treatments-xml/data/03/F4/DF/03F4DF38FFEC664CFF506CDFB86E51F4.xml

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<document id="61DE0D2E679DD5BEF27C26077DC520A8" ID-CLB-Dataset="304981" ID-DOI="10.11646/zootaxa.5536.2.1" ID-GBIF-Dataset="903c9627-28ff-4bfb-9c49-d911e9673c71" ID-ISSN="1175-5326" ID-Zenodo-Dep="14033618" ID-ZooBank="5F1FBB59-9C69-4E66-9B0D-69A015F30EAA" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1730712964139" checkinUser="plazi" docAuthor="ALEKSEEV, VITALII, MCKELLAR, RYAN C. &amp; BUKEJS, ANDRIS" docDate="2024" docId="03F4DF38FFEC664CFF506CDFB86E51F4" docLanguage="en" docName="zootaxa.5536.2.1.pdf" docOrigin="Zootaxa 5536 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5536.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lasconotus tenebrisilvarum Alekseev et Bukejs 2024, sp. nov." docType="treatment" docVersion="4" lastPageNumber="216" masterDocId="FFCDA740FFE06643FFC76A41BB4B5607" masterDocTitle="A revision and addition to Zopheridae (Coleoptera: Tenebrionoidea) in Baltic amber: possible connections between modern Holarctic distributions and Eocene amber forests" masterLastPageNumber="247" masterPageNumber="201" pageNumber="213" updateTime="1730817957888" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7F5A3D3A47CE0C847DA227F13520D074">A revision and addition to Zopheridae (Coleoptera: Tenebrionoidea) in Baltic amber: possible connections between modern Holarctic distributions and Eocene amber forests</mods:title>
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<mods:namePart id="574CEA6BBBF002B17F77785E8EB01300">ALEKSEEV, VITALII</mods:namePart>
<mods:nameIdentifier id="77C8A822A56FEBECE9F622EBB1BBF6B9" type="ORCID">0000-0003-4390-5443</mods:nameIdentifier>
<mods:affiliation id="AB8A8F5BFFD401A62FD63709014431FF">Immanuel Kant Baltic Federal University, Nevskogo 14, Kaliningrad 236016, Russia.</mods:affiliation>
<mods:nameIdentifier id="C09F281C12ACC551E9E0B07641F718A5" type="email">alekseew0802@yahoo.com</mods:nameIdentifier>
</mods:name>
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<mods:role id="E706B2BE1EE00927DB026CEAFFE5E597">
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<mods:namePart id="B484FE27D1B0213EC92228AE3B62978E">MCKELLAR, RYAN C.</mods:namePart>
<mods:nameIdentifier id="CFD6701D6B3E1A445F7EC753A74E8218" type="ORCID">0000-0002-7451-4368</mods:nameIdentifier>
<mods:affiliation id="C2B204D6EB5C792D1BE97EE6F757B45B">Royal Saskatchewan Museum, 2445 Albert St., Regina, SK, S 4 P 4 W 7, Canada. &amp; Biology Department, University of Regina, Regina, SK, S 4 S 0 A 2, Canada; &amp; Department of Ecology &amp; Evolutionary Biology, University of Kansas, Lawrence, Kansas, 66045, USA;</mods:affiliation>
<mods:nameIdentifier id="B412452AD4D00F9CD91801861677CF2B" type="email">ryan.mckellar@gov.sk.ca</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="E1541EBF7C6CDCE6EF2F1BF2546B544F">BUKEJS, ANDRIS</mods:namePart>
<mods:nameIdentifier id="8FD441F6F66D0B0F4CD3A78BE865D61E" type="ORCID">0000-0001-7165-3023</mods:nameIdentifier>
<mods:affiliation id="4EA2029FD9E3D2DE0BBE80320D8F963A">Institute of Life Sciences and Technologies, Daugavpils University, Vienības 13, Daugavpils LV- 5401, Latvia.</mods:affiliation>
<mods:nameIdentifier id="15D62A58194F3E3364714ACB32F34523" type="email">carabidae@inbox.lv</mods:nameIdentifier>
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<mods:date id="2FC03DB0913B4AE3A6F4A777E7347B15">2024</mods:date>
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<treatment id="03F4DF38FFEC664CFF506CDFB86E51F4" ID-DOI="http://doi.org/10.5281/zenodo.14040778" ID-Zenodo-Dep="14040778" LSID="urn:lsid:plazi:treatment:03F4DF38FFEC664CFF506CDFB86E51F4" httpUri="http://treatment.plazi.org/id/03F4DF38FFEC664CFF506CDFB86E51F4" lastPageId="15" lastPageNumber="216" pageId="12" pageNumber="213">
<subSubSection id="C3473DA5FFEC664FFF506CDFB87450B0" box="[151,831,1693,1720]" pageId="12" pageNumber="213" type="nomenclature">
<paragraph id="8BE26E2EFFEC664FFF506CDFB87450B0" blockId="12.[151,831,1693,1755]" box="[151,831,1693,1720]" pageId="12" pageNumber="213">
<heading id="D0AAD942FFEC664FFF506CDFB87450B0" bold="true" box="[151,831,1693,1720]" fontSize="11" level="1" pageId="12" pageNumber="213" reason="1">
<emphasis id="B929B23CFFEC664FFF506CDFB87450B0" bold="true" box="[151,831,1693,1720]" pageId="12" pageNumber="213">
<taxonomicName id="4C5D15ADFFEC664FFF506CDFB99250B0" authority="Alekseev et Bukejs" authorityName="Alekseev et Bukejs" authorityYear="2024" box="[151,729,1693,1720]" class="Insecta" family="Zopheridae" genus="Lasconotus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="tenebrisilvarum" status="sp. nov.">
<emphasis id="B929B23CFFEC664FFF506CDFBAA350BF" bold="true" box="[151,488,1693,1720]" italics="true" pageId="12" pageNumber="213">Lasconotus tenebrisilvarum</emphasis>
Alekseev et Bukejs
</taxonomicName>
<taxonomicNameLabel id="A21A0F47FFEC664FFD266CDCB87450B0" box="[737,831,1693,1719]" pageId="12" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFEC664FFF506C80BA6D50DC" box="[151,294,1729,1755]" pageId="12" pageNumber="213" type="description">
<paragraph id="8BE26E2EFFEC664FFF506C80BA6D50DC" blockId="12.[151,831,1693,1755]" box="[151,294,1729,1755]" pageId="12" pageNumber="213">
(
<figureCitation id="136672ABFFEC664FFF586C80BA5550DC" box="[159,286,1729,1755]" captionStart-0="FIGURES 1113" captionStart-1="FIGURES 1416" captionStartId-0="13.[151,264,1920,1944]" captionStartId-1="16.[151,264,1499,1523]" captionTargetBox-0="[151,1435,179,1895]" captionTargetBox-1="[151,1435,179,1473]" captionTargetId-0="figure-17@13.[151,1435,179,1895]" captionTargetId-1="figure-21@16.[151,1435,179,1473]" captionTargetPageId-0="13" captionTargetPageId-1="16" captionText-0="FIGURES 1113. Lasconotus tenebrisilvarum sp. nov., holotype, No. RSKM_PAL_A73 [RSM]: 11—habitus, dorsal view; 12—habitus, ventral view; 13—habitus, right lateral view. Scale bar = 0.5 mm." captionText-1="FIGURES 1416. Lasconotus tenebrisilvarum sp. nov., holotype, No. RSKM_PAL_A73 [RSM]: 14— habitus, left lateral view; 15—habitus, frontal view; 16— details of head and thorax, ventral view. Abbreviations: a1a11—antennomeres 111, respectively; pp—prosternal process; tmp—terminal maxillary palpomere. Scale bars = 0.5 mm for Figs 14 and 15; 0.25 mm for Fig. 16." figureDoi-0="http://doi.org/10.5281/zenodo.14033630" figureDoi-1="http://doi.org/10.5281/zenodo.14033634" httpUri-0="https://zenodo.org/record/14033630/files/figure.png" httpUri-1="https://zenodo.org/record/14033634/files/figure.png" pageId="12" pageNumber="213">Figs 1116</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFEC664DFF506D48BBA2571A" lastPageId="14" lastPageNumber="215" pageId="12" pageNumber="213" type="materials_examined">
<paragraph id="8BE26E2EFFEC664FFF506D48BEDF51B3" blockId="12.[151,1437,1801,1972]" pageId="12" pageNumber="213">
<emphasis id="B929B23CFFEC664FFF506D48BA055124" bold="true" box="[151,334,1801,1827]" pageId="12" pageNumber="213">Type material.</emphasis>
<materialsCitation id="3B356473FFEC664FFE9A6D48BE355140" collectionCode="RSM" country="Russia" county="Peninsula" determinerName="Alekseev et Bukejs" location="Yantarny" municipality="Sambian" pageId="12" pageNumber="213" specimenCode="RSKM_PAL_A73, JDC-13074" specimenCount="1" specimenCount-adult="1" stateProvince="Kaliningrad Region" typeStatus="holotype">
<typeStatus id="54E6D08CFFEC664FFE9A6D48BA815124" box="[349,458,1801,1827]" pageId="12" pageNumber="213" type="holotype">Holotype</typeStatus>
: No.
