<documentid="54976E42438440ACD2190D0DEDCADB59"ID-CLB-Dataset="21841"ID-DOI="10.5852/ejt.2012.12"ID-GBIF-Dataset="5217c4a1-a0dd-478e-bd0c-95df35bad69b"ID-ISSN="2118-9773"ID-Zenodo-Dep="3857751"IM.bibliography_approvedBy="juliana"IM.materialsCitations_approvedBy="juliana"IM.metadata_approvedBy="felipe"IM.tables_approvedBy="juliana"IM.taxonomicNames_approvedBy="juliana"IM.treatmentCitations_approvedBy="juliana"IM.treatments_approvedBy="juliana"checkinTime="1590516617597"checkinUser="carolina"docAuthor="Oliver, P. Graham"docDate="2012"docId="546087F47C49FFF38749224FFDB7F8BD"docLanguage="en"docName="ejt-2012-12.pdf.imf"docOrigin="European Journal of Taxonomy 12"docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1"docStyleId="EF2B578F1D15862ADE45B0C07C620911"docStyleName="EJT.2018-.journal_article.type1"docStyleVersion="14"docTitle="Ptilomyax hadalis Oliver 2012, sp. nov."docType="treatment"docVersion="11"lastPageNumber="20"masterDocId="A859FF8C7C58FFE08535240CFFB0FFB5"masterDocTitle="Taxonomy of some Galeommatoidea (Mollusca, Bivalvia) associated with deep-sea echinoids: A reassessment of the bivalve genera Axinodon Verrill & Bush, 1898 and Kelliola Dall, 1899 with descriptions of new genera Syssitomya gen. nov. and Ptilomyax gen. nov."masterLastPageNumber="24"masterPageNumber="1"pageNumber="18"updateTime="1736532211476"updateUser="ExternalLinkService"zenodo-license-document="CC-BY-3.0"zenodo-license-figures="CC-BY-3.0"zenodo-license-treatments="UNSPECIFIED">
<mods:titleid="95C5288A2440089D2F6C9D57647ECD7B">Taxonomy of some Galeommatoidea (Mollusca, Bivalvia) associated with deep-sea echinoids: A reassessment of the bivalve genera Axinodon Verrill & Bush, 1898 and Kelliola Dall, 1899 with descriptions of new genera Syssitomya gen. nov. and Ptilomyax gen. nov.</mods:title>
<bibRefCitationid="B8584B137C49FFF18479228BFD88F914"author="MironovA. N."box="[332,568,1671,1697]"journalOrPublisher="Zoologicheskii Zhurnal"pageId="17"pageNumber="18"pagination="721 - 726"part="57"refId="ref9628"refString="MironovA. N. 1978. The most deep-sea species of sea urchins (Echinoidea: Pourtalesiidae). Zoologicheskii Zhurnal 57 (5): 721 - 726."title="The most deep-sea species of sea urchins (Echinoidea: Pourtalesiidae)"type="journal article"year="1978">Mironov, 1978: 722</bibRefCitation>
<collectingDateid="B833E9CA7C49FFF1841123DDFE5CF85E"box="[292,492,2000,2027]"pageId="17"pageNumber="18"value="1975-03-21"valueMax="1975-03-22"valueMin="1975-03-21">21-22 Mar. 1975</collectingDate>
<materialsCitationid="6CA13CBF7C4BFFF385DF2507FB6DFEFC"ID-GBIF-Occurrence="2625650304"collectingDate="1975-03-21"collectingDateMax="1975-03-22"collectingDateMin="1975-03-21"collectionCode="ZMMU"collectorName="A. N. Mironov"elevation="7335.0"latitude="-5.616667"location="RV Vityaz"longLatPrecision="930"longitude="131.125"municipality="Moscow"pageId="19"pageNumber="20"specimenCode="ZMMU Ld-3043"specimenCount="1"typeStatus="holotype">
<materialsCitationid="6CA13CBF7C4BFFF385DF257DFB53FE1A"ID-GBIF-Occurrence="5004498304"collectingDate="1975-03-21"collectingDateMax="1975-03-22"collectingDateMin="1975-03-21"collectionCode="ZMMU"collectorName="A. N. Mironov"elevation="7335.0"latitude="-5.616667"location="RV Vityaz"longLatPrecision="930"longitude="131.125"municipality="Moscow"pageId="19"pageNumber="20"specimenCode="Ld-3044"specimenCount="14"typeStatus="paratype">
<materialsCitationid="6CA13CBF7C4BFFF381DD2599FBA1FE62"ID-GBIF-Occurrence="5004498302"collectingDate="1975-03-21"collectingDateMax="1975-03-22"collectingDateMin="1975-03-21"collectionCode="NMW.Z"collectorName="A. N. Mironov"elevation="7335.0"latitude="-5.616667"location="RV Vityaz"longLatPrecision="930"longitude="131.125"municipality="Moscow"pageId="19"pageNumber="20"specimenCode="2012.013"specimenCount="3"typeStatus="paratype">
<paragraphid="DC7636E27C4BFFF3858825F3FC8BFD3D"blockId="19.[189,1399,511,649]"pageId="19"pageNumber="20">The material has been poorly preserved, thus the shells are considerably softened, and the tissues are rather contracted. No attempt has been made to examine the hinge, as the valves are so soft that they do not survive being separated. However, the distinctive external form of the shell and the anatomy are sufficient to describe it and place it at the family level.</paragraph>
<figureCitationid="44F22A677C4BFFF3842926F2FEC2FCAC"box="[284,370,766,793]"captionStart="Fig"captionStartId="18.[189,232,1965,1991]"captionTargetBox="[189,1399,266,1925]"captionTargetId="figure@18.[189,1399,266,1925]"captionTargetPageId="18"captionText="Fig. 10. Ptilomyax hadalis sp. nov. A. Holotype attached to echinoid spines. B. Paratype, left side. C. Juvenile paratype lacking “wings”. D. Anatomy of a paratype as viewed from the left side."figureDoi="http://doi.org/10.5281/zenodo.3857771"httpUri="https://zenodo.org/record/3857771/files/figure.png"pageId="19"pageNumber="20">Fig. 10</figureCitation>
