treatments-xml/data/2A/6E/8C/2A6E8C766E5D9F56FFE8FD86FE0866EE.xml

<document id="11C0023081717469410CF03FD5BDAACB" ID-CLB-Dataset="141500" ID-DOI="10.11646/zootaxa.5187.1.10" ID-GBIF-Dataset="82a205da-89e5-499c-bca0-21c0fc64bc11" ID-ISSN="1175-5326" ID-Zenodo-Dep="7077946" ID-ZooBank="435C5036-F986-41B0-AEC7-4408B673BC33" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1663143070382" checkinUser="plazi" docAuthor="Seniczak, Stanisław, Ivan, Otilia, Kaczmarek, Sławomir, Koziróg, Katarzyna Faleńczyk- &amp; Seniczak, Anna" docDate="2022" docId="2A6E8C766E5D9F56FFE8FD86FE0866EE" docLanguage="en" docName="zootaxa.5187.1.10.pdf" docOrigin="Zootaxa 5187 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Punctoribates ghilarovi Shaldybina 1969" docType="treatment" docVersion="7" lastPageNumber="157" masterDocId="D657F40E6E5F9F5EFF7FFFC0FFFA6743" masterDocTitle="Morphological ontogeny of Punctoribates ghilarovi (Acari, Oribatida, Punctoribatidae)" masterLastPageNumber="168" masterPageNumber="149" pageNumber="151" updateTime="1699380253751" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="C52404D1C1461E5B5FD25045248CE60D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="14D03A6ABEEB148B3FC84400C69CFF08">
<mods:title id="EBDCBA0EF20FEBCE99A161137BAB6BBF">Morphological ontogeny of Punctoribates ghilarovi (Acari, Oribatida, Punctoribatidae)</mods:title>
</mods:titleInfo>
<mods:name id="C7CB8D80DCF264644EFD670AC56294F5" type="personal">
<mods:role id="A115E5B4251F5B3C9185F7EE3F0DD4BB">
<mods:roleTerm id="ADBCC9775A7F8823B3E5406E5D9E5A45">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="222AFA64DFBF337E8B0EB34BEA41DE62">Seniczak, Stanisław</mods:namePart>
</mods:name>
<mods:name id="9AD6756DF77EF977F055F911DDD77314" type="personal">
<mods:role id="E77985A851B3C3D8A7569A46E020C016">
<mods:roleTerm id="86D2D92A95325EE4A4C9B7999184AD60">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4033690F496070EF759658BDF2A1D3EB">Ivan, Otilia</mods:namePart>
</mods:name>
<mods:name id="EE6E4F0A345A27924FD877F5B1140C3F" type="personal">
<mods:role id="BDC9D419BA35A0A929B1F580DD40199A">
<mods:roleTerm id="D5121D330A1BF0870C080B7090E8BFBD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A007DC04EE57AA07B99AAF8F5AE58899">Kaczmarek, Sławomir</mods:namePart>
</mods:name>
<mods:name id="B7995327E4B0F861AD038E62D8611E7A" type="personal">
<mods:role id="B2CCCE5CBC91ADF66299EF73C048666F">
<mods:roleTerm id="013968A958DD2677DD3CE06AC9E875F3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4E699B715B867E0E578379A543E31639">Koziróg, Katarzyna Faleńczyk-</mods:namePart>
</mods:name>
<mods:name id="D3999AED71246A7A1CE7EC49E4FFC683" type="personal">
<mods:role id="6BA0F58EAB32F64598C54F0F5C63166A">
<mods:roleTerm id="32FF452BD7BF797EC366CC416B56C016">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FC5102170D9D6E9D3761E89CDE9A21B0">Seniczak, Anna</mods:namePart>
</mods:name>
<mods:typeOfResource id="E7AEB177B4632D84640C181A91DF9496">text</mods:typeOfResource>
<mods:relatedItem id="F9BAB96C27DF29BE83C5BADCC3FEA820" type="host">
<mods:titleInfo id="A55C1EF8B6BB0C6C374FE9AA66EE7500">
<mods:title id="D96F52A72469E587E98E2EC3C2A31583">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="49D8FA6B06B7BB63695664FFB653CE3C">
<mods:date id="59E5582E4F86F29779257C721FBE99E9">2022</mods:date>
<mods:detail id="43C98604138F63C26EEDD11274849A9B" type="pubDate">
<mods:number id="D7AC4217272D77C3604E151965A96962">2022-09-14</mods:number>
</mods:detail>
<mods:detail id="729A6D977A984E61C1E48E1679C05D14" type="volume">
<mods:number id="9C47B3603710C682D28C88F1E66EA7F1">5187</mods:number>
</mods:detail>
<mods:detail id="1082041B7ED23A5EA5C37656963A9B2D" type="issue">
<mods:number id="D7ACB6FF59EFE5542275164133D476B2">1</mods:number>
</mods:detail>
<mods:extent id="77B82786CA413459D5F476A9484EEA9C" unit="page">
<mods:start id="356499BD3932B5120CE24E7FF5F2D738">149</mods:start>
<mods:end id="D3C607F37205F3223E416B7C9DAEE79E">168</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="3E533DF323C8563877067E7FC9415A17">journal article</mods:classification>
<mods:identifier id="0FFE4F21763CFEDE1EC38F6A089EA8FB" type="CLB-Dataset">141500</mods:identifier>
<mods:identifier id="F81E06099543B325206CC6BD0D39CDA8" type="DOI">10.11646/zootaxa.5187.1.10</mods:identifier>
<mods:identifier id="941D654936A3279E3C2B086F322E86C6" type="GBIF-Dataset">82a205da-89e5-499c-bca0-21c0fc64bc11</mods:identifier>
<mods:identifier id="7340B2DD063E544A09D310B3CC6E61FF" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A001D04194FB1069FD7360AB127848CC" type="Zenodo-Dep">7077946</mods:identifier>
<mods:identifier id="38B6D2EBDF07E35191F44E472731F715" type="ZooBank">435C5036-F986-41B0-AEC7-4408B673BC33</mods:identifier>
</mods:mods>
<treatment id="2A6E8C766E5D9F56FFE8FD86FE0866EE" ID-DOI="http://doi.org/10.5281/zenodo.7077948" ID-GBIF-Taxon="200289895" ID-Zenodo-Dep="7077948" LSID="urn:lsid:plazi:treatment:2A6E8C766E5D9F56FFE8FD86FE0866EE" httpUri="http://treatment.plazi.org/id/2A6E8C766E5D9F56FFE8FD86FE0866EE" lastPageId="8" lastPageNumber="157" pageId="2" pageNumber="151">
<subSubSection id="EADD6EEB6E5D9F5CFFE8FD86FD686523" box="[151,658,582,609]" pageId="2" pageNumber="151" type="nomenclature">
<paragraph id="A2783D606E5D9F5CFFE8FD86FD686523" blockId="2.[151,658,582,645]" box="[151,658,582,609]" pageId="2" pageNumber="151">
<heading id="F9308A0C6E5D9F5CFFE8FD86FD686523" bold="true" box="[151,658,582,609]" fontSize="11" level="1" pageId="2" pageNumber="151" reason="1">
<taxonomicName id="65C746E36E5D9F5CFFE8FD86FD686523" ID-CoL="7WPDB" authority="Shaldybina, 1969" authorityName="Shaldybina" authorityYear="1969" box="[151,658,582,609]" class="Arachnida" family="Punctoribatidae" genus="Punctoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="151" phylum="Arthropoda" rank="species" species="ghilarovi">
<emphasis id="90B3E1726E5D9F5CFFE8FD86FD686523" bold="true" box="[151,658,582,609]" pageId="2" pageNumber="151">
<emphasis id="90B3E1726E5D9F5CFFE8FD86FE496522" bold="true" box="[151,435,582,609]" italics="true" pageId="2" pageNumber="151">Punctoribates ghilarovi</emphasis>
<bibRefCitation id="C65640916E5D9F5CFEC4FD86FD686523" author="Shaldybina, E. S." box="[443,658,582,608]" pageId="2" pageNumber="151" pagination="53 - 68" refId="ref7718" refString="Shaldybina, E. S. (1969) New species of beetle mites of the genus Punctoribates Berlese, 1908 of the Gorky Region. Notes of the Gorky Pedagogical Institute, Biological series, 99, 53 - 68. [in Russian]" type="journal article" year="1969">Shaldybina, 1969</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="EADD6EEB6E5D9F59FFE8FDABFDEB614B" lastPageId="7" lastPageNumber="156" pageId="2" pageNumber="151" type="description">
<paragraph id="A2783D606E5D9F5CFFE8FDABFEDA65C6" blockId="2.[151,658,582,645]" box="[151,288,618,645]" pageId="2" pageNumber="151">
(
<figureCitation id="3AFC21E56E5D9F5CFFE0FDABFEED65C6" box="[159,279,618,645]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-10="FIGURE 11" captionStart-11="FIGURE 12" captionStart-12="FIGURE 13" captionStart-13="FIGURE 14" captionStart-14="FIGURE 15" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStart-8="FIGURE 9" captionStart-9="FIGURE 10" captionStartId-0="3.[151,250,1902,1926]" captionStartId-1="4.[151,250,1867,1891]" captionStartId-10="13.[151,250,1058,1082]" captionStartId-11="13.[151,250,1952,1976]" captionStartId-12="14.[151,250,1981,2005]" captionStartId-13="15.[151,250,1979,2003]" captionStartId-14="16.[151,250,1785,1809]" captionStartId-2="5.[151,250,1636,1660]" captionStartId-3="6.[151,250,1735,1759]" captionStartId-4="7.[151,250,1411,1435]" captionStartId-5="8.[151,250,1385,1409]" captionStartId-6="9.[151,250,1394,1418]" captionStartId-7="10.[151,250,1867,1891]" captionStartId-8="11.[151,250,1930,1954]" captionStartId-9="12.[151,250,1937,1961]" captionTargetBox-0="[219,1372,202,1874]" captionTargetBox-1="[228,1358,185,1844]" captionTargetBox-10="[286,1283,182,1035]" captionTargetBox-11="[292,1295,1108,1929]" captionTargetBox-12="[263,1314,197,1958]" captionTargetBox-13="[300,1282,189,1929]" captionTargetBox-14="[161,1427,181,1749]" captionTargetBox-2="[185,1404,181,1611]" captionTargetBox-3="[170,1416,777,1712]" captionTargetBox-4="[170,1416,453,1388]" captionTargetBox-5="[188,1399,453,1362]" captionTargetBox-6="[158,1429,417,1371]" captionTargetBox-7="[158,1429,445,1843]" captionTargetBox-8="[266,1318,182,1904]" captionTargetBox-9="[317,1272,186,1909]" captionTargetId-0="figure-19@3.[216,1372,198,1876]" captionTargetId-1="figure-32@4.[227,1359,181,1844]" captionTargetId-10="figure-19@13.[285,1302,181,1035]" captionTargetId-11="figure-40@13.[292,1295,1107,1929]" captionTargetId-12="figure-32@14.[261,1326,181,1958]" captionTargetId-13="figure-19@15.[300,1286,181,1955]" captionTargetId-14="figure-32@16.[161,1427,181,1762]" captionTargetId-2="figure-19@5.[181,1406,181,1612]" captionTargetId-3="figure-378@6.[170,1416,777,1712]" captionTargetId-4="figure-171@7.[170,1416,453,1388]" captionTargetId-5="figure-146@8.[188,1399,453,1362]" captionTargetId-6="figure-120@9.[158,1429,417,1371]" captionTargetId-7="figure-153@10.[158,1429,443,1843]" captionTargetId-8="figure-19@11.[265,1323,181,1907]" captionTargetId-9="figure-32@12.[315,1272,181,1913]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-10="13" captionTargetPageId-11="13" captionTargetPageId-12="14" captionTargetPageId-13="15" captionTargetPageId-14="16" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="10" captionTargetPageId-8="11" captionTargetPageId-9="12" captionText-0="FIGURE 1. Punctoribates ghilarovi, female, dorsal aspect, legs partially drawn, scale bar 50 μm." captionText-1="FIGURE 2. Punctoribates ghilarovi, female, ventral aspect, legs partially drawn, scale bar 50 μm." captionText-10="FIGURE 11. Punctoribates ghilarovi, lateral aspect, legs partially drawn, scale bars 20 μm. (a) Larva, (b) tritonymph." captionText-11="FIGURE 12. Punctoribates ghilarovi, leg segments of larva (part of femur to tarsus), right side, antiaxial aspect, seta on the opposite side not illustrated is indicated in the legend, scale bar 10 μm. (a) Leg I, tarsus (pl’); (b) leg II; (c) leg III." captionText-12="FIGURE 13. Punctoribates ghilarovi, ventral part of hysterosoma, legs partially drawn, scale bars 20 μm. (a) Deutonymph, (b) tritonymph." captionText-13="FIGURE 14. Punctoribates ghilarovi, tritonymph, dorsal aspect, legs partially drawn, scale bar 20 μm." captionText-14="FIGURE 15. Punctoribates ghilarovi, leg segments of tritonymph (part of femur to tarsus), right side, antiaxial aspect, seta on the opposite side not illustrated is indicated in the legend, scale bars 10 μm. (a) Leg I, tarsus (pl’); 
)
</paragraph>
<paragraph id="A2783D606E5D9F5CFFE8FD73FEF3658F" blockId="2.[151,1437,691,1041]" box="[151,265,691,716]" pageId="2" pageNumber="151">
<emphasis id="90B3E1726E5D9F5CFFE8FD73FEF3658F" box="[151,265,691,716]" italics="true" pageId="2" pageNumber="151">Diagnosis</emphasis>
</paragraph>
<paragraph id="A2783D606E5D9F5CFFB8FD17FC40641E" blockId="2.[151,1437,691,1041]" pageId="2" pageNumber="151">
Adult of medium size (325–421), with characters of 
<taxonomicName id="65C746E36E5D9F5CFC71FD17FC5165B2" authorityName="Berlese" authorityYear="1908" box="[782,939,727,753]" class="Arachnida" family="Punctoribatidae" genus="Punctoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="90B3E1726E5D9F5CFC71FD17FC5165B2" box="[782,939,727,753]" italics="true" pageId="2" pageNumber="151">Punctoribates</emphasis>
</taxonomicName>
given by 
<bibRefCitation id="C65640916E5D9F5CFB67FD17FA6265B2" author="Behan-Pelletier, V. M. &amp; Eamer, B." box="[1048,1432,727,753]" pageId="2" pageNumber="151" pagination="73 - 110" refId="ref6593" refString="Behan-Pelletier, V. M. &amp; Eamer, B. (2008) Mycobatidae (Acari, Oribatida) of North America. The Canadian Entomologist., 140, 73 - 110. https: // doi. org / 10.4039 / n 07 - 027" type="journal article" year="2008">Behan-Pelletier and Eamer (2008)</bibRefCitation>
. Bothridial seta clavate, anterior tectum of notogaster straight. Notogaster with seven pairs of microsetae, including 
<emphasis id="90B3E1726E5D9F5CFFE8FCDFFF59647B" box="[151,163,799,824]" italics="true" pageId="2" pageNumber="151">c</emphasis>
<subScript id="3E433F256E5D9F5CFFDBFCF1FF56647C" attach="right" box="[164,172,817,831]" fontSize="6" pageId="2" pageNumber="151">2</subScript>
, and three pairs of short 
<emphasis id="90B3E1726E5D9F5CFEB2FCDFFE21647B" box="[461,475,799,824]" italics="true" pageId="2" pageNumber="151">p</emphasis>
-series. Four pairs of porose areas present, 
<emphasis id="90B3E1726E5D9F5CFCB8FCDFFC1C647A" box="[967,998,799,825]" italics="true" pageId="2" pageNumber="151">Aa</emphasis>
clearly larger than other porose areas. Aggenital, adanal and anal setae short. All tibiae with dorsal projection.
