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<document id="8D43CDB16A9ABBA3E17340DDDE235511" ID-CLB-Dataset="304811" ID-DOI="10.5252/geodiversitas2024v46a15" ID-GBIF-Dataset="fe16163e-5e9d-4c80-89c8-fce3fb2e064f" ID-ISSN="1638-9395" ID-Zenodo-Dep="13985760" ID-ZooBank="urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1729758589633" checkinUser="plazi" docAuthor="Merle, Didier, Pacaud, Jean-Michel, Ledon, Daniel &amp; Goret, Bernard" docDate="2024" docId="038E87DAFFE9E416FC68FC60FC66F858" docLanguage="en" docName="geodiversitas.2024.46.15.pdf" docOrigin="Geodiversitas 46 (15)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Trubatsa calviniacensis Merle, Pacaud, Ledon &amp; Goret, 2024, n. sp." docType="treatment" docUuid="287FDBCE-A766-4813-9403-2ABDD9B439E3" docUuidSource="ZooBank" docVersion="9" lastPageNumber="509" masterDocId="FFB7FFA2FFE6E406FFB8FFD6FF81FFB3" masterDocTitle="New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe" masterLastPageNumber="551" masterPageNumber="495" pageNumber="508" updateTime="1732586620909" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="7B2CE6B2D0F6F22532BCC389494F89F1">New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe</mods:title>
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</mods:titleInfo>
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<mods:namePart id="9D6A78CEC068BFB0573C130263FA6C5B">Merle, Didier</mods:namePart>
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</mods:name>
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<mods:namePart id="3235ABADF24796F8F7FAF251AE5BA15A">Pacaud, Jean-Michel</mods:namePart>
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</mods:name>
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<mods:namePart id="C5714C12FE6F236B2943A8825E00B6C7">Ledon, Daniel</mods:namePart>
<mods:affiliation id="E9816A1B5513B666924585AD2362D35D">Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr</mods:affiliation>
<mods:nameIdentifier id="63472AD71102D71BDDFBEA93CB54EA56" type="email">didier.merle@mnhn.fr</mods:nameIdentifier>
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<mods:namePart id="3B376385E07BD966002716116FAB506C">Goret, Bernard</mods:namePart>
<mods:affiliation id="BF60FE0B21ACACDDA5DFF964BA261BD3">Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com</mods:affiliation>
<mods:nameIdentifier id="A59DB26CEDB007D300A78E851AEDB520" type="email">bernardgoret@hotmail.com</mods:nameIdentifier>
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<mods:title id="9BB999BED4AE60FD1753430348D85375">Geodiversitas</mods:title>
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<mods:date id="2A72D2011C5E5A95D7BE4C2F61FF95D4">2024</mods:date>
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<mods:classification id="11AA4140B9421EA7322C9753C28935B7">journal article</mods:classification>
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<mods:identifier id="35E902BC05B4CB917BF685420ADBBA70" type="DOI">10.5252/geodiversitas2024v46a15</mods:identifier>
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<treatment id="038E87DAFFE9E416FC68FC60FC66F858" ID-DOI="http://doi.org/10.5281/zenodo.13989270" ID-GBIF-Taxon="244735075" ID-Zenodo-Dep="13989270" LSID="urn:lsid:zoobank.org:act:287FDBCE-A766-4813-9403-2ABDD9B439E3" httpUri="http://treatment.plazi.org/id/038E87DAFFE9E416FC68FC60FC66F858" lastPageId="16" lastPageNumber="509" pageId="15" pageNumber="508" scope_class="Gastropoda" scope_family="Muricidae" scope_phylum="Mollusca" scope_subFamily="Typhinae">
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<subSubSection id="C33D6547FFE9E409FC68FC60FA8DFC7C" box="[976,1292,949,976]" pageId="15" pageNumber="508" type="nomenclature">
<paragraph id="8B9836CCFFE9E409FC68FC60FA8DFC7C" blockId="15.[976,1292,949,1007]" box="[976,1292,949,976]" pageId="15" pageNumber="508">
<heading id="D0D081A0FFE9E409FC68FC60FA8DFC7C" box="[976,1292,949,976]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="508" reason="2">
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<taxonomicName id="4C274D4FFFE9E409FC68FC60FB4AFC63" ID-CoL="CN5MF" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[976,1227,950,976]" class="Gastropoda" family="Muricidae" genus="Trubatsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="508" phylum="Mollusca" rank="species" species="calviniacensis" status="sp. nov.">
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<emphasis id="B953EADEFFE9E409FC68FC60FB4AFC63" bold="true" box="[976,1227,950,976]" italics="true" pageId="15" pageNumber="508">Trubatsa calviniacensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFE9E409FB69FC63FA8DFC7C" box="[1233,1292,949,975]" pageId="15" pageNumber="508" rank="species">n. sp.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="C33D6547FFE9E409FBB3FC03FA31FB99" pageId="15" pageNumber="508" type="description">
