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<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="1313–2970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik &amp; Sierwald, Petra" docDate="2009" docId="1772122E7831FFF1FF013EE8AE52EEEC" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Riotintobolus anomalus Wesener &amp; Enghoff &amp; Sierwald 2009, sp. n." docType="treatment" docUuid="D393DC52-5565-41B3-BA6E-D8ED7FAE46BA" docUuidSource="ZooBank" docVersion="11" lastPageNumber="56" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="55" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wesener, Thomas</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. &amp; Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. &amp; B 54 D 3623309</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Enghoff, Henrik</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sierwald, Petra</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-09-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>19</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>19</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>128</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
<mods:identifier type="ISSN">1313–2970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791413" ID-GBIF-Taxon="163747644" ID-Zenodo-Dep="3791413" LSID="urn:lsid:zoobank.org:act:D393DC52-5565-41B3-BA6E-D8ED7FAE46BA" httpUri="http://treatment.plazi.org/id/1772122E7831FFF1FF013EE8AE52EEEC" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55">
<subSubSection box="[140,609,412,439]" pageId="54" pageNumber="55" type="nomenclature">
<paragraph blockId="54.[140,994,412,475]" box="[140,609,412,439]" pageId="54" pageNumber="55">
<heading box="[140,609,412,439]" fontSize="11" level="2" pageId="54" pageNumber="55" reason="3">
<emphasis bold="true" box="[140,609,412,439]" pageId="54" pageNumber="55">
<taxonomicName authority="Wesener" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[140,532,412,439]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis bold="true" box="[140,424,412,439]" italics="true" pageId="54" pageNumber="55">Riotintobolus anomalus</emphasis>
Wesener
</taxonomicName>
,
<taxonomicNameLabel box="[544,609,413,439]" pageId="54" pageNumber="55" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[140,994,448,475]" pageId="54" pageNumber="55" type="description">
<paragraph blockId="54.[140,994,412,475]" box="[140,994,448,475]" pageId="54" pageNumber="55">
<uri box="[140,994,448,475]" pageId="54" pageNumber="55">
urn:lsid:zoobank.org:act:
<uuid box="[419,994,448,475]" pageId="54" pageNumber="55">D393DC52-5565-41B3-BA6E-D8ED7FAE46BA</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection pageId="54" pageNumber="55" type="materials_examined">
<paragraph blockId="54.[140,1115,518,1636]" pageId="54" pageNumber="55">
<emphasis bold="true" box="[140,371,518,544]" pageId="54" pageNumber="55">
<emphasis box="[140,241,518,544]" inLineHeading="true" pageId="54" pageNumber="55" reason="1">Material</emphasis>
examined:
</emphasis>
<materialsCitation ID-GBIF-Occurrence="2608818321" collectingDate="2002-02-11" collectingDateMax="2002-02-15" collectingDateMin="2002-02-11" collectionCode="CAS" collectorName="Fisher &amp; Griswold" country="Madagascar" elevation="200" latitude="-25.581667" location="Reserve Speciale de Cap Sainte Marie" longLatPrecision="21" longitude="45.168335" municipality="Province" pageId="54" pageNumber="55" specimenCode="BLF 5499" specimenCount="1" stateProvince="Toliara" typeStatus="holotype">
<emphasis box="[378,481,519,545]" italics="true" pageId="54" pageNumber="55">
<typeStatus box="[378,475,519,545]" pageId="54" pageNumber="55">Holotype</typeStatus>
:
</emphasis>
1 ♂ (
<quantity box="[547,630,518,545]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="54" pageNumber="55" unit="mm" value="45.0">45 mm</quantity>
long),
<collectionCode box="[711,764,519,545]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="54" pageNumber="55" type="Museum">CAS</collectionCode>
<specimenCode box="[771,890,518,545]" collectionCode="BLF" pageId="54" pageNumber="55">BLF 5499</specimenCode>
,
<collectingCountry box="[901,1035,519,545]" name="Madagascar" pageId="54" pageNumber="55">Madagascar</collectingCountry>
,
<collectingMunicipality pageId="54" pageNumber="55">Province</collectingMunicipality>
<collectingRegion box="[193,275,554,580]" country="Madagascar" name="Toliara" pageId="54" pageNumber="55">Toliara</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:1772122E7831FFF1FF013EE8AE52EEEC:9A04F5E37831FFF0FE923D5EAD95E815" box="[287,716,554,580]" country="Madagascar" latitude="-25.581667" longLatPrecision="21" longitude="45.168335" municipality="Province" name="Reserve Speciale de Cap Sainte Marie" pageId="54" pageNumber="55" stateProvince="Toliara">Réserve Spéciale de Cap Sainte Marie</location>
,
<locationDeviation box="[729,1035,554,580]" location="Marovato" pageId="54" pageNumber="55">
<quantity box="[729,825,554,580]" metricMagnitude="4" metricUnit="m" metricValue="1.23" pageId="54" pageNumber="55" unit="km" value="12.3">12.3 km</quantity>
262°W Marovato
</locationDeviation>
, spiny forest thicket,
<quantity box="[301,378,589,616]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="54" pageNumber="55" unit="m" value="200.0">
<elevation box="[301,378,589,616]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="54" pageNumber="55" unit="m" value="200.0">200 m</elevation>
</quantity>
,
<geoCoordinate box="[389,528,589,616]" degrees="25" direction="south" minutes="34" orientation="latitude" pageId="54" pageNumber="55" precision="15" seconds="54" value="-25.581667">25°34’54” S</geoCoordinate>
,
<geoCoordinate box="[539,667,589,616]" degrees="45" direction="east" minutes="10" orientation="longitude" pageId="54" pageNumber="55" precision="15" seconds="6" value="45.168335">45°10’6” E</geoCoordinate>
, leg.
