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<document id="5D9CA22D1C70B290B2D1D4D45D621C66" ID="10.11646/zootaxa.4258.2.7SLASH10910" ID-CLB-Dataset="33153" ID-DOI="10.11646/zootaxa.4258.2.7" ID-GBIF-Dataset="afeddac3-13cf-48ea-a7b3-f7e7618fe7c3" ID-ISSN="1175-5326" ID-Zenodo-Dep="569461" ID-ZooBank="ACE70044-C5DF-4F7C-8C0E-8224E323D7AD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1493282659784" checkinUser="plazi" docAuthor="Smith-Vaniz, William F." docDate="2017" docId="543B2D494312FFDE31A0FD0ECDE2FE9D" docLanguage="en" docName="zootaxa.4258.2.7SLASH10910.pdf" docOrigin="Zootaxa 4258 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Adelotremus deloachi Smith-Vaniz, 2017, new species" docType="treatment" docVersion="8" lastPageNumber="186" masterDocId="A80255314313FFD93137FFE1CE2EFFF9" masterDocTitle="A new species of the fangblenny Adelotremus from Indonesia, with supplemental description of A. leptus (Teleostei: Blenniidae: Nemophini)" masterLastPageNumber="186" masterPageNumber="179" pageNumber="180" updateTime="1698439104488" updateUser="ExternalLinkService">
<mods:mods id="4A9E560B96185EE1DF5616E2FDCB2C49" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D2B1F3BE5E8836BFA31038A18CF29D90">
<mods:title id="FA946869F3721F829772699C5AA102B9">A new species of the fangblenny Adelotremus from Indonesia, with supplemental description of A. leptus (Teleostei: Blenniidae: Nemophini)</mods:title>
</mods:titleInfo>
<mods:name id="D627993B09DBE1E101496EED63A0FFFB" type="personal">
<mods:role id="CB0D75580F16B934938C30A8059E4532">
<mods:roleTerm id="B77FEB37CEC87F42F6C540CE50825839">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B42E1276BB4215BE09237378C48D8882">Smith-Vaniz, William F.</mods:namePart>
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<mods:typeOfResource id="547D3273FBA7F7CC7B8D2B1A16D901CE">text</mods:typeOfResource>
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<mods:title id="4422C52210152DF671A0909D223C47EC">Zootaxa</mods:title>
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<mods:part id="E4A3E452BA6F74C10516ABDBC721A6DF">
<mods:date id="78FA68394682C255F12D384462F34801">2017</mods:date>
<mods:detail id="AB2C732A18D6FF3F603109A1935C684F" type="volume">
<mods:number id="FA92A577C81AB4F135AE2ADF000671A6">4258</mods:number>
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<mods:classification id="94AE3B0DAEC35BB7A8C51D60D3B9FF64">journal article</mods:classification>
<mods:identifier id="0BA146BE944A873EEFDFF70AC694ABE8" type="CLB-Dataset">33153</mods:identifier>
<mods:identifier id="97C11A2BF1A5139C4696739BB5D6F77D" type="DOI">10.11646/zootaxa.4258.2.7</mods:identifier>
<mods:identifier id="AA8DFAA230A1275C2E613A7B20059643" type="GBIF-Dataset">afeddac3-13cf-48ea-a7b3-f7e7618fe7c3</mods:identifier>
<mods:identifier id="869D0C09D3C7BF8E4A6ED348C73EE44F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="789B7799022EB4EFEE41AFA635842082" type="Zenodo-Dep">569461</mods:identifier>
<mods:identifier id="52B57E596F3DFB411AEE2A1059FB0D9E" type="ZooBank">ACE70044-C5DF-4F7C-8C0E-8224E323D7AD</mods:identifier>
</mods:mods>
<treatment id="543B2D494312FFDE31A0FD0ECDE2FE9D" ID-DOI="http://doi.org/10.5281/zenodo.6007643" ID-GBIF-Taxon="128810316" ID-Zenodo-Dep="6007643" LSID="urn:lsid:plazi:treatment:543B2D494312FFDE31A0FD0ECDE2FE9D" httpUri="http://treatment.plazi.org/id/543B2D494312FFDE31A0FD0ECDE2FE9D" lastPageId="7" lastPageNumber="186" pageId="1" pageNumber="180">
<subSubSection id="9488CFD44312FFD831A0FD0ECF43FCD2" pageId="1" pageNumber="180" type="nomenclature">
<paragraph id="DC2D9C5F4312FFD831A0FD0ECC1BFCF3" blockId="1.[151,565,751,847]" box="[151,565,751,778]" pageId="1" pageNumber="180">
<heading id="87652B334312FFD831A0FD0ECC1BFCF3" bold="true" box="[151,565,751,778]" fontSize="11" level="1" pageId="1" pageNumber="180" reason="1">
<emphasis id="EEE6404D4312FFD831A0FD0ECC1BFCF3" bold="true" box="[151,565,751,778]" pageId="1" pageNumber="180">
<taxonomicName id="1B92E7DC4312FFD831A0FD0ECFB3FCF0" box="[151,413,751,777]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="180" phylum="Chordata" rank="species" species="deloachi" status="sp. nov.">
<emphasis id="EEE6404D4312FFD831A0FD0ECFB3FCF0" bold="true" box="[151,413,751,777]" italics="true" pageId="1" pageNumber="180">Adelotremus deloachi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F5D5FD364312FFD83092FD11CC1BFCF3" box="[421,565,752,778]" pageId="1" pageNumber="180" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="DC2D9C5F4312FFD831A0FCF3CF43FCD2" blockId="1.[151,565,751,847]" box="[151,365,786,811]" pageId="1" pageNumber="180">Spotfin fangblenny</paragraph>
</subSubSection>
<subSubSection id="9488CFD44312FFD831A0FCD7CD50FBDE" pageId="1" pageNumber="180" type="description">
<paragraph id="DC2D9C5F4312FFD831A0FCD7CF53FCB6" blockId="1.[151,565,751,847]" box="[151,381,822,847]" pageId="1" pageNumber="180">
<figureCitation id="44A980DA4312FFD831A0FCD7CF30FCB6" box="[151,286,822,847]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,250,1447,1469]" captionStartId-1="4.[151,250,1471,1493]" captionStartId-2="4.[151,250,1979,2001]" captionStartId-3="5.[151,250,1923,1945]" captionTargetBox-0="[192,1402,197,1427]" captionTargetBox-1="[217,1369,193,1450]" captionTargetBox-2="[159,1427,1574,1958]" captionTargetBox-3="[407,1188,208,1900]" captionTargetId-0="figure@3.[192,1402,197,1427]" captionTargetId-1="figure@4.[217,1369,193,1450]" captionTargetId-2="figure@4.[159,1427,1574,1958]" captionTargetId-3="figure@5.[384,1206,193,1909]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Adelotremus deloachi holotype, ZMA 23004, male, 32.0 mm SL, Bali, Indonesia. Photograph by Ned DeLoach." captionText-1="FIGURE 2. Adelotremus deloachi paratype, USNM 438965, female, 29.4 mm SL, Lembeh Strait, Indonesia. Photograph by Ned DeLoach." captionText-2="FIGURE 3. Adelotremus deloachi holotype, (right side reversed). Photograph by Mark V. Erdmann." captionText-3="FIGURE 4. Cephalic sensory pores in species of Adelotremus: (above) A. deloachi, holotype, male, 32.0 mm SL, ZMA 23004, Bali, Indonesia; (below) A. leptus, holotype, female, 34.5 mm SL, ZMA 23004, Red Sea, Egypt, modified from Smith-Vaniz and Rose (2009, fig. 5). Small arrows identify ventralmost supraorbital and dorsalmost preopercular pores in both lateral and dorsal views." httpUri-0="https://zenodo.org/record/569462/files/figure.png" httpUri-1="https://zenodo.org/record/569463/files/figure.png" httpUri-2="https://zenodo.org/record/569464/files/figure.png" httpUri-3="https://zenodo.org/record/569465/files/figure.png" pageId="1" pageNumber="180">Figures 1–4</figureCitation>
;
<tableCitation id="9110A9E44312FFD8301CFCD7CF53FCB6" box="[299,381,822,847]" captionStart="TABLE 1" captionStartId="2.[151,239,982,1005]" captionText="TABLE 1. Selected characters of the genus Petroscirtes and the two species of Adelotremus 1" pageId="1" pageNumber="180">Table 1</tableCitation>
</paragraph>
<paragraph id="DC2D9C5F4312FFD831A0FC9ECA0DFC42" blockId="1.[151,1437,894,2035]" pageId="1" pageNumber="180">
<materialsCitation id="6CFA96024312FFD831A0FC9ECA0DFC42" ID-GBIF-Occurrence="1499294092" collectingDate="2012-04-26" collectionCode="ZMA" collectorName="Mark V." country="Indonesia" elevation="17" latitude="-8.288889" location="Seraya" longLatPrecision="21" longitude="115.60722" pageId="1" pageNumber="180" specimenCount="1" stateProvince="Bali">
H
<emphasis id="EEE6404D4312FFD8319CFC9ECF27FC6E" bold="true" box="[171,265,895,919]" pageId="1" pageNumber="180">olotype.</emphasis>
<collectionCode id="BA83049A4312FFD83027FC9ECF60FC6E" box="[272,334,895,919]" country="Netherlands" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="1" pageNumber="180">ZMA</collectionCode>
23004, 32.0 mm SL, male,
<collectingCountry id="A485DCCF4312FFD833BFFC9FCCD6FC6E" box="[648,760,894,919]" name="Indonesia" pageId="1" pageNumber="180">Indonesia</collectingCountry>
,
<collectingRegion id="1E5652BD4312FFD83234FC9FCD1AFC6E" box="[771,820,894,919]" country="Indonesia" name="Bali" pageId="1" pageNumber="180">Bali</collectingRegion>
, off
<location id="D94DCA844312FFD83250FC9ECD98FC6E" LSID="urn:lsid:plazi:treatment:543B2D494312FFDE31A0FD0ECDE2FE9D:D94DCA844312FFD83250FC9ECD98FC6E" box="[871,950,895,919]" country="Indonesia" latitude="-8.288889" longLatPrecision="21" longitude="115.60722" name="Seraya" pageId="1" pageNumber="180" stateProvince="Bali">Seraya</location>
, ca.
