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<document id="6DBD9D425B71C889525288A8351FC81E" ID-CLB-Dataset="5563" ID-DOI="10.11646/zootaxa.4990.3.1" ID-GBIF-Dataset="6a6dd89d-6b74-47ed-ba8e-71b526355b69" ID-ISSN="1175-5326" ID-Zenodo-Dep="5026777" ID-ZooBank="70610FEE-4497-4804-817C-CEC2D66DDBFE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624542810433" checkinUser="plazi" docAuthor="Maxwell, Stephen J., Rymer, Tasmin L. &amp; Congdon, Bradley C." docDate="2021" docId="039B87838745FFEFFF1EF0AD114FFCAB" docLanguage="en" docName="zootaxa.4990.3.1.pdf" docOrigin="Zootaxa 4990 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudoterebellum Maxwell &amp; Rymer &amp; Congdon 2021, new genus" docType="treatment" docVersion="9" lastPageNumber="432" masterDocId="FFA2FFFB875AFFF0FF89F03A132EFF92" masterDocTitle="Resolving phylogenetic and classical nomenclature: A revision of Seraphsidae Jung, 1974 (Gastropoda: Neostromboidae)" masterLastPageNumber="453" masterPageNumber="401" pageNumber="432" updateTime="1699020302559" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7CF2EBA7A2E8D96DF5C96BEDC1E220A2">Resolving phylogenetic and classical nomenclature: A revision of Seraphsidae Jung, 1974 (Gastropoda: Neostromboidae)</mods:title>
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<mods:namePart id="EEFC6CA922D8A3F2EA117617479C729F">Maxwell, Stephen J.</mods:namePart>
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<mods:affiliation id="14F030227A8D9C098E1E3B53E6C83329">College of Science and Engineering, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. &amp; Stephen. maxwell @ my. jcu. edu. au; https: // orcid. org / 0000 - 0002 - 4328 - 6171</mods:affiliation>
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<mods:namePart id="7EB3972922B9D08AE3F1F17F402FEF7F">Rymer, Tasmin L.</mods:namePart>
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<mods:affiliation id="94ADF28DFFB26577E668B060CD5D8D13">College of Science and Engineering, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. &amp; Centre for Tropical Environmental and Sustainability Sciences, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. &amp; Tamsin. rymer @ jcu. edu. au; https: // orcid. org / 0000 - 0002 - 9963 - 6345</mods:affiliation>
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<mods:namePart id="14747CB5870FBE6C008BC80A932052D8">Congdon, Bradley C.</mods:namePart>
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<mods:affiliation id="88B9AAB2B4B2EC857467ED0E6EC05133">College of Science and Engineering, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. &amp; Centre for Tropical Environmental and Sustainability Sciences, James Cook University, P. O. Box 6811, Cairns, Queensland 4870, Australia. &amp; brad. congdon @ jcu. edu. au; https: // orcid. org / 0000 - 0002 - 8751 - 0892</mods:affiliation>
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<treatment id="039B87838745FFEFFF1EF0AD114FFCAB" ID-DOI="http://doi.org/10.5281/zenodo.5088444" ID-GBIF-Taxon="183411037" ID-Zenodo-Dep="5088444" LSID="urn:lsid:plazi:treatment:039B87838745FFEFFF1EF0AD114FFCAB" httpUri="http://treatment.plazi.org/id/039B87838745FFEFFF1EF0AD114FFCAB" lastPageNumber="432" pageId="31" pageNumber="432">
<subSubSection id="C328651E8745FFEFFF1EF0AD12D7FF23" box="[151,505,151,178]" pageId="31" pageNumber="432" type="nomenclature">
<paragraph id="8B8D36958745FFEFFF1EF0AD12D7FF23" blockId="31.[151,505,151,249]" box="[151,505,151,178]" pageId="31" pageNumber="432">
<heading id="D0C581F98745FFEFFF1EF0AD12D7FF23" bold="true" box="[151,505,151,178]" fontSize="11" level="1" pageId="31" pageNumber="432" reason="1">
<emphasis id="B946EA878745FFEFFF1EF0AD12D7FF23" bold="true" box="[151,505,151,178]" pageId="31" pageNumber="432">
