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<document ID-DOI="10.5252/z2011n3a6" ID-ISSN="1638-9387" ID-Zenodo-Dep="7671500" checkinTime="1628223569786" checkinUser="felipe" docAuthor="Sanborn, Allen F." docDate="2011" docId="03FD8799FFB8EE4BFCEA325966F0BFAF" docLanguage="en" docName="Zoosystema.33.3.377-418.pdf.imf" docOrigin="Zoosystema 33 (3)" docStyle="DocumentStyle:EB3C2C73672D2E883259A050BD27978B.5:Zoosystema.2011-2012.journal_article" docStyleId="EB3C2C73672D2E883259A050BD27978B" docStyleName="Zoosystema.2011-2012.journal_article" docStyleVersion="5" docTitle="Taphura boulardi Sanborn, 2011, n. sp." docType="treatment" docVersion="3" lastPageNumber="408" masterDocId="FFC4FFE1FFA4EE54FFC4306D675EBC44" masterDocTitle="Checklist of the cicadas of French Guiana including new records and the description of nine new species (Insecta, Hemiptera, Cicadoidea, Cicadidae)" masterLastPageNumber="418" masterPageNumber="377" pageNumber="405" updateTime="1677175843659" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Checklist of the cicadas of French Guiana including new records and the description of nine new species (Insecta, Hemiptera, Cicadoidea, Cicadidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sanborn, Allen F.</mods:namePart>
<mods:affiliation>Department of Biology, Barry University, 11300 NE Second Avenue, Miami Shores, FL 33161 - 6695 (USA) asanborn @ mail. barry. edu</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zoosystema</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2011</mods:date>
<mods:detail type="pubDate">
<mods:number>2011-09-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>33</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>377</mods:start>
<mods:end>418</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5252/z2011n3a6</mods:identifier>
<mods:identifier type="ISSN">1638-9387</mods:identifier>
<mods:identifier type="Zenodo-Dep">7671500</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7670068" ID-Zenodo-Dep="7670068" LSID="urn:lsid:plazi:treatment:03FD8799FFB8EE4BFCEA325966F0BFAF" httpUri="http://treatment.plazi.org/id/03FD8799FFB8EE4BFCEA325966F0BFAF" lastPageId="31" lastPageNumber="408" pageId="28" pageNumber="405">
<subSubSection box="[814,1072,564,591]" pageId="28" pageNumber="405" type="nomenclature">
<paragraph blockId="28.[814,1072,564,623]" box="[814,1072,564,591]" pageId="28" pageNumber="405">
<heading box="[814,1072,564,591]" centered="true" fontSize="11" level="2" pageId="28" pageNumber="405" reason="2">
<taxonomicName ID-CoL="54SNH" authorityName="Sanborn" authorityYear="2011" box="[814,1007,564,591]" class="Insecta" family="Cicadidae" genus="Taphura" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="405" phylum="Arthropoda" rank="species" species="boulardi" status="n. sp.">
<emphasis bold="true" box="[814,1007,564,591]" italics="true" pageId="28" pageNumber="405">Taphura boulardi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1013,1072,564,591]" pageId="28" pageNumber="405" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[857,1030,596,623]" pageId="28" pageNumber="405" type="description">
<paragraph blockId="28.[814,1072,564,623]" box="[857,1030,596,623]" pageId="28" pageNumber="405">
(
<figureCitation box="[865,930,596,623]" captionStart="FIG" captionStartId="30.[113,124,1635,1652]" captionTargetBox="[142,1181,214,1575]" captionTargetId="figure-17@30.[142,1181,214,1575]" captionTargetPageId="30" captionText="FIG. 7. — Taphura boulardi n. sp.: A, holotype ♂ and paratype ♀ habitus; B, holotype ♂ dorsum; C, holotype ♂ timbal; D, holotype ♂ operculum; E, paratype ♀ operculum; F, paratype ♂ lateral view of genitalia; G, paratype ♂ posterior view of genitalia; H, paratype ♀ lateral view of genitalia; I, paratype ♀ ventral view of genitalia. Scale bars: A, 1 cm; B-I, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7671518" httpUri="https://zenodo.org/record/7671518/files/figure.png" pageId="28" pageNumber="405">Fig. 7</figureCitation>
;
<tableCitation box="[940,1020,596,623]" captionStart="TABLE" captionStartId="29.[115,124,219,236]" captionTargetPageId="29" captionText="TABLE 7. — Measurements of Taphura boulardi n. sp. in mm, mean (range). Note: width head including eyes; width of pronotum including suprahumeral plates." pageId="28" pageNumber="405">Table 7</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="405" type="materials_examined">
<paragraph blockId="28.[675,1212,655,864]" pageId="28" pageNumber="405">
