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<document id="F50A8A6319415835E4E52F1067C91434" ID-CLB-Dataset="56018" ID-DOI="10.11646/zootaxa.5125.5.2" ID-GBIF-Dataset="6a677d3b-0f91-4f11-8cf2-64f4ad2a47d2" ID-ISSN="1175-5326" ID-Zenodo-Dep="6457259" ID-ZooBank="39EBA798-29B1-4450-9085-A4FB6D1DB331" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1649834539484" checkinUser="plazi" docAuthor="Makarchenko, Eugenyi A., Semenchenko, Alexander A. &amp; Palatov, Dmitry M." docDate="2022" docId="721787E39454FFEAF5C9668FFBFA6CD8" docLanguage="en" docName="zootaxa.5125.5.2.pdf" docOrigin="Zootaxa 5125 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Diamesa moubayedi Makarchenko et Semenchenko 2022, sp. nov." docType="treatment" docUuid="ECE88876-308D-4C72-9120-F8DECF3CD8D2" docUuidSource="ZooBank" docVersion="7" lastPageNumber="504" masterDocId="8E2EFF9B9440FFFFF55E6618FFBE6F7D" masterDocTitle="Taxonomy of Diamesa steinboecki group (Diptera: Chironomidae: Diamesinae) with description and DNA barcoding of new species. I. Subgroups steinboecki and longipes" masterLastPageNumber="512" masterPageNumber="483" pageNumber="503" updateTime="1699328882303" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="96F441F6D8CB30AEE8D3D57023D64114">Taxonomy of Diamesa steinboecki group (Diptera: Chironomidae: Diamesinae) with description and DNA barcoding of new species. I. Subgroups steinboecki and longipes</mods:title>
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<mods:role id="6ADD7ECBB84D825D66B49B98CC51A05C">
<mods:roleTerm id="5AF7C4592FF67D0A9D83429152FEDB3B">Author</mods:roleTerm>
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<mods:namePart id="B18AB5764AF38D7C586F2E69586614D8">Makarchenko, Eugenyi A.</mods:namePart>
<mods:affiliation id="7CBF7CE76ED65BD93AC72AB88D617E24">Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, 100 let Vladivostoku 159, 690022 Vladivostok, Russia</mods:affiliation>
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</mods:role>
<mods:namePart id="164941FF8F3A43D17FE7DAD1AA5B6287">Semenchenko, Alexander A.</mods:namePart>
<mods:affiliation id="083A9F1C1C30E7CAFC7E1902017BCE89">Laboratory of Ecology and Evolutionary Biology of Aquatic Organisms, Far Eastern Federal University, Suhanova St. 8, 690950 Vladivostok, Russia</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="8989CA1B40DA0A9F3C502BDBB49D400E">Palatov, Dmitry M.</mods:namePart>
<mods:nameIdentifier id="2AE7156A08354AE55023A4BD9296719E" type="ORCID">0000-0002-8826-9316</mods:nameIdentifier>
<mods:affiliation id="C61D8D7C91906BE2A5907F628E471199">A. N. Severtsov Institute of Ecology and Evolution of the Russian Academy of Sciences, Leninskij prosp. 33, 119071 Moscow, Russia triops @ yandex. ru; https: // orcid. org / 0000 - 0002 - 8826 - 9316</mods:affiliation>
<mods:nameIdentifier id="FF4838B4DE985D205A9630243661A42B" type="email">triops@yandex.ru</mods:nameIdentifier>
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<mods:title id="BCC6886BE20B63C5B3D38C117D8A42D0">Zootaxa</mods:title>
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<mods:part id="61E4C51012DB63FD34942DFF0DD7B95F">
<mods:date id="EB75B5941400CFFFDAF8081E42094DFA">2022</mods:date>
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<mods:number id="2386DBDC038880DB351F883B5989CAE5">2022-04-13</mods:number>
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<mods:number id="60444D5C73F79DC05D237592CF2BEF18">5125</mods:number>
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<treatment id="721787E39454FFEAF5C9668FFBFA6CD8" ID-DOI="http://doi.org/10.5281/zenodo.6458142" ID-GBIF-Taxon="194591606" ID-Zenodo-Dep="6458142" LSID="urn:lsid:zoobank.org:act:ECE88876-308D-4C72-9120-F8DECF3CD8D2" httpUri="http://treatment.plazi.org/id/721787E39454FFEAF5C9668FFBFA6CD8" lastPageId="21" lastPageNumber="504" pageId="20" pageNumber="503">
<subSubSection id="B2A4657E9454FFEBF5C9668FFBC06FA8" pageId="20" pageNumber="503" type="nomenclature">
<paragraph id="FA0136F59454FFEBF5C9668FFC3E6FCF" blockId="20.[151,1150,151,250]" box="[151,896,151,178]" pageId="20" pageNumber="503">
<heading id="A14981999454FFEBF5C9668FFC3E6FCF" bold="true" box="[151,896,151,178]" fontSize="11" level="1" pageId="20" pageNumber="503" reason="1">
<emphasis id="C8CAEAE79454FFEBF5C9668FFC3E6FCF" bold="true" box="[151,896,151,178]" pageId="20" pageNumber="503">
<taxonomicName id="3DBE4D769454FFEBF5C9668FFCA86FCC" ID-CoL="9J2PM" authority="Makarchenko et Semenchenko" authorityName="Makarchenko et Semenchenko" authorityYear="2022" box="[151,790,151,178]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="503" phylum="Arthropoda" rank="species" species="moubayedi" status="sp. nov.">
<emphasis id="C8CAEAE79454FFEBF5C9668FFE326FCF" bold="true" box="[151,396,151,178]" italics="true" pageId="20" pageNumber="503">Diamesa moubayedi</emphasis>
Makarchenko
<emphasis id="C8CAEAE79454FFEBF7156680FDDE6FCF" bold="true" box="[587,608,152,178]" italics="true" pageId="20" pageNumber="503">et</emphasis>
Semenchenko
</taxonomicName>
,
<taxonomicNameLabel id="D3F9579C9454FFEBF67C6680FC3E6FCF" box="[802,896,152,178]" pageId="20" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="FA0136F59454FFEBF5C966A3FBC06FA8" blockId="20.[151,1150,151,250]" box="[151,1150,187,213]" pageId="20" pageNumber="503">
http://zoobank.org/NomenclaturalActs/
<uuid id="8E180C209454FFEBF70D66A3FBC06FA8" box="[595,1150,187,213]" pageId="20" pageNumber="503">ECE88876–308D–4C72–9120–F8DECF3CD8D2</uuid>
</paragraph>
</subSubSection>
<subSubSection id="B2A4657E9454FFEBF5C966F8FDEA6F87" box="[151,596,223,250]" pageId="20" pageNumber="503" type="description">
<paragraph id="FA0136F59454FFEBF5C966F8FDEA6F87" blockId="20.[151,1150,151,250]" box="[151,596,223,250]" pageId="20" pageNumber="503">
(