</paragraph>
<paragraph id="A2783D606E5D9F5CFFB8FCA7FDD56352" blockId="2.[151,1437,691,1041]" pageId="2" pageNumber="151">
Juveniles light brown. Most prodorsal setae of medium size, except for long seta 
<emphasis id="90B3E1726E5D9F5CFB08FCA7FB7564C3" box="[1143,1167,871,896]" italics="true" pageId="2" pageNumber="151">ro</emphasis>
in larva, and short and smooth seta 
<emphasis id="90B3E1726E5D9F5CFE59FC4BFEC564E7" box="[294,319,907,932]" italics="true" pageId="2" pageNumber="151">ex</emphasis>
in all instars. The larva with 12 pairs of gastronotal setae, including 
<emphasis id="90B3E1726E5D9F5CFB2FFC4BFBA464E6" box="[1104,1118,907,933]" italics="true" pageId="2" pageNumber="151">h</emphasis>
<subScript id="3E433F256E5D9F5CFB21FC5DFB9C64E8" attach="left" box="[1118,1126,925,939]" fontSize="6" pageId="2" pageNumber="151">3</subScript>
, the nymphs with 15 pairs. Most these setae of medium size and barbed, except for slightly longer 
<emphasis id="90B3E1726E5D9F5CFCC8FC6FFC23648A" box="[951,985,943,969]" italics="true" pageId="2" pageNumber="151">dm</emphasis>
, 
<emphasis id="90B3E1726E5D9F5CFC98FC6FFBF9648A" box="[999,1027,943,969]" italics="true" pageId="2" pageNumber="151">dp</emphasis>
, 
<emphasis id="90B3E1726E5D9F5CFB6EFC6FFBD7648A" box="[1041,1069,943,969]" italics="true" pageId="2" pageNumber="151">lm</emphasis>
, 
<emphasis id="90B3E1726E5D9F5CFB44FC6FFBAB648A" box="[1083,1105,943,969]" italics="true" pageId="2" pageNumber="151">lp</emphasis>
and 
<emphasis id="90B3E1726E5D9F5CFBF8FC6FFB6F648A" box="[1159,1173,943,969]" italics="true" pageId="2" pageNumber="151">h</emphasis>
<subScript id="3E433F256E5D9F5CFBEAFC01FB67648C" attach="left" box="[1173,1181,961,975]" fontSize="6" pageId="2" pageNumber="151">1</subScript>
, and short and smooth 
<emphasis id="90B3E1726E5D9F5CFFE8FC13FF5F64AE" box="[151,165,979,1005]" italics="true" pageId="2" pageNumber="151">h</emphasis>
<quantity id="653F90856E5D9F5CFFDAFC25FF3164AE" box="[165,203,979,1011]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="2" pageNumber="151" unit="in" value="3.0">
<subScript id="3E433F256E5D9F5CFFDAFC25FF5764B0" attach="left" box="[165,173,997,1011]" fontSize="6" pageId="2" pageNumber="151">3</subScript>
in
</quantity>
larva, and also short and smooth 
<emphasis id="90B3E1726E5D9F5CFD39FC13FD9864AE" box="[582,610,979,1005]" italics="true" pageId="2" pageNumber="151">da</emphasis>
and 
<emphasis id="90B3E1726E5D9F5CFDE6FC13FD4164AE" box="[665,699,979,1005]" italics="true" pageId="2" pageNumber="151">dm</emphasis>
, slightly longer 
<emphasis id="90B3E1726E5D9F5CFC0BFC13FC6A64AE" box="[884,912,979,1005]" italics="true" pageId="2" pageNumber="151">dp</emphasis>
, and clearly longer 
<emphasis id="90B3E1726E5D9F5CFB0EFC13FB8564AE" box="[1137,1151,979,1005]" italics="true" pageId="2" pageNumber="151">h</emphasis>
<subScript id="3E433F256E5D9F5CFB00FC25FB7D64B0" attach="left" box="[1151,1159,997,1011]" fontSize="6" pageId="2" pageNumber="151">1</subScript>
, 
<emphasis id="90B3E1726E5D9F5CFBE9FC13FB5E64AE" box="[1174,1188,979,1005]" italics="true" pageId="2" pageNumber="151">h</emphasis>
<subScript id="3E433F256E5D9F5CFBDBFC25FB5664B0" attach="left" box="[1188,1196,997,1011]" fontSize="6" pageId="2" pageNumber="151">3</subScript>
and 
<emphasis id="90B3E1726E5D9F5CFB9CFC13FB0364AE" box="[1251,1273,979,1005]" italics="true" pageId="2" pageNumber="151">lp</emphasis>
in nymphs. In all juveniles, humeral organ present.
</paragraph>
<paragraph id="A2783D606E5D9F5CFFE8FBFFFE85631A" blockId="2.[151,1437,1087,1761]" box="[151,383,1087,1113]" pageId="2" pageNumber="151">
<emphasis id="90B3E1726E5D9F5CFFE8FBFFFE85631A" box="[151,383,1087,1113]" italics="true" pageId="2" pageNumber="151">Morphology of adult</emphasis>
</paragraph>
<paragraph id="A2783D606E5D9F5CFFB8FBA3FDCF62A6" blockId="2.[151,1437,1087,1761]" pageId="2" pageNumber="151">
Adults (
<figureCitation id="3AFC21E56E5D9F5CFE5AFBA3FE68633E" box="[293,402,1123,1149]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStartId-0="3.[151,250,1902,1926]" captionStartId-1="4.[151,250,1867,1891]" captionStartId-2="5.[151,250,1636,1660]" captionStartId-3="6.[151,250,1735,1759]" captionStartId-4="7.[151,250,1411,1435]" captionStartId-5="8.[151,250,1385,1409]" captionStartId-6="9.[151,250,1394,1418]" captionStartId-7="10.[151,250,1867,1891]" captionTargetBox-0="[219,1372,202,1874]" captionTargetBox-1="[228,1358,185,1844]" captionTargetBox-2="[185,1404,181,1611]" captionTargetBox-3="[170,1416,777,1712]" captionTargetBox-4="[170,1416,453,1388]" captionTargetBox-5="[188,1399,453,1362]" captionTargetBox-6="[158,1429,417,1371]" captionTargetBox-7="[158,1429,445,1843]" captionTargetId-0="figure-19@3.[216,1372,198,1876]" captionTargetId-1="figure-32@4.[227,1359,181,1844]" captionTargetId-2="figure-19@5.[181,1406,181,1612]" captionTargetId-3="figure-378@6.[170,1416,777,1712]" captionTargetId-4="figure-171@7.[170,1416,453,1388]" captionTargetId-5="figure-146@8.[188,1399,453,1362]" captionTargetId-6="figure-120@9.[158,1429,417,1371]" captionTargetId-7="figure-153@10.[158,1429,443,1843]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="10" captionText-0="FIGURE 1. Punctoribates ghilarovi, female, dorsal aspect, legs partially drawn, scale bar 50 μm." captionText-1="FIGURE 2. Punctoribates ghilarovi, female, ventral aspect, legs partially drawn, scale bar 50 μm." captionText-2="FIGURE 3. Punctoribates ghilarovi, adult. (a) Lateral aspect, legs partially drawn, scale bar 50 μm; mouthparts, right side, scale bars 10 μm; (b) chelicera (Trägårdh organ in transparent area); (c) palp." captionText-3="FIGURE 4. Punctoribates ghilarovi, adult, SEM micrographs. (a) Dorsal view, (b) lateral view, (c) ventral view, (d) posterior view." captionText-4="FIGURE 5. Punctoribates ghilarovi, adult, SEM micrographs. (a) Dorsal view, anterior and medial part, (b) medial and posterior part, (c) bothridial seta and legs I and II, (d) leg II with dorsal projection on tibia." captionText-5="FIGURE 6. Punctoribates ghilarovi, adult, SEM micrographs. Lateral view, (a), (b) anterior part; ventral view, (c) posterior and medial part, (d) anterior and medial part." captionText-6="FIGURE 7. Punctoribates ghilarovi, tibiae of adult with dorsal projection, SEM micrographs. (a) Leg I, medial view; lateral view, (b) leg III and IV, (c), (d) leg IV." captionText-7="FIGURE 8. Punctoribates ghilarovi, leg segments of adult (part of femur to tarsus), right side, antiaxial aspect, seta on the opposite side not illustrated is indicated in the legend, scale bar 20 μm. (a) Leg I, tarsus (pl’); (b) leg II; (c) leg III; (d) leg IV." figureDoi-0="http://doi.org/10.5281/zenodo.7077950" figureDoi-1="http://doi.org/10.5281/zenodo.7077952" figureDoi-2="http://doi.org/10.5281/zenodo.7077956" figureDoi-3="http://doi.org/10.5281/zenodo.7077958" figureDoi-4="http://doi.org/10.5281/zenodo.7077962" figureDoi-5="http://doi.org/10.5281/zenodo.7077964" figureDoi-6="http://doi.org/10.5281/zenodo.7077966" figureDoi-7="http://doi.org/10.5281/zenodo.7077970" httpUri-0="https://zenodo.org/record/7077950/files/figure.png" httpUri-1="https://zenodo.org/record/7077952/files/figure.png" httpUri-2="https://zenodo.org/record/7077956/files/figure.png" httpUri-3="https://zenodo.org/record/7077958/files/figure.png" httpUri-4="https://zenodo.org/record/7077962/files/figure.png" httpUri-5="https://zenodo.org/record/7077964/files/figure.png" httpUri-6="https://zenodo.org/record/7077966/files/figure.png" httpUri-7="https://zenodo.org/record/7077970/files/figure.png" pageId="2" pageNumber="151">Figs. 1–8</figureCitation>
) similar to those investigated by 
<bibRefCitation id="C65640916E5D9F5CFC64FBA3FC0A633E" author="Shaldybina, E. S." box="[795,1008,1123,1149]" pageId="2" pageNumber="151" pagination="689 - 699" refId="ref7764" refString="Shaldybina, E. S. (1973) New species of oribatid mites of the subfamily Minunthozetinae (Oribatei, Mycobatidae) from the territory of the Soviet Union. Zoologicheskii Zhurnal, 52 (5), 689 - 699. [in Russian]" type="journal article" year="1973">Shaldybina (1973)</bibRefCitation>
, but see 
<emphasis id="90B3E1726E5D9F5CFB23FBA3FB44633E" box="[1116,1214,1123,1149]" italics="true" pageId="2" pageNumber="151">Remarks</emphasis>
. Mean length (and range) of females 367.9±23.2 (337–421, n= 62) and males 345.1±17.8 (325–379, n= 28), mean maximum width (and range) of females 255.8±16.9 (235–289) and males 239.1±12.1 (223–259). Notogaster with seven pairs of microsetae, including 
<emphasis id="90B3E1726E5D9F5CFEF0FB0FFE6163AB" box="[399,411,1231,1256]" italics="true" pageId="2" pageNumber="151">c</emphasis>
<subScript id="3E433F256E5D9F5CFEE4FB21FE5963AC" attach="left" box="[411,419,1249,1263]" fontSize="6" pageId="2" pageNumber="151">2</subScript>
, and three pairs of short 
<emphasis id="90B3E1726E5D9F5CFDC6FB0FFD3D63AB" box="[697,711,1231,1256]" italics="true" pageId="2" pageNumber="151">p</emphasis>
-series (
<figureCitation id="3AFC21E56E5D9F5CFC60FB0FFC9663AA" box="[799,876,1231,1257]" captionStart="FIGURE 1" captionStartId="3.[151,250,1902,1926]" captionTargetBox="[219,1372,202,1874]" captionTargetId="figure-19@3.[216,1372,198,1876]" captionTargetPageId="3" captionText="FIGURE 1. Punctoribates ghilarovi, female, dorsal aspect, legs partially drawn, scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.7077950" httpUri="https://zenodo.org/record/7077950/files/figure.png" pageId="2" pageNumber="151">Figs. 1</figureCitation>
, 
<figureCitation id="3AFC21E56E5D9F5CFC05FB0FFC7063AA" box="[890,906,1231,1257]" captionStart="FIGURE 2" captionStartId="4.[151,250,1867,1891]" captionTargetBox="[228,1358,185,1844]" captionTargetId="figure-32@4.[227,1359,181,1844]" captionTargetPageId="4" captionText="FIGURE 2. Punctoribates ghilarovi, female, ventral aspect, legs partially drawn, scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.7077952" httpUri="https://zenodo.org/record/7077952/files/figure.png" pageId="2" pageNumber="151">2</figureCitation>
, 
<figureCitation id="3AFC21E56E5D9F5CFCE9FB0FFC4963AA" box="[918,947,1231,1257]" captionStart="FIGURE 3" captionStartId="5.[151,250,1636,1660]" captionTargetBox="[185,1404,181,1611]" captionTargetId="figure-19@5.[181,1406,181,1612]" captionTargetPageId="5" captionText="FIGURE 3. Punctoribates ghilarovi, adult. (a) Lateral aspect, legs partially drawn, scale bar 50 μm; mouthparts, right side, scale bars 10 μm; (b) chelicera (Trägårdh organ in transparent area); (c) palp." figureDoi="http://doi.org/10.5281/zenodo.7077956" httpUri="https://zenodo.org/record/7077956/files/figure.png" pageId="2" pageNumber="151">3a</figureCitation>
, 
<figureCitation id="3AFC21E56E5D9F5CFCC1FB0FFC3463AA" box="[958,974,1231,1257]" captionStart="FIGURE 4" captionStartId="6.[151,250,1735,1759]" captionTargetBox="[170,1416,777,1712]" captionTargetId="figure-378@6.[170,1416,777,1712]" captionTargetPageId="6" captionText="FIGURE 4. Punctoribates ghilarovi, adult, SEM micrographs. (a) Dorsal view, (b) lateral view, (c) ventral view, (d) posterior view." figureDoi="http://doi.org/10.5281/zenodo.7077958" httpUri="https://zenodo.org/record/7077958/files/figure.png" pageId="2" pageNumber="151">4</figureCitation>