<specimenCode id="DBFBC655FFEC664FFDD26D48B9A25124" box="[533,745,1801,1827]" pageId="12" pageNumber="213">RSKM_PAL_A73</specimenCode>
[
<collectionCode id="ED4CF6EBFFEC664FFD286D48B8605124" box="[751,811,1801,1827]" country="United Kingdom" httpUri="http://grbio.org/cool/7sv4-pazk" name="Royal Scottish Museum" pageId="12" pageNumber="213">RSM</collectionCode>
] (ex coll. Jonas Damzen
<specimenCode id="DBFBC655FFEC664FFBA86D48BFBE5124" box="[1135,1269,1801,1827]" pageId="12" pageNumber="213">JDC-13074</specimenCode>
); “Holotype /
<taxonomicName id="4C5D15ADFFEC664FFF506D6FBA9A5140" authorityName="Alekseev &amp; Bukejs" authorityYear="2024" box="[151,465,1837,1863]" class="Insecta" family="Zopheridae" genus="Lasconotus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="tenebrisilvarum" status="sp. nov.">
<emphasis id="B929B23CFFEC664FFF506D6FBA9A5140" box="[151,465,1837,1863]" italics="true" pageId="12" pageNumber="213">Lasconotus tenebrisilvarum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A0F47FFEC664FFE1C6D6FB97E514F" box="[475,565,1838,1864]" pageId="12" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
/
<determinerName id="7E3E82EAFFEC664FFD896D6CB8675140" box="[590,812,1837,1864]" pageId="12" pageNumber="213">Alekseev et Bukejs</determinerName>
des. 2024” [red printed label];
<specimenCount id="9D5BA5A7FFEC664FFB5D6D6CBF9E5140" box="[1178,1237,1837,1863]" pageId="12" pageNumber="213" type="adult">adult</specimenCount>
, sex unknown
</materialsCitation>
. A complete beetle with exposed metathoracic wings is included in a transparent, yellow amber piece with dimensions of 10×
<quantity id="4CA5C3CBFFEC664FFF226D34BA6C5197" box="[229,295,1909,1936]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="213" unit="mm" value="5.0">5 mm</quantity>
and a maximum thickness of
<quantity id="4CA5C3CBFFEC664FFD416D34B9875197" box="[646,716,1909,1936]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="213" unit="mm" value="5.0">5 mm</quantity>
, preserved without supplementary fixation. Syninclusions:
<specimenCount id="9D5BA5A7FFEC664FFAB36D37BA4A51B4" count="1" pageId="12" pageNumber="213" type="generic">one specimen</specimenCount>
of a phoretic uropod deutonymph (Acari:
<taxonomicName id="4C5D15ADFFEC664FFD1B6DD8B83551B4" authorityName="G.Canestrini" authorityYear="1891" box="[732,894,1945,1971]" class="Arachnida" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="order">Mesostigmata</taxonomicName>
:
<taxonomicName id="4C5D15ADFFEC664FFC4F6DD8BF4951B4" authorityName="Kramer" authorityYear="1881" box="[904,1026,1945,1971]" class="Arachnida" infraOrder="Uropodina" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="213" phylum="Arthropoda" rank="infraOrder">Uropodina</taxonomicName>
) attached to right side of pronotum.
</paragraph>
<caption id="DF223EA6FFED664EFF506DC1B8F251BB" ID-DOI="http://doi.org/10.5281/zenodo.14033630" ID-Zenodo-Dep="14033630" httpUri="https://zenodo.org/record/14033630/files/figure.png" pageId="13" pageNumber="214" startId="13.[151,264,1920,1944]" targetBox="[151,1435,179,1895]" targetPageId="13" targetType="figure">
<paragraph id="8BE26E2EFFED664EFF506DC1B8F251BB" blockId="13.[151,1436,1920,1981]" pageId="13" pageNumber="214">
<emphasis id="B929B23CFFED664EFF506DC1BA1E519E" bold="true" box="[151,341,1920,1945]" pageId="13" pageNumber="214">FIGURES 1113.</emphasis>
<taxonomicName id="4C5D15ADFFED664EFE9A6DC0B932519E" authorityName="Alekseev &amp; Bukejs" authorityYear="2024" box="[349,633,1921,1945]" class="Insecta" family="Zopheridae" genus="Lasconotus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="tenebrisilvarum" status="sp. nov.">
<emphasis id="B929B23CFFED664EFE9A6DC0B932519E" box="[349,633,1921,1945]" italics="true" pageId="13" pageNumber="214">Lasconotus tenebrisilvarum</emphasis>
</taxonomicName>
<emphasis id="B929B23CFFED664EFD446DC0B99E519E" bold="true" box="[643,725,1921,1945]" pageId="13" pageNumber="214">
<taxonomicNameLabel id="A21A0F47FFED664EFD446DC0B99E519E" box="[643,725,1921,1945]" pageId="13" pageNumber="214" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, No. RSKM_PAL_A73 [RSM]: 11—habitus, dorsal view; 12—habitus, ventral view; 13—habitus, right lateral view. Scale bar = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BE26E2EFFEE664DFF006AD6BAEF56D2" blockId="14.[151,1437,151,1941]" pageId="14" pageNumber="215">
<emphasis id="B929B23CFFEE664DFF006AD6BA3856B6" bold="true" box="[199,371,151,177]" pageId="14" pageNumber="215">
<typeStatus id="54E6D08CFFEE664DFF006AD6BA4956B6" box="[199,258,151,177]" pageId="14" pageNumber="215">Type</typeStatus>
stratum.