<figureCitation id="62852A709454FFEBF5C166F8FEC56F87" box="[159,379,224,250]" captionStart="FIGURES 37–44" captionStartId="14.[151,264,1880,1904]" captionTargetBox="[151,1435,349,1857]" captionTargetId="figure-94@14.[151,1435,345,1857]" captionTargetPageId="14" captionText="FIGURES 37–44. Males of Diamesa zagrosica sp. nov. (37, 41), D. sakartvella Kownacki et Kownacka (38, 42) and D. moubayedi sp. nov. (39–40, 43–44). 37, head; 38, 41, hypopygium in dorsal view; 40, hypopygium in dorsal view without tergite IX; 42, wing; 43, tergite IX; 44, anal point in lateral view." figureDoi="http://doi.org/10.5281/zenodo.6457273" httpUri="https://zenodo.org/record/6457273/files/figure.png" pageId="20" pageNumber="503">Figs. 39–40, 43–44</figureCitation>
,
<figureCitation id="62852A709454FFEBF4D966C7FD9D6F87" box="[391,547,223,250]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="20" pageNumber="503">46–47, 52, 55</figureCitation>
,
<figureCitation id="62852A709454FFEBF77166C7FDF26F87" box="[559,588,223,250]" captionStart="FIGURES 75–78" captionStartId="24.[151,264,1761,1785]" captionTargetBox="[151,1435,653,1737]" captionTargetId="figure-139@24.[151,1435,653,1737]" captionTargetPageId="24" captionText="FIGURES 75–78. Type localities of Diamesa moubayedi sp. nov. (75), Diamesa zagrosica sp. nov. (76), D. steinboecki Goetghebuer (77) and D. praecipua Saether et Willassen (78). 75, glacial karstic spring, Sannyn Mount, alt. 2000 m a. sea l., Central Lebanon (photo by J. Moubayed); 76, Dare Daei River, Zagros Mountains, alt. 2244 m a. sea l., Lorestan Province, Iran (photo by D.M. Palatov); 77, Muttbach Stream, Valais Canton, Swiss Alps (photo by Jean–Louis Lods); 78, Rishi Ganga River, right tributary of the Alaknanda River near town of Badrinath, alt. 3128 m a. s. l., Himalayas, India (photo by D.M. Palatov)." figureDoi="http://doi.org/10.5281/zenodo.6457283" httpUri="https://zenodo.org/record/6457283/files/figure.png" pageId="20" pageNumber="503">75</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B2A4657E9454FFEBF5C9673DFC096E41" box="[151,951,292,316]" pageId="20" pageNumber="503" type="reference_group">
<paragraph id="FA0136F59454FFEBF5C9673DFC096E41" blockId="20.[151,951,292,316]" box="[151,951,292,316]" pageId="20" pageNumber="503">
<treatmentCitationGroup id="DAAE11DB9454FFEBF5C9673DFC096E41" box="[151,951,292,316]" pageId="20" pageNumber="503">
<taxonomicName id="3DBE4D769454FFEBF5C9673DFDD86E41" ID-CoL="359GS" authority="Kownacki et Kownacka" authorityName="Kownacki et Kownacka (Moubayed &amp; Laville" authorityYear="1983" box="[151,614,292,316]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="503" phylum="Arthropoda" rank="species" species="sakartvella">
<emphasis id="C8CAEAE79454FFEBF5C9673DFED76E41" box="[151,361,292,316]" italics="true" pageId="20" pageNumber="503">Diamesa sakartvella</emphasis>
Kownacki et Kownacka
</taxonomicName>
;
<treatmentCitation id="7B1F10E49454FFEBF731673CFC0D6E41" author="Moubayed, Z. &amp; Laville, H." box="[623,947,292,316]" page="222" pageId="20" pageNumber="503" year="1983">
<bibRefCitation id="9E2F4B049454FFEBF731673CFC0D6E41" author="Moubayed, Z. &amp; Laville, H." box="[623,947,292,316]" pageId="20" pageNumber="503" pagination="219 - 228" refId="ref19493" refString="Moubayed, Z. &amp; Laville, H. (1983) Les Chironomides (Diptera) du Liban. I. Premier inventaire faunistique. Annales de Limnologie, 19 (3), 219 - 228. https: // doi. org / 10.1051 / limn / 1983026" type="journal article" year="1983">Moubayed &amp; Laville 1983: 222</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B2A4657E9454FFEBF5C96772FCDD6EB1" pageId="20" pageNumber="503" type="materials_examined">
<paragraph id="FA0136F59454FFEBF5C96772FCDD6EB1" blockId="20.[151,1437,362,1252]" pageId="20" pageNumber="503">
<emphasis id="C8CAEAE79454FFEBF5C96772FEFB6EF9" bold="true" box="[151,325,362,388]" pageId="20" pageNumber="503">Type material.</emphasis>
<materialsCitation id="4AD63CA89454FFEBF4156772FEA96EB1" ID-GBIF-Occurrence="3751071321" collectingDate="2005-08-05" collectorName="J. Moubayed." country="Lebanon" elevation="2000" latitude="33.9025" location="Mount Sannine" longLatPrecision="5" longitude="35.78389" municipality="River" pageId="20" pageNumber="503" specimenCount="1" stateProvince="Beyrouth" typeStatus="holotype">
<typeStatus id="250588579454FFEBF4156772FE086EF9" box="[331,438,362,388]" pageId="20" pageNumber="503" type="holotype">Holotype</typeStatus>
, adult male,
<collectingCountry id="82A976659454FFEBF71B6772FD716EF9" box="[581,719,362,388]" name="Lebanon" pageId="20" pageNumber="503">LEBANON</collectingCountry>
: glacial springs and small cold stream located in the upper basin of the
<collectingRegion id="387AF8179454FFEBF5816796FE9B6ED5" box="[223,293,398,424]" country="Lebanon" name="Beyrouth" pageId="20" pageNumber="503">Beirut</collectingRegion>
<collectingMunicipality id="1A65AC8F9454FFEBF4736796FED36ED5" box="[301,365,398,424]" pageId="20" pageNumber="503">River</collectingMunicipality>
,
<location id="FF61602E9454FFEBF4266797FD996ED5" LSID="urn:lsid:plazi:treatment:721787E39454FFEAF5C9668FFBFA6CD8:FF61602E9454FFEBF4266797FD996ED5" box="[376,551,398,425]" country="Lebanon" latitude="33.9025" longLatPrecision="5" longitude="35.78389" municipality="River" name="Mount Sannine" pageId="20" pageNumber="503" stateProvince="Beyrouth">Mount Sannine</location>
,
<geoCoordinate id="9F8A50329454FFEBF76D6797FD126ED4" box="[563,684,398,425]" degrees="33.9025" direction="north" orientation="latitude" pageId="20" pageNumber="503" precision="5" value="33.9025">N 33.9025</geoCoordinate>
,
<geoCoordinate id="9F8A50329454FFEBF7E66797FCF46ED4" box="[696,842,398,425]" degrees="35.783889" direction="east" orientation="longitude" pageId="20" pageNumber="503" precision="1" value="35.78389">E 35.783889</geoCoordinate>
,
<date id="8E0010359454FFEBF60B6796FC626ED4" box="[853,988,398,425]" pageId="20" pageNumber="503" value="2005-08-05">
<collectingDate id="9E44E9DD9454FFEBF60B6796FC626ED4" box="[853,988,398,425]" pageId="20" pageNumber="503" value="2005-08-05">5.VIII.2005</collectingDate>
</date>
, altitude
<elevation id="7193D1C69454FFEBF11D6796FAF06ED5" box="[1091,1358,398,425]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="503" unit="m" value="2000.0">
<quantity id="3D469B109454FFEBF11D6796FB276ED4" box="[1091,1177,398,425]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="503" unit="m" value="2000.0">2000 m</quantity>
above sea level
</elevation>
, leg.