<materialsCitation collectingDate="2005-06-05" collectingDateMax="2005-06-07" collectingDateMin="2005-06-05" collectionCode="FSCA" collectorName="J. E. Eger &amp; M. T. Messenger" country="French Guiana" elevation="272" latitude="4.5369" location="Roura on Kaw Rd." longLatPrecision="1" longitude="-52.123665" pageId="28" pageNumber="405" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
TYPE MATERIAL. —
<typeStatus box="[856,948,655,679]" pageId="28" pageNumber="405">Holotype</typeStatus>
: “
<collectingCountry box="[963,1166,655,679]" name="French Guiana" pageId="28" pageNumber="405">FRENCH GUIANA</collectingCountry>
: 41 / km SE
<location LSID="urn:lsid:plazi:treatment:03FD8799FFB8EE4BFCEA325966F0BFAF:8E8B6054FFB8EE48FD2332C464C2BE85" box="[743,924,681,705]" country="French Guiana" latitude="4.5369" longLatPrecision="1" longitude="-52.123665" name="Roura on Kaw Rd." pageId="28" pageNumber="405">Roura on Kaw Rd.</location>
, /
<date box="[944,1072,681,705]" pageId="28" pageNumber="405" value="2005-06-05" valueMax="2005-06-07" valueMin="2005-06-05">
<collectingDate box="[944,1072,681,705]" pageId="28" pageNumber="405" value="2005-06-05" valueMax="2005-06-07" valueMin="2005-06-05">5-7-VI-2005</collectingDate>
</date>
,
<collectorName box="[1080,1169,681,705]" pageId="28" pageNumber="405">J. E. Eger</collectorName>
/ &amp;
<collectorName box="[675,834,708,732]" pageId="28" pageNumber="405">M.T. Messenger</collectorName>
, coll. //
<geoCoordinate box="[908,1035,708,732]" degrees="04" direction="north" minutes="32.214" orientation="latitude" pageId="28" pageNumber="405" precision="1" value="4.5369">N04°32.214’</geoCoordinate>
/
<geoCoordinate box="[1051,1196,708,732]" degrees="052" direction="west" minutes="07.420" orientation="longitude" pageId="28" pageNumber="405" precision="1" value="-52.123665">W052°07.420’</geoCoordinate>
/
<quantity box="[675,739,734,758]" metricMagnitude="2" metricUnit="m" metricValue="2.7199999999999998" pageId="28" pageNumber="405" unit="m" value="272.0">
<elevation box="[675,739,734,758]" metricMagnitude="2" metricUnit="m" metricValue="2.7199999999999998" pageId="28" pageNumber="405" unit="m" value="272.0">272 m</elevation>
</quantity>
MV Light”,
<specimenCount box="[867,908,734,758]" pageId="28" pageNumber="405" type="male">1 ♂</specimenCount>
(
<collectionCode box="[921,986,734,758]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="28" pageNumber="405" type="Museum">FSCA</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph blockId="28.[675,1212,655,864]" pageId="28" pageNumber="405">
<typeStatus box="[677,774,760,784]" pageId="28" pageNumber="405">Paratypes</typeStatus>
: same data as
<typeStatus box="[920,1010,760,784]" pageId="28" pageNumber="405">holotype</typeStatus>
,
<specimenCount box="[1023,1064,760,785]" pageId="28" pageNumber="405" type="male">1 ♂</specimenCount>
,
<specimenCount box="[1079,1118,760,785]" pageId="28" pageNumber="405" type="female">1 ♀</specimenCount>
(
<collectionCode box="[1133,1199,760,784]" pageId="28" pageNumber="405">AFSC</collectionCode>
),
<specimenCount box="[678,717,787,811]" pageId="28" pageNumber="405" type="male">1 ♂</specimenCount>
(
<collectionCode box="[733,819,787,811]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="28" pageNumber="405">MNHN</collectionCode>
(
<collectionCode box="[826,863,787,811]" pageId="28" pageNumber="405">EH</collectionCode>
)16226). — “FR.
<collectingCountry box="[1044,1146,787,811]" name="Guyana" pageId="28" pageNumber="405">GUIANA</collectingCountry>
: Hwy. N2 to / Regina,
<quantity box="[835,900,813,837]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="28" pageNumber="405" unit="km" value="45.0">45 km</quantity>
. S of / Cayenne,
<date box="[1063,1171,813,837]" pageId="28" pageNumber="405" value="1986-05-31">31-V-1986</date>
/ E. G. Riley &amp; D. A. Rider”,
<specimenCount box="[929,968,840,864]" pageId="28" pageNumber="405" type="male">1 ♂</specimenCount>
(
<collectionCode box="[981,1052,840,864]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33726" lsid="urn:lsid:biocol.org:col:33726" name="Louisiana State Arthropod Museum" pageId="28" pageNumber="405">LSAM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="405" type="etymology">
<paragraph blockId="28.[674,1209,889,966]" pageId="28" pageNumber="405">
ETYMOLOGY. — The species is named in honor of Michel Boulard who described many new cicada species for
<collectingCountry name="French Guiana" pageId="28" pageNumber="405">French Guiana</collectingCountry>
and the other regions of the world.