<collectorName box="[724,794,589,615]" pageId="54" pageNumber="55">Fisher</collectorName>
,
<collectorName box="[804,908,589,615]" pageId="54" pageNumber="55">Griswold</collectorName>
et al.,
<collectingMethod box="[982,1103,589,616]" pageId="54" pageNumber="55">pitfall trap</collectingMethod>
,
<date box="[140,309,624,651]" pageId="54" pageNumber="55" value="2002-02-11" valueMax="2002-02-15" valueMin="2002-02-11">
<collectingDate box="[140,309,624,651]" pageId="54" pageNumber="55" value="2002-02-11" valueMax="2002-02-15" valueMin="2002-02-11">11–15.II.2002</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2608818319" box="[322,1044,624,651]" collectingDate="2002-02-11" collectingDateMax="2002-02-15" collectingDateMin="2002-02-11" collectionCode="CAS" collectorName="Fisher &amp; Griswold" country="Madagascar" elevation="200" latitude="-25.581667" location="Reserve Speciale de Cap Sainte Marie" longLatPrecision="21" longitude="45.168335" municipality="Province" pageId="54" pageNumber="55" specimenCode="BLF 5499" specimenCount="1" specimenCount-female="1" stateProvince="Toliara" typeStatus="paratype">
<emphasis box="[322,428,625,650]" italics="true" pageId="54" pageNumber="55">
<typeStatus box="[322,424,625,650]" pageId="54" pageNumber="55">Paratypes</typeStatus>
:
</emphasis>
&gt;25 ♂ and
<specimenCount box="[569,590,625,651]" pageId="54" pageNumber="55" type="female">♀</specimenCount>
,
<collectionCode box="[605,658,624,650]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="54" pageNumber="55" type="Museum">CAS</collectionCode>
<specimenCode box="[666,786,624,651]" collectionCode="BLF" pageId="54" pageNumber="55">BLF 5499</specimenCode>
, same data as previous
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608818341" collectingDate="2002-02-13" collectingDateMax="2002-02-19" collectingDateMin="2002-02-13" collectionCode="CAS" collectorName="Fisher &amp; Griswold" country="Madagascar" elevation="160" latitude="-25.594444" location="Reserve Speciale de Cap Sainte Marie" longLatPrecision="21" longitude="45.146946" municipality="Province" pageId="54" pageNumber="55" specimenCode="BLF 5651" specimenCount="3" specimenCount-female="3" stateProvince="Toliara" typeStatus="paratype">
2 ♂,
<specimenCount box="[140,183,660,686]" pageId="54" pageNumber="55" type="female">3 ♀</specimenCount>
,
<collectionCode box="[197,250,660,686]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="54" pageNumber="55" type="Museum">CAS</collectionCode>
<specimenCode box="[257,375,659,686]" collectionCode="BLF" pageId="54" pageNumber="55">BLF 5651</specimenCode>
, same data as previous, but
<quantity box="[689,766,659,686]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="54" pageNumber="55" unit="m" value="160.0">
<elevation box="[689,766,659,686]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="54" pageNumber="55" unit="m" value="160.0">160 m</elevation>
</quantity>
,
<geoCoordinate box="[777,917,659,686]" degrees="25" direction="south" minutes="35" orientation="latitude" pageId="54" pageNumber="55" precision="15" seconds="40" value="-25.594444">25°35’40” S</geoCoordinate>
,
<geoCoordinate box="[929,1057,659,686]" degrees="45" direction="east" minutes="8" orientation="longitude" pageId="54" pageNumber="55" precision="15" seconds="49" value="45.146946">45°8’49” E</geoCoordinate>
, leg.
