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239 lines
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<document ID-DOI="10.1093/zoolinnean/zlab112" ID-GBIF-Dataset="eb68c3ad-c51a-4828-8a1f-314ed0ba6454" ID-ISSN="0024-4082" ID-Zenodo-Dep="7196247" ID-ZooBank="1C9A93CC-F5BE-427B-95B4-B2B9A1F51B46" checkinTime="1665470485720" checkinUser="plazi" docAuthor="Damaška, Albert František, Konstantinov, Alexander & Fikáček, Martin" docDate="2022" docId="7220879B5C5F770DFE8E3D19EA6348B2" docLanguage="en" docName="zlab112.pdf" docOrigin="Zoological Journal of the Linnean Society 196" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-.journal_article" docStyleVersion="7" docTitle="Benedictus SCHERER 1969" docType="treatment" docVersion="7" lastPageNumber="656" masterDocId="8E19FFE35C567707FFBC3951E8624848" masterDocTitle="Multiple origins of moss-inhabiting flea beetles (Coleoptera: Chrysomelidae): molecular phylogeny, overview of genera and a new genus from Africa" masterLastPageNumber="676" masterPageNumber="647" pageNumber="656" updateTime="1665702766640" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Multiple origins of moss-inhabiting flea beetles (Coleoptera: Chrysomelidae): molecular phylogeny, overview of genera and a new genus from Africa</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Damaška, Albert František</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Konstantinov, Alexander</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fikáček, Martin</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zoological Journal of the Linnean Society</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-02-07</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>196</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>647</mods:start>
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<mods:end>676</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1093/zoolinnean/zlab112</mods:identifier>
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<mods:identifier type="GBIF-Dataset">eb68c3ad-c51a-4828-8a1f-314ed0ba6454</mods:identifier>
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<mods:identifier type="ISSN">0024-4082</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7196247</mods:identifier>
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<mods:identifier type="ZooBank">1C9A93CC-F5BE-427B-95B4-B2B9A1F51B46</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7184353" ID-GBIF-Taxon="202593533" ID-Zenodo-Dep="7184353" LSID="urn:lsid:plazi:treatment:7220879B5C5F770DFE8E3D19EA6348B2" httpUri="http://treatment.plazi.org/id/7220879B5C5F770DFE8E3D19EA6348B2" lastPageId="10" lastPageNumber="656" pageId="9" pageNumber="656">
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<subSubSection box="[306,635,1095,1120]" pageId="9" pageNumber="656" type="nomenclature">
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<paragraph blockId="9.[306,635,1095,1120]" box="[306,635,1095,1120]" pageId="9" pageNumber="656">
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<heading box="[306,635,1095,1120]" centered="true" fontSize="9" level="2" pageId="9" pageNumber="656" reason="2">
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<taxonomicName authority="SCHERER, 1969" authorityName="SCHERER" authorityYear="1969" box="[306,635,1095,1120]" class="Insecta" family="Chrysomelidae" genus="Benedictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="656" phylum="Arthropoda" rank="genus">
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<emphasis box="[306,452,1096,1119]" italics="true" pageId="9" pageNumber="656">BENEDICTUS</emphasis>
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SCHERER, 1969
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="10" lastPageNumber="657" pageId="9" pageNumber="656" type="description">
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<paragraph blockId="9.[428,513,1134,1159]" box="[428,513,1134,1159]" pageId="9" pageNumber="656">
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(
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<figureCitation box="[437,505,1134,1159]" captionStart="Figure 4" captionStartId="8.[145,223,1817,1839]" captionTargetBox="[147,1425,822,1776]" captionTargetId="figure-229@8.[145,1428,805,1777]" captionTargetPageId="8" captionText="Figure 4. Moss-inhabiting flea beetle genera distributed in Asia. A A, Paraminotella nigrita; B, Benedictus shivalayanicus; C, Paraminota lauribina; D, Ivalia korakundah; E, Cangshanaltica nigra; F, Baoshanaltica minuta; G, Phaelota viridipennis; H, Clavicornaltica dali." figureDoi="http://doi.org/10.5281/zenodo.7196259" httpUri="https://zenodo.org/record/7196259/files/figure.png" pageId="9" pageNumber="656">FIG. 4</figureCitation>
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)
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</paragraph>
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<paragraph blockId="9.[163,750,1174,1196]" box="[163,750,1174,1196]" pageId="9" pageNumber="656">
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<emphasis box="[163,307,1175,1196]" italics="true" pageId="9" pageNumber="656">
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<typeStatus box="[163,216,1175,1196]" pageId="9" pageNumber="656">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName authority="Scherer, 1969" authorityName="Scherer" authorityYear="1969" box="[326,746,1174,1196]" class="Insecta" family="Chrysomelidae" genus="Benedictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="656" phylum="Arthropoda" rank="species" species="elisabethae">
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<emphasis box="[326,582,1174,1195]" italics="true" pageId="9" pageNumber="656">Benedictus elisabethae</emphasis>
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Scherer, 1969
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</taxonomicName>
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.
