treatments-xml/data/72/05/87/720587AC3160D90044B26A2AFD3518CD.xml

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<mods:title id="25FCA52CD99D0D8C86FB819AC0556E22">Revision of Hypoxys Amyot &amp; Serville, 1843 stat. rest. (Heteroptera Pentatomidae)</mods:title>
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<mods:namePart id="511C1335D1B9132766685A13841D7033">Nunes, Benedito Mendes</mods:namePart>
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<mods:namePart id="4ECF3F76BFEE4C90DB66294243CB62E8">Campos, Lourival Dias</mods:namePart>
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<mods:namePart id="B2721DE29FFB8EDD5EA59E0EAE562278">Mendonca, Maria Thayane Da Silva</mods:namePart>
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<mods:namePart id="905170FF2E44BF09686D3387C3483DA4">Cunha, Eduardo Victor De Paiva</mods:namePart>
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<heading id="A15B81D63160D90244B26A2AFD661D60" bold="true" box="[151,762,1859,1886]" fontSize="11" level="1" pageId="29" pageNumber="430" reason="1">
<emphasis id="C8D8EAA83160D90244B26A2AFD661D60" bold="true" box="[151,762,1859,1886]" pageId="29" pageNumber="430">
<taxonomicName id="3DAC4D393160D90244B26A2AFDF01D63" authority="(Breddin, 1904)" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1904" box="[151,620,1859,1886]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="leptacanthus" status="comb. nov.">
<emphasis id="C8D8EAA83160D90244B26A2AFE3E1D60" bold="true" box="[151,418,1859,1886]" italics="true" pageId="29" pageNumber="430">Hypoxys leptacanthus</emphasis>
(Breddin, 1904)
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<taxonomicNameLabel id="D3EB57D33160D90246566A2AFD661D60" box="[627,762,1860,1886]" pageId="29" pageNumber="430" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="FA1336BA3160D90244B26A09FE541DBC" blockId="29.[151,762,1859,1922]" box="[151,456,1895,1922]" pageId="29" pageNumber="430">
(
<figureCitation id="62972A3F3160D90244BA6A09FF671DBC" box="[159,251,1895,1922]" captionStart="FIGURE 15" captionStartId="30.[151,250,1667,1692]" captionTargetBox="[171,1415,201,1606]" captionTargetId="figure@30.[151,1436,181,1641]" captionTargetPageId="30" captionText="FIGURE 15. H. leptacanthus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677899" httpUri="https://zenodo.org/record/3677899/files/figure.png" pageId="29" pageNumber="430">Figs. 15</figureCitation>
A–E; 20 E–F; 22)
</paragraph>
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<subSubSection id="B2B665313160D90244B26ACBFB301DC5" pageId="29" pageNumber="430" type="reference_group">
<paragraph id="FA1336BA3160D90244B26ACBFBD91D83" blockId="29.[151,1437,1956,2043]" box="[151,1093,1956,1981]" pageId="29" pageNumber="430">
<treatmentCitationGroup id="DABC11943160D90244B26ACBFBD91D83" box="[151,1093,1956,1981]" pageId="29" pageNumber="430">
<treatmentCitation id="7B0D10AB3160D90244B26ACBFDAA1D83" author="Breddin, G." box="[151,566,1956,1981]" page="139" pageId="29" pageNumber="430" year="1904">
<taxonomicName id="3DAC4D393160D90244B26ACBFDAA1D83" authority="Breddin, 1904 a: 139" authorityName="Breddin" authorityPageNumber="139" authorityYear="1904" box="[151,566,1956,1981]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="leptacantha">
<emphasis id="C8D8EAA83160D90244B26ACBFEC11D83" box="[151,349,1957,1981]" italics="true" pageId="29" pageNumber="430">Edessa leptacantha</emphasis>
<bibRefCitation id="9E3D4B4B3160D90245466ACAFDAA1D83" author="Breddin, G." box="[355,566,1956,1981]" pageId="29" pageNumber="430" pagination="135 - 153" refId="ref24439" refString="Breddin, G. (1904 a) Beitrage zur Systematik der Rhynchoten. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin, 1904, 135 - 153." type="journal article" year="1904">Breddin, 1904a: 139</bibRefCitation>
</taxonomicName>
</treatmentCitation>
; Bergroth, 1908: 179; 
<treatmentCitation id="7B0D10AB3160D902473F6ACAFC791D83" author="Kirkaldy, G. W." box="[794,997,1956,1981]" page="159" pageId="29" pageNumber="430" year="1909">
<bibRefCitation id="9E3D4B4B3160D902473F6ACAFC791D83" author="Kirkaldy, G. W." box="[794,997,1956,1981]" pageId="29" pageNumber="430" refId="ref25016" refString="Kirkaldy, G. W. (1909) Catalogue of the Hemiptera (Heteroptera). Vol. 1. Cimicidae. Felix L. Dames ed., Berlin, XL + 392 pp." type="book" year="1909">Kirkaldy, 1909: 159</bibRefCitation>
</treatmentCitation>
(catalog)
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</paragraph>
<paragraph id="FA1336BA3160D90244B26AAAFB301DC5" blockId="29.[151,1437,1956,2043]" pageId="29" pageNumber="430">
<materialsCitation id="4AC43CE73160D90244B26AAAFBDC1DC4" ID-GBIF-Occurrence="2570056005" collectionCode="SDEI" country="Peru" location="Amazonas" pageId="29" pageNumber="430" specimenCount="2" typeStatus="lectotype">
<typeStatus id="251788183160D90244B26AAAFF601DE2" box="[151,252,1988,2012]" pageId="29" pageNumber="430" type="lectotype">Lectotype</typeStatus>
male. 