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="405" type="distribution">
<paragraph blockId="28.[677,1209,991,1042]" pageId="28" pageNumber="405">
DISTRIBUTION. — The species is only known from
<collectingCountry box="[679,827,1018,1042]" name="French Guiana" pageId="28" pageNumber="405">French Guiana</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="405" type="diagnosis">
<paragraph blockId="28.[677,1212,1067,1699]" pageId="28" pageNumber="405">
DIAGNOSIS. — The male genitalia, particularly the pointed upper pygofer lobe, quickly separate male specimens of
<taxonomicName authorityName="Sanborn" authorityYear="2011" box="[814,923,1119,1143]" class="Insecta" family="Cicadidae" genus="Taphura" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="405" phylum="Arthropoda" rank="species" species="boulardi" status="n. sp.">
<emphasis box="[814,923,1119,1143]" italics="true" pageId="28" pageNumber="405">T. boulardi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[932,988,1120,1144]" pageId="28" pageNumber="405" rank="species">n. sp.</taxonomicNameLabel>
from other species of
<taxonomicName authorityName="Stal" authorityYear="1862" box="[678,757,1146,1170]" class="Insecta" family="Cicadidae" genus="Taphura" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="405" phylum="Arthropoda" rank="genus">
<emphasis box="[678,757,1146,1170]" italics="true" pageId="28" pageNumber="405">Taphura</emphasis>
</taxonomicName>
as each species has highly unique genitalia. All species are generally similar in size and appearance. The spots on the head differentiate the species from all but
<taxonomicName authorityName="Sanborn" authorityYear="2011" box="[677,790,1225,1249]" class="Insecta" family="Cicadidae" genus="Taphura" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="405" phylum="Arthropoda" rank="species" species="maculata" status="n. sp.">
<emphasis box="[677,790,1225,1249]" italics="true" pageId="28" pageNumber="405">T. maculata</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[797,853,1225,1249]" pageId="28" pageNumber="405" rank="species">n. sp.</taxonomicNameLabel>
, the spots are connected into a “W” on the dorsal head of
<taxonomicName authorityName="Sanborn" authorityYear="2011" box="[892,961,1252,1275]" class="Insecta" family="Cicadidae" genus="Taphura" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="405" phylum="Arthropoda" rank="species" species="egeri" status="n. sp.">
<emphasis box="[892,961,1252,1275]" italics="true" pageId="28" pageNumber="405">T. egeri</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[968,1023,1252,1276]" pageId="28" pageNumber="405" rank="species">n. sp.</taxonomicNameLabel>
and the remaining
<taxonomicName box="[677,832,1278,1302]" pageId="28" pageNumber="405">
<emphasis box="[677,758,1278,1302]" italics="true" pageId="28" pageNumber="405">Taphura</emphasis>
species
</taxonomicName>
have transverse piceous marks across the dorsal head; if this transverse mark is incomplete, there are black marks laterally between the lateral ocelli and eye which are absent in
<taxonomicName authorityName="Sanborn" authorityYear="2011" box="[957,1062,1358,1382]" class="Insecta" family="Cicadidae" genus="Taphura" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="405" phylum="Arthropoda" rank="species" species="boulardi" status="n. sp.">
<emphasis box="[957,1062,1358,1382]" italics="true" pageId="28" pageNumber="405">T. boulardi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1070,1124,1358,1382]" pageId="28" pageNumber="405" rank="species">n. sp.</taxonomicNameLabel>
<taxonomicName authorityName="Sanborn" authorityYear="2011" class="Insecta" family="Cicadidae" genus="Taphura" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="405" phylum="Arthropoda" rank="species" species="maculata" status="n. sp.">
<emphasis italics="true" pageId="28" pageNumber="405">Taphura maculata</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[770,824,1384,1408]" pageId="28" pageNumber="405" rank="species">n. sp.</taxonomicNameLabel>
specimens are smaller and lack piceous markings on the lateral disc of the pronotum. Females can be differentiated from the other species of
<taxonomicName pageId="28" pageNumber="405">
<emphasis box="[1130,1209,1437,1461]" italics="true" pageId="28" pageNumber="405">Taphura</emphasis>
species
</taxonomicName>
except
<taxonomicName authority="(Stal, 1854)" baseAuthorityName="Stal" baseAuthorityYear="1854" box="[819,1036,1463,1488]" class="Insecta" family="Cicadidae" genus="Cicada" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="405" phylum="Arthropoda" rank="species" species="misella">
<emphasis box="[819,912,1463,1487]" italics="true" pageId="28" pageNumber="405">T. misella</emphasis>
(
<bibRefCitation author="STAL C." box="[926,1029,1464,1488]" pageId="28" pageNumber="405" pagination="231 - 255" refId="ref30770" refString="STAL C. 1854. - Nya Hemiptera. Ofversigt af Svenska Vetenskaps Akademien Forhandlingar 11: 231 - 255." type="journal article" year="1854">Stål, 1854</bibRefCitation>
)
</taxonomicName>
in the anal styles that are found at the widest point of gonapophysis X, while the broadest portion of gonapophysis X is beyond the anal styles in the other species of
