012">
Notogaster – Notogastral tubercle present. Distance between medial notogastral condyles comparatively short (distinctly shorter than distance between setae 
<emphasis box="[1212,1279,368,392]" italics="true" pageId="18" pageNumber="1012">lm–lm</emphasis>
). Thirteen pairs of notogastral setae present: seta 
<emphasis box="[814,826,403,426]" italics="true" pageId="18" pageNumber="1012">c</emphasis>
(114–118) with phylliform basal part and attenuate mediodistal part and flexible tip, roughened, straight, directed anteromediad; 
<emphasis box="[1185,1198,437,461]" italics="true" pageId="18" pageNumber="1012">h</emphasis>
<subScript attach="left" box="[1199,1205,449,466]" fontSize="7" pageId="18" pageNumber="1012">3</subScript>
, 
<emphasis box="[1223,1236,438,461]" italics="true" pageId="18" pageNumber="1012">p</emphasis>
<subScript attach="left" box="[1236,1245,449,466]" fontSize="7" pageId="18" pageNumber="1012">1</subScript>
– 
<emphasis box="[1260,1273,438,461]" italics="true" pageId="18" pageNumber="1012">p</emphasis>
<subScript attach="left" box="[1273,1279,449,466]" fontSize="7" pageId="18" pageNumber="1012">3</subScript>
(57–65) and others (82–90) thickened, barbed. Opisthonotal gland opening and all lyrifissures distinct.
</paragraph>
<paragraph blockId="18.[491,1530,229,1290]" pageId="18" pageNumber="1012">
Gnathosoma – Mostly similar to 
<taxonomicName authority="Ermilov &amp; Martens, 2021" authorityName="Ermilov &amp; Martens" authorityYear="2021" box="[890,1056,506,530]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="1012" phylum="Arthropoda" rank="species" species="becki" status="n. sp.">
<emphasis box="[890,1056,506,530]" italics="true" pageId="18" pageNumber="1012">Leobodes becki</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1064,1125,506,530]" pageId="18" pageNumber="1012">
<taxonomicNameLabel box="[1064,1125,506,530]" pageId="18" pageNumber="1012" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Subcapitulum size: 143–151 × 123–131. Subcapitular setae (
<emphasis box="[808,821,541,564]" italics="true" pageId="18" pageNumber="1012">a</emphasis>
: 16; 
<emphasis box="[878,897,541,564]" italics="true" pageId="18" pageNumber="1012">m</emphasis>
, 
<emphasis box="[910,923,540,564]" italics="true" pageId="18" pageNumber="1012">h</emphasis>
: 65–69) with attenuate tip, slightly barbed; 
<emphasis box="[1391,1411,541,564]" italics="true" pageId="18" pageNumber="1012">a</emphasis>
thinnest. Palp length: 73–77. Length of postpalpal seta: 10. Chelicera length: 180–188. Length of 
<emphasis box="[1451,1489,575,599]" italics="true" pageId="18" pageNumber="1012">cha</emphasis>
: 53–57; 
<emphasis box="[572,610,609,633]" italics="true" pageId="18" pageNumber="1012">chb</emphasis>
: 22–24.
</paragraph>
<paragraph blockId="18.[491,1530,229,1290]" pageId="18" pageNumber="1012">
Epimeral and lateral podosomal regions – Epimeral setae 
<emphasis box="[1138,1165,644,668]" italics="true" pageId="18" pageNumber="1012">1b</emphasis>
, 
<emphasis box="[1178,1204,644,668]" italics="true" pageId="18" pageNumber="1012">3b</emphasis>
, 
<emphasis box="[1217,1249,644,668]" italics="true" pageId="18" pageNumber="1012">4a</emphasis>
(41–49), 
<emphasis box="[1346,1371,644,668]" italics="true" pageId="18" pageNumber="1012">3c</emphasis>
, 
<emphasis box="[1383,1408,644,668]" italics="true" pageId="18" pageNumber="1012">4c</emphasis>
(61–69) thickened, barbed; other setae (24–32) thin, with attenuate tip, slightly barbed; 
<emphasis box="[1352,1377,678,702]" italics="true" pageId="18" pageNumber="1012">4c</emphasis>
thickest. Ventrolateral tubercle on epimere II distinct. Discidium with triangular tip.
</paragraph>
<paragraph blockId="18.[491,1530,229,1290]" pageId="18" pageNumber="1012">
Anogenital region – Genital seta (28–32) thin, with attenuate tip, slightly barbed. Aggenital (36–41), anal (36–41) and adanal (
<emphasis box="[870,898,782,806]" italics="true" pageId="18" pageNumber="1012">ad</emphasis>
<subScript attach="left" box="[898,907,794,811]" fontSize="7" pageId="18" pageNumber="1012">1</subScript>
, 
<emphasis box="[924,952,782,806]" italics="true" pageId="18" pageNumber="1012">ad</emphasis>
<subScript attach="left" box="[952,958,794,811]" fontSize="7" pageId="18" pageNumber="1012">2</subScript>
: 49–53; 
<emphasis box="[1066,1094,782,806]" italics="true" pageId="18" pageNumber="1012">ad</emphasis>
<subScript attach="left" box="[1094,1100,794,811]" fontSize="7" pageId="18" pageNumber="1012">3</subScript>
: 36–41) setae thickened, barbed. Adanal lyrifissure distinct.