<collectorName box="[140,210,695,721]" pageId="54" pageNumber="55">Fisher</collectorName>
,
<collectorName box="[221,325,695,721]" pageId="54" pageNumber="55">Griswold</collectorName>
et al.,
<collectingMethod box="[401,539,695,721]" pageId="54" pageNumber="55">malaise trap</collectingMethod>
,
<date box="[551,721,695,721]" pageId="54" pageNumber="55" value="2002-02-13" valueMax="2002-02-19" valueMin="2002-02-13">
<collectingDate box="[551,721,695,721]" pageId="54" pageNumber="55" value="2002-02-13" valueMax="2002-02-19" valueMin="2002-02-13">13–19.II.2002</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph blockId="54.[140,1115,518,1636]" pageId="54" pageNumber="55">
<emphasis bold="true" box="[185,487,729,755]" pageId="54" pageNumber="55">
<emphasis box="[185,257,729,755]" inLineHeading="true" pageId="54" pageNumber="55" reason="1">Other</emphasis>
material examined
</emphasis>
:
<materialsCitation ID-GBIF-Occurrence="2608818308" collectingDate="2002-02-11" collectingDateMax="2002-02-15" collectingDateMin="2002-02-11" collectionCode="CAS" collectorName="Fisher &amp; Griswold" elevation="200" latitude="-25.581667" location="Reserve Speciale de Cap Sainte Marie" longLatPrecision="21" longitude="45.168335" pageId="54" pageNumber="55" specimenCode="BLF 5503" specimenCount="1" specimenCount-female="1">
<specimenCount box="[502,546,730,756]" pageId="54" pageNumber="55" type="female">1 ♀</specimenCount>
,
<collectionCode box="[562,615,730,756]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="54" pageNumber="55" type="Museum">CAS</collectionCode>
<specimenCode box="[623,742,730,757]" collectionCode="BLF" pageId="54" pageNumber="55">BLF 5503</specimenCode>
,
<location LSID="urn:lsid:plazi:treatment:1772122E7831FFF1FF013EE8AE52EEEC:9A04F5E37831FFF0FD7E3DAEAF8BE946" latitude="-25.581667" longLatPrecision="21" longitude="45.168335" name="Reserve Speciale de Cap Sainte Marie" pageId="54" pageNumber="55">Réserve Spéciale de Cap Sainte Marie</location>
, spiny forest thicket,
<quantity box="[454,532,765,792]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="54" pageNumber="55" unit="m" value="200.0">
<elevation box="[454,532,765,792]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="54" pageNumber="55" unit="m" value="200.0">200 m</elevation>
</quantity>
,
<geoCoordinate box="[544,685,765,792]" degrees="25" direction="south" minutes="34" orientation="latitude" pageId="54" pageNumber="55" precision="15" seconds="54" value="-25.581667">25°34’54” S</geoCoordinate>
,
<geoCoordinate box="[697,826,765,792]" degrees="45" direction="east" minutes="10" orientation="longitude" pageId="54" pageNumber="55" precision="15" seconds="6" value="45.168335">45°10’6” E</geoCoordinate>
, leg.
<collectorName box="[886,956,765,791]" pageId="54" pageNumber="55">Fisher</collectorName>
,
<collectorName box="[968,1075,765,791]" pageId="54" pageNumber="55">Griswold</collectorName>
, et al.,
<date box="[181,350,800,827]" pageId="54" pageNumber="55" value="2002-02-11" valueMax="2002-02-15" valueMin="2002-02-11">
<collectingDate box="[181,350,800,827]" pageId="54" pageNumber="55" value="2002-02-11" valueMax="2002-02-15" valueMin="2002-02-11">11–15.II.2002</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608818344" collectingDate="2002-02-21" collectingDateMax="2002-02-25" collectingDateMin="2002-02-21" collectionCode="CAS" collectorName="Fisher &amp; Griswold" country="Madagascar" elevation="80" latitude="-24.65361" location="Mahafaly Plateau" longLatPrecision="21" longitude="43.996666" municipality="Province" pageId="54" pageNumber="55" specimenCode="BLF 5758" specimenCount="1" stateProvince="Toliara">
1 ♂,
<collectionCode box="[419,472,800,826]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="54" pageNumber="55" type="Museum">CAS</collectionCode>
<specimenCode box="[479,598,800,827]" collectionCode="BLF" pageId="54" pageNumber="55">BLF 5758</specimenCode>
,
<collectingCountry box="[609,743,800,826]" name="Madagascar" pageId="54" pageNumber="55">Madagascar</collectingCountry>
,
<collectingMunicipality box="[754,852,800,826]" pageId="54" pageNumber="55">Province</collectingMunicipality>