<taxonomicName authorityName="Stal" authorityYear="1862" box="[1124,1203,1543,1567]" class="Insecta" family="Cicadidae" genus="Taphura" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="405" phylum="Arthropoda" rank="genus">
<emphasis box="[1124,1203,1543,1567]" italics="true" pageId="28" pageNumber="405">Taphura</emphasis>
</taxonomicName>
. Sternite VII is doubly notched in
<taxonomicName authorityName="Sanborn" authorityYear="2011" box="[1004,1108,1569,1593]" class="Insecta" family="Cicadidae" genus="Taphura" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="405" phylum="Arthropoda" rank="species" species="boulardi" status="n. sp.">
<emphasis box="[1004,1108,1569,1593]" italics="true" pageId="28" pageNumber="405">T. boulardi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1115,1168,1569,1593]" pageId="28" pageNumber="405" rank="species">n. sp.</taxonomicNameLabel>
and
<taxonomicName authorityName="Sanborn" authorityYear="2011" box="[677,746,1596,1619]" class="Insecta" family="Cicadidae" genus="Taphura" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="405" phylum="Arthropoda" rank="species" species="egeri" status="n. sp.">
<emphasis box="[677,746,1596,1619]" italics="true" pageId="28" pageNumber="405">T. egeri</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[752,806,1596,1620]" pageId="28" pageNumber="405" rank="species">n. sp.</taxonomicNameLabel>
but a single V-shaped notch in
<taxonomicName baseAuthorityName="Stal" baseAuthorityYear="1854" box="[1119,1210,1596,1620]" class="Insecta" family="Cicadidae" genus="Cicada" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="405" phylum="Arthropoda" rank="species" species="misella">
<emphasis box="[1119,1210,1596,1620]" italics="true" pageId="28" pageNumber="405">T. misella</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Sanborn" authorityYear="2011" box="[720,834,1622,1646]" class="Insecta" family="Cicadidae" genus="Taphura" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="405" phylum="Arthropoda" rank="species" species="maculata" status="n.sp.">
<emphasis box="[720,834,1622,1646]" italics="true" pageId="28" pageNumber="405">T. maculata</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[841,888,1622,1646]" pageId="28" pageNumber="405" rank="species">n.sp.</taxonomicNameLabel>
Female
<taxonomicName authorityName="Sanborn" authorityYear="2011" box="[971,1077,1622,1646]" class="Insecta" family="Cicadidae" genus="Taphura" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="405" phylum="Arthropoda" rank="species" species="boulardi" status="n. sp.">
<emphasis box="[971,1077,1622,1646]" italics="true" pageId="28" pageNumber="405">T. boulardi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1085,1139,1622,1646]" pageId="28" pageNumber="405" rank="species">n. sp.</taxonomicNameLabel>
can be differentiated from female
<taxonomicName authorityName="Sanborn" authorityYear="2011" box="[936,1004,1649,1672]" class="Insecta" family="Cicadidae" genus="Taphura" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="405" phylum="Arthropoda" rank="species" species="egeri" status="n. sp.">
<emphasis box="[936,1004,1649,1672]" italics="true" pageId="28" pageNumber="405">T. egeri</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1010,1063,1649,1673]" pageId="28" pageNumber="405" rank="species">n. sp.</taxonomicNameLabel>
by the absence of longitudinal stripes on the postclypeus.
</paragraph>
</subSubSection>
<caption pageId="29" pageNumber="406" startId="29.[115,124,219,236]" targetBox="[113,1198,318,474]" targetIsTable="true" targetPageId="29">
<paragraph blockId="29.[115,1209,219,259]" pageId="29" pageNumber="406">
TABLE 7. — Measurements of
<taxonomicName authorityName="Sanborn" authorityYear="2011" box="[369,511,219,236]" class="Insecta" family="Cicadidae" genus="Taphura" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="406" phylum="Arthropoda" rank="species" species="boulardi" status="n. sp.">
<emphasis box="[369,511,219,236]" italics="true" pageId="29" pageNumber="406">Taphura boulardi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[518,568,219,236]" pageId="29" pageNumber="406" rank="species">n. sp.</taxonomicNameLabel>
in mm, mean (range). Note: width head including eyes; width of pronotum including suprahumeral plates.
</paragraph>
</caption>
<paragraph pageId="29" pageNumber="406">
<table box="[113,1198,318,474]" colsContinueFrom="25.[113,1198,318,474]" colsContinueIn="31.[113,1198,318,474]" gridcols="8" gridrows="5" pageId="29" pageNumber="406">
<tr box="[113,1198,318,337]" gridrow="0" pageId="29" pageNumber="406" rowspan-0="1" rowspan-1="1" rowspan-4="1" rowspan-5="1">
<th box="[401,504,318,337]" gridcol="2" gridrow="0" pageId="29" pageNumber="406">
<emphasis bold="true" box="[409,495,318,337]" pageId="29" pageNumber="406">Tegmina</emphasis>
</th>
<th box="[546,632,318,337]" gridcol="3" gridrow="0" pageId="29" pageNumber="406">
<emphasis bold="true" box="[546,632,318,337]" pageId="29" pageNumber="406">Tegmina</emphasis>
</th>
<th box="[950,1050,318,337]" gridcol="6" gridrow="0" pageId="29" pageNumber="406">
<emphasis bold="true" box="[950,1050,318,337]" pageId="29" pageNumber="406">Pronotum</emphasis>
</th>
<th box="[1077,1198,318,337]" gridcol="7" gridrow="0" pageId="29" pageNumber="406">
<emphasis bold="true" box="[1077,1198,318,337]" pageId="29" pageNumber="406">Mesonotum</emphasis>
