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149 lines
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<document id="13C9EC794E1870FEC63364B2B71383A5" ID-CLB-Dataset="30626" ID-DOI="10.11646/zootaxa.4387.3.4" ID-GBIF-Dataset="43de8956-0706-4f8d-b5a9-f51c30c276d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="1187697" ID-ZooBank="B53A6B20-5386-4E5A-A00F-A559EDF640DF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1520059483769" checkinUser="plazi" docAuthor="Fernández, Mercedes, Fernicola, Juan Carlos, Cerdeño, Esperanza & Reguero, Marcelo A." docDate="2018" docId="03EC879BCF5F8316C192296E8F42FBE6" docLanguage="en" docName="zootaxa.4387.3.4.pdf" docOrigin="Zootaxa 4387 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Protypotherium globosum Ameghino 1891" docType="treatment" docVersion="7" lastPageNumber="485" masterDocId="FFD5FFE3CF53831AC10528498E40FFD5" masterDocTitle="Identification of type materials of the species of Protypotherium Ameghino, 1885 and Patriarchus Ameghino, 1889 (Notoungulata: Interatheriidae) erected by Florentino Ameghino" masterLastPageNumber="498" masterPageNumber="473" pageNumber="485" updateTime="1698526258101" updateUser="ExternalLinkService">
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<mods:title id="26EF98408E5E47389892E36DF1BA7314">Identification of type materials of the species of Protypotherium Ameghino, 1885 and Patriarchus Ameghino, 1889 (Notoungulata: Interatheriidae) erected by Florentino Ameghino</mods:title>
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<mods:namePart id="4C0E47B860B48ADAE9B115C9E2CCBC79">Fernández, Mercedes</mods:namePart>
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<mods:namePart id="A8C06068ED1B73CB4BEF419DF1FB8897">Fernicola, Juan Carlos</mods:namePart>
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<mods:name id="8ECBF6C15C90A12DE01D759739B5E26A" type="personal">
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<mods:namePart id="A115B4EB83DFF3DB6EA05686ECC11006">Cerdeño, Esperanza</mods:namePart>
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<mods:namePart id="6FE54040E9DAECE7DEC1ED189F35C448">Reguero, Marcelo A.</mods:namePart>
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<mods:title id="677C6A29822F1113683F2F559F69C11D">Zootaxa</mods:title>
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<mods:date id="5F770939FFFBB9C9302A6D1321DA5C60">2018</mods:date>
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<mods:number id="CF422492DD03C3F9E4462A38CDC24ED1">2018-02-28</mods:number>
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<mods:number id="09556914DE6F4E8C36A4F914C5AA923E">4387</mods:number>
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<mods:number id="DE6761EC2B387CD7F48808330A53F592">3</mods:number>
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<mods:identifier id="9E499B513561E965DDFE34B10A3E6F34" type="DOI">10.11646/zootaxa.4387.3.4</mods:identifier>
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<treatment id="03EC879BCF5F8316C192296E8F42FBE6" ID-DOI="http://doi.org/10.5281/zenodo.5986689" ID-GBIF-Taxon="141264029" ID-Zenodo-Dep="5986689" LSID="urn:lsid:plazi:treatment:03EC879BCF5F8316C192296E8F42FBE6" httpUri="http://treatment.plazi.org/id/03EC879BCF5F8316C192296E8F42FBE6" lastPageNumber="485" pageId="12" pageNumber="485">
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<subSubSection id="C35F6506CF5F8316C192296E8CE3FE97" box="[151,675,295,322]" pageId="12" pageNumber="485" type="nomenclature">
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<paragraph id="8BFA368DCF5F8316C192296E8CE3FE97" blockId="12.[151,675,295,322]" box="[151,675,295,322]" pageId="12" pageNumber="485">
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<heading id="D0B281E1CF5F8316C192296E8CE3FE97" bold="true" box="[151,675,295,322]" fontSize="11" level="1" pageId="12" pageNumber="485" reason="1">
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<taxonomicName id="4C454D0ECF5F8316C192296E8CE3FE97" authority="Ameghino, 1891" authorityName="Ameghino" authorityYear="1891" box="[151,675,295,322]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="485" phylum="Chordata" rank="species" species="globosum">
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<emphasis id="B931EA9FCF5F8316C192296E8CE3FE97" bold="true" box="[151,675,295,322]" pageId="12" pageNumber="485">
