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<mods:title id="3C5385EF40A17F5ABD4EB8966E052E25">Taxonomy of the Mexican species of Belonuchus Nordmann (Coleoptera: Staphylinidae)</mods:title>
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<mods:namePart id="2C70AC602F93650E00BBA9CE356DAB36">Asiain, Julieta</mods:namePart>
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<heading id="D0D081F377607F4C81AFA4C6FD14FCF0" bold="true" box="[151,651,762,789]" fontSize="11" level="2" pageId="79" pageNumber="80" reason="4">
<taxonomicName id="4C274D1C77607F4C81AFA4C6FD14FCF0" ID-CoL="LC9T" authority="Bernhauer, 1918" authorityName="Bernhauer" authorityYear="1918" box="[151,651,762,789]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="nigerrimus">
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<emphasis id="B953EA8D77607F4C81AFA4C6FE29FCF0" bold="true" box="[151,438,762,789]" italics="true" pageId="79" pageNumber="80">Belonuchus nigerrimus</emphasis>
<bibRefCitation id="EFB64B6E77607F4C8085A4C6FD14FCF0" author="Bernhauer, M." box="[445,651,762,788]" pageId="79" pageNumber="80" pagination="225 - 226" refId="ref111961" refString="Bernhauer, M. (1918) Vier neue Belonuchus aus Mexiko [cont.]. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 67, 225 - 226." type="journal article" year="1918">Bernhauer, 1918</bibRefCitation>
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<subSubSection id="C33D651477607F4C81AFA522FE75FCDC" box="[151,490,798,824]" pageId="79" pageNumber="80" type="description">
<paragraph id="8B98369F77607F4C81AFA522FE75FCDC" blockId="79.[151,651,762,824]" box="[151,490,798,824]" pageId="79" pageNumber="80">
<figureCitation id="131C2A1A77607F4C81AFA522FF6CFCDC" box="[151,243,798,824]" captionStart="FIGURE 8" captionStartId="102.[151,250,1716,1740]" captionTargetBox="[174,1414,190,1687]" captionTargetId="figure-25@102.[163,1425,181,1692]" captionTargetPageId="102" captionText="FIGURE 8. Dorsal view of males of: a) Belonuchus aenigmaticus (paratype), b) B. nigerrimus, c) B. optatus, d) B. platypterus, e) B. juanmorronei, f) B. abnormalis, g) B. pectinipes (paralectotype), h) B. viridipennis. Scale bars = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6630903" httpUri="https://zenodo.org/record/6630903/files/figure.png" pageId="79" pageNumber="80">Figs. 8b</figureCitation>
, 
<figureCitation id="131C2A1A77607F4C81C7A522FEB8FCDC" box="[255,295,798,824]" captionStart="FIGURE 10" captionStartId="104.[151,250,1839,1863]" captionTargetBox="[176,1418,194,1806]" captionTargetId="figure-25@104.[152,1435,181,1816]" captionTargetPageId="104" captionText="FIGURE 10. Dorsal view of the head of males of: a) Belonuchus julietitae, b) B. singularis, c) B. connexus, d) B. platypterus, e) B. bidens, f) B. nigerrimus, g) B. optatus, h) B. cyanipennis, i) B. godmani. Dorsal view of the head and pronotum of the holotype female of: j) B. optatus." figureDoi="http://doi.org/10.5281/zenodo.6630909" httpUri="https://zenodo.org/record/6630909/files/figure.png" pageId="79" pageNumber="80">10f</figureCitation>
, 
<figureCitation id="131C2A1A77607F4C800BA522FEC1FCDC" box="[307,350,798,824]" captionStart="FIGURE 16" captionStartId="110.[151,250,1818,1842]" captionTargetBox="[183,1402,184,1781]" captionTargetId="figure-25@110.[170,1417,181,1794]" captionTargetPageId="110" captionText="FIGURE 16. Male pregenital sternite of: a) Belonuchus moroni, b) B. dichrous, c) B. flavipennis, d) B. aenigmaticus, e) B. nigerrimus, f) B. juanmorronei, g) B. godmani, h) B. magnistylus, i) B. trochanterinus, j) B. rufipennis, k) B. basiventris, l) B. connexus, m) B. xanthomelas, n) B. ephippiatus, o) B. optatus." figureDoi="http://doi.org/10.5281/zenodo.6630923" httpUri="https://zenodo.org/record/6630923/files/figure.png" pageId="79" pageNumber="80">16e</figureCitation>
, 
<figureCitation id="131C2A1A77607F4C8051A522FE0EFCDC" box="[361,401,798,824]" captionStart="FIGURE 20" captionStartId="114.[151,250,1910,1934]" captionTargetBox="[172,1414,182,1881]" captionTargetId="figure-25@114.[170,1417,181,1886]" captionTargetPageId="114" captionText="FIGURE 20. Male genital sternite of: a) Belonuchus cifuentesi, b) B. dichrous, c) B. flavipennis, d) B. aenigmaticus, e) B. iridescens, f) B. nigerrimus, g) B. optatus, h) B. platypterus, i) B. juanmorronei, j) B. abnormalis, k) B. pectinipes, l) B. viridipennis, m) Belonuchus sp. Q, n) B. cyanipennis, o) B. godmani. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6630929" httpUri="https://zenodo.org/record/6630929/files/figure.png" pageId="79" pageNumber="80">20f</figureCitation>
, 
<figureCitation id="131C2A1A77607F4C80A5A522FE5DFCDC" box="[413,450,798,824]" captionStart="FIGURE 24" captionStartId="118.[151,250,1786,1810]" captionTargetBox="[164,1436,193,1751]" captionTargetId="figure-25@118.[151,1436,181,1762]" captionTargetPageId="118" captionText="FIGURE 24. Ventral (left) and lateral (right) view (except e) of the aedeagus of: a) Belonuchus rufiventris, b) B. xanthomelas (at center, detail of the apex of the median lobe), c) B. ephippiatus, d) B. moroni, e) B. cifuentesi (left: detail of paramere, right: ventral view; from Rodríguez and Navarrete-Heredia, 2016), f) B. dichrous, g) B. flavipennis, h) B. aenigmaticus, i) B. iridescens, j) B. nigerrimus, k) B. optatus, l) B. platypterus. Scale bar = 0.5 mm (all figures, except e)." figureDoi="http://doi.org/10.5281/zenodo.6630941" httpUri="https://zenodo.org/record/6630941/files/figure.png" pageId="79" pageNumber="80">24j</figureCitation>
, 
<figureCitation id="131C2A1A77607F4C80F6A522FE75FCDC" box="[462,490,798,824]" captionStart="FIGURE 34" captionStartId="128.[151,250,992,1016]" captionTargetBox="[158,1428,188,960]" captionTargetId="figure-25@128.[151,1436,181,968]" captionTargetPageId="128" captionText="FIGURE 34. Geographical distribution in Mexico of Belonuchus nigerrimus." figureDoi="http://doi.org/10.5281/zenodo.6630961" httpUri="https://zenodo.org/record/6630961/files/figure.png" pageId="79" pageNumber="80">34</figureCitation>
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<paragraph id="8B98369F77607F4C81AFA55FFDF3FC9E" blockId="79.[151,620,866,891]" box="[151,620,866,891]" pageId="79" pageNumber="80">
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<treatmentCitation id="0A86108E77607F4C81AFA55FFDF8FC9E" author="Bernhauer, M." box="[151,615,866,891]" page="225" pageId="79" pageNumber="80" year="1918">
<taxonomicName id="4C274D1C77607F4C81AFA55FFDF8FC9E" ID-CoL="LC9T" authority="Bernhauer, 1918: 225" authorityName="Bernhauer" authorityPageNumber="225" authorityYear="1918" box="[151,615,866,891]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="nigerrimus">
<emphasis id="B953EA8D77607F4C81AFA55FFE1CFC9E" box="[151,387,867,891]" italics="true" pageId="79" pageNumber="80">Belonuchus nigerrimus</emphasis>
<bibRefCitation id="EFB64B6E77607F4C80B1A55FFDF8FC9E" author="Bernhauer, M." box="[393,615,866,891]" pageId="79" pageNumber="80" pagination="225 - 226" refId="ref111961" refString="Bernhauer, M. (1918) Vier neue Belonuchus aus Mexiko [cont.]. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 67, 225 - 226." type="journal article" year="1918">Bernhauer, 1918: 225</bibRefCitation>
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.
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<paragraph id="8B98369F77607F4C81AFA595FA08FC27" blockId="79.[151,1437,937,2044]" box="[151,1431,937,963]" pageId="79" pageNumber="80">
Total body length 
<quantity id="4CDF9B7A77607F4C805BA595FE55FC27" box="[355,458,937,963]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="79" pageNumber="80" unit="mm" value="13.3">13.3 mm</quantity>
. Coloration dark-brown, almost black, with elytra slightly darker than remaining body.
