treatments-xml/data/72/2C/87/722C87C6FFCAFFEF7AB5FF52576DAAEC.xml

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<document ID-DOI="10.26107/RBZ-2023-0005" ID-ISSN="2345-7600" ID-Zenodo-Dep="7815677" ID-ZooBank="BA361B52-E36A-4C15-AB6B-FECA9CEF58AF" approvalRequired="6" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="5" checkinTime="1681181651611" checkinUser="felipe" docAuthor="Ng, Peter K. L. &amp; Tri, Ngo Van" docDate="2023" docId="722C87C6FFCAFFEF7AB5FF52576DAAEC" docLanguage="en" docName="RafflesBZool.71.7086.pdf" docOrigin="Raffles Bulletin of Zoology 71" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.4:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="4" docTitle="Dacuomon quynhi Ng &amp; Tri 2023, new species" docType="treatment" docVersion="3" lastPageNumber="76" masterDocId="8E15FFBEFFC8FFE97A4AFFEF561CAC12" masterDocTitle="A new genus and three new species of freshwater crab (Crustacea: Brachyura: Potamidae) from central and northern Vietnam" masterLastPageNumber="86" masterPageNumber="70" pageNumber="72" updateTime="1681184872437" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new genus and three new species of freshwater crab (Crustacea: Brachyura: Potamidae) from central and northern Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ng, Peter K. L.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tri, Ngo Van</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Raffles Bulletin of Zoology</mods:title>
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<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-03</mods:number>
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<mods:detail type="volume">
<mods:number>71</mods:number>
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<mods:start>70</mods:start>
<mods:end>86</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.26107/RBZ-2023-0005</mods:identifier>
<mods:identifier type="ISSN">2345-7600</mods:identifier>
<mods:identifier type="Zenodo-Dep">7815677</mods:identifier>
<mods:identifier type="ZooBank">BA361B52-E36A-4C15-AB6B-FECA9CEF58AF</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:722C87C6FFCAFFEF7AB5FF52576DAAEC" httpUri="http://treatment.plazi.org/id/722C87C6FFCAFFEF7AB5FF52576DAAEC" lastPageId="6" lastPageNumber="76" pageId="2" pageNumber="72">
<subSubSection box="[255,611,188,213]" pageId="2" pageNumber="72" type="nomenclature">
<paragraph blockId="2.[255,611,188,245]" box="[255,611,188,213]" pageId="2" pageNumber="72">
<heading bold="true" box="[255,611,188,213]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="72" reason="2">
<emphasis bold="true" box="[255,611,188,213]" pageId="2" pageNumber="72">
<taxonomicName authority="Ng &amp; Tri, 2023" authorityName="Ng &amp; Tri" authorityYear="2023" box="[255,462,188,212]" class="Malacostraca" family="Potamidae" genus="Dacuomon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="72" phylum="Arthropoda" rank="species" species="quynhi" status="new species">
<emphasis bold="true" box="[255,462,188,212]" italics="true" pageId="2" pageNumber="72">Dacuomon quynhi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[477,611,189,213]" pageId="2" pageNumber="72" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[373,492,221,245]" pageId="2" pageNumber="72" type="description">
<paragraph blockId="2.[255,611,188,245]" box="[373,492,221,245]" pageId="2" pageNumber="72">
(
<figureCitation box="[381,484,221,245]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="3.[136,171,1700,1722]" captionStartId-1="4.[113,148,2036,2058]" captionStartId-2="5.[136,171,2035,2057]" captionStartId-3="6.[113,148,1055,1077]" captionTargetBox-0="[136,1474,189,1684]" captionTargetBox-1="[115,1448,189,2020]" captionTargetBox-2="[138,1471,189,2018]" captionTargetBox-3="[114,1461,189,1038]" captionTargetId-0="figure-7@3.[136,1474,189,1684]" captionTargetId-1="figure-13@4.[115,1448,189,2020]" captionTargetId-2="figure-7@5.[138,1471,189,2018]" captionTargetId-3="figure-12@6.[90,1510,179,1062]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="Fig. 1. Dacuomon quynhi, new genus, new species, colour in life. A, B, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam; CE, paratype male (23.9 × 20.4 mm) (ZRC 2022.0048), Phu Yên Province, Vietnam; F, type locality, Xuan Lam commune, Phu Yên Province, Vietnam." captionText-1="Fig. 2. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, subfrontal view of cephalothorax showing setose third maxillipeds; E, frontal view showing antennules, antennae and epistome; F, outer view of chelae." captionText-2="Fig. 3. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, left third maxilliped; B, ventral view of cephalothorax and pleon; B, male pleon; D, sternopleonal cavity showing G1 in situ; E, left G1 (ventral view); F, left G1 (dorsal view); G, terminal segment of left G1 (ventral view); H, terminal segment of left G1 (dorsal view); I, left G2." captionText-3="Fig. 4. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, left G1 (ventral view); B, left G1 (dorsal view); C, terminal segment of left G1 (ventral view); D, terminal segment of left G1 (dorsal view); E, left G2. Scales: A, D = 1.0 mm B, C = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7816165" figureDoi-1="http://doi.org/10.5281/zenodo.7816167" figureDoi-2="http://doi.org/10.5281/zenodo.7816169" figureDoi-3="http://doi.org/10.5281/zenodo.7816171" httpUri-0="https://zenodo.org/record/7816165/files/figure.png" httpUri-1="https://zenodo.org/record/7816167/files/figure.png" httpUri-2="https://zenodo.org/record/7816169/files/figure.png" httpUri-3="https://zenodo.org/record/7816171/files/figure.png" pageId="2" pageNumber="72">Figs. 14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="72" type="materials_examined">
<paragraph blockId="2.[113,754,285,501]" pageId="2" pageNumber="72">
<emphasis bold="true" box="[113,349,285,309]" pageId="2" pageNumber="72">Material examined.</emphasis>
<materialsCitation collectingDate="2022-01-13" collectionCode="V, T" country="Vietnam" elevation="160" latitude="13.461991" location="Xuan Lam" longLatPrecision="1" longitude="109.11926" municipality="Dong Xuan district" pageId="2" pageNumber="72" specimenCode="ZRC 2022.0047" specimenCount="1" stateProvince="Phu Yen" typeStatus="holotype">
<typeStatus box="[359,467,285,309]" pageId="2" pageNumber="72">Holotype</typeStatus>
: male (27.7 ×
<quantity box="[641,745,285,309]" metricMagnitude="-2" metricUnit="m" metricValue="2.38" pageId="2" pageNumber="72" unit="mm" value="23.8">23.8 mm</quantity>
) (
<specimenCode box="[121,302,317,341]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="72">ZRC 2022.0047</specimenCode>
), intermittent small stream in
<location LSID="urn:lsid:plazi:treatment:722C87C6FFCAFFEF7AB5FF52576DAAEC:FF5A600BFFCAFFEB7836FED254ECAD46" box="[636,752,317,340]" country="Vietnam" latitude="13.461991" longLatPrecision="1" longitude="109.11926" municipality="Dong Xuan district" name="Xuan Lam" pageId="2" pageNumber="72" stateProvince="Phu Yen">Xuan Lam</location>
commune,
<collectingMunicipality box="[235,449,349,373]" pageId="2" pageNumber="72">Dong Xuan district</collectingMunicipality>
,
<collectingRegion box="[463,666,349,373]" country="Vietnam" name="Phu Yen" pageId="2" pageNumber="72">Phu Yên Province</collectingRegion>
, central
<collectingCountry box="[113,208,381,405]" name="Vietnam" pageId="2" pageNumber="72">Vietnam</collectingCountry>
, ca.