<paragraph id="8B9836CCFFE9E409FBB3FC03FB51FC5C" blockId="15.[976,1292,949,1007]" box="[1035,1232,980,1007]" pageId="15" pageNumber="508">
(
<figureCitation id="131C2A49FFE9E409FBACFC02FB11FC5C" box="[1044,1168,980,1007]" captionStart="FIG" captionStartId="14.[132,143,1183,1200]" captionTargetBox="[132,1455,215,1138]" captionTargetId="figure-499@14.[698,1267,213,769]" captionTargetPageId="14" captionText="FIG. 5. — Spiral cords in Trubatsa ganensis n. sp. (A-D), T. parisiensis (d’Orbigny, 1850) (E, F) and T. calviniacensis n. sp. (G, H): A, B, holotype, MNHN.F.A90520 (Varone coll.), Gan (Tuilerie), Pyrénées-Atlantiques, Aquitaine Basin, France, early Eocene (Ypresian); C, paratype, MNHN.F.A90518 (Merle coll.), Gan (Tuilerie); D, paratype, MNHN.F.A90517 (Merle coll.), Gan (Tuilerie); E, F, Alan Morton coll., Upper Barton Bed H, Barton-Highcliff, United Kingdom, middle Eocene (Bartonian); G, H, holotype, MNHN.F.A25590 (Pacaud coll.) Cauvigny (Châteaurouge), Oise, Paris Basin, France, middle Eocene (Lutetian). Scale bars: 5 mm. Credits: A-D, G, H, P. Loubry (MNHN/CNRS); E, F, A. Morton." figureDoi="http://doi.org/10.5281/zenodo.13985770" httpUri="https://zenodo.org/record/13985770/files/figure.png" pageId="15" pageNumber="508">Figs 5G, H</figureCitation>
;
<figureCitation id="131C2A49FFE9E409FB24FC02FB48FC5C" box="[1180,1225,980,1007]" captionStart="FIG" captionStartId="44.[132,143,1530,1547]" captionTargetBox="[132,1455,215,1484]" captionTargetId="figure-374@44.[700,1257,764,1304]" captionTargetPageId="44" captionText="FIG. 25. — New species of Muricopsis Bucquoy &amp; Dautzenberg, 1882, Favartia Jousseaume, 1880, Dermomurex Monterosato, 1890, Trubatsa Dall, 1889 and Timbellus De Gregorio, 1885: A, B, Muricopsis pontileviensis n. sp.: A, holotype, MNHN.F.A71120 (Dollfus coll.), Pontlevoy, France, Langhian; B, paratype, MNHN.F.A77743, (Dollfus coll.), Pontlevoy; C, D, M. neraudeaui n. sp.: C, holotype, MNHN.F.A77746 (Néraudeau sample), Saint-Jacques-de-la-Lande, France, Messinian;D, paratype,MNHN.F.A77747 (Néraudeau sample),Saint-Jacques-de-la-Lande;E, F, Favartia jansseni n. sp.:E, holotype,RGM.225979,Winterswijk,The Netherlands, Langhian; F, paratype,MNHN.F.A77748 (Ledon coll.),Winterswijk;G, H, Dermomurex (Gracilimurex) ligerianus n. sp.: G, holotype,MNHN.F.A33456 (Dollfus coll.), Pontlevoy; H, paratype, MNHN.F.A33457 (Dollfus coll.); I-K, Trubatsa ganensis n. sp.: I, holotype, MNHN.A90520 (Varone coll.), Gan (Tuilerie), France, Ypresian; J, paratype, MNHN.F.A90518 (Merle coll.), Gan (Tuilerie); K, paratype, MNHN.F.A90517 (Merle coll.), Gan (Tuilerie); L, T. calviniacensis n. sp., holotype,MNHN.F.A25590 (Pacaud coll.), Cauvigny (Châteaurouge),France, Lutetian; M-O, Timbellus magnificus n. sp.: M, holotype, MNHN.F.A90522 (Pacaud coll.), Baron (Sablière Heudebert), France, Bartonian; N, paratype, MNHN.F.A90525 (Schtrock coll.), Saint-Vaast-lès-Mello (Barisseuse), France, Bartonian; O, paratype, MNHN.F.A90523 (Pacaud coll.), Baron (Sablière Heudebert). Scale bars: A-L, 5 mm; M-O, 20 mm." figureDoi="http://doi.org/10.5281/zenodo.13985823" httpUri="https://zenodo.org/record/13985823/files/figure.png" pageId="15" pageNumber="508">25L</figureCitation>
)
</paragraph>
<paragraph id="8B9836CCFFE9E409FC95FBCFFA31FB99" blockId="15.[813,1456,1049,1066]" box="[813,1456,1049,1066]" pageId="15" pageNumber="508">
<uri id="FFB63ACEFFE9E409FC95FBCFFA31FB99" box="[813,1456,1049,1066]" pageId="15" pageNumber="508">
urn:lsid:zoobank.org:act:
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</subSubSection>
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<taxonomicName id="4C274D4FFFE9E409FC95FB8DFC66FBC1" ID-CoL="8YPH6" baseAuthorityName="d'Orbigny" baseAuthorityYear="1850" box="[813,999,1115,1139]" class="Gastropoda" family="Muricidae" genus="Trubatsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="508" phylum="Mollusca" rank="species" species="parisiensis">
<emphasis id="B953EADEFFE9E409FC95FB8DFC66FBC1" box="[813,999,1115,1139]" italics="true" pageId="15" pageNumber="508">Trubatsa parisiensis</emphasis>
</taxonomicName>
–
<treatmentCitation id="0A8610DDFFE9E409FBBDFB8DFA8CFBC1" author="MERLE D. &amp; PACAUD J. - M." box="[1029,1293,1114,1139]" page="16" pageId="15" pageNumber="508" year="2019">
<bibRefCitation id="EFB64B3DFFE9E409FBBDFB8DFA8CFBC1" author="MERLE D. &amp; PACAUD J. - M." box="[1029,1293,1114,1139]" pageId="15" pageNumber="508" pagination="11 - 22" refId="ref47064" refString="MERLE D. &amp; PACAUD J. - M. 2019. - Five new muricid species (Mollusca, Gastropoda) from the Eocene of France, Senegal and Pakistan. Xenophora Taxonomy 25: 11 - 22." type="journal article" year="2019">Merle &amp; Pacaud 2019: 16</bibRefCitation>
</treatmentCitation>
, 17, text-fig 3.5 [
<emphasis id="B953EADEFFE9E409FC8DFBA0FCD8FB3E" box="[821,857,1142,1165]" italics="true" pageId="15" pageNumber="508">non</emphasis>
d’Orbigny, 1850].