<collectingCountry id="82BB762A3160D90245606AAAFEEA1DE5" box="[325,374,1988,2011]" name="Peru" pageId="29" pageNumber="430">Peru</collectingCountry>
, 
<location id="FF7360613160D90245A46AAAFE701DE2" LSID="urn:lsid:plazi:treatment:720587AC3160D90044B26A2AFD3518CD:FF7360613160D90245A46AAAFE701DE2" box="[385,492,1988,2012]" country="Peru" name="Amazonas" pageId="29" pageNumber="430">Amazonas</location>
(
<collectionCode id="9CBDAE7F3160D90245D96AAAFDA61DE2" box="[508,570,1988,2012]" pageId="29" pageNumber="430">SDEI</collectionCode>
). Designated by 
<bibRefCitation id="9E3D4B4B3160D90246CC6AADFC291DE2" author="Gaedike, H." box="[745,949,1987,2012]" pageId="29" pageNumber="430" pagination="79 - 159" refId="ref24984" refString="Gaedike, H. (1971) Katalog der in Sammlungen des ehemaligen Deutschen Entomologischen Institutes aufbewahrten Typen- V. Heteroptera. Beitrage zur Entomologie, 21, 79 - 159." type="journal article" year="1971">Gaedike (1971: 90)</bibRefCitation>
. Examined. Gaedike mentioned more than one specimen but did not mentioned gender of 
<typeStatus id="251788183160D902465C6A8CFD451DC5" box="[633,729,2018,2043]" pageId="29" pageNumber="430" type="lectotype">lectotype</typeStatus>
. So, we are designating the male as
</materialsCitation>
<typeStatus id="251788183160D90240626A8DFB301DC5" box="[1095,1196,2019,2043]" pageId="29" pageNumber="430" type="lectotype">Lectotype</typeStatus>
</paragraph>
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<caption id="AED366323163D90144B26BEDFD0F1CFE" ID-DOI="http://doi.org/10.5281/zenodo.3677899" ID-Zenodo-Dep="3677899" httpUri="https://zenodo.org/record/3677899/files/figure.png" pageId="30" pageNumber="431" startId="30.[151,250,1667,1692]" targetBox="[171,1415,201,1606]" targetPageId="30">
<paragraph id="FA1336BA3163D90144B26BEDFD0F1CFE" blockId="30.[151,1436,1667,1728]" pageId="30" pageNumber="431">
<emphasis id="C8D8EAA83163D90144B26BEDFE831CA2" bold="true" box="[151,287,1667,1692]" pageId="30" pageNumber="431">FIGURE 15.</emphasis>
<taxonomicName id="3DAC4D393163D90145006BEAFE591CA2" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1904" box="[293,453,1668,1692]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="431" phylum="Arthropoda" rank="species" species="leptacanthus">
<emphasis id="C8D8EAA83163D90145006BEAFE591CA2" box="[293,453,1668,1692]" italics="true" pageId="30" pageNumber="431">H. leptacanthus</emphasis>
</taxonomicName>
. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm).