<geoCoordinate id="B9A6FA984312FFD832DFFC9ECA72FC6E" box="[1000,1116,895,919]" direction="south" orientation="latitude" pageId="1" pageNumber="180" precision="15" value="-8.288889">8°17'20&quot;S</geoCoordinate>
,
<geoCoordinate id="B9A6FA984312FFD8355FFC9FCAD6FC6E" box="[1128,1272,894,919]" direction="east" orientation="longitude" pageId="1" pageNumber="180" precision="15" value="115.60722">115°36'26&quot;E</geoCoordinate>
, on sand slope in
<quantity id="1B6A31BA4312FFD83183FC42CEC1FC43" box="[180,239,931,955]" metricMagnitude="1" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="180" unit="m" value="17.0">
<elevation id="57BF7B6C4312FFD83183FC42CEC1FC43" box="[180,239,931,955]" metricMagnitude="1" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="180" unit="m" value="17.0">17 m</elevation>
</quantity>
, collected with clove oil and hand-net,
<collectorName id="7167F9894312FFD83390FC43CD2DFC43" box="[679,771,930,955]" pageId="1" pageNumber="180">Mark V.</collectorName>
Erdmann,
<date id="A82CBA9F4312FFD83249FC43CA30FC42" box="[894,1054,930,955]" pageId="1" pageNumber="180" value="2012-04-26">
<collectingDate id="B86843774312FFD83249FC43CA30FC42" box="[894,1054,930,955]" pageId="1" pageNumber="180" value="2012-04-26">26 April 2012</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="DC2D9C5F4312FFD831F0FC26CD50FBDE" blockId="1.[151,1437,894,2035]" pageId="1" pageNumber="180">
<materialsCitation id="6CFA96024312FFD831F0FC26CBB6FBFA" ID-GBIF-Occurrence="1499294086" collectingDate="2015-11-17" collectionCode="ZMA" country="Indonesia" county="Ned DeLoach" elevation="11" latitude="1.4811112" location="Lembeh Strait" longLatPrecision="21" longitude="125.25305" municipality="Aer Bajo One" pageId="1" pageNumber="180" specimenCode="ZMA 23208, 29.4" specimenCount="1" typeStatus="paratype">
<emphasis id="EEE6404D4312FFD831F0FC26CF68FC26" bold="true" box="[199,326,967,991]" pageId="1" pageNumber="180">
<typeStatus id="032922FD4312FFD831F0FC26CF6FFC26" box="[199,321,967,991]" pageId="1" pageNumber="180" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCode id="8C3434244312FFD8307BFC26CC3BFC26" box="[332,533,967,991]" pageId="1" pageNumber="180">
<collectionCode id="BA83049A4312FFD8307BFC26CFA4FC26" box="[332,394,967,991]" country="Netherlands" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="1" pageNumber="180">ZMA</collectionCode>
23208,
<quantity id="1B6A31BA4312FFD830D3FC26CC3BFC26" box="[484,533,967,991]" metricMagnitude="-2" metricUnit="m" metricValue="2.94" pageId="1" pageNumber="180" unit="mm" value="29.4">29.4</quantity>
</specimenCode>
mm SL, female,
<collectingCountry id="A485DCCF4312FFD833E1FC27CD68FC26" box="[726,838,966,991]" name="Indonesia" pageId="1" pageNumber="180">Indonesia</collectingCountry>
,
<location id="D94DCA844312FFD83267FC27CDDFFC26" LSID="urn:lsid:plazi:treatment:543B2D494312FFDE31A0FD0ECDE2FE9D:D94DCA844312FFD83267FC27CDDFFC26" box="[848,1009,966,991]" country="Indonesia" county="Ned DeLoach" latitude="1.4811112" longLatPrecision="21" longitude="125.25305" municipality="Aer Bajo One" name="Lembeh Strait" pageId="1" pageNumber="180">Lembeh Strait</location>
, &quot;
<collectingMunicipality id="3C4906254312FFD83532FC26CA8AFC26" box="[1029,1188,966,991]" pageId="1" pageNumber="180">Aer Bajo One</collectingMunicipality>
&quot; dive site,
<geoCoordinate id="B9A6FA984312FFD83428FC27CBB9FC26" box="[1311,1431,966,991]" direction="north" orientation="latitude" pageId="1" pageNumber="180" precision="15" value="1.4811112">1°28'52&quot;N</geoCoordinate>
,
<geoCoordinate id="B9A6FA984312FFD831A0FC0BCF09FBFA" box="[151,295,1002,1027]" direction="east" orientation="longitude" pageId="1" pageNumber="180" precision="15" value="125.25305">125°15'11&quot;E</geoCoordinate>
, on black sand slope in
<quantity id="1B6A31BA4312FFD8337BFC0ACC9BFBFB" box="[588,693,1003,1027]" metricMagnitude="1" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="1" pageNumber="180" unit="m" value="11.0" valueMax="12.0" valueMin="10.0">
<elevation id="57BF7B6C4312FFD8337BFC0ACC9BFBFB" box="[588,693,1003,1027]" metricMagnitude="1" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="1" pageNumber="180" unit="m" value="11.0" valueMax="12.0" valueMin="10.0">10‒12 m</elevation>
</quantity>
, collected with plastic bag,
<collectingCounty id="354CE4D34312FFD83536FC0BCA8AFBFA" box="[1025,1188,1002,1027]" pageId="1" pageNumber="180">Ned DeLoach</collectingCounty>
,
<date id="A82CBA9F4312FFD83583FC0ACBB6FBFA" box="[1204,1432,1002,1027]" pageId="1" pageNumber="180" value="2015-11-17">
<collectingDate id="B86843774312FFD83583FC0ACBB6FBFA" box="[1204,1432,1002,1027]" pageId="1" pageNumber="180" value="2015-11-17">17 November 2015</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="6CFA96024312FFD831A0FBEECD54FBDE" ID-GBIF-Occurrence="1499294132" box="[151,890,1038,1063]" collectingDate="2015-11-17" collectionCode="USNM" country="Indonesia" county="Ned DeLoach" elevation="11" latitude="1.4811112" location="Lembeh Strait" longLatPrecision="21" longitude="125.25305" municipality="Aer Bajo One" pageId="1" pageNumber="180" specimenCode="USNM 438965, 34.9" specimenCount="1" typeStatus="paratype">
<specimenCode id="8C3434244312FFD831A0FBEECFACFBDE" box="[151,386,1038,1063]" pageId="1" pageNumber="180">
<collectionCode id="BA83049A4312FFD831A0FBEECEC6FBDE" box="[151,232,1039,1063]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="180">USNM</collectionCode>
438965,
<quantity id="1B6A31BA4312FFD83066FBEECFACFBDE" box="[337,386,1039,1063]" metricMagnitude="-2" metricUnit="m" metricValue="3.4899999999999998" pageId="1" pageNumber="180" unit="mm" value="34.9">34.9</quantity>
</specimenCode>
mm SL, male, same data as female
<typeStatus id="032922FD4312FFD83221FBEECD54FBDE" box="[790,890,1039,1063]" pageId="1" pageNumber="180" type="paratype">paratype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9488CFD44312FFD831F0FBD3CA9EFB6A" pageId="1" pageNumber="180" type="diagnosis">
<paragraph id="DC2D9C5F4312FFD831F0FBD3CA9EFB6A" blockId="1.[151,1437,894,2035]" pageId="1" pageNumber="180">
<emphasis id="EEE6404D4312FFD831F0FBD3CF6FFBB2" bold="true" box="[199,321,1074,1099]" pageId="1" pageNumber="180">Diagnosis.</emphasis>
A species of
<taxonomicName id="1B92E7DC4312FFD830EBFBD2CC47FBB2" box="[476,617,1075,1099]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="180" phylum="Chordata" rank="genus">
<emphasis id="EEE6404D4312FFD830EBFBD2CC47FBB2" box="[476,617,1075,1099]" italics="true" pageId="1" pageNumber="180">Adelotremus</emphasis>
</taxonomicName>
distinguished from its only known congener,
<taxonomicName id="1B92E7DC4312FFD83547FBD2CAFDFBB2" box="[1136,1235,1075,1099]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="180" phylum="Chordata" rank="species" species="leptus">
<emphasis id="EEE6404D4312FFD83547FBD2CAFDFBB2" box="[1136,1235,1075,1099]" italics="true" pageId="1" pageNumber="180">A. leptus</emphasis>
</taxonomicName>
, by the following characters: Dorsal-fin spines X (vs. IX); vertebrae 12 + 23 (vs. 13 + 19); and mid-dorsal supratemporal pores 2 (vs. 1). Males have a conspicuous blue ocellus between dorsal-fin spines that is absent in females.
