<documentid="3F18EFC897D1CCC1632F2BC0CDA46A00"ID-DOI="10.11646/zootaxa.5570.3.7"ID-ISSN="1175-5326"ID-Zenodo-Dep="14745461"ID-ZooBank="41BC11D8-E0D7-40D6-837E-DF48B4A42CA5"IM.bibliography_approvedBy="felipe"IM.illustrations_approvedBy="felipe"IM.materialsCitations_approvedBy="diego"IM.metadata_approvedBy="felipe"IM.taxonomicNames_approvedBy="diego"IM.treatmentCitations_approvedBy="felipe"IM.treatments_approvedBy="diego"checkinTime="1737968622267"checkinUser="plazi"docAuthor="Minkina, Łukasz, Skelley, Paul, Jákl, Stanislav, Král, David & Li, Xiao-Feng"docDate="2025"docId="637487AB235B045F578464ADFE9CFB18"docLanguage="en"docName="zootaxa.5570.3.7.pdf"docOrigin="Zootaxa 5570 (3)"docSource="https://doi.org/10.11646/zootaxa.5570.3.7"docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article"docStyleId="647186512141C8FC8976D5BCC54AEB7D"docStyleName="Zootaxa.2013-.journal_article"docStyleVersion="9"docTitle="Rhyparus malaitaensis Minkina, Skelley, JAkl, KrAl & Li 2025, new species"docType="treatment"docVersion="3"lastPageNumber="572"masterDocId="9F4DFFD3235D04565713643AFFA6FFDC"masterDocTitle="Review of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) in the Solomon Islands with description of two new species"masterLastPageNumber="573"masterPageNumber="563"pageNumber="569"updateTime="1738230843662"updateUser="diego"zenodo-license-document="CLOSED">
<mods:titleid="309DAE1E1E16A8CDA09BA1FDEFA8A0AC">Review of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) in the Solomon Islands with description of two new species</mods:title>
<mods:affiliationid="383D7D2377DC2555B4C9541E255D7D3F">Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 128 00, Praha 2, Czechia.</mods:affiliation>
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. Body large-sized for the genus, distinctly elongate, not distinctly convex, flattened medially; matt; apparently almost glabrous, except head and longitudinal costae on pronotum and eltyra partly clothed with very short and thin, yellowish macrosetae. Dark brownish; antennae, tarsomeres and mouth parts pale brownish.
<figureCitationid="73E62A38235B0450560367CDFEC1FBCD"box="[272,359,1015,1041]"captionStart="FIGURES 4"captionStartId="8.[151,264,1820,1844]"captionTargetBox="[156,1431,601,1792]"captionTargetId="figure-195@8.[151,1435,597,1797]"captionTargetPageId="8"captionText="FIGURES 4A–G. Details of morphology. A, Rhyparus guadalcanalensis, new species, male, holotype, head; B, Rhyparus guadalcanalensis, new species, male, holotype, metatibia; C, Rhyparus helophoroides Fairmaire, 1893, male, metatibia; D, Rhyparus malaitaensis, new species, female, holotype, head; E, Rhyparus malaitaensis, new species, female, holotype, pygidium and last abdominal ventrite; F, Rhyparus argopuroensis Minkina & Jákl, 2024, female, holotype, pygidium and last abdominal ventrite; G, Rhyparus bacanensis Minkina, Anichtchenko, Vasiljeva & Skelley, 2022, female, holotype, pygidium and last abdominal ventrite."figureDoi="http://doi.org/10.5281/zenodo.14745471"httpUri="https://zenodo.org/record/14745471/files/figure.png"pageId="6"pageNumber="569">Fig. 4D</figureCitation>
) transversely sub-hexagonal; surface very weakly shiny, tops of ridges moderately shiny; clypeus trapezoidal in outline, anteriorly truncate, on each side upturned as obtuse, weak tooth, laterally sinuous; genae distinctly more excavate than eyes; clypeal disc distinctly convex, ringed by a deep groove; convexity with a pair of quite distinct, very short, ridges that converge anteriorly, nearly whole surface with distinct, fine punctures bearing small macrosetae. Frons with four distinct, longitudinal ridges with similar structure as ridges on clypeal convexity. Head covered by regularly spaced, dense, moderately large punctures bearing short macrosetae.