, 
<figureCitation id="3AFC21E56E5D9F5CFCA5FB0EFBEB63AA" box="[986,1041,1230,1257]" captionStart="FIGURE 5" captionStartId="7.[151,250,1411,1435]" captionTargetBox="[170,1416,453,1388]" captionTargetId="figure-171@7.[170,1416,453,1388]" captionTargetPageId="7" captionText="FIGURE 5. Punctoribates ghilarovi, adult, SEM micrographs. (a) Dorsal view, anterior and medial part, (b) medial and posterior part, (c) bothridial seta and legs I and II, (d) leg II with dorsal projection on tibia." figureDoi="http://doi.org/10.5281/zenodo.7077962" httpUri="https://zenodo.org/record/7077962/files/figure.png" pageId="2" pageNumber="151">5a–c</figureCitation>
, 
<figureCitation id="3AFC21E56E5D9F5CFB63FB0FFBC063AA" box="[1052,1082,1231,1257]" captionStart="FIGURE 6" captionStartId="8.[151,250,1385,1409]" captionTargetBox="[188,1399,453,1362]" captionTargetId="figure-146@8.[188,1399,453,1362]" captionTargetPageId="8" captionText="FIGURE 6. Punctoribates ghilarovi, adult, SEM micrographs. Lateral view, (a), (b) anterior part; ventral view, (c) posterior and medial part, (d) anterior and medial part." figureDoi="http://doi.org/10.5281/zenodo.7077964" httpUri="https://zenodo.org/record/7077964/files/figure.png" pageId="2" pageNumber="151">6c</figureCitation>
). Pedotectum I strongly convex dorsally (
<figureCitation id="3AFC21E56E5D9F5CFE7EFB33FEA7624E" box="[257,349,1267,1293]" captionStart="FIGURE 3" captionStartId="5.[151,250,1636,1660]" captionTargetBox="[185,1404,181,1611]" captionTargetId="figure-19@5.[181,1406,181,1612]" captionTargetPageId="5" captionText="FIGURE 3. Punctoribates ghilarovi, adult. (a) Lateral aspect, legs partially drawn, scale bar 50 μm; mouthparts, right side, scale bars 10 μm; (b) chelicera (Trägårdh organ in transparent area); (c) palp." figureDoi="http://doi.org/10.5281/zenodo.7077956" httpUri="https://zenodo.org/record/7077956/files/figure.png" pageId="2" pageNumber="151">Figs. 3a</figureCitation>
, 
<figureCitation id="3AFC21E56E5D9F5CFE16FB33FE7D624E" box="[361,391,1267,1293]" captionStart="FIGURE 4" captionStartId="6.[151,250,1735,1759]" captionTargetBox="[170,1416,777,1712]" captionTargetId="figure-378@6.[170,1416,777,1712]" captionTargetPageId="6" captionText="FIGURE 4. Punctoribates ghilarovi, adult, SEM micrographs. (a) Dorsal view, (b) lateral view, (c) ventral view, (d) posterior view." figureDoi="http://doi.org/10.5281/zenodo.7077958" httpUri="https://zenodo.org/record/7077958/files/figure.png" pageId="2" pageNumber="151">4b</figureCitation>
, 
<figureCitation id="3AFC21E56E5D9F5CFEECFB33FE20624E" box="[403,474,1267,1293]" captionStart="FIGURE 6" captionStartId="8.[151,250,1385,1409]" captionTargetBox="[188,1399,453,1362]" captionTargetId="figure-146@8.[188,1399,453,1362]" captionTargetPageId="8" captionText="FIGURE 6. Punctoribates ghilarovi, adult, SEM micrographs. Lateral view, (a), (b) anterior part; ventral view, (c) posterior and medial part, (d) anterior and medial part." figureDoi="http://doi.org/10.5281/zenodo.7077964" httpUri="https://zenodo.org/record/7077964/files/figure.png" pageId="2" pageNumber="151">6a, 6b</figureCitation>
). Hypostomal setae 
<emphasis id="90B3E1726E5D9F5CFDC2FB33FD31624E" box="[701,715,1267,1293]" italics="true" pageId="2" pageNumber="151">h</emphasis>
, 
<emphasis id="90B3E1726E5D9F5CFDA6FB33FD17624F" box="[729,749,1267,1292]" italics="true" pageId="2" pageNumber="151">m</emphasis>
and 
<emphasis id="90B3E1726E5D9F5CFC5BFB33FCC8624F" box="[804,818,1267,1292]" italics="true" pageId="2" pageNumber="151">a</emphasis>
short and smooth (
<figureCitation id="3AFC21E56E5D9F5CFB75FB33FBA3624E" box="[1034,1113,1267,1293]" captionStart="FIGURE 2" captionStartId="4.[151,250,1867,1891]" captionTargetBox="[228,1358,185,1844]" captionTargetId="figure-32@4.[227,1359,181,1844]" captionTargetPageId="4" captionText="FIGURE 2. Punctoribates ghilarovi, female, ventral aspect, legs partially drawn, scale bar 50 μm." figureDoi="http://doi.org/10.5281/zenodo.7077952" httpUri="https://zenodo.org/record/7077952/files/figure.png" pageId="2" pageNumber="151">Figs. 2</figureCitation>
, 
<figureCitation id="3AFC21E56E5D9F5CFB1AFB33FB78624E" box="[1125,1154,1267,1293]" captionStart="FIGURE 4" captionStartId="6.[151,250,1735,1759]" captionTargetBox="[170,1416,777,1712]" captionTargetId="figure-378@6.[170,1416,777,1712]" captionTargetPageId="6" captionText="FIGURE 4. Punctoribates ghilarovi, adult, SEM micrographs. (a) Dorsal view, (b) lateral view, (c) ventral view, (d) posterior view." figureDoi="http://doi.org/10.5281/zenodo.7077958" httpUri="https://zenodo.org/record/7077958/files/figure.png" pageId="2" pageNumber="151">4c</figureCitation>
, 
<figureCitation id="3AFC21E56E5D9F5CFBF1FB33FB57624E" box="[1166,1197,1267,1293]" captionStart="FIGURE 6" captionStartId="8.[151,250,1385,1409]" captionTargetBox="[188,1399,453,1362]" captionTargetId="figure-146@8.[188,1399,453,1362]" captionTargetPageId="8" captionText="FIGURE 6. Punctoribates ghilarovi, adult, SEM micrographs. Lateral view, (a), (b) anterior part; ventral view, (c) posterior and medial part, (d) anterior and medial part." figureDoi="http://doi.org/10.5281/zenodo.7077964" httpUri="https://zenodo.org/record/7077964/files/figure.png" pageId="2" pageNumber="151">6d</figureCitation>
). Chelicera small (90 x 30), setae 
<emphasis id="90B3E1726E5D9F5CFE61FAD7FEBC6272" box="[286,326,1303,1329]" italics="true" pageId="2" pageNumber="151">cha</emphasis>
longer and thicker than 
<emphasis id="90B3E1726E5D9F5CFD28FAD7FD856272" box="[599,639,1303,1329]" italics="true" pageId="2" pageNumber="151">chb</emphasis>
, both barbed (
<figureCitation id="3AFC21E56E5D9F5CFC5DFAD7FC8F6272" box="[802,885,1303,1329]" captionStart="FIGURE 3" captionStartId="5.[151,250,1636,1660]" captionTargetBox="[185,1404,181,1611]" captionTargetId="figure-19@5.[181,1406,181,1612]" captionTargetPageId="5" captionText="FIGURE 3. Punctoribates ghilarovi, adult. (a) Lateral aspect, legs partially drawn, scale bar 50 μm; mouthparts, right side, scale bars 10 μm; (b) chelicera (Trägårdh organ in transparent area); (c) palp." figureDoi="http://doi.org/10.5281/zenodo.7077956" httpUri="https://zenodo.org/record/7077956/files/figure.png" pageId="2" pageNumber="151">Fig. 3b</figureCitation>
). Palp short (55), with most setae relatively long, 
<emphasis id="90B3E1726E5D9F5CFFE8FAFBFF446217" box="[151,190,1339,1364]" italics="true" pageId="2" pageNumber="151">sup</emphasis>
, 
<emphasis id="90B3E1726E5D9F5CFFB3FAFBFF106216" box="[204,234,1339,1365]" italics="true" pageId="2" pageNumber="151">inf</emphasis>
and 
<emphasis id="90B3E1726E5D9F5CFE5EFAFBFEC26216" box="[289,312,1339,1365]" italics="true" pageId="2" pageNumber="151">l”</emphasis>
on tibia finely barbed, other setae smooth (
<figureCitation id="3AFC21E56E5D9F5CFC5EFAFBFC886216" box="[801,882,1339,1365]" captionStart="FIGURE 3" captionStartId="5.[151,250,1636,1660]" captionTargetBox="[185,1404,181,1611]" captionTargetId="figure-19@5.[181,1406,181,1612]" captionTargetPageId="5" captionText="FIGURE 3. Punctoribates ghilarovi, adult. (a) Lateral aspect, legs partially drawn, scale bar 50 μm; mouthparts, right side, scale bars 10 μm; (b) chelicera (Trägårdh organ in transparent area); (c) palp." figureDoi="http://doi.org/10.5281/zenodo.7077956" httpUri="https://zenodo.org/record/7077956/files/figure.png" pageId="2" pageNumber="151">Fig. 3c</figureCitation>
), formula of palp setae 0-2-1-3-9(1). Trochanters III and IV and all femora flattened, porose areas on inner parts of trochanters III and IV and all femora, femur IV also with transverse folds, all tibiae with dorsal projections (
<figureCitation id="3AFC21E56E5D9F5CFC3DFA43FC5A62DE" box="[834,928,1411,1437]" captionStart="FIGURE 4" captionStartId="6.[151,250,1735,1759]" captionTargetBox="[170,1416,777,1712]" captionTargetId="figure-378@6.[170,1416,777,1712]" captionTargetPageId="6" captionText="FIGURE 4. Punctoribates ghilarovi, adult, SEM micrographs. (a) Dorsal view, (b) lateral view, (c) ventral view, (d) posterior view." figureDoi="http://doi.org/10.5281/zenodo.7077958" httpUri="https://zenodo.org/record/7077958/files/figure.png" pageId="2" pageNumber="151">Figs. 4b</figureCitation>
, 
<figureCitation id="3AFC21E56E5D9F5CFCD2FA42FC3162DE" box="[941,971,1410,1437]" captionStart="FIGURE 5" captionStartId="7.[151,250,1411,1435]" captionTargetBox="[170,1416,453,1388]" captionTargetId="figure-171@7.[170,1416,453,1388]" captionTargetPageId="7" captionText="FIGURE 5. Punctoribates ghilarovi, adult, SEM micrographs. (a) Dorsal view, anterior and medial part, (b) medial and posterior part, (c) bothridial seta and legs I and II, (d) leg II with dorsal projection on tibia." figureDoi="http://doi.org/10.5281/zenodo.7077962" httpUri="https://zenodo.org/record/7077962/files/figure.png" pageId="2" pageNumber="151">5d</figureCitation>
, 
<figureCitation id="3AFC21E56E5D9F5CFCA7FA43FBE562DE" box="[984,1055,1411,1437]" captionStart="FIGURE 6" captionStartId="8.[151,250,1385,1409]" captionTargetBox="[188,1399,453,1362]" captionTargetId="figure-146@8.[188,1399,453,1362]" captionTargetPageId="8" captionText="FIGURE 6. Punctoribates ghilarovi, adult, SEM micrographs. Lateral view, (a), (b) anterior part; ventral view, (c) posterior and medial part, (d) anterior and medial part." figureDoi="http://doi.org/10.5281/zenodo.7077964" httpUri="https://zenodo.org/record/7077964/files/figure.png" pageId="2" pageNumber="151">6a, 6b</figureCitation>
, 
<figureCitation id="3AFC21E56E5D9F5CFB53FA43FBC662DE" box="[1068,1084,1411,1437]" captionStart="FIGURE 7" captionStartId="9.[151,250,1394,1418]" captionTargetBox="[158,1429,417,1371]" captionTargetId="figure-120@9.[158,1429,417,1371]" captionTargetPageId="9" captionText="FIGURE 7. Punctoribates ghilarovi, tibiae of adult with dorsal projection, SEM micrographs. (a) Leg I, medial view; lateral view, (b) leg III and IV, (c), (d) leg IV." figureDoi="http://doi.org/10.5281/zenodo.7077966" httpUri="https://zenodo.org/record/7077966/files/figure.png" pageId="2" pageNumber="151">7</figureCitation>
, 
<figureCitation id="3AFC21E56E5D9F5CFB37FA43FBA262DE" box="[1096,1112,1411,1437]" captionStart="FIGURE 8" captionStartId="10.[151,250,1867,1891]" captionTargetBox="[158,1429,445,1843]" captionTargetId="figure-153@10.[158,1429,443,1843]" captionTargetPageId="10" captionText="FIGURE 8. Punctoribates ghilarovi, leg segments of adult (part of femur to tarsus), right side, antiaxial aspect, seta on the opposite side not illustrated is indicated in the legend, scale bar 20 μm. (a) Leg I, tarsus (pl’); (b) leg II; (c) leg III; (d) leg IV." figureDoi="http://doi.org/10.5281/zenodo.7077970" httpUri="https://zenodo.org/record/7077970/files/figure.png" pageId="2" pageNumber="151">8</figureCitation>
). Formulae of leg setae [trochanter to tarsus (+ solenidia)]: I – 1-5-3(1)-4(2)-20(2); II – 1-5-3(1)-4(1)-15(2); III – 2-3-1(1)-3(1)-15; IV – 1-2-2- 3(1)-12. Leg tarsi heterotridactylous.