<geoCoordinate box="[261,421,379,404]" degrees="13" direction="north" minutes="27" orientation="latitude" pageId="2" pageNumber="72" precision="1" seconds="43.17" value="13.461991">13°2743.17″N</geoCoordinate>
<geoCoordinate box="[430,603,379,404]" degrees="109" direction="east" minutes="07" orientation="longitude" pageId="2" pageNumber="72" precision="1" seconds="09.34" value="109.11926">109°0709.34″E</geoCoordinate>
,
<elevation box="[617,746,381,405]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="72" unit="m" value="160.0">
<quantity box="[617,687,381,405]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="72" unit="m" value="160.0">160 m</quantity>
a.s.l.
</elevation>
, coll.
<collectionCode box="[169,192,413,436]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="72" type="Museum">V</collectionCode>
.
<collectionCode box="[195,214,413,436]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="72">T</collectionCode>
. Ngo,
<date box="[288,474,413,437]" pageId="2" pageNumber="72" value="2022-01-13">
<collectingDate box="[288,474,413,437]" pageId="2" pageNumber="72" value="2022-01-13">13 January 2022</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation collectingDate="2022-01-13" collectionCode="ZRC" country="Vietnam" elevation="160" latitude="13.461991" location="Xuan Lam" longLatPrecision="1" longitude="109.11926" municipality="Dong Xuan district" pageId="2" pageNumber="72" specimenCode="ZRC 2022.0048" specimenCount="2" specimenCount-male="2" stateProvince="Phu Yen" typeStatus="paratype">
<typeStatus box="[486,594,413,437]" pageId="2" pageNumber="72">Paratypes</typeStatus>
:
<specimenCount box="[606,690,413,437]" pageId="2" pageNumber="72" type="male">2 males</specimenCount>
(23.9 ×
<quantity box="[137,235,445,469]" metricMagnitude="-2" metricUnit="m" metricValue="2.04" pageId="2" pageNumber="72" unit="mm" value="20.4">20.4 mm</quantity>
, 22.9 ×
<quantity box="[326,423,445,469]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="2" pageNumber="72" unit="mm" value="19.5">19.5 mm</quantity>
) (
<specimenCode box="[448,627,445,469]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="72">ZRC 2022.0048</specimenCode>
), same data as holotype
</materialsCitation>
.
</paragraph>
<paragraph blockId="2.[113,752,541,725]" pageId="2" pageNumber="72">
<emphasis bold="true" box="[115,379,541,565]" pageId="2" pageNumber="72">Comparative material.</emphasis>
<taxonomicName authority="(Dang, 1975)" baseAuthorityName="Dang" baseAuthorityYear="1975" class="Malacostraca" family="Potamidae" genus="Kukrimon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="72" phylum="Arthropoda" rank="species" species="cucphuongense">
<emphasis box="[387,669,541,565]" italics="true" pageId="2" pageNumber="72">Kukrimon cucphuongense</emphasis>
(
<bibRefCitation author="Dang NT" pageId="2" pageNumber="72" pagination="65 - 78" refId="ref8269" refString="Dang NT (1975) Phan loai tom cua nuoc ngot mien bac Viet Nam [= The identities of North Vietnamese freshwater shrimp and crabs]. Tap san Sinh Vat - Dia Hoc [= Journal of Biology and Geology, National Institute of Science, Hanoi], 13 (3): 65 - 78. [In Vietnamese with a French summary]" type="journal article" year="1975">Đăng, 1975</bibRefCitation>
)
</taxonomicName>
,
<materialsCitation collectingDate="1997-09-16" collectionCode="ZRC" collectorName="D. C. J. Yeo &amp; H. H. Ng" country="Vietnam" location="Cuc Phuong National Park" pageId="2" pageNumber="72" specimenCode="ZRC 2022.0058" specimenCount="11" specimenCount-female="2" specimenCount-juvenile="3" specimenCount-male="6" stateProvince="Ninh Binh">
<specimenCount box="[190,274,573,597]" pageId="2" pageNumber="72" type="male">6 males</specimenCount>
(largest 22.3 ×
<quantity box="[448,548,573,597]" metricMagnitude="-2" metricUnit="m" metricValue="1.92" pageId="2" pageNumber="72" unit="mm" value="19.2">19.2 mm</quantity>
),
<specimenCount box="[568,673,573,597]" pageId="2" pageNumber="72" type="female">2 females</specimenCount>
(larger 18.3 ×
<quantity box="[193,294,605,629]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" pageId="2" pageNumber="72" unit="mm" value="15.6">15.6 mm</quantity>
),
<specimenCount box="[315,434,605,629]" pageId="2" pageNumber="72" type="juvenile">3 juveniles</specimenCount>
(
<specimenCode box="[450,631,605,629]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="72">ZRC 2022.0058</specimenCode>
), stream in
<location LSID="urn:lsid:plazi:treatment:722C87C6FFCAFFEF7AB5FF52576DAAEC:FF5A600BFFCAFFEB7A3BFD925780AE87" box="[113,412,637,661]" country="Vietnam" name="Cuc Phuong National Park" pageId="2" pageNumber="72" stateProvince="Ninh Binh">Cuc Phuong National Park</location>
, about
<quantity box="[495,566,637,661]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="72" unit="km" value="16.0">16 km</quantity>
from main gate,
<collectingRegion box="[113,329,669,693]" country="Vietnam" name="Ninh Binh" pageId="2" pageNumber="72">Ninh Binh Province</collectingRegion>
, northeastern
<collectingCountry box="[477,572,669,693]" name="Vietnam" pageId="2" pageNumber="72">Vietnam</collectingCountry>
, coll.