</th>
</tr>
<tr box="[113,1198,344,363]" gridrow="1" pageId="29" pageNumber="406" rowspan-0="1">
<td box="[254,376,344,363]" gridcol="1" gridrow="1" pageId="29" pageNumber="406">
<emphasis bold="true" box="[254,376,344,363]" pageId="29" pageNumber="406">Body length</emphasis>
</td>
<td box="[401,504,344,363]" gridcol="2" gridrow="1" pageId="29" pageNumber="406">
<emphasis bold="true" box="[420,484,344,363]" pageId="29" pageNumber="406">length</emphasis>
</td>
<td box="[546,632,344,363]" gridcol="3" gridrow="1" pageId="29" pageNumber="406">
<emphasis bold="true" box="[561,617,344,363]" pageId="29" pageNumber="406">width</emphasis>
</td>
<td box="[665,788,344,363]" gridcol="4" gridrow="1" pageId="29" pageNumber="406">
<emphasis bold="true" box="[665,788,344,363]" pageId="29" pageNumber="406">Head length</emphasis>
</td>
<td box="[805,921,344,363]" gridcol="5" gridrow="1" pageId="29" pageNumber="406">
<emphasis bold="true" box="[805,921,344,363]" pageId="29" pageNumber="406">Head width</emphasis>
</td>
<td box="[950,1050,344,363]" gridcol="6" gridrow="1" pageId="29" pageNumber="406">
<emphasis bold="true" box="[972,1028,344,363]" pageId="29" pageNumber="406">width</emphasis>
</td>
<td box="[1077,1198,344,363]" gridcol="7" gridrow="1" pageId="29" pageNumber="406">
<emphasis bold="true" box="[1109,1165,344,363]" pageId="29" pageNumber="406">width</emphasis>
</td>
</tr>
<tr box="[113,1198,379,399]" gridrow="2" pageId="29" pageNumber="406">
<th box="[113,193,379,399]" gridcol="0" gridrow="2" pageId="29" pageNumber="406">Males</th>
<td box="[254,376,379,399]" gridcol="1" gridrow="2" pageId="29" pageNumber="406">12.06</td>
<td box="[401,504,379,399]" gridcol="2" gridrow="2" pageId="29" pageNumber="406">15.05</td>
<td box="[546,632,379,399]" gridcol="3" gridrow="2" pageId="29" pageNumber="406">5.7</td>
<td box="[665,788,379,399]" gridcol="4" gridrow="2" pageId="29" pageNumber="406">1.5</td>
<td box="[805,921,379,399]" gridcol="5" gridrow="2" pageId="29" pageNumber="406">3.6</td>
<td box="[950,1050,379,399]" gridcol="6" gridrow="2" pageId="29" pageNumber="406">3.75</td>
<td box="[1077,1198,379,399]" gridcol="7" gridrow="2" pageId="29" pageNumber="406">3.35</td>
</tr>
<tr box="[113,1198,404,424]" gridrow="3" pageId="29" pageNumber="406">
<th box="[113,193,404,424]" gridcol="0" gridrow="3" pageId="29" pageNumber="406">(N = 4)</th>
<td box="[254,376,404,424]" gridcol="1" gridrow="3" pageId="29" pageNumber="406">(11.3-13.0)</td>
<td box="[401,504,404,424]" gridcol="2" gridrow="3" pageId="29" pageNumber="406">(14.1-16.0)</td>
<td box="[546,632,404,424]" gridcol="3" gridrow="3" pageId="29" pageNumber="406">(5.0-6.0)</td>
<td box="[665,788,404,424]" gridcol="4" gridrow="3" pageId="29" pageNumber="406">(1.3-1.6)</td>
<td box="[805,921,404,424]" gridcol="5" gridrow="3" pageId="29" pageNumber="406">(3.3-3.8)</td>
<td box="[950,1050,404,424]" gridcol="6" gridrow="3" pageId="29" pageNumber="406">(3.6-3.9)</td>
<td box="[1077,1198,404,424]" gridcol="7" gridrow="3" pageId="29" pageNumber="406">(3.3-3.4)</td>
</tr>
<tr box="[113,1198,428,474]" gridrow="4" pageId="29" pageNumber="406">
<th box="[113,193,428,474]" gridcol="0" gridrow="4" pageId="29" pageNumber="406">Females (N = 1)</th>
<td box="[254,376,428,474]" gridcol="1" gridrow="4" pageId="29" pageNumber="406">13.0</td>
<td box="[401,504,428,474]" gridcol="2" gridrow="4" pageId="29" pageNumber="406">14.1</td>
<td box="[546,632,428,474]" gridcol="3" gridrow="4" pageId="29" pageNumber="406">5.9</td>
<td box="[665,788,428,474]" gridcol="4" gridrow="4" pageId="29" pageNumber="406">1.6</td>
<td box="[805,921,428,474]" gridcol="5" gridrow="4" pageId="29" pageNumber="406">3.8</td>
<td box="[950,1050,428,474]" gridcol="6" gridrow="4" pageId="29" pageNumber="406">3.9</td>
<td box="[1077,1198,428,474]" gridcol="7" gridrow="4" pageId="29" pageNumber="406">3.4</td>
</tr>
</table>
</paragraph>
<subSubSection lastPageId="31" lastPageNumber="408" pageId="29" pageNumber="406" type="description">
<paragraph blockId="29.[111,648,533,1700]" box="[113,258,533,560]" pageId="29" pageNumber="406">DESCRIPTION</paragraph>
<paragraph blockId="29.[111,648,533,1700]" pageId="29" pageNumber="406">
Ground color tawny and ferruginous, may have been altered by chemical exposure during capture or transport. One
<typeStatus box="[311,403,628,655]" pageId="29" pageNumber="406">paratype</typeStatus>
with greenish tinge.
</paragraph>
<paragraph blockId="29.[111,648,533,1700]" pageId="29" pageNumber="406">
Head slightly wider than mesonotum, tawny with fuscous spots posterior to median ocellus on vertex posterior to epicranial suture anterior arm at level of median ocellus and posterolateral to lateral ocelli. Ocelli golden or marked with rosaceus. Eyes golden. Ventral head ground color. Head covered with sparse, short, golden pile. Postclypeus ground color with eight transverse grooves, darker on either side of midline in
<specimenCount box="[290,433,913,940]" pageId="29" pageNumber="406" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
. Anteclypeus ground color. Rostrum with ochraceous mentum, labium tawny becoming darker to piceous tip, reaching to middle trochanters. Scape and annulus on proximal pedicel ground color, remaining antennal segments fuscous. Sparse, long golden pile on postclypeus and anteclypeus.