</emphasis>
Baltic amber from Eocene amber-bearing Blaue Erde deposits; estimated age: middlelate Eocene (Standke 1998).
</paragraph>
<paragraph id="8BE26E2EFFEE664DFF006A9EBBA2571A" blockId="14.[151,1437,151,1941]" pageId="14" pageNumber="215">
<emphasis id="B929B23CFFEE664DFF006A9EBA2E56FF" bold="true" box="[199,357,222,249]" pageId="14" pageNumber="215">Type locality.</emphasis>
<location id="8E8238F5FFEE664DFEAD6A9EBA8456FE" LSID="urn:lsid:plazi:treatment:03F4DF38FFEC664CFF506CDFB86E51F4:8E8238F5FFEE664DFEAD6A9EBA8456FE" box="[362,463,223,249]" country="Russia" county="Peninsula" municipality="Sambian" name="Yantarny" pageId="14" pageNumber="215" stateProvince="Kaliningrad Region">Yantarny</location>
settlement (formerly
<location id="8E8238F5FFEE664DFD076A9EB80256FE" LSID="urn:lsid:plazi:treatment:03F4DF38FFEC664CFF506CDFB86E51F4:8E8238F5FFEE664DFD076A9EB80256FE" box="[704,841,223,249]" country="Russia" county="Peninsula" municipality="Sambian" name="Palmnicken" pageId="14" pageNumber="215" stateProvince="Kaliningrad Region">Palmnicken</location>
),
<collectingMunicipality id="6B86F454FFEE664DFC9C6A9EB8F656FE" box="[859,957,223,249]" pageId="14" pageNumber="215">Sambian</collectingMunicipality>
(Samland)
<collectingCounty id="628316A2FFEE664DFBF96A9EBFE556FE" box="[1086,1198,223,249]" pageId="14" pageNumber="215">Peninsula</collectingCounty>
,
<collectingRegion id="4999A0CCFFEE664DFB7F6A9EBED356FE" box="[1208,1432,223,249]" country="Russia" name="Kaliningrad" pageId="14" pageNumber="215">Kaliningrad Region</collectingRegion>
,
<collectingCountry id="F34A2EBEFFEE664DFF506B42BBAE571A" box="[151,229,259,285]" name="Russia" pageId="14" pageNumber="215">Russia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFEE664CFF006B67BE6D57F4" lastPageId="15" lastPageNumber="216" pageId="14" pageNumber="215" type="description">
<paragraph id="8BE26E2EFFEE664DFF006B67BB8157AA" blockId="14.[151,1437,151,1941]" pageId="14" pageNumber="215">
<emphasis id="B929B23CFFEE664DFF006B67BA1A5747" bold="true" box="[199,337,294,320]" pageId="14" pageNumber="215">Description</emphasis>
. Measurements: total body length (including visible part of head)
<quantity id="4CA5C3CBFFEE664DFBF86B67BFDC5746" box="[1087,1175,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="14" pageNumber="215" unit="mm" value="3.5">3.5 mm</quantity>
, maximum body width (across elytra)
<quantity id="4CA5C3CBFFEE664DFEFA6B0BBAEF5762" box="[317,420,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="14" pageNumber="215" unit="mm" value="1.15">1.15 mm</quantity>
; head length
<quantity id="4CA5C3CBFFEE664DFDF06B0AB9C55762" box="[567,654,331,357]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="215" unit="mm" value="0.4">0.4 mm</quantity>
, head width (across eyes)
<quantity id="4CA5C3CBFFEE664DFC746B0ABF475762" box="[947,1036,331,357]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="215" unit="mm" value="0.8">0.8 mm</quantity>
; pronotal length
<quantity id="4CA5C3CBFFEE664DFB0F6B0ABE545762" box="[1224,1311,331,357]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="215" unit="mm" value="0.8">0.8 mm</quantity>
, maximum pronotal width 1.0 mm, basal pronotal width 1.0 mm; elytral length (along elytral suture, including scutellum)
<quantity id="4CA5C3CBFFEE664DFABE6B2EBB8D57AA" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="14" pageNumber="215" unit="mm" value="2.3">2.3 mm</quantity>
.
</paragraph>
<paragraph id="8BE26E2EFFEE664DFF006BF6BF09541E" blockId="14.[151,1437,151,1941]" pageId="14" pageNumber="215">Body subparallel-sided, elongate, about 3.1× as long as wide, weakly convex dorsally and ventrally, unicolorous black (as preserved). Pubescence: head and pronotum with fine, curved setae; elytra sparsely covered with slightly longer, semierect, narrow, uniform setae arranged in rows, and not forming patches.</paragraph>
<paragraph id="8BE26E2EFFEE664DFF006862BF415232" blockId="14.[151,1437,151,1941]" pageId="14" pageNumber="215">Head prognathous, transverse; densely granulose dorsally, with granules rather large near center of frons, about 2× as wide as diameter of one eye facet, distance between granules distinctly smaller than diameter of one granule, each granule bearing short, curved, narrow seta. Anterior margin of clypeus widely rounded. Compound eyes large, nearly hemispherical, strongly convex, without interfacetal setation. Frons with strong, complete supraocular carinae. Antennal insertions concealed dorsally. Antennal groove ventrad eye absent. Antennae short, extending to about middle of pronotal length; 11-segmented with distinct, loose, flattened, 3-segmented club; sparsely covered with short, semierect setae; scape and pedicel wide, cylindrical, about 1.5× as long as wide; antennomere 3 conical, slightly dilated apically, about 1.4× longer than antennomere 4, narrower than pedicel; antennomeres 48 slightly elongate, subcylindrical to subtrapezoidal, slightly dilated apically, subequal in length; antennomere 9 dilated apically, strongly transverse, 2.2× as wide as long, about 2.0× wider than antennomere 8; antennomere 10 widest, strongly transverse, 2.3× as wide as long, about 1.3× wider than antennomere 9; antennomere 11 widely oval, transverse, about 1.3× as wide as long, distinctly narrower than antennomere 10, finely setose, with widely rounded apex. Maxillary palpi with four palpomeres; palpomere 1 smallest, subtriangular; palpomeres 2 and 3 subquadratic, nearly as long as wide, subequal in size and shape; palpomere 4 elongate oval, truncate, about as long as previous palpomeres combined. Terminal labial palpomere subtriangular, acute apically.</paragraph>
<paragraph id="8BE26E2EFFEE664DFF006E7EB957502A" blockId="14.[151,1437,151,1941]" pageId="14" pageNumber="215">Pronotum slightly transverse, 1.2× as wide as long, widest at base, almost parallel-sided, slightly narrowed anteriad, maximum pronotal width / basal pronotal width = 1.0; pronotal disc weakly convex, with two pairs of sharp, uninterrupted, sublateral carinae; pronotal surface densely covered with granules, bearing curved setae, distance between granules smaller than diameter of one granule. Outer pair of pronotal sublateral carinae subparallel, slightly curved, and almost forming subrectangular area together with anterior and posterior pronotal margins; inner pair of pronotal carinae curved, forming longitudinal oval median area with truncated ends, interrupted anteriorly and posteriorly. Lateral margins finely crenulate, raised, forming distinct longitudinal depression between the lateral margin and first pair of pronotal sublateral carina; anterior pronotal margin arcuate, slightly raised medially; posterior margin rounded. Anterior pronotal angles prominent, acute, slightly rounded; posterior angles almost rectangular and not projecting posteriorly. Prothoracic notosternal suture complete. Hypomera densely and coarsely granulose, distance between granules smaller than diameter of one granule. Prosternum convex; densely covered with small granules. Intercoxal prosternal process elongate, with rounded apex, narrow, about 0.7× as wide as diameter of procoxa, with fine median carina in apical portion, not expanded apically beyond midpoint of procoxa. Procoxal cavities narrowly open posteriorly.</paragraph>