<collectorName box="[634,752,669,693]" pageId="2" pageNumber="72">D.C.J. Yeo</collectorName>
&amp;
<collectorName box="[142,238,701,725]" pageId="2" pageNumber="72">H.H. Ng</collectorName>
,
<date box="[251,462,701,725]" pageId="2" pageNumber="72" value="1997-09-16">
<collectingDate box="[251,462,701,725]" pageId="2" pageNumber="72" value="1997-09-16">16 September 1997</collectingDate>
</date>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection box="[113,386,765,789]" pageId="2" pageNumber="72" type="diagnosis">
<paragraph blockId="2.[113,386,765,789]" box="[113,386,765,789]" pageId="2" pageNumber="72">
<emphasis bold="true" box="[113,230,765,789]" pageId="2" pageNumber="72">Diagnosis.</emphasis>
As for genus.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="76" pageId="2" pageNumber="72" type="description">
<paragraph blockId="2.[112,753,829,1941]" pageId="2" pageNumber="72">
<emphasis bold="true" box="[113,437,829,853]" pageId="2" pageNumber="72">
Description of male
<typeStatus box="[334,432,829,853]" pageId="2" pageNumber="72">holotype</typeStatus>
.
</emphasis>
Carapace rounded, wider than long (ratio 1.16); dorsal surface gently convex in frontal view, glabrous, regions not prominently defined or inflated (
<figureCitation captionStart="Fig" captionStartId="3.[136,171,1700,1722]" captionTargetBox="[136,1474,189,1684]" captionTargetId="figure-7@3.[136,1474,189,1684]" captionTargetPageId="3" captionText="Fig. 1. Dacuomon quynhi, new genus, new species, colour in life. A, B, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam; CE, paratype male (23.9 × 20.4 mm) (ZRC 2022.0048), Phu Yên Province, Vietnam; F, type locality, Xuan Lam commune, Phu Yên Province, Vietnam." figureDoi="http://doi.org/10.5281/zenodo.7816165" httpUri="https://zenodo.org/record/7816165/files/figure.png" pageId="2" pageNumber="72">Figs. 1A, B</figureCitation>
,
<figureCitation box="[186,251,925,949]" captionStart="Fig" captionStartId="4.[113,148,2036,2058]" captionTargetBox="[115,1448,189,2020]" captionTargetId="figure-13@4.[115,1448,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, subfrontal view of cephalothorax showing setose third maxillipeds; E, frontal view showing antennules, antennae and epistome; F, outer view of chelae." figureDoi="http://doi.org/10.5281/zenodo.7816167" httpUri="https://zenodo.org/record/7816167/files/figure.png" pageId="2" pageNumber="72">2AD</figureCitation>
). Frontal regions with low granules, postorbital regions almost smooth; lateral surface of anterolateral and branchial regions covered with low granules, not prominent; mesogastric, urogastric, cardiac and intestinal regions almost smooth; suborbital, subhepatic and subbranchial regions with low granules and striae; pterygostomial regions with low granules (
<figureCitation box="[264,377,1117,1141]" captionStart="Fig" captionStartId="4.[113,148,2036,2058]" captionTargetBox="[115,1448,189,2020]" captionTargetId="figure-13@4.[115,1448,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, subfrontal view of cephalothorax showing setose third maxillipeds; E, frontal view showing antennules, antennae and epistome; F, outer view of chelae." figureDoi="http://doi.org/10.5281/zenodo.7816167" httpUri="https://zenodo.org/record/7816167/files/figure.png" pageId="2" pageNumber="72">Fig. 2BD</figureCitation>
). Epigastric cristae visible, rugose, not sharp, separated by shallow median Y-shaped furrow; epigastric cristae just anterior to postorbital cristae, separated from it by short furrow; postorbital crista distinct, sharp, raised, subparallel to frontal margin, outer edge lower, separated by small fissure from strong crista leading to anterolateral margin (
<figureCitation box="[343,453,1309,1333]" captionStart="Fig" captionStartId="4.[113,148,2036,2058]" captionTargetBox="[115,1448,189,2020]" captionTargetId="figure-13@4.[115,1448,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, subfrontal view of cephalothorax showing setose third maxillipeds; E, frontal view showing antennules, antennae and epistome; F, outer view of chelae." figureDoi="http://doi.org/10.5281/zenodo.7816167" httpUri="https://zenodo.org/record/7816167/files/figure.png" pageId="2" pageNumber="72">Fig. 2B, D</figureCitation>
). Cervical grooves shallow, not reaching lateral margins, barely connected to H-shaped median gastric groove (
<figureCitation box="[388,475,1373,1397]" captionStart="Fig" captionStartId="4.[113,148,2036,2058]" captionTargetBox="[115,1448,189,2020]" captionTargetId="figure-13@4.[115,1448,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, subfrontal view of cephalothorax showing setose third maxillipeds; E, frontal view showing antennules, antennae and epistome; F, outer view of chelae." figureDoi="http://doi.org/10.5281/zenodo.7816167" httpUri="https://zenodo.org/record/7816167/files/figure.png" pageId="2" pageNumber="72">Fig. 2B</figureCitation>
). Frontal margin gently sinuous in frontal and dorsal views (
<figureCitation box="[518,636,1405,1429]" captionStart="Fig" captionStartId="4.[113,148,2036,2058]" captionTargetBox="[115,1448,189,2020]" captionTargetId="figure-13@4.[115,1448,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, subfrontal view of cephalothorax showing setose third maxillipeds; E, frontal view showing antennules, antennae and epistome; F, outer view of chelae." figureDoi="http://doi.org/10.5281/zenodo.7816167" httpUri="https://zenodo.org/record/7816167/files/figure.png" pageId="2" pageNumber="72">Fig. 2B, D</figureCitation>
). External orbital tooth distinct, triangular, outer margin more than twice length of inner margin, demarcated from rest of anterolateral margin by distinct V-shaped cleft; epibranchial tooth small, sharp (
<figureCitation box="[185,263,1533,1557]" captionStart="Fig" captionStartId="4.[113,148,2036,2058]" captionTargetBox="[115,1448,189,2020]" captionTargetId="figure-13@4.[115,1448,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, subfrontal view of cephalothorax showing setose third maxillipeds; E, frontal view showing antennules, antennae and epistome; F, outer view of chelae." figureDoi="http://doi.org/10.5281/zenodo.7816167" httpUri="https://zenodo.org/record/7816167/files/figure.png" pageId="2" pageNumber="72">Fig. 2B</figureCitation>
). Anterolateral margins convex, cristate, lined with low granules, appearing entire (