<collectorName id="574B53239454FFEBF0D46797FEA96EB1" pageId="20" pageNumber="503">J. Moubayed.</collectorName>
</materialsCitation>
<materialsCitation id="4AD63CA89454FFEBF44367ABFCDE6EB1" ID-GBIF-Occurrence="3751071319" box="[285,864,434,461]" collectingDate="2005-08-05" collectorName="J. Moubayed." country="Lebanon" elevation="2000" latitude="33.9025" location="Mount Sannine" longLatPrecision="5" longitude="35.78389" municipality="River" pageId="20" pageNumber="503" specimenCount="6" specimenCount-adult="6" stateProvince="Beyrouth" typeStatus="paratype">
<typeStatus id="250588579454FFEBF44367ABFE306EB0" box="[285,398,435,461]" pageId="20" pageNumber="503" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="ECB8FD7C9454FFEBF4C767AAFE586EB1" box="[409,486,434,460]" pageId="20" pageNumber="503" type="adult">6 adult</specimenCount>
males, the same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B2A4657E9454FFEBF59967CEFE206D69" pageId="20" pageNumber="503" type="etymology">
<paragraph id="FA0136F59454FFEBF59967CEFE206D69" blockId="20.[151,1437,362,1252]" pageId="20" pageNumber="503">
<emphasis id="C8CAEAE79454FFEBF59967CEFE1B6E8D" bold="true" box="[199,421,470,496]" pageId="20" pageNumber="503">Derivatio nominis.</emphasis>
The species is named in honor of the French chironomid taxonomist Joel Moubayed who collected
<typeStatus id="250588579454FFEBF45D67E3FE8D6D68" box="[259,307,507,533]" pageId="20" pageNumber="503">type</typeStatus>
material.
</paragraph>
</subSubSection>
<subSubSection id="B2A4657E9454FFEAF5996406FE526E60" lastPageId="21" lastPageNumber="504" pageId="20" pageNumber="503" type="description">
<paragraph id="FA0136F59454FFEBF5996406FEEF6D45" blockId="20.[151,1437,362,1252]" box="[199,337,542,568]" pageId="20" pageNumber="503">
<emphasis id="C8CAEAE79454FFEBF5996406FEEF6D45" bold="true" box="[199,337,542,568]" pageId="20" pageNumber="503">Description</emphasis>
</paragraph>
<paragraph id="FA0136F59454FFEBF599645AFA256D21" blockId="20.[151,1437,362,1252]" box="[199,1435,578,605]" pageId="20" pageNumber="503">
<emphasis id="C8CAEAE79454FFEBF599645AFEF86D20" bold="true" box="[199,326,578,605]" italics="true" pageId="20" pageNumber="503">Adult male</emphasis>
(n = 5, except when otherwise stated). Total length
<quantity id="3D469B109454FFEBF6D5645AFC416D20" box="[907,1023,578,605]" metricMagnitude="0" metricUnit="m" metricValue="3.9" metricValueMax="4.1" metricValueMin="3.7" pageId="20" pageNumber="503" unit="m" value="3.9" valueMax="4.1" valueMin="3.7">3.7–4.1 m</quantity>
. Total length/wing length 1.13–1.23.
</paragraph>
<paragraph id="FA0136F59454FFEBF599647EFDB16DD9" blockId="20.[151,1437,362,1252]" pageId="20" pageNumber="503">Coloration. Brown to dark brown. Head, thorax and abdomen brown to dark brown. Legs brown to dark brown. Wings grey, with brownish veins.</paragraph>
<paragraph id="FA0136F59454FFEBF59964B6FE3B6CB9" blockId="20.[151,1437,362,1252]" pageId="20" pageNumber="503">
Head. Eyes hairy, reniform. Temporal setae including 6–8 frontals, 7–9 orbitals, 5–8 verticals and 4–5 postorbitals. Frontal tubercles 28 μm height. Clypeus with 5–11 setae. Antenna with 8 flagellomeres and reduced plume of setae (
<figureCitation id="62852A709454FFEBF58164EEFE8D6C6D" box="[223,307,758,784]" captionStart="FIGURES 37–44" captionStartId="14.[151,264,1880,1904]" captionTargetBox="[151,1435,349,1857]" captionTargetId="figure-94@14.[151,1435,345,1857]" captionTargetPageId="14" captionText="FIGURES 37–44. Males of Diamesa zagrosica sp. nov. (37, 41), D. sakartvella Kownacki et Kownacka (38, 42) and D. moubayedi sp. nov. (39–40, 43–44). 37, head; 38, 41, hypopygium in dorsal view; 40, hypopygium in dorsal view without tergite IX; 42, wing; 43, tergite IX; 44, anal point in lateral view." figureDoi="http://doi.org/10.5281/zenodo.6457273" httpUri="https://zenodo.org/record/6457273/files/figure.png" pageId="20" pageNumber="503">Fig. 39</figureCitation>
); number and length of these setae on 1–7 flagellomeres respectively (n=2): 3 (32–40 μm), 3 (56–68 μm), 3 (64–68 μm), 0, 2 (60–64 μm) 0, 0; terminal flagellomere with 5 setae, 80–88 μm long in basal part and with 2 subapical setae, 20–24 μm long. Length of 1–8 flagellomeres (μm): 100–104, 44–56, 48, 36–38, 32–44, 30–36, 30–32, 200–204; AR 0.58–0.61. Palpomere length (μm): 40–44, 92–104, 140–164, 116–132, 164–180. Palpomere
<quantity id="3D469B109454FFEBF0D0657AFF136CDD" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="20" pageNumber="503" unit="in" value="3.0">3 in</quantity>
distal part with sensilla capitata with diameter 24 μm. Head width/palpal length 0.90–0.97. Antennal length/palpal length 0.88–0.97.