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</paragraph>
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<paragraph blockId="9.[163,779,1245,1298]" pageId="9" pageNumber="656">
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<emphasis box="[163,426,1245,1267]" italics="true" pageId="9" pageNumber="656">Synonymy: Himalalta</emphasis>
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Medvedev, 1990 (synonymized by
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<bibRefCitation author="Sprecher-Uebersax E & Konstantinov AS & Prathapan KD & Doberl M" box="[197,544,1276,1298]" pageId="9" pageNumber="656" pagination="475 - 514" refId="ref22634" refString="Sprecher-Uebersax E, Konstantinov AS, Prathapan KD, Doberl M. 2009. Revision of the genus Benedictus Scherer (Insecta: Coleoptera: Chrysomelidae: Galerucinae). In: Biodiversitat und Naturausstattung im Himalaya, Band III. Erfurt: Verein der Freunde & Forderer des Naturkundemuseums Erfurt, 475 - 514." type="book chapter" year="2009">
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Sprecher-Uebersax
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<emphasis box="[421,469,1276,1298]" italics="true" pageId="9" pageNumber="656">et al</emphasis>
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., 2009
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</bibRefCitation>
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).
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</paragraph>
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<paragraph blockId="9.[163,779,1344,1549]" pageId="9" pageNumber="656">
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<emphasis box="[163,436,1344,1365]" italics="true" pageId="9" pageNumber="656">Phylogenetic position:</emphasis>
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The genus belongs to the
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<taxonomicName authorityName="Jacoby" authorityYear="1885" box="[163,270,1374,1395]" class="Insecta" family="Chrysomelidae" genus="Manobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="656" phylum="Arthropoda" rank="genus">
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<emphasis box="[163,270,1374,1395]" italics="true" pageId="9" pageNumber="656">Manobia</emphasis>
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</taxonomicName>
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generic group, together with the winged and leaf-surface-living genera
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<taxonomicName authority="Jacoby, 1885" authorityName="Jacoby" authorityYear="1885" class="Insecta" family="Chrysomelidae" genus="Aphthonoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="656" phylum="Arthropoda" rank="genus">
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<emphasis box="[597,779,1405,1427]" italics="true" pageId="9" pageNumber="656">Aphthonoides</emphasis>
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Jacoby, 1885
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</taxonomicName>
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,
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<taxonomicName authority="Jacoby, 1885" authorityName="Jacoby" authorityYear="1885" box="[326,586,1436,1458]" class="Insecta" family="Chrysomelidae" genus="Manobia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="656" phylum="Arthropoda" rank="genus">
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<emphasis box="[326,430,1436,1457]" italics="true" pageId="9" pageNumber="656">Manobia</emphasis>
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Jacoby, 1885
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</taxonomicName>
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and
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<taxonomicName authority="Chevrolat, 1836" authorityName="Chevrolat" authorityYear="1836" class="Insecta" family="Chrysomelidae" genus="Phyllotreta" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="656" phylum="Arthropoda" rank="genus">
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<emphasis box="[648,779,1436,1457]" italics="true" pageId="9" pageNumber="656">Phyllotreta</emphasis>
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Chevrolat, 1836
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</taxonomicName>
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. The sister-group of
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<taxonomicName authorityName="SCHERER" authorityYear="1969" box="[583,708,1466,1487]" class="Insecta" family="Chrysomelidae" genus="Benedictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="656" phylum="Arthropoda" rank="genus">
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<emphasis box="[583,708,1466,1487]" italics="true" pageId="9" pageNumber="656">Benedictus</emphasis>
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</taxonomicName>
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is the morphologically similar new African moss-inhabiting genus
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<taxonomicName authorityName="Damaška & Konstantinov & Fikáček" authorityYear="2022" box="[237,392,1528,1549]" class="Insecta" family="Chrysomelidae" genus="Benedictoides" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="656" phylum="Arthropoda" rank="genus">
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<emphasis box="[237,392,1528,1549]" italics="true" pageId="9" pageNumber="656">Benedictoides</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph blockId="9.[163,779,1590,1888]" lastBlockId="9.[827,1442,197,372]" pageId="9" pageNumber="656">