H). Inequivalve in adult, lateral dorsal margins of right valve elevated and rolled over projecting beyond left valve. Outline slightly inequilateral, beaks slightly prosogyrate, just behind the mid line, anterior more expanded than posterior; dorsal margins more or less straight, those in adult right valve projecting laterally, becoming alate; forming distinct lateral sinuses with adjacent anterior and posterior margins; lateral margins broadly rounded, anterior more so; ventral margin curved. Ligament small, posterior, deeply sunken; dorsal margin with thickened periostracal “ligament”. Hinge plate not examined. Sculpture almost smooth with faint commarginal lines. Prodissoconch I,
<figureCitationid="44F22A677C4BFFF38474206EFE1CFBC9"box="[321,428,1122,1148]"captionStart="Fig"captionStartId="18.[189,232,1965,1991]"captionTargetBox="[189,1399,266,1925]"captionTargetId="figure@18.[189,1399,266,1925]"captionTargetPageId="18"captionText="Fig. 10. Ptilomyax hadalis sp. nov. A. Holotype attached to echinoid spines. B. Paratype, left side. C. Juvenile paratype lacking “wings”. D. Anatomy of a paratype as viewed from the left side."figureDoi="http://doi.org/10.5281/zenodo.3857771"httpUri="https://zenodo.org/record/3857771/files/figure.png"pageId="19"pageNumber="20">Fig. 10D</figureCitation>
) Mantle edge mostly unfused with extensive anterior-ventral pedal aperture; fused with terminal of gill axis; exhalant aperture small. Anterior-ventral inner margin raised into series of prominent evenly spaced transverse ridges. Anterior and posterior adductor muscles of similar size, oval in section. Posterior pedal retractor and anterior pedal muscles present. Ctenidium of a single demibranch, ascending and descending lamellae of almost equal size. Labial palps small. Foot with well developed toe, sole flattened and apparently with tissue projections (these may be an artefact of preservation); heel distinct, byssus functional producing numerous thin filaments.
shares many common features with other montacutids associated with echinoids including: a large anterior pedal aperture, mantle fusion limited to junction with gill axis, ctenidium with a single demibranch, foot with prominent toe with a sole and with a functional byssus, ligament internal and weak hinge with poorly developed teeth (
<bibRefCitationid="B8584B137C4BFFF387272273FD09F92C"author="Oldfield"box="[530,697,1663,1689]"firstAuthor="Oldfield"pageId="19"pageNumber="20"pagination="255 - 295"refId="ref9844"refString="Oldfield E. 1961. The functional morphology of Kellia suborbicularis (Montagu), Montacuta ferruginosa (Montagu) and M. substriata (Montagu), (Mollusca, Lamellibranchiata). Proceedings of the Malacological Society of London 34 (5): 255 - 295."type="journal article"year="1961">Oldfield 1961</bibRefCitation>
;
<bibRefCitationid="B8584B137C4BFFF387F12273FC20F92F"author="Ockelmann"box="[708,912,1663,1690]"firstAuthor="Ockelmann"pageId="19"pageNumber="20"pagination="211 - 221"refId="ref9716"refString="Ockelmann K. W. 1965. Redescription, distribution, biology, and dimorphous sperm of Montacuta tenella Loven (Mollusca, Leptonacea). Ophelia 2 (1): 211 - 221."type="journal article"year="1965">Ockelmann 1965</bibRefCitation>
;
<bibRefCitationid="B8584B137C4BFFF386A92273FB83F92C"author="Ponder"box="[924,1075,1663,1689]"firstAuthor="Ponder"pageId="19"pageNumber="20"pagination="125 - 131"refId="ref10044"refString="Ponder W. F. 1968. Three commensal bivalves from New Zealand. Records of the Dominion Museum New Zealand 6: 125 - 131."type="journal article"year="1968">Ponder 1968</bibRefCitation>
;
<bibRefCitationid="B8584B137C4BFFF3810A2273FA87F92C"author="Jespersen"box="[1087,1335,1663,1689]"etAl="et al."firstAuthor="Jespersen"pageId="19"pageNumber="20"pagination="3 - 19"refId="ref9429"refString="Jespersen A., Lutzen J. & Nielsen C. 2004. On three species and two new genera (Montacutella and Brachiomya) of galeommatid bivalves from the irregular Sea Urchin Brissus latecarinatus with emphasis on their reproduction. Zoologischer Anzeiger 243: 3 - 19. http: // dx. doi. org / 10.1016 / j. jcz. 2004.04.001"type="journal article"year="2004">
exhibit an extraordinary range of shell morphologies as can be seen in
<bibRefCitationid="B8584B137C4BFFF381F222AFFAC6F90B"author="Chavan"box="[1223,1398,1699,1726]"firstAuthor="Chavan"pageId="19"pageNumber="20"pagination="518 - 537"refId="ref8709"refString="Chavan A. 1969. Superfamily Leptonacea Gray, 1847. In: Moore R. C. (ed.) Treatise on Invertebrate Zoology Part. N., Mollusca 6, Bivalvia 2: 518 - 537. University of Kansas and Geological Society of America, Lawrence, Kansas; Boulder, Colorado."type="journal article"year="1969">Chavan (1969)</bibRefCitation>
but none have projecting dorsal margins. This unique feature is sufficient to warrant the creation of a new genus and new species.