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<emphasis id="B931EA9FCF5F8316C192296E8F92FE94" bold="true" box="[151,466,295,321]" italics="true" pageId="12" pageNumber="485">Protypotherium globosum</emphasis>
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<bibRefCitation id="EFD44B7CCF5F8316C0DC29618CE3FE97" box="[473,675,296,322]" pageId="12" pageNumber="485" refId="ref17076" refString="Ameghino, F. (1891) Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural, 1 (5), 289 - 328." type="journal article">Ameghino, 1891</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35F6506CF5F8316C19229278A85FE52" box="[151,1221,366,391]" pageId="12" pageNumber="485" type="materials_examined">
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<paragraph id="8BFA368DCF5F8316C19229278A85FE52" blockId="12.[151,1437,366,1075]" box="[151,1221,366,391]" pageId="12" pageNumber="485">
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<emphasis id="B931EA9FCF5F8316C19229278F4BFE52" bold="true" box="[151,267,366,391]" pageId="12" pageNumber="485">Holotype.</emphasis>
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MACN-A 4049 (
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<figureCitation id="137E2A08CF5F8316C0D729278C25FE52" box="[466,613,366,391]" captionStart="FIGURE 2" captionStartId="5.[151,249,1632,1654]" captionTargetBox="[308,1273,213,1595]" captionTargetId="figure@5.[305,1281,193,1611]" captionTargetPageId="5" captionText="FIGURE 2. Protypotherium attenuatum, syntype MACN-A 524, left maxilla with P2–M3 but lacking P1, in occlusal view (A); syntype MACN-A 628, right fragmented mandible with alveoli of i1–3 and c (broken)–m1 but lacking m2–3, in labial (B) and occlusal (C) views; reproductions of figures 21 (D), 20 (E) and 20a (F) from Ameghino (1889, plate 14). Protypotherium obstructum, holotype MACN-A 1677, lower premolar in labial, oclusal and lingual views (G); reproduction of figure 19 (H) from Ameghino (1889, plate 15). Protypotherium claudum, holotype MACN-A 551, right mandibular fragment with alveolus of p2 andp3–m2, in labial (I) and occlusal (J)views; reproductions of figures 22 (K) and 22a (L) from Ameghino (1889, plate 14). Protypotherium globosum, holotype MACN-A 4049, skull, in ventral (M) and dorsal (N) views. Protypotherium convexidens, holotype MACN-A 4054, right maxillary fragment with P3–M2 (broken), in occlusal view (O). Protypotherium diversidens, holotype MACN-A 4052, right maxillary fragment with alveolus of P1 and P2–M3, in occlusal view (P). Protypotheriumcompressidens, holotype MACN-A 4029, left maxillaryfragment withP3–M3, in occlusal view (Q); holotype MACN-A 4030, left mandibular fragment with p4 (talonid)–m3, in occlusal view (R). Protypotherium lineare, holotype MACN-A4039, right mandibular fragment with alveolus of p1 and p2–m1, in occlusal view (S); holotype MACN-A 4038, left mandibular fragment with p3–m2, in occlusal view (T). Scale bar = 10 mm." httpUri="https://zenodo.org/record/1187701/files/figure.png" pageId="12" pageNumber="485">Figs. 2 M–N</figureCitation>
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), an almost complete skull lacking the left premaxilla.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35F6506CF5F8316C1C229DB8FF9FD3A" pageId="12" pageNumber="485" type="diagnosis">
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<paragraph id="8BFA368DCF5F8316C1C229DB8ABAFDEE" blockId="12.[151,1437,366,1075]" pageId="12" pageNumber="485">
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<emphasis id="B931EA9FCF5F8316C1C229DB8F7AFE7E" bold="true" box="[199,314,402,427]" pageId="12" pageNumber="485">Diagnosis</emphasis>
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(
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<bibRefCitation id="EFD44B7CCF5F8316C04F29DB8C02FE7E" box="[330,578,402,427]" pageId="12" pageNumber="485" refId="ref17076" refString="Ameghino, F. (1891) Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural, 1 (5), 289 - 328." type="journal article">Ameghino 1891: 292</bibRefCitation>
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). “
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<emphasis id="B931EA9FCF5F8316C36629DD8A50FE7E" box="[611,1040,402,427]" italics="true" pageId="12" pageNumber="485">Tamaño bastante más pequeño que el</emphasis>