</paragraph>
<paragraph id="FA0136F59454FFEBF59965D6FCB26B70" blockId="20.[151,1437,362,1252]" pageId="20" pageNumber="503">
Thorax. Antepronotum with 12–14 ventrolateral setae, 52–88 μm long. Dorsocentrals 20, 72–120 μm long; prealars 5, 52–116 μm long. Scutellum with
<emphasis id="C8CAEAE79454FFEBF7D665EBFD1C6B71" box="[648,674,1011,1036]" italics="true" pageId="20" pageNumber="503">ca</emphasis>
20 setae.
</paragraph>
<paragraph id="FA0136F59454FFEBF599620EFE586B04" blockId="20.[151,1437,362,1252]" pageId="20" pageNumber="503">
Wing. Length
<quantity id="3D469B109454FFEBF430620EFDAB6B4C" box="[366,533,1046,1073]" metricMagnitude="-3" metricUnit="m" metricValue="3.26" metricValueMax="3.44" metricValueMin="3.08" pageId="20" pageNumber="503" unit="mm" value="3.26" valueMax="3.44" valueMin="3.08">3.08–3.44 mm</quantity>
, width 0.96–1.0 mm. Anal lobe slightly reduced, angular. Squama with 24–26 setae (in 1–2 rows), 56–112 μm long. R and R
<subScript id="663A34B09454FFEBF7F26255FD0A6B26" attach="left" box="[684,692,1101,1115]" fontSize="6" pageId="20" pageNumber="503">1</subScript>
with 26–27 setae, R
<subScript id="663A34B09454FFEBF6FF6255FC056B26" attach="left" box="[929,955,1101,1115]" fontSize="6" pageId="20" pageNumber="503">4+5</subScript>
with 5–6 (in distal half). Costal extension 76–82 μm. RM/MCu 2.5–3.0.
</paragraph>
<paragraph id="FA0136F59454FFEBF599629BFF416B99" blockId="20.[151,1437,362,1252]" pageId="20" pageNumber="503">
Legs. Spur of front tibia 38–40 µm long. Spurs of mid tibia 40–44 µm and 40–48 µm. Spurs of hind tibia 60–68 µm and 40–44 µm long. Hind tibial comb with 21–25 setae. Length (μm) and proportions of leg segments are as in
<tableCitation id="B73C034E9454FFEBF5C962D2FF456B99" box="[151,251,1226,1252]" captionStart="TABLE 10" captionStartId="20.[151,239,1300,1324]" captionTargetPageId="20" captionText="TABLE 10. Lengths (in µm) and proportions of leg segments of Diamesa moubayedi sp. nov., male (n=5)" pageId="20" pageNumber="503">Table 10</tableCitation>
.
</paragraph>
<caption id="AEC1667D9454FFEBF5C9630CFAB86A51" box="[151,1286,1300,1325]" pageId="20" pageNumber="503" startId="20.[151,239,1300,1324]" targetBox="[151,1367,1340,1725]" targetIsTable="true" targetPageId="20">
<paragraph id="FA0136F59454FFEBF5C9630CFAB86A51" blockId="20.[151,1286,1300,1325]" box="[151,1286,1300,1325]" pageId="20" pageNumber="503">
<emphasis id="C8CAEAE79454FFEBF5C9630CFEA96A51" bold="true" box="[151,279,1300,1324]" pageId="20" pageNumber="503">TABLE 10.</emphasis>
Lengths (in µm) and proportions of leg segments of
<taxonomicName id="3DBE4D769454FFEBF613630DFB996A51" authority="Makarchenko &amp; Semenchenko &amp; Palatov, 2022" authorityName="Makarchenko &amp; Semenchenko &amp; Palatov" authorityYear="2022" box="[845,1063,1300,1324]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="503" phylum="Arthropoda" rank="species" species="moubayedi" status="sp. nov.">
<emphasis id="C8CAEAE79454FFEBF613630DFB996A51" box="[845,1063,1300,1324]" italics="true" pageId="20" pageNumber="503">Diamesa moubayedi</emphasis>
</taxonomicName>
<emphasis id="C8CAEAE79454FFEBF170630CFB3D6A51" bold="true" box="[1070,1155,1300,1324]" pageId="20" pageNumber="503">
<taxonomicNameLabel id="D3F9579C9454FFEBF170630CFB3D6A51" box="[1070,1155,1300,1324]" pageId="20" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male (n=5)
</paragraph>
</caption>
<paragraph id="FA0136F59454FFEBF5B16324FB4269CA" pageId="20" pageNumber="503">
<table id="88BEC45594540000F5C96324FAE969C0" box="[151,1367,1340,1725]" colsContinueFrom="19.[151,1366,733,1117]" gridcols="9" gridrows="9" pageId="20" pageNumber="503">
<tr id="448E34B794540000F5C96324FAE96A26" box="[151,1367,1340,1371]" gridrow="0" pageId="20" pageNumber="503" rowspan-0="1" rowspan-7="1">
<th id="075F5DCB94540000F5B16324FEDF6A26" box="[239,353,1340,1371]" gridcol="1" gridrow="0" pageId="20" pageNumber="503">fe</th>
<th id="075F5DCB94540000F4C26324FDB06A26" box="[412,526,1340,1371]" gridcol="2" gridrow="0" pageId="20" pageNumber="503">ti</th>
<th id="075F5DCB94540000F7176324FD056A26" box="[585,699,1340,1371]" gridcol="3" gridrow="0" pageId="20" pageNumber="503">ta1</th>
<th id="075F5DCB94540000F7A86324FCF16A26" box="[758,847,1340,1371]" gridcol="4" gridrow="0" pageId="20" pageNumber="503">ta2</th>
<th id="075F5DCB94540000F6DD6324FC426A26" box="[899,1020,1340,1371]" gridcol="5" gridrow="0" pageId="20" pageNumber="503">ta3</th>
<th id="075F5DCB94540000F10E6324FBD46A26" box="[1104,1130,1340,1371]" gridcol="6" gridrow="0" pageId="20" pageNumber="503">ta4</th>
<th id="075F5DCB94540000F1A06324FAE96A26" box="[1278,1367,1340,1371]" gridcol="8" gridrow="0" pageId="20" pageNumber="503">ta 5</th>
</tr>
<tr id="448E34B794540000F5C9637CFAE96AFF" box="[151,1367,1380,1410]" gridrow="1" pageId="20" pageNumber="503">
<th id="075F5DCB94540000F5C9637CFF056AFF" box="[151,187,1380,1410]" gridcol="0" gridrow="1" pageId="20" pageNumber="503">P1</th>
<td id="075F5DCB94540000F5B1637CFEDF6AFF" box="[239,353,1380,1410]" gridcol="1" gridrow="1" pageId="20" pageNumber="503">1920–2040</td>
<td id="075F5DCB94540000F4C2637CFDB06AFF" box="[412,526,1380,1410]" gridcol="2" gridrow="1" pageId="20" pageNumber="503">1840–1960</td>