<paragraph id="8BE26E2EFFEE664DFF006C76B8B75056" blockId="14.[151,1437,151,1941]" box="[199,1020,1591,1617]" pageId="14" pageNumber="215">Scutellar shield minute, drop-shaped, slightly transverse, dilated apically.</paragraph>
<paragraph id="8BE26E2EFFEE664DFF006C1ABFCA5192" blockId="14.[151,1437,151,1941]" pageId="14" pageNumber="215">Elytra almost parallel-sided in anterior two-thirds of their lengths, narrowed in posterior one-third, rounded at apex; about 2.0× as long as wide combined; carinate and convex; 2.8× longer than pronotum. Elytral base slightly wider than pronotal posterior margin. Humeral angles rounded. Elytra striate-punctate. Scutellary striole present. Elytral striae with rows of elongate granules; odd elytral intervals (1, 3, 5, 7) carinate; elytral carinae similarly raised along their entire lengths. Epipleura present, well-developed, widest in basal half, narrowing posteriad, incomplete, apparently reaching abdominal ventrite 5, densely covered with round granules. Mesoventrite convex, densely and coarsely punctate. Mesocoxal cavities closed. Metaventrite with disc almost flat, longer than abdominal ventrite 1, with dense and coarse punctation; discrimen distinct in posterior two-thirds. Relative length ratio of pro- to meso- to metaventrite to abdomen approximately equal to 10:6:11:26. Metathoracic wings present.</paragraph>
<paragraph id="8BE26E2EFFEF664CFF006AD6B84A5780" blockId="15.[151,1437,151,2035]" pageId="15" pageNumber="216">Legs rather short and robust, finely punctate and pubescent. Procoxa nearly rounded, narrowly separated by 0.6× diameter of procoxa; mesocoxae hemispherical, narrowly separated by about 0.3× diameter of mesocoxa; metacoxae oval, transverse, separated by 0.5× longitudinal diameter of metacoxa. Femora elongate oval, dilated medially, flattened; femora and tibiae subequal in length. Tibiae straight, dilated apically, with fringe of stout setae and two spines apically. Tarsi slender, about 0.6× as long as tibia; tarsal formula 4-4-4; tarsomeres with sparse, fine setation ventrally; tarsomere 4 longest, longer than metatarsomeres 13 combined, widened distally, slightly curved. Claws simple, large, about 0.3× as long as tarsomere 4.</paragraph>
<paragraph id="8BE26E2EFFEF664CFF006BD0BE6D57F4" blockId="15.[151,1437,151,2035]" pageId="15" pageNumber="216">Abdomen with five visible ventrites, abdominal sutures distinct throughout length; densely covered with small punctation. Relative length ratios of ventrites 15 equal to 10:5:4:3:3. Abdominal ventrite 1 with small, triangular intercoxal process. Abdominal ventrite 5 with widely rounded apical margin, and with preapical groove.</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFEF664CFF006BBDBE505598" pageId="15" pageNumber="216" type="diagnosis">
<paragraph id="8BE26E2EFFEF664CFF006BBDBFC35532" blockId="15.[151,1437,151,2035]" pageId="15" pageNumber="216">
<emphasis id="B929B23CFFEF664CFF006BBDBA805411" bold="true" box="[199,459,508,534]" pageId="15" pageNumber="216">Differential diagnosis.</emphasis>
<taxonomicName id="4C5D15ADFFEF664CFE176BBCB84E5411" authorityName="Alekseev &amp; Bukejs" authorityYear="2024" box="[464,773,508,534]" class="Insecta" family="Zopheridae" genus="Lasconotus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="tenebrisilvarum" status="sp. nov.">
<emphasis id="B929B23CFFEF664CFE176BBCB84E5411" box="[464,773,508,534]" italics="true" pageId="15" pageNumber="216">Lasconotus tenebrisilvarum</emphasis>
</taxonomicName>
<emphasis id="B929B23CFFEF664CFCCD6BBDB8295411" bold="true" box="[778,866,508,534]" pageId="15" pageNumber="216">
<taxonomicNameLabel id="A21A0F47FFEF664CFCCD6BBDB8295411" box="[778,866,508,534]" pageId="15" pageNumber="216" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from all extant congeners in the combination of the following characters: elytra convex (in contrast to concave elytra of several North American representatives); elytra with elytral carinae similarly raised along entire lengths; pronotum with two pairs of distinct and uninterrupted sublateral carinae; anterior margin of pronotum without distinctive paired tufts of long, golden setae; raised lateral margin of pronotum forming distinct longitudinal depression between lateral margin and first pair of pronotal carina; pronotal margins almost parallel; posterior pronotal angles almost rectangular and not projecting posteriorly; and elytra at base only slightly wider than pronotum. Additionally,
<taxonomicName id="4C5D15ADFFEF664CFC946892BF6954EB" authorityName="Alekseev &amp; Bukejs" authorityYear="2024" box="[851,1058,722,748]" class="Insecta" family="Zopheridae" genus="Lasconotus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="tenebrisilvarum" status="sp. nov.">
<emphasis id="B929B23CFFEF664CFC946892BF6954EB" box="[851,1058,722,748]" italics="true" pageId="15" pageNumber="216">L. tenebrisilvarum</emphasis>
</taxonomicName>
<emphasis id="B929B23CFFEF664CFBEE6893BFC954EB" bold="true" box="[1065,1154,722,748]" pageId="15" pageNumber="216">
<taxonomicNameLabel id="A21A0F47FFEF664CFBEE6893BFC954EB" box="[1065,1154,722,748]" pageId="15" pageNumber="216" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from the majority of the genus representatives in possessing narrowly open procoxal cavities, a character shared with two North American species,
<taxonomicName id="4C5D15ADFFEF664CFEAC695AB8545533" authority="Kingsolver et al., 2006" authorityName="Kingsolver" authorityYear="2006" box="[363,799,794,820]" class="Insecta" family="Zopheridae" genus="Lasconotus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="fitzgibbonae">
<emphasis id="B929B23CFFEF664CFEAC695AB9585533" box="[363,531,794,820]" italics="true" pageId="15" pageNumber="216">L. fitzgibbonae</emphasis>
<bibRefCitation id="EFCC13DFFFEF664CFDDD695BB8545533" author="Kingsolver, J. M. &amp; Stephan, K. &amp; Moser, J. C." box="[538,799,794,820]" pageId="15" pageNumber="216" pagination="53 - 56" refId="ref26243" refString="Kingsolver, J. M., Stephan, K. &amp; Moser, J. C. (2006) A new species of Lasconotus (Coleoptera: Colydiidae) from Arizona and South Dakota, U. S. A. Entomological News, 117 (1), 53 - 56. https: // doi. org / 10.3157 / 0013 - 872 X (2006) 117 [53: ANSOLC] 2.0. CO; 2" type="journal article" year="2006">
Kingsolver
<emphasis id="B929B23CFFEF664CFD59695AB9845533" box="[670,719,794,820]" italics="true" pageId="15" pageNumber="216">et al</emphasis>
., 2006
</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C5D15ADFFEF664CFC9E695ABFCF5532" authority="Hinton, 1935" authorityName="Hinton" authorityYear="1935" box="[857,1156,794,821]" class="Insecta" family="Zopheridae" genus="Lasconotus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="coronatus">
<emphasis id="B929B23CFFEF664CFC9E695AB8AD5533" box="[857,998,795,820]" italics="true" pageId="15" pageNumber="216">L. coronatus</emphasis>
Hinton, 1935
</taxonomicName>
.