</paragraph>
</caption>
<subSubSection id="B2B665313163D90144B26A7FFE1F1D6E" pageId="30" pageNumber="431" type="materials_examined">
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<emphasis id="C8D8EAA83163D90144B26A7FFE181D12" bold="true" box="[151,388,1809,1836]" pageId="30" pageNumber="431">
<collectingRegion id="3868F8583163D90144B26A7FFF631D12" box="[151,255,1809,1836]" pageId="30" pageNumber="431">Material</collectingRegion>
examined.
</emphasis>
<materialsCitation id="4AC43CE73163D90145AA6A7CFE1C1D6E" ID-GBIF-Occurrence="2570056022" collectionCode="RMNH" country="Peru" location="Oxapampa" pageId="30" pageNumber="431" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Oxapampa">
<emphasis id="C8D8EAA83163D90145AA6A7CFDE61D12" bold="true" box="[399,634,1809,1836]" pageId="30" pageNumber="431">
<collectingCountry id="82BB762A3163D90145AA6A7CFE411D12" box="[399,477,1810,1836]" name="Peru" pageId="30" pageNumber="431">PERU</collectingCountry>
. 
<collectingRegion id="3868F8583163D90145C86A7FFDEA1D12" box="[493,630,1809,1836]" pageId="30" pageNumber="431">Oxapampa</collectingRegion>
:
</emphasis>
<specimenCount id="ECAAFD333163D90146A06A7DFD061D13" box="[645,666,1811,1837]" pageId="30" pageNumber="431" type="male">♂</specimenCount>
<specimenCount id="ECAAFD333163D90146816A7DFD251D13" box="[676,697,1811,1837]" pageId="30" pageNumber="431" type="female">♀</specimenCount>
<taxonomicName id="3DAC4D393163D90146E16A7CFBC81D12" authority="Breddin, 1904" authorityName="Breddin" authorityYear="1904" box="[708,1108,1810,1836]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="431" phylum="Arthropoda" rank="species" species="leptacantha">
<emphasis id="C8D8EAA83163D90146E16A7CFC3E1D12" box="[708,930,1810,1836]" italics="true" pageId="30" pageNumber="431">Edessa leptacantha</emphasis>
Breddin, 1904
</taxonomicName>
, Comp.w. 
<typeStatus id="251788183163D90140F46A7DFA8B1D13" box="[1233,1303,1811,1837]" pageId="30" pageNumber="431">TYPE</typeStatus>
Fernandes, JAM 1999 (
<collectionCode id="9CBDAE7F3163D90145386A58FEE41D6E" box="[285,376,1846,1872]" country="Netherlands" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="431" type="Museum">RMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B2B665313163D90144E26A34FAD61D82" pageId="30" pageNumber="431" type="description">
<paragraph id="FA1336BA3163D90144E26A34FAD61D82" blockId="30.[151,1437,1809,2017]" pageId="30" pageNumber="431">
<emphasis id="C8D8EAA83163D90144E26A34FEE21D4A" bold="true" box="[199,382,1882,1908]" pageId="30" pageNumber="431">Measurements.</emphasis>
total length: 15.4–17.1; head length: 1.7–2.0; head width: 2.8–3.1; pronotal width: 10.3–11.6; abdominal width: 7.8–9.1; length of antennal segments. I: 0.8–1.0; II: 1.6–1.9; III: 1.0–1.3; IV: 3.6–4.4; V: 4.2–4.7; interocular distance: 1.4–1.6; scutellum length: 7.0–8.0; scutellum width: 5.0–5.8; pronotal length: 3.1–3.6.