</paragraph>
<paragraph id="A2783D606E5D9F5CFFB8FA2FFE6B61A2" blockId="2.[151,1437,1087,1761]" pageId="2" pageNumber="151">
<emphasis id="90B3E1726E5D9F5CFFB8FA2FFED3614A" box="[199,297,1519,1545]" italics="true" pageId="2" pageNumber="151">Remarks</emphasis>
. The range of body length of adults of 
<taxonomicName id="65C746E36E5D9F5CFDA1FA2FFC99614A" authorityName="Shaldybina" authorityYear="1969" box="[734,867,1519,1545]" class="Arachnida" family="Punctoribatidae" genus="Punctoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="151" phylum="Arthropoda" rank="species" species="ghilarovi">
<emphasis id="90B3E1726E5D9F5CFDA1FA2FFD15614B" box="[734,751,1519,1544]" italics="true" pageId="2" pageNumber="151">P</emphasis>
. 
<emphasis id="90B3E1726E5D9F5CFD82FA2FFC99614A" box="[765,867,1519,1545]" italics="true" pageId="2" pageNumber="151">ghilarovi</emphasis>
</taxonomicName>
investigated herein is slightly larger, and the range of width is slightly smaller than those investigated by 
<bibRefCitation id="C65640916E5D9F5CFD8EF9D3FC3A616E" author="Shaldybina, E. S." box="[753,960,1555,1581]" pageId="2" pageNumber="151" pagination="53 - 68" refId="ref7718" refString="Shaldybina, E. S. (1969) New species of beetle mites of the genus Punctoribates Berlese, 1908 of the Gorky Region. Notes of the Gorky Pedagogical Institute, Biological series, 99, 53 - 68. [in Russian]" type="journal article" year="1969">Shaldybina (1969)</bibRefCitation>
– length 370–430, width 274–344, sex not investigated, but the distribution of notogastral setae and porose areas are similar as in 
<bibRefCitation id="C65640916E5D9F5CFB2BF9F7FADC6112" author="Shaldybina, E. S." box="[1108,1318,1591,1617]" pageId="2" pageNumber="151" pagination="53 - 68" refId="ref7718" refString="Shaldybina, E. S. (1969) New species of beetle mites of the genus Punctoribates Berlese, 1908 of the Gorky Region. Notes of the Gorky Pedagogical Institute, Biological series, 99, 53 - 68. [in Russian]" type="journal article" year="1969">Shaldybina (1969)</bibRefCitation>
. However, in our some individuals studied herein, the porose area 
<emphasis id="90B3E1726E5D9F5CFC77F99BFCD26136" box="[776,808,1627,1653]" italics="true" pageId="2" pageNumber="151">A1</emphasis>
is larger than on other side, but smaller than the porose area 
<emphasis id="90B3E1726E5D9F5CFFB2F9BFFF1661DA" box="[205,236,1663,1689]" italics="true" pageId="2" pageNumber="151">Aa</emphasis>
. In our individuals, the shape of leg segments, setae and solenidia is similar to those investigated by 
<bibRefCitation id="C65640916E5D9F5CFA1EF9BFFEC261FE" author="Shaldybina, E. S." pageId="2" pageNumber="151" pagination="53 - 68" refId="ref7718" refString="Shaldybina, E. S. (1969) New species of beetle mites of the genus Punctoribates Berlese, 1908 of the Gorky Region. Notes of the Gorky Pedagogical Institute, Biological series, 99, 53 - 68. [in Russian]" type="journal article" year="1969">Shaldybina (1969)</bibRefCitation>
, except for anterodorsal projections on tibiae that are slightly smaller than in the adults investigated by 
<bibRefCitation id="C65640916E5D9F5CFFC5F907FE7761A2" author="Shaldybina, E. S." box="[186,397,1735,1761]" pageId="2" pageNumber="151" pagination="53 - 68" refId="ref7718" refString="Shaldybina, E. S. (1969) New species of beetle mites of the genus Punctoribates Berlese, 1908 of the Gorky Region. Notes of the Gorky Pedagogical Institute, Biological series, 99, 53 - 68. [in Russian]" type="journal article" year="1969">Shaldybina (1969)</bibRefCitation>
.
</paragraph>
<paragraph id="A2783D606E5D9F5CFFE8F8CFFE59606A" blockId="2.[151,1437,1807,2013]" box="[151,419,1807,1833]" pageId="2" pageNumber="151">
<emphasis id="90B3E1726E5D9F5CFFE8F8CFFE59606A" box="[151,419,1807,1833]" italics="true" pageId="2" pageNumber="151">Description of juveniles</emphasis>
</paragraph>
<paragraph id="A2783D606E5D9F5CFFB8F8F3FF2E609E" blockId="2.[151,1437,1807,2013]" pageId="2" pageNumber="151">
Larva oval in dorsal view (
<figureCitation id="3AFC21E56E5D9F5CFE83F8F3FDB8600E" box="[508,578,1843,1869]" captionStart="FIGURE 9" captionStartId="11.[151,250,1930,1954]" captionTargetBox="[266,1318,182,1904]" captionTargetId="figure-19@11.[265,1323,181,1907]" captionTargetPageId="11" captionText="FIGURE 9. Punctoribates ghilarovi, larva. (a) Dorsal aspect, legs partially drawn, scale bar 20 μm, (b) shape of seta dp (enlarged)." figureDoi="http://doi.org/10.5281/zenodo.7077972" httpUri="https://zenodo.org/record/7077972/files/figure.png" pageId="2" pageNumber="151">Fig. 9</figureCitation>
), light brown. Prodorsum subtriangular, prodorsal seta 
<emphasis id="90B3E1726E5D9F5CFBC8F8F3FB35600F" box="[1207,1231,1843,1868]" italics="true" pageId="2" pageNumber="151">ro</emphasis>
long, 
<emphasis id="90B3E1726E5D9F5CFA66F8F3FAD5600F" box="[1305,1327,1843,1868]" italics="true" pageId="2" pageNumber="151">in</emphasis>
and 
<emphasis id="90B3E1726E5D9F5CFA17F8F3FA86600E" box="[1384,1404,1843,1869]" italics="true" pageId="2" pageNumber="151">le</emphasis>
of medium size (
<tableCitation id="EF4508DB6E5D9F5CFE4BF897FE7D6032" box="[308,391,1879,1905]" captionStart="TABLE 1" captionStartId="1.[151,239,729,753]" captionTargetPageId="1" captionText="TABLE 1. Measurements of some morphological characters of juvenile stages and adult of Punctoribates ghilarovi (mean measurements of 2–5 specimens in μm); nd – not developed." httpUri="http://table.plazi.org/id/F6B86DE86E5E9F5FFFE8FD19FC906456" pageId="2" pageNumber="151" tableUuid="F6B86DE86E5E9F5FFFE8FD19FC906456">Table 1</tableCitation>
) and 
<emphasis id="90B3E1726E5D9F5CFEB6F897FE186033" box="[457,482,1879,1904]" italics="true" pageId="2" pageNumber="151">ex</emphasis>
short; most barbed, 
<emphasis id="90B3E1726E5D9F5CFDB7F897FD1B6033" box="[712,737,1879,1904]" italics="true" pageId="2" pageNumber="151">ex</emphasis>
smooth. Mutual distance between setal pair 
<emphasis id="90B3E1726E5D9F5CFBAAF897FB136032" box="[1237,1257,1879,1905]" italics="true" pageId="2" pageNumber="151">le</emphasis>
about 1.5 times longer than between pair 
<emphasis id="90B3E1726E5D9F5CFEC5F8BBFE2860D7" box="[442,466,1915,1940]" italics="true" pageId="2" pageNumber="151">ro</emphasis>
, between setal pair 
<emphasis id="90B3E1726E5D9F5CFDC8F8BBFD3760D7" box="[695,717,1915,1940]" italics="true" pageId="2" pageNumber="151">in</emphasis>
nearly three times longer than between pair 
<emphasis id="90B3E1726E5D9F5CFBB3F8BBFB1E60D7" box="[1228,1252,1915,1940]" italics="true" pageId="2" pageNumber="151">ro</emphasis>
, pair 
<emphasis id="90B3E1726E5D9F5CFA58F8BBFAC160D6" box="[1319,1339,1915,1941]" italics="true" pageId="2" pageNumber="151">le</emphasis>
inserted approximately midway between pairs 
<emphasis id="90B3E1726E5D9F5CFD36F85FFD9B60FB" box="[585,609,1951,1976]" italics="true" pageId="2" pageNumber="151">ro</emphasis>
and 
<emphasis id="90B3E1726E5D9F5CFDE5F85FFD4A60FB" box="[666,688,1951,1976]" italics="true" pageId="2" pageNumber="151">in</emphasis>
. Opening of bothridium oval, bothridial seta clavate, with barbed head.
</paragraph>
<caption id="F6B86DE86E5C9F5DFFE8F8AEFB8C60C4" ID-DOI="http://doi.org/10.5281/zenodo.7077950" ID-Zenodo-Dep="7077950" box="[151,1142,1902,1927]" httpUri="https://zenodo.org/record/7077950/files/figure.png" pageId="3" pageNumber="152" startId="3.[151,250,1902,1926]" targetBox="[219,1372,202,1874]" targetPageId="3">
<paragraph id="A2783D606E5C9F5DFFE8F8AEFB8C60C4" blockId="3.[151,1142,1902,1927]" box="[151,1142,1902,1927]" pageId="3" pageNumber="152">
<emphasis id="90B3E1726E5C9F5DFFE8F8AEFEE960C5" bold="true" box="[151,275,1902,1926]" pageId="3" pageNumber="152">FIGURE 1.</emphasis>
<taxonomicName id="65C746E36E5C9F5DFE66F8AEFDF060C5" authorityName="Shaldybina" authorityYear="1969" box="[281,522,1902,1926]" class="Arachnida" family="Punctoribatidae" genus="Punctoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="152" phylum="Arthropoda" rank="species" species="ghilarovi">
<emphasis id="90B3E1726E5C9F5DFE66F8AEFDF060C5" box="[281,522,1902,1926]" italics="true" pageId="3" pageNumber="152">Punctoribates ghilarovi</emphasis>
</taxonomicName>
, female, dorsal aspect, legs partially drawn, scale bar 50 μm.
</paragraph>
</caption>
<caption id="F6B86DE86E5B9F5AFFE8F88BFB846027" ID-DOI="http://doi.org/10.5281/zenodo.7077952" ID-Zenodo-Dep="7077952" box="[151,1150,1867,1892]" httpUri="https://zenodo.org/record/7077952/files/figure.png" pageId="4" pageNumber="153" startId="4.[151,250,1867,1891]" targetBox="[228,1358,185,1844]" targetPageId="4">
<paragraph id="A2783D606E5B9F5AFFE8F88BFB846027" blockId="4.[151,1150,1867,1892]" box="[151,1150,1867,1892]" pageId="4" pageNumber="153">
<emphasis id="90B3E1726E5B9F5AFFE8F88BFEE96027" bold="true" box="[151,275,1867,1892]" pageId="4" pageNumber="153">FIGURE 2.</emphasis>
<taxonomicName id="65C746E36E5B9F5AFE66F88CFDF06027" authorityName="Shaldybina" authorityYear="1969" box="[281,522,1868,1892]" class="Arachnida" family="Punctoribatidae" genus="Punctoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="153" phylum="Arthropoda" rank="species" species="ghilarovi">
<emphasis id="90B3E1726E5B9F5AFE66F88CFDF06027" box="[281,522,1868,1892]" italics="true" pageId="4" pageNumber="153">Punctoribates ghilarovi</emphasis>
</taxonomicName>
, female, ventral aspect, legs partially drawn, scale bar 50 μm.
</paragraph>
</caption>
<caption id="F6B86DE86E5A9F5BFFE8F9A4FC9661E3" ID-DOI="http://doi.org/10.5281/zenodo.7077956" ID-Zenodo-Dep="7077956" httpUri="https://zenodo.org/record/7077956/files/figure.png" pageId="5" pageNumber="154" startId="5.[151,250,1636,1660]" targetBox="[185,1404,181,1611]" targetPageId="5">
<paragraph id="A2783D606E5A9F5BFFE8F9A4FC9661E3" blockId="5.[151,1437,1636,1696]" pageId="5" pageNumber="154">
<emphasis id="90B3E1726E5A9F5BFFE8F9A4FEE8613F" bold="true" box="[151,274,1636,1660]" pageId="5" pageNumber="154">FIGURE 3.</emphasis>
<taxonomicName id="65C746E36E5A9F5BFE68F9A4FDFC613F" authorityName="Shaldybina" authorityYear="1969" box="[279,518,1636,1660]" class="Arachnida" family="Punctoribatidae" genus="Punctoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="154" phylum="Arthropoda" rank="species" species="ghilarovi">
<emphasis id="90B3E1726E5A9F5BFE68F9A4FDFC613F" box="[279,518,1636,1660]" italics="true" pageId="5" pageNumber="154">Punctoribates ghilarovi</emphasis>
</taxonomicName>
, adult. (a) Lateral aspect, legs partially drawn, scale bar 50 μm; mouthparts, right side, scale bars 10 μm; (b) chelicera (Trägårdh organ in transparent area); (c) palp.