</paragraph>
</subSubSection>
<subSubSection id="9488CFD44312FFDB31F0FB7FCC05F851" lastPageId="2" lastPageNumber="181" pageId="1" pageNumber="180" type="description">
<paragraph id="DC2D9C5F4312FFD831F0FB7FCF90F852" blockId="1.[151,1437,894,2035]" pageId="1" pageNumber="180">
<emphasis id="EEE6404D4312FFD831F0FB7FCF77FB4E" bold="true" box="[199,345,1182,1207]" pageId="1" pageNumber="180">Description.</emphasis>
Where counts differ, those of the
<typeStatus id="032922FD4312FFD833C7FB7FCD7CFB4E" box="[752,850,1182,1207]" pageId="1" pageNumber="180" type="holotype">holotype</typeStatus>
are given first, followed by the larger (male) and smaller (female)
<typeStatus id="032922FD4312FFD83061FB22CFEBFB22" box="[342,453,1219,1243]" pageId="1" pageNumber="180" type="paratype">paratypes</typeStatus>
. Dorsal fin X, 19; last ray broadly attached by membrane to caudal peduncle; first spine with small flap on anterior margin, length of spine slightly shorter than second spine, and with spines 2‒4 subequal in length. Anal fin II, 19; last ray broadly attached by membrane to caudal peduncle. Caudal fin with 6 procurrent rays (3 dorsal + 3 ventral), 11 segmented rays (6 dorsal + 5 ventral), all rays unbranched and outermost dorsal and ventral soft ray elongate; hypural 5 absent; epurals 1. Pectoral-fin rays 13 (both sides). Pelvic fin I, 3. Vertebrae: precaudal 12 + caudal 23; dorsal-fin spine pterygiophores broadly contacting neural spines (see Smith-Vaniz &amp; Rose 2007, figs. 2‒3A). Posteriomost epineurals and pleural ribs on vertebra 13. Upper and lower jaws each with posterior recurved canines (premaxillary canines much smaller than dentary canines) on each side. Incisor teeth broad based and firmly attached, 30 (30, 29) in upper jaw and 26 (30, 28) in lower jaw. Cranial bones ornamented with numerous small pits. Dentaries connected by a tight interdigitating joint at ventral midline. Infraorbital bones 3, including dermophenotic; second infraorbital slender, elongate and tapering to a point posteriorly (see Smith-
<bibRefCitation id="B803E1AE4312FFD831A0F9AFCF56F99E" author="Vaniz" box="[151,376,1614,1639]" pageId="1" pageNumber="180" refString="Smith-Vaniz, W. F. &amp; Rose, J. M. (2012) Adelotremus leptus, a new genus and species of sabertooth blenny from the Red Sea (Teleostei: Blenniidae: Nemophini). Zootaxa, 3249, 39 - 46." type="journal article" year="2012">Vaniz &amp; Rose 2012</bibRefCitation>
, fig. 4A); wide gap between second infraorbital and dermosphenotic indicating the loss of an infraorbital, which corresponds to absence of infraorbital pores in postorbital region of head (
<figureCitation id="44A980DA4312FFD83583F993CAD4F972" box="[1204,1274,1650,1675]" captionStart="FIGURE 4" captionStartId="5.[151,250,1923,1945]" captionTargetBox="[407,1188,208,1900]" captionTargetId="figure@5.[384,1206,193,1909]" captionTargetPageId="5" captionText="FIGURE 4. Cephalic sensory pores in species of Adelotremus: (above) A. deloachi, holotype, male, 32.0 mm SL, ZMA 23004, Bali, Indonesia; (below) A. leptus, holotype, female, 34.5 mm SL, ZMA 23004, Red Sea, Egypt, modified from Smith-Vaniz and Rose (2009, fig. 5). Small arrows identify ventralmost supraorbital and dorsalmost preopercular pores in both lateral and dorsal views." httpUri="https://zenodo.org/record/569465/files/figure.png" pageId="1" pageNumber="180">Fig. 4</figureCitation>
). Gill opening with lateral flap only; ventral margin of gill opening opposite level of dorsalmost 5th or 6th pectoral-fin ray. The cephalic sensory pore system includes 3 dentary pores, 5 preopercular pores, 3 ventral infraorbital pores and the absence of posterior infraorbital pores, 1 pair of interorbital pores, 3 supratemporal pores (mid-dorsal pair and 1 lateral), and 3 posttemporal pores. No lateral-line tubes or associated pores present. Each mid-dorsal supratemporal pore has a minute cirrus, the last posttemporal and the 1st dentary pores each have a well-developed, slender cirrus; no cirri are associated with the preopercular and supraorbital pores and there is no orbital cirrus. Anterior nostril consists of a very short tube without a small flap on posterior margin; posterior nostril with only a slightly raised rim. Swim bladder absent.
</paragraph>
<paragraph id="DC2D9C5F4312FFDB31F0F857CB69FE71" blockId="1.[151,1437,894,2035]" lastBlockId="2.[151,1437,151,824]" lastPageId="2" lastPageNumber="181" pageId="1" pageNumber="180">
Proportions of the 32.0 mm SL male
<typeStatus id="032922FD4312FFD8335CF857CCE3F836" box="[619,717,1974,1999]" pageId="1" pageNumber="180" type="holotype">holotype</typeStatus>
are given first, followed by the male and then female
<typeStatus id="032922FD4312FFD83406F856CBB2F836" box="[1329,1436,1975,1999]" pageId="1" pageNumber="180" type="paratype">paratypes</typeStatus>
(in parentheses) as percent SL: Standard length in mm 32.0 (34.9; 29.4); head length 23.4 (24.1; 25.5); eye diameter 7.2 (7.4; 8.3); preanal length 51.2 (49.9; 55.1); dorsal-fin length 80.0 (81.9; 79.6); anal-fin length 44.4 (43.1; 39.8); depth at anal-fin origin 11.1 (11.4; 11.4); longest outer caudal-fin rays, upper 31.6 (23.8; 25.5) and lower 24.7 (21.8; 30.5); inner caudal-fin ray 18.1 (16.9; 18.1); pelvic fin 7.0 (5.9; 7.3); length of first dorsal-fin spine -- (14.6, 8.8); second spine 12.8 (18.6, 11.6); third spine 13.8 (21.8, 12.4) forth spine 13.4 (21.8, 12.4), spine length measurements only approximate because of small size and curvature of spines. Measurements of the single female and two males suggest that the heights of the anterior dorsal-fin spines are sexually dimorphic, which is not too surprising because males have a prominent ocellus anteriorly on the dorsal fin that is absent in females.
</paragraph>
<paragraph id="DC2D9C5F4311FFDB31F0FE75CFCEFCC1" blockId="2.[151,1437,151,824]" pageId="2" pageNumber="181">
Life coloration (
<figureCitation id="44A980DA4311FFDB30B7FE75CFC3FE55" box="[384,493,404,429]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1447,1469]" captionStartId-1="4.[151,250,1471,1493]" captionStartId-2="4.[151,250,1979,2001]" captionTargetBox-0="[192,1402,197,1427]" captionTargetBox-1="[217,1369,193,1450]" captionTargetBox-2="[159,1427,1574,1958]" captionTargetId-0="figure@3.[192,1402,197,1427]" captionTargetId-1="figure@4.[217,1369,193,1450]" captionTargetId-2="figure@4.[159,1427,1574,1958]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="4" captionText-0="FIGURE 1. Adelotremus deloachi holotype, ZMA 23004, male, 32.0 mm SL, Bali, Indonesia. Photograph by Ned DeLoach." captionText-1="FIGURE 2. Adelotremus deloachi paratype, USNM 438965, female, 29.4 mm SL, Lembeh Strait, Indonesia. Photograph by Ned DeLoach." captionText-2="FIGURE 3. Adelotremus deloachi holotype, (right side reversed). Photograph by Mark V. Erdmann." httpUri-0="https://zenodo.org/record/569462/files/figure.png" httpUri-1="https://zenodo.org/record/569463/files/figure.png" httpUri-2="https://zenodo.org/record/569464/files/figure.png" pageId="2" pageNumber="181">Figs. 1‒3</figureCitation>
). The most striking feature of the life coloration is the prominent ocellus of males. The ocellus is almost as large as the eye, has a narrow white ring, and is mostly confined to membrane of first two spines; the ocellus is oblong (shape and ocellus width influenced by relative spread of dorsal fin, compare
<figureCitation id="44A980DA4311FFDB3478FE3DCBB2FE0D" box="[1359,1436,476,501]" captionStart="FIGURE 1" captionStartId="3.[151,250,1447,1469]" captionTargetBox="[192,1402,197,1427]" captionTargetId="figure@3.[192,1402,197,1427]" captionTargetPageId="3" captionText="FIGURE 1. Adelotremus deloachi holotype, ZMA 23004, male, 32.0 mm SL, Bali, Indonesia. Photograph by Ned DeLoach." httpUri="https://zenodo.org/record/569462/files/figure.png" pageId="2" pageNumber="181">Figs. 1</figureCitation>
and
<figureCitation id="44A980DA4311FFDB31F0FDE1CEFBFDE1" box="[199,213,512,536]" captionStart="FIGURE 3" captionStartId="4.[151,250,1979,2001]" captionTargetBox="[159,1427,1574,1958]" captionTargetId="figure@4.[159,1427,1574,1958]" captionTargetPageId="4" captionText="FIGURE 3. Adelotremus deloachi holotype, (right side reversed). Photograph by Mark V. Erdmann." httpUri="https://zenodo.org/record/569464/files/figure.png" pageId="2" pageNumber="181">3</figureCitation>
of presumably the same individual), and appears black or bright blue depending on light conditions; except for the ocellus, membranes of first three dorsal-fin spines are golden olive, and 3rd spine bordered by narrow white margin; ground color of remainder of the dorsal fin of males light brown, and segmented rays with small, alternating, dark brown and white spots. In contrast to males, the dorsal fin of females anteriorly is densely and uniformly brown speckled and the segmented rays lack alternating spots. Other features of the coloration are a brown to black mid-lateral stripe, approximately width of pupil, extending from snout through middle of eye to base of caudal fin; stripe slightly expanded in front of gill opening; remainder of body varying from mostly white ventrally to tan and speckled with various shades of brown, including series of about eight light blotches; pelvic fin white; and iris golden brown.