</paragraph>
<paragraph blockId="29.[111,648,533,1700]" pageId="29" pageNumber="406">
Dorsal thorax ground color. Pronotal disc light ferruginous.Piceous mark along either side of ochraceous midline, curving anteriorly and posteriorly before margins, anterior extension almost completely covering anterior disc to paramedian fissure, posterior extension continues laterad to posterior lateral fissure, extending through lateral fissure in some
<typeStatus box="[174,279,1356,1383]" pageId="29" pageNumber="406">paratypes</typeStatus>
. Piceous marks on anterior disc between paramedian and lateral fissures and anterior margin and centrally of disc lateral to lateral fissure, proportion of discs covered with piceous varies in
<typeStatus box="[113,218,1483,1510]" pageId="29" pageNumber="406">paratypes</typeStatus>
. Pronotal collar ochraceous, lateral portion absent. Mesonotum and metanotum ground color. Sparse, long golden pile between anterior arms and laterally between arms of cruciform elevation. Ventral thoracic segments ground color except darker spot on posterior katepisternum 2, spot absent in some
<typeStatus box="[333,438,1673,1700]" pageId="29" pageNumber="406">paratypes</typeStatus>
.
</paragraph>
<paragraph blockId="29.[674,1212,533,1700]" pageId="29" pageNumber="406">Tegmina and wings hyaline, tegmina with eight apical cells, hind wing with five apical cells. Venation ground color, becoming testaceous distally, except testaceous anal vein 2 + 3. Infuscation spot at apex of apical cell 1. Basal membrane grayish testaceous. Hind wing venation ochraceous proximally, testaceous distally, except testaceous anal vein 3. Vanal fold, anal cell 3 and anal cell 2 along anal veins 2 and 3 gray.</paragraph>
<paragraph blockId="29.[674,1212,533,1700]" pageId="29" pageNumber="406">Legs ground color, tarsi becoming fuscous distally. Linear mark on proximal anterior fore coxae, triangular on middle and hind coxae, smaller linear mark on lateral coxae. Fore femora with four fuscous oblique spines, becoming smaller distally. Pretarsal claws ground color proximally, fuscous distally. Tibial spurs and comb ground color.</paragraph>
<paragraph blockId="29.[674,1212,533,1700]" pageId="29" pageNumber="406">Opercula ground color. Male operculum with lateral margin straight curving to obliquely angled sinuate posterior margin to rounded medial margin and straight anteromedial margin, reaching to middle of tympanal cavity. Medial margin extending to medial margin of meracanthus. Meracanthus ground color, not reaching anteromedial margin of operculum. Female operculum with notched lateral margin,slightly curved posterior margin, terminating medially at lateral meracanthus reaching to middle of tympanal cavity. Long golden pile on operculum surface and radiating from margin.</paragraph>
<paragraph blockId="29.[674,1212,533,1700]" pageId="29" pageNumber="406">Abdomen ground color with short golden pile. Timbal with six ribs.</paragraph>
<paragraph blockId="29.[674,1212,533,1700]" lastBlockId="31.[111,649,533,1003]" lastPageId="31" lastPageNumber="408" pageId="29" pageNumber="406">
Male pygofer ground color with light fuscous dorsal beak and margin between dorsal beak and upper lobe of pygofer, ground color in some
<typeStatus pageId="29" pageNumber="406">paratypes</typeStatus>
. Sternite VIII ground color with long golden pile. Pygofer upper lobe broad at base, sides parallel to dorsal beak, anterior margin slightly sinuate extending to point at level of dorsal beak with long pile, posterior margin half the length of anterior margin curving and joining transversely to anterior margin. Median uncus lobes very small around fuscous anal styles. Aedeagus dark tawny, proportionately large.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7671518" ID-Zenodo-Dep="7671518" httpUri="https://zenodo.org/record/7671518/files/figure.png" pageId="30" pageNumber="407" startId="30.[113,124,1635,1652]" targetBox="[142,1181,214,1575]" targetPageId="30">
<paragraph blockId="30.[113,1210,1635,1698]" pageId="30" pageNumber="407">
FIG. 7. —
<taxonomicName authorityName="Sanborn" authorityYear="2011" box="[193,331,1635,1652]" class="Insecta" family="Cicadidae" genus="Taphura" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="407" phylum="Arthropoda" rank="species" species="boulardi" status="n. sp.">
<emphasis box="[193,331,1635,1652]" italics="true" pageId="30" pageNumber="407">Taphura boulardi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[336,386,1635,1652]" pageId="30" pageNumber="407" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis bold="true" box="[392,405,1635,1652]" pageId="30" pageNumber="407">A</emphasis>
, holotype ♂ and paratype ♀ habitus;
<emphasis bold="true" box="[717,730,1635,1652]" pageId="30" pageNumber="407">B</emphasis>
, holotype ♂ dorsum;
<emphasis bold="true" box="[911,925,1635,1652]" pageId="30" pageNumber="407">C</emphasis>
, holotype ♂ timbal;
<emphasis bold="true" box="[1093,1107,1635,1652]" pageId="30" pageNumber="407">D</emphasis>
, holotype ♂ operculum;
<emphasis bold="true" box="[212,224,1658,1675]" pageId="30" pageNumber="407">E</emphasis>
, paratype ♀ operculum;
<emphasis bold="true" box="[430,441,1658,1675]" pageId="30" pageNumber="407">F</emphasis>
, paratype ♂ lateral view of genitalia;
<emphasis bold="true" box="[748,762,1658,1675]" pageId="30" pageNumber="407">G</emphasis>
, paratype ♂ posterior view of genitalia;
<emphasis bold="true" box="[1094,1108,1658,1675]" pageId="30" pageNumber="407">H</emphasis>
, paratype ♀ lateral view of genitalia;
<emphasis bold="true" box="[313,319,1680,1697]" pageId="30" pageNumber="407">I</emphasis>
, paratype ♀ ventral view of genitalia. Scale bars: A, 1 cm; B-I, 1 mm.