</paragraph>
</caption>
<paragraph id="A2783D606E5A9F5BFFB8F90EFB7660A7" blockId="5.[151,1437,1742,2021]" pageId="5" pageNumber="154">
Gastronotum of larva with 12 pairs of setae, including 
<emphasis id="90B3E1726E5A9F5BFC41F90EFCB661AB" box="[830,844,1742,1768]" italics="true" pageId="5" pageNumber="154">h</emphasis>
<subScript id="3E433F256E5A9F5BFC33F921FCAE61AC" attach="left" box="[844,852,1761,1775]" fontSize="6" pageId="5" pageNumber="154">3</subScript>
inserted lateral to medial part of anal valves (
<figureCitation id="3AFC21E56E5A9F5BFA1BF90EFF5D604F" captionStart="FIGURE 9" captionStartId="11.[151,250,1930,1954]" captionTargetBox="[266,1318,182,1904]" captionTargetId="figure-19@11.[265,1323,181,1907]" captionTargetPageId="11" captionText="FIGURE 9. Punctoribates ghilarovi, larva. (a) Dorsal aspect, legs partially drawn, scale bar 20 μm, (b) shape of seta dp (enlarged)." figureDoi="http://doi.org/10.5281/zenodo.7077972" httpUri="https://zenodo.org/record/7077972/files/figure.png" pageId="5" pageNumber="154">Figs. 9</figureCitation>
, 
<figureCitation id="3AFC21E56E5A9F5BFFCBF932FF25604F" box="[180,223,1778,1804]" captionStart="FIGURE 10" captionStartId="12.[151,250,1937,1961]" captionTargetBox="[317,1272,186,1909]" captionTargetId="figure-32@12.[315,1272,181,1913]" captionTargetPageId="12" captionText="FIGURE 10. Punctoribates ghilarovi, ventral part of hysterosoma, legs partially drawn, scale bars 20 μm. (a) Larva, (b) prot- onymph." figureDoi="http://doi.org/10.5281/zenodo.7077974" httpUri="https://zenodo.org/record/7077974/files/figure.png" pageId="5" pageNumber="154">10a</figureCitation>
, 
<figureCitation id="3AFC21E56E5A9F5BFF94F932FEEE604F" box="[235,276,1778,1804]" captionStart="FIGURE 11" captionStartId="13.[151,250,1058,1082]" captionTargetBox="[286,1283,182,1035]" captionTargetId="figure-19@13.[285,1302,181,1035]" captionTargetPageId="13" captionText="FIGURE 11. Punctoribates ghilarovi, lateral aspect, legs partially drawn, scale bars 20 μm. (a) Larva, (b) tritonymph." figureDoi="http://doi.org/10.5281/zenodo.7077976" httpUri="https://zenodo.org/record/7077976/files/figure.png" pageId="5" pageNumber="154">11a</figureCitation>
), most setae of medium size and barbed, except for slightly longer 
<emphasis id="90B3E1726E5A9F5BFB73F932FBD4604F" box="[1036,1070,1778,1804]" italics="true" pageId="5" pageNumber="154">dm</emphasis>
, 
<emphasis id="90B3E1726E5A9F5BFB42F932FBA3604F" box="[1085,1113,1778,1804]" italics="true" pageId="5" pageNumber="154">dp</emphasis>
, 
<emphasis id="90B3E1726E5A9F5BFB18F932FB79604F" box="[1127,1155,1778,1804]" italics="true" pageId="5" pageNumber="154">lm</emphasis>
, 
<emphasis id="90B3E1726E5A9F5BFBEDF932FB52604F" box="[1170,1192,1778,1804]" italics="true" pageId="5" pageNumber="154">lp</emphasis>
and 
<emphasis id="90B3E1726E5A9F5BFBA0F932FB17604F" box="[1247,1261,1778,1804]" italics="true" pageId="5" pageNumber="154">h</emphasis>
<subScript id="3E433F256E5A9F5BFB92F8C5FB0F6050" attach="left" box="[1261,1269,1797,1811]" fontSize="6" pageId="5" pageNumber="154">1</subScript>
, and short and smooth 
<emphasis id="90B3E1726E5A9F5BFF8CF8D6FEFB6073" box="[243,257,1814,1840]" italics="true" pageId="5" pageNumber="154">h</emphasis>
<subScript id="3E433F256E5A9F5BFE7EF8E9FEF36074" attach="left" box="[257,265,1833,1847]" fontSize="6" pageId="5" pageNumber="154">3</subScript>
. Pygidial shield poorly developed, with setae of 
<emphasis id="90B3E1726E5A9F5BFC3EF8D6FCB56073" box="[833,847,1814,1840]" italics="true" pageId="5" pageNumber="154">d</emphasis>
-series and 
<emphasis id="90B3E1726E5A9F5BFCABF8D6FC186073" box="[980,994,1814,1840]" italics="true" pageId="5" pageNumber="154">h</emphasis>
<subScript id="3E433F256E5A9F5BFC9DF8E9FC106074" attach="left" box="[994,1002,1833,1847]" fontSize="6" pageId="5" pageNumber="154">1</subScript>
, other setae inserted on unsclerotized integument. Seta 
<emphasis id="90B3E1726E5A9F5BFE24F8FBFE9D6017" box="[347,359,1851,1876]" italics="true" pageId="5" pageNumber="154">c</emphasis>
<subScript id="3E433F256E5A9F5BFE17F88DFE8A6018" attach="left" box="[360,368,1869,1883]" fontSize="6" pageId="5" pageNumber="154">3</subScript>
longer than 
<emphasis id="90B3E1726E5A9F5BFE82F8FBFDF36017" box="[509,521,1851,1876]" italics="true" pageId="5" pageNumber="154">c</emphasis>
<subScript id="3E433F256E5A9F5BFD75F88DFDE86018" attach="left" box="[522,530,1869,1883]" fontSize="6" pageId="5" pageNumber="154">1</subScript>
and 
<emphasis id="90B3E1726E5A9F5BFD36F8FBFDAF6017" box="[585,597,1851,1876]" italics="true" pageId="5" pageNumber="154">c</emphasis>
<subScript id="3E433F256E5A9F5BFD29F88DFDA46018" attach="right" box="[598,606,1869,1883]" fontSize="6" pageId="5" pageNumber="154">2</subScript>
, length of setae of 
<emphasis id="90B3E1726E5A9F5BFC49F8FAFCBE6017" box="[822,836,1850,1876]" italics="true" pageId="5" pageNumber="154">d</emphasis>
- and 
<emphasis id="90B3E1726E5A9F5BFCFAF8FAFC776017" box="[901,909,1850,1876]" italics="true" pageId="5" pageNumber="154">l</emphasis>
-series slightly increasing from anterior to posterior, and setae of 
<emphasis id="90B3E1726E5A9F5BFE0BF89EFE78603B" box="[372,386,1886,1912]" italics="true" pageId="5" pageNumber="154">h</emphasis>
-series increasing from posterior to anterior (
<tableCitation id="EF4508DB6E5A9F5BFC00F89EFC2F603B" box="[895,981,1886,1912]" captionStart="TABLE 1" captionStartId="1.[151,239,729,753]" captionTargetPageId="1" captionText="TABLE 1. Measurements of some morphological characters of juvenile stages and adult of Punctoribates ghilarovi (mean measurements of 2–5 specimens in μm); nd – not developed." httpUri="http://table.plazi.org/id/F6B86DE86E5E9F5FFFE8FD19FC906456" pageId="5" pageNumber="154" tableUuid="F6B86DE86E5E9F5FFFE8FD19FC906456">Table 1</tableCitation>
). Cupules 
<emphasis id="90B3E1726E5A9F5BFB2CF89FFB93603B" box="[1107,1129,1887,1912]" italics="true" pageId="5" pageNumber="154">ia</emphasis>
posterior to seta 
<emphasis id="90B3E1726E5A9F5BFA4FF89FFAC6603B" box="[1328,1340,1887,1912]" italics="true" pageId="5" pageNumber="154">c</emphasis>
<subScript id="3E433F256E5A9F5BFA43F8B1FABE603C" attach="left" box="[1340,1348,1905,1919]" fontSize="6" pageId="5" pageNumber="154">3</subScript>
, 
<emphasis id="90B3E1726E5A9F5BFA2BF89FFA8A603B" box="[1364,1392,1887,1912]" italics="true" pageId="5" pageNumber="154">im</emphasis>
not observed, 
<emphasis id="90B3E1726E5A9F5BFE72F843FED960DF" box="[269,291,1923,1948]" italics="true" pageId="5" pageNumber="154">ip</emphasis>
between setae 
<emphasis id="90B3E1726E5A9F5BFEABF842FE1860DF" box="[468,482,1922,1948]" italics="true" pageId="5" pageNumber="154">h</emphasis>
<subScript id="3E433F256E5A9F5BFE9DF855FE1060E0" attach="left" box="[482,490,1941,1955]" fontSize="6" pageId="5" pageNumber="154">1</subScript>
and 
<emphasis id="90B3E1726E5A9F5BFD59F842FDCE60DF" box="[550,564,1922,1948]" italics="true" pageId="5" pageNumber="154">h</emphasis>
<subScript id="3E433F256E5A9F5BFD4BF855FDC660E0" attach="left" box="[564,572,1941,1955]" fontSize="6" pageId="5" pageNumber="154">2</subScript>
, 
<emphasis id="90B3E1726E5A9F5BFD33F842FD9860DF" box="[588,610,1922,1948]" italics="true" pageId="5" pageNumber="154">ih</emphasis>
lateral to anterior part of anal opening. Opisthonotal gland opening 
<emphasis id="90B3E1726E5A9F5BFA07F842FA6660DF" box="[1400,1436,1922,1948]" italics="true" pageId="5" pageNumber="154">gla</emphasis>
lateral to seta 
<emphasis id="90B3E1726E5A9F5BFE4BF866FEB06083" box="[308,330,1958,1984]" italics="true" pageId="5" pageNumber="154">lp</emphasis>
, with dark sclerotized surrounding, humeral organ anterior to seta 
<emphasis id="90B3E1726E5A9F5BFB46F867FBBF6083" box="[1081,1093,1959,1984]" italics="true" pageId="5" pageNumber="154">c</emphasis>
<subScript id="3E433F256E5A9F5BFB3AF879FBB76084" attach="left" box="[1093,1101,1977,1991]" fontSize="6" pageId="5" pageNumber="154">3</subScript>
(
<figureCitation id="3AFC21E56E5A9F5BFB25F866FB406083" box="[1114,1210,1958,1984]" captionStart="FIGURE 11" captionStartId="13.[151,250,1058,1082]" captionTargetBox="[286,1283,182,1035]" captionTargetId="figure-19@13.[285,1302,181,1035]" captionTargetPageId="13" captionText="FIGURE 11. Punctoribates ghilarovi, lateral aspect, legs partially drawn, scale bars 20 μm. (a) Larva, (b) tritonymph." figureDoi="http://doi.org/10.5281/zenodo.7077976" httpUri="https://zenodo.org/record/7077976/files/figure.png" pageId="5" pageNumber="154">Fig. 11a</figureCitation>
). Paraproctal valves (segment PS) glabrous. Most leg setae barbed, seta 
<emphasis id="90B3E1726E5A9F5BFDA7F80AFD0D60A7" box="[728,759,1994,2020]" italics="true" pageId="5" pageNumber="154">ft”</emphasis>
on tarsus I relatively long (
<figureCitation id="3AFC21E56E5A9F5BFB53F80AFB8560A7" box="[1068,1151,1994,2020]" captionStart="FIGURE 12" captionStartId="13.[151,250,1952,1976]" captionTargetBox="[292,1295,1108,1929]" captionTargetId="figure-40@13.[292,1295,1107,1929]" captionTargetPageId="13" captionText="FIGURE 12. Punctoribates ghilarovi, leg segments of larva (part of femur to tarsus), right side, antiaxial aspect, seta on the opposite side not illustrated is indicated in the legend, scale bar 10 μm. (a) Leg I, tarsus (pl’); (b) leg II; (c) leg III." figureDoi="http://doi.org/10.5281/zenodo.7077978" httpUri="https://zenodo.org/record/7077978/files/figure.png" pageId="5" pageNumber="154">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="A2783D606E599F58FFB8FF57FCA565B2" blockId="6.[151,1437,151,753]" pageId="6" pageNumber="155">
Shape and colour of protonymph, prodorsal setae and bothridium as in larva, but bothridial setae relatively thinner, and gastronotum with 15 pairs of setae because 
<emphasis id="90B3E1726E599F58FD9DFF7BFD0A6797" box="[738,752,187,212]" italics="true" pageId="6" pageNumber="155">p</emphasis>
-series added (
<figureCitation id="3AFC21E56E599F58FCECFF7BFC0E6796" box="[915,1012,187,213]" captionStart="FIGURE 10" captionStartId="12.[151,250,1937,1961]" captionTargetBox="[317,1272,186,1909]" captionTargetId="figure-32@12.[315,1272,181,1913]" captionTargetPageId="12" captionText="FIGURE 10. Punctoribates ghilarovi, ventral part of hysterosoma, legs partially drawn, scale bars 20 μm. (a) Larva, (b) prot- onymph." figureDoi="http://doi.org/10.5281/zenodo.7077974" httpUri="https://zenodo.org/record/7077974/files/figure.png" pageId="6" pageNumber="155">Fig. 10b</figureCitation>
), and retaining in subsequent nymphs (
<figureCitation id="3AFC21E56E599F58FFE0FF1FFEBD67BA" box="[159,327,223,249]" captionStart="FIGURE 13" captionStartId="14.[151,250,1981,2005]" captionTargetBox="[263,1314,197,1958]" captionTargetId="figure-32@14.[261,1326,181,1958]" captionTargetPageId="14" captionText="FIGURE 13. Punctoribates ghilarovi, ventral part of hysterosoma, legs partially drawn, scale bars 20 μm. (a) Deutonymph, (b) tritonymph." figureDoi="http://doi.org/10.5281/zenodo.7077983" httpUri="https://zenodo.org/record/7077983/files/figure.png" pageId="6" pageNumber="155">Figs. 13a, 13b</figureCitation>
). Most gastronotal setae of medium size (
<tableCitation id="EF4508DB6E599F58FC56FF1FFC8567BA" box="[809,895,223,249]" captionStart="TABLE 1" captionStartId="1.[151,239,729,753]" captionTargetPageId="1" captionText="TABLE 1. Measurements of some morphological characters of juvenile stages and adult of Punctoribates ghilarovi (mean measurements of 2–5 specimens in μm); nd – not developed." httpUri="http://table.plazi.org/id/F6B86DE86E5E9F5FFFE8FD19FC906456" pageId="6" pageNumber="155" tableUuid="F6B86DE86E5E9F5FFFE8FD19FC906456">Table 1</tableCitation>
). In protonymph, setae of 
<emphasis id="90B3E1726E599F58FBCAFF1FFB3967BB" box="[1205,1219,223,248]" italics="true" pageId="6" pageNumber="155">p</emphasis>
-series shorter than other posterior setae, and length increasing from 
<emphasis id="90B3E1726E599F58FDB8FEC3FD2F665F" box="[711,725,259,284]" italics="true" pageId="6" pageNumber="155">p</emphasis>
<subScript id="3E433F256E599F58FDAAFED5FD276660" attach="left" box="[725,733,277,291]" fontSize="6" pageId="6" pageNumber="155">3</subScript>
to 
<emphasis id="90B3E1726E599F58FC7AFEC3FCE9665F" box="[773,787,259,284]" italics="true" pageId="6" pageNumber="155">p</emphasis>
<subScript id="3E433F256E599F58FC6CFED5FCE16660" attach="left" box="[787,795,277,291]" fontSize="6" pageId="6" pageNumber="155">1</subScript>
(
<emphasis id="90B3E1726E599F58FC56FEC3FCCD665F" box="[809,823,259,284]" italics="true" pageId="6" pageNumber="155">p</emphasis>
<subScript id="3E433F256E599F58FC48FED5FCC56660" attach="left" box="[823,831,277,291]" fontSize="6" pageId="6" pageNumber="155">3</subScript>
smooth, other setae barbed), but in other nymphs all setae of 