</paragraph>
<paragraphid="EB6236BD235B045057D460EAFE53FA39"blockId="6.[151,1437,259,1977]"pageId="6"pageNumber="569">Pronotum very weakly shiny, tops of costae distinctly shiny; with eight distinct costae and seven intercostae, lateral margin with two rounded lobes. Anterior lobe very weakly more prominent laterally and distinctly narrower than median lobe, anterior lobes are the widest part of pronotum. Costae of middle, third and fourth pair entire from posterior to anterior margins; median pair very gently convergent anteriorly, distinctly convergent in the middle of anterior third; second pair of costae distinctly interrupted in anterior half; all costae on each side with very small punctures bearing very small macrosetae. All intercostae anteriorly without additional short costae. Median intercostae with dense punctures, distinctly concentrated medially into a longitudinal line, all intercostae posteriorly with distinct, dense punctation.</paragraph>
<paragraphid="EB6236BD235B045057D461D5FD95F9D5"blockId="6.[151,1437,259,1977]"box="[199,563,1519,1545]"pageId="6"pageNumber="569">Scutellum almost imperceptible.</paragraph>
<paragraphid="EB6236BD235B045057D46229FD4FF8F5"blockId="6.[151,1437,259,1977]"pageId="6"pageNumber="569">Elytra matt, tops of costae and preapical glandular area distinctly shiny. Each elytron with six elevated, distinctly convex costae, and five flat intercostae. Costae on each side with very small punctures bearing very small setae. Intercostae 1–4 with two distinct rows of punctures, medially smooth; basally third intercostae with short costal remnant and additional median row of small sparse punctures extending to apical third of elytra; humeral region of fourth intercosta with dense, irregularly spaced additional punctures; fifth intercosta with one row of punctures. Preapical glandular area relatively small. External caudal bulb distinctly reduced, area between external caudal bulb and sides of elytra not divided; external and mediointernal caudal bulbs not divided; medio-internal caudal bulb shortened, transversely rounded, internally stretched.</paragraph>
<figureCitationid="73E62A38235B04505651636DFE30F8AD"box="[322,406,1879,1905]"captionStart="FIGURES 4"captionStartId="8.[151,264,1820,1844]"captionTargetBox="[156,1431,601,1792]"captionTargetId="figure-195@8.[151,1435,597,1797]"captionTargetPageId="8"captionText="FIGURES 4A–G. Details of morphology. A, Rhyparus guadalcanalensis, new species, male, holotype, head; B, Rhyparus guadalcanalensis, new species, male, holotype, metatibia; C, Rhyparus helophoroides Fairmaire, 1893, male, metatibia; D, Rhyparus malaitaensis, new species, female, holotype, head; E, Rhyparus malaitaensis, new species, female, holotype, pygidium and last abdominal ventrite; F, Rhyparus argopuroensis Minkina & Jákl, 2024, female, holotype, pygidium and last abdominal ventrite; G, Rhyparus bacanensis Minkina, Anichtchenko, Vasiljeva & Skelley, 2022, female, holotype, pygidium and last abdominal ventrite."figureDoi="http://doi.org/10.5281/zenodo.14745471"httpUri="https://zenodo.org/record/14745471/files/figure.png"pageId="6"pageNumber="569">Fig. 4E</figureCitation>
) triangular with acute apex, distinctly bent inwards in lateral view (not perpendicular to body axis), with dense, irregularly spaced punctation, with weak longitudinal carina in the middle and deep excision on each side.
<taxonomicNameid="2CDD4D3E235A0451564A6301FDE6F88F"authorityName="Minkina & Skelley & Jákl & Král & Li"authorityYear="2025"box="[345,576,1851,1875]"class="Insecta"family="Scarabaeidae"genus="Rhyparus"kingdom="Animalia"order="Coleoptera"pageId="7"pageNumber="570"phylum="Arthropoda"rank="species"species="malaitaensis"status="sp. nov.">
<figureCitationid="73E62A38235A04515633639FFED1F863"box="[288,375,1957,1983]"captionStart="FIGURES 3"captionStartId="7.[151,264,1850,1874]"captionTargetBox="[156,1431,186,1825]"captionTargetId="figure-21@7.[151,1435,181,1827]"captionTargetPageId="7"captionText="FIGURES 3A–C. Rhyparus malaitaensis, new species, female, holotype. A, dorsal view; B, ventral view; C, lateral view."figureDoi="http://doi.org/10.5281/zenodo.14745469"httpUri="https://zenodo.org/record/14745469/files/figure.png"pageId="7"pageNumber="570">Fig. 3B</figureCitation>
) moderately shiny. Metaventral plate flattened in the middle; with distinct, narrow, quite deep longitudinal furrow on posterior half; punctation of plate dense, quite regularly spaced, slightly irregular in size; all punctures bearing short macrosetae. Abdominal ventrites weakly shiny, on sides with rows of punctures; anteriorly with an additional punctured furrow. Last abdominal ventrite shortened medially for reception of pygidium, with dense punctures, which are about one and half time larger than on penultimate ventrite; anterior half with dense, very deep punctures (pits) separated by narrow longitudinal ridges; posterior part in the middle with very deep groove, located very slightly below apex of pygidium. Meso- and metafemora with two very indistinct tubercles on posterior border; all femora shiny, with regular, very distinct, rather small, very dense punctation; all punctures bearing small macrosetae.