</paragraph>
</caption>
<caption pageId="31" pageNumber="408" startId="31.[113,124,219,236]" targetBox="[113,1198,318,474]" targetIsTable="true" targetPageId="31">
<paragraph blockId="31.[113,1209,219,259]" pageId="31" pageNumber="408">
TABLE 8. — Measurements of
<taxonomicName authorityName="Sanborn" authorityYear="2011" box="[356,468,219,236]" class="Insecta" family="Cicadidae" genus="Taphura" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="408" phylum="Arthropoda" rank="species" species="egeri" status="n. sp.">
<emphasis box="[356,468,219,236]" italics="true" pageId="31" pageNumber="408">Taphura egeri</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[473,520,219,236]" pageId="31" pageNumber="408" rank="species">n. sp.</taxonomicNameLabel>
in mm, mean (range). Note: width head including eyes; width of pronotum including suprahumeral plates.
</paragraph>
</caption>
<paragraph pageId="31" pageNumber="408">
<table box="[113,1198,318,474]" colsContinueFrom="29.[113,1198,318,474]" colsContinueIn="35.[113,1198,318,474]" gridcols="8" gridrows="6" pageId="31" pageNumber="408">
<tr box="[113,1198,318,337]" gridrow="0" pageId="31" pageNumber="408" rowspan-0="1" rowspan-1="1" rowspan-4="1" rowspan-5="1">
<th box="[401,503,318,337]" gridcol="2" gridrow="0" pageId="31" pageNumber="408">
<emphasis bold="true" box="[409,495,318,337]" pageId="31" pageNumber="408">Tegmina</emphasis>
</th>
<th box="[546,632,318,337]" gridcol="3" gridrow="0" pageId="31" pageNumber="408">
<emphasis bold="true" box="[546,632,318,337]" pageId="31" pageNumber="408">Tegmina</emphasis>
</th>
<th box="[950,1050,318,337]" gridcol="6" gridrow="0" pageId="31" pageNumber="408">
<emphasis bold="true" box="[950,1050,318,337]" pageId="31" pageNumber="408">Pronotum</emphasis>
</th>
<th box="[1077,1198,318,337]" gridcol="7" gridrow="0" pageId="31" pageNumber="408">
<emphasis bold="true" box="[1077,1198,318,337]" pageId="31" pageNumber="408">Mesonotum</emphasis>
</th>
</tr>
<tr box="[113,1198,344,363]" gridrow="1" pageId="31" pageNumber="408" rowspan-0="1">
<td box="[254,376,344,363]" gridcol="1" gridrow="1" pageId="31" pageNumber="408">
<emphasis bold="true" box="[254,376,344,363]" pageId="31" pageNumber="408">Body length</emphasis>
</td>
<td box="[401,503,344,363]" gridcol="2" gridrow="1" pageId="31" pageNumber="408">
<emphasis bold="true" box="[420,484,344,363]" pageId="31" pageNumber="408">length</emphasis>
</td>
<td box="[546,632,344,363]" gridcol="3" gridrow="1" pageId="31" pageNumber="408">
<emphasis bold="true" box="[561,617,344,363]" pageId="31" pageNumber="408">width</emphasis>
</td>
<td box="[665,788,344,363]" gridcol="4" gridrow="1" pageId="31" pageNumber="408">
<emphasis bold="true" box="[665,788,344,363]" pageId="31" pageNumber="408">Head length</emphasis>
</td>
<td box="[805,921,344,363]" gridcol="5" gridrow="1" pageId="31" pageNumber="408">
<emphasis bold="true" box="[805,921,344,363]" pageId="31" pageNumber="408">Head width</emphasis>
</td>
<td box="[950,1050,344,363]" gridcol="6" gridrow="1" pageId="31" pageNumber="408">
<emphasis bold="true" box="[972,1028,344,363]" pageId="31" pageNumber="408">width</emphasis>
</td>
<td box="[1077,1198,344,363]" gridcol="7" gridrow="1" pageId="31" pageNumber="408">
<emphasis bold="true" box="[1109,1165,344,363]" pageId="31" pageNumber="408">width</emphasis>
</td>
</tr>
<tr box="[113,1198,378,398]" gridrow="2" pageId="31" pageNumber="408">
<th box="[113,193,378,398]" gridcol="0" gridrow="2" pageId="31" pageNumber="408">Males</th>
<td box="[254,376,378,398]" gridcol="1" gridrow="2" pageId="31" pageNumber="408">13.0</td>
<td box="[401,503,378,398]" gridcol="2" gridrow="2" pageId="31" pageNumber="408">17.19</td>
<td box="[546,632,378,398]" gridcol="3" gridrow="2" pageId="31" pageNumber="408">5.72</td>
<td box="[665,788,378,398]" gridcol="4" gridrow="2" pageId="31" pageNumber="408">2.06</td>