</paragraph>
<paragraph id="DC2D9C5F4311FFDB31F0FC89CAABFC5C" blockId="2.[151,1436,871,933]" pageId="2" pageNumber="181">In alcohol-preserved specimens the color pattern, except for the ocellus of males, is essentially the same as in life, consisting of the brown stripe, and speckles and blotches of various shades of brown.</paragraph>
<caption id="88EDCCD74311FFDB31A0FC37CA5AFC19" box="[151,1140,980,1005]" pageId="2" pageNumber="181">
<paragraph id="DC2D9C5F4311FFDB31A0FC37CA5AFC19" blockId="2.[151,1140,980,1005]" box="[151,1140,980,1005]" pageId="2" pageNumber="181">
<emphasis id="EEE6404D4311FFDB31A0FC37CF24FC14" bold="true" box="[151,266,982,1005]" pageId="2" pageNumber="181">TABLE 1.</emphasis>
Selected characters of the genus
<taxonomicName id="1B92E7DC4311FFDB335AFC37CCC3FC14" box="[621,749,982,1005]" class="Actinopterygii" family="Blenniidae" genus="Petroscirtes" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="181" phylum="Chordata" rank="genus">
<emphasis id="EEE6404D4311FFDB335AFC37CCC3FC14" box="[621,749,982,1005]" italics="true" pageId="2" pageNumber="181">Petroscirtes</emphasis>
</taxonomicName>
and the two species of
<taxonomicName id="1B92E7DC4311FFDB32D0FC34CA43FC14" box="[999,1133,981,1005]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="181" phylum="Chordata" rank="genus">
<emphasis id="EEE6404D4311FFDB32D0FC34CA43FC14" box="[999,1133,981,1005]" italics="true" pageId="2" pageNumber="181">Adelotremus</emphasis>
</taxonomicName>
1
</paragraph>
</caption>
<paragraph id="DC2D9C5F4311FFDB31ACFBEDCAF1F8B7" pageId="2" pageNumber="181">
<table id="AE926EFF4311002631ACFBE8CB52F8B7" box="[155,1404,1033,1870]" gridcols="4" gridrows="17" pageId="2" pageNumber="181">
<tr id="62A29E1D4311002631ACFBE8CB52FBC7" box="[155,1404,1033,1086]" gridrow="0" pageId="2" pageNumber="181">
<th id="2173F7614311002631ACFBE8CFF1FBC7" box="[155,479,1033,1086]" gridcol="0" gridrow="0" pageId="2" pageNumber="181">
Characters
<emphasis id="EEE6404D4311FFDB3035FBE8CF27FBEC" bold="true" box="[258,265,1033,1045]" pageId="2" pageNumber="181">¹</emphasis>
</th>
<th id="2173F761431100263320FBE8CD09FBC7" box="[535,807,1033,1086]" gridcol="1" gridrow="0" pageId="2" pageNumber="181">
<taxonomicName id="1B92E7DC4311FFDB3320FBEBCCBFFBE6" box="[535,657,1034,1055]" class="Actinopterygii" family="Blenniidae" genus="Petroscirtes" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="181" phylum="Chordata" rank="genus">
<emphasis id="EEE6404D4311FFDB3320FBEBCCBFFBE6" box="[535,657,1034,1055]" italics="true" pageId="2" pageNumber="181">Petroscirtes</emphasis>
</taxonomicName>
spp. (n = 11 species)
</th>
<th id="2173F761431100263271FBE8CAA0FBC7" box="[838,1166,1033,1086]" gridcol="2" gridrow="0" pageId="2" pageNumber="181">
<taxonomicName id="1B92E7DC4311FFDB3271FBE8CA29FBE7" box="[838,1031,1033,1054]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="181" phylum="Chordata" rank="species" species="leptus">
<emphasis id="EEE6404D4311FFDB3271FBE8CA29FBE7" box="[838,1031,1033,1054]" italics="true" pageId="2" pageNumber="181">Adelotremus leptus</emphasis>
</taxonomicName>
(n = 1 specimen)
</th>
<th id="2173F761431100263597FBE8CB52FBC7" box="[1184,1404,1033,1086]" gridcol="3" gridrow="0" pageId="2" pageNumber="181">
<taxonomicName id="1B92E7DC4311FFDB3597FBE8CB52FBE7" box="[1184,1404,1033,1054]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="181" phylum="Chordata" rank="species" species="deloachi">
<emphasis id="EEE6404D4311FFDB3597FBE8CB52FBE7" box="[1184,1404,1033,1054]" italics="true" pageId="2" pageNumber="181">Adelotremus deloachi</emphasis>
</taxonomicName>
(n = 3 specimens)
</th>
</tr>
<tr id="62A29E1D4311002631ACFBB0CB52FB7F" box="[155,1404,1105,1158]" gridrow="1" pageId="2" pageNumber="181">
<th id="2173F7614311002631ACFBB0CFF1FB7F" box="[155,479,1105,1158]" gridcol="0" gridrow="1" pageId="2" pageNumber="181">Gill opening ventral margin</th>
<td id="2173F761431100263320FBB0CD09FB7F" box="[535,807,1105,1158]" gridcol="1" gridrow="1" pageId="2" pageNumber="181">entirely above pectoral fin</td>
<td id="2173F761431100263271FBB0CAA0FB7F" box="[838,1166,1105,1158]" gridcol="2" gridrow="1" pageId="2" pageNumber="181">opposite level of dorsalmost 5th or 6th pectoral-fin ray</td>
<td id="2173F761431100263597FBB0CB52FB7F" box="[1184,1404,1105,1158]" gridcol="3" gridrow="1" pageId="2" pageNumber="181">
like
<taxonomicName id="1B92E7DC4311FFDB35FBFBB3CB0BFB9F" box="[1228,1317,1105,1127]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="181" phylum="Chordata" rank="species" species="leptus">
<emphasis id="EEE6404D4311FFDB35FBFBB3CB0BFB9F" box="[1228,1317,1105,1127]" italics="true" pageId="2" pageNumber="181">A. leptus</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="62A29E1D4311002631ACFB78CB52FB37" box="[155,1404,1177,1230]" gridrow="2" pageId="2" pageNumber="181">
<th id="2173F7614311002631ACFB78CFF1FB37" box="[155,479,1177,1230]" gridcol="0" gridrow="2" pageId="2" pageNumber="181">Dorsal-fin spine pterygiophores</th>
<td id="2173F761431100263320FB78CD09FB37" box="[535,807,1177,1230]" gridcol="1" gridrow="2" pageId="2" pageNumber="181">not broadly contacting neural spines</td>
<td id="2173F761431100263271FB78CAA0FB37" box="[838,1166,1177,1230]" gridcol="2" gridrow="2" pageId="2" pageNumber="181">broadly contacting neural spines</td>
<td id="2173F761431100263597FB78CB52FB37" box="[1184,1404,1177,1230]" gridcol="3" gridrow="2" pageId="2" pageNumber="181">
like
<taxonomicName id="1B92E7DC4311FFDB35FBFB7BCB0BFB57" box="[1228,1317,1177,1199]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="181" phylum="Chordata" rank="species" species="leptus">
<emphasis id="EEE6404D4311FFDB35FBFB7BCB0BFB57" box="[1228,1317,1177,1199]" italics="true" pageId="2" pageNumber="181">A. leptus</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="62A29E1D4311002631ACFB00CB52FB0E" box="[155,1404,1249,1271]" gridrow="3" pageId="2" pageNumber="181">
<th id="2173F7614311002631ACFB00CFF1FB0E" box="[155,479,1249,1271]" gridcol="0" gridrow="3" pageId="2" pageNumber="181">Infraorbital bones</th>
<td id="2173F761431100263320FB00CD09FB0E" box="[535,807,1249,1271]" gridcol="1" gridrow="3" pageId="2" pageNumber="181">4</td>
<td id="2173F761431100263271FB00CAA0FB0E" box="[838,1166,1249,1271]" gridcol="2" gridrow="3" pageId="2" pageNumber="181">3 (3rd infraorbital absent)</td>
<td id="2173F761431100263597FB00CB52FB0E" box="[1184,1404,1249,1271]" gridcol="3" gridrow="3" pageId="2" pageNumber="181">
like
<taxonomicName id="1B92E7DC4311FFDB35FBFB03CB0BFB0F" box="[1228,1317,1249,1271]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="181" phylum="Chordata" rank="species" species="leptus">
<emphasis id="EEE6404D4311FFDB35FBFB03CB0BFB0F" box="[1228,1317,1249,1271]" italics="true" pageId="2" pageNumber="181">A. leptus</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="62A29E1D4311002631ACFAEBCB52FAD9" box="[155,1404,1290,1312]" gridrow="4" pageId="2" pageNumber="181">
<th id="2173F7614311002631ACFAEBCFF1FAD9" box="[155,479,1290,1312]" gridcol="0" gridrow="4" pageId="2" pageNumber="181">Posterior infraorbital pores</th>
<td id="2173F761431100263320FAEBCD09FAD9" box="[535,807,1290,1312]" gridcol="1" gridrow="4" pageId="2" pageNumber="181">present 3–4</td>
<td id="2173F761431100263271FAEBCAA0FAD9" box="[838,1166,1290,1312]" gridcol="2" gridrow="4" pageId="2" pageNumber="181">absent</td>
<td id="2173F761431100263597FAEBCB52FAD9" box="[1184,1404,1290,1312]" gridcol="3" gridrow="4" pageId="2" pageNumber="181">
like
<taxonomicName id="1B92E7DC4311FFDB35FBFAEACB0BFAD9" box="[1228,1317,1291,1312]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="181" phylum="Chordata" rank="species" species="leptus">
<emphasis id="EEE6404D4311FFDB35FBFAEACB0BFAD9" box="[1228,1317,1291,1312]" italics="true" pageId="2" pageNumber="181">A. leptus</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="62A29E1D4311002631ACFAD5CB52FAB3" box="[155,1404,1332,1354]" gridrow="5" pageId="2" pageNumber="181">
<th id="2173F7614311002631ACFAD5CFF1FAB3" box="[155,479,1332,1354]" gridcol="0" gridrow="5" pageId="2" pageNumber="181">Interorbital pores</th>
<td id="2173F761431100263320FAD5CD09FAB3" box="[535,807,1332,1354]" gridcol="1" gridrow="5" pageId="2" pageNumber="181">4</td>
<td id="2173F761431100263271FAD5CAA0FAB3" box="[838,1166,1332,1354]" gridcol="2" gridrow="5" pageId="2" pageNumber="181">2</td>
<td id="2173F761431100263597FAD5CB52FAB3" box="[1184,1404,1332,1354]" gridcol="3" gridrow="5" pageId="2" pageNumber="181">
like
<taxonomicName id="1B92E7DC4311FFDB35FBFAD5CB0BFAB0" box="[1228,1317,1332,1353]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="181" phylum="Chordata" rank="species" species="leptus">