<taxonomicName id="4C324D168745FFEFFF1EF0AD125EFF20" ID-CoL="8KTGT" authority="Maxwell &amp; Rymer &amp; Congdon, 2021" authorityName="Maxwell &amp; Rymer &amp; Congdon" authorityYear="2021" box="[151,368,151,178]" class="Gastropoda" family="Seraphsidae" genus="Pseudoterebellum" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="432" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B946EA878745FFEFFF1EF0AD125EFF20" bold="true" box="[151,368,151,178]" italics="true" pageId="31" pageNumber="432">Pseudoterebellum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27557FC8745FFEFFEFEF0AD12D7FF23" box="[375,505,151,177]" pageId="31" pageNumber="432" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
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<subSubSection id="C328651E8745FFEFFF1EF0811205FF6B" pageId="31" pageNumber="432" type="description">
<paragraph id="8B8D36958745FFEFFF1EF0811253FF44" blockId="31.[151,505,151,249]" box="[151,381,187,214]" pageId="31" pageNumber="432">
(
<taxonomicName id="4C324D168745FFEFFF16F0811258FF44" authority="Maxwell &amp; Rymer &amp; Congdon, 2021" authorityName="Maxwell &amp; Rymer &amp; Congdon" authorityYear="2021" box="[159,374,187,214]" class="Gastropoda" family="Seraphsidae" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="425" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Pseudoterebellinae">Pseudoterebellinae</taxonomicName>
)
</paragraph>
<paragraph id="8B8D36958745FFEFFF1EF0E51205FF6B" blockId="31.[151,505,151,249]" box="[151,299,223,249]" pageId="31" pageNumber="432">
<figureCitation id="13092A108745FFEFFF1EF0E51205FF6B" box="[151,299,223,249]" captionStart="FIGURE 8" captionStartId="32.[151,250,1577,1601]" captionTargetBox="[204,1373,360,1524]" captionTargetId="figure-88@32.[180,1407,344,1553]" captionTargetPageId="32" captionText="FIGURE 8. Pseudoterebellum new genus and Terebellum Röding, 1798. A. Pseudoterebellum diversiornatum (Eames, 1952), holotype, NHMUK GG68239, length 10.1 mm (Jung 1974, pl. 14, figs. 1214). B. Pseudoterebellum oympiae (Rolando, 2001), holotype, MGPD 28168, length 20.9 mm (Rolando 2001, fig. 2). C. Pseudoterebellum saudiensis (Abbass, 1972), holotype, ASU S53, length 25 mm (Abbass 1972, pl. V, figs. 1, 2). D. Terebellum cinctum Martin, 1916, holotype, RGM St10337, length 34.7 mm (Jung 1974, pl. 13, figs. 4, 6). E. Terebellum obtusum Sowerby, 1840, holotype, NHMUK GG21000, length 42.2 mm (Jung 1974, pl. 13, figs. 17, 18). F. Terebellum papilliferum Martin, 1916, lectotype, RGM St10339, length 28 mm (Jung 1974, pl. 13, figs. 9, 11)." figureDoi="http://doi.org/10.5281/zenodo.5026799" httpUri="https://zenodo.org/record/5026799/files/figure.png" pageId="31" pageNumber="432">Figure 8AC</figureCitation>
</paragraph>
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<subSubSection id="C328651E8745FFEFFF1EF11D115DFEF7" pageId="31" pageNumber="432" type="materials_examined">
<paragraph id="8B8D36958745FFEFFF1EF11D10A4FED0" blockId="31.[151,1436,295,825]" box="[151,906,295,322]" pageId="31" pageNumber="432">
<emphasis id="B946EA878745FFEFFF1EF11D13F7FED3" bold="true" box="[151,217,295,321]" pageId="31" pageNumber="432">
<typeStatus id="548988378745FFEFFF1EF11D13FAFED3" box="[151,212,295,321]" pageId="31" pageNumber="432">Type</typeStatus>
.
</emphasis>
<taxonomicName id="4C324D168745FFEFFF69F11D11C0FED3" authority="Eames, 1952, p. 74" authorityName="Eames" authorityPageNumber="74" authorityYear="1952" box="[224,750,295,322]" class="Gastropoda" family="Seraphsidae" genus="Terebellum" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="432" phylum="Mollusca" rank="species" species="diversiornatum">
<emphasis id="B946EA878745FFEFFF69F11D1120FED3" box="[224,526,295,321]" italics="true" pageId="31" pageNumber="432">Terebellum diversiornatum</emphasis>
<bibRefCitation id="EFA34B648745FFEFFD9CF1121186FED0" author="Eames, F. E." box="[533,680,295,322]" pageId="31" pageNumber="432" pagination="1 - 168" refId="ref34832" refString="Eames, F. E. (1952) A contribution to the study of the Eocene in Western Pakistan and Western India C. The description of the Scaphopoda and Gastropoda from standard sections in the Rakhi Nala and Zinda Pir areas of the Western Piunjab and in the Kohat District. Philosophical Transactions of the Royal Society of London, 236, 1 - 168. https: // doi. org / 10.1098 / rstb. 1952.0001" type="journal article" year="1952">Eames, 1952</bibRefCitation>
, p. 74
</taxonomicName>
, pl. 3, fig. 77.