<figureCitation box="[502,584,1565,1589]" captionStart="Fig" captionStartId="4.[113,148,2036,2058]" captionTargetBox="[115,1448,189,2020]" captionTargetId="figure-13@4.[115,1448,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, subfrontal view of cephalothorax showing setose third maxillipeds; E, frontal view showing antennules, antennae and epistome; F, outer view of chelae." figureDoi="http://doi.org/10.5281/zenodo.7816167" httpUri="https://zenodo.org/record/7816167/files/figure.png" pageId="2" pageNumber="72">Fig. 2B</figureCitation>
). Posterolateral margin gently sinuous, converging towards convex posterior carapace margin (
<figureCitation box="[318,408,1629,1653]" captionStart="Fig" captionStartId="4.[113,148,2036,2058]" captionTargetBox="[115,1448,189,2020]" captionTargetId="figure-13@4.[115,1448,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, subfrontal view of cephalothorax showing setose third maxillipeds; E, frontal view showing antennules, antennae and epistome; F, outer view of chelae." figureDoi="http://doi.org/10.5281/zenodo.7816167" httpUri="https://zenodo.org/record/7816167/files/figure.png" pageId="2" pageNumber="72">Fig. 2B</figureCitation>
). Orbits subovate; eye filling orbital space; eye peduncle relatively short, stout; cornea large, round, pigmented (
<figureCitation box="[393,509,1693,1717]" captionStart="Fig" captionStartId="4.[113,148,2036,2058]" captionTargetBox="[115,1448,189,2020]" captionTargetId="figure-13@4.[115,1448,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, subfrontal view of cephalothorax showing setose third maxillipeds; E, frontal view showing antennules, antennae and epistome; F, outer view of chelae." figureDoi="http://doi.org/10.5281/zenodo.7816167" httpUri="https://zenodo.org/record/7816167/files/figure.png" pageId="2" pageNumber="72">Fig. 2BD</figureCitation>
). Supraorbital margin slightly sinuous, entire (
<figureCitation box="[364,441,1725,1749]" captionStart="Fig" captionStartId="4.[113,148,2036,2058]" captionTargetBox="[115,1448,189,2020]" captionTargetId="figure-13@4.[115,1448,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, subfrontal view of cephalothorax showing setose third maxillipeds; E, frontal view showing antennules, antennae and epistome; F, outer view of chelae." figureDoi="http://doi.org/10.5281/zenodo.7816167" httpUri="https://zenodo.org/record/7816167/files/figure.png" pageId="2" pageNumber="72">Fig. 2B</figureCitation>
). Suborbital margin concave, complete, lined with small, low granules; inner angle with low tooth (
<figureCitation box="[225,333,1789,1813]" captionStart="Fig" captionStartId="4.[113,148,2036,2058]" captionTargetBox="[115,1448,189,2020]" captionTargetId="figure-13@4.[115,1448,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, subfrontal view of cephalothorax showing setose third maxillipeds; E, frontal view showing antennules, antennae and epistome; F, outer view of chelae." figureDoi="http://doi.org/10.5281/zenodo.7816167" httpUri="https://zenodo.org/record/7816167/files/figure.png" pageId="2" pageNumber="72">Fig. 2C, D</figureCitation>
). Antennae short, stretching across base of eyes; antennules short, folding transversely in rectangular fossa (
<figureCitation box="[184,298,1853,1877]" captionStart="Fig" captionStartId="4.[113,148,2036,2058]" captionTargetBox="[115,1448,189,2020]" captionTargetId="figure-13@4.[115,1448,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, subfrontal view of cephalothorax showing setose third maxillipeds; E, frontal view showing antennules, antennae and epistome; F, outer view of chelae." figureDoi="http://doi.org/10.5281/zenodo.7816167" httpUri="https://zenodo.org/record/7816167/files/figure.png" pageId="2" pageNumber="72">Fig. 2CE</figureCitation>
). Posterior margin of epistome with low, broadly triangular median structure, lateral margins gently concave (
<figureCitation box="[217,330,1917,1941]" captionStart="Fig" captionStartId="4.[113,148,2036,2058]" captionTargetBox="[115,1448,189,2020]" captionTargetId="figure-13@4.[115,1448,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, subfrontal view of cephalothorax showing setose third maxillipeds; E, frontal view showing antennules, antennae and epistome; F, outer view of chelae." figureDoi="http://doi.org/10.5281/zenodo.7816167" httpUri="https://zenodo.org/record/7816167/files/figure.png" pageId="2" pageNumber="72">Fig. 2CE</figureCitation>
).
</paragraph>
<paragraph blockId="2.[113,752,1981,2101]" lastBlockId="2.[812,1451,189,309]" pageId="2" pageNumber="72">
Third maxillipeds covering most of buccal cavity when closed; covered with relatively dense long stiff simple setae which do not obscure surface; ischium subrectangular, with distinct oblique median groove, surface mostly smooth; merus subquadrate, surface almost smooth, margins cristate, anteroexternal angle rounded; exopod slender, reaching to about half length of merus, with elongate flagellum that reaches across width of merus (
<figureCitation box="[1159,1289,285,309]" captionStart="Fig" captionStartId="3.[136,171,1700,1722]" captionTargetBox="[136,1474,189,1684]" captionTargetId="figure-7@3.[136,1474,189,1684]" captionTargetPageId="3" captionText="Fig. 1. Dacuomon quynhi, new genus, new species, colour in life. A, B, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam; CE, paratype male (23.9 × 20.4 mm) (ZRC 2022.0048), Phu Yên Province, Vietnam; F, type locality, Xuan Lam commune, Phu Yên Province, Vietnam." figureDoi="http://doi.org/10.5281/zenodo.7816165" httpUri="https://zenodo.org/record/7816165/files/figure.png" pageId="2" pageNumber="72">Figs. 1B, D</figureCitation>
,
<figureCitation box="[1304,1335,285,309]" captionStart="Fig" captionStartId="5.[136,171,2035,2057]" captionTargetBox="[138,1471,189,2018]" captionTargetId="figure-7@5.[138,1471,189,2018]" captionTargetPageId="5" captionText="Fig. 3. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, left third maxilliped; B, ventral view of cephalothorax and pleon; B, male pleon; D, sternopleonal cavity showing G1 in situ; E, left G1 (ventral view); F, left G1 (dorsal view); G, terminal segment of left G1 (ventral view); H, terminal segment of left G1 (dorsal view); I, left G2." figureDoi="http://doi.org/10.5281/zenodo.7816169" httpUri="https://zenodo.org/record/7816169/files/figure.png" pageId="2" pageNumber="72">3A</figureCitation>
).