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<materialsCitation id="3B4F3C91FFE9E409FC95FB73FBD4FABF" ID-GBIF-Occurrence="4958671420" collectionCode="MNHN" collectorName="Pacaud" country="France" county="Lutetian" location="Paris Basin" municipality="Eocene" pageId="15" pageNumber="508" specimenCount="1" typeStatus="holotype">
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. —
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.
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•
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,
<location id="8EF86017FFE9E409FAA0FB73FAC6FB0E" LSID="urn:lsid:plazi:treatment:038E87DAFFE9E416FC68FC60FC66F858:8EF86017FFE9E409FAA0FB73FAC6FB0E" box="[1304,1351,1189,1213]" country="France" county="Lutetian" municipality="Eocene" name="Oise" pageId="15" pageNumber="508">Oise</location>
,
<location id="8EF86017FFE9E409FAE8FB73FA2FFB0E" LSID="urn:lsid:plazi:treatment:038E87DAFFE9E416FC68FC60FC66F858:8EF86017FFE9E409FAE8FB73FA2FFB0E" box="[1360,1454,1189,1213]" country="France" county="Lutetian" municipality="Eocene" name="Cauvigny" pageId="15" pageNumber="508">Cauvigny</location>
(
<location id="8EF86017FFE9E409FC8AFB69FC42FB64" LSID="urn:lsid:plazi:treatment:038E87DAFFE9E416FC68FC60FC66F858:8EF86017FFE9E409FC8AFB69FC42FB64" box="[818,963,1215,1239]" country="France" county="Lutetian" municipality="Eocene" name="Chateaurouge" pageId="15" pageNumber="508">Châteaurouge</location>
);
<location id="8EF86017FFE9E409FC6DFB69FBA7FB64" LSID="urn:lsid:plazi:treatment:038E87DAFFE9E416FC68FC60FC66F858:8EF86017FFE9E409FC6DFB69FBA7FB64" box="[981,1062,1215,1239]" country="France" county="Lutetian" municipality="Eocene" name="Calcaire" pageId="15" pageNumber="508">Calcaire</location>
grossier moyen (biozone NP15); middle
<collectingMunicipality id="6BFCACB6FFE9E409FCEAFB0CFC18FB42" box="[850,921,1242,1265]" pageId="15" pageNumber="508">Eocene</collectingMunicipality>
(middle
<collectingCounty id="62F94E40FFE9E409FC4EFB0FFBCCFB42" box="[1014,1101,1241,1265]" pageId="15" pageNumber="508">Lutetian</collectingCounty>
);
<collectionCode id="ED36AE09FFE9E409FBD9FB0CFB34FB42" box="[1121,1205,1242,1265]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="508">MNHN</collectionCode>
.F.A25590 (
<collectorName id="26D2531AFFE9E409FA90FB0FFAEEFB42" box="[1320,1391,1241,1265]" pageId="15" pageNumber="508">Pacaud</collectorName>
coll.), H:
<quantity id="4CDF9B29FFE9E409FCF7FB22FC16FAB8" box="[847,919,1268,1292]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="15" pageNumber="508" unit="mm" value="17.0">17 mm</quantity>
(
<figureCitation id="131C2A49FFE9E409FC1CFB22FB92FAB8" box="[932,1043,1268,1292]" captionStart="FIG" captionStartId="14.[132,143,1183,1200]" captionTargetBox="[132,1455,215,1138]" captionTargetId="figure-499@14.[698,1267,213,769]" captionTargetPageId="14" captionText="FIG. 5. — Spiral cords in Trubatsa ganensis n. sp. (A-D), T. parisiensis (d’Orbigny, 1850) (E, F) and T. calviniacensis n. sp. (G, H): A, B, holotype, MNHN.F.A90520 (Varone coll.), Gan (Tuilerie), Pyrénées-Atlantiques, Aquitaine Basin, France, early Eocene (Ypresian); C, paratype, MNHN.F.A90518 (Merle coll.), Gan (Tuilerie); D, paratype, MNHN.F.A90517 (Merle coll.), Gan (Tuilerie); E, F, Alan Morton coll., Upper Barton Bed H, Barton-Highcliff, United Kingdom, middle Eocene (Bartonian); G, H, holotype, MNHN.F.A25590 (Pacaud coll.) Cauvigny (Châteaurouge), Oise, Paris Basin, France, middle Eocene (Lutetian). Scale bars: 5 mm. Credits: A-D, G, H, P. Loubry (MNHN/CNRS); E, F, A. Morton." figureDoi="http://doi.org/10.5281/zenodo.13985770" httpUri="https://zenodo.org/record/13985770/files/figure.png" pageId="15" pageNumber="508">Figs 5G, H</figureCitation>
;