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<taxonomicName id="4C454D0ECF5F8316C51C29DB8F73FE1B" authority="(Moreno) Ameghino. Se" authorityName="Ameghino. Se" baseAuthorityName="Moreno" baseAuthorityYear="1882" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="485" phylum="Chordata" rank="species" species="australe">
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Protypotherium australe
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<emphasis id="B931EA9FCF5F8316C43729DA8F73FE1B" italics="true" pageId="12" pageNumber="485">(Moreno) Ameghino. Se</emphasis>
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</taxonomicName>
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<emphasis id="B931EA9FCF5F8316C03829FF8AA6FDEE" italics="true" pageId="12" pageNumber="485">distingue por los parietales, que se levantan bastante más arriba que los frontales, de superficie globosa, cortos y que atrás descienden hacia abajo de una manera casi repentina. Frente muy ancha y plana; nasales cortos. Longitud del cráneo, 8 centímetros. Diámetro transverso máximo, 53 milímetros. Diámetro interorbitario de los frontales, 3 centímetros. Longitud de la serie dentaria superior, 45 milímetros</emphasis>
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”.
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</paragraph>
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<paragraph id="8BFA368DCF5F8316C1C22A0F8FF9FD3A" blockId="12.[151,1437,366,1075]" pageId="12" pageNumber="485">
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English translation. Considerably smaller size than
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<taxonomicName id="4C454D0ECF5F8316C21C2A0F8B06FD8B" authority="(Moreno) Ameghino. It" authorityName="Ameghino. It" baseAuthorityName="Moreno" baseAuthorityYear="1882" box="[793,1350,582,607]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="485" phylum="Chordata" rank="species" species="australe">
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<emphasis id="B931EA9FCF5F8316C21C2A0F8A6FFD8A" box="[793,1071,582,607]" italics="true" pageId="12" pageNumber="485">Protypotherium australe</emphasis>
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(Moreno) Ameghino. It
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</taxonomicName>
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can be distinguished by its short parietals that rise much higher than the frontals, with a globular surface, and suddenly descend downwards posteriorly. Forehead very broad and flat; nasals short. Length of the skull, 8 centimetres. Maximum transverse diameter, 53 millimetres. Interorbital diameter of the frontals, 3 centimetres. Length of upper tooth row, 45 millimetres.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35F6506CF5F8316C1C22AB38F42FBE6" pageId="12" pageNumber="485" type="discussion">
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<paragraph id="8BFA368DCF5F8316C1C22AB38D5BFC3F" blockId="12.[151,1437,366,1075]" pageId="12" pageNumber="485">
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<emphasis id="B931EA9FCF5F8316C1C22AB38F0FFCC6" bold="true" box="[199,335,762,787]" pageId="12" pageNumber="485">Comments.</emphasis>
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<bibRefCitation id="EFD44B7CCF5F8316C05B2AB38C6EFCC6" box="[350,558,762,787]" pageId="12" pageNumber="485" refId="ref17076" refString="Ameghino, F. (1891) Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural, 1 (5), 289 - 328." type="journal article">Ameghino (1891)</bibRefCitation>
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described this taxon without any illustration. According to Ameghino’s catalogue and to
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<bibRefCitation id="EFD44B7CCF5F8316C0642B568C45FCE2" box="[353,517,798,823]" pageId="12" pageNumber="485" refId="ref18248" refString="Mones A. (1986) Palaeovertebrata Sudamericana. Catalogo Sistematico de los Vertebrados Fosiles de America del Sur, Parte I. Preliminary List and Bibliography. Courier Forschungsinstitut Senckenberg, 82, 1 - 625." type="journal article">Mones (1986)</bibRefCitation>
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, the incomplete skull MACN-A 4049 is the