<emphasis id="EEE6404D4311FFDB35FBFAD5CB0BFAB0" box="[1228,1317,1332,1353]" italics="true" pageId="2" pageNumber="181">A. leptus</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="62A29E1D4311002631ACFABCCB52FA49" box="[155,1404,1373,1456]" gridrow="6" pageId="2" pageNumber="181">
<th id="2173F7614311002631ACFABCCFF1FA49" box="[155,479,1373,1456]" gridcol="0" gridrow="6" pageId="2" pageNumber="181">Preopercular pores</th>
<td id="2173F761431100263320FABCCD09FA49" box="[535,807,1373,1456]" gridcol="1" gridrow="6" pageId="2" pageNumber="181">6 (pores series continuous)</td>
<td id="2173F761431100263271FABCCAA0FA49" box="[838,1166,1373,1456]" gridcol="2" gridrow="6" pageId="2" pageNumber="181">5 (not continuous, wide gap between dorsalmost pore and the others, see Fig. 4)</td>
<td id="2173F761431100263597FABCCB52FA49" box="[1184,1404,1373,1456]" gridcol="3" gridrow="6" pageId="2" pageNumber="181">
like
<taxonomicName id="1B92E7DC4311FFDB35FBFABFCB0BFA8B" box="[1228,1317,1373,1395]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="181" phylum="Chordata" rank="species" species="leptus">
<emphasis id="EEE6404D4311FFDB35FBFABFCB0BFA8B" box="[1228,1317,1373,1395]" italics="true" pageId="2" pageNumber="181">A. leptus</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="62A29E1D4311002631ACFA25CB52FA23" box="[155,1404,1476,1498]" gridrow="7" pageId="2" pageNumber="181">
<th id="2173F7614311002631ACFA25CFF1FA23" box="[155,479,1476,1498]" gridcol="0" gridrow="7" pageId="2" pageNumber="181">Mid-dorsal supratemporal pores</th>
<td id="2173F761431100263320FA25CD09FA23" box="[535,807,1476,1498]" gridcol="1" gridrow="7" pageId="2" pageNumber="181">1 or 2</td>
<td id="2173F761431100263271FA25CAA0FA23" box="[838,1166,1476,1498]" gridcol="2" gridrow="7" pageId="2" pageNumber="181">1</td>
<td id="2173F761431100263597FA25CB52FA23" box="[1184,1404,1476,1498]" gridcol="3" gridrow="7" pageId="2" pageNumber="181">2</td>
</tr>
<tr id="62A29E1D4311002631ACFA0CCB52F9FA" box="[155,1404,1517,1539]" gridrow="8" pageId="2" pageNumber="181">
<th id="2173F7614311002631ACFA0CCFF1F9FA" box="[155,479,1517,1539]" gridcol="0" gridrow="8" pageId="2" pageNumber="181">Lateral line tubes or pores</th>
<td id="2173F761431100263320FA0CCD09F9FA" box="[535,807,1517,1539]" gridcol="1" gridrow="8" pageId="2" pageNumber="181">present</td>
<td id="2173F761431100263271FA0CCAA0F9FA" box="[838,1166,1517,1539]" gridcol="2" gridrow="8" pageId="2" pageNumber="181">absent</td>
<td id="2173F761431100263597FA0CCB52F9FA" box="[1184,1404,1517,1539]" gridcol="3" gridrow="8" pageId="2" pageNumber="181">
like
<taxonomicName id="1B92E7DC4311FFDB35FBFA0FCB0BF9FB" box="[1228,1317,1517,1539]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="181" phylum="Chordata" rank="species" species="leptus">
<emphasis id="EEE6404D4311FFDB35FBFA0FCB0BF9FB" box="[1228,1317,1517,1539]" italics="true" pageId="2" pageNumber="181">A. leptus</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="62A29E1D4311002631ACF9F7CB52F9D5" box="[155,1404,1558,1580]" gridrow="9" pageId="2" pageNumber="181">
<th id="2173F7614311002631ACF9F7CFF1F9D5" box="[155,479,1558,1580]" gridcol="0" gridrow="9" pageId="2" pageNumber="181">Orbital cirri</th>
<td id="2173F761431100263320F9F7CD09F9D5" box="[535,807,1558,1580]" gridcol="1" gridrow="9" pageId="2" pageNumber="181">present or absent</td>
<td id="2173F761431100263271F9F7CAA0F9D5" box="[838,1166,1558,1580]" gridcol="2" gridrow="9" pageId="2" pageNumber="181">absent</td>
<td id="2173F761431100263597F9F7CB52F9D5" box="[1184,1404,1558,1580]" gridcol="3" gridrow="9" pageId="2" pageNumber="181">
like
<taxonomicName id="1B92E7DC4311FFDB35FBF9F6CB0BF9D5" box="[1228,1317,1559,1580]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="181" phylum="Chordata" rank="species" species="leptus">
<emphasis id="EEE6404D4311FFDB35FBF9F6CB0BF9D5" box="[1228,1317,1559,1580]" italics="true" pageId="2" pageNumber="181">A. leptus</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="62A29E1D4311002631ACF9A1CB52F9AF" box="[155,1404,1600,1622]" gridrow="10" pageId="2" pageNumber="181">
<th id="2173F7614311002631ACF9A1CFF1F9AF" box="[155,479,1600,1622]" gridcol="0" gridrow="10" pageId="2" pageNumber="181">Dorsal fin spines</th>
<td id="2173F761431100263320F9A1CD09F9AF" box="[535,807,1600,1622]" gridcol="1" gridrow="10" pageId="2" pageNumber="181">10–12</td>
<td id="2173F761431100263271F9A1CAA0F9AF" box="[838,1166,1600,1622]" gridcol="2" gridrow="10" pageId="2" pageNumber="181">9</td>
<td id="2173F761431100263597F9A1CB52F9AF" box="[1184,1404,1600,1622]" gridcol="3" gridrow="10" pageId="2" pageNumber="181">10</td>
</tr>
<tr id="62A29E1D4311002631ACF988CB52F986" box="[155,1404,1641,1663]" gridrow="11" pageId="2" pageNumber="181">
<th id="2173F7614311002631ACF988CFF1F986" box="[155,479,1641,1663]" gridcol="0" gridrow="11" pageId="2" pageNumber="181">Dorsal fin segmented rays</th>
<td id="2173F761431100263320F988CD09F986" box="[535,807,1641,1663]" gridcol="1" gridrow="11" pageId="2" pageNumber="181">14–21</td>
<td id="2173F761431100263271F988CAA0F986" box="[838,1166,1641,1663]" gridcol="2" gridrow="11" pageId="2" pageNumber="181">19</td>
<td id="2173F761431100263597F988CB52F986" box="[1184,1404,1641,1663]" gridcol="3" gridrow="11" pageId="2" pageNumber="181">
like
<taxonomicName id="1B92E7DC4311FFDB35FBF98BCB0BF987" box="[1228,1317,1641,1663]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="181" phylum="Chordata" rank="species" species="leptus">
<emphasis id="EEE6404D4311FFDB35FBF98BCB0BF987" box="[1228,1317,1641,1663]" italics="true" pageId="2" pageNumber="181">A. leptus</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="62A29E1D4311002631ACF973CB52F951" box="[155,1404,1682,1704]" gridrow="12" pageId="2" pageNumber="181">
<th id="2173F7614311002631ACF973CFF1F951" box="[155,479,1682,1704]" gridcol="0" gridrow="12" pageId="2" pageNumber="181">Anal fin segmented rays</th>
<td id="2173F761431100263320F973CD09F951" box="[535,807,1682,1704]" gridcol="1" gridrow="12" pageId="2" pageNumber="181">14–21</td>
<td id="2173F761431100263271F973CAA0F951" box="[838,1166,1682,1704]" gridcol="2" gridrow="12" pageId="2" pageNumber="181">19</td>
<td id="2173F761431100263597F973CB52F951" box="[1184,1404,1682,1704]" gridcol="3" gridrow="12" pageId="2" pageNumber="181">
like
<taxonomicName id="1B92E7DC4311FFDB35FBF972CB0BF951" box="[1228,1317,1683,1704]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="181" phylum="Chordata" rank="species" species="leptus">
<emphasis id="EEE6404D4311FFDB35FBF972CB0BF951" box="[1228,1317,1683,1704]" italics="true" pageId="2" pageNumber="181">A. leptus</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="62A29E1D4311002631ACF95DCB52F92B" box="[155,1404,1724,1746]" gridrow="13" pageId="2" pageNumber="181">
<th id="2173F7614311002631ACF95DCFF1F92B" box="[155,479,1724,1746]" gridcol="0" gridrow="13" pageId="2" pageNumber="181">vertebrae (caudal /precaudal)</th>
<td id="2173F761431100263320F95DCD09F92B" box="[535,807,1724,1746]" gridcol="1" gridrow="13" pageId="2" pageNumber="181">11–13/19–25</td>
<td id="2173F761431100263271F95DCAA0F92B" box="[838,1166,1724,1746]" gridcol="2" gridrow="13" pageId="2" pageNumber="181">13/19</td>
<td id="2173F761431100263597F95DCB52F92B" box="[1184,1404,1724,1746]" gridcol="3" gridrow="13" pageId="2" pageNumber="181">12/23</td>
</tr>
<tr id="62A29E1D4311002631ACF904CB52F902" box="[155,1404,1765,1787]" gridrow="14" pageId="2" pageNumber="181">
<th id="2173F7614311002631ACF904CFF1F902" box="[155,479,1765,1787]" gridcol="0" gridrow="14" pageId="2" pageNumber="181">Pectoral fin rays</th>
<td id="2173F761431100263320F904CD09F902" box="[535,807,1765,1787]" gridcol="1" gridrow="14" pageId="2" pageNumber="181">13–16</td>
<td id="2173F761431100263271F904CAA0F902" box="[838,1166,1765,1787]" gridcol="2" gridrow="14" pageId="2" pageNumber="181">13</td>
<td id="2173F761431100263597F904CB52F902" box="[1184,1404,1765,1787]" gridcol="3" gridrow="14" pageId="2" pageNumber="181">
like
<taxonomicName id="1B92E7DC4311FFDB35FBF907CB0BF903" box="[1228,1317,1765,1787]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="181" phylum="Chordata" rank="species" species="leptus">
<emphasis id="EEE6404D4311FFDB35FBF907CB0BF903" box="[1228,1317,1765,1787]" italics="true" pageId="2" pageNumber="181">A. leptus</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="62A29E1D4311002631ACF8EFCB52F8DD" box="[155,1404,1806,1828]" gridrow="15" pageId="2" pageNumber="181">
<th id="2173F7614311002631ACF8EFCFF1F8DD" box="[155,479,1806,1828]" gridcol="0" gridrow="15" pageId="2" pageNumber="181">Premaxillary incisors in adults</th>
<td id="2173F761431100263320F8EFCD09F8DD" box="[535,807,1806,1828]" gridcol="1" gridrow="15" pageId="2" pageNumber="181">26–45</td>