</paragraph>
<paragraph id="8B8D36958745FFEFFF4EF171115DFEF7" blockId="31.[151,1436,295,825]" box="[199,627,331,357]" pageId="31" pageNumber="432">
<emphasis id="B946EA878745FFEFFF4EF1711115FEF7" bold="true" box="[199,571,331,357]" pageId="31" pageNumber="432">RegNum Registration Number.</emphasis>
683.
</paragraph>
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<subSubSection id="C328651E8745FFEFFF4EF155111AFE1B" box="[199,564,367,393]" pageId="31" pageNumber="432" type="description">
<paragraph id="8B8D36958745FFEFFF4EF155111AFE1B" blockId="31.[151,1436,295,825]" box="[199,564,367,393]" pageId="31" pageNumber="432">
<emphasis id="B946EA878745FFEFFF4EF15512E9FE1B" bold="true" box="[199,455,367,393]" pageId="31" pageNumber="432">Reference Phylogeny.</emphasis>
<figureCitation id="13092A108745FFEFFE47F1551100FE1B" box="[462,558,367,393]" captionStart="FIGURE 3" captionStartId="5.[151,250,833,857]" captionTargetBox="[155,1432,185,804]" captionTargetId="figure-19@5.[151,1436,181,809]" captionTargetPageId="5" captionText="FIGURE 3. The clade Seraphsidae Jung, 1974 derived from character analysis and contextualised with the fossil record showing the ancestral Seraphsinae and crown clade Pseudoterebellinae both new subfamilies." figureDoi="http://doi.org/10.5281/zenodo.5026783" httpUri="https://zenodo.org/record/5026783/files/figure.png" pageId="31" pageNumber="432">Figure 3</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C328651E8745FFEFFF4EF1A911D1FE64" pageId="31" pageNumber="432" type="discussion">
<paragraph id="8B8D36958745FFEFFF4EF1A911D1FE64" blockId="31.[151,1436,295,825]" pageId="31" pageNumber="432">
<emphasis id="B946EA878745FFEFFF4EF1A91268FE3F" bold="true" box="[199,326,403,429]" pageId="31" pageNumber="432">Definition.</emphasis>
The maximum clade consisting of
<taxonomicName id="4C324D168745FFEFFD47F1A910B9FE3F" authority="Maxwell &amp; Rymer &amp; Congdon, 2021" authorityName="Maxwell &amp; Rymer &amp; Congdon" authorityYear="2021" box="[718,919,403,429]" class="Gastropoda" family="Seraphsidae" genus="Pseudoterebellum" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="433" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B946EA878745FFEFFD47F1A910B9FE3F" box="[718,919,403,429]" italics="true" pageId="31" pageNumber="432">Pseudoterebellum</emphasis>
</taxonomicName>
<taxonomicName id="4C324D168745FFEFFC17F1A917DDFE3C" authority="(Eames, 1952)" baseAuthorityName="Eames" baseAuthorityYear="1952" box="[926,1267,403,430]" class="Gastropoda" family="Seraphsidae" genus="Terebellum" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="432" phylum="Mollusca" rank="species" species="diversiornatum">
<emphasis id="B946EA878745FFEFFC17F1A91767FE3F" box="[926,1097,403,429]" italics="true" pageId="31" pageNumber="432">diversiornatum</emphasis>
(
<bibRefCitation id="EFA34B648745FFEFFBD1F1A917C5FE3C" author="Eames, F. E." box="[1112,1259,403,430]" pageId="31" pageNumber="432" pagination="1 - 168" refId="ref34832" refString="Eames, F. E. (1952) A contribution to the study of the Eocene in Western Pakistan and Western India C. The description of the Scaphopoda and Gastropoda from standard sections in the Rakhi Nala and Zinda Pir areas of the Western Piunjab and in the Kohat District. Philosophical Transactions of the Royal Society of London, 236, 1 - 168. https: // doi. org / 10.1098 / rstb. 1952.0001" type="journal article" year="1952">Eames, 1952</bibRefCitation>