</paragraph>
<paragraph blockId="2.[811,1452,349,821]" pageId="2" pageNumber="72">
Chelipeds slightly asymmetrical, right larger (
<figureCitation box="[1319,1436,349,373]" captionStart="Fig" captionStartId="4.[113,148,2036,2058]" captionTargetBox="[115,1448,189,2020]" captionTargetId="figure-13@4.[115,1448,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, subfrontal view of cephalothorax showing setose third maxillipeds; E, frontal view showing antennules, antennae and epistome; F, outer view of chelae." figureDoi="http://doi.org/10.5281/zenodo.7816167" httpUri="https://zenodo.org/record/7816167/files/figure.png" pageId="2" pageNumber="72">Fig. 2A, F</figureCitation>
). Margins of basis-ischium and merus with numerous long setae. Anterior margin of basis-ischium lined with several small low tubercles and granules; margins of merus lined with prominent low tubercles, with larger tubercle on subdistal surface (
<figureCitation box="[904,986,509,533]" captionStart="Fig" captionStartId="5.[136,171,2035,2057]" captionTargetBox="[138,1471,189,2018]" captionTargetId="figure-7@5.[138,1471,189,2018]" captionTargetPageId="5" captionText="Fig. 3. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, left third maxilliped; B, ventral view of cephalothorax and pleon; B, male pleon; D, sternopleonal cavity showing G1 in situ; E, left G1 (ventral view); F, left G1 (dorsal view); G, terminal segment of left G1 (ventral view); H, terminal segment of left G1 (dorsal view); I, left G2." figureDoi="http://doi.org/10.5281/zenodo.7816169" httpUri="https://zenodo.org/record/7816169/files/figure.png" pageId="2" pageNumber="72">Fig. 3C</figureCitation>
). Outer surface of carpus covered with low tubercles, inner distal angle with large sharp tooth and small sub-basal tubercle (
<figureCitation box="[1020,1103,573,597]" captionStart="Fig" captionStartId="4.[113,148,2036,2058]" captionTargetBox="[115,1448,189,2020]" captionTargetId="figure-13@4.[115,1448,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, subfrontal view of cephalothorax showing setose third maxillipeds; E, frontal view showing antennules, antennae and epistome; F, outer view of chelae." figureDoi="http://doi.org/10.5281/zenodo.7816167" httpUri="https://zenodo.org/record/7816167/files/figure.png" pageId="2" pageNumber="72">Fig. 2A</figureCitation>
). Outer surface of chela covered with prominent tubercles and granules (
<figureCitation box="[1240,1319,605,629]" captionStart="Fig" captionStartId="4.[113,148,2036,2058]" captionTargetBox="[115,1448,189,2020]" captionTargetId="figure-13@4.[115,1448,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, subfrontal view of cephalothorax showing setose third maxillipeds; E, frontal view showing antennules, antennae and epistome; F, outer view of chelae." figureDoi="http://doi.org/10.5281/zenodo.7816167" httpUri="https://zenodo.org/record/7816167/files/figure.png" pageId="2" pageNumber="72">Fig. 2F</figureCitation>
). Fingers of major chela long, gently curved, longer than palm, outer surface of both fingers lined with 2 rows of pits; cutting edges of both fingers with variously sized sharp teeth and denticles; dorsal margin of dactylus with low tubercles and granules (
<figureCitation box="[920,998,765,789]" captionStart="Fig" captionStartId="4.[113,148,2036,2058]" captionTargetBox="[115,1448,189,2020]" captionTargetId="figure-13@4.[115,1448,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, subfrontal view of cephalothorax showing setose third maxillipeds; E, frontal view showing antennules, antennae and epistome; F, outer view of chelae." figureDoi="http://doi.org/10.5281/zenodo.7816167" httpUri="https://zenodo.org/record/7816167/files/figure.png" pageId="2" pageNumber="72">Fig. 2F</figureCitation>
). Fingers of minor chela similar to major chela in form but more slender (
<figureCitation box="[1169,1252,797,821]" captionStart="Fig" captionStartId="4.[113,148,2036,2058]" captionTargetBox="[115,1448,189,2020]" captionTargetId="figure-13@4.[115,1448,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, subfrontal view of cephalothorax showing setose third maxillipeds; E, frontal view showing antennules, antennae and epistome; F, outer view of chelae." figureDoi="http://doi.org/10.5281/zenodo.7816167" httpUri="https://zenodo.org/record/7816167/files/figure.png" pageId="2" pageNumber="72">Fig. 2F</figureCitation>
).
</paragraph>
<paragraph blockId="2.[811,1453,861,1269]" pageId="2" pageNumber="72">
Ambulatory legs relatively, slender; second pair longest, last pair shortest (
<figureCitation box="[964,1062,893,917]" captionStart="Fig" captionStartId="3.[136,171,1700,1722]" captionTargetBox="[136,1474,189,1684]" captionTargetId="figure-7@3.[136,1474,189,1684]" captionTargetPageId="3" captionText="Fig. 1. Dacuomon quynhi, new genus, new species, colour in life. A, B, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam; CE, paratype male (23.9 × 20.4 mm) (ZRC 2022.0048), Phu Yên Province, Vietnam; F, type locality, Xuan Lam commune, Phu Yên Province, Vietnam." figureDoi="http://doi.org/10.5281/zenodo.7816165" httpUri="https://zenodo.org/record/7816165/files/figure.png" pageId="2" pageNumber="72">Figs. 1A</figureCitation>
,
<figureCitation box="[1079,1111,893,917]" captionStart="Fig" captionStartId="4.[113,148,2036,2058]" captionTargetBox="[115,1448,189,2020]" captionTargetId="figure-13@4.[115,1448,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, subfrontal view of cephalothorax showing setose third maxillipeds; E, frontal view showing antennules, antennae and epistome; F, outer view of chelae." figureDoi="http://doi.org/10.5281/zenodo.7816167" httpUri="https://zenodo.org/record/7816167/files/figure.png" pageId="2" pageNumber="72">2A</figureCitation>
). Merus relatively short, outer surface gently rugose, dorsal margin uneven, subcristate, without subdistal spine or tooth; carpus rugose, dorsal margin cristate, outer surface with low submedian crista on first to third legs, that on fourth leg smooth; dorsal margin of propodus with crista, outer surface with low, submedian crista; dactylus relatively long, gently curved, quadrate in cross section, margins with short, sharp chitinous spines; margins of merus, carpus and propodus of first ambulatory leg with dense, long stiff simple setae which partially obscure surface; setae on other legs less dense and distinctly shorter (
<figureCitation box="[821,916,1245,1269]" captionStart="Fig" captionStartId="3.[136,171,1700,1722]" captionTargetBox="[136,1474,189,1684]" captionTargetId="figure-7@3.[136,1474,189,1684]" captionTargetPageId="3" captionText="Fig. 1. Dacuomon quynhi, new genus, new species, colour in life. A, B, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam; CE, paratype male (23.9 × 20.4 mm) (ZRC 2022.0048), Phu Yên Province, Vietnam; F, type locality, Xuan Lam commune, Phu Yên Province, Vietnam." figureDoi="http://doi.org/10.5281/zenodo.7816165" httpUri="https://zenodo.org/record/7816165/files/figure.png" pageId="2" pageNumber="72">Figs. 1B</figureCitation>
,
<figureCitation box="[931,962,1245,1269]" captionStart="Fig" captionStartId="4.[113,148,2036,2058]" captionTargetBox="[115,1448,189,2020]" captionTargetId="figure-13@4.[115,1448,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, subfrontal view of cephalothorax showing setose third maxillipeds; E, frontal view showing antennules, antennae and epistome; F, outer view of chelae." figureDoi="http://doi.org/10.5281/zenodo.7816167" httpUri="https://zenodo.org/record/7816167/files/figure.png" pageId="2" pageNumber="72">2A</figureCitation>
).
