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<document id="FC66D336B01002C31CED3F4DDC67BF53" ID-DOI="10.5281/zenodo.193965" ID-GBIF-Dataset="38d7b0ca-ac3c-4c22-8a28-fa4034b43e03" ID-ISSN="1175-5326" ID-Zenodo-Dep="193965" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460181890217" checkinUser="plazi" docAuthor="Larsson, Karolina &amp; Willems, Wim" docDate="2010" docId="552687E0FFA00D044FFDCE51FAB8FC53" docLanguage="en" docName="zt02396p018.pdf" docOrigin="Zootaxa 2396" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Stenostomum heebuktense Larsson &amp; Willems, 2010, n. sp." docType="treatment" docVersion="9" lastPageNumber="9" masterDocId="A91FFF98FFA60D0C4F6AC953FFEEFFB4" masterDocTitle="Report on freshwater Catenulida (Platyhelminthes) from Sweden with the description of four new species" masterLastPageNumber="18" masterPageNumber="1" pageNumber="7" updateTime="1698236207144" updateUser="plazi">
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<mods:title id="A97A731B549C0BE9693222D6F599FA23">Report on freshwater Catenulida (Platyhelminthes) from Sweden with the description of four new species</mods:title>
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<mods:namePart id="B1A94E85495F73C85CDC6FE202F5931F">Larsson, Karolina</mods:namePart>
</mods:name>
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<mods:namePart id="9C89BB1BC4BBB9E1359FFE4750177662">Willems, Wim</mods:namePart>
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<mods:date id="964B886EE3AF3A2A597AC37C7F886AE0">2010</mods:date>
<mods:detail id="76101CBFB568590DFA2CE731EB2306CC" type="volume">
<mods:number id="103C0A8D460958FBAB856BDC046E655C">2396</mods:number>
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<mods:identifier id="F630E67C3DD9983F39BA1FECADF039C8" type="DOI">10.5281/zenodo.193965</mods:identifier>
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<mods:identifier id="CD3AF2ABC0FAFCF4D8CF132976C75A58" type="ISSN">1175-5326</mods:identifier>
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<treatment id="552687E0FFA00D044FFDCE51FAB8FC53" ID-DOI="http://doi.org/10.5281/zenodo.5670602" ID-GBIF-Taxon="124956604" ID-Zenodo-Dep="5670602" LSID="urn:lsid:plazi:treatment:552687E0FFA00D044FFDCE51FAB8FC53" httpUri="http://treatment.plazi.org/id/552687E0FFA00D044FFDCE51FAB8FC53" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
<subSubSection id="9595657DFFA00D0A4FFDCE51FF01F88B" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="DD3036F6FFA00D0A4FFDCE51FDCFF8A8" blockId="6.[151,545,1794,1855]" box="[151,545,1794,1820]" pageId="6" pageNumber="7">
<heading id="8678819AFFA00D0A4FFDCE51FDCFF8A8" bold="true" box="[151,545,1794,1820]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<emphasis id="EFFBEAE4FFA00D0A4FFDCE51FDCFF8A8" bold="true" box="[151,545,1794,1820]" pageId="6" pageNumber="7">
<taxonomicName id="1A8F4D75FFA00D0A4FFDCE51FE39F8A8" ID-CoL="525KT" box="[151,471,1794,1820]" class="Catenulida" family="Stenostomidae" genus="Stenostomum" kingdom="Animalia" order="Gentianales" pageId="6" pageNumber="7" phylum="Platyhelminthes" rank="species" species="heebuktense" status="sp. nov.">