</paragraph>
<paragraph id="8BE26E2EFFEF664CFF00697CBE505598" blockId="15.[151,1437,151,2035]" pageId="15" pageNumber="216">
The new species can be easily distinguished from other extinct
<taxonomicName id="4C5D15ADFFEF664CFC57697CBF405550" authorityName="Erichson" authorityYear="1842" box="[912,1035,829,855]" class="Insecta" family="Colydiinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="family">Colydiinae</taxonomicName>
found in Baltic amber based on the combination of generic characters, most obviously in the presence of 11-segmented antennae with a 3-segmented club, pronotum with two pairs of sublateral carinae, presence of supraocular carina, and carinate elytra.
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFEF664CFF0069E9BE015228" pageId="15" pageNumber="216" type="etymology">
<paragraph id="8BE26E2EFFEF664CFF0069E9BE015228" blockId="15.[151,1437,151,2035]" pageId="15" pageNumber="216">
<emphasis id="B929B23CFFEF664CFF0069E9BAD655C5" bold="true" box="[199,413,936,962]" pageId="15" pageNumber="216">Derivatio nominis</emphasis>
. The name of the new species
<taxonomicName id="4C5D15ADFFEF664CFD3569E8B8E855C4" authorityName="Alekseev &amp; Bukejs" authorityYear="2024" box="[754,931,937,963]" class="Insecta" family="Zopheridae" genus="Lasconotus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="tenebrisilvarum">
<emphasis id="B929B23CFFEF664CFD3569E8B8E855C4" box="[754,931,937,963]" italics="true" pageId="15" pageNumber="216">tenebrisilvarum</emphasis>
</taxonomicName>
is derived from the Latin “
<emphasis id="B929B23CFFEF664CFB1569E8BEC455C4" box="[1234,1423,937,963]" italics="true" pageId="15" pageNumber="216">tenebricae silvae</emphasis>
” in genitive case, meaning “of dark forests”. This refers to the region of the specimens origin in two ways: (1) referring to Eocene forest paleohabitats, and (2) to the former Eastern Prussia, a historical area with the informal anthem “Land der dunklen Wälder“ (land of dark forests), written by Erich Hannighofer and Herbert Brust.
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFEF664CFF006E76B946506D" pageId="15" pageNumber="216" type="distribution">
<paragraph id="8BE26E2EFFEF664CFF006E76B8F05303" blockId="15.[151,1437,151,2035]" pageId="15" pageNumber="216">
<emphasis id="B929B23CFFEF664CFF006E76BAB65255" bold="true" box="[199,509,1079,1106]" pageId="15" pageNumber="216">Distribution of congeners.</emphasis>
The genus
<taxonomicName id="4C5D15ADFFEF664CFDB96E79B9B55256" authorityName="Erichson" authorityYear="1845" box="[638,766,1080,1105]" class="Insecta" family="Zopheridae" genus="Lasconotus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFEF664CFDB96E79B9B55256" box="[638,766,1080,1105]" italics="true" pageId="15" pageNumber="216">Lasconotus</emphasis>
</taxonomicName>
is cosmopolitan, being represented in the New World by 37 species (
<bibRefCitation id="EFCC13DFFFEF664CFF306E1ABAE35272" author="Ivie, M. A. &amp; Lord, N. P. &amp; Foley, I. A." box="[247,424,1115,1142]" pageId="15" pageNumber="216" pagination="755 - 788" refId="ref25945" refString="Ivie, M. A., Lord, N. P. &amp; Foley, I. A. (2016) Colydiine genera (Coleoptera: Zopheridae: Colydiinae) of the new world: a key and nomenclatural acts 30 years in the making. The Coleopterists Bulletin, 70 (4), 755 - 788. https: // doi. org / 10.1649 / 0010 - 065 X- 70.4.755" type="journal article" year="2016">
Ivie
<emphasis id="B929B23CFFEF664CFEEB6E1DBA155272" box="[300,350,1115,1141]" italics="true" pageId="15" pageNumber="216">et al</emphasis>
. 2016
</bibRefCitation>
), and in the Palearctic by 8 species (
<bibRefCitation id="EFCC13DFFFEF664CFC856E1CBF1E5271" author="Slipinski, S. A. &amp; Schuh, R." box="[834,1109,1115,1144]" pageId="15" pageNumber="216" refId="ref28217" refString="Slipinski, S. A. &amp; Schuh, R. (2008) Zopheridae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 5. Apollo Books, Stenstrup, pp. 78 - 87." type="book" year="2008">Ślipiński &amp; Schuh 2008</bibRefCitation>
;
<bibRefCitation id="EFCC13DFFFEF664CFB986E1ABF925271" author="Aoki, J." box="[1119,1241,1115,1142]" pageId="15" pageNumber="216" pagination="97 - 102" refId="ref24561" refString="Aoki, J. (2011) Revised status of a colydiid species known as &quot; Lasconotus niponius (Lewis) &quot; (Coleoptera, Zopheridae). Elytra, New Series, 1 (1), 97 - 102." type="journal article" year="2011">Aoki 2011</bibRefCitation>
,
<bibRefCitation id="EFCC13DFFFEF664CFB236E1DBE6A5271" author="Aoki, J." box="[1252,1313,1116,1142]" pageId="15" pageNumber="216" pagination="69 - 74" refId="ref24603" refString="Aoki, J. (2018) Japanese species of the genus Lasconotus (Coleoptera: Zopheridae: Colydiinae), with description of a new species from Ishigaki-jima Island, the Ryukyu Islands. Species Diversity, 23, 69 - 74. https: // doi. org / 10.12782 / specdiv. 23.69" type="journal article" year="2018">2018</bibRefCitation>
). Only one species,
<taxonomicName id="4C5D15ADFFEF664CFF326EC1B971529E" authority="(Wankowicz, 1867)" baseAuthorityName="Wankowicz" baseAuthorityYear="1867" box="[245,570,1151,1177]" class="Insecta" family="Zopheridae" genus="Lasconotus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="jelskii">
<emphasis id="B929B23CFFEF664CFF326EC1BA1E529E" box="[245,341,1151,1177]" italics="true" pageId="15" pageNumber="216">L. jelskii</emphasis>
(Wankowicz, 1867)
</taxonomicName>
, is known as a primeval forest relict in Eastern and Northern Europe, whereas the most diverse Palearctic fauna has been reported from
<collectingCountry id="F34A2EBEFFEF664CFCD36EE2B81F52BA" box="[788,852,1187,1213]" name="Japan" pageId="15" pageNumber="216">Japan</collectingCountry>
(4 species:
<taxonomicName id="4C5D15ADFFEF664CFC106EE5BFF252BA" authority="Aoki, 2018" authorityName="Aoki" authorityYear="2018" box="[983,1209,1187,1213]" class="Insecta" family="Zopheridae" genus="Lasconotus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="akitai">
<emphasis id="B929B23CFFEF664CFC106EE5BF7F52BA" box="[983,1076,1187,1213]" italics="true" pageId="15" pageNumber="216">L. akitai</emphasis>