</paragraph>
<paragraph blockId="18.[491,1530,229,1290]" pageId="18" pageNumber="1012">
Legs – Mostly similar to 
<taxonomicName authority="Ermilov &amp; Martens, 2021" authorityName="Ermilov &amp; Martens" authorityYear="2021" box="[809,976,851,875]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="1012" phylum="Arthropoda" rank="species" species="becki" status="n. sp.">
<emphasis box="[809,976,851,875]" italics="true" pageId="18" pageNumber="1012">Leobodes becki</emphasis>
</taxonomicName>
<emphasis bold="true" box="[984,1046,852,876]" pageId="18" pageNumber="1012">
<taxonomicNameLabel box="[984,1046,852,876]" pageId="18" pageNumber="1012" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, but seta 
<emphasis box="[1154,1167,851,875]" italics="true" pageId="18" pageNumber="1012">l′</emphasis>
on trochanter III and 
<emphasis box="[1411,1429,851,875]" italics="true" pageId="18" pageNumber="1012">v′</emphasis>
on trochanters III, IV long, thick.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="1012" type="materials_examined">
<paragraph blockId="18.[491,1530,229,1290]" box="[531,1487,920,945]" pageId="18" pageNumber="1012">
<emphasis bold="true" box="[531,744,921,945]" pageId="18" pageNumber="1012">Material examined</emphasis>
— 
<collectingCountry box="[782,848,920,944]" name="Nepal" pageId="18" pageNumber="1012">Nepal</collectingCountry>
, Panchthar District, Dhorpar Kharka, 
<geoCoordinate box="[1269,1350,920,944]" degrees="27" direction="north" minutes="05" orientation="latitude" pageId="18" pageNumber="1012" precision="925" value="27.083334">27°05′N</geoCoordinate>
, 
<geoCoordinate box="[1358,1436,920,944]" degrees="87" direction="east" minutes="55" orientation="longitude" pageId="18" pageNumber="1012" precision="925" value="87.916664">87°55′E</geoCoordinate>
, 2700
</paragraph>
<paragraph blockId="18.[491,1530,229,1290]" pageId="18" pageNumber="1012">
m a.s.l., soil-litter in mature 
<emphasis box="[790,1076,955,979]" italics="true" pageId="18" pageNumber="1012">Rododendron -Lithocarpus</emphasis>
forest (sample No. 324), 
<date box="[1330,1470,955,979]" pageId="18" pageNumber="1012" value="1988-04-13" valueMax="1988-04-16" valueMin="1988-04-13">13–16.IV.1988</date>
(collected by J. Martens and W. Schawaller).
</paragraph>
<paragraph blockId="18.[491,1530,229,1290]" pageId="18" pageNumber="1012">
<emphasis bold="true" box="[531,711,1024,1048]" pageId="18" pageNumber="1012">Type deposition</emphasis>
— The 
<typeStatus box="[807,901,1024,1048]" pageId="18" pageNumber="1012">holotype</typeStatus>
and 
<specimenCount box="[961,1094,1025,1049]" pageId="18" pageNumber="1012" type="generic" typeStatus="paratype">one paratype</specimenCount>
are deposited in the collection of the 
<collectionCode box="[491,571,1058,1082]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="18" pageNumber="1012">SMNH</collectionCode>
; 
<specimenCount box="[582,712,1058,1083]" pageId="18" pageNumber="1012" type="generic" typeStatus="paratypes">15 paratypes</specimenCount>
are deposited in the collection of the 
<collectionCode box="[1136,1230,1058,1082]" pageId="18" pageNumber="1012">TSUMZ</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="1012" type="etymology">
<paragraph blockId="18.[491,1530,229,1290]" pageId="18" pageNumber="1012">
<emphasis bold="true" box="[531,654,1093,1117]" pageId="18" pageNumber="1012">Etymology</emphasis>
— The specific name is dedicated to Dr. Wolfgang Schawaller, well known entomologist, active participant and collector of the J. Martens expeditions to various parts of Europe and Asia, until recently working at State Museum of Natural History, Stuttgart, 
<collectingCountry box="[491,587,1196,1220]" name="Germany" pageId="18" pageNumber="1012">Germany</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="1012" type="discussion">
<paragraph blockId="18.[491,1530,229,1290]" pageId="18" pageNumber="1012">
<emphasis bold="true" box="[531,634,1231,1255]" pageId="18" pageNumber="1012">Remarks</emphasis>
— 
<taxonomicName authority="Ermilov &amp; Martens, 2021" authorityName="Ermilov &amp; Martens" authorityYear="2021" box="[676,916,1231,1255]" class="Arachnida" family="Nippobodidae" genus="Leobodes" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="1012" phylum="Arthropoda" rank="species" species="schawalleri" status="n. sp.">
<emphasis box="[676,916,1231,1255]" italics="true" pageId="18" pageNumber="1012">Leobodes schawalleri</emphasis>
</taxonomicName>
<emphasis bold="true" box="[916,976,1231,1255]" pageId="18" pageNumber="1012">
<taxonomicNameLabel box="[916,976,1231,1255]" pageId="18" pageNumber="1012" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs from all representatives of the genus by the presence of 13 pairs (versus 10) of notogastral setae.
</paragraph>
</subSubSection>
</treatment>
</document>