</paragraph>
</subSubSection>
<subSubSection id="B2B665313163D90044E26AABFC441BC9" lastPageId="31" lastPageNumber="432" pageId="30" pageNumber="431" type="diagnosis">
<paragraph id="FA1336BA3163D90044E26AABFC441BC9" blockId="30.[151,1437,1809,2017]" lastBlockId="31.[151,1437,153,755]" lastPageId="31" lastPageNumber="432" pageId="30" pageNumber="431">
<emphasis id="C8D8EAA83163D90144E26AABFEDE1DDE" bold="true" box="[199,322,1989,2016]" pageId="30" pageNumber="431">Diagnosis.</emphasis>
Pronotum with small, elongate, lateral black spots including posterolateral angles. Anterolateral margin of pronotum without solid black stripe and not punctured. Posterolateral angles developed, as long as interocular distance. Anterior margin of scutellum yellow (
<figureCitation id="62972A3F3162D900472B6DD3FCEF1AE9" box="[782,883,189,215]" captionStart="FIGURE 20" captionStartId="36.[151,250,1749,1774]" captionTargetBox="[181,1415,196,1699]" captionTargetId="figure@36.[151,1436,181,1723]" captionTargetPageId="36" captionText="FIGURE 20. Dorsal and ventral view. H. brachyacanthus A–B; H. infulatus C–D; H. leptacanthus E–F; H. tragelaphus G–H (Scale = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.3677909" httpUri="https://zenodo.org/record/3677909/files/figure.png" pageId="31" pageNumber="432">Fig. 20E</figureCitation>
). Dorsal surface of abdomen brown. Connexivum green without spots, except distal margin of segment VII, dorsally urotergite 8 with inner black stripes continuing in free part of laterotergites 8 (
<figureCitation id="62972A3F3162D90045D36C6BFDC11B21" box="[502,605,261,287]" captionStart="FIGURE 20" captionStartId="36.[151,250,1749,1774]" captionTargetBox="[181,1415,196,1699]" captionTargetId="figure@36.[151,1436,181,1723]" captionTargetPageId="36" captionText="FIGURE 20. Dorsal and ventral view. H. brachyacanthus A–B; H. infulatus C–D; H. leptacanthus E–F; H. tragelaphus G–H (Scale = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.3677909" httpUri="https://zenodo.org/record/3677909/files/figure.png" pageId="31" pageNumber="432">Fig. 20E</figureCitation>
). Ventrally apex of pseudosutures without rounded dark spots (
<figureCitation id="62972A3F3162D900410F6C6BFA0C1B21" box="[1322,1424,261,287]" captionStart="FIGURE 20" captionStartId="36.[151,250,1749,1774]" captionTargetBox="[181,1415,196,1699]" captionTargetId="figure@36.[151,1436,181,1723]" captionTargetPageId="36" captionText="FIGURE 20. Dorsal and ventral view. H. brachyacanthus A–B; H. infulatus C–D; H. leptacanthus E–F; H. tragelaphus G–H (Scale = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.3677909" httpUri="https://zenodo.org/record/3677909/files/figure.png" pageId="31" pageNumber="432">Fig. 20F</figureCitation>
). Male (
<figureCitation id="62972A3F3162D90044C46C47FEB31B7D" box="[225,303,296,323]" captionStart="FIGURE 15" captionStartId="30.[151,250,1667,1692]" captionTargetBox="[171,1415,201,1606]" captionTargetId="figure@30.[151,1436,181,1641]" captionTargetPageId="30" captionText="FIGURE 15. H. leptacanthus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677899" httpUri="https://zenodo.org/record/3677899/files/figure.png" pageId="31" pageNumber="432">Fig. 15</figureCitation>
A–C). Pygophore with dorsal rim barely excavated; lateral strongly projected posteriorly, surpassing posterolateral angles and expansions of ventral rim (
<figureCitation id="62972A3F3162D90046C36C23FCF51B59" box="[742,873,332,359]" captionStart="FIGURE 15" captionStartId="30.[151,250,1667,1692]" captionTargetBox="[171,1415,201,1606]" captionTargetId="figure@30.[151,1436,181,1641]" captionTargetPageId="30" captionText="FIGURE 15. H. leptacanthus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677899" httpUri="https://zenodo.org/record/3677899/files/figure.png" pageId="31" pageNumber="432">Fig. 15A,B</figureCitation>
). Paramere with anterior lobe strongly developed, almost reaching base of proctiger; posterior lobe strongly developed and acuminated (
<figureCitation id="62972A3F3162D900406B6C1FFB281BB5" box="[1102,1204,368,395]" captionStart="FIGURE 15" captionStartId="30.[151,250,1667,1692]" captionTargetBox="[171,1415,201,1606]" captionTargetId="figure@30.[151,1436,181,1641]" captionTargetPageId="30" captionText="FIGURE 15. H. leptacanthus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677899" httpUri="https://zenodo.org/record/3677899/files/figure.png" pageId="31" pageNumber="432">Fig. 15B</figureCitation>