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<paragraph id="8B98369F77607F4C81FFA5F1FAC3FB07" blockId="79.[151,1437,937,2044]" pageId="79" pageNumber="80">
<emphasis id="B953EA8D77607F4C81FFA5F1FE8FFC03" bold="true" box="[199,272,973,999]" pageId="79" pageNumber="80">Head:</emphasis>
transverse, ratio length/width 0.76. Dorsal surface with sulcate longitudinal midline less visible behind front and more visible at front, latter with a foveate area that extends laterad (
<figureCitation id="131C2A1A77607F4C82D6A5CDFBD4FBE8" box="[1006,1099,1009,1036]" captionStart="FIGURE 10" captionStartId="104.[151,250,1839,1863]" captionTargetBox="[176,1418,194,1806]" captionTargetId="figure-25@104.[152,1435,181,1816]" captionTargetPageId="104" captionText="FIGURE 10. Dorsal view of the head of males of: a) Belonuchus julietitae, b) B. singularis, c) B. connexus, d) B. platypterus, e) B. bidens, f) B. nigerrimus, g) B. optatus, h) B. cyanipennis, i) B. godmani. Dorsal view of the head and pronotum of the holotype female of: j) B. optatus." figureDoi="http://doi.org/10.5281/zenodo.6630909" httpUri="https://zenodo.org/record/6630909/files/figure.png" pageId="79" pageNumber="80">Fig. 10f</figureCitation>
). Eyes nearly 0.5 times the cephalic lateral length, weakly protruding laterally. First antennomere almost as long as antennomeres 2–3 combined, 2 slightly shorter than 3, 4–5 slightly longer than wide, 6–10 almost as long as wide. Mandibles 1.4 times longer than head; each mandible with basal tooth and middle tooth widely separated from each other; mandibular channel well developed, external margin visible and internal margin strongly carinate at base, reaching level of middle tooth, then becoming less visible like impressed line. Apical palpomere of maxillary and labial palpi almost twice as long as preapical palpomere. Head 1.13 times wider than pronotum. Neck with wrinkled microsculpture ventrally.
</paragraph>
<paragraph id="8B98369F77607F4C81FFA2D1FAF6F9C3" blockId="79.[151,1437,937,2044]" pageId="79" pageNumber="80">
<emphasis id="B953EA8D77607F4C81FFA2D1FEBFFAE3" bold="true" box="[199,288,1261,1287]" pageId="79" pageNumber="80">Thorax</emphasis>
: dorsal rows of pronotum with six punctures, with posterior punctures more separated from previous punctures; ratio length/width 1.02; almost as wide at anterior corners as at posterior corners (ratio 1.05). Elytra and scutellum with moderately dense fine setae. Prosternum with elevated transverse area near anterior margin, delimited backwards by a clearly visible transverse line. Intercoxal process of mesoventrite shield-shaped, moderately acute and weakly projecting posteriad; transverse discal ridge broadly U-shaped, not meeting margin of intercoxal process laterally. Profemur with external row of spines initiating near base, extending near level of the few apical spines of internal margin, then interrupted as if two or three spines were lacking, ending in two apical spines; spines black, thick and moderately large. Metacoxa with coarse spines moderately dense; metatrochanter with spines as on coxa, but denser; metafemur without row of spines, but with some coarse setae. Tarsomeres flattened dorsally.
</paragraph>
<paragraph id="8B98369F77607F4C81FFA00DFDC1F8A3" blockId="79.[151,1437,937,2044]" pageId="79" pageNumber="80">
<emphasis id="B953EA8D77607F4C81FFA00DFEA3F9AF" bold="true" box="[199,316,1585,1611]" pageId="79" pageNumber="80">Abdomen</emphasis>
: first three visible tergites with posterior basal transverse carina well developed; adjacent depressed area narrow but visible, with sparse wide-superficial punctures combined with fine punctures, tergite 4 also with some wide-superficial punctures at base, remaining surface of each tergite covered with moderately dense, fine punctures. Sternites with punctures similar to those of tergites, but at anterior corners with mesh-like microsculpture looking rough in comparison with that of remaining of their surface, where it is smoother. Male pregenital sternite moderately emarginate at posterior margin (
<figureCitation id="131C2A1A77607F4C8345A0D9FD42F8E4" box="[637,733,1765,1792]" captionStart="FIGURE 16" captionStartId="110.[151,250,1818,1842]" captionTargetBox="[183,1402,184,1781]" captionTargetId="figure-25@110.[170,1417,181,1794]" captionTargetPageId="110" captionText="FIGURE 16. Male pregenital sternite of: a) Belonuchus moroni, b) B. dichrous, c) B. flavipennis, d) B. aenigmaticus, e) B. nigerrimus, f) B. juanmorronei, g) B. godmani, h) B. magnistylus, i) B. trochanterinus, j) B. rufipennis, k) B. basiventris, l) B. connexus, m) B. xanthomelas, n) B. ephippiatus, o) B. optatus." figureDoi="http://doi.org/10.5281/zenodo.6630923" httpUri="https://zenodo.org/record/6630923/files/figure.png" pageId="79" pageNumber="80">Fig. 16e</figureCitation>
). Male genital sternite short (2.14 times longer than wide), very asymmetrical, anterior portion occupying 30% and posterior portion 70% of its length, apical emargination very deep (
<figureCitation id="131C2A1A77607F4C81E3A111FEA5F8AC" box="[219,314,1837,1864]" captionStart="FIGURE 20" captionStartId="114.[151,250,1910,1934]" captionTargetBox="[172,1414,182,1881]" captionTargetId="figure-25@114.[170,1417,181,1886]" captionTargetPageId="114" captionText="FIGURE 20. Male genital sternite of: a) Belonuchus cifuentesi, b) B. dichrous, c) B. flavipennis, d) B. aenigmaticus, e) B. iridescens, f) B. nigerrimus, g) B. optatus, h) B. platypterus, i) B. juanmorronei, j) B. abnormalis, k) B. pectinipes, l) B. viridipennis, m) Belonuchus sp. Q, n) B. cyanipennis, o) B. godmani. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6630929" httpUri="https://zenodo.org/record/6630929/files/figure.png" pageId="79" pageNumber="80">Fig. 20f</figureCitation>
). Abdominal styli slender.
</paragraph>
<paragraph id="8B98369F77607F4C81FFA16DFCA1F874" blockId="79.[151,1437,937,2044]" pageId="79" pageNumber="80">
<emphasis id="B953EA8D77607F4C81FFA16DFEA5F88F" bold="true" box="[199,314,1873,1899]" pageId="79" pageNumber="80">Aedeagus</emphasis>
: length 
<quantity id="4CDF9B7A77607F4C80A1A16DFE6CF88F" box="[409,499,1873,1900]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="79" pageNumber="80" unit="mm" value="1.5">1.5 mm</quantity>
; elongate in apical half and ovoid in basal half; apex somewhat convex; basal half notably wider than apical half; internal sac visible (
<figureCitation id="131C2A1A77607F4C83EEA149FCADF874" box="[726,818,1909,1936]" captionStart="FIGURE 24" captionStartId="118.[151,250,1786,1810]" captionTargetBox="[164,1436,193,1751]" captionTargetId="figure-25@118.[151,1436,181,1762]" captionTargetPageId="118" captionText="FIGURE 24. Ventral (left) and lateral (right) view (except e) of the aedeagus of: a) Belonuchus rufiventris, b) B. xanthomelas (at center, detail of the apex of the median lobe), c) B. ephippiatus, d) B. moroni, e) B. cifuentesi (left: detail of paramere, right: ventral view; from Rodríguez and Navarrete-Heredia, 2016), f) B. dichrous, g) B. flavipennis, h) B. aenigmaticus, i) B. iridescens, j) B. nigerrimus, k) B. optatus, l) B. platypterus. Scale bar = 0.5 mm (all figures, except e)." figureDoi="http://doi.org/10.5281/zenodo.6630941" httpUri="https://zenodo.org/record/6630941/files/figure.png" pageId="79" pageNumber="80">Fig. 24j</figureCitation>
).
</paragraph>
<paragraph id="8B98369F77607F5381FFA1A5FEC5FEA5" blockId="79.[151,1437,937,2044]" lastBlockId="80.[151,1437,150,1581]" lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80">
<emphasis id="B953EA8D77607F4C81FFA1A5FED8F857" bold="true" box="[199,327,1945,1971]" pageId="79" pageNumber="80">Variability</emphasis>
. There is the following variation in ratios and measurements: ratio length/width of head in males 0.79 (range 0.71–0.85), in females 0.825 (range 0.82–0.84); ratio mandibular length/cephalic length in males 1.13 (range 0.88–1.35), in females 0.98 (range 0.94–1.02); ratio cephalic width/pronotal width in males 1.15 (range 1.07–1.27), in females 1.11 (range 1.09–1.13); ratio length/width of pronotum in males 1.10 (range 1.03–1.15), in females 1.12 (range 1.05–1.16); ratio anterior width/posterior width of pronotum in males 1.09 (range 1.06–1.14), in females 1.07 (range 1.05–1.09). The 
<typeStatus id="549C883D777F7F538358A6E3FD68FF1D" box="[608,759,223,249]" pageId="80" pageNumber="81" type="paralectotype">paralectotype</typeStatus>
is of paler color than the 
<typeStatus id="549C883D777F7F53851EA6E3FB0FFF1D" box="[1062,1168,223,249]" pageId="80" pageNumber="81" type="lectotype">lectotype</typeStatus>
, dark-brown, pronotum with dorsal rows with five (left) and six (right) punctures; external row of spines in profemur with spines smaller than in 
<typeStatus id="549C883D777F7F5381D3A71BFECAFEA5" box="[235,341,295,321]" pageId="80" pageNumber="81" type="lectotype">lectotype</typeStatus>
.