<collectingRegion box="[858,940,800,826]" country="Madagascar" name="Toliara" pageId="54" pageNumber="55">Toliara</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:1772122E7831FFF1FF013EE8AE52EEEC:9A04F5E37831FFF0FC3B3C54AFE7E90F" country="Madagascar" latitude="-24.65361" longLatPrecision="21" longitude="43.996666" municipality="Province" name="Mahafaly Plateau" pageId="54" pageNumber="55" stateProvince="Toliara">Mahafaly Plateau</location>
,
<locationDeviation box="[201,498,835,862]" location="Itampolo" pageId="54" pageNumber="55">
<quantity box="[201,282,835,861]" metricMagnitude="3" metricUnit="m" metricValue="6.2" pageId="54" pageNumber="55" unit="km" value="6.2">6.2 km</quantity>
74°ENE Itampolo
</locationDeviation>
, spiny forest thicket,
<quantity box="[737,799,836,862]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="54" pageNumber="55" unit="m" value="80.0">
<elevation box="[737,799,836,862]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="54" pageNumber="55" unit="m" value="80.0">80 m</elevation>
</quantity>
,
<geoCoordinate box="[810,949,836,862]" degrees="24" direction="south" minutes="39" orientation="latitude" pageId="54" pageNumber="55" precision="15" seconds="13" value="-24.65361">24°39’13” S</geoCoordinate>
,
<geoCoordinate box="[961,1103,835,862]" degrees="43" direction="east" minutes="59" orientation="longitude" pageId="54" pageNumber="55" precision="15" seconds="48" value="43.996666">43°59’48” E</geoCoordinate>
, leg.
<collectorName box="[185,255,871,897]" pageId="54" pageNumber="55">Fisher</collectorName>
,
<collectorName box="[265,369,871,897]" pageId="54" pageNumber="55">Griswold</collectorName>
et al., sifted litter,
<date box="[575,744,870,897]" pageId="54" pageNumber="55" value="2002-02-21" valueMax="2002-02-25" valueMin="2002-02-21">
<collectingDate box="[575,744,870,897]" pageId="54" pageNumber="55" value="2002-02-21" valueMax="2002-02-25" valueMin="2002-02-21">21–25.II.2002</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608818284" collectingDate="2002-02-21" collectingDateMax="2002-02-25" collectingDateMin="2002-02-21" collectionCode="CAS" collectorName="Fisher &amp; Griswold" country="Madagascar" elevation="80" latitude="-24.65361" location="Mahafaly Plateau" longLatPrecision="21" longitude="43.996666" municipality="Province" pageId="54" pageNumber="55" specimenCode="BLF 5762" specimenCount="1" stateProvince="Toliara">
2 ♂,
<collectionCode box="[810,863,871,897]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="54" pageNumber="55" type="Museum">CAS</collectionCode>
<specimenCode box="[870,987,870,897]" collectionCode="BLF" pageId="54" pageNumber="55">BLF 5762</specimenCode>
, same data as previous, but hand collecting
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608818226" collectingDate="2002-02-21" collectingDateMax="2002-02-25" collectingDateMin="2002-02-21" collectionCode="CAS" collectorName="Fisher &amp; Griswold" country="Madagascar" elevation="80" latitude="-24.65361" location="Mahafaly Plateau" longLatPrecision="21" longitude="43.996666" municipality="Province" pageId="54" pageNumber="55" specimenCode="BLF 5763" specimenCount="6" specimenCount-female="6" stateProvince="Toliara">
13 ♂,
<specimenCount box="[575,617,906,932]" pageId="54" pageNumber="55" type="female">6 ♀</specimenCount>
,
<collectionCode box="[631,684,906,932]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="54" pageNumber="55" type="Museum">CAS</collectionCode>
<specimenCode box="[690,806,906,933]" collectionCode="BLF" pageId="54" pageNumber="55">BLF 5763</specimenCode>
, same data as previous,
<collectingMethod pageId="54" pageNumber="55">but pitfall trap</collectingMethod>
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608818257" box="[274,1102,941,968]" collectionCode="MNHN" collectorName="Petit" country="Madagascar" location="Madagascar" pageId="54" pageNumber="55" specimenCount="1">
1 ♂,
<collectionCode box="[331,430,942,968]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="54" pageNumber="55">MNHN</collectionCode>
,
<collectingCountry box="[442,576,941,967]" name="Madagascar" pageId="54" pageNumber="55">Madagascar</collectingCountry>
, leg.