<emphasis id="90B3E1726E599F58FF89FEE7FEFE6603" box="[246,260,295,320]" italics="true" pageId="6" pageNumber="155">p</emphasis>
-series of similar length and barbed (
<figureCitation id="3AFC21E56E599F58FDDDFEE7FCF46602" box="[674,782,295,321]" captionStart="FIGURE 10" captionStartId="12.[151,250,1937,1961]" captionTargetBox="[317,1272,186,1909]" captionTargetId="figure-32@12.[315,1272,181,1913]" captionTargetPageId="12" captionText="FIGURE 10. Punctoribates ghilarovi, ventral part of hysterosoma, legs partially drawn, scale bars 20 μm. (a) Larva, (b) prot- onymph." figureDoi="http://doi.org/10.5281/zenodo.7077974" httpUri="https://zenodo.org/record/7077974/files/figure.png" pageId="6" pageNumber="155">Figs. 10b</figureCitation>
, 
<figureCitation id="3AFC21E56E599F58FC65FEE7FC846602" box="[794,894,295,321]" captionStart="FIGURE 13" captionStartId="14.[151,250,1981,2005]" captionTargetBox="[263,1314,197,1958]" captionTargetId="figure-32@14.[261,1326,181,1958]" captionTargetPageId="14" captionText="FIGURE 13. Punctoribates ghilarovi, ventral part of hysterosoma, legs partially drawn, scale bars 20 μm. (a) Deutonymph, (b) tritonymph." figureDoi="http://doi.org/10.5281/zenodo.7077983" httpUri="https://zenodo.org/record/7077983/files/figure.png" pageId="6" pageNumber="155">13a, 13b</figureCitation>
). In all nymphs, gastronotal shield present, with 10 pairs of setae (
<emphasis id="90B3E1726E599F58FE1BFE8BFE886626" box="[356,370,331,357]" italics="true" pageId="6" pageNumber="155">d</emphasis>
-, 
<emphasis id="90B3E1726E599F58FEF4FE8BFE696626" box="[395,403,331,357]" italics="true" pageId="6" pageNumber="155">l</emphasis>
-, 
<emphasis id="90B3E1726E599F58FED4FE8BFE436626" box="[427,441,331,357]" italics="true" pageId="6" pageNumber="155">h</emphasis>
-series, 
<emphasis id="90B3E1726E599F58FD6DFE8BFDDA6627" box="[530,544,331,356]" italics="true" pageId="6" pageNumber="155">p</emphasis>
<subScript id="3E433F256E599F58FD5FFE9DFDD26628" attach="left" box="[544,552,349,363]" fontSize="6" pageId="6" pageNumber="155">1</subScript>
), in deutonymph and tritonymph setae 
<emphasis id="90B3E1726E599F58FC96FE8BFC0D6627" box="[1001,1015,331,356]" italics="true" pageId="6" pageNumber="155">p</emphasis>
<subScript id="3E433F256E599F58FC88FE9DFC056628" attach="left" box="[1015,1023,349,363]" fontSize="6" pageId="6" pageNumber="155">2</subScript>
and 
<emphasis id="90B3E1726E599F58FB47FE8BFBBC6627" box="[1080,1094,331,356]" italics="true" pageId="6" pageNumber="155">p</emphasis>
<subScript id="3E433F256E599F58FB39FE9DFBB46628" attach="left" box="[1094,1102,349,363]" fontSize="6" pageId="6" pageNumber="155">3</subScript>
inserted on ventral shield. In protonymph, one pair of genital setae appeared on genital valves, two pairs added in deutonymph and tritonymph each (
<figureCitation id="3AFC21E56E599F58FFA2FE53FEB166EE" box="[221,331,403,429]" captionStart="FIGURE 10" captionStartId="12.[151,250,1937,1961]" captionTargetBox="[317,1272,186,1909]" captionTargetId="figure-32@12.[315,1272,181,1913]" captionTargetPageId="12" captionText="FIGURE 10. Punctoribates ghilarovi, ventral part of hysterosoma, legs partially drawn, scale bars 20 μm. (a) Larva, (b) prot- onymph." figureDoi="http://doi.org/10.5281/zenodo.7077974" httpUri="https://zenodo.org/record/7077974/files/figure.png" pageId="6" pageNumber="155">Figs. 10b</figureCitation>
, 
<figureCitation id="3AFC21E56E599F58FE25FE53FE3A66EE" box="[346,448,403,429]" captionStart="FIGURE 13" captionStartId="14.[151,250,1981,2005]" captionTargetBox="[263,1314,197,1958]" captionTargetId="figure-32@14.[261,1326,181,1958]" captionTargetPageId="14" captionText="FIGURE 13. Punctoribates ghilarovi, ventral part of hysterosoma, legs partially drawn, scale bars 20 μm. (a) Deutonymph, (b) tritonymph." figureDoi="http://doi.org/10.5281/zenodo.7077983" httpUri="https://zenodo.org/record/7077983/files/figure.png" pageId="6" pageNumber="155">13a, 13b</figureCitation>
), all short and smooth. In deutonymph, one pair of aggenital setae and three pairs of adanal setae appeared, and remaining in tritonymph; all short and smooth. In protonymph and deutonymph, anal valves glabrous, in tritonymph two pairs of short and smooth anal setae present. In tritonymph, setae of 
<emphasis id="90B3E1726E599F58FA57FE1BFACE66B7" box="[1320,1332,475,500]" italics="true" pageId="6" pageNumber="155">c</emphasis>
-series of medium size and barbed, inserted on unsclerotized integument, 
<emphasis id="90B3E1726E599F58FC17FE3FFC8E655B" box="[872,884,511,536]" italics="true" pageId="6" pageNumber="155">c</emphasis>
<subScript id="3E433F256E599F58FC0BFDD1FC86655C" attach="left" box="[884,892,529,543]" fontSize="6" pageId="6" pageNumber="155">3</subScript>
slightly longer than 
<emphasis id="90B3E1726E599F58FB18FE3FFB89655B" box="[1127,1139,511,536]" italics="true" pageId="6" pageNumber="155">c</emphasis>
<subScript id="3E433F256E599F58FB0BFDD1FB86655C" attach="left" box="[1140,1148,529,543]" fontSize="6" pageId="6" pageNumber="155">1</subScript>
and 
<emphasis id="90B3E1726E599F58FBCBFE3FFB3A655B" box="[1204,1216,511,536]" italics="true" pageId="6" pageNumber="155">c</emphasis>
<subScript id="3E433F256E599F58FBBFFDD1FB32655C" attach="left" box="[1216,1224,529,543]" fontSize="6" pageId="6" pageNumber="155">2</subScript>
(
<figureCitation id="3AFC21E56E599F58FBA6FE3FFAD6655A" box="[1241,1324,511,537]" captionStart="FIGURE 14" captionStartId="15.[151,250,1979,2003]" captionTargetBox="[300,1282,189,1929]" captionTargetId="figure-19@15.[300,1286,181,1955]" captionTargetPageId="15" captionText="FIGURE 14. Punctoribates ghilarovi, tritonymph, dorsal aspect, legs partially drawn, scale bar 20 μm." figureDoi="http://doi.org/10.5281/zenodo.7077985" httpUri="https://zenodo.org/record/7077985/files/figure.png" pageId="6" pageNumber="155">Fig. 14</figureCitation>
, 
<tableCitation id="EF4508DB6E599F58FA47FE3FFA71655A" box="[1336,1419,511,537]" captionStart="TABLE 1" captionStartId="1.[151,239,729,753]" captionTargetPageId="1" captionText="TABLE 1. Measurements of some morphological characters of juvenile stages and adult of Punctoribates ghilarovi (mean measurements of 2–5 specimens in μm); nd – not developed." httpUri="http://table.plazi.org/id/F6B86DE86E5E9F5FFFE8FD19FC906456" pageId="6" pageNumber="155" tableUuid="F6B86DE86E5E9F5FFFE8FD19FC906456">Table 1</tableCitation>
). Other gastronotal setae also of medium size and barbed, except for short and smooth 
<emphasis id="90B3E1726E599F58FB31FDE3FB90657E" box="[1102,1130,547,573]" italics="true" pageId="6" pageNumber="155">da</emphasis>
and 
<emphasis id="90B3E1726E599F58FBDFFDE3FB38657E" box="[1184,1218,547,573]" italics="true" pageId="6" pageNumber="155">dm</emphasis>
, slightly longer 
<emphasis id="90B3E1726E599F58FA06FDE3FA6F657E" box="[1401,1429,547,573]" italics="true" pageId="6" pageNumber="155">dp</emphasis>
, and clearly longer 
<emphasis id="90B3E1726E599F58FE18FD87FE8F6522" box="[359,373,583,609]" italics="true" pageId="6" pageNumber="155">h</emphasis>
<subScript id="3E433F256E599F58FE0AFD99FE876524" attach="left" box="[373,381,601,615]" fontSize="6" pageId="6" pageNumber="155">1</subScript>
, 
<emphasis id="90B3E1726E599F58FEF4FD87FE636522" box="[395,409,583,609]" italics="true" pageId="6" pageNumber="155">h</emphasis>
<subScript id="3E433F256E599F58FEE6FD99FE5B6524" attach="left" box="[409,417,601,615]" fontSize="6" pageId="6" pageNumber="155">3</subScript>
and 
<emphasis id="90B3E1726E599F58FEA8FD87FE176522" box="[471,493,583,609]" italics="true" pageId="6" pageNumber="155">lp</emphasis>
. In tritonymph, cupules 
<emphasis id="90B3E1726E599F58FD80FD87FCEF6523" box="[767,789,583,608]" italics="true" pageId="6" pageNumber="155">ia</emphasis>
and 
<emphasis id="90B3E1726E599F58FC34FD87FC9D6523" box="[843,871,583,608]" italics="true" pageId="6" pageNumber="155">im</emphasis>
as in larva, 
<emphasis id="90B3E1726E599F58FC91FD87FBFE6523" box="[1006,1028,583,608]" italics="true" pageId="6" pageNumber="155">ip</emphasis>
anterolateral to seta 
<emphasis id="90B3E1726E599F58FB94FD87FB036523" box="[1259,1273,583,608]" italics="true" pageId="6" pageNumber="155">p</emphasis>
<subScript id="3E433F256E599F58FB86FD99FAFB6524" attach="left" box="[1273,1281,601,615]" fontSize="6" pageId="6" pageNumber="155">1</subScript>
, 
<emphasis id="90B3E1726E599F58FA70FD87FAC96522" box="[1295,1331,583,609]" italics="true" pageId="6" pageNumber="155">iad</emphasis>
lateral to anterior part of anal valves, and 
<emphasis id="90B3E1726E599F58FD7FFDABFDDB65C7" box="[512,545,619,644]" italics="true" pageId="6" pageNumber="155">ips</emphasis>
and 
<emphasis id="90B3E1726E599F58FD28FDABFD9765C6" box="[599,621,619,645]" italics="true" pageId="6" pageNumber="155">ih</emphasis>
pushed lateral and anterolateral to 
<emphasis id="90B3E1726E599F58FC89FDABFBE065C6" box="[1014,1050,619,645]" italics="true" pageId="6" pageNumber="155">iad</emphasis>
, respectively (
<figureCitation id="3AFC21E56E599F58FBC0FDABFAD065C6" box="[1215,1322,619,645]" captionStart="FIGURE 10" captionStartId="12.[151,250,1937,1961]" captionTargetBox="[317,1272,186,1909]" captionTargetId="figure-32@12.[315,1272,181,1913]" captionTargetPageId="12" captionText="FIGURE 10. Punctoribates ghilarovi, ventral part of hysterosoma, legs partially drawn, scale bars 20 μm. (a) Larva, (b) prot- onymph." figureDoi="http://doi.org/10.5281/zenodo.7077974" httpUri="https://zenodo.org/record/7077974/files/figure.png" pageId="6" pageNumber="155">Figs. 10b</figureCitation>
, 
<figureCitation id="3AFC21E56E599F58FA49FDABFA9B65C6" box="[1334,1377,619,645]" captionStart="FIGURE 11" captionStartId="13.[151,250,1058,1082]" captionTargetBox="[286,1283,182,1035]" captionTargetId="figure-19@13.[285,1302,181,1035]" captionTargetPageId="13" captionText="FIGURE 11. Punctoribates ghilarovi, lateral aspect, legs partially drawn, scale bars 20 μm. (a) Larva, (b) tritonymph." figureDoi="http://doi.org/10.5281/zenodo.7077976" httpUri="https://zenodo.org/record/7077976/files/figure.png" pageId="6" pageNumber="155">11b</figureCitation>
, 
<figureCitation id="3AFC21E56E599F58FA12FDABFF3E65EA" captionStart="FIGURE 13" captionStartId="14.[151,250,1981,2005]" captionTargetBox="[263,1314,197,1958]" captionTargetId="figure-32@14.[261,1326,181,1958]" captionTargetPageId="14" captionText="FIGURE 13. Punctoribates ghilarovi, ventral part of hysterosoma, legs partially drawn, scale bars 20 μm. (a) Deutonymph, (b) tritonymph." figureDoi="http://doi.org/10.5281/zenodo.7077983" httpUri="https://zenodo.org/record/7077983/files/figure.png" pageId="6" pageNumber="155">13a, 13b</figureCitation>
). Humeral organ large, rounded, porose, located lateral to seta 
<emphasis id="90B3E1726E599F58FCF8FD4FFC6965EB" box="[903,915,655,680]" italics="true" pageId="6" pageNumber="155">c</emphasis>
<subScript id="3E433F256E599F58FCECFD61FC6165EC" attach="left" box="[915,923,673,687]" fontSize="6" pageId="6" pageNumber="155">3</subScript>
, opisthonotal gland opening lateral to seta 
<emphasis id="90B3E1726E599F58FA00FD4FFA6F65EA" box="[1407,1429,655,681]" italics="true" pageId="6" pageNumber="155">lp</emphasis>
, with dark sclerotized surrounding (
<figureCitation id="3AFC21E56E599F58FD60FD73FD85658E" box="[543,639,691,717]" captionStart="FIGURE 11" captionStartId="13.[151,250,1058,1082]" captionTargetBox="[286,1283,182,1035]" captionTargetId="figure-19@13.[285,1302,181,1035]" captionTargetPageId="13" captionText="FIGURE 11. Punctoribates ghilarovi, lateral aspect, legs partially drawn, scale bars 20 μm. (a) Larva, (b) tritonymph." figureDoi="http://doi.org/10.5281/zenodo.7077976" httpUri="https://zenodo.org/record/7077976/files/figure.png" pageId="6" pageNumber="155">Fig. 11b</figureCitation>
). Legs of tritonymph stocky, tibiae without dorsal projections, most leg setae finely barbed, seta 
<emphasis id="90B3E1726E599F58FED4FD17FE3065B2" box="[427,458,727,753]" italics="true" pageId="6" pageNumber="155">ft”</emphasis>
on tarsus I relatively long (
<figureCitation id="3AFC21E56E599F58FD80FD17FCA865B2" box="[767,850,726,753]" captionStart="FIGURE 15" captionStartId="16.[151,250,1785,1809]" captionTargetBox="[161,1427,181,1749]" captionTargetId="figure-32@16.[161,1427,181,1762]" captionTargetPageId="16" captionText="FIGURE 15. Punctoribates ghilarovi, leg segments of tritonymph (part of femur to tarsus), right side, antiaxial aspect, seta on the opposite side not illustrated is indicated in the legend, scale bars 10 μm. (a) Leg I, tarsus (pl’); (b) leg II; (c) leg III; (d) leg IV." figureDoi="http://doi.org/10.5281/zenodo.7077987" httpUri="https://zenodo.org/record/7077987/files/figure.png" pageId="6" pageNumber="155">Fig. 15</figureCitation>
).