<emphasis id="EFFBEAE4FFA00D0A4FFDCE51FE39F8A8" bold="true" box="[151,471,1794,1820]" italics="true" pageId="6" pageNumber="7">Stenostomum heebuktense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F4C8579FFFA00D0A4EB4CE51FDCFF8A8" box="[478,545,1794,1820]" pageId="6" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="DD3036F6FFA00D0A4FFDCE76FF01F88B" blockId="6.[151,545,1794,1855]" box="[151,239,1829,1855]" pageId="6" pageNumber="7">
(
<figureCitation id="45B42A73FFA00D0A4FF5CE76FF08F88B" box="[159,230,1829,1855]" captionStart="FIGURE 4" captionStartId="7.[151,255,1646,1670]" captionTargetBox="[151,1435,474,1623]" captionTargetId="figure@7.[151,1436,455,1624]" captionTargetPageId="7" captionText="FIGURE 4. Stenostomum heebuktense n. sp. (A – B: photographs of live specimens; C: habitus of live specimen; D: live individual illustrating pharynx constriction; E: sagittal reconstruction of the pharynx from the left hand side (pharynx appears shorter in comparison with live animals due to constriction during fixation); F: anterior end of sagittal section)." httpUri="https://zenodo.org/record/193968/files/figure.png" pageId="6" pageNumber="7">Fig. 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="9595657DFFA00D0A4FFDCE23FD9DF87B" pageId="6" pageNumber="7" type="description">
<paragraph id="DD3036F6FFA00D0A4FFDCE23FD19F833" blockId="6.[151,759,1904,1927]" box="[151,759,1904,1927]" pageId="6" pageNumber="7">
<emphasis id="EFFBEAE4FFA00D0A4FFDCE23FE70F833" box="[151,414,1904,1927]" italics="true" pageId="6" pageNumber="7">
Synonyms:
<taxonomicName id="1A8F4D75FFA00D0A4E7BCE23FE70F833" box="[273,414,1904,1927]" class="Catenulida" family="Stenostomidae" genus="Stenostomum" kingdom="Animalia" order="Gentianales" pageId="6" pageNumber="7" phylum="Platyhelminthes" rank="genus">Stenostomum</taxonomicName>
</emphasis>
bigmouth (Larsson
<emphasis id="EFFBEAE4FFA00D0A4D1FCE23FD4AF833" box="[629,676,1904,1927]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
. 2008.)
</paragraph>
<paragraph id="DD3036F6FFA00D0A4FFDCEE6FD9DF87B" blockId="6.[151,1435,1973,2036]" box="[151,627,1973,1999]" pageId="6" pageNumber="7">
<emphasis id="EFFBEAE4FFA00D0A4FFDCEE6FEF9F87B" bold="true" box="[151,279,1973,1999]" pageId="6" pageNumber="7">Localities.</emphasis>
Loc. 9 (
<typeStatus id="02348854FFA00D0A4E12CEE6FE42F87B" box="[376,428,1973,1999]" pageId="6" pageNumber="7">type</typeStatus>
locality) and 33.
</paragraph>
</subSubSection>
<subSubSection id="9595657DFFA00D0B4FAECE89FCE6FF48" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="DD3036F6FFA00D0B4FAECE89FCE6FF48" blockId="6.[151,1435,1973,2036]" lastBlockId="7.[151,1437,152,402]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<emphasis id="EFFBEAE4FFA00D0A4FAECE89FED6F840" bold="true" box="[196,312,2010,2036]" pageId="6" pageNumber="7">Material.</emphasis>
Several individuals studied alive. Photograph (fig. 4A) designated
<typeStatus id="02348854FFA00D0A4B25CE89FB5BF840" box="[1103,1205,2010,2036]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
(