<td id="075F5DCB94540000F717637CFD056AFF" box="[585,699,1380,1410]" gridcol="3" gridrow="1" pageId="20" pageNumber="503">1200–1320</td>
<td id="075F5DCB94540000F7A8637CFCF16AFF" box="[758,847,1380,1410]" gridcol="4" gridrow="1" pageId="20" pageNumber="503">560–680</td>
<td id="075F5DCB94540000F6DD637CFC426AFF" box="[899,1020,1380,1410]" gridcol="5" gridrow="1" pageId="20" pageNumber="503">320–380</td>
<td id="075F5DCB94540000F10E637CFB6C6AFF" box="[1104,1234,1380,1410]" colspan="2" colspanRight="1" gridcol="6" gridrow="1" pageId="20" pageNumber="503">98–131</td>
<td id="075F5DCB94540000F1A0637CFAE96AFF" box="[1278,1367,1380,1410]" gridcol="8" gridrow="1" pageId="20" pageNumber="503">156–164</td>
</tr>
<tr id="448E34B794540000F5C96395FAE96AD6" box="[151,1367,1421,1451]" gridrow="2" pageId="20" pageNumber="503">
<th id="075F5DCB94540000F5C96395FF056AD6" box="[151,187,1421,1451]" gridcol="0" gridrow="2" pageId="20" pageNumber="503">P 2</th>
<td id="075F5DCB94540000F5B16395FEDF6AD6" box="[239,353,1421,1451]" gridcol="1" gridrow="2" pageId="20" pageNumber="503">1720–1960</td>
<td id="075F5DCB94540000F4C26395FDB06AD6" box="[412,526,1421,1451]" gridcol="2" gridrow="2" pageId="20" pageNumber="503">1520–1640</td>
<td id="075F5DCB94540000F7176395FD056AD6" box="[585,699,1421,1451]" gridcol="3" gridrow="2" pageId="20" pageNumber="503">640–780</td>
<td id="075F5DCB94540000F7A86395FCF16AD6" box="[758,847,1421,1451]" gridcol="4" gridrow="2" pageId="20" pageNumber="503">320–400</td>
<td id="075F5DCB94540000F6DD6395FC426AD6" box="[899,1020,1421,1451]" gridcol="5" gridrow="2" pageId="20" pageNumber="503">220–240</td>
<td id="075F5DCB94540000F10E6395FB6C6AD6" box="[1104,1234,1421,1451]" colspan="2" colspanRight="1" gridcol="6" gridrow="2" pageId="20" pageNumber="503">98–131</td>
<td id="075F5DCB94540000F1A06395FAE96AD6" box="[1278,1367,1421,1451]" gridcol="8" gridrow="2" pageId="20" pageNumber="503">139–164</td>
</tr>
<tr id="448E34B794540000F5C963ADFAE96AAE" box="[151,1367,1461,1491]" gridrow="3" pageId="20" pageNumber="503">
<th id="075F5DCB94540000F5C963ADFF056AAE" box="[151,187,1461,1491]" gridcol="0" gridrow="3" pageId="20" pageNumber="503">P3</th>
<td id="075F5DCB94540000F5B163ADFEDF6AAE" box="[239,353,1461,1491]" gridcol="1" gridrow="3" pageId="20" pageNumber="503">1940–2100</td>
<td id="075F5DCB94540000F4C263ADFDB06AAE" box="[412,526,1461,1491]" gridcol="2" gridrow="3" pageId="20" pageNumber="503">1769–1960</td>
<td id="075F5DCB94540000F71763ADFD056AAE" box="[585,699,1461,1491]" gridcol="3" gridrow="3" pageId="20" pageNumber="503">1120–1300</td>
<td id="075F5DCB94540000F7A863ADFCF16AAE" box="[758,847,1461,1491]" gridcol="4" gridrow="3" pageId="20" pageNumber="503">600–660</td>
<td id="075F5DCB94540000F6DD63ADFC426AAE" box="[899,1020,1461,1491]" gridcol="5" gridrow="3" pageId="20" pageNumber="503">340–360</td>
<td id="075F5DCB94540000F10E63ADFB6C6AAE" box="[1104,1234,1461,1491]" colspan="2" colspanRight="1" gridcol="6" gridrow="3" pageId="20" pageNumber="503">123–131</td>
<td id="075F5DCB94540000F1A063ADFAE96AAE" box="[1278,1367,1461,1491]" gridcol="8" gridrow="3" pageId="20" pageNumber="503">164</td>
</tr>
<tr id="448E34B794540000F5C963E6FAE9696B" box="[151,1367,1534,1558]" gridrow="4" pageId="20" pageNumber="503">
<th id="075F5DCB94540000F5C963E6FAE9696B" box="[151,1367,1534,1558]" colspan="9" colspanRight="8" gridcol="0" gridrow="4" pageId="20" pageNumber="503">continued</th>
</tr>
<tr id="448E34B794540000F5C9603FFAE96942" box="[151,1367,1575,1599]" gridrow="5" pageId="20" pageNumber="503" rowspan-0="1" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-8="1">
<td id="075F5DCB94540000F5B1603FFEDF6942" box="[239,353,1575,1599]" gridcol="1" gridrow="5" pageId="20" pageNumber="503">LR</td>
<td id="075F5DCB94540000F717603FFD056942" box="[585,699,1575,1599]" gridcol="3" gridrow="5" pageId="20" pageNumber="503">BV</td>
<td id="075F5DCB94540000F6DD603FFC426942" box="[899,1020,1575,1599]" gridcol="5" gridrow="5" pageId="20" pageNumber="503">SV</td>
<td id="075F5DCB94540000F1EE603FFB6C6942" box="[1200,1234,1575,1599]" gridcol="7" gridrow="5" pageId="20" pageNumber="503">BR</td>
</tr>
<tr id="448E34B794540000F5C96057FAE96910" box="[151,1367,1615,1645]" gridrow="6" pageId="20" pageNumber="503" rowspan-4="1" rowspan-6="1">
<th id="075F5DCB94540000F5C96057FF056910" box="[151,187,1615,1645]" gridcol="0" gridrow="6" pageId="20" pageNumber="503">P 1</th>
<td id="075F5DCB94540000F5B16057FDB06910" box="[239,526,1615,1645]" colspan="2" colspanRight="1" gridcol="1" gridrow="6" pageId="20" pageNumber="503">0.64–0.70</td>
<td id="075F5DCB94540000F7176057FD056910" box="[585,699,1615,1645]" gridcol="3" gridrow="6" pageId="20" pageNumber="503">3.78–4.41</td>
<td id="075F5DCB94540000F6DD6057FC426910" box="[899,1020,1615,1645]" gridcol="5" gridrow="6" pageId="20" pageNumber="503">3.0–3.17</td>
<td id="075F5DCB94540000F1EE6057FAE96910" box="[1200,1367,1615,1645]" colspan="2" colspanRight="1" gridcol="7" gridrow="6" pageId="20" pageNumber="503">1.1–1.2</td>
</tr>
<tr id="448E34B794540000F5C9606FFAE969E8" box="[151,1367,1655,1685]" gridrow="7" pageId="20" pageNumber="503" rowspan-4="1" rowspan-6="1" rowspan-8="1">