<collectorName box="[634,689,941,967]" pageId="54" pageNumber="55">Petit</collectorName>
, 24–1922, envoi vi, entrée 24–1922
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2608818264" collectingDate="2003-01-21" collectingDateMax="2003-01-28" collectingDateMin="2003-01-21" collectionCode="CASENT" collectorName="Frontier Wilderness Project" country="Madagascar" elevation="30" latitude="-23.049017" location="Lake Ranobe" longLatPrecision="1" longitude="43.610584" municipality="Province" pageId="54" pageNumber="55" specimenCode="CASENT 9032805" specimenCount="1" stateProvince="Toliara">
1 ♂,
<specimenCode box="[195,419,976,1003]" collectionCode="CASENT" pageId="54" pageNumber="55">CASENT 9032805</specimenCode>
,
<collectingMunicipality box="[429,527,976,1002]" pageId="54" pageNumber="55">Province</collectingMunicipality>
<collectingRegion box="[531,613,976,1002]" country="Madagascar" name="Toliara" pageId="54" pageNumber="55">Toliara</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:1772122E7831FFF1FF013EE8AE52EEEC:9A04F5E37831FFF0FDE23CA4AC5CE9BB" box="[623,773,976,1002]" country="Madagascar" latitude="-23.049017" longLatPrecision="1" longitude="43.610584" municipality="Province" name="Lake Ranobe" pageId="54" pageNumber="55" stateProvince="Toliara">Lake Ranobe</location>
, spiny forest,
<quantity box="[925,986,976,1003]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="54" pageNumber="55" unit="m" value="30.0">
<elevation box="[925,986,976,1003]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="54" pageNumber="55" unit="m" value="30.0">30 m</elevation>
</quantity>
,
<geoCoordinate degrees="23" direction="south" minutes="2.941" orientation="latitude" pageId="54" pageNumber="55" precision="1" value="-23.049017">23°2.941’ S</geoCoordinate>
,
<geoCoordinate box="[169,322,1011,1038]" degrees="43" direction="east" minutes="36.635" orientation="longitude" pageId="54" pageNumber="55" precision="1" value="43.610584">43°36.635’ E</geoCoordinate>
, leg.
<collectorName box="[380,693,1011,1037]" pageId="54" pageNumber="55">Frontier Wilderness Project</collectorName>
,
<date box="[705,865,1012,1038]" pageId="54" pageNumber="55" value="2003-01-21" valueMax="2003-01-28" valueMin="2003-01-21">
<collectingDate box="[705,865,1012,1038]" pageId="54" pageNumber="55" value="2003-01-21" valueMax="2003-01-28" valueMin="2003-01-21">21–28.I.2003</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="54" pageNumber="55" type="diagnosis">
<paragraph blockId="54.[140,1115,518,1636]" pageId="54" pageNumber="55">
<emphasis bold="true" box="[185,454,1046,1072]" pageId="54" pageNumber="55">
<emphasis box="[185,324,1046,1072]" inLineHeading="true" pageId="54" pageNumber="55" reason="1">Differential</emphasis>
diagnosis:
</emphasis>
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[465,604,1047,1073]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species">
<emphasis box="[465,604,1047,1073]" italics="true" pageId="54" pageNumber="55">R. anomalus</emphasis>
</taxonomicName>
is the only known species of
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[964,1107,1047,1073]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis box="[964,1107,1047,1073]" italics="true" pageId="54" pageNumber="55">Riotintobolus</emphasis>
</taxonomicName>
where a sharp-edged preanal process and well-developed lips are absent. Furthermore it is the only species with male tarsal pads on every leg pair. The gonopods of this species are distinct but in some aspects similar to those of
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[756,923,1152,1178]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species">
<emphasis box="[756,923,1152,1178]" italics="true" pageId="54" pageNumber="55">R. mandenensis</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[979,1100,1153,1178]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species">
<emphasis box="[979,1100,1153,1178]" italics="true" pageId="54" pageNumber="55">R. minutus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55" type="description">
<paragraph blockId="54.[140,1115,518,1636]" pageId="54" pageNumber="55">
<emphasis bold="true" box="[185,335,1187,1213]" pageId="54" pageNumber="55">
<emphasis box="[185,330,1187,1213]" inLineHeading="true" pageId="54" pageNumber="55" reason="1">Description</emphasis>
.