<collectionCode id="BB9EAE33FFA00D0A4BAFCE89FAF2F840" box="[1221,1308,2010,2036]" httpUri="http://grbio.org/cool/8z6v-f8is" name="Saskatchewan Museum of Natural History" pageId="6" pageNumber="7">SMNH</collectionCode>
no. 7370), one serially-sectioned specimen designated as
<typeStatus id="02348854FFA10D0B4D8FC9CBFCBFFF06" box="[741,849,152,178]" pageId="7" pageNumber="8" type="paratype">paratype</typeStatus>
(
<collectionCode id="BB9EAE33FFA10D0B4C08C9CBFC53FF06" box="[866,957,152,178]" httpUri="http://grbio.org/cool/8z6v-f8is" name="Saskatchewan Museum of Natural History" pageId="7" pageNumber="8">SMNH</collectionCode>
no. 7371). There are DNA-sequences (accession numbers:
<accessionNumber id="C2DCAB15FFA10D0B4EE5C9EEFDE8FF63" box="[399,518,189,215]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ196330" pageId="7" pageNumber="8">FJ196330</accessionNumber>
,
<accessionNumber id="C2DCAB15FFA10D0B4D79C9EEFD64FF63" box="[531,650,189,215]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ196341" pageId="7" pageNumber="8">FJ196341</accessionNumber>
,
<accessionNumber id="C2DCAB15FFA10D0B4DFCC9EEFCE0FF63" box="[662,782,189,215]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ384929" pageId="7" pageNumber="8">FJ384929</accessionNumber>
,
<accessionNumber id="C2DCAB15FFA10D0B4C70C9EEFC7FFF63" box="[794,913,189,215]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ384887" pageId="7" pageNumber="8">FJ384887</accessionNumber>
,
<accessionNumber id="C2DCAB15FFA10D0B4CF4C9EEFBFBFF63" box="[926,1045,189,215]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ384831" pageId="7" pageNumber="8">FJ384831</accessionNumber>
,
<accessionNumber id="C2DCAB15FFA10D0B4B48C9EEFB75FF63" box="[1058,1179,189,215]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ384959" pageId="7" pageNumber="8">FJ384959</accessionNumber>
), which will facilitate unambiguous identification (see Larsson
<emphasis id="EFFBEAE4FFA10D0B4D10C9B1FD5BFF4F" box="[634,693,226,251]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
2008).
</paragraph>
</subSubSection>
<subSubSection id="9595657DFFA10D0B4FAEC854FBDEFE96" box="[196,1072,263,290]" pageId="7" pageNumber="8" type="etymology">
<paragraph id="DD3036F6FFA10D0B4FAEC854FBDEFE96" blockId="7.[151,1437,152,402]" box="[196,1072,263,290]" pageId="7" pageNumber="8">
<emphasis id="EFFBEAE4FFA10D0B4FAEC854FEBFFE95" bold="true" box="[196,337,263,289]" pageId="7" pageNumber="8">Etymology.</emphasis>
Species name refers to the
<typeStatus id="02348854FFA10D0B4DF9C85BFD2BFE96" box="[659,709,264,290]" pageId="7" pageNumber="8">type</typeStatus>
locality Heebukten, Bohuslän.
</paragraph>
</subSubSection>
<subSubSection id="9595657DFFA10D044FAEC87EFC66FDB6" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
<paragraph id="DD3036F6FFA10D0B4FAEC87EFC24FE26" blockId="7.[151,1437,152,402]" pageId="7" pageNumber="8">
<emphasis id="EFFBEAE4FFA10D0B4FAEC87EFE8AFEF3" bold="true" box="[196,356,301,327]" pageId="7" pageNumber="8">Description.</emphasis>
The animals are 0.5 mm; animals with one or two zooids are found. They are white in incident light with a dark and spotted gut caused by ingested material. Excretophores are absent. The body is evenly shaped and cylindrical with the posterior end slightly tapering.