<td id="2173F761431100263271F8EFCAA0F8DD" box="[838,1166,1806,1828]" gridcol="2" gridrow="15" pageId="2" pageNumber="181">26</td>
<td id="2173F761431100263597F8EFCB52F8DD" box="[1184,1404,1806,1828]" gridcol="3" gridrow="15" pageId="2" pageNumber="181">29–30</td>
</tr>
<tr id="62A29E1D4311002631ACF8D9CB52F8B7" box="[155,1404,1848,1870]" gridrow="16" pageId="2" pageNumber="181">
<th id="2173F7614311002631ACF8D9CFF1F8B7" box="[155,479,1848,1870]" gridcol="0" gridrow="16" pageId="2" pageNumber="181">Dentary incisors in adults</th>
<td id="2173F761431100263320F8D9CD09F8B7" box="[535,807,1848,1870]" gridcol="1" gridrow="16" pageId="2" pageNumber="181">28–48</td>
<td id="2173F761431100263271F8D9CAA0F8B7" box="[838,1166,1848,1870]" gridcol="2" gridrow="16" pageId="2" pageNumber="181">27</td>
<td id="2173F761431100263597F8D9CB52F8B7" box="[1184,1404,1848,1870]" gridcol="3" gridrow="16" pageId="2" pageNumber="181">26–30</td>
</tr>
</table>
</paragraph>
<paragraph id="DC2D9C5F4311FFDB31A0F88ECC05F851" blockId="2.[151,1436,1903,1960]" pageId="2" pageNumber="181">
1Cephalic pore terminology follows
<bibRefCitation id="B803E1AE4311FFDB3315F890CD12F871" author="Smith-Vaniz" box="[546,828,1905,1928]" pageId="2" pageNumber="181" refString="Smith-Vaniz, W. F. (1976) The saber-toothed blennies, tribe Nemophini (Pisces: Blenniidae). Academy of Natural Sciences of Philadelphia, Monograph, 16, 1 - 196." type="journal article" year="1976">Smith-Vaniz (1976, fig.1)</bibRefCitation>
; also see figs. 10–13 for cephalic pore configuration in representative species of
<emphasis id="EEE6404D4311FFDB3093F870CC05F851" box="[420,555,1937,1960]" italics="true" pageId="2" pageNumber="181">
<taxonomicName id="1B92E7DC4311FFDB3093F870CC09F851" box="[420,551,1937,1960]" class="Actinopterygii" family="Blenniidae" genus="Petroscirtes" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="181" phylum="Chordata" rank="genus">Petroscirtes</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<caption id="88EDCCD74310FFDA31A0FA46CBA7FA44" box="[151,1417,1447,1469]" httpUri="https://zenodo.org/record/569462/files/figure.png" pageId="3" pageNumber="182" targetBox="[192,1402,197,1427]" targetPageId="3">
<paragraph id="DC2D9C5F4310FFDA31A0FA46CBA7FA44" blockId="3.[151,1417,1447,1469]" box="[151,1417,1447,1469]" pageId="3" pageNumber="182">
<emphasis id="EEE6404D4310FFDA31A0FA46CF3CFA45" bold="true" box="[151,274,1447,1469]" pageId="3" pageNumber="182">FIGURE 1.</emphasis>
<taxonomicName id="1B92E7DC4310FFDA302EFA46CFDBFA45" box="[281,501,1447,1468]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="182" phylum="Chordata" rank="species" species="deloachi">
<emphasis id="EEE6404D4310FFDA302EFA46CFDBFA45" box="[281,501,1447,1468]" italics="true" pageId="3" pageNumber="182">Adelotremus deloachi</emphasis>
</taxonomicName>
holotype, ZMA 23004, male, 32.0 mm SL, Bali, Indonesia. Photograph by Ned DeLoach.
</paragraph>
</caption>
<subSubSection id="9488CFD44310FFDA31F0FA02CBB3FA04" box="[199,1437,1507,1533]" pageId="3" pageNumber="182" type="distribution">
<paragraph id="DC2D9C5F4310FFDA31F0FA02CBB3FA04" blockId="3.[151,1437,1507,2037]" box="[199,1437,1507,1533]" pageId="3" pageNumber="182">
<emphasis id="EEE6404D4310FFDA31F0FA02CF77FA05" bold="true" box="[199,345,1507,1532]" pageId="3" pageNumber="182">Distribution</emphasis>
. Known only from
<collectingRegion id="1E5652BD4310FFDA3302FA05CC4AFA04" box="[565,612,1508,1533]" country="Indonesia" name="Bali" pageId="3" pageNumber="182">Bali</collectingRegion>
and Lembeh Strait but undoubtedly more widely distributed in
<collectingCountry id="A485DCCF4310FFDA341FFA05CBB7FA04" box="[1320,1433,1508,1533]" name="Indonesia" pageId="3" pageNumber="182">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="9488CFD44310FFDA31F0F9E8CD3DF974" pageId="3" pageNumber="182" type="etymology">
<paragraph id="DC2D9C5F4310FFDA31F0F9E8CD3DF974" blockId="3.[151,1437,1507,2037]" pageId="3" pageNumber="182">
<emphasis id="EEE6404D4310FFDA31F0F9E8CF60F9D8" bold="true" box="[199,334,1545,1569]" pageId="3" pageNumber="182">Etymology.</emphasis>
This new species is named
<emphasis id="EEE6404D4310FFDA33A1F9E9CDA3F9D9" box="[662,909,1544,1568]" italics="true" pageId="3" pageNumber="182">
Aledotremus
<taxonomicName id="1B92E7DC4310FFDA321AF9E9CDA3F9D9" box="[813,909,1544,1568]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="182" phylum="Chordata" rank="species" species="deloachi">deloachi</taxonomicName>
</emphasis>
in honor of Ned DeLoach in recognition and appreciation of his books, magazine articles and photographs that celebrate the beauty and diversity of reef fishes, all of which have encouraged numerous divers and fish watchers to become more aware of the importance of protecting the threatened marine environment and fauna.
</paragraph>
</subSubSection>
<subSubSection id="9488CFD44310FFDE31F0F978CDE2FE9D" lastPageId="7" lastPageNumber="186" pageId="3" pageNumber="182" type="discussion">
<paragraph id="DC2D9C5F4310FFDF31F0F978CA8FFA7B" blockId="3.[151,1437,1507,2037]" lastBlockId="6.[151,1437,1277,2022]" lastPageId="6" lastPageNumber="185" pageId="3" pageNumber="182">
<emphasis id="EEE6404D4310FFDA31F0F978CF15F948" bold="true" box="[199,315,1689,1713]" pageId="3" pageNumber="182">Remarks.</emphasis>
Discovery of a second new species of
<taxonomicName id="1B92E7DC4310FFDA33C3F979CDAFF949" box="[756,897,1688,1712]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="182" phylum="Chordata" rank="genus">
<emphasis id="EEE6404D4310FFDA33C3F979CDAFF949" box="[756,897,1688,1712]" italics="true" pageId="3" pageNumber="182">Adelotremus</emphasis>
</taxonomicName>
from
<collectingCountry id="A485DCCF4310FFDA32F1F979CA1DF948" box="[966,1075,1688,1713]" name="Indonesia" pageId="3" pageNumber="182">Indonesia</collectingCountry>
began while Ned DeLoach was conducting underwater fish photography in
<date id="A82CBA9F4310FFDA33BCF95DCD29F92C" box="[651,775,1724,1749]" pageId="3" pageNumber="182" value="2012-04">April 2012</date>
at the Seraya dive resort near Tulamben,
<collectingRegion id="1E5652BD4310FFDA35DDF95DCB35F92C" box="[1258,1307,1724,1749]" country="Indonesia" name="Bali" pageId="3" pageNumber="182">Bali</collectingRegion>
,
<collectingCountry id="A485DCCF4310FFDA341FF95DCBB7F92C" box="[1320,1433,1724,1749]" name="Indonesia" pageId="3" pageNumber="182">Indonesia</collectingCountry>
. Ned was heading down a slope at a site known as Big Tree, located less than five minutes west of the resort, when he noticed a small pencil-thin blenny, with its beautiful dorsal fin fully spread, sticking halfway out of a hole on the sandy substrata in
<quantity id="1B6A31BA4310FFDA305CF8C9CF8BF8B9" box="[363,421,1832,1857]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="182" unit="m" value="16.0">16 m</quantity>
(
<quantity id="1B6A31BA4310FFDA3082F8C9CFDCF8B8" box="[437,498,1832,1857]" metricMagnitude="1" metricUnit="m" metricValue="1.58496" pageId="3" pageNumber="182" unit="ft" value="52.0">52 ft</quantity>
.). Because the fish was so close when first seen, it spooked and disappeared down into its hole. Ned patiently waited for 40 minutes about one meter from the hole before the small blenny reappeared far enough out of the hole to get a good photograph (
<figureCitation id="44A980DA4310FFDA3231F891CD64F870" box="[774,842,1904,1929]" captionStart="FIGURE 1" captionStartId="3.[151,250,1447,1469]" captionTargetBox="[192,1402,197,1427]" captionTargetId="figure@3.[192,1402,197,1427]" captionTargetPageId="3" captionText="FIGURE 1. Adelotremus deloachi holotype, ZMA 23004, male, 32.0 mm SL, Bali, Indonesia. Photograph by Ned DeLoach." httpUri="https://zenodo.org/record/569462/files/figure.png" pageId="3" pageNumber="182">Fig. 1</figureCitation>
). Realizing that the fish was unusual, Ned sent the photographic image to several ichthyologists but none of them could identify the blenny. Ned mentioned that the dive guides had returned to check on him several times because he had remained for such a long time at the same site while trying to photograph the blenny, and therefore they would probably remember the exact location. Mark Erdmann, who then lived in
<collectingRegion id="1E5652BD4315FFDF30EDFB1CCC25FAEF" box="[474,523,1277,1302]" country="Indonesia" name="Bali" pageId="6" pageNumber="185">Bali</collectingRegion>
, kindly visited the dive resort at my request during the same month of the initial discovery to try to collect the mystery blenny. With the help of the original dive guides, Mark was able to find and collect with clove oil what was probably the same individual. The dive guides said they believed they had seen the same species only a few other times at different sites and always in the same kind of habitat.