</paragraph>
<paragraph id="8B98369F777F7F5381FFA776FC2DFDD9" blockId="80.[151,1437,150,1581]" pageId="80" pageNumber="81">
<emphasis id="B953EA8D777F7F5381FFA776FE56FE81" bold="true" box="[199,457,330,357]" pageId="80" pageNumber="81">Taxonomic comments</emphasis>
. Of the group 
<taxonomicName id="4C274D1C777F7F538351A777FD78FE81" baseAuthorityName="Sharp" baseAuthorityYear="1885" box="[617,743,331,357]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="platypterus">
<emphasis id="B953EA8D777F7F538351A777FD78FE81" box="[617,743,331,357]" italics="true" pageId="80" pageNumber="81">platypterus</emphasis>
</taxonomicName>
only 
<taxonomicName id="4C274D1C777F7F53821DA777FC22FE80" authorityName="Bernhauer" authorityYear="1918" box="[805,957,331,356]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="nigerrimus">
<emphasis id="B953EA8D777F7F53821DA777FC22FE80" box="[805,957,331,356]" italics="true" pageId="80" pageNumber="81">B. nigerrimus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C274D1C777F7F5382CAA777FB37FE80" authorityName="Márquez &amp; Asiain" authorityYear="2022" box="[1010,1192,331,356]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="juanmorronei">
<emphasis id="B953EA8D777F7F5382CAA777FB37FE80" box="[1010,1192,331,356]" italics="true" pageId="80" pageNumber="81">B. juanmorronei</emphasis>
</taxonomicName>
are completely black, but 
<taxonomicName id="4C274D1C777F7F5381FAA753FEC2FE6C" authorityName="Bernhauer" authorityYear="1918" box="[194,349,367,392]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="nigerrimus">
<emphasis id="B953EA8D777F7F5381FAA753FEC2FE6C" box="[194,349,367,392]" italics="true" pageId="80" pageNumber="81">B. nigerrimus</emphasis>
</taxonomicName>
has a transverse head, mandibles as long or longer than head, eyes almost 0.5 times the cephalic lateral length, and a different genital sternite and aedeagus than in 
<taxonomicName id="4C274D1C777F7F53824DA7AFFBB3FE48" authorityName="Márquez &amp; Asiain" authorityYear="2022" box="[885,1068,403,428]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="juanmorronei">
<emphasis id="B953EA8D777F7F53824DA7AFFBB3FE48" box="[885,1068,403,428]" italics="true" pageId="80" pageNumber="81">B. juanmorronei</emphasis>
</taxonomicName>
, which also has an almost square head, slightly widened backwards, mandibles shorter than head and eyes less than 0.5 times the cephalic lateral length. Other black species, such as 
<taxonomicName id="4C274D1C777F7F53831FA7E7FD0AFE10" authorityName="Marquez, Asiain &amp; Navarrete-Heredia" authorityYear="2018" box="[551,661,475,500]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="moroni">
<emphasis id="B953EA8D777F7F53831FA7E7FD0AFE10" box="[551,661,475,500]" italics="true" pageId="80" pageNumber="81">B. moroni</emphasis>
</taxonomicName>
, belong to different groups, making it easier to distinguish them using group characters. 
<typeStatus id="549C883D777F7F538059A7C3FE4EFDFD" box="[353,465,511,537]" pageId="80" pageNumber="81" type="lectotype">Lectotype</typeStatus>
of this species is designated here taking into account Article 74.7 of the ICZN and its recommendations, to establish more clearly the identity of this species.
</paragraph>
</subSubSection>
<subSubSection id="C33D6514777F7F5381FFA47BFF66F9C9" pageId="80" pageNumber="81" type="materials_examined">
<paragraph id="8B98369F777F7F5381FFA47BFF66F9C9" blockId="80.[151,1437,150,1581]" pageId="80" pageNumber="81">
<emphasis id="B953EA8D777F7F5381FFA47BFD17FD84" bold="true" box="[199,648,582,609]" pageId="80" pageNumber="81">
Type material examined (
<specimenCount id="9D21FD16777F7F5380C3A47AFDE4FD84" box="[507,635,582,608]" pageId="80" pageNumber="81" type="male">two males</specimenCount>
):
</emphasis>
<materialsCitation id="3B4F3CC2777F7F5383A8A47BFDA9FD4D" collectionCode="FMNH" collectorName="M. Bernhauer Collection" country="Mexico" county="Jalapa" determinerName="Det. Bernh" location="Chicago" municipality="Typus" pageId="80" pageNumber="81" specimenCount="2" stateProvince="Mexico" typeStatus="lectotype">
<emphasis id="B953EA8D777F7F5383A8A47BFC99FD85" bold="true" box="[656,774,583,609]" pageId="80" pageNumber="81">
<typeStatus id="549C883D777F7F5383A8A47BFC99FD85" box="[656,774,583,609]" pageId="80" pageNumber="81" type="lectotype">Lectotype</typeStatus>
</emphasis>
(male, 
<collectionCode id="ED36AE5A777F7F538266A47BFC2AFD85" box="[862,949,583,609]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="80" pageNumber="81" type="Museum">FMNH</collectionCode>
): “ 
<collectingCounty id="62F94E13777F7F5382EAA47BFB82FD85" box="[978,1053,583,609]" pageId="80" pageNumber="81">Jalapa</collectingCounty>
, 
<collectingRegion id="49E3F87D777F7F538512A47BFBE0FD85" box="[1066,1151,583,609]" country="Mexico" name="Mexico" pageId="80" pageNumber="81">Mexico</collectingRegion>
/ 
<emphasis id="B953EA8D777F7F5385A8A47BFB78FD84" box="[1168,1255,583,608]" italics="true" pageId="80" pageNumber="81">gagates</emphasis>
Er. 
<determinerName id="7E44DA5B777F7F538420A47BFA0BFD85" box="[1304,1428,583,609]" pageId="80" pageNumber="81">Det. Bernh</determinerName>
/ 
<taxonomicName id="4C274D1C777F7F5381AFA457FEFBFD61" authority="Bernh." authorityName="Bernhauer" authorityYear="1918" box="[151,356,619,645]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="nigerrimus">
<emphasis id="B953EA8D777F7F5381AFA457FE8DFD60" box="[151,274,619,644]" italics="true" pageId="80" pageNumber="81">nigerrimus</emphasis>
Bernh.
</taxonomicName>
<collectingMunicipality id="6BFCACE5777F7F538051A457FE33FD61" box="[361,428,619,645]" pageId="80" pageNumber="81">Typus</collectingMunicipality>
/ 
<location id="8EF86044777F7F538083A457FD87FD61" LSID="urn:lsid:plazi:treatment:038E878977607F5381AFA4C6FF66F9C9:8EF86044777F7F538083A457FD87FD61" box="[443,536,619,645]" country="Mexico" county="Jalapa" municipality="Typus" name="Chicago" pageId="80" pageNumber="81" stateProvince="Mexico">Chicago</location>
NHMus 
<collectorName id="26D25349777F7F538347A457FC0CFD61" box="[639,915,619,645]" pageId="80" pageNumber="81">M. Bernhauer Collection</collectorName>
/ 
<typeStatus id="549C883D777F7F53829AA457FB8DFD61" box="[930,1042,619,645]" pageId="80" pageNumber="81" type="lectotype">Lectotype</typeStatus>
<taxonomicName id="4C274D1C777F7F538520A457FF4DFD4D" authority="Bernhauer, 1918" authorityName="Bernhauer" authorityYear="1918" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="nigerrimus">
<emphasis id="B953EA8D777F7F538520A457FA83FD60" box="[1048,1308,619,645]" italics="true" pageId="80" pageNumber="81">Belonuchus nigerrimus</emphasis>
<bibRefCitation id="EFB64B6E777F7F53841AA457FF4DFD4D" author="Bernhauer, M." pageId="80" pageNumber="81" pagination="225 - 226" refId="ref111961" refString="Bernhauer, M. (1918) Vier neue Belonuchus aus Mexiko [cont.]. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 67, 225 - 226." type="journal article" year="1918">Bernhauer, 1918</bibRefCitation>
</taxonomicName>
, Márquez &amp; Asiain des. 2022”
</materialsCitation>
. 