</paragraph>
<caption id="89F0667EFFA10D0B4FFDCF3DFA7EF971" httpUri="https://zenodo.org/record/193968/files/figure.png" pageId="7" pageNumber="8" targetBox="[151,1435,474,1623]" targetPageId="7">
<paragraph id="DD3036F6FFA10D0B4FFDCF3DFA7EF971" blockId="7.[151,1436,1646,1733]" pageId="7" pageNumber="8">
<emphasis id="EFFBEAE4FFA10D0B4FFDCF3DFEF7F932" bold="true" box="[151,281,1646,1670]" pageId="7" pageNumber="8">FIGURE 4.</emphasis>
<taxonomicName id="1A8F4D75FFA10D0B4E4ACF3DFDD9F931" box="[288,567,1646,1669]" class="Catenulida" family="Stenostomidae" genus="Stenostomum" kingdom="Animalia" order="Gentianales" pageId="7" pageNumber="8" phylum="Platyhelminthes" rank="species" species="heebuktense" status="sp. nov.">
<emphasis id="EFFBEAE4FFA10D0B4E4ACF3DFDD9F931" box="[288,567,1646,1669]" italics="true" pageId="7" pageNumber="8">Stenostomum heebuktense</emphasis>
</taxonomicName>
<emphasis id="EFFBEAE4FFA10D0B4D54CF3DFD94F932" bold="true" box="[574,634,1646,1670]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="F4C8579FFFA10D0B4D54CF3DFD94F932" box="[574,634,1646,1670]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(A–B: photographs of live specimens; C: habitus of live specimen; D: live individual illustrating pharynx constriction; E: sagittal reconstruction of the pharynx from the left hand side (pharynx appears shorter in comparison with live animals due to constriction during fixation); F: anterior end of sagittal section).
</paragraph>
</caption>
<paragraph id="DD3036F6FFA10D0B4FAECFA7FE87F817" blockId="7.[151,1437,1780,2030]" pageId="7" pageNumber="8">The epithelium is covered with short cilia and basally comprises secretory vesicles, possibly rhabdoids. The body wall musculature consists of very weak, hardly visible inner circular and very strong outer longitudinal muscles. The small ciliated pits are lateral and situated a bit close to the anterior tip. The anterior brain lobes are clearly visible and dentate but not divided into separate ganglia. Refractile organs are absent.</paragraph>
<paragraph id="DD3036F6FFA10D044FAECEFDFD3CFE26" blockId="7.[151,1437,1780,2030]" lastBlockId="8.[151,1437,152,999]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">The large mouth opening is located at 1/4 and is surrounded by small eosinophilic glands (fig. 4 C, E: phg), creating a lip-like structure in live animals. The pharynx is very long and extremely conspicuous in live animals, because of its bluish colour (under incident light) and continuous contractions whereby its width is reduced by 2/3 (fig. 4 D). Its total length is slightly variable, ranging between 2/3 and 1/2 of the total body length. Both the pharynx’ distal (fig. 4 E: sph1) and proximal (fig. 4 E: sph2) ends have a sphincter. The distal one is extremely thick and situated very close to the mouth. In between both sphincters the pharynx is surrounded by inner longitudinal and outer circular muscles. It is connected to the body wall by very strong radial muscles (fig. 4 E rm). The pharynx lumen is lined with a high, nucleated epithelium, which is slightly degenerated forming a so-called pseudociliation.</paragraph>
<paragraph id="DD3036F6FFAE0D044FAEC8CEFED6FE68" blockId="8.[151,1437,152,999]" pageId="8" pageNumber="9">The protonephridium is visible in live specimens, and the nephridiopore is situated terminally at the posterior end.</paragraph>
<paragraph id="DD3036F6FFAE0D044FAEC8BBFC66FDB6" blockId="8.[151,1437,152,999]" box="[196,904,488,514]" pageId="8" pageNumber="9">Individuals with a developed genital system were not found.</paragraph>
</subSubSection>
<subSubSection id="9595657DFFAE0D044FAECB5EFD79FDC6" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="DD3036F6FFAE0D044FAECB5EFD79FDC6" blockId="8.[151,1437,152,999]" pageId="8" pageNumber="9">
<emphasis id="EFFBEAE4FFAE0D044FAECB5EFEA5FD93" bold="true" box="[196,331,525,551]" pageId="8" pageNumber="9">Diagnosis.</emphasis>
<taxonomicName id="1A8F4D75FFAE0D044E3ECB5EFD79FD92" box="[340,663,525,550]" class="Catenulida" family="Stenostomidae" genus="Stenostomum" kingdom="Animalia" order="Gentianales" pageId="8" pageNumber="9" phylum="Platyhelminthes" rank="species" species="heebuktense" status="sp. nov.">
<emphasis id="EFFBEAE4FFAE0D044E3ECB5EFD79FD92" box="[340,663,525,550]" italics="true" pageId="8" pageNumber="9">Stenostomum heebuktense</emphasis>
</taxonomicName>
<emphasis id="EFFBEAE4FFAE0D044DF5CB5EFD08FD93" bold="true" box="[671,742,525,551]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="F4C8579FFFAE0D044DF5CB5EFD08FD93" box="[671,742,525,551]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: species of
<taxonomicName id="1A8F4D75FFAE0D044C17CB5EFBCEFD92" box="[893,1056,525,550]" class="Catenulida" family="Stenostomidae" genus="Stenostomum" kingdom="Animalia" order="Gentianales" pageId="8" pageNumber="9" phylum="Platyhelminthes" rank="genus">
<emphasis id="EFFBEAE4FFAE0D044C17CB5EFBCEFD92" box="[893,1056,525,550]" italics="true" pageId="8" pageNumber="9">Stenostomum</emphasis>
</taxonomicName>
with one or two zooids, small ciliated pits, anterior brain lobes with a dentate appearance, an extremely long and muscular pharynx. Refractile organs and excretophores absent.