</emphasis>
<emphasis box="[342,494,1188,1213]" italics="true" pageId="54" pageNumber="55">Measurements</emphasis>
: male
<typeStatus box="[569,668,1187,1213]" pageId="54" pageNumber="55">holotype</typeStatus>
with 45 podous and no apodous rings, circa
<quantity box="[201,284,1222,1249]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="54" pageNumber="55" unit="mm" value="45.0">45 mm</quantity>
long (broken),
<quantity box="[465,556,1223,1249]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="54" pageNumber="55" unit="mm" value="4.3">4.3 mm</quantity>
wide. Females with 44 or 45 body rings and no apodous ring, up to
<quantity box="[370,453,1258,1285]" metricMagnitude="-2" metricUnit="m" metricValue="5.1" pageId="54" pageNumber="55" unit="mm" value="51.0">51 mm</quantity>
long,
<quantity box="[525,615,1258,1284]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="54" pageNumber="55" unit="mm" value="4.6">4.6 mm</quantity>
wide.
</paragraph>
<paragraph blockId="54.[140,1115,518,1636]" pageId="54" pageNumber="55">
<emphasis box="[185,300,1293,1319]" italics="true" pageId="54" pageNumber="55">Coloration</emphasis>
light grey with darker posterior margins. Openings of ozopores highlighted with a black spot. Collum and rings dorsally with an extraordinary wide bloodred or rust-red stripe. Antennae and legs dark grey.
<emphasis box="[721,764,1364,1389]" italics="true" pageId="54" pageNumber="55">Eyes</emphasis>
with 30–34 ocelli arranged in five rows.
<emphasis box="[259,362,1399,1425]" italics="true" pageId="54" pageNumber="55">Antennae</emphasis>
short, protruding in both sexes back to body ring 2 (
<figureCitation box="[990,1095,1399,1425]" captionStart="Figure 29" captionStartId="55.[140,212,595,617]" captionTargetBox="[146,1108,172,572]" captionTargetId="figure@55.[140,1108,165,575]" captionTargetPageId="55" captionText="Figure 29. Riotintobolus anomalus sp. n., A–C, male (MNHN), D–G male holotype, A telson B right antenna C left midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F posterior gonopods, anterior view G posterior gonopods, posterior view. av = anal valves; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767310" httpUri="https://zenodo.org/record/3767310/files/figure.png" pageId="54" pageNumber="55">Fig. 29B</figureCitation>
).
<emphasis box="[140,185,1435,1460]" italics="true" pageId="54" pageNumber="55">Legs</emphasis>
reach 0.7 times body diameter, tarsi 3 and beyond in males with tarsal pads (
<figureCitation captionStart="Figure 29" captionStartId="55.[140,212,595,617]" captionTargetBox="[146,1108,172,572]" captionTargetId="figure@55.[140,1108,165,575]" captionTargetPageId="55" captionText="Figure 29. Riotintobolus anomalus sp. n., A–C, male (MNHN), D–G male holotype, A telson B right antenna C left midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F posterior gonopods, anterior view G posterior gonopods, posterior view. av = anal valves; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767310" httpUri="https://zenodo.org/record/3767310/files/figure.png" pageId="54" pageNumber="55">Fig. 29C</figureCitation>
).
<emphasis box="[214,374,1469,1495]" italics="true" pageId="54" pageNumber="55">Preanal process</emphasis>
absent, not protruding (
<figureCitation box="[654,758,1469,1495]" captionStart="Figure 29" captionStartId="55.[140,212,595,617]" captionTargetBox="[146,1108,172,572]" captionTargetId="figure@55.[140,1108,165,575]" captionTargetPageId="55" captionText="Figure 29. Riotintobolus anomalus sp. n., A–C, male (MNHN), D–G male holotype, A telson B right antenna C left midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F posterior gonopods, anterior view G posterior gonopods, posterior view. av = anal valves; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767310" httpUri="https://zenodo.org/record/3767310/files/figure.png" pageId="54" pageNumber="55">Fig. 29A</figureCitation>
). Anal valves small, lips weakly developed (
<figureCitation box="[269,372,1504,1530]" captionStart="Figure 29" captionStartId="55.[140,212,595,617]" captionTargetBox="[146,1108,172,572]" captionTargetId="figure@55.[140,1108,165,575]" captionTargetPageId="55" captionText="Figure 29. Riotintobolus anomalus sp. n., A–C, male (MNHN), D–G male holotype, A telson B right antenna C left midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F posterior gonopods, anterior view G posterior gonopods, posterior view. av = anal valves; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767310" httpUri="https://zenodo.org/record/3767310/files/figure.png" pageId="54" pageNumber="55">Fig. 29A</figureCitation>
).