<figureCitation id="131C2A49FFE9E409FBA6FB22FBC9FABF" box="[1054,1096,1268,1292]" captionStart="FIG" captionStartId="44.[132,143,1530,1547]" captionTargetBox="[132,1455,215,1484]" captionTargetId="figure-374@44.[700,1257,764,1304]" captionTargetPageId="44" captionText="FIG. 25. — New species of Muricopsis Bucquoy &amp; Dautzenberg, 1882, Favartia Jousseaume, 1880, Dermomurex Monterosato, 1890, Trubatsa Dall, 1889 and Timbellus De Gregorio, 1885: A, B, Muricopsis pontileviensis n. sp.: A, holotype, MNHN.F.A71120 (Dollfus coll.), Pontlevoy, France, Langhian; B, paratype, MNHN.F.A77743, (Dollfus coll.), Pontlevoy; C, D, M. neraudeaui n. sp.: C, holotype, MNHN.F.A77746 (Néraudeau sample), Saint-Jacques-de-la-Lande, France, Messinian;D, paratype,MNHN.F.A77747 (Néraudeau sample),Saint-Jacques-de-la-Lande;E, F, Favartia jansseni n. sp.:E, holotype,RGM.225979,Winterswijk,The Netherlands, Langhian; F, paratype,MNHN.F.A77748 (Ledon coll.),Winterswijk;G, H, Dermomurex (Gracilimurex) ligerianus n. sp.: G, holotype,MNHN.F.A33456 (Dollfus coll.), Pontlevoy; H, paratype, MNHN.F.A33457 (Dollfus coll.); I-K, Trubatsa ganensis n. sp.: I, holotype, MNHN.A90520 (Varone coll.), Gan (Tuilerie), France, Ypresian; J, paratype, MNHN.F.A90518 (Merle coll.), Gan (Tuilerie); K, paratype, MNHN.F.A90517 (Merle coll.), Gan (Tuilerie); L, T. calviniacensis n. sp., holotype,MNHN.F.A25590 (Pacaud coll.), Cauvigny (Châteaurouge),France, Lutetian; M-O, Timbellus magnificus n. sp.: M, holotype, MNHN.F.A90522 (Pacaud coll.), Baron (Sablière Heudebert), France, Bartonian; N, paratype, MNHN.F.A90525 (Schtrock coll.), Saint-Vaast-lès-Mello (Barisseuse), France, Bartonian; O, paratype, MNHN.F.A90523 (Pacaud coll.), Baron (Sablière Heudebert). Scale bars: A-L, 5 mm; M-O, 20 mm." figureDoi="http://doi.org/10.5281/zenodo.13985823" httpUri="https://zenodo.org/record/13985823/files/figure.png" pageId="15" pageNumber="508">25L</figureCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFE9E409FC95FAF2FBDCFAE5" pageId="15" pageNumber="508" type="etymology">
<paragraph id="8B9836CCFFE9E409FC95FAF2FBDCFAE5" blockId="15.[813,1456,1316,1366]" pageId="15" pageNumber="508">
<smallCapsWord id="8D7EA010FFE9E409FC95FAF2FC27FA89" baselines="1333,1334" box="[813,934,1316,1339]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="15" pageNumber="508">ETYMOLOGY</smallCapsWord>
. — From
<emphasis id="B953EADEFFE9E409FBB6FAF2FBFDFA8F" box="[1038,1148,1316,1340]" italics="true" pageId="15" pageNumber="508">Calviniacus</emphasis>
, the Latin name of the
<typeStatus id="549C886EFFE9E409FADCFAF2FA0FFA8F" box="[1380,1422,1316,1340]" pageId="15" pageNumber="508">type</typeStatus>
locality Cauvigny (Oise,
<collectingCountry id="F330765CFFE9E409FBB5FAE8FBD2FAE5" box="[1037,1107,1342,1366]" name="France" pageId="15" pageNumber="508">France</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFE9E409FC95FAB8FBB5FA59" pageId="15" pageNumber="508" type="materials_examined">
<paragraph id="8B9836CCFFE9E409FC95FAB8FA1AFA13" blockId="15.[813,1455,1390,1440]" pageId="15" pageNumber="508">
<smallCapsWord id="8D7EA010FFE9E409FC95FAB8FCDFFA30" baselines="1407,1407" box="[813,862,1390,1413]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="15" pageNumber="508">
<typeStatus id="549C886EFFE9E409FC95FAB8FCDFFA30" box="[813,862,1390,1413]" pageId="15" pageNumber="508">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D7EA010FFE9E409FCDEFAA5FC43FA37" baselines="1408" box="[870,962,1395,1412]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="horizon" pageId="15" pageNumber="508">HORIZON</smallCapsWord>
. — Calcaire grossier moyen (biozone NP15), see
<bibRefCitation id="EFB64B3DFFE9E409FC96FA5EFB80FA13" author="GELY J. - P. &amp; LORENZ C." box="[814,1025,1416,1440]" pageId="15" pageNumber="508" pagination="713 - 747" refId="ref44561" refString="GELY J. - P. &amp; LORENZ C. 1991. - Analyse sequentielle de l'Eocene et de l'Oligocene du Bassin parisien (France). Revue de l'institut francais du petrole 46 (6): 713 - 747." type="journal article" year="1991">Gély &amp; Lorenz (1991</bibRefCitation>
: pl. 1); middle Eocene (middle Lutetian).