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<emphasis box="[163,500,1590,1612]" italics="true" pageId="9" pageNumber="656">Diversity and distribution:</emphasis>
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Twenty-eight species of
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<taxonomicName authorityName="SCHERER" authorityYear="1969" box="[195,329,1621,1642]" class="Insecta" family="Chrysomelidae" genus="Benedictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="656" phylum="Arthropoda" rank="genus">
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<emphasis box="[195,329,1621,1642]" italics="true" pageId="9" pageNumber="656">Benedictus</emphasis>
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</taxonomicName>
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are described. The genus is mainly distributed in the Oriental region, where it clearly shows a ‘Himalayan’ pattern of distribution, with the centre of diversity in the Himalayas and adjacent lower mountain ranges in
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<collectingCountry box="[510,625,1744,1766]" name="Thailand" pageId="9" pageNumber="656">Thailand</collectingCountry>
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and
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<collectingCountry box="[695,773,1744,1766]" name="China" pageId="9" pageNumber="656">China</collectingCountry>
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. It also reaches the
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<collectingCountry box="[415,559,1774,1796]" name="Philippines" pageId="9" pageNumber="656">Philippines</collectingCountry>
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(Luzon). Species of
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<taxonomicName authorityName="SCHERER" authorityYear="1969" box="[197,335,1805,1826]" class="Insecta" family="Chrysomelidae" genus="Benedictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="656" phylum="Arthropoda" rank="genus">
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<emphasis box="[197,335,1805,1826]" italics="true" pageId="9" pageNumber="656">Benedictus</emphasis>
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</taxonomicName>
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are not yet known from any high mountain areas of Sundaland (like Mt. Kinabalu on Borneo or the mountain ranges of Sumatra), but a single species is known from southern
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<collectingCountry box="[1325,1389,197,219]" name="India" pageId="9" pageNumber="656">India</collectingCountry>
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and one from New
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<collectingCountry box="[999,1088,228,250]" name="Guinea-Bissau" pageId="9" pageNumber="656">Guinea</collectingCountry>
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. However, the Papuan species (
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<taxonomicName authority="Sprecher-Uebersax et al., 2009" authorityName="Sprecher-Uebersax" authorityYear="2009" box="[836,1401,258,280]" class="Insecta" family="Chrysomelidae" genus="Benedictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="656" phylum="Arthropoda" rank="species" species="andersoni">
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<emphasis box="[836,996,258,280]" italics="true" pageId="9" pageNumber="656">B. andersoni</emphasis>
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<bibRefCitation author="Sprecher-Uebersax E & Konstantinov AS & Prathapan KD & Doberl M" box="[1007,1401,258,280]" pageId="9" pageNumber="656" pagination="475 - 514" refId="ref22634" refString="Sprecher-Uebersax E, Konstantinov AS, Prathapan KD, Doberl M. 2009. Revision of the genus Benedictus Scherer (Insecta: Coleoptera: Chrysomelidae: Galerucinae). In: Biodiversitat und Naturausstattung im Himalaya, Band III. Erfurt: Verein der Freunde & Forderer des Naturkundemuseums Erfurt, 475 - 514." type="book chapter" year="2009">
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Sprecher-Uebersax
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<emphasis box="[1258,1323,258,280]" italics="true" pageId="9" pageNumber="656">et al.</emphasis>
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, 2009
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</bibRefCitation>
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</taxonomicName>
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) is highly divergent from other
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<taxonomicName box="[1158,1376,289,311]" pageId="9" pageNumber="656">
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<emphasis box="[1158,1285,289,310]" italics="true" pageId="9" pageNumber="656">Benedictus</emphasis>
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species
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</taxonomicName>
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in its morphology and its generic status shall be, therefore, considered carefully.
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</paragraph>
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<paragraph blockId="9.[827,1442,421,474]" pageId="9" pageNumber="656">
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<emphasis box="[827,954,422,443]" italics="true" pageId="9" pageNumber="656">Revisions:</emphasis>
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The genus was revised by SprecherUebersax
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<emphasis box="[941,990,452,474]" italics="true" pageId="9" pageNumber="656">et al</emphasis>
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. (2009).