</paragraph>
<paragraph blockId="54.[140,1115,518,1636]" lastBlockId="55.[140,1108,765,1213]" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55">
<emphasis box="[185,373,1539,1566]" italics="true" pageId="54" pageNumber="55">Anterior gonopod</emphasis>
sternite apically swollen into a wide lobe with a triangular, wellrounded tip (
<figureCitation box="[289,396,1575,1601]" captionStart="Figure 29" captionStartId="55.[140,212,595,617]" captionTargetBox="[146,1108,172,572]" captionTargetId="figure@55.[140,1108,165,575]" captionTargetPageId="55" captionText="Figure 29. Riotintobolus anomalus sp. n., A–C, male (MNHN), D–G male holotype, A telson B right antenna C left midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F posterior gonopods, anterior view G posterior gonopods, posterior view. av = anal valves; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767310" httpUri="https://zenodo.org/record/3767310/files/figure.png" pageId="54" pageNumber="55">Fig. 29D</figureCitation>
). Sternite tip almost as high as mesal coxite process, latter long, relatively slender, longer than sternite (
<figureCitation box="[575,681,1610,1636]" captionStart="Figure 29" captionStartId="55.[140,212,595,617]" captionTargetBox="[146,1108,172,572]" captionTargetId="figure@55.[140,1108,165,575]" captionTargetPageId="55" captionText="Figure 29. Riotintobolus anomalus sp. n., A–C, male (MNHN), D–G male holotype, A telson B right antenna C left midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F posterior gonopods, anterior view G posterior gonopods, posterior view. av = anal valves; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767310" httpUri="https://zenodo.org/record/3767310/files/figure.png" pageId="54" pageNumber="55">Fig. 29D</figureCitation>
). Telopodite on posterior side apically with a large triangular retrorse process (
<figureCitation box="[601,705,765,791]" captionStart="Figure 29" captionStartId="55.[140,212,595,617]" captionTargetBox="[146,1108,172,572]" captionTargetId="figure@55.[140,1108,165,575]" captionTargetPageId="55" captionText="Figure 29. Riotintobolus anomalus sp. n., A–C, male (MNHN), D–G male holotype, A telson B right antenna C left midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F posterior gonopods, anterior view G posterior gonopods, posterior view. av = anal valves; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767310" httpUri="https://zenodo.org/record/3767310/files/figure.png" pageId="55" pageNumber="56">Fig. 29E</figureCitation>
), projecting above coxite and telopodite margins (
<figureCitation box="[325,428,800,826]" captionStart="Figure 29" captionStartId="55.[140,212,595,617]" captionTargetBox="[146,1108,172,572]" captionTargetId="figure@55.[140,1108,165,575]" captionTargetPageId="55" captionText="Figure 29. Riotintobolus anomalus sp. n., A–C, male (MNHN), D–G male holotype, A telson B right antenna C left midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F posterior gonopods, anterior view G posterior gonopods, posterior view. av = anal valves; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767310" httpUri="https://zenodo.org/record/3767310/files/figure.png" pageId="55" pageNumber="56">Fig. 29E</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767310" ID-Zenodo-Dep="3767310" httpUri="https://zenodo.org/record/3767310/files/figure.png" pageId="55" pageNumber="56" startId="55.[140,212,595,617]" targetBox="[146,1108,172,572]" targetPageId="55">
<paragraph pageId="55" pageNumber="56">
<emphasis bold="true" box="[140,252,595,617]" pageId="55" pageNumber="56">Figure 29.</emphasis>
<taxonomicName authority="Wesener &amp; Enghoff &amp; Sierwald, 2009" authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[259,468,595,616]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" status="sp. n.">
<emphasis box="[259,468,595,616]" italics="true" pageId="55" pageNumber="56">Riotintobolus anomalus</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[475,527,596,617]" pageId="55" pageNumber="56" rank="species">sp. n.</taxonomicNameLabel>
, A–C, male (
<emphasis bold="true" box="[654,731,595,616]" pageId="55" pageNumber="56">MNHN</emphasis>
), D–G male holotype,
<emphasis bold="true" box="[952,971,595,617]" pageId="55" pageNumber="56">A</emphasis>
telson
<emphasis bold="true" box="[1039,1056,595,617]" pageId="55" pageNumber="56">B</emphasis>
right antenna
<emphasis bold="true" box="[222,241,627,649]" pageId="55" pageNumber="56">C</emphasis>
left midbody leg
<emphasis bold="true" box="[413,432,627,649]" pageId="55" pageNumber="56">D</emphasis>
anterior gonopod, anterior view
<emphasis bold="true" box="[750,765,627,649]" pageId="55" pageNumber="56">E</emphasis>
anterior gonopod, posterior view
<emphasis bold="true" box="[1093,1107,627,649]" pageId="55" pageNumber="56">F</emphasis>
posterior gonopods, anterior view