<th id="075F5DCB94540000F5C9606FFF0569E8" box="[151,187,1655,1685]" gridcol="0" gridrow="7" pageId="20" pageNumber="503">P2</th>
<td id="075F5DCB94540000F5B1606FFDB069E8" box="[239,526,1655,1685]" colspan="2" colspanRight="1" gridcol="1" gridrow="7" pageId="20" pageNumber="503">0.42–0.48</td>
<td id="075F5DCB94540000F717606FFD0569E8" box="[585,699,1655,1685]" gridcol="3" gridrow="7" pageId="20" pageNumber="503">3.38–4.99</td>
<td id="075F5DCB94540000F6DD606FFC4269E8" box="[899,1020,1655,1685]" gridcol="5" gridrow="7" pageId="20" pageNumber="503">4.62–5.06</td>
<td id="075F5DCB94540000F1EE606FFB6C69E8" box="[1200,1234,1655,1685]" gridcol="7" gridrow="7" pageId="20" pageNumber="503">1.1</td>
</tr>
<tr id="448E34B794540000F5C96087FAE969C0" box="[151,1367,1695,1725]" gridrow="8" pageId="20" pageNumber="503" rowspan-4="1" rowspan-6="1">
<th id="075F5DCB94540000F5C96087FF0569C0" box="[151,187,1695,1725]" gridcol="0" gridrow="8" pageId="20" pageNumber="503">P3</th>
<td id="075F5DCB94540000F5B16087FDB069C0" box="[239,526,1695,1725]" colspan="2" colspanRight="1" gridcol="1" gridrow="8" pageId="20" pageNumber="503">0.64–0.66</td>
<td id="075F5DCB94540000F7176087FD0569C0" box="[585,699,1695,1725]" gridcol="3" gridrow="8" pageId="20" pageNumber="503">3.93–4.14</td>
<td id="075F5DCB94540000F6DD6087FC4269C0" box="[899,1020,1695,1725]" gridcol="5" gridrow="8" pageId="20" pageNumber="503">3.12–3.30</td>
<td id="075F5DCB94540000F1EE6087FAE969C0" box="[1200,1367,1695,1725]" colspan="2" colspanRight="1" gridcol="7" gridrow="8" pageId="20" pageNumber="503">1.1–1.2</td>
</tr>
</table>
</paragraph>
<paragraph id="FA0136F59454FFEAF59960FFFEB16F84" blockId="20.[151,1436,1767,2010]" lastBlockId="21.[151,1437,151,933]" lastPageId="21" lastPageNumber="504" pageId="20" pageNumber="503">
Hypopygium (
<figureCitation id="62852A709454FFEBF43560FFFDA1687C" box="[363,543,1767,1793]" captionStart="FIGURES 37–44" captionStartId="14.[151,264,1880,1904]" captionTargetBox="[151,1435,349,1857]" captionTargetId="figure-94@14.[151,1435,345,1857]" captionTargetPageId="14" captionText="FIGURES 37–44. Males of Diamesa zagrosica sp. nov. (37, 41), D. sakartvella Kownacki et Kownacka (38, 42) and D. moubayedi sp. nov. (39–40, 43–44). 37, head; 38, 41, hypopygium in dorsal view; 40, hypopygium in dorsal view without tergite IX; 42, wing; 43, tergite IX; 44, anal point in lateral view." figureDoi="http://doi.org/10.5281/zenodo.6457273" httpUri="https://zenodo.org/record/6457273/files/figure.png" pageId="20" pageNumber="503">Figs. 40, 43–44</figureCitation>
,
<figureCitation id="62852A709454FFEBF77260FFFD72687F" box="[556,716,1767,1794]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="20" pageNumber="503">46–47, 52, 55</figureCitation>
). Tergite IX densely covered with strong macrotrichia apices of which are directed anteriorly, with 7–15 setae, 32–48 µm long and with hitinized, narrow (12–14 µm) and naked anal point, 148–160 µm long, when viewed from side it is slightly curved in distal part (
<figureCitation id="62852A709454FFEBF1DD6137FAB06834" box="[1155,1294,1839,1865]" captionStart="FIGURES 37–44" captionStartId="14.[151,264,1880,1904]" captionTargetBox="[151,1435,349,1857]" captionTargetId="figure-94@14.[151,1435,345,1857]" captionTargetPageId="14" captionText="FIGURES 37–44. Males of Diamesa zagrosica sp. nov. (37, 41), D. sakartvella Kownacki et Kownacka (38, 42) and D. moubayedi sp. nov. (39–40, 43–44). 37, head; 38, 41, hypopygium in dorsal view; 40, hypopygium in dorsal view without tergite IX; 42, wing; 43, tergite IX; 44, anal point in lateral view." figureDoi="http://doi.org/10.5281/zenodo.6457273" httpUri="https://zenodo.org/record/6457273/files/figure.png" pageId="20" pageNumber="503">Figs. 43–44</figureCitation>
). Laterosternite IX with 8–10 setae, 28–32 µm long. Transverse sternopodeme (TSA) triangular, 104–140 µm high, 148–184 µm wide at the base (
<figureCitation id="62852A709454FFEBF4D5616FFE5E68EF" box="[395,480,1911,1938]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="20" pageNumber="503">Fig. 52</figureCitation>
); TSA height/TSA width 0.80–0.89. Aedeagal lobe 164–176 µm long; phallapodeme sclerotized, 140–156 µm long. Gonocoxite 490–492 µm long; superior volsellae angular, along margin with some setae (
<figureCitation id="62852A709454FFEBF58161A7FE8D68A7" box="[223,307,1983,2010]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="20" pageNumber="503">Fig. 55</figureCitation>
). Gonostylus 200–240 µm long, slightly curved, expanded in the distal half along the outer edge and with rounded–angular projection basally (
<figureCitation id="62852A709455FFEAF728668FFD736FCC" box="[630,717,151,177]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="21" pageNumber="504">Fig. 47</figureCitation>
); in apical part with megaseta in form of wide terminal spine, 12 μm long and tooth the same size, next to it there is two setae approximately of the same length (
<figureCitation id="62852A709455FFEAF05466A3FADE6FAB" box="[1290,1376,187,214]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="21" pageNumber="504">Fig. 46</figureCitation>
). HR 2.14–2.23.