</paragraph>
<paragraph id="8B9836CCFFE9E409FC95FA6EFBB5FA59" blockId="15.[813,1455,1464,1514]" pageId="15" pageNumber="508">
<smallCapsWord id="8D7EA010FFE9E409FC95FA6EFCDCFA7D" baselines="1481,1482" box="[813,861,1464,1487]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="15" pageNumber="508">
<typeStatus id="549C886EFFE9E409FC95FA6EFCDCFA7D" box="[813,861,1464,1487]" pageId="15" pageNumber="508">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D7EA010FFE9E409FCDAFA6BFC3CFA7D" baselines="1482" box="[866,957,1469,1486]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="locality" pageId="15" pageNumber="508">LOCALITY</smallCapsWord>
. —
<collectingCountry id="F330765CFFE9E409FC53FA6EFBAEFA7C" box="[1003,1071,1464,1487]" name="France" pageId="15" pageNumber="508">France</collectingCountry>
, Paris Basin, Oise, Cauvigny (Châteaurouge), see
<bibRefCitation id="EFB64B3DFFE9E409FC2BFA04FBA9FA59" author="FRITEL P. H." box="[915,1064,1490,1514]" pageId="15" pageNumber="508" refId="ref44321" refString="FRITEL P. H. 1910. - Guide geologique et paleontologique de la region parisienne dans un rayon de 100 kilometres. Deyrolle, Paris, 356 p." type="book" year="1910">Fritel 1910: 85</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFE9E409FC95F9D4FAC0F9A9" box="[813,1345,1538,1562]" pageId="15" pageNumber="508" type="distribution">
<paragraph id="8B9836CCFFE9E409FC95F9D4FAC0F9A9" blockId="15.[813,1345,1538,1562]" box="[813,1345,1538,1562]" pageId="15" pageNumber="508">
<smallCapsWord id="8D7EA010FFE9E409FC95F9D4FC3BF9AB" baselines="1555,1556" box="[813,954,1538,1561]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Distribution" pageId="15" pageNumber="508">DISTRIBUTION</smallCapsWord>
. — Only known from the
<typeStatus id="549C886EFFE9E409FB7CF9D4FB6FF9A9" box="[1220,1262,1538,1562]" pageId="15" pageNumber="508">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFE9E416FC95F985FC66F858" lastPageId="16" lastPageNumber="509" pageId="15" pageNumber="508" type="description">
<paragraph id="8B9836CCFFE9E409FC95F985FC3CF9D8" blockId="15.[811,1457,1619,2028]" box="[813,957,1619,1645]" pageId="15" pageNumber="508">
<smallCapsWord id="8D7EA010FFE9E409FC95F985FC3CF9D8" baselines="1638,1638" box="[813,957,1619,1645]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="15" pageNumber="508">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="8B9836CCFFE9E416FC95F9A4FED7F95F" blockId="15.[811,1457,1619,2028]" lastBlockId="16.[131,777,1650,1772]" lastPageId="16" lastPageNumber="509" pageId="15" pageNumber="508">
Bulbous protoconch of two whorls; pointed apex.Teleoconch up to
<quantity id="4CDF9B29FFE9E409FCD6F944FC52F91E" box="[878,979,1682,1709]" metricMagnitude="-2" metricUnit="m" metricValue="1.78" pageId="15" pageNumber="508" unit="mm" value="17.8">17.8 mm</quantity>
in height, up to
<quantity id="4CDF9B29FFE9E409FB29F944FB69F91E" box="[1169,1256,1682,1709]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="15" pageNumber="508" unit="mm" value="9.2">9.2 mm</quantity>
in width, biconic in profile, composed of six whorls. High spire. Last whorl representing up to 70% total height. Spiral angle 62°. Suture weakly impressed. Axial sculpture consisting of four slightly spiny varices per whorl. On spire, anal tube (P1) and short spine (P2). Anal tube forming angle of approximately 45° with shell axis. Anal tube (P1) totally integrated in varices eliminating shoulder. Short P2 spine on first whorls. No callus at base of anal tube. On convex part of last whorl and abapically to anal tube, five spinelets (P2, P3, P4, P5 and P6). Intervarical spaces with weak P2 to P5 cords. Ovate aperture up to 30% of diameter, up to 74% of height of last whorl (including siphonal canal). Edge, erect, smooth. peristome complete. Siphonal canal closed, occupying 47% of apertural lenght.