<emphasis bold="true" box="[465,485,659,681]" pageId="55" pageNumber="56">G</emphasis>
posterior gonopods, posterior view. av = anal valves; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph blockId="55.[140,1108,765,1213]" pageId="55" pageNumber="56">
<emphasis box="[185,386,835,861]" italics="true" pageId="55" pageNumber="56">Posterior gonopods</emphasis>
telopodite laterally with a single, large, finger-shaped process (
<figureCitation box="[148,249,871,897]" captionStart="Figure 29" captionStartId="55.[140,212,595,617]" captionTargetBox="[146,1108,172,572]" captionTargetId="figure@55.[140,1108,165,575]" captionTargetPageId="55" captionText="Figure 29. Riotintobolus anomalus sp. n., A–C, male (MNHN), D–G male holotype, A telson B right antenna C left midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F posterior gonopods, anterior view G posterior gonopods, posterior view. av = anal valves; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767310" httpUri="https://zenodo.org/record/3767310/files/figure.png" pageId="55" pageNumber="56">Fig. 29F</figureCitation>
). Sperm canal discharging into process. Apically, a large membrane present, which extends into two lobes, apically twice as wide as basally (
<figureCitation box="[851,958,906,932]" captionStart="Figure 29" captionStartId="55.[140,212,595,617]" captionTargetBox="[146,1108,172,572]" captionTargetId="figure@55.[140,1108,165,575]" captionTargetPageId="55" captionText="Figure 29. Riotintobolus anomalus sp. n., A–C, male (MNHN), D–G male holotype, A telson B right antenna C left midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F posterior gonopods, anterior view G posterior gonopods, posterior view. av = anal valves; Cx = coxite; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767310" httpUri="https://zenodo.org/record/3767310/files/figure.png" pageId="55" pageNumber="56">Fig. 29G</figureCitation>
).
</paragraph>
<paragraph blockId="55.[140,1108,765,1213]" pageId="55" pageNumber="56">
<emphasis bold="true" box="[185,457,941,967]" pageId="55" pageNumber="56">
<emphasis box="[185,333,941,967]" inLineHeading="true" pageId="55" pageNumber="56" reason="1">Intraspecific</emphasis>
variation:
</emphasis>
no variation inside or in between the Mahafaly and Cap Sainte Marie populations was observed.
</paragraph>
</subSubSection>
<subSubSection pageId="55" pageNumber="56" type="distribution">
<paragraph blockId="55.[140,1108,765,1213]" pageId="55" pageNumber="56">
<emphasis bold="true" box="[185,504,1011,1037]" pageId="55" pageNumber="56">
<emphasis box="[185,338,1011,1037]" inLineHeading="true" pageId="55" pageNumber="56" reason="1">Distribution</emphasis>
and ecology:
</emphasis>
this species is currently known from the Mahafaly plateau, the Cap Sainte Marie, and a third, distant location at the Lake Ranobe. All localities are probably the driest places on
<collectingCountry box="[672,806,1082,1108]" name="Madagascar" pageId="55" pageNumber="56">Madagascar</collectingCountry>
(
<bibRefCitation author="Battistini" box="[822,1000,1082,1108]" firstAuthor="Battistini" pageId="55" pageNumber="56" pagination="1 - 27" refId="ref56767" refString="Battistini R (1972) Madagascar relief and main types of landscape. In: Battistini R, Richard- Vindard G (Eds) Biogeography and Ecology in Madagascar. Monographiae Biologicae 21. The Hague, 1 - 27." type="book chapter" year="1972">Battistini 1972</bibRefCitation>
,
<bibRefCitation author="Moat &amp; Smith" firstAuthor="Moat" pageId="55" pageNumber="56" refId="ref57536" refString="Moat J, Smith P (2007) Atlas of the Vegetation of Madagascar. Kew Publishing, London, 124 pp." type="book" year="2007">Moat &amp; Smith 2007</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="55" pageNumber="56" type="etymology">
<paragraph blockId="55.[140,1108,765,1213]" pageId="55" pageNumber="56">
<emphasis bold="true" box="[185,323,1152,1178]" pageId="55" pageNumber="56">
<emphasis box="[185,318,1152,1178]" inLineHeading="true" pageId="55" pageNumber="56" reason="1">Etymology</emphasis>
:
</emphasis>
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[330,433,1152,1178]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species">
<emphasis box="[330,433,1152,1178]" italics="true" pageId="55" pageNumber="56">anomalus</emphasis>
</taxonomicName>
, adjective, refers to the unusual (for
<taxonomicName authorityName="Wesener &amp; Enghoff &amp; Sierwald" authorityYear="2009" box="[850,993,1152,1178]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="55" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis box="[850,993,1152,1178]" italics="true" pageId="55" pageNumber="56">Riotintobolus</emphasis>
</taxonomicName>
) telson of the species.
</paragraph>
</subSubSection>
</treatment>
</document>
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