<td box="[805,921,378,398]" gridcol="5" gridrow="2" pageId="31" pageNumber="408">3.97</td>
<td box="[950,1050,378,398]" gridcol="6" gridrow="2" pageId="31" pageNumber="408">4.34</td>
<td box="[1077,1198,378,398]" gridcol="7" gridrow="2" pageId="31" pageNumber="408">3.71</td>
</tr>
<tr box="[113,1198,404,424]" gridrow="3" pageId="31" pageNumber="408">
<th box="[113,193,404,424]" gridcol="0" gridrow="3" pageId="31" pageNumber="408">(N = 10)</th>
<td box="[254,376,404,424]" gridcol="1" gridrow="3" pageId="31" pageNumber="408">(11.8-14.0)</td>
<td box="[401,503,404,424]" gridcol="2" gridrow="3" pageId="31" pageNumber="408">(15.7-18.0)</td>
<td box="[546,632,404,424]" gridcol="3" gridrow="3" pageId="31" pageNumber="408">(5.3-6.0)</td>
<td box="[665,788,404,424]" gridcol="4" gridrow="3" pageId="31" pageNumber="408">(2.0-2.2)</td>
<td box="[805,921,404,424]" gridcol="5" gridrow="3" pageId="31" pageNumber="408">(3.8-4.1)</td>
<td box="[950,1050,404,424]" gridcol="6" gridrow="3" pageId="31" pageNumber="408">(4.2-4.4)</td>
<td box="[1077,1198,404,424]" gridcol="7" gridrow="3" pageId="31" pageNumber="408">(3.6-3.8)</td>
</tr>
<tr box="[113,1198,428,448]" gridrow="4" pageId="31" pageNumber="408">
<th box="[113,193,428,448]" gridcol="0" gridrow="4" pageId="31" pageNumber="408">Females</th>
<td box="[254,376,428,448]" gridcol="1" gridrow="4" pageId="31" pageNumber="408">14.17</td>
<td box="[401,503,428,448]" gridcol="2" gridrow="4" pageId="31" pageNumber="408">17.93</td>
<td box="[546,632,428,448]" gridcol="3" gridrow="4" pageId="31" pageNumber="408">6.1</td>
<td box="[665,788,428,448]" gridcol="4" gridrow="4" pageId="31" pageNumber="408">2.07</td>
<td box="[805,921,428,448]" gridcol="5" gridrow="4" pageId="31" pageNumber="408">4.35</td>
<td box="[950,1050,428,448]" gridcol="6" gridrow="4" pageId="31" pageNumber="408">4.43</td>
<td box="[1077,1198,428,448]" gridcol="7" gridrow="4" pageId="31" pageNumber="408">3.0</td>
</tr>
<tr box="[113,1198,454,474]" gridrow="5" pageId="31" pageNumber="408">
<th box="[113,193,454,474]" gridcol="0" gridrow="5" pageId="31" pageNumber="408">(N = 10)</th>
<td box="[254,376,454,474]" gridcol="1" gridrow="5" pageId="31" pageNumber="408">(13.1-15.2)</td>
<td box="[401,503,454,474]" gridcol="2" gridrow="5" pageId="31" pageNumber="408">(16.7-18.8)</td>
<td box="[546,632,454,474]" gridcol="3" gridrow="5" pageId="31" pageNumber="408">(5.8-6.6)</td>
<td box="[665,788,454,474]" gridcol="4" gridrow="5" pageId="31" pageNumber="408">(2.0-2.1)</td>
<td box="[805,921,454,474]" gridcol="5" gridrow="5" pageId="31" pageNumber="408">(4.2-4.6)</td>
<td box="[950,1050,454,474]" gridcol="6" gridrow="5" pageId="31" pageNumber="408">(4.3-4.6)</td>
<td box="[1077,1198,454,474]" gridcol="7" gridrow="5" pageId="31" pageNumber="408">(3.7-4.0)</td>
</tr>
</table>
</paragraph>
<paragraph blockId="31.[111,649,533,1003]" pageId="31" pageNumber="408">Female genitalia ground color covered with short golden pile, gonapophyses darker. Sternite VII with a single notch, sinuate posterior margin curving and angled lateral portion of posterior margin. Dorsal beak fuscous extending to level of fuscous anal styles. Dorsal beak produces approximate right angle with posterior margin of abdominal segment 9. Ovipositor sheath with long golden pile, extending beyond dorsal beak.</paragraph>
<paragraph blockId="31.[111,649,533,1003]" box="[136,430,976,1003]" pageId="31" pageNumber="408">
Measurements: see
<tableCitation box="[343,426,976,1003]" captionStart="TABLE" captionStartId="29.[115,124,219,236]" captionTargetPageId="29" captionText="TABLE 7. — Measurements of Taphura boulardi n. sp. in mm, mean (range). Note: width head including eyes; width of pronotum including suprahumeral plates." pageId="31" pageNumber="408">Table 7</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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