</paragraph>
<caption id="DF586644FFF6E416FF3CFA6DFDE4F98E" ID-DOI="http://doi.org/10.5281/zenodo.13985772" ID-Zenodo-Dep="13985772" httpUri="https://zenodo.org/record/13985772/files/figure.png" pageId="16" pageNumber="509" startId="16.[132,143,1467,1484]" targetBox="[132,1455,215,1424]" targetPageId="16" targetType="figure">
<paragraph id="8B9836CCFFF6E416FF3CFA6DFDE4F98E" blockId="16.[132,1457,1467,1597]" pageId="16" pageNumber="509">
<smallCapsWord id="8D7EA010FFF6E416FF3CFA6DFF1CFA78" baselines="1480,1480" box="[132,157,1467,1484]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="16" pageNumber="509">FIG</smallCapsWord>
. 6. — Neotype, historical figures, spiral cords and denticles of the outer lip in
<taxonomicName id="4C274D4FFFF6E416FC8CFA6DFBB8FA7F" authority="(Lamarck, 1822)" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[820,1081,1467,1484]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="16" pageNumber="509" phylum="Mollusca" rank="species" species="tripteroides">
<emphasis id="B953EADEFFF6E416FC8CFA6DFC29FA7F" box="[820,936,1467,1484]" italics="true" pageId="16" pageNumber="509">T. tripteroides</emphasis>
(
<bibRefCitation id="EFB64B3DFFF6E416FC0BFA6DFBB5FA7F" author="LAMARCK J. - B." box="[947,1076,1467,1484]" pageId="16" pageNumber="509" refId="ref45696" refString="LAMARCK J. - B. 1822. - Histoire naturelle des animaux sans vertebres presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent et Supplement a divers types de gasteropodes et de Trachelipodes comprenant l'indication des coquilles fossiles qui ne furent point citees dans leurs genres respectifs. Tome 7. Lamarck, Paris, 711 p." type="book" year="1822">Lamarck, 1822</bibRefCitation>
)
</taxonomicName>
:
<emphasis id="B953EADEFFF6E416FBF9FA6DFBE2FA7F" bold="true" box="[1089,1123,1467,1484]" pageId="16" pageNumber="509">A -D</emphasis>
, morphotype with rounded varices and poorly distinguishable P1 spine;
<emphasis id="B953EADEFFF6E416FE2FFA04FE39FA50" bold="true" box="[407,440,1490,1507]" pageId="16" pageNumber="509">E -H</emphasis>
, morphotype with easily distinguishable P1 spine:
<emphasis id="B953EADEFFF6E416FCDEFA04FCF2FA50" bold="true" box="[870,883,1490,1507]" pageId="16" pageNumber="509">A</emphasis>
, original figure of
<bibRefCitation id="EFB64B3DFFF6E416FBB4FA04FB08FA50" author="LAMARCK J. - B." box="[1036,1161,1490,1507]" pageId="16" pageNumber="509" pagination="222 - 228" refId="ref45602" refString="LAMARCK J. - B. 1805. - Sur les fossiles des environs de Paris. Explication des planches relatives aux coquilles fossiles des environs de Paris. Annales du Museum d'Histoire Naturelle 6 (33): 222 - 228." type="journal article" year="1805">Lamarck (1805</bibRefCitation>
: pl. 3, fig. 4);
<emphasis id="B953EADEFFF6E416FB47FA04FA8DFA50" bold="true" box="[1279,1292,1490,1507]" pageId="16" pageNumber="509">B</emphasis>
,
<emphasis id="B953EADEFFF6E416FAA0FA04FAA7FA50" bold="true" box="[1304,1318,1490,1507]" pageId="16" pageNumber="509">C</emphasis>
, neotype designated herein, MNHN.F.A90546 (Club géologique d'Île-de-France leg.), Thiverval-Grignon (Falunière), Yvelines, Paris Basin, France, middle Eocene (Lutetian);
<emphasis id="B953EADEFFF6E416FF3CFA29FF13F9A3" bold="true" box="[132,146,1535,1552]" pageId="16" pageNumber="509">D</emphasis>
, MNHN.F.A90552 (Pacaud coll.), Chaussy (Les Garennes),Paris Basin, France,middle Eocene (Lutetian);
<emphasis id="B953EADEFFF6E416FC44FA29FB89F9A3" bold="true" box="[1020,1032,1535,1552]" pageId="16" pageNumber="509">E</emphasis>
,
<emphasis id="B953EADEFFF6E416FBAAFA29FB9CF9A3" bold="true" box="[1042,1053,1535,1552]" pageId="16" pageNumber="509">F</emphasis>
, figured specimen of
<bibRefCitation id="EFB64B3DFFF6E416FB68FA29FAD5F9A3" author="DESHAYES G. - P." box="[1232,1364,1535,1552]" pageId="16" pageNumber="509" pagination="79 - 106" refId="ref43930" refString="DESHAYES G. - P. 1835. - Description des coquilles fossiles des environs de Paris. Tome 2. Livraisons 40 - 45, l'auteur, Bechet Jeune, Baudouin freres, Treuttel &amp; Wurtz, Paris: 499 - 780; Atlas 2, Levrault, Paris: pl. 79 - 106. https: // doi. org / 10.5962 / bhl. title. 52303" type="book chapter" year="1835">Deshayes (1835</bibRefCitation>