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<typeStatus id="54FE882FCF5F8316C5132B568A06FCE2" box="[1046,1094,799,823]" pageId="12" pageNumber="485">type</typeStatus>
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material of the species. This specimen matches the original diagnosis (
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<bibRefCitation id="EFD44B7CCF5F8316C36A2B0B8D68FC8E" box="[623,808,834,859]" pageId="12" pageNumber="485" refId="ref17076" refString="Ameghino, F. (1891) Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural, 1 (5), 289 - 328." type="journal article">Ameghino 1891</bibRefCitation>
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) both in morphology (e.g., the most posterior region of the frontals and the anterior portion of the parietals form a convex surface that rapidly descends posteriorly, frontals are wide and flat, and nasals and parietals are short) and measurements (length of the skull =
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<quantity id="4CBD9B68CF5F8316C5B22BC28B5FFC77" box="[1207,1311,907,931]" metricMagnitude="-2" metricUnit="m" metricValue="8.42" pageId="12" pageNumber="485" unit="mm" value="84.2">84.2 mm</quantity>
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; maximum transverse diameter =
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<quantity id="4CBD9B68CF5F8316C0972BE78FB9FC13" box="[402,505,942,967]" metricMagnitude="-2" metricUnit="m" metricValue="5.34" pageId="12" pageNumber="485" unit="mm" value="53.4">53.4 mm</quantity>
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; interorbital frontal diameter =
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<quantity id="4CBD9B68CF5F8316C25B2BE68D85FC13" box="[862,965,943,967]" metricMagnitude="-2" metricUnit="m" metricValue="3.01" pageId="12" pageNumber="485" unit="mm" value="30.1">30.1 mm</quantity>
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; length of the upper tooth row =
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<quantity id="4CBD9B68CF5F8316C4392BE78BD0FC12" box="[1340,1424,942,967]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="12" pageNumber="485" unit="mm" value="45.0">45 mm</quantity>
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), and it is herein confirmed as the
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<typeStatus id="54FE882FCF5F8316C3002B9B8C27FC3E" box="[517,615,978,1003]" pageId="12" pageNumber="485" type="holotype">holotype</typeStatus>
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of
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<taxonomicName id="4C454D0ECF5F8316C3892B9D8D54FC3E" authorityName="Ameghino" authorityYear="1891" box="[652,788,978,1003]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="12" pageNumber="485" phylum="Chordata" rank="species" species="globosum">
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<emphasis id="B931EA9FCF5F8316C3892B9D8D54FC3E" box="[652,788,978,1003]" italics="true" pageId="12" pageNumber="485">P. globosum</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BFA368DCF5F8316C1C22BBF8F42FBE6" blockId="12.[151,1437,366,1075]" pageId="12" pageNumber="485">
|
|||
|
|
<materialsCitation id="3B2D3CD0CF5F8316C1C22BBF8F42FBE6" ID-GBIF-Occurrence="1828873149" collectionCode="SALMA" location="Santa Cruz Province" pageId="12" pageNumber="485" specimenCount="1" stateProvince="Santa Cruz Province" typeStatus="holotype">
|
|||
|
|
<emphasis id="B931EA9FCF5F8316C1C22BBF8DBEFBDA" bold="true" box="[199,1022,1014,1039]" pageId="12" pageNumber="485">
|
|||
|
|
Chronological and geographical distribution of the
|
|||
|
|
<typeStatus id="54FE882FCF5F8316C24C2BBF8D3DFBDA" box="[841,893,1014,1039]" pageId="12" pageNumber="485">type</typeStatus>
|
|||
|
|
specimen.
|
|||
|
|
</emphasis>
|
|||
|
|
Santacrucian
|
|||
|
|
<collectionCode id="ED54AE48CF5F8316C5A32BBE8B4BFBDA" box="[1190,1291,1015,1039]" pageId="12" pageNumber="485">SALMA</collectionCode>
|
|||
|
|
,
|
|||
|
|
<collectingRegion id="4981F86FCF5F8316C41E2BBE8EBFFBE6" country="Argentina" name="Santa Cruz" pageId="12" pageNumber="485">
|
|||
|
|
Santa
|
|||
|
|
<collectingRegion id="4981F86FCF5F8316C4632BBE8EBFFBE6" inferredBySuffix="true" pageId="12" pageNumber="485">Cruz Province</collectingRegion>
|
|||
|
|
</collectingRegion>
|
|||
|
|
.
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|