. Punctation of pronotum can be more or less dense. Sexual dimorphism evident in the shape of meso- and metatibiae, which are weakly sinuate on inner edge and are distinctly inwardly hooked at apex; additionally last abdominal ventrite and pygidium in males are not modified.
<taxonomicNameid="2CDD4D3E2355045E55696326FC33F8E8"authorityName="Minkina & Skelley & Jákl & Král & Li"authorityYear="2025"box="[634,917,1820,1844]"class="Insecta"family="Scarabaeidae"genus="Rhyparus"kingdom="Animalia"order="Coleoptera"pageId="8"pageNumber="571"phylum="Arthropoda"rank="species"species="guadalcanalensis"status="sp. nov.">
<taxonomicNameid="2CDD4D3E2355045E52286326FEEEF884"authorityName="Minkina & Skelley & Jákl & Král & Li"authorityYear="2025"class="Insecta"family="Scarabaeidae"genus="Rhyparus"kingdom="Animalia"order="Coleoptera"pageId="8"pageNumber="571"phylum="Arthropoda"rank="species"species="guadalcanalensis"status="sp. nov.">
<bibRefCitationid="8F4C4B4C2355045E532A637AFB45F885"author="Fairmaire, L."box="[1081,1251,1856,1881]"pageId="8"pageNumber="571"pagination="144 - 146"refId="ref7731"refString="Fairmaire, L. (1893) Description de quatre especes nouvelles du genre Rhyparus, Westw. (Scarabaeidae: Coprini). Note XVII. Notes from the Leyden Museum, 15, 144 - 146."type="journal article"year="1893">Fairmaire, 1893</bibRefCitation>
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, male, metatibia; D,
<taxonomicNameid="2CDD4D3E2355045E57A9635EFE02F8A0"authorityName="Minkina & Skelley & Jákl & Král & Li"authorityYear="2025"box="[186,420,1892,1916]"class="Insecta"family="Scarabaeidae"genus="Rhyparus"kingdom="Animalia"order="Coleoptera"pageId="8"pageNumber="571"phylum="Arthropoda"rank="species"species="malaitaensis"status="sp. nov.">
<taxonomicNameid="2CDD4D3E2355045E5473635EFBEDF8A0"authorityName="Minkina & Skelley & Jákl & Král & Li"authorityYear="2025"box="[864,1099,1892,1916]"class="Insecta"family="Scarabaeidae"genus="Rhyparus"kingdom="Animalia"order="Coleoptera"pageId="8"pageNumber="571"phylum="Arthropoda"rank="species"species="malaitaensis"status="sp. nov.">
, female, holotype, pygidium and last abdominal ventrite; F,
<taxonomicNameid="2CDD4D3E2355045E552E63B2FB87F87D"authority="Minkina & Jakl, 2024"authorityName="Minkina & Jakl"authorityYear="2024"box="[573,1057,1928,1953]"class="Insecta"family="Scarabaeidae"genus="Rhyparus"kingdom="Animalia"order="Coleoptera"pageId="8"pageNumber="571"phylum="Arthropoda"rank="species"species="argopuroensis">
<bibRefCitationid="8F4C4B4C2355045E542E63B2FB87F87D"author="Minkina, L. & JAkl, S."box="[829,1057,1928,1953]"pageId="8"pageNumber="571"pagination="131 - 152"refId="ref7932"refString="Minkina, L. & JAkl, S. (2024) Seven new Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) species. Studies and Reports, Taxonomic Series, 20 (1), 131 - 152."type="journal article"year="2024">Minkina & Jákl, 2024</bibRefCitation>
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, female, holotype, pygidium and last abdominal ventrite; G,
<taxonomicNameid="2CDD4D3E2355045E56946396FBD3F819"authority="Minkina, Anichtchenko, Vasiljeva & Skelley, 2022"authorityName="Minkina, Anichtchenko, Vasiljeva & Skelley"authorityYear="2022"box="[391,1141,1964,1989]"class="Insecta"family="Scarabaeidae"genus="Rhyparus"kingdom="Animalia"order="Coleoptera"pageId="8"pageNumber="571"phylum="Arthropoda"rank="species"species="bacanensis">