</paragraph>
</subSubSection>
<subSubSection id="9595657DFFAE0D044FAECB2EFAB8FC53" pageId="8" pageNumber="9" type="discussion">
<paragraph id="DD3036F6FFAE0D044FAECB2EFC7AFD56" blockId="8.[151,1437,152,999]" pageId="8" pageNumber="9">
<emphasis id="EFFBEAE4FFAE0D044FAECB2EFEBEFD23" bold="true" box="[196,336,637,663]" pageId="8" pageNumber="9">Discussion.</emphasis>
Based on the previously mentioned diagnostic characters this species can be assigned to the taxon
<taxonomicName id="1A8F4D75FFAE0D044F8CCBF1FE62FD0F" box="[230,396,674,699]" class="Catenulida" family="Stenostomidae" genus="Stenostomum" kingdom="Animalia" order="Gentianales" pageId="8" pageNumber="9" phylum="Platyhelminthes" rank="genus">
<emphasis id="EFFBEAE4FFAE0D044F8CCBF1FE62FD0F" box="[230,396,674,699]" italics="true" pageId="8" pageNumber="9">Stenostomum</emphasis>
</taxonomicName>
. However, this species is characterized by the presence of a remarkable and very conspicuous pharynx, absent in all other species of
<taxonomicName id="1A8F4D75FFAE0D044D98CB9BFC63FD55" box="[754,909,712,737]" class="Catenulida" family="Stenostomidae" genus="Stenostomum" kingdom="Animalia" order="Gentianales" pageId="8" pageNumber="9" phylum="Platyhelminthes" rank="genus">
<emphasis id="EFFBEAE4FFAE0D044D98CB9BFC63FD55" box="[754,909,712,737]" italics="true" pageId="8" pageNumber="9">Stenostomum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="DD3036F6FFAE0D044FAECBBEFD30FC28" blockId="8.[151,1437,152,999]" pageId="8" pageNumber="9">
This very long and barrel-shaped, muscular pharynx is highly characteristic for this species and, at first sight, in live animals, looks similar to the muscular gut (“Muskeldarm” in
<bibRefCitation id="B91E4B07FFAE0D044B69CA41FB76FC98" author="Luther" box="[1027,1176,786,812]" pageId="8" pageNumber="17" refString="Luther, A. (1960) Die Turbellarien Ostfennoskandiens. Fauna Fennica, 7, 1 - 155." type="journal article" year="1960">Luther 1960</bibRefCitation>
), present in species of
<taxonomicName id="1A8F4D75FFAE0D044FFDCA6BFDFEFCE6" authority="Luther, 1960" authorityName="Luther" authorityYear="1960" box="[151,528,824,850]" class="Catenulida" family="Stenostomidae" genus="Myostenostomum" kingdom="Animalia" pageId="8" pageNumber="9" phylum="Platyhelminthes" rank="genus">
<emphasis id="EFFBEAE4FFAE0D044FFDCA6BFE87FCE5" box="[151,361,824,849]" italics="true" pageId="8" pageNumber="9">Myostenostomum</emphasis>
<bibRefCitation id="B91E4B07FFAE0D044E1BCA6BFDFEFCE6" author="Luther" box="[369,528,824,850]" pageId="8" pageNumber="17" refString="Luther, A. (1960) Die Turbellarien Ostfennoskandiens. Fauna Fennica, 7, 1 - 155." type="journal article" year="1960">Luther, 1960</bibRefCitation>
</taxonomicName>