)
</taxonomicName>
and all species that share a more recent common ancestor with it than with
<taxonomicName id="4C324D168745FFEFFCB5F18D1787FE40" authority="(Brander, 1766)" baseAuthorityName="Brander" baseAuthorityYear="1766" box="[828,1193,439,466]" class="Gastropoda" family="Seraphsidae" genus="Seraphs" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="432" phylum="Mollusca" rank="species" species="sopitus">
<emphasis id="B946EA878745FFEFFCB5F18D10C2FE43" box="[828,1004,439,465]" italics="true" pageId="31" pageNumber="432">Seraphs sopitus</emphasis>
(
<bibRefCitation id="EFA34B648745FFEFFC75F18D178FFE40" author="Brander, G." box="[1020,1185,439,466]" pageId="31" pageNumber="432" refId="ref32698" refString="Brander, G. (1766) Fossilia Hantoniensia Collecta, et in Museao Britannico Deposita. s. n., London, 43 pp. https: // doi. org / 10.5962 / bhl. title. 146466" type="book" year="1766">Brander, 1766</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C324D168745FFEFFB3CF18D123CFE67" authority="Jung, 1974" authorityName="Jung" authorityYear="1974" class="Gastropoda" family="Seraphsidae" genus="Paraseraphs" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="432" phylum="Mollusca" rank="species" species="tetanus">
<emphasis id="B946EA878745FFEFFB3CF18D16B2FE43" box="[1205,1436,439,465]" italics="true" pageId="31" pageNumber="432">Paraseraphs tetanus</emphasis>
<bibRefCitation id="EFA34B648745FFEFFF1EF1E6123CFE67" author="Jung, P." box="[151,274,475,502]" pageId="31" pageNumber="432" pagination="1 - 72" refId="ref36677" refString="Jung, P. (1974) A revision of the family Seraphsidae (Gastropoda: Strombacea). Palaeontographica Americana, 8, 1 - 72." type="journal article" year="1974">Jung, 1974</bibRefCitation>
</taxonomicName>
or
<taxonomicName id="4C324D168745FFEFFEBEF1E111D4FE64" authority="terebellum (Linnaeus, 1758)" authorityName="terebellum (Linnaeus" authorityYear="1758" baseAuthorityName="Terebellum" baseAuthorityYear="1758" box="[311,762,475,502]" class="Gastropoda" family="Seraphsidae" genus="Terebellum" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="432" phylum="Mollusca" rank="species" species="terebellum">
<emphasis id="B946EA878745FFEFFEBEF1E1111FFE67" box="[311,561,475,501]" italics="true" pageId="31" pageNumber="432">Terebellum terebellum</emphasis>
(
<bibRefCitation id="EFA34B648745FFEFFDC9F1E111DCFE64" author="Linnaeus, C." box="[576,754,475,502]" pageId="31" pageNumber="432" refId="ref37629" refString="Linnaeus, C. (1758) Systema Naturae per Regna Tria Naturae Secundum Classes, Ordines, Genera, Species, cum Caracteribus, Differentiis, Synonymis, Locis. Vol. 1. 10 th Edition. Reformata. Laurentii Salvii, Holmiae, 824 pp. https: // doi. org / 10.5962 / bhl. title. 542" type="book" year="1758">Linnaeus, 1758</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C328651E8745FFEFFF4EF1C5115DFD14" pageId="31" pageNumber="432" type="diagnosis">
<paragraph id="8B8D36958745FFEFFF4EF1C510BDFDAF" blockId="31.[151,1436,295,825]" pageId="31" pageNumber="432">
<emphasis id="B946EA878745FFEFFF4EF1C5126CFD8B" bold="true" box="[199,322,511,537]" pageId="31" pageNumber="432">Diagnosis.</emphasis>
The shell is evolute, lacking a posterior canal, and is widest near the posterior. The labrum of the shell is protocyrt. The suture is channeled. The columella is smooth.