</paragraph>
<paragraph blockId="2.[811,1454,1309,1781]" pageId="2" pageNumber="72">
Thoracic sternum, notably sternites 3 and 4, as long as broad, surface almost smooth (
<figureCitation box="[1157,1278,1341,1365]" captionStart="Fig" captionStartId="5.[136,171,2035,2057]" captionTargetBox="[138,1471,189,2018]" captionTargetId="figure-7@5.[138,1471,189,2018]" captionTargetPageId="5" captionText="Fig. 3. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, left third maxilliped; B, ventral view of cephalothorax and pleon; B, male pleon; D, sternopleonal cavity showing G1 in situ; E, left G1 (ventral view); F, left G1 (dorsal view); G, terminal segment of left G1 (ventral view); H, terminal segment of left G1 (dorsal view); I, left G2." figureDoi="http://doi.org/10.5281/zenodo.7816169" httpUri="https://zenodo.org/record/7816169/files/figure.png" pageId="2" pageNumber="72">Fig. 3C, D</figureCitation>
). Sternites 1, 2 completely fused to form broadly triangular plate; separated from sternite 3 by distinct, gently concave suture (directed anteriorly); sternites 3, 4 completely fused without discernible groove or suture, only lateral notches visible (
<figureCitation box="[1346,1435,1469,1493]" captionStart="Fig" captionStartId="5.[136,171,2035,2057]" captionTargetBox="[138,1471,189,2018]" captionTargetId="figure-7@5.[138,1471,189,2018]" captionTargetPageId="5" captionText="Fig. 3. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, left third maxilliped; B, ventral view of cephalothorax and pleon; B, male pleon; D, sternopleonal cavity showing G1 in situ; E, left G1 (ventral view); F, left G1 (dorsal view); G, terminal segment of left G1 (ventral view); H, terminal segment of left G1 (dorsal view); I, left G2." figureDoi="http://doi.org/10.5281/zenodo.7816169" httpUri="https://zenodo.org/record/7816169/files/figure.png" pageId="2" pageNumber="72">Fig. 3C</figureCitation>
). Sutures 4/5, 5/6, 6/7 and 7/8 medially interrupted; median longitudinal groove on sternites 7 and 8 (
<figureCitation box="[1280,1370,1533,1557]" captionStart="Fig" captionStartId="5.[136,171,2035,2057]" captionTargetBox="[138,1471,189,2018]" captionTargetId="figure-7@5.[138,1471,189,2018]" captionTargetPageId="5" captionText="Fig. 3. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, left third maxilliped; B, ventral view of cephalothorax and pleon; B, male pleon; D, sternopleonal cavity showing G1 in situ; E, left G1 (ventral view); F, left G1 (dorsal view); G, terminal segment of left G1 (ventral view); H, terminal segment of left G1 (dorsal view); I, left G2." figureDoi="http://doi.org/10.5281/zenodo.7816169" httpUri="https://zenodo.org/record/7816169/files/figure.png" pageId="2" pageNumber="72">Fig. 3D</figureCitation>
). Penis coxal, emerging on condyle of coxa of fourth ambulatory leg. Sternopleonal cavity deep, anterior edge reaching imaginary line connecting median part of cheliped coxae (
<figureCitation box="[1349,1437,1629,1653]" captionStart="Fig" captionStartId="5.[136,171,2035,2057]" captionTargetBox="[138,1471,189,2018]" captionTargetId="figure-7@5.[138,1471,189,2018]" captionTargetPageId="5" captionText="Fig. 3. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, left third maxilliped; B, ventral view of cephalothorax and pleon; B, male pleon; D, sternopleonal cavity showing G1 in situ; E, left G1 (ventral view); F, left G1 (dorsal view); G, terminal segment of left G1 (ventral view); H, terminal segment of left G1 (dorsal view); I, left G2." figureDoi="http://doi.org/10.5281/zenodo.7816169" httpUri="https://zenodo.org/record/7816169/files/figure.png" pageId="2" pageNumber="72">Fig. 3C</figureCitation>
). Sternite 8 not visible when pleon closed. Male press-button pleonal locking mechanism composed of pair of relatively low peg-like tubercles, positioned on posterior quarter of sternite 5 (
<figureCitation box="[930,1014,1757,1781]" captionStart="Fig" captionStartId="5.[136,171,2035,2057]" captionTargetBox="[138,1471,189,2018]" captionTargetId="figure-7@5.[138,1471,189,2018]" captionTargetPageId="5" captionText="Fig. 3. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, left third maxilliped; B, ventral view of cephalothorax and pleon; B, male pleon; D, sternopleonal cavity showing G1 in situ; E, left G1 (ventral view); F, left G1 (dorsal view); G, terminal segment of left G1 (ventral view); H, terminal segment of left G1 (dorsal view); I, left G2." figureDoi="http://doi.org/10.5281/zenodo.7816169" httpUri="https://zenodo.org/record/7816169/files/figure.png" pageId="2" pageNumber="72">Fig. 3D</figureCitation>
).
</paragraph>
<paragraph blockId="2.[811,1453,1821,2069]" pageId="2" pageNumber="72">
Pleon broadly triangular, all somites and telson free; telson triangular, as long as broad, lateral margins gently sinuous; somite 6 transversely trapezoidal, much wider than long, lateral margin gently convex; somites 35 trapezoidal, gradually decreasing in width, increasing in length anteriorly; somite 3 widest; somites 1 and 2 subrectangular, wide, reaching to bases of coxae of fourth ambulatory legs (
<figureCitation captionStart="Fig" captionStartId="5.[136,171,2035,2057]" captionTargetBox="[138,1471,189,2018]" captionTargetId="figure-7@5.[138,1471,189,2018]" captionTargetPageId="5" captionText="Fig. 3. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, left third maxilliped; B, ventral view of cephalothorax and pleon; B, male pleon; D, sternopleonal cavity showing G1 in situ; E, left G1 (ventral view); F, left G1 (dorsal view); G, terminal segment of left G1 (ventral view); H, terminal segment of left G1 (dorsal view); I, left G2." figureDoi="http://doi.org/10.5281/zenodo.7816169" httpUri="https://zenodo.org/record/7816169/files/figure.png" pageId="2" pageNumber="72">Fig. 3B, C</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7816165" ID-Zenodo-Dep="7816165" httpUri="https://zenodo.org/record/7816165/files/figure.png" pageId="3" pageNumber="73" startId="3.[136,171,1700,1722]" targetBox="[136,1474,189,1684]" targetPageId="3">
<paragraph blockId="3.[136,1474,1700,1779]" pageId="3" pageNumber="73">
Fig. 1.