<materialsCitation id="3B4F3CC2777F7F53837AA4B3FC9BFD29" collectionCode="FMNH" country="Mexico" location="Mexiko" pageId="80" pageNumber="81" specimenCount="1" stateProvince="Mexico" typeStatus="paralectotype">
<emphasis id="B953EA8D777F7F53837AA4B3FD78FD4D" bold="true" box="[578,743,655,681]" pageId="80" pageNumber="81">
<typeStatus id="549C883D777F7F53837AA4B3FD78FD4D" box="[578,743,655,681]" pageId="80" pageNumber="81" type="paralectotype">Paralectotype</typeStatus>
</emphasis>
: “Jalapa, 
<collectingRegion id="49E3F87D777F7F538261A4B3FC2BFD4D" box="[857,948,655,681]" country="Mexico" name="Mexico" pageId="80" pageNumber="81">Mexiko</collectingRegion>
. Dabberl/ 
<taxonomicName id="4C274D1C777F7F538513A4B3FAC5FD4D" authority=", Brnh. Cotypus" authorityName="Brnh. Cotypus" box="[1067,1370,655,681]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="nigerrimus">
<emphasis id="B953EA8D777F7F538513A4B3FB39FD4C" box="[1067,1190,655,680]" italics="true" pageId="80" pageNumber="81">nigerrimus</emphasis>
, Brnh. Cotypus
</taxonomicName>
/ Chicago NHMus M. Bernhauer Collection” (m 1, 
<collectionCode id="ED36AE5A777F7F53839EA48FFD63FD29" box="[678,764,691,717]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="80" pageNumber="81" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B4F3CC2777F7F538237A48EFE18FC65" collectingDate="2013-05-05" collectingDateMax="2017-07-05" collectingDateMin="2013-05-05" collectionCode="CC-UAEH" collectorName="J. Marquez &amp; A. Lora &amp; N. Gutierrez" country="Mexico" elevation="1647" latitude="21.141832" location="Santa Monica" longLatPrecision="86" longitude="-98.23333" municipality="Tianguistengo" pageId="80" pageNumber="81" specimenCount="12" specimenCount-female="1" specimenCount-male="11" stateProvince="Hidalgo">
<emphasis id="B953EA8D777F7F538237A48EFEF8FD14" bold="true" pageId="80" pageNumber="81">
Additional material examined (
<specimenCount id="9D21FD16777F7F5385BEA48EFB70FD29" box="[1158,1263,690,717]" pageId="80" pageNumber="81" type="male">11 males</specimenCount>
, 
<specimenCount id="9D21FD16777F7F5385C4A48EFA10FD28" box="[1276,1423,690,716]" pageId="80" pageNumber="81" type="female">five females</specimenCount>
): 
<collectingCountry id="F330760F777F7F5381AFA4EAFF6DFD14" box="[151,242,726,752]" name="Mexico" pageId="80" pageNumber="81">Mexico</collectingCountry>
: 
<collectingRegion id="49E3F87D777F7F5381C7A4EAFEFDFD14" box="[255,354,726,752]" country="Mexico" name="Hidalgo" pageId="80" pageNumber="81">Hidalgo</collectingRegion>
:
</emphasis>
“La Misión, camino al río Moctezuma, 
<geoCoordinate id="EE135058777F7F538214A4EBFC20FD15" box="[812,959,727,753]" degrees="21" direction="north" minutes="8" orientation="latitude" pageId="80" pageNumber="81" precision="1" seconds="30.6" value="21.141832">N21°8’30.6”</geoCoordinate>
, 
<geoCoordinate id="EE135058777F7F5382F2A4EAFBFBFD15" box="[970,1124,726,753]" degrees="99" direction="west" minutes="5" orientation="longitude" pageId="80" pageNumber="81" precision="1" seconds="49.1" value="-99.09698">W99°5’49.1”</geoCoordinate>
, bosque tropical subcaducifolio, en NTP-80 (calamar), 
<date id="FF99105F777F7F5380E0A4C7FD26FCF1" box="[472,697,762,789]" pageId="80" pageNumber="81" value="2017-06-23" valueMax="2017-07-05" valueMin="2017-06-23">
<collectingDate id="EFDDE9B7777F7F5380E0A4C7FD26FCF1" box="[472,697,762,789]" pageId="80" pageNumber="81" value="2017-06-23" valueMax="2017-07-05" valueMin="2017-06-23">23-VI a 5-VII-2017</collectingDate>
</date>
, 
<collectorName id="26D25349777F7F5383FDA4C7FCDCFCF1" box="[709,835,763,789]" pageId="80" pageNumber="81">J. Márquez</collectorName>
y 
<collectorName id="26D25349777F7F538266A4C7FC2AFCF1" box="[862,949,763,789]" pageId="80" pageNumber="81">A. Lora</collectorName>
cols.” (1, 
<collectionCode id="ED36AE5A777F7F538513A4C7FB32FCF1" box="[1067,1197,763,789]" pageId="80" pageNumber="81">CC-UAEH</collectionCode>
). “Tenango de Doria, camino a La Viejita, bosque mesófilo de montaña, 
<quantity id="4CDF9B7A777F7F5383E9A523FCB6FCDD" box="[721,809,799,825]" metricMagnitude="3" metricUnit="m" metricValue="1.6469999999999998" pageId="80" pageNumber="81" unit="m" value="1647.0">
<elevation id="000AD1AC777F7F5383E9A523FCB6FCDD" box="[721,809,799,825]" metricMagnitude="3" metricUnit="m" metricValue="1.6469999999999998" pageId="80" pageNumber="81" unit="m" value="1647.0">1647 m</elevation>
</quantity>
, 
<geoCoordinate id="EE135058777F7F53820DA523FC37FCDD" box="[821,936,799,825]" degrees="20" direction="north" minutes="19.0" orientation="latitude" pageId="80" pageNumber="81" precision="92" value="20.316668">N20°19.0’</geoCoordinate>
37.9”, 
<geoCoordinate id="EE135058777F7F5382C3A523FBE8FCDD" box="[1019,1143,799,825]" degrees="98" direction="west" minutes="14.0" orientation="longitude" pageId="80" pageNumber="81" precision="92" value="-98.23333">W98°14.0’</geoCoordinate>
39.7”, trampa de intercepción de vuelo #3, 
<date id="FF99105F777F7F538063A57EFE62FCB9" box="[347,509,834,861]" pageId="80" pageNumber="81" value="2013-05-05" valueMax="2013-05-10" valueMin="2013-05-05">
<collectingDate id="EFDDE9B7777F7F538063A57EFE62FCB9" box="[347,509,834,861]" pageId="80" pageNumber="81" value="2013-05-05" valueMax="2013-05-10" valueMin="2013-05-05">5 a 10-V-2013</collectingDate>
</date>
, 
<collectorName id="26D25349777F7F53833EA57FFD0DFCB9" box="[518,658,835,861]" pageId="80" pageNumber="81">N. Gutiérrez</collectorName>
col.” (1, 
<collectionCode id="ED36AE5A777F7F5383CFA57FFCE6FCB9" box="[759,889,835,861]" pageId="80" pageNumber="81">CC-UAEH</collectionCode>
). “ 
<collectingMunicipality id="6BFCACE5777F7F5382ABA57FFBAAFCB9" box="[915,1077,835,861]" pageId="80" pageNumber="81">Tianguistengo</collectingMunicipality>
camino a 
<location id="8EF86044777F7F53859EA57FFADDFCB9" LSID="urn:lsid:plazi:treatment:038E878977607F5381AFA4C6FF66F9C9:8EF86044777F7F53859EA57FFADDFCB9" box="[1190,1346,835,861]" country="Mexico" latitude="21.141832" longLatPrecision="86" longitude="-98.23333" municipality="Tianguistengo" name="Santa Monica" pageId="80" pageNumber="81" stateProvince="Hidalgo">Santa Mónica</location>
, bosque mesófilo de montaña
</materialsCitation>
, 
<materialsCitation id="3B4F3CC2777F7F5380ACA55BFF65FC2D" collectingDate="2009-04-13" collectionCode="CC-UAEH" collectorName="J. Sanchez &amp; M. Rivero &amp; Vargas &amp; Sanchez" country="Mexico" elevation="1868" latitude="20.718166" location="La Cabana" longLatPrecision="1" longitude="-98.661194" municipality="Tlanchinol" pageId="80" pageNumber="81" specimenCount="1" stateProvince="Hidalgo">
<quantity id="4CDF9B7A777F7F5380ACA55BFE73FC65" box="[404,492,871,897]" metricMagnitude="3" metricUnit="m" metricValue="1.8679999999999999" pageId="80" pageNumber="81" unit="m" value="1868.0">
<elevation id="000AD1AC777F7F5380ACA55BFE73FC65" box="[404,492,871,897]" metricMagnitude="3" metricUnit="m" metricValue="1.8679999999999999" pageId="80" pageNumber="81" unit="m" value="1868.0">1868 m</elevation>
</quantity>
, 
<geoCoordinate id="EE135058777F7F5380C1A55AFD14FC65" box="[505,651,870,897]" degrees="20" direction="north" minutes="43" orientation="latitude" pageId="80" pageNumber="81" precision="1" seconds="5.4" value="20.718166">N20°43’5.4”</geoCoordinate>
, 
<geoCoordinate id="EE135058777F7F5383A0A55BFCA1FC65" box="[664,830,871,897]" degrees="98" direction="west" minutes="39" orientation="longitude" pageId="80" pageNumber="81" precision="1" seconds="40.3" value="-98.661194">W98°39’40.3”</geoCoordinate>
, trampa de intercepción de vuelo, 7 al 
<date id="FF99105F777F7F5385C1A55BFAE2FC65" box="[1273,1405,871,897]" pageId="80" pageNumber="81" value="2009-04-13">
<collectingDate id="EFDDE9B7777F7F5385C1A55BFAE2FC65" box="[1273,1405,871,897]" pageId="80" pageNumber="81" value="2009-04-13">13-IV-2009</collectingDate>
</date>
, 
<collectorName id="26D25349777F7F5384B2A55BFF68FC41" pageId="80" pageNumber="81">J. Sánchez</collectorName>
, 
<collectorName id="26D25349777F7F53803CA5B7FEE4FC41" box="[260,379,907,933]" pageId="80" pageNumber="81">M. Rivero</collectorName>
, M. 