</paragraph>
<paragraph id="FA0136F59455FFEAF599671BFE526E60" blockId="21.[151,1437,151,933]" box="[199,492,258,285]" pageId="21" pageNumber="504">
<emphasis id="C8CAEAE79455FFEAF599671BFEBA6E60" bold="true" box="[199,260,259,285]" italics="true" pageId="21" pageNumber="504">Pupa</emphasis>
and
<emphasis id="C8CAEAE79455FFEAF464671AFECB6E60" bold="true" box="[314,373,258,285]" italics="true" pageId="21" pageNumber="504">larva</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="B2A4657E9455FFEAF599673EFACB6DD4" pageId="21" pageNumber="504" type="diagnosis">
<paragraph id="FA0136F59455FFEAF599673EFACB6DD4" blockId="21.[151,1437,151,933]" pageId="21" pageNumber="504">
<emphasis id="C8CAEAE79455FFEAF599673EFEFC6E3D" bold="true" box="[199,322,294,320]" pageId="21" pageNumber="504">Diagnosis.</emphasis>
The male of
<taxonomicName id="3DBE4D769455FFEAF48D673FFDCE6E3C" authority="Makarchenko &amp; Semenchenko &amp; Palatov, 2022" authorityName="Makarchenko &amp; Semenchenko &amp; Palatov" authorityYear="2022" box="[467,624,295,321]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="504" phylum="Arthropoda" rank="species" species="moubayedi" status="sp. nov.">
<emphasis id="C8CAEAE79455FFEAF48D673FFDCE6E3C" box="[467,624,295,321]" italics="true" pageId="21" pageNumber="504">D. moubayedi</emphasis>
</taxonomicName>
<emphasis id="C8CAEAE79455FFEAF728673FFD716E3C" bold="true" box="[630,719,295,321]" pageId="21" pageNumber="504">
<taxonomicNameLabel id="D3F9579C9455FFEAF728673FFD716E3C" box="[630,719,295,321]" pageId="21" pageNumber="504" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is closely related in the subgroup to
<taxonomicName id="3DBE4D769455FFEAF13B673FFB4D6E3D" authority="Makarchenko &amp; Semenchenko &amp; Palatov, 2022" authorityName="Makarchenko &amp; Semenchenko &amp; Palatov" authorityYear="2022" box="[1125,1267,295,320]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="504" phylum="Arthropoda" rank="species" species="zagrosica" status="sp. nov.">
<emphasis id="C8CAEAE79455FFEAF13B673FFB4D6E3D" box="[1125,1267,295,320]" italics="true" pageId="21" pageNumber="504">D. zagrosica</emphasis>
</taxonomicName>
<emphasis id="C8CAEAE79455FFEAF1A7673FFAF26E3C" bold="true" box="[1273,1356,295,321]" pageId="21" pageNumber="504">
<taxonomicNameLabel id="D3F9579C9455FFEAF1A7673FFAF26E3C" box="[1273,1356,295,321]" pageId="21" pageNumber="504" rank="species">sp. nov</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="3DBE4D769455FFEAF0DF673FFEAD6E18" authorityName="Kownacki et Kownacka (Moubayed &amp; Laville" authorityYear="1983" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="504" phylum="Arthropoda" rank="species" species="sakartvella">
<emphasis id="C8CAEAE79455FFEAF0DF673FFEAD6E18" italics="true" pageId="21" pageNumber="504">D. sakartvella</emphasis>
</taxonomicName>
but this species has longest body (
<quantity id="3D469B109455FFEAF7C36753FC946E18" box="[669,810,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="4.1" metricValueMin="3.7" pageId="21" pageNumber="504" unit="mm" value="3.9" valueMax="4.1" valueMin="3.7">3.7–4.1 mm</quantity>
); transverse sternopodeme is triangular shape (
<figureCitation id="62852A709455FFEAF0656753FA316E18" box="[1339,1423,330,357]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="21" pageNumber="504">Fig. 52</figureCitation>
), of great height (104–140 µm) and not very wide basally (148–184 µm), the ratio of its height to width is 0.80–0.89; superior volsellae angular (
<figureCitation id="62852A709455FFEAF496678BFDA56ED0" box="[456,539,402,429]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="21" pageNumber="504">Fig. 55</figureCitation>
); gonostylus expanded in the distal half along the outer edge and with rounded– angular projection basally (
<figureCitation id="62852A709455FFEAF49267AFFDA76EAF" box="[460,537,439,466]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="21" pageNumber="504">Fig.47</figureCitation>
). Total length of
<taxonomicName id="3DBE4D769455FFEAF78567AFFCD46EAD" authority="Makarchenko &amp; Semenchenko &amp; Palatov, 2022" authorityName="Makarchenko &amp; Semenchenko &amp; Palatov" authorityYear="2022" box="[731,874,439,464]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="504" phylum="Arthropoda" rank="species" species="zagrosica" status="sp. nov.">
<emphasis id="C8CAEAE79455FFEAF78567AFFCD46EAD" box="[731,874,439,464]" italics="true" pageId="21" pageNumber="504">D. zagrosica</emphasis>
</taxonomicName>
<emphasis id="C8CAEAE79455FFEAF62C67AFFC726EAC" bold="true" box="[882,972,439,465]" pageId="21" pageNumber="504">
<taxonomicNameLabel id="D3F9579C9455FFEAF62C67AFFC726EAC" box="[882,972,439,465]" pageId="21" pageNumber="504" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<quantity id="3D469B109455FFEAF68A67AFFBE16EAC" box="[980,1119,439,465]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.8" metricValueMin="2.3" pageId="21" pageNumber="504" unit="mm" value="2.55" valueMax="2.8" valueMin="2.3">2.3–2.8 mm</quantity>
; transverse sternopodeme is regular triangular shape (
<figureCitation id="62852A709455FFEAF4F367C3FD886E88" box="[429,566,474,501]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="21" pageNumber="504">Figs. 53–54</figureCitation>
), of medium height (84–108 µm) and not very wide basally (162–172 µm), the ratio of its height to width is 0.49–0.67; superior volsellae rounded–angular (
<figureCitation id="62852A709455FFEAF6B167E7FBFD6D64" box="[1007,1091,510,537]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="21" pageNumber="504">Fig. 56</figureCitation>
); gonostylus slightly expanded in the distal half along the outer edge and with small rounded projection basally (
<figureCitation id="62852A709455FFEAF164643BFB036D43" box="[1082,1213,547,574]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="21" pageNumber="504">Figs.48–49</figureCitation>
). Total length of
<taxonomicName id="3DBE4D769455FFEAF0DF643BFEAD6D1C" authorityName="Kownacki et Kownacka (Moubayed &amp; Laville" authorityYear="1983" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="504" phylum="Arthropoda" rank="species" species="sakartvella">