<bibRefCitation id="EFCC13DFFFEF664CFBFF6EE2BFF252BA" author="Aoki, J." box="[1080,1209,1187,1213]" pageId="15" pageNumber="216" pagination="69 - 74" refId="ref24603" refString="Aoki, J. (2018) Japanese species of the genus Lasconotus (Coleoptera: Zopheridae: Colydiinae), with description of a new species from Ishigaki-jima Island, the Ryukyu Islands. Species Diversity, 23, 69 - 74. https: // doi. org / 10.12782 / specdiv. 23.69" type="journal article" year="2018">Aoki, 2018</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C5D15ADFFEF664CFB036EE5BB9052E6" authority="(Lewis, 1879)" baseAuthorityName="Lewis" baseAuthorityYear="1879" class="Insecta" family="Zopheridae" genus="Lasconotus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="niponius">
<emphasis id="B929B23CFFEF664CFB036EE5BE0A52BA" box="[1220,1345,1188,1213]" italics="true" pageId="15" pageNumber="216">L. niponius</emphasis>
(Lewis, 1879)
</taxonomicName>
;
<taxonomicName id="4C5D15ADFFEF664CFF2F6E86BA9152E6" authority="Aoki, 2011" authorityName="Aoki" authorityYear="2011" box="[232,474,1223,1249]" class="Insecta" family="Zopheridae" genus="Lasconotus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="okadai">
<emphasis id="B929B23CFFEF664CFF2F6E86BA1852E6" box="[232,339,1223,1249]" italics="true" pageId="15" pageNumber="216">L. okadai</emphasis>
<bibRefCitation id="EFCC13DFFFEF664CFE9E6E86BA9152E6" author="Aoki, J." box="[345,474,1223,1249]" pageId="15" pageNumber="216" pagination="97 - 102" refId="ref24561" refString="Aoki, J. (2011) Revised status of a colydiid species known as &quot; Lasconotus niponius (Lewis) &quot; (Coleoptera, Zopheridae). Elytra, New Series, 1 (1), 97 - 102." type="journal article" year="2011">Aoki, 2011</bibRefCitation>
</taxonomicName>
; and
<taxonomicName id="4C5D15ADFFEF664CFDD06E86B83B52E6" authority="(Sharp, 1885))" authorityName="" baseAuthorityName="Sharp" baseAuthorityYear="1885" box="[535,880,1222,1249]" class="Insecta" family="Zopheridae" genus="Lasconotus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="sculpturatus">
<emphasis id="B929B23CFFEF664CFDD06E86B98B52E6" box="[535,704,1223,1249]" italics="true" pageId="15" pageNumber="216">L. sculpturatus</emphasis>
(Sharp, 1885))
</taxonomicName>
. The fossil species described in the current paper confirms the presence of the genus in the Paleogene Western Palearctic.
</paragraph>
<paragraph id="8BE26E2EFFEF664CFF006F4FB946506D" blockId="15.[151,1437,151,2035]" pageId="15" pageNumber="216">
The single Recent European species is boreal and the locality in Puszcza Białowieska (
<collectingCountry id="F34A2EBEFFEF664CFB456F4FBF98532F" box="[1154,1235,1294,1320]" name="Poland" pageId="15" pageNumber="216">Poland</collectingCountry>
/
<collectingCountry id="F34A2EBEFFEF664CFB1B6F4FBE78532F" box="[1244,1331,1294,1320]" name="Belarus" pageId="15" pageNumber="216">Belarus</collectingCountry>
), situated at about
<geoCoordinate id="EE6908E9FFEF664CFF3A6F73BA2B534A" box="[253,352,1330,1357]" degrees="52" direction="north" minutes="30" orientation="latitude" pageId="15" pageNumber="216" precision="925" value="52.5">52°30´N</geoCoordinate>
, is one of the southernmost known points of the groups present-day distribution (
<bibRefCitation id="EFCC13DFFFEF664CFADD6F73BA0A5377" author="Nikitsky, N. B. &amp; Slipinski, S. A." pageId="15" pageNumber="216" pagination="43 - 45" refId="ref27087" refString="Nikitsky, N. B. &amp; Slipinski, S. A. (2003) New species of thi genus Lasconotus Erichson, 1845 (Coleoptera: Zopheridae) from Middle Asia and Kazakhstan. Byulleten' Moskovskogo Obshchestva Ispytatelei Prirody. Otdel Biologicheskii, 108 (6), 43 - 45. [in Russian]" type="journal article" year="2003">Nikitsky &amp; Slipinski 2003</bibRefCitation>
). Like the situation with fossil erotylid species from Rovno amber,
<taxonomicName id="4C5D15ADFFEF664CFB896F14BA015394" authority="Alekseev et Bukejs, 2023" authorityName="Alekseev et Bukejs" authorityYear="2023" class="Insecta" family="Erotylidae" genus="Zavaljus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="lyubarskyi">
<emphasis id="B929B23CFFEF664CFB896F14BE665368" box="[1102,1325,1365,1391]" italics="true" pageId="15" pageNumber="216">Zavaljus lyubarskyi</emphasis>
<bibRefCitation id="EFCC13DFFFEF664CFAF36F14BA015394" author="Alekseev, V. I. &amp; Bukejs, A." pageId="15" pageNumber="216" pagination="315 - 319" refId="ref24145" refString="Alekseev, V. I. &amp; Bukejs, A. (2023) A new extinct species of Zavaljus Reitter (Coleoptera: Erotylidae) from Rovno amber: boreal distribution range since the Eocene. Historical Biology, 35 (3), 315 - 319. https: // doi. org / 10.1080 / 08912963.2022.2036141" type="journal article" year="2023">Alekseev et Bukejs, 2023</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFCC13DFFFEF664CFE9C6F38B9345394" author="Alekseev, V. I. &amp; Bukejs, A." box="[347,639,1401,1427]" pageId="15" pageNumber="216" pagination="315 - 319" refId="ref24145" refString="Alekseev, V. I. &amp; Bukejs, A. (2023) A new extinct species of Zavaljus Reitter (Coleoptera: Erotylidae) from Rovno amber: boreal distribution range since the Eocene. Historical Biology, 35 (3), 315 - 319. https: // doi. org / 10.1080 / 08912963.2022.2036141" type="journal article" year="2023">Alekseev &amp; Bukejs 2023</bibRefCitation>
), the southern limit of the present-day distribution range of the genus
<taxonomicName id="4C5D15ADFFEF664CFF506FDFBA5C53B0" authorityName="Erichson" authorityYear="1845" box="[151,279,1438,1463]" class="Insecta" family="Zopheridae" genus="Lasconotus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFEF664CFF506FDFBA5C53B0" box="[151,279,1438,1463]" italics="true" pageId="15" pageNumber="216">Lasconotus</emphasis>
</taxonomicName>
in Europe almost coincides with the late Eocene record at approximately
<geoCoordinate id="EE6908E9FFEF664CFB956FDCBFDC53BF" box="[1106,1175,1437,1464]" degrees="51" direction="north" orientation="latitude" pageId="15" pageNumber="216" precision="55555" value="51.0">51° N</geoCoordinate>
. On the other hand, the modern boreal distribution of
<taxonomicName id="4C5D15ADFFEF664CFE206F83B92C53DC" authorityName="Erichson" authorityYear="1845" box="[487,615,1474,1499]" class="Insecta" family="Zopheridae" genus="Lasconotus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFEF664CFE206F83B92C53DC" box="[487,615,1474,1499]" italics="true" pageId="15" pageNumber="216">Lasconotus</emphasis>
</taxonomicName>