</paragraph>
<caption id="F6B86DE86E599F58FFE8F907FF346040" ID-DOI="http://doi.org/10.5281/zenodo.7077958" ID-Zenodo-Dep="7077958" httpUri="https://zenodo.org/record/7077958/files/figure.png" pageId="6" pageNumber="155" startId="6.[151,250,1735,1759]" targetBox="[170,1416,777,1712]" targetPageId="6">
<paragraph id="A2783D606E599F58FFE8F907FF346040" blockId="6.[151,1436,1735,1795]" pageId="6" pageNumber="155">
<emphasis id="90B3E1726E599F58FFE8F907FEE9619C" bold="true" box="[151,275,1735,1759]" pageId="6" pageNumber="155">FIGURE 4.</emphasis>
<taxonomicName id="65C746E36E599F58FE65F907FDF1619C" authorityName="Shaldybina" authorityYear="1969" box="[282,523,1735,1759]" class="Arachnida" family="Punctoribatidae" genus="Punctoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="155" phylum="Arthropoda" rank="species" species="ghilarovi">
<emphasis id="90B3E1726E599F58FE65F907FDF1619C" box="[282,523,1735,1759]" italics="true" pageId="6" pageNumber="155">Punctoribates ghilarovi</emphasis>
</taxonomicName>
, adult, SEM micrographs. (a) Dorsal view, (b) lateral view, (c) ventral view, (d) posterior view.
</paragraph>
</caption>
<paragraph id="A2783D606E599F58FFE8F8F2FD9B600F" blockId="6.[151,1436,1842,2012]" box="[151,609,1842,1868]" pageId="6" pageNumber="155">
<emphasis id="90B3E1726E599F58FFE8F8F2FD9B600F" box="[151,609,1842,1868]" italics="true" pageId="6" pageNumber="155">Summary of ontogenetic transformations</emphasis>
</paragraph>
<paragraph id="A2783D606E599F59FFB8F896FE3766EE" blockId="6.[151,1436,1842,2012]" lastBlockId="7.[151,1437,150,429]" lastPageId="7" lastPageNumber="156" pageId="6" pageNumber="155">
In the larva, the prodorsal seta 
<emphasis id="90B3E1726E599F58FD54F896FDB9602C" box="[555,579,1878,1903]" italics="true" pageId="6" pageNumber="155">ro</emphasis>
is long and setae 
<emphasis id="90B3E1726E599F58FC6CF896FCDD6033" box="[787,807,1878,1904]" italics="true" pageId="6" pageNumber="155">le</emphasis>
and 
<emphasis id="90B3E1726E599F58FC1EF896FC8D602C" box="[865,887,1878,1903]" italics="true" pageId="6" pageNumber="155">in</emphasis>
are of medium size, in the nymphs all setae are of medium size, whereas in the adult seta 
<emphasis id="90B3E1726E599F58FD0BF8BAFD7060D0" box="[628,650,1914,1939]" italics="true" pageId="6" pageNumber="155">in</emphasis>
is long, and other setae are of medium size. Seta 
<emphasis id="90B3E1726E599F58FBBFF8BAFB2360D0" box="[1216,1241,1914,1939]" italics="true" pageId="6" pageNumber="155">ex</emphasis>
is short in all instars. The bothridium is oval in all juveniles, whereas in the adult it is rounded and covered by anterior tectum of notogaster. In all instars, the bothridial seta is clavate, with barbed head, but in the larva the head is thicker than in other instars. The larva has 12 pairs of gastronotal setae, including 
<emphasis id="90B3E1726E589F59FCF0FF57FC6767F2" box="[911,925,151,177]" italics="true" pageId="7" pageNumber="156">h</emphasis>
<subScript id="3E433F256E589F59FCE2FF69FC5F67F4" attach="left" box="[925,933,169,183]" fontSize="6" pageId="7" pageNumber="156">3</subScript>
, the nymphs have 15 pairs (
<emphasis id="90B3E1726E589F59FB97FF57FB0C67F3" box="[1256,1270,151,176]" italics="true" pageId="7" pageNumber="156">p</emphasis>
-series appears in the protonymph), whereas the notogaster of adult loses five pairs of setae (
<emphasis id="90B3E1726E589F59FC86FF7BFBFF6797" box="[1017,1029,187,212]" italics="true" pageId="7" pageNumber="156">c</emphasis>
<subScript id="3E433F256E589F59FB79FF0DFBF46798" attach="right" box="[1030,1038,205,219]" fontSize="6" pageId="7" pageNumber="156">1</subScript>
, 
<emphasis id="90B3E1726E589F59FB63FF7BFBD26797" box="[1052,1064,187,212]" italics="true" pageId="7" pageNumber="156">c</emphasis>
<subScript id="3E433F256E589F59FB57FF0DFBCA6798" attach="left" box="[1064,1072,205,219]" fontSize="6" pageId="7" pageNumber="156">3</subScript>
and 
<emphasis id="90B3E1726E589F59FB18FF7BFB8F6796" box="[1127,1141,187,213]" italics="true" pageId="7" pageNumber="156">d</emphasis>
-series), so the seven pairs of microsetae remain and three pairs of short setae of 
<emphasis id="90B3E1726E589F59FD88FF1FFCFF67BB" box="[759,773,223,248]" italics="true" pageId="7" pageNumber="156">p</emphasis>
-series, including 
<emphasis id="90B3E1726E589F59FCB1FF1FFC2067BB" box="[974,986,223,248]" italics="true" pageId="7" pageNumber="156">c</emphasis>
<subScript id="3E433F256E589F59FCA4FF31FC1967BC" attach="right" box="[987,995,241,255]" fontSize="6" pageId="7" pageNumber="156">2</subScript>
. The formula of gastronotal setae in 
<taxonomicName id="65C746E36E589F59FAFBFF1FFF07665E" authorityName="Shaldybina" authorityYear="1969" class="Arachnida" family="Punctoribatidae" genus="Punctoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="156" phylum="Arthropoda" rank="species" species="ghilarovi">
<emphasis id="90B3E1726E589F59FAFBFF1FFA6F67BB" box="[1412,1429,223,248]" italics="true" pageId="7" pageNumber="156">P</emphasis>
. 
<emphasis id="90B3E1726E589F59FFE8FEC3FF07665E" box="[151,253,259,285]" italics="true" pageId="7" pageNumber="156">ghilarovi</emphasis>
</taxonomicName>
is 12-15-15-15-10 (from larva to adult), the formula of epimeral setae are 3-1-2 (larva, including scaliform 
<emphasis id="90B3E1726E589F59FFE8FEE7FF486602" box="[151,178,295,321]" italics="true" pageId="7" pageNumber="156">1c</emphasis>
), 3-1-2-1 (protonymph), 3-1-2-2 (deutonymph) and 3-1-3-3 (tritonymph and adult). The formula of genital setae is 1-3-5-6 (protonymph to adult), formula of aggenital setae is 1-1-1 (deutonymph to adult), and setal formula of segments PS-AN is 03333-0333-022. The ontogeny of leg setae and solenidia of 
<taxonomicName id="65C746E36E589F59FB61FEAFFB5866CA" authorityName="Shaldybina" authorityYear="1969" box="[1054,1186,367,393]" class="Arachnida" family="Punctoribatidae" genus="Punctoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="156" phylum="Arthropoda" rank="species" species="ghilarovi">
<emphasis id="90B3E1726E589F59FB61FEAFFBD566CB" box="[1054,1071,367,392]" italics="true" pageId="7" pageNumber="156">P</emphasis>
. 
<emphasis id="90B3E1726E589F59FB43FEAFFB5866CA" box="[1084,1186,367,393]" italics="true" pageId="7" pageNumber="156">ghilarovi</emphasis>
</taxonomicName>
is as in 
<taxonomicName id="65C746E36E589F59FB83FEAFFE3366EE" authority="(Seniczak et al. 2020 a)" baseAuthorityName="Seniczak" baseAuthorityYear="2020" class="Arachnida" family="Punctoribatidae" genus="Punctoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="156" phylum="Arthropoda" rank="species" species="astrachanicus">
<emphasis id="90B3E1726E589F59FB83FEAFFAF766CB" box="[1276,1293,367,392]" italics="true" pageId="7" pageNumber="156">P</emphasis>
. 
<emphasis id="90B3E1726E589F59FA64FEAFFF4666EF" italics="true" pageId="7" pageNumber="156">astrachanicus</emphasis>
(
<bibRefCitation id="C65640916E589F59FFB4FE53FE3866EE" author="Seniczak, S. &amp; Ivan O. &amp; Marquardt, T. &amp; Seniczak, A." box="[203,450,403,429]" pageId="7" pageNumber="156" pagination="1" refId="ref7460" refString="Seniczak, S., Ivan O., Marquardt, T. &amp; Seniczak, A. (2020 a) Morphological ontogeny of Punctoribates astrachanicus (Acari: Oribatida: Punctoribatidae), and comments on Punctoribates Berlese. Zootaxa, 4900 (1), 043 - 061. https: // doi. org / 10.11646 / zootaxa. 4900.1.6" type="journal article" year="2020">
Seniczak 
<emphasis id="90B3E1726E589F59FE46FE53FE9066EE" box="[313,362,403,429]" italics="true" pageId="7" pageNumber="156">et al</emphasis>
. 2020a
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<caption id="F6B86DE86E589F59FFE8FA43FC0062FC" ID-DOI="http://doi.org/10.5281/zenodo.7077962" ID-Zenodo-Dep="7077962" httpUri="https://zenodo.org/record/7077962/files/figure.png" pageId="7" pageNumber="156" startId="7.[151,250,1411,1435]" targetBox="[170,1416,453,1388]" targetPageId="7">
<paragraph id="A2783D606E589F59FFE8FA43FC0062FC" blockId="7.[151,1436,1411,1472]" pageId="7" pageNumber="156">
<emphasis id="90B3E1726E589F59FFE8FA43FEE962DF" bold="true" box="[151,275,1411,1436]" pageId="7" pageNumber="156">FIGURE 5.</emphasis>
<taxonomicName id="65C746E36E589F59FE66FA43FDF062D8" authorityName="Shaldybina" authorityYear="1969" box="[281,522,1411,1435]" class="Arachnida" family="Punctoribatidae" genus="Punctoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="156" phylum="Arthropoda" rank="species" species="ghilarovi">
<emphasis id="90B3E1726E589F59FE66FA43FDF062D8" box="[281,522,1411,1435]" italics="true" pageId="7" pageNumber="156">Punctoribates ghilarovi</emphasis>
</taxonomicName>
, adult, SEM micrographs. (a) Dorsal view, anterior and medial part, (b) medial and posterior part, (c) bothridial seta and legs I and II, (d) leg II with dorsal projection on tibia.