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</paragraph>
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<paragraph blockId="9.[827,1444,514,1272]" pageId="9" pageNumber="656">
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<emphasis box="[827,1223,514,535]" italics="true" pageId="9" pageNumber="656">Morphological characteristics:</emphasis>
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The beetles are generally tiny (
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<quantity box="[1026,1128,545,566]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="9" pageNumber="656" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">1–3 mm</quantity>
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) with relatively distinct morphological diagnosis (
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<bibRefCitation author="Sprecher-Uebersax E & Konstantinov AS & Prathapan KD & Doberl M" pageId="9" pageNumber="656" pagination="475 - 514" refId="ref22634" refString="Sprecher-Uebersax E, Konstantinov AS, Prathapan KD, Doberl M. 2009. Revision of the genus Benedictus Scherer (Insecta: Coleoptera: Chrysomelidae: Galerucinae). In: Biodiversitat und Naturausstattung im Himalaya, Band III. Erfurt: Verein der Freunde & Forderer des Naturkundemuseums Erfurt, 475 - 514." type="book chapter" year="2009">
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Sprecher-Uebersax
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<emphasis box="[1376,1436,576,598]" italics="true" pageId="9" pageNumber="656">et al.</emphasis>
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, 2009
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</bibRefCitation>
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). Body elongate to oval, colour usually brown, in some species light brown with dark colour pattern on elytra. Head with distinct supracallinal sulci and frontal calli, antennae usually long, reaching almost the midlength of elytra. Pronotum massive, convex, in some species longer than broad or subquadrate, with distinct anterior pronotal edges. Pronotal disc bearing, usually, an antebasal transverse impression with a stripe of distinct punctures, sometimes, impression shallow or even absent. Some species have short, basal, longitudinal, pronotal impressions. Procoxal cavities open posteriorly. Elytra usually convex and (in most species) strongly punctate; punctures usually forming 11 longitudinal striae. Wings absent. Humeral calli usually absent or feeble, sometimes surrounded with a row of punctures anteriorly. Metatibiae straight, bearing an apical spur. Abdomen forming a flat process reaching metacoxae. Aedeagus usually long and slender, spermatheca in most species with a long and strongly curved duct, sometimes with many coils. Vaginal palpi parallel, with apices close together.
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</paragraph>
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<paragraph blockId="9.[827,1443,1313,1550]" pageId="9" pageNumber="656">
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<emphasis box="[827,924,1313,1334]" italics="true" pageId="9" pageNumber="656">Ecology:</emphasis>
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Many species of
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<taxonomicName authorityName="SCHERER" authorityYear="1969" box="[1138,1267,1313,1334]" class="Insecta" family="Chrysomelidae" genus="Benedictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="656" phylum="Arthropoda" rank="genus">
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<emphasis box="[1138,1267,1313,1334]" italics="true" pageId="9" pageNumber="656">Benedictus</emphasis>
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</taxonomicName>
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were found by sifting moss or leaf litter at high altitudes, sometimes even around
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<quantity box="[975,1058,1375,1396]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="656" unit="m" value="4000.0">4000 m</quantity>
|
|||
|
|
a.s.l. Thus, members of the genus are believed to be moss-inhabiting (
|
|||
|
|
<bibRefCitation author="Sprecher-Uebersax E & Konstantinov AS & Prathapan KD & Doberl M" pageId="9" pageNumber="656" pagination="475 - 514" refId="ref22634" refString="Sprecher-Uebersax E, Konstantinov AS, Prathapan KD, Doberl M. 2009. Revision of the genus Benedictus Scherer (Insecta: Coleoptera: Chrysomelidae: Galerucinae). In: Biodiversitat und Naturausstattung im Himalaya, Band III. Erfurt: Verein der Freunde & Forderer des Naturkundemuseums Erfurt, 475 - 514." type="book chapter" year="2009">
|
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|
|
Sprecher-Uebersax
|
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|
|
<emphasis box="[827,886,1436,1457]" italics="true" pageId="9" pageNumber="656">et al.</emphasis>
|
|||
|
|
, 2009
|
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|
|
</bibRefCitation>
|
|||
|
|
). Interestingly, one species was recently also found in mid-elevation mountain habitats of
|
|||
|
|
<collectingCountry box="[827,958,1498,1519]" name="Hong Kong" pageId="9" pageNumber="656">Hong Kong</collectingCountry>