: pl. 82, fig. 1-2);
<emphasis id="B953EADEFFF6E416FF08F9C0FF3FF994" bold="true" box="[176,190,1558,1575]" pageId="16" pageNumber="509">G</emphasis>
,
<emphasis id="B953EADEFFF6E416FF70F9C0FF57F994" bold="true" box="[200,214,1558,1575]" pageId="16" pageNumber="509">H</emphasis>
, figured specimen of
<bibRefCitation id="EFB64B3DFFF6E416FE34F9C0FDF5F994" author="COSSMANN M. &amp; PISSARRO G." box="[396,628,1558,1575]" pageId="16" pageNumber="509" pagination="26 - 45" refId="ref43883" refString="COSSMANN M. &amp; PISSARRO G. 1911. - Iconographie complete des coquilles fossiles de l'Eocene du Bassin de Paris. Tome 2. Hermann, Paris: pl. 26 - 45. https: // www. biodiversitylibrary. org / page / 36319938" type="book chapter" year="1911">Cossmann &amp; Pissarro (1911</bibRefCitation>
: pl. 35, fig. 169-1; MNHN.F.J10350 (Cossmann coll.), Chaussy (Les Garennes). Scale bars: 20 mm. Credits: B-D, P. Loubry MNHN/CNRS; G, H, J. Mouchart.
</paragraph>
</caption>
<paragraph id="8B9836CCFFF6E416FF3CF8C4FE9EF899" blockId="16.[132,775,1810,2028]" box="[132,287,1810,1836]" pageId="16" pageNumber="509">
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</paragraph>
<paragraph id="8B9836CCFFF6E416FF3CF8E7FC66F858" blockId="16.[132,775,1810,2028]" lastBlockId="16.[811,1457,1650,2027]" pageId="16" pageNumber="509">
<bibRefCitation id="EFB64B3DFFF6E416FF3CF8E7FE03F8FF" author="MERLE D. &amp; PACAUD J. - M." box="[132,386,1841,1868]" pageId="16" pageNumber="509" pagination="11 - 22" refId="ref47064" refString="MERLE D. &amp; PACAUD J. - M. 2019. - Five new muricid species (Mollusca, Gastropoda) from the Eocene of France, Senegal and Pakistan. Xenophora Taxonomy 25: 11 - 22." type="journal article" year="2019">Merle &amp; Pacaud (2019</bibRefCitation>
: pl. 3, fig. 4) attributed a specimen (MNHN.F.A25590) from the middle Lutetian of Cauvigny (Paris Basin) to
<taxonomicName id="4C274D4FFFF6E416FED0F8A4FD4BF83F" ID-CoL="8YPH6" authority="(d'Orbigny, 1850)" baseAuthorityName="d'Orbigny" baseAuthorityYear="1850" box="[360,714,1905,1932]" class="Gastropoda" family="Muricidae" genus="Trubatsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="16" pageNumber="509" phylum="Mollusca" rank="species" species="parisiensis">
<emphasis id="B953EADEFFF6E416FED0F8A4FE73F838" box="[360,498,1906,1931]" italics="true" pageId="16" pageNumber="509">T. parisiensis</emphasis>
(d’Orbigny, 1850)
</taxonomicName>
. Like
<taxonomicName id="4C274D4FFFF6E416FF3DF844FF79F818" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[133,248,1938,1963]" class="Gastropoda" family="Muricidae" genus="Trubatsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="16" pageNumber="509" phylum="Mollusca" rank="species" species="ganensis" status="n. sp.">
<emphasis id="B953EADEFFF6E416FF3DF844FF79F818" box="[133,248,1938,1963]" italics="true" pageId="16" pageNumber="509">T. ganensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFF6E416FF47F844FEBDF81F" box="[255,316,1938,1964]" pageId="16" pageNumber="509" rank="species">n. sp.</taxonomicNameLabel>
it has a paucispiral protoconch and lacks partition. Therefore, it can easily be distinguished from
<taxonomicName id="4C274D4FFFF6E416FF3CF804FE8DF858" ID-CoL="8YPH6" baseAuthorityName="d'Orbigny" baseAuthorityYear="1850" box="[132,268,2002,2027]" class="Gastropoda" family="Muricidae" genus="Trubatsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="16" pageNumber="509" phylum="Mollusca" rank="species" species="parisiensis">
<emphasis id="B953EADEFFF6E416FF3CF804FE8DF858" box="[132,268,2002,2027]" italics="true" pageId="16" pageNumber="509">T. parisiensis</emphasis>
</taxonomicName>
from the
<collectingRegion id="49E3F82EFFF6E416FECFF807FE72F858" box="[375,499,2001,2027]" country="United Kingdom" name="Hampshire" pageId="16" pageNumber="509">Hampshire</collectingRegion>
Basin which has a multispiral protoconch.