). Proctiger with tufts of short setae widely separated, but not as much as in 
<taxonomicName id="3DAC4D393162D90046CE6CFBFC3C1B91" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1904" box="[747,928,404,431]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="432" phylum="Arthropoda" rank="species" species="oxyacanthus">
<emphasis id="C8D8EAA83162D90046CE6CFBFC3C1B91" bold="true" box="[747,928,404,431]" italics="true" pageId="31" pageNumber="432">H. oxyacanthus</emphasis>
</taxonomicName>
. Ventral rim with expansions well developed, barely tumid, and rounded (
<figureCitation id="62972A3F3162D90045EB6CD7FDA91BED" box="[462,565,440,467]" captionStart="FIGURE 15" captionStartId="30.[151,250,1667,1692]" captionTargetBox="[171,1415,201,1606]" captionTargetId="figure@30.[151,1436,181,1641]" captionTargetPageId="30" captionText="FIGURE 15. H. leptacanthus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677899" httpUri="https://zenodo.org/record/3677899/files/figure.png" pageId="31" pageNumber="432">Fig. 15C</figureCitation>
). Female (
<figureCitation id="62972A3F3162D900468E6CD7FC8F1BED" box="[683,787,440,467]" captionStart="FIGURE 15" captionStartId="30.[151,250,1667,1692]" captionTargetBox="[171,1415,201,1606]" captionTargetId="figure@30.[151,1436,181,1641]" captionTargetPageId="30" captionText="FIGURE 15. H. leptacanthus. A–C Male, pygophore; A—dorsal view; B—posterior view; C—ventral view. D—Female, genital plates. E—metasternal process (Scale = 1mm)." figureDoi="http://doi.org/10.5281/zenodo.3677899" httpUri="https://zenodo.org/record/3677899/files/figure.png" pageId="31" pageNumber="432">Fig. 15D</figureCitation>
). Gonocoxites 8 inner angle covering base of gonocoxites 9. Laterotergites 9 wide, lightly surpassing sclerite uniting laterotergites 8.
</paragraph>
</subSubSection>
<subSubSection id="B2B665313162D90044E26F6EFF4F18F1" pageId="31" pageNumber="432" type="discussion">
<paragraph id="FA1336BA3162D90044E26F6EFF4F18F1" blockId="31.[151,1437,153,755]" pageId="31" pageNumber="432">
<emphasis id="C8D8EAA83162D90044E26F6EFED31825" bold="true" box="[199,335,512,539]" pageId="31" pageNumber="432">Comments.</emphasis>
<bibRefCitation id="9E3D4B4B3162D900457C6F6FFD881825" author="Breddin, G." box="[345,532,513,539]" pageId="31" pageNumber="432" pagination="135 - 153" refId="ref24439" refString="Breddin, G. (1904 a) Beitrage zur Systematik der Rhynchoten. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin, 1904, 135 - 153." type="journal article" year="1904">Breddin (1904a)</bibRefCitation>
considered 
<taxonomicName id="3DAC4D393162D90046876F6FFCE31825" authorityName="Breddin" authorityYear="1904" box="[674,895,513,539]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="432" phylum="Arthropoda" rank="species" species="leptacantha">
<emphasis id="C8D8EAA83162D90046876F6FFCE31825" box="[674,895,513,539]" italics="true" pageId="31" pageNumber="432">Edessa leptacantha</emphasis>
</taxonomicName>
related to 
<taxonomicName id="3DAC4D393162D90047DA6F6CFB221825" authorityName="Breddin" authorityYear="1904" box="[1023,1214,513,539]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="432" phylum="Arthropoda" rank="species" species="pachyacantha">
<emphasis id="C8D8EAA83162D90047DA6F6CFB8C1825" box="[1023,1040,514,539]" italics="true" pageId="31" pageNumber="432">E</emphasis>
. 
<emphasis id="C8D8EAA83162D90040046F6FFB221825" box="[1057,1214,513,539]" italics="true" pageId="31" pageNumber="432">pachyacantha</emphasis>
</taxonomicName>
(= 
<taxonomicName id="3DAC4D393162D90040C46F6CFA101825" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1903" box="[1249,1420,513,539]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="432" phylum="Arthropoda" rank="species" species="tragelaphus">
<emphasis id="C8D8EAA83162D90040C46F6CFA101825" box="[1249,1420,513,539]" italics="true" pageId="31" pageNumber="432">H. tragelaphus</emphasis>
</taxonomicName>
). These species can be differentiated by humeral angles globose in 
<taxonomicName id="3DAC4D393162D90047516F48FBAC1801" authorityName="Breddin" authorityYear="1904" box="[884,1072,549,575]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="432" phylum="Arthropoda" rank="species" species="pachyacantha">
<emphasis id="C8D8EAA83162D90047516F48FC191801" box="[884,901,550,575]" italics="true" pageId="31" pageNumber="432">E</emphasis>
. 