<collectorName id="26D25349777F7F538088A5B7FE63FC41" box="[432,508,907,933]" pageId="80" pageNumber="81">Vargas</collectorName>
y J. 
<collectorName id="26D25349777F7F53830DA5B7FD0DFC41" box="[565,658,907,933]" pageId="80" pageNumber="81">Sánchez</collectorName>
cols.” (1, 
<collectionCode id="ED36AE5A777F7F538233A5B7FC12FC41" box="[779,909,907,933]" pageId="80" pageNumber="81">CC-UAEH</collectionCode>
). “ 
<collectingMunicipality id="6BFCACE5777F7F538292A5B7FBB6FC41" box="[938,1065,907,933]" pageId="80" pageNumber="81">Tlanchinol</collectingMunicipality>
, 
<location id="8EF86044777F7F53850EA5B7FB2CFC41" LSID="urn:lsid:plazi:treatment:038E878977607F5381AFA4C6FF66F9C9:8EF86044777F7F53850EA5B7FB2CFC41" box="[1078,1203,907,933]" country="Mexico" latitude="20.718166" longLatPrecision="1" longitude="-98.661194" municipality="Tlanchinol" name="La Cabana" pageId="80" pageNumber="81" stateProvince="Hidalgo">La Cabaña</location>
, bosque mesófilo de montaña
</materialsCitation>
, 
<materialsCitation id="3B4F3CC2777F7F53803FA592FAB0FC09" collectingDate="2006-03-18" collectingDateMax="2006-03-25" collectingDateMin="2006-03-18" collectionCode="CC-UAEH" collectorName="C. Ortiz &amp; M. C. Pedraza" country="Mexico" county="Tlanchinol" elevation="1485" latitude="21.022617" location="El Pozo" longLatPrecision="1" longitude="-98.6426" municipality="Apantlazol" pageId="80" pageNumber="81" specimenCount="1" stateProvince="Hidalgo">
<quantity id="4CDF9B7A777F7F53803FA592FEC0FC2D" box="[263,351,942,969]" metricMagnitude="3" metricUnit="m" metricValue="1.4849999999999999" pageId="80" pageNumber="81" unit="m" value="1485.0">
<elevation id="000AD1AC777F7F53803FA592FEC0FC2D" box="[263,351,942,969]" metricMagnitude="3" metricUnit="m" metricValue="1.4849999999999999" pageId="80" pageNumber="81" unit="m" value="1485.0">1485 m</elevation>
</quantity>
, 
<geoCoordinate id="EE135058777F7F538053A592FE73FC2D" box="[363,492,942,969]" degrees="21" direction="north" minutes="1.357" orientation="latitude" pageId="80" pageNumber="81" precision="1" value="21.022617">N21°1.357’</geoCoordinate>
<geoCoordinate id="EE135058777F7F5380CCA592FD10FC2D" box="[500,655,942,969]" degrees="98" direction="west" minutes="38.556" orientation="longitude" pageId="80" pageNumber="81" precision="1" value="-98.6426">W98°38.556’</geoCoordinate>
, trampa de intercepción de vuelo #5, 
<date id="FF99105F777F7F538501A593FB69FC2D" box="[1081,1270,942,969]" pageId="80" pageNumber="81" value="2006-03-18" valueMax="2006-03-25" valueMin="2006-03-18">
<collectingDate id="EFDDE9B7777F7F538501A593FB69FC2D" box="[1081,1270,942,969]" pageId="80" pageNumber="81" value="2006-03-18" valueMax="2006-03-25" valueMin="2006-03-18">18 a 25-III-2006</collectingDate>
</date>
, 
<collectorName id="26D25349777F7F53843BA593FAC0FC2D" box="[1283,1375,943,969]" pageId="80" pageNumber="81">C. Ortiz</collectorName>
y 
<collectorName id="26D25349777F7F538444A593FE88FC09" pageId="80" pageNumber="81">M. C. Pedraza</collectorName>
” (1, 
<collectionCode id="ED36AE5A777F7F53806AA5EFFE4BFC09" box="[338,468,979,1005]" pageId="80" pageNumber="81">CC-UAEH</collectionCode>
). “ 
<collectingCounty id="62F94E13777F7F5380CBA5EFFDF0FC09" box="[499,623,979,1005]" pageId="80" pageNumber="81">Tlanchinol</collectingCounty>
camino a 
<collectingMunicipality id="6BFCACE5777F7F5383D2A5EFFCF7FC09" box="[746,872,979,1005]" pageId="80" pageNumber="81">Apantlazol</collectingMunicipality>
, 
<location id="8EF86044777F7F53824EA5EFFC4BFC09" LSID="urn:lsid:plazi:treatment:038E878977607F5381AFA4C6FF66F9C9:8EF86044777F7F53824EA5EFFC4BFC09" box="[886,980,979,1005]" country="Mexico" county="Tlanchinol" latitude="21.022617" longLatPrecision="1" longitude="-98.6426" municipality="Apantlazol" name="El Pozo" pageId="80" pageNumber="81" stateProvince="Hidalgo">El Pozo</location>
, bosque mesófilo de montaña
</materialsCitation>
, 
<materialsCitation id="3B4F3CC2777F7F538405A5EFFE1AFB45" collectingDate="2011-05-10" collectingDateMax="2011-10-22" collectingDateMin="2011-05-10" collectionCode="CC-UAEH" collectorName="J. Marquez" country="Mexico" county="Mismos" elevation="1391" latitude="20.985203" location="Same" longLatPrecision="1" longitude="-98.62774" municipality="Same" pageId="80" pageNumber="81" specimenCount="1" stateProvince="Hidalgo">
<quantity id="4CDF9B7A777F7F538405A5EFFA08FC09" box="[1341,1431,979,1005]" metricMagnitude="3" metricUnit="m" metricValue="1.391" pageId="80" pageNumber="81" unit="m" value="1391.0">
<elevation id="000AD1AC777F7F538405A5EFFA08FC09" box="[1341,1431,979,1005]" metricMagnitude="3" metricUnit="m" metricValue="1.391" pageId="80" pageNumber="81" unit="m" value="1391.0">1391 m</elevation>
</quantity>
, 
<geoCoordinate id="EE135058777F7F5381AFA5CAFEA8FBF5" box="[151,311,1014,1041]" degrees="20" direction="north" minutes="59" orientation="latitude" pageId="80" pageNumber="81" precision="1" seconds="6.73" value="20.985203">N20°59’6.73”</geoCoordinate>
, 
<geoCoordinate id="EE135058777F7F53807BA5CBFE67FBF5" box="[323,504,1015,1041]" degrees="98" direction="west" minutes="37" orientation="longitude" pageId="80" pageNumber="81" precision="1" seconds="39.86" value="-98.62774">W98°37’39.86”</geoCoordinate>
, NTP-80 (calamar) #2, 22-IV al 
<date id="FF99105F777F7F53825FA5CBFC7DFBF5" box="[871,994,1015,1041]" pageId="80" pageNumber="81" value="2011-05-10">
<collectingDate id="EFDDE9B7777F7F53825FA5CBFC7DFBF5" box="[871,994,1015,1041]" pageId="80" pageNumber="81" value="2011-05-10">10-V-2011</collectingDate>
</date>
, 
<collectorName id="26D25349777F7F5382D5A5CBFBF5FBF5" box="[1005,1130,1015,1041]" pageId="80" pageNumber="81">J. Márquez</collectorName>
col.” (1, 
<collectionCode id="ED36AE5A777F7F5385EBA5CBFACAFBF5" box="[1235,1365,1015,1041]" pageId="80" pageNumber="81">CC-UAEH</collectionCode>
). 