<bibRefCitationid="8F4C4B4C2355045E557B6396FBD3F819"author="Minkina, L. & Anichtchenko, A. & Vasiljeva, A. & Skelley, P."box="[616,1141,1964,1989]"pageId="8"pageNumber="571"pagination="81 - 101"refId="ref7869"refString="Minkina, L., Anichtchenko, A., Vasiljeva, A. & Skelley, P. (2022) Four new species of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) with modified pygidium and last abdominal ventrite in females. Baltic Journal of Coleopterology, 22 (1), 81 - 101."type="journal article"year="2022">Minkina, Anichtchenko, Vasiljeva & Skelley, 2022</bibRefCitation>
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, female, holotype, pygidium and last abdominal ventrite.
<taxonomicNameid="2CDD4D3E2354045F56C264ACFD69FF6C"authorityName="Minkina & Skelley & Jákl & Král & Li"authorityYear="2025"box="[465,719,150,176]"class="Insecta"family="Scarabaeidae"genus="Rhyparus"kingdom="Animalia"order="Coleoptera"pageId="9"pageNumber="572"phylum="Arthropoda"rank="species"species="malaitaensis"status="sp. nov.">
belongs to the group of species with females having a modified pygidium and last abdominal ventrite. Due to this modification, females are always easily distinguishable from other
by the relatively large body (for a representatives of the genus), with a body length in nearly all cases more than 6.0 mm, elongate body, relatively weakly developed, rounded lateral lobes of pronotum, and weakly developed caudal bulbs.
<taxonomicNameid="2CDD4D3E2354045F56F66555FD32FE54"authorityName="Minkina & Skelley & Jákl & Král & Li"authorityYear="2025"box="[485,660,366,392]"class="Insecta"family="Scarabaeidae"genus="Rhyparus"kingdom="Animalia"order="Coleoptera"pageId="9"pageNumber="572"phylum="Arthropoda"rank="species"species="malaitaensis"status="new species">
has the apex acute, shaped like a distinct triangle (not truncate). Of all species in the group with a modified female pygidium, only
<taxonomicNameid="2CDD4D3E2354045F54FF65A9FE16FE0C"authority="Minkina, Anichtchenko, Vasiljeva & Skelley 2022"authorityName="Minkina, Anichtchenko, Vasiljeva & Skelley"authorityYear="2022"class="Insecta"family="Scarabaeidae"genus="Rhyparus"kingdom="Animalia"order="Coleoptera"pageId="9"pageNumber="572"phylum="Arthropoda"rank="species"species="bacanensis">
<bibRefCitationid="8F4C4B4C2354045F539E65A8FE16FE0C"author="Minkina, L. & Anichtchenko, A. & Vasiljeva, A. & Skelley, P."pageId="9"pageNumber="572"pagination="81 - 101"refId="ref7869"refString="Minkina, L., Anichtchenko, A., Vasiljeva, A. & Skelley, P. (2022) Four new species of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) with modified pygidium and last abdominal ventrite in females. Baltic Journal of Coleopterology, 22 (1), 81 - 101."type="journal article"year="2022">Minkina, Anichtchenko, Vasiljeva & Skelley 2022</bibRefCitation>
</taxonomicName>
and
<taxonomicNameid="2CDD4D3E2354045F56F7658DFC3EFE0C"authority="Minkina & Jakl, 2024"authorityName="Minkina & Jakl"authorityYear="2024"box="[484,920,438,464]"class="Insecta"family="Scarabaeidae"genus="Rhyparus"kingdom="Animalia"order="Coleoptera"pageId="9"pageNumber="572"phylum="Arthropoda"rank="species"species="argopuroensis">
<bibRefCitationid="8F4C4B4C2354045F55B4658CFC3EFE0C"author="Minkina, L. & JAkl, S."box="[679,920,438,464]"pageId="9"pageNumber="572"pagination="131 - 152"refId="ref7932"refString="Minkina, L. & JAkl, S. (2024) Seven new Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) species. Studies and Reports, Taxonomic Series, 20 (1), 131 - 152."type="journal article"year="2024">Minkina & Jákl, 2024</bibRefCitation>