</paragraph>
</caption>
<paragraph id="A2783D606E589F59FFE8FA2EFDEB614B" blockId="7.[151,1437,1518,2012]" box="[151,529,1518,1544]" pageId="7" pageNumber="156">
<emphasis id="90B3E1726E589F59FFE8FA2EFDEB614B" box="[151,529,1518,1544]" italics="true" pageId="7" pageNumber="156">Distribution, ecology and biology</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="EADD6EEB6E589F59FFB8F9D2FCC861DB" pageId="7" pageNumber="156" type="reference_group">
<paragraph id="A2783D606E589F59FFB8F9D2FCC861DB" blockId="7.[151,1437,1518,2012]" pageId="7" pageNumber="156">
<taxonomicName id="65C746E36E589F59FFB8F9D2FE38616F" authorityName="Shaldybina" authorityYear="1969" box="[199,450,1554,1580]" class="Arachnida" family="Punctoribatidae" genus="Punctoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="156" phylum="Arthropoda" rank="species" species="gilarovi">
<emphasis id="90B3E1726E589F59FFB8F9D2FE38616F" box="[199,450,1554,1580]" italics="true" pageId="7" pageNumber="156">Punctoribates gilarovi</emphasis>
</taxonomicName>
is a species with Palaearctic distribution, i. e. distributed in east of occidental Palaearctic and 
<collectingCountry id="DAD07DF06E589F59FFB9F9F6FEC96113" box="[198,307,1590,1616]" name="Mongolia" pageId="7" pageNumber="156">Mongolia</collectingCountry>
(
<bibRefCitation id="C65640916E589F59FE3EF9F6FD5D6113" author="Murvanidze, M. &amp; Mumladze, L." box="[321,679,1590,1616]" pageId="7" pageNumber="156" pagination="1 - 81" refId="ref6985" refString="Murvanidze, M. &amp; Mumladze, L. (2016) Annotated checklist of Georgian oribatid mites. Zootaxa, 4089 (1), 1 - 81. https: // doi. org / 10.11646 / zootaxa. 4089.1.1" type="journal article" year="2016">Murvanidze &amp; Mumladze 2016</bibRefCitation>
; 
<bibRefCitation id="C65640916E589F59FDCEF9F6FCC46113" author="Subias, L. S." box="[689,830,1590,1616]" pageId="7" pageNumber="156" refId="ref7853" refString="Subias, L. S. (2004, updated in 2022) Listado sistematico, sinonimico y biogeografico de los Acaros Oribatidos (Acariformes, Oribatida) del mundo (1758 - 2002). Graellsia, 60 (numero extraordinario), 3 - 305. 17 ª actualizacion, 537 pp. (accessed April 2022) https: // doi. org / 10.3989 / graellsia. 2004. v 60. iextra. 218" type="book" year="2004">Subías 2004</bibRefCitation>
, 2022). This species was described from Gorky region (
<collectingRegion id="6003F3826E589F59FFE0F99AFE976137" box="[159,365,1626,1652]" country="Russia" name="Nizhniy Novgorod" pageId="7" pageNumber="156">Nizhny Novgorod</collectingRegion>
, 
<collectingCountry id="DAD07DF06E589F59FE08F99AFE3C6137" box="[375,454,1626,1652]" name="Russia" pageId="7" pageNumber="156">Russia</collectingCountry>
), and then was recorded from Caucasus (
<bibRefCitation id="C65640916E589F59FCF5F99AFB326137" author="Shtanchaeva, U. Ya. &amp; Subias, L. S." box="[906,1224,1626,1652]" pageId="7" pageNumber="156" refId="ref7811" refString="Shtanchaeva, U. Ya., Subias, L. S. (2010) Catalogue of oribatid mites of the Caucasus. Russian Academy of Sciences, Nauka Publ. Makhachkala, pp. 276. [in Russian]" type="book" year="2010">Shtanchaeva &amp; Subías 2010</bibRefCitation>
), 
<collectingCountry id="DAD07DF06E589F59FBA5F99AFAD26137" box="[1242,1320,1626,1652]" name="Poland" pageId="7" pageNumber="156">Poland</collectingCountry>
(
<bibRefCitation id="C65640916E589F59FA4AF99AFE7261DB" author="Niedbala, W. &amp; Olszanowski, Z." pageId="7" pageNumber="156" pagination="79 - 93" refId="ref7086" refString="Niedbala, W. &amp; Olszanowski, Z. (2008) Mechowce (Oribatida). In: Bogdanowicz W., Chudzicka E., Pilipiuk I., Skibinska E. (Eds.), Fauna of Poland; characteristics and checklist of species. Warszawa, Muzeum i Institut Zoologii PAN: pp. 79 - 93." type="book chapter" year="2008">Niedbała &amp; Olszanowski 2008</bibRefCitation>
), and 
<collectingCountry id="DAD07DF06E589F59FEB5F9BEFDD561DB" box="[458,559,1662,1688]" name="Romania" pageId="7" pageNumber="156">Romania</collectingCountry>
(
<bibRefCitation id="C65640916E589F59FD41F9BEFCDD61DB" author="Ivan, O. &amp; Vasiliu, N. A." box="[574,807,1662,1688]" pageId="7" pageNumber="156" pagination="29 - 40" refId="ref6880" refString="Ivan, O. &amp; Vasiliu, N. A. (2010) Fauna of oribatid mites (Acari, Oribatida) from the Movile Cave area (Dobrogea, Romania), Travaux de l'Institut de Speologie &quot; Emile Racovitza &quot;, 49, 29 - 40." type="journal article" year="2010">Ivan &amp; Vasiliu 2010</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="EADD6EEB6E589F56FFB8F962FE0866EE" lastPageId="8" lastPageNumber="157" pageId="7" pageNumber="156" type="biology_ecology">
<paragraph id="A2783D606E589F59FFB8F962FCD0606B" blockId="7.[151,1437,1518,2012]" pageId="7" pageNumber="156">
Ecology of 
<taxonomicName id="65C746E36E589F59FE38F962FE3D61FF" authorityName="Shaldybina" authorityYear="1969" box="[327,455,1698,1724]" class="Arachnida" family="Punctoribatidae" genus="Punctoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="156" phylum="Arthropoda" rank="species" species="ghilarovi">
<emphasis id="90B3E1726E589F59FE38F962FE3D61FF" box="[327,455,1698,1724]" italics="true" pageId="7" pageNumber="156">P. ghilarovi</emphasis>
</taxonomicName>
is poorly known so far. 
<bibRefCitation id="C65640916E589F59FDB0F962FC6761FF" author="Shaldybina, E. S." box="[719,925,1698,1724]" pageId="7" pageNumber="156" pagination="53 - 68" refId="ref7718" refString="Shaldybina, E. S. (1969) New species of beetle mites of the genus Punctoribates Berlese, 1908 of the Gorky Region. Notes of the Gorky Pedagogical Institute, Biological series, 99, 53 - 68. [in Russian]" type="journal article" year="1969">Shaldybina (1969)</bibRefCitation>
found this species in damp and bog meadows, while 
<bibRefCitation id="C65640916E589F59FFA3F906FD9A61A3" author="Murvanidze, M. &amp; Mumladze, L." box="[220,608,1734,1760]" pageId="7" pageNumber="156" pagination="1 - 81" refId="ref6985" refString="Murvanidze, M. &amp; Mumladze, L. (2016) Annotated checklist of Georgian oribatid mites. Zootaxa, 4089 (1), 1 - 81. https: // doi. org / 10.11646 / zootaxa. 4089.1.1" type="journal article" year="2016">Murvanidze and Mumladze (2016)</bibRefCitation>
reported it from forest soils. In 
<collectingCountry id="DAD07DF06E589F59FCBFF906FBDF61A3" box="[960,1061,1734,1760]" name="Romania" pageId="7" pageNumber="156">Romania</collectingCountry>
<taxonomicName id="65C746E36E589F59FB54F906FB5661A3" authorityName="Shaldybina" authorityYear="1969" box="[1067,1196,1734,1760]" class="Arachnida" family="Punctoribatidae" genus="Punctoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="156" phylum="Arthropoda" rank="species" species="ghilarovi">
<emphasis id="90B3E1726E589F59FB54F906FB5661A3" box="[1067,1196,1734,1760]" italics="true" pageId="7" pageNumber="156">P. ghilarovi</emphasis>
</taxonomicName>
was firstly found in a closed cave (
<bibRefCitation id="C65640916E589F59FE5AF92AFDF26047" author="Ivan, O. &amp; Vasiliu, N. A." box="[293,520,1770,1796]" pageId="7" pageNumber="156" pagination="29 - 40" refId="ref6880" refString="Ivan, O. &amp; Vasiliu, N. A. (2010) Fauna of oribatid mites (Acari, Oribatida) from the Movile Cave area (Dobrogea, Romania), Travaux de l'Institut de Speologie &quot; Emile Racovitza &quot;, 49, 29 - 40." type="journal article" year="2010">Ivan &amp; Vasiliu 2010</bibRefCitation>
), then in a forest robinia plantation (
<bibRefCitation id="C65640916E589F59FCEAF92AFB7F6047" author="Calugar, A. &amp; Ivan, O." box="[917,1157,1770,1796]" pageId="7" pageNumber="156" pagination="97 - 103" refId="ref6719" refString="Calugar, A. &amp; Ivan, O. (2013) Diversity and distribution of the edaphic mites (Acari: Gamasina, Oribatida) in some forest plantations from the Central Moldavian Plateau, Scientific papers, Agronomy series, Iasi, 56 (2), 97 - 103." type="journal article" year="2013">Călugăr &amp; Ivan 2013</bibRefCitation>
). All localities (including Murighiol, 
<tableCitation id="EF4508DB6E589F59FE69F8CEFE9E606B" box="[278,356,1806,1832]" captionStart="TABLE 1" captionStartId="1.[151,239,729,753]" captionTargetPageId="1" captionText="TABLE 1. Measurements of some morphological characters of juvenile stages and adult of Punctoribates ghilarovi (mean measurements of 2–5 specimens in μm); nd – not developed." httpUri="http://table.plazi.org/id/F6B86DE86E5E9F5FFFE8FD19FC906456" pageId="7" pageNumber="156" tableUuid="F6B86DE86E5E9F5FFFE8FD19FC906456">Table1</tableCitation>
) are situated in eastern part of 
<collectingCountry id="DAD07DF06E589F59FDC1F8CEFCDD606B" box="[702,807,1806,1832]" name="Romania" pageId="7" pageNumber="156">Romania</collectingCountry>
.
</paragraph>
<paragraph id="A2783D606E589F56FFB8F8F2FDA567BA" blockId="7.[151,1437,1518,2012]" lastBlockId="8.[151,1436,150,429]" lastPageId="8" lastPageNumber="157" pageId="7" pageNumber="156">
In this study, 
<taxonomicName id="65C746E36E589F59FE23F8F2FE1A600F" authorityName="Shaldybina" authorityYear="1969" box="[348,480,1842,1868]" class="Arachnida" family="Punctoribatidae" genus="Punctoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="156" phylum="Arthropoda" rank="species" species="ghilarovi">
<emphasis id="90B3E1726E589F59FE23F8F2FE976008" box="[348,365,1842,1867]" italics="true" pageId="7" pageNumber="156">P</emphasis>
. 
<emphasis id="90B3E1726E589F59FE05F8F2FE1A600F" box="[378,480,1842,1868]" italics="true" pageId="7" pageNumber="156">ghilarovi</emphasis>
</taxonomicName>
was the most abundant in halophilous meadow (100.2 indiv./ 
<quantity id="653F90856E589F59FBF6F8F1FB2C600F" box="[1161,1238,1841,1868]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="156" unit="cm" value="500.0">500cm</quantity>
<superScript id="55B290286E589F59FBAAF8F1FB27607C" attach="left" box="[1237,1245,1841,1855]" fontSize="6" pageId="7" pageNumber="156">3</superScript>
), and the density was lower in poplar plantation (44.4 indiv./ 
<quantity id="653F90856E589F59FD08F895FD3E6033" box="[631,708,1877,1904]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="156" unit="cm" value="500.0">500cm</quantity>
<superScript id="55B290286E589F59FDBBF895FD366020" attach="left" box="[708,716,1877,1891]" fontSize="6" pageId="7" pageNumber="156">3</superScript>
) and robinia plantation (11.2 indiv./ 
<quantity id="653F90856E589F59FB22F895FB506033" box="[1117,1194,1877,1904]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="156" unit="cm" value="500.0">500cm</quantity>
<superScript id="55B290286E589F59FBD5F895FB486020" attach="left" box="[1194,1202,1877,1891]" fontSize="6" pageId="7" pageNumber="156">3</superScript>
). The juveniles of 
<taxonomicName id="65C746E36E589F59FAFBF896FF0760D7" authorityName="Shaldybina" authorityYear="1969" class="Arachnida" family="Punctoribatidae" genus="Punctoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="156" phylum="Arthropoda" rank="species" species="ghilarovi">
<emphasis id="90B3E1726E589F59FAFBF896FA6F602C" box="[1412,1429,1878,1903]" italics="true" pageId="7" pageNumber="156">P</emphasis>
. 
<emphasis id="90B3E1726E589F59FFE8F8BAFF0760D7" box="[151,253,1914,1940]" italics="true" pageId="7" pageNumber="156">ghilarovi</emphasis>
</taxonomicName>
were present only in forest poplar plantation, where they constituted 9% of population. The stage structure of this population was the following: 
<specimenCount id="B4C1F6E96E589F59FD38F85EFD3D60FB" box="[583,711,1950,1976]" pageId="7" pageNumber="156" type="larva">nine larvae</specimenCount>
, 3 protonymphs, 6 deutonymphs, 2 tritonymphs and 
<specimenCount id="B4C1F6E96E589F59FA59F85EFA6660FB" box="[1318,1436,1950,1976]" pageId="7" pageNumber="156" type="adult">202 adults</specimenCount>
(total number from five replicates). In all habitats, the sex ratio (females to males) of species varied (1:0.36–1:0.58, mean 1:0.45), as well as the number of gravid females (32–76%). In poplar plantation and halophilous meadow females carried 1–3 large eggs, whereas in robinia plantation they were carried 1–6 large eggs (175 x 76), which comprised 48% of the length of females.
</paragraph>
<paragraph id="A2783D606E579F56FFB8FEC3FE0866EE" blockId="8.[151,1436,150,429]" pageId="8" pageNumber="157">
Among the investigated habitats, the adults inhabiting halophilous meadow were the smallest, and those inhabiting robinia plantation were the largest, and the adults living in poplar plantation were of medium size (
<tableCitation id="EF4508DB6E579F56FA23FEE7FF526626" captionStart="TABLE 2" captionStartId="8.[151,239,1493,1517]" captionTargetPageId="8" captionText="TABLE 2. Sex ratio, number of gravid females and eggs, and mean body length and width ± standard deviation (and range) of Punctoribates ghilarovi in μm in different sampling sites from Romania." httpUri="http://table.plazi.org/id/F6B86DE86E579F56FFE8FA15FBFC6152" pageId="8" pageNumber="157" tableUuid="F6B86DE86E579F56FFE8FA15FBFC6152">Table 2</tableCitation>
). The adults from robinia plantation were significanlty larger than those in other habitats, whereas only females from poplar plantation were significantly larger that those from halophilous meadow. In all habitats, females were significantly larger than males.
</paragraph>
</subSubSection>
</treatment>
</document>