<collectingCounty id="62F94E13777F7F53845FA5CBFF59FBD1" pageId="80" pageNumber="81">Mismos</collectingCounty>
datos excepto: “NTP-80 (calamar) #5” (2, 
<collectionCode id="ED36AE5A777F7F53838AA227FCABFBD1" box="[690,820,1051,1077]" pageId="80" pageNumber="81">CC-UAEH</collectionCode>
). 
<collectingMunicipality id="6BFCACE5777F7F53827FA227FC1AFBD1" box="[839,901,1051,1077]" pageId="80" pageNumber="81">Same</collectingMunicipality>
data, except: “trampa de intercepción de vuelo #2” (1, 
<collectionCode id="ED36AE5A777F7F5381D6A203FEEFFBBD" box="[238,368,1087,1113]" pageId="80" pageNumber="81">CC-UAEH</collectionCode>
). 
<location id="8EF86044777F7F5380BAA203FE5FFBBD" LSID="urn:lsid:plazi:treatment:038E878977607F5381AFA4C6FF66F9C9:8EF86044777F7F5380BAA203FE5FFBBD" box="[386,448,1087,1113]" country="Mexico" county="Mismos" latitude="20.985203" longLatPrecision="1" longitude="-98.62774" municipality="Same" name="Same" pageId="80" pageNumber="81" stateProvince="Hidalgo">Same</location>
data, except: “trampa de intercepción de vuelo #3, 1-X al 
<date id="FF99105F777F7F538561A203FB4AFBBD" box="[1113,1237,1087,1113]" pageId="80" pageNumber="81" value="2011-10-22">
<collectingDate id="EFDDE9B7777F7F538561A203FB4AFBBD" box="[1113,1237,1087,1113]" pageId="80" pageNumber="81" value="2011-10-22">22-X-2011</collectingDate>
</date>
, 
<collectorName id="26D25349777F7F5385D9A203FAC0FBBD" box="[1249,1375,1087,1113]" pageId="80" pageNumber="81">J. Márquez</collectorName>
col.” (1, 
<collectionCode id="ED36AE5A777F7F538185A25FFEA0FB99" box="[189,319,1123,1149]" pageId="80" pageNumber="81">CC-UAEH</collectionCode>
). Same data, except: “22-IV al 
<date id="FF99105F777F7F5383A6A25FFC86FB99" box="[670,793,1123,1149]" pageId="80" pageNumber="81" value="2011-05-10">
<collectingDate id="EFDDE9B7777F7F5383A6A25FFC86FB99" box="[670,793,1123,1149]" pageId="80" pageNumber="81" value="2011-05-10">10-V-2011</collectingDate>
</date>
” (1, 
<collectionCode id="ED36AE5A777F7F538277A25FFC4EFB99" box="[847,977,1123,1149]" pageId="80" pageNumber="81">CC-UAEH</collectionCode>
). “ 
<location id="8EF86044777F7F5382D5A25FFBE8FB99" LSID="urn:lsid:plazi:treatment:038E878977607F5381AFA4C6FF66F9C9:8EF86044777F7F5382D5A25FFBE8FB99" box="[1005,1143,1123,1149]" country="Mexico" county="Mismos" latitude="20.985203" longLatPrecision="1" longitude="-98.62774" municipality="Same" name="Zacualtipan" pageId="80" pageNumber="81" stateProvince="Hidalgo">Zacualtipán</location>
, camino a 
<location id="8EF86044777F7F5385C8A25FFAD3FB99" LSID="urn:lsid:plazi:treatment:038E878977607F5381AFA4C6FF66F9C9:8EF86044777F7F5385C8A25FFAD3FB99" box="[1264,1356,1123,1149]" country="Mexico" county="Mismos" latitude="20.985203" longLatPrecision="1" longitude="-98.62774" municipality="Same" name="Tizapan" pageId="80" pageNumber="81" stateProvince="Hidalgo">Tizapán</location>
, boque mesófilo de montaña
</materialsCitation>
, 
<materialsCitation id="3B4F3CC2777F7F5380A9A2BBFD97FB21" collectingDate="2011-04-28" collectionCode="CC-UAEH" collectorName="J. Marquez" country="Mexico" elevation="1690" latitude="20.651266" longLatPrecision="1" longitude="-98.600914" pageId="80" pageNumber="81" specimenCount="1" stateProvince="Hidalgo">
<quantity id="4CDF9B7A777F7F5380A9A2BBFE76FB45" box="[401,489,1159,1185]" metricMagnitude="3" metricUnit="m" metricValue="1.69" pageId="80" pageNumber="81" unit="m" value="1690.0">
<elevation id="000AD1AC777F7F5380A9A2BBFE76FB45" box="[401,489,1159,1185]" metricMagnitude="3" metricUnit="m" metricValue="1.69" pageId="80" pageNumber="81" unit="m" value="1690.0">1690 m</elevation>
</quantity>
, 
<geoCoordinate id="EE135058777F7F5380CDA2BBFD16FB45" box="[501,649,1159,1185]" degrees="20" direction="north" minutes="39.076" orientation="latitude" pageId="80" pageNumber="81" precision="1" value="20.651266">N20°39.076’</geoCoordinate>
, 
<geoCoordinate id="EE135058777F7F5383ADA2BAFCB0FB45" box="[661,815,1158,1185]" degrees="98" direction="west" minutes="36.055" orientation="longitude" pageId="80" pageNumber="81" precision="1" value="-98.600914">W98°36.055’</geoCoordinate>
, trampa de intercepción de vuelo #2, 14 al 
<date id="FF99105F777F7F53842DA2BBFA07FB45" box="[1301,1432,1159,1185]" pageId="80" pageNumber="81" value="2011-04-28">
<collectingDate id="EFDDE9B7777F7F53842DA2BBFA07FB45" box="[1301,1432,1159,1185]" pageId="80" pageNumber="81" value="2011-04-28">28-IV-2011</collectingDate>
</date>
, 
<collectorName id="26D25349777F7F5381AFA297FE8BFB21" box="[151,276,1195,1221]" pageId="80" pageNumber="81">J. Márquez</collectorName>
col.” (1, 
<collectionCode id="ED36AE5A777F7F538043A297FE62FB21" box="[379,509,1195,1221]" pageId="80" pageNumber="81">CC-UAEH</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B4F3CC2777F7F538336A297FAC6FB21" box="[526,1369,1195,1221]" country="Mexico" location="Plan de Ayala" municipality="Cordoba" pageId="80" pageNumber="81" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
<emphasis id="B953EA8D777F7F538336A297FD1AFB21" bold="true" box="[526,645,1195,1221]" pageId="80" pageNumber="81">
<collectingRegion id="49E3F87D777F7F538336A297FD1FFB21" box="[526,640,1195,1221]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="80" pageNumber="81">Veracruz</collectingRegion>
:
</emphasis>
“ 
<collectingMunicipality id="6BFCACE5777F7F5383ADA297FD64FB21" box="[661,763,1195,1221]" pageId="80" pageNumber="81">Córdoba</collectingMunicipality>
, camino a 
<location id="8EF86044777F7F538249A297FB93FB21" LSID="urn:lsid:plazi:treatment:038E878977607F5381AFA4C6FF66F9C9:8EF86044777F7F538249A297FB93FB21" box="[881,1036,1195,1221]" country="Mexico" municipality="Cordoba" name="Plan de Ayala" pageId="80" pageNumber="81" stateProvince="Veracruz de Ignacio de la Llave">Plan de Ayala</location>
, bosque mesófilo de montaña
</materialsCitation>
, 
<materialsCitation id="3B4F3CC2777F7F53845CA296FCEBFAE9" collectingDate="1999-06" collectionCode="MAAS" collectorName="J. Asiain &amp; Q. Santiago &amp; Marquez" country="Mexico" elevation="1352" latitude="18.954224" location="Coxcontla" longLatPrecision="14" longitude="-96.97806" municipality="Huatusco" pageId="80" pageNumber="81" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
<quantity id="4CDF9B7A777F7F53845CA296FF30FB0D" metricMagnitude="3" metricUnit="m" metricValue="1.3519999999999999" pageId="80" pageNumber="81" unit="m" value="1352.0">
<elevation id="000AD1AC777F7F53845CA296FF30FB0D" metricMagnitude="3" metricUnit="m" metricValue="1.3519999999999999" pageId="80" pageNumber="81" unit="m" value="1352.0">1352 m</elevation>
</quantity>
, 
<geoCoordinate id="EE135058777F7F538184A2F2FEC2FB0D" box="[188,349,1230,1257]" degrees="18" direction="north" minutes="57" orientation="latitude" pageId="80" pageNumber="81" precision="1" seconds="15.2" value="18.954224">N18°57’15.2”</geoCoordinate>
, 
<geoCoordinate id="EE135058777F7F538050A2F2FE65FB0D" box="[360,506,1230,1257]" degrees="96" direction="west" minutes="58" orientation="longitude" pageId="80" pageNumber="81" precision="15" seconds="41" value="-96.97806">W96°58’41”</geoCoordinate>
, NTP-80 (calamar), V a 
<date id="FF99105F777F7F53822AA2F3FCECFB0D" box="[786,883,1231,1257]" pageId="80" pageNumber="81" value="1999-06">
<collectingDate id="EFDDE9B7777F7F53822AA2F3FCECFB0D" box="[786,883,1231,1257]" pageId="80" pageNumber="81" value="1999-06">VI-1999</collectingDate>
</date>
, 
<collectorName id="26D25349777F7F538247A2F3FC7CFB0D" box="[895,995,1231,1257]" pageId="80" pageNumber="81">J. Asiain</collectorName>
, 
<collectorName id="26D25349777F7F5382D7A2F3FBE8FB0D" box="[1007,1143,1231,1257]" pageId="80" pageNumber="81">Q. Santiago</collectorName>
, J. 