is evident only for Europe. Representatives of the genus inhabit different latitudes in North America from
<collectingCountry id="F34A2EBEFFEF664CFDCF6FA5B91753F9" box="[520,604,1508,1534]" name="Canada" pageId="15" pageNumber="216">Canada</collectingCountry>
to
<collectingCountry id="F34A2EBEFFEF664CFD466FA5B99C53F9" box="[641,727,1508,1534]" name="Mexico" pageId="15" pageNumber="216">Mexico</collectingCountry>
inclusive, and they are also known in different localities in the southern Palearctic (e.g., Central Asia,
<collectingCountry id="F34A2EBEFFEF664CFD806C49B9C65025" box="[583,653,1544,1570]" name="Nepal" pageId="15" pageNumber="216">Nepal</collectingCountry>
,
<collectingCountry id="F34A2EBEFFEF664CFD506C49B8615025" box="[663,810,1544,1570]" name="Saudi Arabia" pageId="15" pageNumber="216">Saudi Arabia</collectingCountry>
,
<collectingCountry id="F34A2EBEFFEF664CFCF36C49B8325025" box="[820,889,1544,1570]" name="Japan" pageId="15" pageNumber="216">Japan</collectingCountry>
). The present-day boreal distribution of the genus in Europe is probably determined not by climatic reasons and preferences of the genus, but by historical factors related to Quaternary glaciations.
</paragraph>
</subSubSection>
<subSubSection id="C3473DA5FFEF664CFF006C35B86E51F4" pageId="15" pageNumber="216" type="biology_ecology">
<paragraph id="8BE26E2EFFEF664CFF006C35B86E51F4" blockId="15.[151,1437,151,2035]" pageId="15" pageNumber="216">
<emphasis id="B929B23CFFEF664CFF006C35BA835089" bold="true" box="[199,456,1651,1678]" pageId="15" pageNumber="216">Ecology of congeners.</emphasis>
The ecology of representatives of the genus
<taxonomicName id="4C5D15ADFFEF664CFC736C35BF7F508A" authorityName="Erichson" authorityYear="1845" box="[948,1076,1652,1677]" class="Insecta" family="Zopheridae" genus="Lasconotus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFEF664CFC736C35BF7F508A" box="[948,1076,1652,1677]" italics="true" pageId="15" pageNumber="216">Lasconotus</emphasis>
</taxonomicName>
is insufficiently known. Reports from North America, Europe and Asia show association of different species with pinacean conifers (
<taxonomicName id="4C5D15ADFFEF664CFAED6CD9BE2150B6" box="[1322,1386,1688,1713]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="15" pageNumber="216" phylum="Tracheophyta" rank="genus">
<emphasis id="B929B23CFFEF664CFAED6CD9BE2150B6" box="[1322,1386,1688,1713]" italics="true" pageId="15" pageNumber="216">Pinus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5D15ADFFEF664CFF506CFDBB9C50D2" box="[151,215,1724,1749]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="15" pageNumber="216" phylum="Tracheophyta" rank="genus">
<emphasis id="B929B23CFFEF664CFF506CFDBB9C50D2" box="[151,215,1724,1749]" italics="true" pageId="15" pageNumber="216">Picea</emphasis>
</taxonomicName>
), where the beetles are noted under bark in the galleries of bark beetles (
<taxonomicName id="4C5D15ADFFEF664CFC3F6CFABFD650D2" box="[1016,1181,1723,1749]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="family">Curculionidae</taxonomicName>
:
<taxonomicName id="4C5D15ADFFEF664CFB606CFABE5450D2" box="[1191,1311,1723,1749]" class="Insecta" family="Scolytinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="family">Scolytinae</taxonomicName>
) belonging to the representatives of genera
<taxonomicName id="4C5D15ADFFEF664CFDCD6C9EB96750FF" authorityName="De Geer" authorityYear="1775" box="[522,556,1759,1784]" class="Insecta" family="Curculionidae" genus="Ips" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFEF664CFDCD6C9EB96750FF" box="[522,556,1759,1784]" italics="true" pageId="15" pageNumber="216">Ips</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5D15ADFFEF664CFDAE6C9EB9B350FE" box="[617,760,1759,1785]" class="Insecta" family="Curculionidae" genus="Polygraphus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFEF664CFDAE6C9EB9B350FE" box="[617,760,1759,1785]" italics="true" pageId="15" pageNumber="216">Polygraphus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFCC13DFFFEF664CFCCD6C9EB8D350FE" author="Saalas, U." box="[778,920,1759,1785]" pageId="15" pageNumber="216" pagination="1 - 726" refId="ref27681" refString="Saalas, U. (1923) Die Fichtenkafer Finnlands. II. Studien uber die Entwicklungstadien, Lebensweise und geographische Verbreitung der an Picea excelsa Link. lebenden Coleopteren nebst einer Larvenbestimmungstabelle. Annales Academiae Scientarium Fennicae, Serie A, 22 (1), 1 - 726." type="journal article" year="1923">Saalas 1923</bibRefCitation>
;
<bibRefCitation id="EFCC13DFFFEF664CFC616C9EBF1D50FE" author="Hackwell, G. A." box="[934,1110,1759,1785]" pageId="15" pageNumber="216" pagination="62 - 65" refId="ref25802" refString="Hackwell, G. A. (1973) Biology of Lasconotus subcostulatus (Coleoptera: Colydiidae) with special reference to feeding behavior. Annals of the Entomological Society of America, 66 (1), 62 - 65. https: // doi. org / 10.1093 / aesa / 66.1.62" type="journal article" year="1973">Hackwell 1973</bibRefCitation>
;
<bibRefCitation id="EFCC13DFFFEF664CFBA26C9EBEDC50FE" author="Nikitsky, N. B. &amp; Slipinski, S. A." box="[1125,1431,1759,1785]" pageId="15" pageNumber="216" pagination="43 - 45" refId="ref27087" refString="Nikitsky, N. B. &amp; Slipinski, S. A. (2003) New species of thi genus Lasconotus Erichson, 1845 (Coleoptera: Zopheridae) from Middle Asia and Kazakhstan. Byulleten' Moskovskogo Obshchestva Ispytatelei Prirody. Otdel Biologicheskii, 108 (6), 43 - 45. [in Russian]" type="journal article" year="2003">Nikitsky &amp; Slipinski 2003</bibRefCitation>
;
<bibRefCitation id="EFCC13DFFFEF664CFF506D43BAD2511B" author="Kingsolver, J. M. &amp; Stephan, K. &amp; Moser, J. C." box="[151,409,1794,1821]" pageId="15" pageNumber="216" pagination="53 - 56" refId="ref26243" refString="Kingsolver, J. M., Stephan, K. &amp; Moser, J. C. (2006) A new species of Lasconotus (Coleoptera: Colydiidae) from Arizona and South Dakota, U. S. A. Entomological News, 117 (1), 53 - 56. https: // doi. org / 10.3157 / 0013 - 872 X (2006) 117 [53: ANSOLC] 2.0. CO; 2" type="journal article" year="2006">
Kingsolver
<emphasis id="B929B23CFFEF664CFEDC6D42BA05511B" box="[283,334,1794,1820]" italics="true" pageId="15" pageNumber="216">et al</emphasis>
. 2006
</bibRefCitation>
).