<bibRefCitationid="8F4C4B4C2354045F578465E0FED6FE28"author="Minkina, L. & Anichtchenko, A. & Vasiljeva, A. & Skelley, P."box="[151,368,474,500]"pageId="9"pageNumber="572"pagination="81 - 101"refId="ref7869"refString="Minkina, L., Anichtchenko, A., Vasiljeva, A. & Skelley, P. (2022) Four new species of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) with modified pygidium and last abdominal ventrite in females. Baltic Journal of Coleopterology, 22 (1), 81 - 101."type="journal article"year="2022">
<bibRefCitationid="8F4C4B4C2354045F556365E0FCF8FE28"author="Minkina, L. & JAkl, S."box="[624,862,474,500]"pageId="9"pageNumber="572"pagination="131 - 152"refId="ref7932"refString="Minkina, L. & JAkl, S. (2024) Seven new Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) species. Studies and Reports, Taxonomic Series, 20 (1), 131 - 152."type="journal article"year="2024">Minkina & Jákl 2024</bibRefCitation>
), but they have different last abdominal ventrites. In
<taxonomicNameid="2CDD4D3E2354045F578465C5FEE4FDC4"authorityName="Minkina & Skelley & Jákl & Král & Li"authorityYear="2025"box="[151,322,510,536]"class="Insecta"family="Scarabaeidae"genus="Rhyparus"kingdom="Animalia"order="Coleoptera"pageId="9"pageNumber="572"phylum="Arthropoda"rank="species"species="malaitaensis"status="new species">
the pits in the anterior furrow are about 1.5–2.5 times longer than wider; in
<taxonomicNameid="2CDD4D3E2354045F521065C5FA3AFDC4"authorityName="Minkina, Anichtchenko, Vasiljeva & Skelley"authorityYear="2022"box="[1283,1436,510,536]"class="Insecta"family="Scarabaeidae"genus="Rhyparus"kingdom="Animalia"order="Coleoptera"pageId="9"pageNumber="572"phylum="Arthropoda"rank="species"species="bacanensis">
, has much smaller pits, similar to large punctures.
<taxonomicNameid="2CDD4D3E2354045F5607667CFDA3FDBC"authorityName="Minkina, Anichtchenko, Vasiljeva & Skelley"authorityYear="2022"box="[276,517,582,608]"class="Insecta"family="Scarabaeidae"genus="Rhyparus"kingdom="Animalia"order="Coleoptera"pageId="9"pageNumber="572"phylum="Arthropoda"rank="species"species="bacanensis">
have apical part of last abdominal ventrite located distinctly below the pygidial apex;
<taxonomicNameid="2CDD4D3E2354045F57CE6651FE2DFD58"authorityName="Minkina & Skelley & Jákl & Král & Li"authorityYear="2025"box="[221,395,618,644]"class="Insecta"family="Scarabaeidae"genus="Rhyparus"kingdom="Animalia"order="Coleoptera"pageId="9"pageNumber="572"phylum="Arthropoda"rank="species"species="malaitaensis"status="new species and r.">
<taxonomicNameid="2CDD4D3E2354045F554E6651FDD3FD58"authorityName="Minkina & Jakl"authorityYear="2024"box="[605,629,619,644]"class="Insecta"family="Scarabaeidae"genus="Rhyparus"kingdom="Animalia"order="Coleoptera"pageId="9"pageNumber="572"phylum="Arthropoda"rank="species"species="argopuroensis">
have apical part of last abdominal ventrite located very slightly below the pygidial apex. Female pygidia of these three species are not easy to distinguish without having specimens of all species for comparison: proportions and prominence (especially median carina of pygidium) are slightly different. Another feature that distinguishes females of
<taxonomicNameid="2CDD4D3E2354045F547166EDFBB6FD2C"authorityName="Minkina & Skelley & Jákl & Král & Li"authorityYear="2025"box="[866,1040,726,752]"class="Insecta"family="Scarabaeidae"genus="Rhyparus"kingdom="Animalia"order="Coleoptera"pageId="9"pageNumber="572"phylum="Arthropoda"rank="species"species="malaitaensis"status="new species and r.">