. However, this muscular gut has a completely different anatomy, as can be seen on sectioned individuals. It consists of two half-circular large muscle bundles on each side of the distal part of the pharynx (
<bibRefCitation id="B91E4B07FFAE0D044EEFCAD1FDF5FC28" author="Luther" box="[389,539,898,924]" pageId="8" pageNumber="17" refString="Luther, A. (1960) Die Turbellarien Ostfennoskandiens. Fauna Fennica, 7, 1 - 155." type="journal article" year="1960">Luther 1960</bibRefCitation>
;
<bibRefCitation id="B91E4B07FFAE0D044D4DCAD1FD3EFC28" author="Rogozin" box="[551,720,898,924]" pageId="8" pageNumber="18" refString="Rogozin, A. G. (1992) Short revison of the genus Myostenostomum (Turbellaria, Catenulida). Zoologicheskii Zhurnal, 71, 5 - 11." type="journal article" year="1992">Rogozin 1992</bibRefCitation>
).
</paragraph>
<paragraph id="DD3036F6FFAE0D044FAECAFBFAB8FC53" blockId="8.[151,1437,152,999]" pageId="8" pageNumber="9">
According to the molecular analyses of Swedish
<taxonomicName id="1A8F4D75FFAE0D044C75CAFBFC4DFC76" box="[799,931,936,962]" class="Turbellaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Catenulida" pageId="8" pageNumber="9" phylum="Platyhelminthes" rank="order">Catenulida</taxonomicName>
in Larsson
<emphasis id="EFFBEAE4FFAE0D044B5FCAFBFB82FC75" box="[1077,1132,936,961]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. 2008 (see figure 2), the closest relatives of
<taxonomicName id="1A8F4D75FFAE0D044E12CA9EFDCAFC52" box="[376,548,973,998]" class="Catenulida" family="Stenostomidae" genus="Stenostomum" kingdom="Animalia" order="Gentianales" pageId="8" pageNumber="9" phylum="Platyhelminthes" rank="species" species="heebuktense">
<emphasis id="EFFBEAE4FFAE0D044E12CA9EFDCAFC52" box="[376,548,973,998]" italics="true" pageId="8" pageNumber="9">S. heebuktense</emphasis>
</taxonomicName>
are two other newly described species,
<taxonomicName id="1A8F4D75FFAE0D044C92CA9EFB49FC52" box="[1016,1191,973,998]" class="Catenulida" family="Stenostomidae" genus="Stenostomum" kingdom="Animalia" order="Gentianales" pageId="8" pageNumber="9" phylum="Platyhelminthes" rank="species" species="handoelense">
<emphasis id="EFFBEAE4FFAE0D044C92CA9EFB49FC52" box="[1016,1191,973,998]" italics="true" pageId="8" pageNumber="9">S. handoelense</emphasis>
</taxonomicName>
and
<taxonomicName id="1A8F4D75FFAE0D044B8ACA9EFAA1FC52" box="[1248,1359,973,998]" class="Catenulida" family="Stenostomidae" genus="Stenostomum" kingdom="Animalia" order="Gentianales" pageId="8" pageNumber="9" phylum="Platyhelminthes" rank="species" species="steveoi">
<emphasis id="EFFBEAE4FFAE0D044B8ACA9EFAA1FC52" box="[1248,1359,973,998]" italics="true" pageId="8" pageNumber="9">S. steveoi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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