<emphasis id="C8CAEAE79455FFEAF0DF643BFEAD6D1C" italics="true" pageId="21" pageNumber="504">D. sakartvella</emphasis>
</taxonomicName>
<quantity id="3D469B109455FFEAF442645FFE196D1C" box="[284,423,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.1" metricValueMin="2.9" pageId="21" pageNumber="504" unit="mm" value="3.0" valueMax="3.1" valueMin="2.9">2.9–3.1 mm</quantity>
;
<emphasis id="C8CAEAE79455FFEAF4ED645FFE056D1D" box="[435,443,583,608]" italics="true" pageId="21" pageNumber="504">t</emphasis>
ransverse sternopodeme is wide triangular (
<figureCitation id="62852A709455FFEAF6F6645FFB8C6D1C" box="[936,1074,582,609]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="21" pageNumber="504">Figs. 50–51</figureCitation>
), of low height (68–88 µm) and very wide basally (172–188 µm), the ratio of its height to width is 0.36–0.50; superior volsellae rounded (
<figureCitation id="62852A709455FFEAF0656473FA316DF8" box="[1339,1423,618,645]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="21" pageNumber="504">Fig. 57</figureCitation>
); gonostylus almost not expanded in the distal half along the outer edge and without projection basally (
<figureCitation id="62852A709455FFEAF04F6496FAD76DD4" box="[1297,1385,654,681]" captionStart="FIGURES 45–57" captionStartId="17.[151,264,1903,1927]" captionTargetBox="[156,1426,417,1862]" captionTargetId="figure-125@17.[151,1436,417,1880]" captionTargetPageId="17" captionText="FIGURES 45–57. Some details of male’s hypopygium of Diamesa sakartvella Kownacki et Kownacka (45, 50–51, 57), D. moubayedi sp. nov. (46–47, 52, 55), D. zagrosica sp. nov. (48–49, 53–54, 56). 45–49, gonostylus; 46, apex of gonostylus; 50–54, transverse sternapodeme; 55–57, superior volsellae." figureDoi="http://doi.org/10.5281/zenodo.6457277" httpUri="https://zenodo.org/record/6457277/files/figure.png" pageId="21" pageNumber="504">Figs.45</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B2A4657E9455FFEAF59964ABFDFD6C6B" pageId="21" pageNumber="504" type="biology_ecology">
<paragraph id="FA0136F59455FFEAF59964ABFDFD6C6B" blockId="21.[151,1437,151,933]" pageId="21" pageNumber="504">
<emphasis id="C8CAEAE79455FFEAF59964ABFE926DB0" bold="true" box="[199,300,691,717]" pageId="21" pageNumber="504">Ecology.</emphasis>
Adult males were collected in karstic springs and streams, located at an altitude of
<quantity id="3D469B109455FFEAF1AD64ABFAF36DB0" box="[1267,1357,691,717]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="21" pageNumber="504" unit="m" value="2000.0">2000 m</quantity>
. Water calcareous, conductivity 320–360 μS/cm, water temperature 8–10 °C during summer, about 4–8 °C in winter and spring (pers. comm. by J. Moubayed).
</paragraph>
</subSubSection>
<subSubSection id="B2A4657E9455FFEAF5996506FBFA6CD8" pageId="21" pageNumber="504" type="distribution">
<paragraph id="FA0136F59455FFEAF5996506FE856C20" blockId="21.[151,1437,151,933]" pageId="21" pageNumber="504">
<emphasis id="C8CAEAE79455FFEAF5996506FEDF6C45" bold="true" box="[199,353,798,824]" pageId="21" pageNumber="504">Distribution.</emphasis>
Known only from the
<typeStatus id="250588579455FFEAF7056507FD356C44" box="[603,651,799,825]" pageId="21" pageNumber="504">type</typeStatus>
locality—karstic springs and streams of Sannyn Mount (Central
<collectingCountry id="82A976659455FFEAF0056507FF7D6C20" name="Lebanon" pageId="21" pageNumber="504">Lebanon</collectingCountry>
) (
<figureCitation id="62852A709455FFEAF584655BFE906C20" box="[218,302,834,861]" captionStart="FIGURES 75–78" captionStartId="24.[151,264,1761,1785]" captionTargetBox="[151,1435,653,1737]" captionTargetId="figure-139@24.[151,1435,653,1737]" captionTargetPageId="24" captionText="FIGURES 75–78. Type localities of Diamesa moubayedi sp. nov. (75), Diamesa zagrosica sp. nov. (76), D. steinboecki Goetghebuer (77) and D. praecipua Saether et Willassen (78). 75, glacial karstic spring, Sannyn Mount, alt. 2000 m a. sea l., Central Lebanon (photo by J. Moubayed); 76, Dare Daei River, Zagros Mountains, alt. 2244 m a. sea l., Lorestan Province, Iran (photo by D.M. Palatov); 77, Muttbach Stream, Valais Canton, Swiss Alps (photo by Jean–Louis Lods); 78, Rishi Ganga River, right tributary of the Alaknanda River near town of Badrinath, alt. 3128 m a. s. l., Himalayas, India (photo by D.M. Palatov)." figureDoi="http://doi.org/10.5281/zenodo.6457283" httpUri="https://zenodo.org/record/6457283/files/figure.png" pageId="21" pageNumber="504">Fig. 75</figureCitation>
).
</paragraph>
<paragraph id="FA0136F59455FFEAF599657FFBFA6CD8" blockId="21.[151,1437,151,933]" pageId="21" pageNumber="504">
Previously, this species, after the identification of B. Serra-Tosio (pers. comm. by J. Moubayed), was recorded for
<collectingCountry id="82A976659455FFEAF5E16593FE9F6CD8" box="[191,289,907,933]" name="Lebanon" pageId="21" pageNumber="504">Lebanon</collectingCountry>
as
<taxonomicName id="3DBE4D769455FFEAF4186593FB816CD8" authority="Kownacki et Kownacka (Moubayed &amp; Laville 1983)" authorityName="Kownacki et Kownacka (Moubayed &amp; Laville" authorityYear="1983" box="[326,1087,907,933]" class="Insecta" family="Chironomidae" genus="Diamesa" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="504" phylum="Arthropoda" rank="species" species="sakartvella">
<emphasis id="C8CAEAE79455FFEAF4186593FE5A6CD8" box="[326,484,907,933]" italics="true" pageId="21" pageNumber="504">D. sakartvella</emphasis>
Kownacki et Kownacka (
<bibRefCitation id="9E2F4B049455FFEAF6596593FB896CD8" author="Moubayed, Z. &amp; Laville, H." box="[775,1079,907,933]" pageId="21" pageNumber="504" pagination="219 - 228" refId="ref19493" refString="Moubayed, Z. &amp; Laville, H. (1983) Les Chironomides (Diptera) du Liban. I. Premier inventaire faunistique. Annales de Limnologie, 19 (3), 219 - 228. https: // doi. org / 10.1051 / limn / 1983026" type="journal article" year="1983">Moubayed &amp; Laville 1983</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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