<emphasis id="C8D8EAA83162D90047B66F4BFBAC1801" box="[915,1072,549,575]" italics="true" pageId="31" pageNumber="432">pachyacantha</emphasis>
</taxonomicName>
(
<figureCitation id="62972A3F3162D900401A6F4BFB341801" box="[1087,1192,549,575]" captionStart="FIGURE 20" captionStartId="36.[151,250,1749,1774]" captionTargetBox="[181,1415,196,1699]" captionTargetId="figure@36.[151,1436,181,1723]" captionTargetPageId="36" captionText="FIGURE 20. Dorsal and ventral view. H. brachyacanthus A–B; H. infulatus C–D; H. leptacanthus E–F; H. tragelaphus G–H (Scale = 5mm)." figureDoi="http://doi.org/10.5281/zenodo.3677909" httpUri="https://zenodo.org/record/3677909/files/figure.png" pageId="31" pageNumber="432">Fig. 20G</figureCitation>
). Other characters are subtle: expansions of ventral rim are more rounded, setae of lateral excavation of proctiger shorter, and paramere has anterior and posterior lobes more developed in 
<taxonomicName id="3DAC4D393162D90046F16F00FC1918B9" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1904" box="[724,901,621,647]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="432" phylum="Arthropoda" rank="species" species="leptacanthus">
<emphasis id="C8D8EAA83162D90046F16F00FC1918B9" box="[724,901,621,647]" italics="true" pageId="31" pageNumber="432">H. leptacanthus</emphasis>
</taxonomicName>
than in 
<taxonomicName id="3DAC4D393162D90047C56F00FB1418B9" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1903" box="[992,1160,621,647]" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="432" phylum="Arthropoda" rank="species" species="tragelaphus">
<emphasis id="C8D8EAA83162D90047C56F00FB1418B9" box="[992,1160,621,647]" italics="true" pageId="31" pageNumber="432">H. tragelaphus</emphasis>
</taxonomicName>
. The female of 
<taxonomicName id="3DAC4D393162D900411C6F00FF721895" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1904" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="432" phylum="Arthropoda" rank="species" species="leptacanthus">
<emphasis id="C8D8EAA83162D900411C6F00FF721895" italics="true" pageId="31" pageNumber="432">H. leptacanthus</emphasis>
</taxonomicName>
has four parallel narrow black stripes dorsally on seventh and eighth segments not found in 
<taxonomicName id="3DAC4D393162D90041386FFCFF5018F1" authorityName="Nunes &amp; Campos &amp; Mendonca &amp; Cunha &amp; Fernandes" authorityYear="2020" baseAuthorityName="Breddin" baseAuthorityYear="1903" class="Insecta" genus="Hypoxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="432" phylum="Arthropoda" rank="species" species="tragelaphus">
<emphasis id="C8D8EAA83162D90041386FFCFF5018F1" italics="true" pageId="31" pageNumber="432">H. tragelaphus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="B2B665313162D90044E26FB6FD3518CD" box="[199,681,728,755]" pageId="31" pageNumber="432" type="distribution">
<paragraph id="FA1336BA3162D90044E26FB6FD3518CD" blockId="31.[151,1437,153,755]" box="[199,681,728,755]" pageId="31" pageNumber="432">
<emphasis id="C8D8EAA83162D90044E26FB6FE5718CD" bold="true" box="[199,459,728,755]" pageId="31" pageNumber="432">
Distribution (
<figureCitation id="62972A3F3162D900454D6FB6FE2218CD" box="[360,446,728,755]" captionStart="FIGURE 23" captionStartId="39.[151,250,1206,1231]" captionTargetBox="[155,1431,192,1174]" captionTargetId="figure@39.[151,1436,181,1181]" captionTargetPageId="39" captionText="FIGURE 23. Distributional map of the species of H. oxyacanthus group." figureDoi="http://doi.org/10.5281/zenodo.3677915" httpUri="https://zenodo.org/record/3677915/files/figure.png" pageId="31" pageNumber="432">Fig. 23</figureCitation>
).
</emphasis>
<collectingCountry id="82BB762A3162D90045F76FB7FD8218CD" box="[466,542,729,755]" name="Peru" pageId="31" pageNumber="432">PERU</collectingCountry>
: Oxapampa.
</paragraph>
</subSubSection>
</treatment>
</document>