<collectorName id="26D25349777F7F5385A5A2F3FA9CFB0D" box="[1181,1283,1231,1257]" pageId="80" pageNumber="81">Márquez</collectorName>
” (1, 
<collectionCode id="ED36AE5A777F7F538403A2F3FA0EFB0D" box="[1339,1425,1231,1257]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="80" pageNumber="81" type="Herbarium">MAAS</collectionCode>
). “ 
<collectingMunicipality id="6BFCACE5777F7F538199A2CFFE8FFAE9" box="[161,272,1267,1293]" pageId="80" pageNumber="81">Huatusco</collectingMunicipality>
, 
<location id="8EF86044777F7F538027A2CFFE0AFAE9" LSID="urn:lsid:plazi:treatment:038E878977607F5381AFA4C6FF66F9C9:8EF86044777F7F538027A2CFFE0AFAE9" box="[287,405,1267,1293]" country="Mexico" latitude="18.954224" longLatPrecision="14" longitude="-96.97806" municipality="Huatusco" name="Coxcontla" pageId="80" pageNumber="81" stateProvince="Veracruz de Ignacio de la Llave">Coxcontla</location>
, bosque mesófilo de montaña perturbado
</materialsCitation>
, 
<materialsCitation id="3B4F3CC2777F7F5382BBA2CEFC17FA9D" collectingDate="1995-01-14" collectingDateMax="1995-08-26" collectingDateMin="1995-01-14" collectionCode="QJSJ" collectorName="M. A. Pensado &amp; Q. Santiago" country="Mexico" county="Same" elevation="1250" location="Same" municipality="Same" pageId="80" pageNumber="81" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
<quantity id="4CDF9B7A777F7F5382BBA2CEFC42FAE9" box="[899,989,1266,1293]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="80" pageNumber="81" unit="m" value="1250.0">
<elevation id="000AD1AC777F7F5382BBA2CEFC42FAE9" box="[899,989,1266,1293]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="80" pageNumber="81" unit="m" value="1250.0">1250 m</elevation>
</quantity>
, NTP-80 (calamar), 
<date id="FF99105F777F7F5385F5A2CFFA08FAE9" box="[1229,1431,1266,1293]" pageId="80" pageNumber="81" value="1995-01-14" valueMax="1995-02-11" valueMin="1995-01-14">
<collectingDate id="EFDDE9B7777F7F5385F5A2CFFA08FAE9" box="[1229,1431,1266,1293]" pageId="80" pageNumber="81" value="1995-01-14" valueMax="1995-02-11" valueMin="1995-01-14">14-I a 11-II-1995</collectingDate>
</date>
, 
<collectorName id="26D25349777F7F5381AFA32BFEA1FAD5" box="[151,318,1303,1329]" pageId="80" pageNumber="81">M. A. Pensado</collectorName>
y 
<collectorName id="26D25349777F7F538064A32BFE7DFAD5" box="[348,482,1303,1329]" pageId="80" pageNumber="81">Q. Santiago</collectorName>
cols.” (1, 
<collectionCode id="ED36AE5A777F7F538361A32BFD08FAD5" box="[601,663,1303,1329]" pageId="80" pageNumber="81">QJSJ</collectionCode>
). 
<collectingCounty id="62F94E13777F7F538393A32BFD76FAD5" box="[683,745,1303,1329]" pageId="80" pageNumber="81">Same</collectingCounty>
data, except: “ 
<date id="FF99105F777F7F5382AAA32BFB09FAD5" box="[914,1174,1302,1329]" pageId="80" pageNumber="81" value="1995-07-29" valueMax="1995-08-26" valueMin="1995-07-29">
<collectingDate id="EFDDE9B7777F7F5382AAA32BFB09FAD5" box="[914,1174,1302,1329]" pageId="80" pageNumber="81" value="1995-07-29" valueMax="1995-08-26" valueMin="1995-07-29">29-VII a 26-VIII-1995</collectingDate>
</date>
” (1, 
<collectionCode id="ED36AE5A777F7F5385F6A32BFA93FAD5" box="[1230,1292,1303,1329]" pageId="80" pageNumber="81">QJSJ</collectionCode>
). 
<collectingMunicipality id="6BFCACE5777F7F538418A32BFAC1FAD5" box="[1312,1374,1303,1329]" pageId="80" pageNumber="81">Same</collectingMunicipality>
data, except: “ 
<date id="FF99105F777F7F5381C7A307FE27FAB1" box="[255,440,1338,1365]" pageId="80" pageNumber="81" value="1995-07-01" valueMax="1995-07-29" valueMin="1995-07-01">
<collectingDate id="EFDDE9B7777F7F5381C7A307FE27FAB1" box="[255,440,1338,1365]" pageId="80" pageNumber="81" value="1995-07-01" valueMax="1995-07-29" valueMin="1995-07-01">1 a 29-VII-1995</collectingDate>
</date>
, coprotrampa” (1, 
<collectionCode id="ED36AE5A777F7F5383B7A307FD52FAB1" box="[655,717,1339,1365]" pageId="80" pageNumber="81">QJSJ</collectionCode>
). 
<location id="8EF86044777F7F5383DAA307FCBFFAB1" LSID="urn:lsid:plazi:treatment:038E878977607F5381AFA4C6FF66F9C9:8EF86044777F7F5383DAA307FCBFFAB1" box="[738,800,1339,1365]" country="Mexico" county="Same" municipality="Same" name="Same" pageId="80" pageNumber="81" stateProvince="Veracruz de Ignacio de la Llave">Same</location>
data, except: “ 
<date id="FF99105F777F7F5382F3A307FB11FAB1" box="[971,1166,1338,1365]" pageId="80" pageNumber="81" value="1995-05-06" valueMax="1995-06-03" valueMin="1995-05-06">
<collectingDate id="EFDDE9B7777F7F5382F3A307FB11FAB1" box="[971,1166,1338,1365]" pageId="80" pageNumber="81" value="1995-05-06" valueMax="1995-06-03" valueMin="1995-05-06">6-V a 3-VI-1995</collectingDate>
</date>
, NTP-80 (calamar)” (1, 
<collectionCode id="ED36AE5A777F7F5381AFA363FF4AFA9D" box="[151,213,1375,1401]" pageId="80" pageNumber="81">QJSJ</collectionCode>
). “ 
<location id="8EF86044777F7F5381CCA363FE0BFA9D" LSID="urn:lsid:plazi:treatment:038E878977607F5381AFA4C6FF66F9C9:8EF86044777F7F5381CCA363FE0BFA9D" box="[244,404,1375,1401]" country="Mexico" county="Same" municipality="Same" name="Huayacocotla" pageId="80" pageNumber="81" stateProvince="Veracruz de Ignacio de la Llave">Huayacocotla</location>
, 
<location id="8EF86044777F7F538098A363FDAEFA9D" LSID="urn:lsid:plazi:treatment:038E878977607F5381AFA4C6FF66F9C9:8EF86044777F7F538098A363FDAEFA9D" box="[416,561,1375,1401]" country="Mexico" county="Same" municipality="Same" name="San Antonio" pageId="80" pageNumber="81" stateProvince="Veracruz de Ignacio de la Llave">San Antonio</location>
, bosque mesófilo de montaña
</materialsCitation>
, 
<materialsCitation id="3B4F3CC2777F7F5382ADA363FE81FA25" collectingDate="2006-03-06" collectingDateMax="2006-04-02" collectingDateMin="2006-03-06" collectionCode="CC-UAEH" collectorName="F. Ramirez" country="Mexico" elevation="1434" latitude="20.67561" location="Helechales" longLatPrecision="1" longitude="-98.39075" municipality="Huayacocotla" pageId="80" pageNumber="81" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
<quantity id="4CDF9B7A777F7F5382ADA363FC71FA9D" box="[917,1006,1375,1401]" metricMagnitude="3" metricUnit="m" metricValue="1.434" pageId="80" pageNumber="81" unit="m" value="1434.0">
<elevation id="000AD1AC777F7F5382ADA363FC71FA9D" box="[917,1006,1375,1401]" metricMagnitude="3" metricUnit="m" metricValue="1.434" pageId="80" pageNumber="81" unit="m" value="1434.0">1434 m</elevation>
</quantity>
, 
<geoCoordinate id="EE135058777F7F5382C4A363FB03FA9D" box="[1020,1180,1375,1401]" degrees="20" direction="north" minutes="40" orientation="latitude" pageId="80" pageNumber="81" precision="1" seconds="32.2" value="20.67561">N20°40’32.2”</geoCoordinate>