<taxonomicNameid="2CDD4D3E2354045F53CE66EDFB53FD2C"authorityName="Minkina & Jakl"authorityYear="2024"box="[1245,1269,727,752]"class="Insecta"family="Scarabaeidae"genus="Rhyparus"kingdom="Animalia"order="Coleoptera"pageId="9"pageNumber="572"phylum="Arthropoda"rank="species"species="argopuroensis">
<taxonomicNameid="2CDD4D3E2354045F57C766C1FEC8FCC8"authorityName="Minkina, Anichtchenko, Vasiljeva & Skelley"authorityYear="2022"box="[212,366,762,788]"class="Insecta"family="Scarabaeidae"genus="Rhyparus"kingdom="Animalia"order="Coleoptera"pageId="9"pageNumber="572"phylum="Arthropoda"rank="species"species="bacanensis">
<taxonomicNameid="2CDD4D3E2354045F55FA66C0FC71FCC8"authorityName="Minkina, Anichtchenko, Vasiljeva & Skelley"authorityYear="2022"box="[745,983,762,788]"class="Insecta"family="Scarabaeidae"genus="Rhyparus"kingdom="Animalia"order="Coleoptera"pageId="9"pageNumber="572"phylum="Arthropoda"rank="species"species="bacanensis">
have distinct concavities on each side of the median longitudinal furrow, which are lacking on
<taxonomicNameid="2CDD4D3E2354045F55E16725FC39FCE4"authorityName="Minkina & Skelley & Jákl & Král & Li"authorityYear="2025"box="[754,927,798,824]"class="Insecta"family="Scarabaeidae"genus="Rhyparus"kingdom="Animalia"order="Coleoptera"pageId="9"pageNumber="572"phylum="Arthropoda"rank="species"species="malaitaensis"status="new species and r.">
<taxonomicNameid="2CDD4D3E2354045F537A6725FB27FCE4"authorityName="Minkina & Jakl"authorityYear="2024"box="[1129,1153,799,824]"class="Insecta"family="Scarabaeidae"genus="Rhyparus"kingdom="Animalia"order="Coleoptera"pageId="9"pageNumber="572"phylum="Arthropoda"rank="species"species="argopuroensis">
Males of this group with modified female pygidia are difficult to distinguish. Males of
<taxonomicNameid="2CDD4D3E2354045F53BC6779FAF8FC80"authorityName="Minkina & Skelley & Jákl & Král & Li"authorityYear="2025"box="[1199,1374,834,861]"class="Insecta"family="Scarabaeidae"genus="Rhyparus"kingdom="Animalia"order="Coleoptera"pageId="9"pageNumber="572"phylum="Arthropoda"rank="species"species="malaitaensis"status="new species">
can be recognised by comparing the following combination of characters with those presented in Table 1 and Table 2 of
<bibRefCitationid="8F4C4B4C2354045F567467B0FDC0FC78"author="Minkina, L. & Anichtchenko, A. & Vasiljeva, A. & Skelley, P."box="[359,614,906,932]"pageId="9"pageNumber="572"pagination="81 - 101"refId="ref7869"refString="Minkina, L., Anichtchenko, A., Vasiljeva, A. & Skelley, P. (2022) Four new species of the genus Rhyparus Westwood, 1845 (Coleoptera: Scarabaeidae: Aphodiinae) with modified pygidium and last abdominal ventrite in females. Baltic Journal of Coleopterology, 22 (1), 81 - 101."type="journal article"year="2022">
: all costae shiny, intercostae of pronotum weakly shiny, intercostae of elytra matt; body distinctly elongate, lateral lobes of pronotum weakly developed, sinuation between them not so distinct, anterior lateral lobe of pronotum triangular, very slightly higher than posteriorly; posterior lateral lobe of pronotum widely, regularly rounded; punctation of median intercostae of pronotum dense, regularly spaced, distinctly concentrate in the middle into longitudinal line, punctures coarse here; elytral costae distinctly convex, relatively narrow; some additional fine punctures are observed on all intercostae; third intercostae with three distinct rows of moderately coarse punctures.