</paragraph>
<caption id="88EDCCD74317FFDD31A0FA5ECF02FA0D" httpUri="https://zenodo.org/record/569463/files/figure.png" pageId="4" pageNumber="183" targetBox="[217,1369,193,1450]" targetPageId="4">
<paragraph id="DC2D9C5F4317FFDD31A0FA5ECF02FA0D" blockId="4.[151,1435,1471,1524]" pageId="4" pageNumber="183">
<emphasis id="EEE6404D4317FFDD31A0FA5ECF3AFA2D" bold="true" box="[151,276,1471,1493]" pageId="4" pageNumber="183">FIGURE 2.</emphasis>
<taxonomicName id="1B92E7DC4317FFDD302BFA21CFD4FA2C" box="[284,506,1472,1493]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="183" phylum="Chordata" rank="species" species="deloachi">
<emphasis id="EEE6404D4317FFDD302BFA21CFD4FA2C" box="[284,506,1472,1493]" italics="true" pageId="4" pageNumber="183">Adelotremus deloachi</emphasis>
</taxonomicName>
paratype, USNM 438965, female, 29.4 mm SL, Lembeh Strait, Indonesia. Photograph by Ned DeLoach.
</paragraph>
</caption>
<caption id="88EDCCD74317FFDD31A0F85ACABFF828" box="[151,1169,1979,2001]" httpUri="https://zenodo.org/record/569464/files/figure.png" pageId="4" pageNumber="183" targetBox="[159,1427,1574,1958]" targetPageId="4">
<paragraph id="DC2D9C5F4317FFDD31A0F85ACABFF828" blockId="4.[151,1169,1979,2001]" box="[151,1169,1979,2001]" pageId="4" pageNumber="183">
<emphasis id="EEE6404D4317FFDD31A0F85ACF3CF829" bold="true" box="[151,274,1979,2001]" pageId="4" pageNumber="183">FIGURE 3.</emphasis>
<taxonomicName id="1B92E7DC4317FFDD302EF85ACFDBF829" box="[281,501,1979,2000]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="183" phylum="Chordata" rank="species" species="deloachi">
<emphasis id="EEE6404D4317FFDD302EF85ACFDBF829" box="[281,501,1979,2000]" italics="true" pageId="4" pageNumber="183">Adelotremus deloachi</emphasis>
</taxonomicName>
holotype, (right side reversed). Photograph by Mark V. Erdmann.
</paragraph>
</caption>
<caption id="88EDCCD74316FFDC31A0F862CF33F80C" httpUri="https://zenodo.org/record/569465/files/figure.png" pageId="5" pageNumber="184" targetBox="[407,1188,208,1900]" targetPageId="5">
<paragraph id="DC2D9C5F4316FFDC31A0F862CF33F80C" blockId="5.[151,1436,1923,2037]" pageId="5" pageNumber="184">
<emphasis id="EEE6404D4316FFDC31A0F862CF3CF861" bold="true" box="[151,274,1923,1945]" pageId="5" pageNumber="184">FIGURE 4.</emphasis>
Cephalic sensory pores in species of
<taxonomicName id="1B92E7DC4316FFDC33BAF862CD22F861" box="[653,780,1923,1944]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="184" phylum="Chordata" rank="genus">
<emphasis id="EEE6404D4316FFDC33BAF862CD22F861" box="[653,780,1923,1944]" italics="true" pageId="5" pageNumber="184">Adelotremus</emphasis>
</taxonomicName>
: (above)
<taxonomicName id="1B92E7DC4316FFDC325BF865CDF1F861" box="[876,991,1923,1945]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="184" phylum="Chordata" rank="species" species="deloachi">
<emphasis id="EEE6404D4316FFDC325BF865CDF1F861" box="[876,991,1923,1945]" italics="true" pageId="5" pageNumber="184">A. deloachi</emphasis>
</taxonomicName>
, holotype, male, 32.0 mm SL, ZMA 23004, Bali, Indonesia; (below)
<taxonomicName id="1B92E7DC4316FFDC30A1F842CFDEF84E" box="[406,496,1954,1976]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="184" phylum="Chordata" rank="species" species="leptus">
<emphasis id="EEE6404D4316FFDC30A1F842CFDEF84E" box="[406,496,1954,1976]" italics="true" pageId="5" pageNumber="184">A. leptus</emphasis>
</taxonomicName>
, holotype, female, 34.5 mm SL, ZMA 23004, Red Sea, Egypt, modified from Smith-Vaniz and Rose (2009, fig. 5). Small arrows identify ventralmost supraorbital and dorsalmost preopercular pores in both lateral and dorsal views.
</paragraph>
</caption>
<caption id="88EDCCD74315FFDF31A0FB40CEF2FB2F" httpUri="https://zenodo.org/record/569466/files/figure.png" pageId="6" pageNumber="185" targetBox="[244,1335,179,1155]" targetPageId="6">
<paragraph id="DC2D9C5F4315FFDF31A0FB40CEF2FB2F" blockId="6.[151,1435,1185,1238]" pageId="6" pageNumber="185">
<emphasis id="EEE6404D4315FFDF31A0FB40CF3DFB4E" bold="true" box="[151,275,1185,1207]" pageId="6" pageNumber="185">FIGURE 5.</emphasis>
<taxonomicName id="1B92E7DC4315FFDF302DFB43CFF2FB4E" box="[282,476,1186,1207]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="185" phylum="Chordata" rank="species" species="leptus">
<emphasis id="EEE6404D4315FFDF302DFB43CFF2FB4E" box="[282,476,1186,1207]" italics="true" pageId="6" pageNumber="185">Adelotremus leptus</emphasis>
</taxonomicName>
, male, close-up of dorsal-fin ocellus, Red Sea, Marsa Abu Dabab, Egypt. Photograph by Bart Hazes.
</paragraph>
</caption>
<paragraph id="DC2D9C5F4315FFDF31F0FA6CCA29F9CF" blockId="6.[151,1437,1277,2022]" pageId="6" pageNumber="185">
Three individuals of the new species of
<taxonomicName id="1B92E7DC4315FFDF33A2FA6CCD0CFA5C" box="[661,802,1421,1445]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="185" phylum="Chordata" rank="genus">
<emphasis id="EEE6404D4315FFDF33A2FA6CCD0CFA5C" box="[661,802,1421,1445]" italics="true" pageId="6" pageNumber="185">Adelotremus</emphasis>
</taxonomicName>
were subsequently observed on
<date id="A82CBA9F4315FFDF35ADFA6CCB59FA5F" box="[1178,1399,1421,1446]" pageId="6" pageNumber="185" value="2015-11-16">16 November 2015</date>
by Ned and Anna Deloach at Lembeh Strait, Indonesia, on a black sand slope in
<quantity id="1B6A31BA4315FFDF352CFA53CAAAFA30" box="[1051,1156,1458,1482]" metricMagnitude="1" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="6" pageNumber="185" unit="m" value="11.0" valueMax="12.0" valueMin="10.0">10‒12 m</quantity>
. The following day Ned returned to the same site and was able to photograph several of the blennies and, with the help of dive guide Abdulrahman Tampilang, collected two specimens, one of each sex. A large zip-lock bag was used to capture the fish when they retreated inside hollow soda-like tubes sticking out of the sand.
</paragraph>
<paragraph id="DC2D9C5F4315FFDF31F0F9A3CCBDF867" blockId="6.[151,1437,1277,2022]" pageId="6" pageNumber="185">
<materialsCitation id="6CFA96024315FFDF31F0F9A3CD47F8F7" ID-GBIF-Occurrence="1499294130" collectingDate="2015-06" collectorName="Notes &amp; The &amp; Marsa el At &amp; Sharm el Sheikh" country="Egypt" latitude="25.338" location="Red Sea" longLatPrecision="74" longitude="34.739" pageId="6" pageNumber="185" specimenCount="2" stateProvince="Red Sea" typeStatus="holotype">
<emphasis id="EEE6404D4315FFDF31F0F9A3CC33F9A3" bold="true" box="[199,541,1601,1626]" pageId="6" pageNumber="185">
<collectorName id="7167F9894315FFDF31F0F9A3CF24F9A3" box="[199,266,1602,1626]" pageId="6" pageNumber="185">Notes</collectorName>
<collectedFrom id="6F8EDFA24315FFDF3025F9A3CC38F9A3" box="[274,534,1601,1626]" pageId="6" pageNumber="185">
on
<taxonomicName id="1B92E7DC4315FFDF3000F9A0CC38F9A3" box="[311,534,1601,1626]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="185" phylum="Chordata" rank="species" species="leptus">
<emphasis id="EEE6404D4315FFDF3000F9A0CC38F9A3" bold="true" box="[311,534,1601,1626]" italics="true" pageId="6" pageNumber="185">Adelotremus leptus</emphasis>
</taxonomicName>
</collectedFrom>
.