, 
<geoCoordinate id="EE135058777F7F538592A363FACFFA9D" box="[1194,1360,1375,1401]" degrees="98" direction="west" minutes="23" orientation="longitude" pageId="80" pageNumber="81" precision="1" seconds="26.7" value="-98.39075">W98°23’26.7”</geoCoordinate>
, NTP- 80 (calamar) #3, 
<date id="FF99105F777F7F538061A3BFFD81FA79" box="[345,542,1411,1437]" pageId="80" pageNumber="81" value="2006-03-06" valueMax="2006-04-02" valueMin="2006-03-06">
<collectingDate id="EFDDE9B7777F7F538061A3BFFD81FA79" box="[345,542,1411,1437]" pageId="80" pageNumber="81" value="2006-03-06" valueMax="2006-04-02" valueMin="2006-03-06">6-III a 2-IV-2006</collectingDate>
</date>
, 
<collectorName id="26D25349777F7F538312A3BFFD3AFA79" box="[554,677,1411,1437]" pageId="80" pageNumber="81">F. Ramírez</collectorName>
col.” (1, 
<collectionCode id="ED36AE5A777F7F538228A3BFFC0DFA79" box="[784,914,1411,1437]" pageId="80" pageNumber="81">CC-UAEH</collectionCode>
). “ 
<collectingMunicipality id="6BFCACE5777F7F538296A3BFFBD1FA79" box="[942,1102,1411,1437]" pageId="80" pageNumber="81">Huayacocotla</collectingMunicipality>
, 
<location id="8EF86044777F7F538562A3BFFB47FA79" LSID="urn:lsid:plazi:treatment:038E878977607F5381AFA4C6FF66F9C9:8EF86044777F7F538562A3BFFB47FA79" box="[1114,1240,1411,1437]" country="Mexico" latitude="20.67561" longLatPrecision="1" longitude="-98.39075" municipality="Huayacocotla" name="Helechales" pageId="80" pageNumber="81" stateProvince="Veracruz de Ignacio de la Llave">Helechales</location>
, bosque mesófilo de montaña
</materialsCitation>
, 
<materialsCitation id="3B4F3CC2777F7F538014A39BFAFCFA01" collectingDate="2011-04-02" collectionCode="CC-UAEH" collectorName="J. Marquez &amp; I. Martinez" country="Mexico" county="Totutla" elevation="1936" latitude="20.625" location="Rancho Zacuapam" longLatPrecision="126" longitude="-98.46" municipality="Mata Oscura" pageId="80" pageNumber="81" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
<quantity id="4CDF9B7A777F7F538014A39BFE1AFA25" box="[300,389,1447,1473]" metricMagnitude="3" metricUnit="m" metricValue="1.936" pageId="80" pageNumber="81" unit="m" value="1936.0">
<elevation id="000AD1AC777F7F538014A39BFE1AFA25" box="[300,389,1447,1473]" metricMagnitude="3" metricUnit="m" metricValue="1.936" pageId="80" pageNumber="81" unit="m" value="1936.0">1936 m</elevation>
</quantity>
, 
<geoCoordinate id="EE135058777F7F5380ACA39AFD95FA25" box="[404,522,1446,1473]" degrees="20" direction="north" minutes="37.5" orientation="latitude" pageId="80" pageNumber="81" precision="92" value="20.625">N20°37.5’</geoCoordinate>
25” 
<geoCoordinate id="EE135058777F7F538303A39BFD26FA25" box="[571,697,1447,1473]" degrees="98" direction="west" minutes="27.6" orientation="longitude" pageId="80" pageNumber="81" precision="92" value="-98.46">W98°27.6’</geoCoordinate>
49”, NTP-80 (calamar) #10, 19-III al 
<date id="FF99105F777F7F538553A39BFB7FFA25" box="[1131,1248,1447,1473]" pageId="80" pageNumber="81" value="2011-04-02">
<collectingDate id="EFDDE9B7777F7F538553A39BFB7FFA25" box="[1131,1248,1447,1473]" pageId="80" pageNumber="81" value="2011-04-02">2-IV-2011</collectingDate>
</date>
, 
<collectorName id="26D25349777F7F5385D6A39BFAF2FA25" box="[1262,1389,1447,1473]" pageId="80" pageNumber="81">J. Márquez</collectorName>
e 
<collectorName id="26D25349777F7F5384B4A39BFF63FA01" pageId="80" pageNumber="81">I. Martínez</collectorName>
cols.” (1, 
<collectionCode id="ED36AE5A777F7F538044A3F7FE61FA01" box="[380,510,1483,1509]" pageId="80" pageNumber="81">CC-UAEH</collectionCode>
). “ 
<collectingCounty id="62F94E13777F7F538326A3F7FDECFA01" box="[542,627,1483,1509]" pageId="80" pageNumber="81">Totutla</collectingCounty>
, 
<collectingMunicipality id="6BFCACE5777F7F5383B9A3F7FC87FA01" box="[641,792,1483,1509]" pageId="80" pageNumber="81">Mata Oscura</collectingMunicipality>
, 
<location id="8EF86044777F7F53821FA3F7FB9FFA01" LSID="urn:lsid:plazi:treatment:038E878977607F5381AFA4C6FF66F9C9:8EF86044777F7F53821FA3F7FB9FFA01" box="[807,1024,1483,1509]" country="Mexico" county="Totutla" latitude="20.625" longLatPrecision="126" longitude="-98.46" municipality="Mata Oscura" name="Rancho Zacuapam" pageId="80" pageNumber="81" stateProvince="Veracruz de Ignacio de la Llave">Rancho Zacuapam</location>
, selva mediana subcaducifolia
</materialsCitation>
, 
<materialsCitation id="3B4F3CC2777F7F53844AA3F7FF6AF9C9" collectingDate="1999-05-18" collectionCode="MAAS" collectorName="J. Asiain &amp; Q. Santiago &amp; J. Marquez" country="Mexico" elevation="864" latitude="19.20639" longLatPrecision="21" longitude="-96.842224" pageId="80" pageNumber="81" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave">
<quantity id="4CDF9B7A777F7F53844AA3F7FF30F9ED" metricMagnitude="2" metricUnit="m" metricValue="8.64" pageId="80" pageNumber="81" unit="m" value="864.0">
<elevation id="000AD1AC777F7F53844AA3F7FF30F9ED" metricMagnitude="2" metricUnit="m" metricValue="8.64" pageId="80" pageNumber="81" unit="m" value="864.0">864 m</elevation>
</quantity>
, 
<geoCoordinate id="EE135058777F7F538186A3D3FED5F9ED" box="[190,330,1519,1545]" degrees="19" direction="north" minutes="12" orientation="latitude" pageId="80" pageNumber="81" precision="15" seconds="23" value="19.20639">N19°12’23”</geoCoordinate>
, 
<geoCoordinate id="EE135058777F7F538060A3D2FE75F9ED" box="[344,490,1518,1545]" degrees="96" direction="west" minutes="50" orientation="longitude" pageId="80" pageNumber="81" precision="15" seconds="32" value="-96.842224">W96°50’32”</geoCoordinate>
, hojarasca de riachuelo, 
<date id="FF99105F777F7F538233A3D3FC19F9ED" box="[779,902,1519,1545]" pageId="80" pageNumber="81" value="1999-05-18">
<collectingDate id="EFDDE9B7777F7F538233A3D3FC19F9ED" box="[779,902,1519,1545]" pageId="80" pageNumber="81" value="1999-05-18">18-V-1999</collectingDate>
</date>
, 
<collectorName id="26D25349777F7F5382ADA3D3FC64F9ED" box="[917,1019,1519,1545]" pageId="80" pageNumber="81">J. Asiain</collectorName>
, 
<collectorName id="26D25349777F7F538532A3D3FB0EF9ED" box="[1034,1169,1519,1545]" pageId="80" pageNumber="81">Q. Santiago</collectorName>
y 
<collectorName id="26D25349777F7F53858CA3D3FAABF9ED" box="[1204,1332,1519,1545]" pageId="80" pageNumber="81">J. Márquez</collectorName>
col.” (1, 
<collectionCode id="ED36AE5A777F7F5381AFA02FFF72F9C9" box="[151,237,1555,1581]" country="Netherlands" lsid="urn:lsid:biocol.org:col:13754" name="Natuurhistorisch Museum Maastricht, Botany Department" pageId="80" pageNumber="81" type="Herbarium">MAAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>