<taxonomicName id="4C274D4FFFF6E416FC40F9A4FB68F93F" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[1016,1257,1650,1676]" class="Gastropoda" family="Muricidae" genus="Trubatsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="16" pageNumber="509" phylum="Mollusca" rank="species" species="calviniacensis" status="sp. nov.">
<emphasis id="B953EADEFFF6E416FC40F9A4FB68F93F" box="[1016,1257,1650,1676]" italics="true" pageId="16" pageNumber="509">Trubatsa calviniacensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFF6E416FB48F9A5FAABF93E" box="[1264,1322,1651,1677]" pageId="16" pageNumber="509" rank="species">n. sp.</taxonomicNameLabel>
differs from
<taxonomicName id="4C274D4FFFF6E416FC93F945FC1EF91F" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[811,927,1683,1708]" class="Gastropoda" family="Muricidae" genus="Trubatsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="16" pageNumber="509" phylum="Mollusca" rank="species" species="ganensis" status="n. sp.">
<emphasis id="B953EADEFFF6E416FC93F945FC1EF91F" box="[811,927,1683,1708]" italics="true" pageId="16" pageNumber="509">T. ganensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFF6E416FC11F945FC67F91E" box="[937,998,1683,1709]" pageId="16" pageNumber="509" rank="species">n. sp.</taxonomicNameLabel>
in being higher spired, the primary cord P6 on the last whorl, weak primary cords P2 to P
<quantity id="4CDF9B29FFF6E416FAE0F964FA05F97F" box="[1368,1412,1714,1740]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="16" pageNumber="509" unit="in" value="4.0">4 in</quantity>
the intervarical spaces, and by having a more ovate aperture. P2 spine is also more marked on the first teleoconch whorls. Although we have only
<specimenCount id="9D21FD45FFF6E416FB8CF8C4FB4FF89F" box="[1076,1230,1810,1836]" count="1" pageId="16" pageNumber="509" type="generic">one specimen</specimenCount>
, the number of shell differences with the Gan specimens and its occurrence in a strongly different paleoenvironment (deep water at Gan (
<bibRefCitation id="EFB64B3DFFF6E416FC8EF8A7FC3DF83F" author="MERLE D." box="[822,956,1905,1932]" pageId="16" pageNumber="509" pagination="257 - 262" refId="ref46465" refString="MERLE D. 1985. - Les communautes fossiles en tant qu'indicateurs paleobiologiques de milieux: exemples pris sur le Cuisien de Gan (Pyrenees-Atlantiques) et l'Egerien de Hongrie. Bulletin du Museum national d'Histoire naturelle, Paris 4 eme ser. 7, sect. C, 24: 257 - 262." type="journal article" year="1985">Merle 1985</bibRefCitation>
;
<bibRefCitation id="EFB64B3DFFF6E416FC76F8A7FB3FF838" author="MERLE D. &amp; ROUX M." box="[974,1214,1905,1932]" pageId="16" pageNumber="509" pagination="225 - 244" refId="ref47176" refString="MERLE D. &amp; ROUX M. 2018. - Stalked crinoids from Gan (Late Ypresian, southwestern France). Exceptional stereom preservation, paleoecology and taxonomic affinities. Swiss Journal of Paleontology 137: 225 - 244. https: // doi. org / 10.1007 / s 13358 - 018 - 0162 - 0" type="journal article" year="2018">
<taxonomicName id="4C274D4FFFF6E416FC76F8A7FB91F838" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[974,1040,1905,1931]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="16" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
&amp; Roux 2018
</bibRefCitation>
) in
<collectingRegion id="49E3F82EFFF6E416FB48F8A7FADEF838" box="[1264,1375,1905,1931]" country="France" name="Aquitaine" pageId="16" pageNumber="509">Aquitaine</collectingRegion>
Basin, versus shallow water in the Paris Basin (
<bibRefCitation id="EFB64B3DFFF6E416FB42F847FAF5F818" author="GELY J. - P." box="[1274,1396,1937,1963]" pageId="16" pageNumber="509" refId="ref44503" refString="GELY J. - P. 2008. - La stratigraphie et la paleogeographie du Lutetien en France, in MERLE D. (ed.), Stratotype Lutetien. Museum national d'Histoire naturelle, Paris; Biotope, Meze; BRGM, Orleans: 76 - 104 (coll. Patrimoine geologique; 1)." type="journal volume" year="2008">Gély 2008</bibRefCitation>
) and also quite different age, highly suggest that it represents a different species.
</paragraph>
</subSubSection>
</treatment>
</document>
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