<taxonomicName id="4C5D15ADFFEF664CFE6B6D42B967511B" authorityName="Erichson" authorityYear="1845" box="[428,556,1795,1820]" class="Insecta" family="Zopheridae" genus="Lasconotus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFEF664CFE6B6D42B967511B" box="[428,556,1795,1820]" italics="true" pageId="15" pageNumber="216">Lasconotus</emphasis>
</taxonomicName>
larvae of instars 12 feed upon fungi, and later (instar 3) they are predaceous on bark beetle larvae (
<bibRefCitation id="EFCC13DFFFEF664CFE536D67B90E5147" author="Hackwell, G. A." box="[404,581,1830,1856]" pageId="15" pageNumber="216" pagination="62 - 65" refId="ref25802" refString="Hackwell, G. A. (1973) Biology of Lasconotus subcostulatus (Coleoptera: Colydiidae) with special reference to feeding behavior. Annals of the Entomological Society of America, 66 (1), 62 - 65. https: // doi. org / 10.1093 / aesa / 66.1.62" type="journal article" year="1973">Hackwell 1973</bibRefCitation>
). The Eocene
<taxonomicName id="4C5D15ADFFEF664CFD206D66BF6B5147" authorityName="Alekseev &amp; Bukejs" authorityYear="2024" box="[743,1056,1830,1856]" class="Insecta" family="Zopheridae" genus="Lasconotus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="tenebrisilvarum" status="sp. nov.">
<emphasis id="B929B23CFFEF664CFD206D66BF6B5147" box="[743,1056,1830,1856]" italics="true" pageId="15" pageNumber="216">Lasconotus tenebrisilvarum</emphasis>
</taxonomicName>
<emphasis id="B929B23CFFEF664CFBEF6D67BFC85147" bold="true" box="[1064,1155,1830,1856]" pageId="15" pageNumber="216">
<taxonomicNameLabel id="A21A0F47FFEF664CFBEF6D67BFC85147" box="[1064,1155,1830,1856]" pageId="15" pageNumber="216" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
could also be associated with an analogous ancient forest community of dead coniferous trees infested by scolytines. While bark beetles of the genus
<taxonomicName id="4C5D15ADFFEF664CFECA6D2FBA94518F" authority="De Geer, 1775" authorityName="De Geer" authorityYear="1775" box="[269,479,1901,1928]" class="Insecta" family="Curculionidae" genus="Ips" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFEF664CFECA6D2FBA645180" box="[269,303,1902,1927]" italics="true" pageId="15" pageNumber="216">Ips</emphasis>
De Geer, 1775
</taxonomicName>
and tribe
<taxonomicName id="4C5D15ADFFEF664CFD916D2CB855518F" authority="Bedel, 1888" authorityName="Bedel" authorityYear="1888" box="[598,798,1901,1928]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="tribe" tribe="Ipini">Ipini Bedel, 1888</taxonomicName>
are unknown from inclusions in Baltic amber, the tribe
<taxonomicName id="4C5D15ADFFEF664CFF506DD0BA9B51AC" authority="Chapuis, 1869" authorityName="Chapuis" authorityYear="1869" box="[151,464,1937,1963]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="tribe" tribe="Polygraphini">Polygraphini Chapuis, 1869</taxonomicName>
(
<taxonomicName id="4C5D15ADFFEF664CFE196DD0B9C851AC" box="[478,643,1937,1963]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="family">Curculionidae</taxonomicName>
:
<taxonomicName id="4C5D15ADFFEF664CFD4A6DD0B84E51AC" box="[653,773,1937,1963]" class="Insecta" family="Scolytinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="family">Scolytinae</taxonomicName>
) is represented in Baltic amber by two described species of the genus
<taxonomicName id="4C5D15ADFFEF664CFEC06DF4B90351C8" authority="Eichhoff, 1864" authorityName="Eichhoff" authorityYear="1864" box="[263,584,1973,1999]" class="Insecta" family="Curculionidae" genus="Carphoborus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFEF664CFEC06DF4BAD751C8" box="[263,412,1973,1999]" italics="true" pageId="15" pageNumber="216">Carphoborus</emphasis>
Eichhoff, 1864
</taxonomicName>
(
<bibRefCitation id="EFCC13DFFFEF664CFD916DF4B9BF51C8" author="Legalov, A. A." box="[598,756,1973,1999]" pageId="15" pageNumber="216" refId="ref26516" refString="Legalov, A. A. (2020) A review of the Curculionoidea (Coleoptera) from European Eocene ambers. Geosciences, 10 (1), 16. https: // doi. org / 10.3390 / geosciences 10010016" type="journal volume" year="2020">Legalov 2020</bibRefCitation>
,
<bibRefCitation id="EFCC13DFFFEF664CFD396DF4B87051C8" author="Legalov, A. A." box="[766,827,1973,1999]" pageId="15" pageNumber="216" pagination="378 - 388" refId="ref26558" refString="Legalov, A. A. (2024) Fossil history of bark-beetles (Coleoptera: Scolytidae) with descriptions of two new species. Historical Biology, 36 (2), 378 - 388. https: // doi. org / 10.1080 / 08912963.2022.2157275" type="journal article" year="2024">2024</bibRefCitation>
). The trophic association with galleries of these beetles could be assumed for
<taxonomicName id="4C5D15ADFFEF664CFE4A6D9BB98F51F4" authorityName="Alekseev &amp; Bukejs" authorityYear="2024" box="[397,708,2009,2035]" class="Insecta" family="Zopheridae" genus="Lasconotus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="tenebrisilvarum" status="sp. nov.">
<emphasis id="B929B23CFFEF664CFE4A6D9BB98F51F4" box="[397,708,2009,2035]" italics="true" pageId="15" pageNumber="216">Lasconotus tenebrisilvarum</emphasis>
</taxonomicName>
<emphasis id="B929B23CFFEF664CFD0B6D98B86E51F4" bold="true" box="[716,805,2009,2035]" pageId="15" pageNumber="216">
<taxonomicNameLabel id="A21A0F47FFEF664CFD0B6D98B86E51F4" box="[716,805,2009,2035]" pageId="15" pageNumber="216" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>