</emphasis>
<collectorName id="7167F9894315FFDF3312F9A0CC7FF9A3" box="[549,593,1601,1626]" pageId="6" pageNumber="185">The</collectorName>
original description of
<taxonomicName id="1B92E7DC4315FFDF326BF9A3CA1DF9A3" box="[860,1075,1602,1626]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="185" phylum="Chordata" rank="species" species="leptus">
<emphasis id="EEE6404D4315FFDF326BF9A3CA1DF9A3" box="[860,1075,1602,1626]" italics="true" pageId="6" pageNumber="185">Adelotremus leptus</emphasis>
</taxonomicName>
was based on the single female
<typeStatus id="032922FD4315FFDF31A0F984CED7F987" box="[151,249,1637,1662]" pageId="6" pageNumber="185" type="holotype">holotype</typeStatus>
collected from the
<collectingRegion id="1E5652BD4315FFDF30EBF984CC13F987" box="[476,573,1637,1662]" country="Egypt" name="Al Bahr al Ahmar" pageId="6" pageNumber="185">Red Sea</collectingRegion>
, at
<collectorName id="7167F9894315FFDF335FF987CCDFF987" box="[616,753,1637,1662]" pageId="6" pageNumber="185">Marsa el At</collectorName>
(
<geoCoordinate id="B9A6FA984315FFDF3235F984CDA4F987" box="[770,906,1637,1662]" direction="north" orientation="latitude" pageId="6" pageNumber="185" precision="15" value="27.910555">27°54'38&quot;N</geoCoordinate>
,
<geoCoordinate id="B9A6FA984315FFDF32A0F987CA37F987" box="[919,1049,1638,1662]" direction="east" orientation="longitude" pageId="6" pageNumber="185" precision="15" value="34.328888">34°19'44&quot;E</geoCoordinate>
) near
<collectorName id="7167F9894315FFDF3553F984CB0BF987" box="[1124,1317,1637,1662]" pageId="6" pageNumber="185">Sharm el Sheikh</collectorName>
,
<collectingCountry id="A485DCCF4315FFDF3405F987CB56F987" box="[1330,1400,1638,1662]" name="Egypt" pageId="6" pageNumber="185">Egypt</collectingCountry>
. In
<date id="A82CBA9F4315FFDF31A0F96BCF27F95B" box="[151,265,1673,1698]" pageId="6" pageNumber="185" value="2015-06">
<collectingDate id="B86843774315FFDF31A0F96BCF27F95B" box="[151,265,1673,1698]" pageId="6" pageNumber="185" value="2015-06">June 2015</collectingDate>
</date>
Bart Hazes discovered this blenny at another
<collectingRegion id="1E5652BD4315FFDF323EF968CD4BF95B" box="[777,869,1673,1698]" country="Egypt" name="Al Bahr al Ahmar" pageId="6" pageNumber="185">Red Sea</collectingRegion>
locality, Marsa Abu Dabab (
<geoCoordinate id="B9A6FA984315FFDF3590F968CB39F95B" box="[1191,1303,1673,1698]" direction="north" orientation="latitude" pageId="6" pageNumber="185" precision="55" value="25.338">25.338°N</geoCoordinate>
,
<geoCoordinate id="B9A6FA984315FFDF3415F96BCBBEF95B" box="[1314,1424,1674,1698]" direction="east" orientation="longitude" pageId="6" pageNumber="185" precision="55" value="34.739">34.739°E</geoCoordinate>
), in a large central area of silt, sand and seagrass, but no rubble. He noted that this locality &quot;is as close as you get to muck diving in the
<collectingRegion id="1E5652BD4315FFDF3041F930CFF7F913" box="[374,473,1745,1770]" country="Egypt" name="Al Bahr al Ahmar" pageId="6" pageNumber="185">Red Sea</collectingRegion>
.&quot; Only males were observed which, like
<taxonomicName id="1B92E7DC4315FFDF329EF933CA07F913" box="[937,1065,1746,1770]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="185" phylum="Chordata" rank="species" species="deloachi">
<emphasis id="EEE6404D4315FFDF329EF933CA07F913" box="[937,1065,1746,1770]" italics="true" pageId="6" pageNumber="185">A. deloachi</emphasis>
</taxonomicName>
, have a beautiful blue ocellus in the dorsal fin (
<figureCitation id="44A980DA4315FFDF300CF914CF8BF8F7" box="[315,421,1781,1806]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="6.[151,250,1185,1207]" captionStartId-1="7.[151,250,1109,1131]" captionTargetBox-0="[244,1335,179,1155]" captionTargetBox-1="[213,1376,411,1092]" captionTargetId-0="figure@6.[244,1335,179,1155]" captionTargetId-1="figure@7.[213,1376,411,1092]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 5. Adelotremus leptus, male, close-up of dorsal-fin ocellus, Red Sea, Marsa Abu Dabab, Egypt. Photograph by Bart Hazes." captionText-1="FIGURE 6. Adelotremus leptus, large male, with elongate outer caudal-fin rays, Red Sea, Mars Abua Dabab, Egypt. Photograph by Bart Hazes." httpUri-0="https://zenodo.org/record/569466/files/figure.png" httpUri-1="https://zenodo.org/record/569467/files/figure.png" pageId="6" pageNumber="185">Figs. 5‒6</figureCitation>
) which is absent in the female
<typeStatus id="032922FD4315FFDF3234F914CD47F8F7" box="[771,873,1781,1806]" pageId="6" pageNumber="185" type="holotype">holotype</typeStatus>
</materialsCitation>
. Based on these two photographs, the ocelli shape and coloration are essentially the same if not virtually identical to that in
<taxonomicName id="1B92E7DC4315FFDF32E2F8FBCA7BF8CB" box="[981,1109,1818,1842]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="185" phylum="Chordata" rank="species" species="deloachi">
<emphasis id="EEE6404D4315FFDF32E2F8FBCA7BF8CB" box="[981,1109,1818,1842]" italics="true" pageId="6" pageNumber="185">A. deloachi</emphasis>
</taxonomicName>
(see discussion of ocellus in above color description). Unlike the female holotype, large males of
<taxonomicName id="1B92E7DC4315FFDF329DF8DFCA23F8AC" box="[938,1037,1853,1877]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="185" phylum="Chordata" rank="species" species="leptus">
<emphasis id="EEE6404D4315FFDF329DF8DFCA23F8AC" box="[938,1037,1853,1877]" italics="true" pageId="6" pageNumber="185">A. leptus</emphasis>
</taxonomicName>
have cirri associated with some of the more posteriorly positioned mandibular pores and several of the preopercular pores, and also have outer caudalfin rays that are much more elongate (
<figureCitation id="44A980DA4315FFDF3376F864CCABF867" box="[577,645,1925,1950]" captionStart="FIGURE 6" captionStartId="7.[151,250,1109,1131]" captionTargetBox="[213,1376,411,1092]" captionTargetId="figure@7.[213,1376,411,1092]" captionTargetPageId="7" captionText="FIGURE 6. Adelotremus leptus, large male, with elongate outer caudal-fin rays, Red Sea, Mars Abua Dabab, Egypt. Photograph by Bart Hazes." httpUri="https://zenodo.org/record/569467/files/figure.png" pageId="6" pageNumber="185">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="DC2D9C5F4315FFDE31F0F848CDD7FF0E" blockId="6.[151,1437,1277,2022]" lastBlockId="7.[151,1437,151,357]" lastPageId="7" lastPageNumber="186" pageId="6" pageNumber="185">
Supratemporal (ST) pores were very difficult to observe and only one lateral temporal (LT) pore could be detected in the
<typeStatus id="032922FD4315FFDF3070F82CCF87F81F" box="[327,425,1997,2022]" pageId="6" pageNumber="185" type="holotype">holotype</typeStatus>
even with high magnification; however, after publication of the original description, reexamination of the
<typeStatus id="032922FD4314FFDE3059FF76CFFEFF49" box="[366,464,151,176]" pageId="7" pageNumber="186" type="holotype">holotype</typeStatus>
with a Zeiss Discovery
<date id="A82CBA9F4314FFDE33EAFF79CD23FF49" box="[733,781,152,176]" pageId="7" pageNumber="186" value="2012-05">V12</date>
stereomicroscope revealed a second LT pore that had been previously missed (
<figureCitation id="44A980DA4314FFDE3044FF5DCF97FF2D" box="[371,441,188,213]" captionStart="FIGURE 4" captionStartId="5.[151,250,1923,1945]" captionTargetBox="[407,1188,208,1900]" captionTargetId="figure@5.[384,1206,193,1909]" captionTargetPageId="5" captionText="FIGURE 4. Cephalic sensory pores in species of Adelotremus: (above) A. deloachi, holotype, male, 32.0 mm SL, ZMA 23004, Bali, Indonesia; (below) A. leptus, holotype, female, 34.5 mm SL, ZMA 23004, Red Sea, Egypt, modified from Smith-Vaniz and Rose (2009, fig. 5). Small arrows identify ventralmost supraorbital and dorsalmost preopercular pores in both lateral and dorsal views." httpUri="https://zenodo.org/record/569465/files/figure.png" pageId="7" pageNumber="186">Fig. 4</figureCitation>
). More specimens are required to determine if
<taxonomicName id="1B92E7DC4314FFDE32FAFF5DCA1EFF2D" box="[973,1072,188,212]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="186" phylum="Chordata" rank="species" species="leptus">
<emphasis id="EEE6404D4314FFDE32FAFF5DCA1EFF2D" box="[973,1072,188,212]" italics="true" pageId="7" pageNumber="186">A. leptus</emphasis>
</taxonomicName>
actually has only two LT pores. In contrast, three LT pores were present in all three specimens of
<taxonomicName id="1B92E7DC4314FFDE3244FF01CDDCFF01" box="[883,1010,224,248]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="186" phylum="Chordata" rank="species" species="deloachi">
<emphasis id="EEE6404D4314FFDE3244FF01CDDCFF01" box="[883,1010,224,248]" italics="true" pageId="7" pageNumber="186">A. deloachi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="DC2D9C5F4314FFDE31F0FEE5CDE2FE9D" blockId="7.[151,1437,151,357]" pageId="7" pageNumber="186">
The slender bodies of the two species of
<taxonomicName id="1B92E7DC4314FFDE33F4FEE5CD7EFEE5" box="[707,848,260,284]" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="186" phylum="Chordata" rank="genus">
<emphasis id="EEE6404D4314FFDE33F4FEE5CD7EFEE5" box="[707,848,260,284]" italics="true" pageId="7" pageNumber="186">Adelotremus</emphasis>
</taxonomicName>
are possibly an adaption for utilization of small abandoned worm holes or other protective shelter. Bart Hazes (pers. com.) reported that at Marsa Abu Dabab
<taxonomicName id="1B92E7DC4314FFDE34B3FEC9CEF4FE9D" class="Actinopterygii" family="Blenniidae" genus="Adelotremus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="186" phylum="Chordata" rank="species" species="leptus">
<emphasis id="EEE6404D4314FFDE34B3FEC9CEF4FE9D" italics="true" pageId="7" pageNumber="186">A. leptus</emphasis>
</taxonomicName>
occupied empty shells of the scaphopod mollusc genus
<taxonomicName id="1B92E7DC4314FFDE3278FEADCDEBFE9D" box="[847,965,332,356]" class="Scaphopoda" family="Dentaliidae" genus="Dentalium" kingdom="Animalia" order="Dentaliida" pageId="7" pageNumber="186" phylum="Mollusca" rank="genus">
<emphasis id="EEE6404D4314FFDE3278FEADCDEBFE9D" box="[847,965,332,356]" italics="true" pageId="7" pageNumber="186">Dentalium</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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