<taxonomicName authority="Ng &amp; Tri, 2023" authorityName="Ng &amp; Tri" authorityYear="2023" box="[209,389,1700,1721]" class="Malacostraca" family="Potamidae" genus="Dacuomon" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="73" phylum="Arthropoda" rank="species" species="quynhi" status="new genus, new species">
<emphasis box="[209,389,1700,1721]" italics="true" pageId="3" pageNumber="73">Dacuomon quynhi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[404,645,1700,1722]" pageId="3" pageNumber="73" rank="species">new genus, new species</taxonomicNameLabel>
, colour in life. A, B, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam; CE, paratype male (23.9 × 20.4 mm) (ZRC 2022.0048), Phu Yên Province, Vietnam; F, type locality, Xuan Lam commune, Phu Yên Province, Vietnam.
</paragraph>
</caption>
<paragraph blockId="3.[135,776,1834,2083]" lastBlockId="3.[835,1476,1834,2050]" pageId="3" pageNumber="73">
G1 relatively slender; subterminal segment widest basally, gradually tapering towards terminal segment, distal quarter neck-like, groove for G2 ventral; subterminal segment distinctly separated from terminal segment by grooves; terminal segment relatively short, subcylindrical, hookshaped, curved outwards, tapering to tip, dorsal flap low, not prominent (
<figureCitation box="[298,420,2026,2051]" captionStart="Fig" captionStartId="5.[136,171,2035,2057]" captionTargetBox="[138,1471,189,2018]" captionTargetId="figure-7@5.[138,1471,189,2018]" captionTargetPageId="5" captionText="Fig. 3. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, left third maxilliped; B, ventral view of cephalothorax and pleon; B, male pleon; D, sternopleonal cavity showing G1 in situ; E, left G1 (ventral view); F, left G1 (dorsal view); G, terminal segment of left G1 (ventral view); H, terminal segment of left G1 (dorsal view); I, left G2." figureDoi="http://doi.org/10.5281/zenodo.7816169" httpUri="https://zenodo.org/record/7816169/files/figure.png" pageId="3" pageNumber="73">Figs. 3EH</figureCitation>
,
<figureCitation box="[430,493,2027,2051]" captionStart="Fig" captionStartId="6.[113,148,1055,1077]" captionTargetBox="[114,1461,189,1038]" captionTargetId="figure-12@6.[90,1510,179,1062]" captionTargetPageId="6" captionText="Fig. 4. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, left G1 (ventral view); B, left G1 (dorsal view); C, terminal segment of left G1 (ventral view); D, terminal segment of left G1 (dorsal view); E, left G2. Scales: A, D = 1.0 mm B, C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7816171" httpUri="https://zenodo.org/record/7816171/files/figure.png" pageId="3" pageNumber="73">4AC</figureCitation>
). G2 elongate, longer than G1; basal segment longer than distal segment (
<figureCitation box="[630,715,2058,2083]" captionStart="Fig" captionStartId="5.[136,171,2035,2057]" captionTargetBox="[138,1471,189,2018]" captionTargetId="figure-7@5.[138,1471,189,2018]" captionTargetPageId="5" captionText="Fig. 3. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, left third maxilliped; B, ventral view of cephalothorax and pleon; B, male pleon; D, sternopleonal cavity showing G1 in situ; E, left G1 (ventral view); F, left G1 (dorsal view); G, terminal segment of left G1 (ventral view); H, terminal segment of left G1 (dorsal view); I, left G2." figureDoi="http://doi.org/10.5281/zenodo.7816169" httpUri="https://zenodo.org/record/7816169/files/figure.png" pageId="3" pageNumber="73">Figs. 3I</figureCitation>
,
<figureCitation box="[726,758,2059,2083]" captionStart="Fig" captionStartId="6.[113,148,1055,1077]" captionTargetBox="[114,1461,189,1038]" captionTargetId="figure-12@6.[90,1510,179,1062]" captionTargetPageId="6" captionText="Fig. 4. Dacuomon quynhi, new genus, new species, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, left G1 (ventral view); B, left G1 (dorsal view); C, terminal segment of left G1 (ventral view); D, terminal segment of left G1 (dorsal view); E, left G2. Scales: A, D = 1.0 mm B, C = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7816171" httpUri="https://zenodo.org/record/7816171/files/figure.png" pageId="3" pageNumber="73">4D</figureCitation>
).
<emphasis bold="true" box="[835,961,1834,1858]" pageId="3" pageNumber="73">Variation.</emphasis>
No female specimens were available. The two smaller male
<typeStatus box="[1058,1171,1867,1891]" pageId="3" pageNumber="73">paratypes</typeStatus>
agree with the
<typeStatus box="[1372,1473,1866,1890]" pageId="3" pageNumber="73">holotype</typeStatus>
in most respects. The postorbital crista in these
<specimenCount pageId="3" pageNumber="73" type="generic">two specimens</specimenCount>
is almost contiguous with the crista connecting to the anterolateral margin, the structures appearing almost confluent. The small fissure separating the cristae is still visible but less pronounced.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7816167" ID-Zenodo-Dep="7816167" httpUri="https://zenodo.org/record/7816167/files/figure.png" pageId="4" pageNumber="74" startId="4.[113,148,2036,2058]" targetBox="[115,1448,189,2020]" targetPageId="4">
<paragraph blockId="4.[113,1451,2036,2115]" pageId="4" pageNumber="74">
Fig. 2.
<taxonomicName authority="Ng &amp; Tri, 2023" authorityName="Ng &amp; Tri" authorityYear="2023" box="[186,363,2036,2057]" class="Malacostraca" family="Potamidae" genus="Dacuomon" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="74" phylum="Arthropoda" rank="species" species="quynhi" status="new genus, new species">
<emphasis box="[186,363,2036,2057]" italics="true" pageId="4" pageNumber="74">Dacuomon quynhi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[376,614,2036,2058]" pageId="4" pageNumber="74" rank="species">new genus, new species</taxonomicNameLabel>
, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of cephalothorax; D, subfrontal view of cephalothorax showing setose third maxillipeds; E, frontal view showing antennules, antennae and epistome; F, outer view of chelae.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7816169" ID-Zenodo-Dep="7816169" httpUri="https://zenodo.org/record/7816169/files/figure.png" pageId="5" pageNumber="75" startId="5.[136,171,2035,2057]" targetBox="[138,1471,189,2018]" targetPageId="5">
<paragraph blockId="5.[136,1474,2035,2115]" pageId="5" pageNumber="75">
Fig. 3.
<taxonomicName authority="Ng &amp; Tri, 2023" authorityName="Ng &amp; Tri" authorityYear="2023" box="[203,380,2035,2057]" class="Malacostraca" family="Potamidae" genus="Dacuomon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="75" phylum="Arthropoda" rank="species" species="quynhi" status="new genus, new species">
<emphasis box="[203,380,2035,2057]" italics="true" pageId="5" pageNumber="75">Dacuomon quynhi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[392,624,2035,2057]" pageId="5" pageNumber="75" rank="species">new genus, new species</taxonomicNameLabel>
, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, left third maxilliped; B, ventral view of cephalothorax and pleon; B, male pleon; D, sternopleonal cavity showing G1 in situ; E, left G1 (ventral view); F, left G1 (dorsal view); G, terminal segment of left G1 (ventral view); H, terminal segment of left G1 (dorsal view); I, left G2.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7816171" ID-Zenodo-Dep="7816171" httpUri="https://zenodo.org/record/7816171/files/figure.png" pageId="6" pageNumber="76" startId="6.[113,148,1055,1077]" targetBox="[114,1461,189,1038]" targetPageId="6">
<paragraph blockId="6.[113,1453,1055,1134]" pageId="6" pageNumber="76">
Fig. 4.