</paragraph>
<paragraph id="8B8D36958745FFEFFF4EF27D115DFD14" blockId="31.[151,1436,295,825]" pageId="31" pageNumber="432">
<emphasis id="B946EA878745FFEFFF4EF27D1249FDF3" bold="true" box="[199,359,583,609]" pageId="31" pageNumber="432">Composition.</emphasis>
<taxonomicName id="4C324D168745FFEFFEE7F27D1119FDF3" authority="Maxwell &amp; Rymer &amp; Congdon, 2021" authorityName="Maxwell &amp; Rymer &amp; Congdon" authorityYear="2021" box="[366,567,583,609]" class="Gastropoda" family="Seraphsidae" genus="Pseudoterebellum" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="433" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B946EA878745FFEFFEE7F27D1119FDF3" box="[366,567,583,609]" italics="true" pageId="31" pageNumber="432">Pseudoterebellum</emphasis>
</taxonomicName>
contains three species detailed herein, and belongs to the
<taxonomicName id="4C324D168745FFEFFB4BF27D16B7FDF3" authority="Maxwell &amp; Rymer &amp; Congdon, 2021" authorityName="Maxwell &amp; Rymer &amp; Congdon" authorityYear="2021" box="[1218,1433,583,609]" class="Gastropoda" family="Seraphsidae" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="425" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Pseudoterebellinae">Pseudoterebellinae</taxonomicName>
, but excludes
<taxonomicName id="4C324D168745FFEFFEA2F2511294FD17" authorityName="Jung" authorityYear="1974" box="[299,442,619,645]" class="Gastropoda" family="Seraphsidae" genus="Paraseraphs" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="432" phylum="Mollusca" rank="genus">
<emphasis id="B946EA878745FFEFFEA2F2511294FD17" box="[299,442,619,645]" italics="true" pageId="31" pageNumber="432">Paraseraphs</emphasis>
</taxonomicName>
and
<taxonomicName id="4C324D168745FFEFFE79F2511142FD17" baseAuthorityName="Jung" baseAuthorityYear="1974" box="[496,620,619,645]" class="Gastropoda" family="Seraphsidae" genus="Terebellum" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="432" phylum="Mollusca" rank="genus">
<emphasis id="B946EA878745FFEFFE79F2511142FD17" box="[496,620,619,645]" italics="true" pageId="31" pageNumber="432">Terebellum</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C328651E8745FFEFFF4EF2B5114FFCAB" pageId="31" pageNumber="432" type="discussion">
<paragraph id="8B8D36958745FFEFFF4EF2B5114FFCAB" blockId="31.[151,1436,295,825]" pageId="31" pageNumber="432">
<emphasis id="B946EA878745FFEFFF4EF2B51215FD3B" bold="true" box="[199,315,655,681]" pageId="31" pageNumber="432">Remarks.</emphasis>
This clade arose in the early Ypresian of the Middle East (Table 7). Members of the complex may have radiated east via the Tethys during the Ypresian into the Indonesian region of the southwestern Pacific, and contained the forebearer of the modern clade
<taxonomicName id="4C324D168745FFEFFD19F2ED1022FD63" baseAuthorityName="Jung" baseAuthorityYear="1974" box="[656,780,727,753]" class="Gastropoda" family="Seraphsidae" genus="Terebellum" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="432" phylum="Mollusca" rank="genus">
<emphasis id="B946EA878745FFEFFD19F2ED1022FD63" box="[656,780,727,753]" italics="true" pageId="31" pageNumber="432">Terebellum</emphasis>
</taxonomicName>
. However, there is a significant geological discontinuity in the geological record from the period of extinction of
<taxonomicName id="4C324D168745FFEFFD79F2C11097FC87" authority="Maxwell &amp; Rymer &amp; Congdon, 2021" authorityName="Maxwell &amp; Rymer &amp; Congdon" authorityYear="2021" box="[752,953,763,789]" class="Gastropoda" family="Seraphsidae" genus="Pseudoterebellum" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="433" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B946EA878745FFEFFD79F2C11097FC87" box="[752,953,763,789]" italics="true" pageId="31" pageNumber="432">Pseudoterebellum</emphasis>
</taxonomicName>
at the end of the Eocene, to the appearance of
<taxonomicName id="4C324D168745FFEFFF3FF325121CFCAB" baseAuthorityName="Jung" baseAuthorityYear="1974" box="[182,306,799,825]" class="Gastropoda" family="Seraphsidae" genus="Terebellum" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="432" phylum="Mollusca" rank="genus">
<emphasis id="B946EA878745FFEFFF3FF325121CFCAB" box="[182,306,799,825]" italics="true" pageId="31" pageNumber="432">Terebellum</emphasis>
</taxonomicName>
during the mid-Oligocene.
</paragraph>
</subSubSection>
</treatment>
</document>