|
|||
|
|
, where it was collected using pitfall traps (
|
|||
|
|
<bibRefCitation author="Damaska AF & Aston P" box="[835,1110,1526,1550]" pageId="9" pageNumber="656" pagination="151 - 161" refId="ref19905" refString="Damaska AF, Aston P. 2019. Leaf litter and moss-inhabiting flea beetles of Hong Kong (Coleoptera: Chrysomelidae: Alticini). Acta Entomologica Musei Nationalis Pragae 59: 151 - 161." type="journal article" year="2019">DamaŠka & Aston, 2019</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph blockId="9.[827,1443,1590,1888]" lastBlockId="10.[145,761,197,250]" lastPageId="10" lastPageNumber="657" pageId="9" pageNumber="656">
|
|||
|
|
<emphasis box="[827,945,1590,1611]" italics="true" pageId="9" pageNumber="656">Remarks:</emphasis>
|
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|
|
The genus resembles the Himalayan and Chinese genera
|
|||
|
|
<taxonomicName authority="Scherer, 1989" authorityName="Scherer" authorityYear="1989" box="[1079,1388,1621,1643]" class="Insecta" family="Chrysomelidae" genus="Loeblaltica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="656" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis box="[1079,1213,1621,1642]" italics="true" pageId="9" pageNumber="656">Loeblaltica</emphasis>
|
|||
|
|
Scherer, 1989
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName authority="Chen, 1933" authorityName="Chen" authorityYear="1933" box="[827,1098,1652,1674]" class="Insecta" family="Chrysomelidae" genus="Microcrepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="656" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis box="[827,960,1652,1673]" italics="true" pageId="9" pageNumber="656">Microcrepis</emphasis>
|
|||
|
|
Chen, 1933
|
|||
|
|
</taxonomicName>
|
|||
|
|
, which are flightless and leaflitter-inhabiting. The genus
|
|||
|
|
<taxonomicName authorityName="Chen" authorityYear="1933" box="[1160,1296,1683,1704]" class="Insecta" family="Chrysomelidae" genus="Microcrepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="656" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis box="[1160,1296,1683,1704]" italics="true" pageId="9" pageNumber="656">Microcrepis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
differs from
|
|||
|
|
<taxonomicName authorityName="SCHERER" authorityYear="1969" box="[827,951,1713,1734]" class="Insecta" family="Chrysomelidae" genus="Benedictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="656" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis box="[827,951,1713,1734]" italics="true" pageId="9" pageNumber="656">Benedictus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
especially in having closed procoxal cavities, compared to the open ones in
|
|||
|
|
<taxonomicName authorityName="SCHERER" authorityYear="1969" box="[1174,1300,1744,1765]" class="Insecta" family="Chrysomelidae" genus="Benedictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="656" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis box="[1174,1300,1744,1765]" italics="true" pageId="9" pageNumber="656">Benedictus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
<taxonomicName authorityName="Scherer" authorityYear="1989" box="[1314,1443,1744,1765]" class="Insecta" family="Chrysomelidae" genus="Loeblaltica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="656" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis box="[1314,1443,1744,1765]" italics="true" pageId="9" pageNumber="656">Loeblaltica</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
can be distinguished from
|
|||
|
|
<taxonomicName authorityName="SCHERER" authorityYear="1969" box="[1127,1251,1774,1795]" class="Insecta" family="Chrysomelidae" genus="Benedictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="656" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis box="[1127,1251,1774,1795]" italics="true" pageId="9" pageNumber="656">Benedictus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by having round metatibia in cross-section, not having a channel dorsally and by having triangular frons (SprecherUebersax
|
|||
|
|
<emphasis box="[947,1007,1866,1888]" italics="true" pageId="9" pageNumber="656">et al.</emphasis>
|
|||
|
|
, 2009).
|
|||
|
|
<taxonomicName authorityName="SCHERER" authorityYear="1969" box="[1103,1232,1866,1887]" class="Insecta" family="Chrysomelidae" genus="Benedictus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="656" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis box="[1103,1232,1866,1887]" italics="true" pageId="9" pageNumber="656">Benedictus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
can be separated from the similar African genus
|
|||
|
|
<taxonomicName authorityName="Damaška & Konstantinov & Fikáček" authorityYear="2022" box="[518,678,197,218]" class="Insecta" family="Chrysomelidae" genus="Benedictoides" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="657" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis box="[518,678,197,218]" italics="true" pageId="10" pageNumber="657">Benedictoides</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, based on the diagnosis provided below.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|