<taxonomicName authority="Ng &amp; Tri, 2023" authorityName="Ng &amp; Tri" authorityYear="2023" box="[186,363,1055,1076]" class="Malacostraca" family="Potamidae" genus="Dacuomon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="76" phylum="Arthropoda" rank="species" species="quynhi" status="new genus, new species">
<emphasis box="[186,363,1055,1076]" italics="true" pageId="6" pageNumber="76">Dacuomon quynhi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[376,614,1055,1077]" pageId="6" pageNumber="76" rank="species">new genus, new species</taxonomicNameLabel>
, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam. A, left G1 (ventral view); B, left G1 (dorsal view); C, terminal segment of left G1 (ventral view); D, terminal segment of left G1 (dorsal view); E, left G2. Scales: A, D = 1.0 mm B, C = 0.5 mm.
</paragraph>
</caption>
<paragraph blockId="6.[112,753,1189,1374]" pageId="6" pageNumber="76">
<emphasis bold="true" box="[113,199,1189,1213]" pageId="6" pageNumber="76">Colour.</emphasis>
In adult specimens, the dorsal surface of the carapace, chelae and ambulatory legs are purplish brown, with the fingers of the cheliped orange (
<figureCitation box="[429,540,1254,1278]" captionStart="Fig" captionStartId="3.[136,171,1700,1722]" captionTargetBox="[136,1474,189,1684]" captionTargetId="figure-7@3.[136,1474,189,1684]" captionTargetPageId="3" captionText="Fig. 1. Dacuomon quynhi, new genus, new species, colour in life. A, B, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam; CE, paratype male (23.9 × 20.4 mm) (ZRC 2022.0048), Phu Yên Province, Vietnam; F, type locality, Xuan Lam commune, Phu Yên Province, Vietnam." figureDoi="http://doi.org/10.5281/zenodo.7816165" httpUri="https://zenodo.org/record/7816165/files/figure.png" pageId="6" pageNumber="76">Fig. 1A, B</figureCitation>
). Smaller specimens have dark grey carapaces with the chelae and ambulatory legs mottled dark and light brown (
<figureCitation box="[451,569,1318,1342]" captionStart="Fig" captionStartId="3.[136,171,1700,1722]" captionTargetBox="[136,1474,189,1684]" captionTargetId="figure-7@3.[136,1474,189,1684]" captionTargetPageId="3" captionText="Fig. 1. Dacuomon quynhi, new genus, new species, colour in life. A, B, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam; CE, paratype male (23.9 × 20.4 mm) (ZRC 2022.0048), Phu Yên Province, Vietnam; F, type locality, Xuan Lam commune, Phu Yên Province, Vietnam." figureDoi="http://doi.org/10.5281/zenodo.7816165" httpUri="https://zenodo.org/record/7816165/files/figure.png" pageId="6" pageNumber="76">Fig. 1C, B</figureCitation>
) and the ventral surfaces are pale yellow to pale pink (
<figureCitation box="[531,615,1350,1374]" captionStart="Fig" captionStartId="3.[136,171,1700,1722]" captionTargetBox="[136,1474,189,1684]" captionTargetId="figure-7@3.[136,1474,189,1684]" captionTargetPageId="3" captionText="Fig. 1. Dacuomon quynhi, new genus, new species, colour in life. A, B, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam; CE, paratype male (23.9 × 20.4 mm) (ZRC 2022.0048), Phu Yên Province, Vietnam; F, type locality, Xuan Lam commune, Phu Yên Province, Vietnam." figureDoi="http://doi.org/10.5281/zenodo.7816165" httpUri="https://zenodo.org/record/7816165/files/figure.png" pageId="6" pageNumber="76">Fig. 1E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="76" type="etymology">
<paragraph blockId="6.[112,753,1414,1598]" pageId="6" pageNumber="76">
<emphasis bold="true" box="[113,243,1414,1438]" pageId="6" pageNumber="76">Etymology.</emphasis>
The species is named after Nguyen Xuan Quynh from the
<collectingRegion box="[218,284,1446,1470]" country="Vietnam" name="Ha Noi" pageId="6" pageNumber="76">Hanoi</collectingRegion>
University of Science, who passed away in early 2022. He was the one who encouraged the first author to start studies on the aquatic fauna of
<collectingCountry box="[630,722,1510,1534]" name="Vietnam" pageId="6" pageNumber="76">Vietnam</collectingCountry>
in the 1990s, and has been a good friend over the years. He will be sorely missed.
</paragraph>
</subSubSection>
<subSubSection box="[113,380,1638,1662]" pageId="6" pageNumber="76" type="discussion">
<paragraph blockId="6.[113,380,1638,1662]" box="[113,380,1638,1662]" pageId="6" pageNumber="76">
<emphasis bold="true" box="[113,224,1638,1662]" pageId="6" pageNumber="76">Remarks.</emphasis>
As for genus.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="76" type="biology_ecology">
<paragraph blockId="6.[113,755,1702,1790]" pageId="6" pageNumber="76">
<emphasis bold="true" box="[113,211,1702,1726]" pageId="6" pageNumber="76">Ecology.</emphasis>
This is a semiterrestrial species, digging burrows under large rocks by the edge of the stream near a planted
<taxonomicName box="[115,189,1766,1789]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="6" pageNumber="76" phylum="Tracheophyta" rank="genus">
<emphasis box="[115,189,1766,1789]" italics="true" pageId="6" pageNumber="76">Acacia</emphasis>
</taxonomicName>
forest (
<figureCitation box="[273,356,1766,1790]" captionStart="Fig" captionStartId="3.[136,171,1700,1722]" captionTargetBox="[136,1474,189,1684]" captionTargetId="figure-7@3.[136,1474,189,1684]" captionTargetPageId="3" captionText="Fig. 1. Dacuomon quynhi, new genus, new species, colour in life. A, B, holotype male (27.7 × 23.8 mm) (ZRC 2022.0047), Phu Yên Province, Vietnam; CE, paratype male (23.9 × 20.4 mm) (ZRC 2022.0048), Phu Yên Province, Vietnam; F, type locality, Xuan Lam commune, Phu Yên Province, Vietnam." figureDoi="http://doi.org/10.5281/zenodo.7816165" httpUri="https://zenodo.org/record/7816165/files/figure.png" pageId="6" pageNumber="76">Fig. 1F</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>