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<document id="2958E71872B203E7E78CE806D1EE7483" ID-DOI="10.11646/zootaxa.5009.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5157635" ID-ZooBank="3E5643F0-BBC2-45FA-83E5-07FEF6ECB690" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628051913103" checkinUser="plazi" docAuthor="Gerecke, Reinhard" docDate="2021" docId="039B1062FF80CB0947C777D9FBCDFD8F" docLanguage="en" docName="zootaxa.5009.1.1.pdf" docOrigin="Zootaxa 5009 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5009.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Atractides protendens K. O. Viets 1955" docType="treatment" docVersion="4" lastPageNumber="39" masterDocId="FFA2681AFFA5CB2F4750774EFFA7FFEE" masterDocTitle="The water mites of the family Hygrobatidae (Acari, Hydrachnidia) in Italy" masterLastPageNumber="85" masterPageNumber="1" pageNumber="38" updateTime="1699033811025" updateUser="plazi">
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<mods:title id="45A2F8C6C7D1DFA2C2070DEC23D1B05E">The water mites of the family Hygrobatidae (Acari, Hydrachnidia) in Italy</mods:title>
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<mods:namePart id="DC411E1E368A492EFA84C2F0A5915A3E">Gerecke, Reinhard</mods:namePart>
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<mods:date id="FCB7459E51EC23689D2398FBF1B8FB8A">2021</mods:date>
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<treatment id="039B1062FF80CB0947C777D9FBCDFD8F" ID-DOI="http://doi.org/10.5281/zenodo.5162200" ID-Zenodo-Dep="5162200" LSID="urn:lsid:plazi:treatment:039B1062FF80CB0947C777D9FBCDFD8F" httpUri="http://treatment.plazi.org/id/039B1062FF80CB0947C777D9FBCDFD8F" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
<subSubSection id="C328F2FFFF80CB0A47C777D9FE91FF3B" pageId="37" pageNumber="38" type="nomenclature">
<paragraph id="8B8DA174FF80CB0A47C777D9FD6CFF5F" blockId="37.[151,715,151,213]" box="[151,715,151,178]" pageId="37" pageNumber="38">
<heading id="D0C51618FF80CB0A47C777D9FD6CFF5F" bold="true" box="[151,715,151,178]" fontSize="11" level="2" pageId="37" pageNumber="38" reason="4">
<taxonomicName id="4C32DAF7FF80CB0A47C777D9FD6CFF5F" authority="K. O. Viets, 1955" authorityName="K. O. Viets" authorityYear="1955" box="[151,715,151,178]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="protendens">
<emphasis id="B9467D66FF80CB0A47C777D9FD6CFF5F" bold="true" box="[151,715,151,178]" pageId="37" pageNumber="38">
<emphasis id="B9467D66FF80CB0A47C777D9FEB6FF5C" bold="true" box="[151,273,151,178]" italics="true" pageId="37" pageNumber="38">Atractides</emphasis>
(
<emphasis id="B9467D66FF80CB0A467377D9FEC6FF5F" bold="true" box="[291,353,151,177]" italics="true" pageId="37" pageNumber="38">s. str.</emphasis>
)
<emphasis id="B9467D66FF80CB0A462277D9FE5FFF5C" bold="true" box="[370,504,151,178]" italics="true" pageId="37" pageNumber="38">protendens</emphasis>
K.O. Viets, 1955
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8DA174FF80CB0A47C777F5FE91FF3B" blockId="37.[151,715,151,213]" box="[151,310,187,213]" pageId="37" pageNumber="38">(Figs 26 G-H)</paragraph>
</subSubSection>
<subSubSection id="C328F2FFFF80CB0947C7764DFBCDFD8F" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38" type="materials_examined">
<paragraph id="8B8DA174FF80CB0A47C7764DFAD9FD1F" blockId="37.[151,1437,259,2014]" pageId="37" pageNumber="38">
Material examined:
<emphasis id="B9467D66FF80CB0A462D764DFE4AFEF3" bold="true" box="[381,493,259,285]" pageId="37" pageNumber="38">
<collectingRegion id="49F66F96FF80CB0A462D764DFE4FFEF3" box="[381,488,259,285]" country="Italy" name="Abruzzo" pageId="37" pageNumber="38">Abruzzo</collectingRegion>
:
</emphasis>
<date id="FF8C87B4FF80CB0A46A6764AFDE4FEF3" box="[502,579,259,286]" pageId="37" pageNumber="38" value="1187-01">I 1187</date>
, 0/2/0;
<emphasis id="B9467D66FF80CB0A45CB764DFCBEFEF3" bold="true" box="[667,793,259,285]" pageId="37" pageNumber="38">
<collectingRegion id="49F66F96FF80CB0A45CB764DFCB3FEF3" box="[667,788,259,285]" country="Italy" name="Basilicata" pageId="37" pageNumber="38">Basilicata</collectingRegion>
:
</emphasis>
<date id="FF8C87B4FF80CB0A4472764AFCC8FEF3" box="[802,879,259,286]" pageId="37" pageNumber="38" value="1089-01">I 1089</date>
, 6/2/0 (1/1/0 slide);
<date id="FF8C87B4FF80CB0A430B764AFB0FFEF3" box="[1115,1192,259,286]" pageId="37" pageNumber="38" value="1093-01">I 1093</date>
, 0/2/0;
<date id="FF8C87B4FF80CB0A4250764AFAE9FEF3" box="[1280,1358,259,286]" pageId="37" pageNumber="38" value="1094-01">I 1094</date>
, 1/1/0;
<date id="FF8C87B4FF80CB0A47C77666FF43FEAF" box="[151,228,295,322]" pageId="37" pageNumber="38" value="1121-01">I 1121</date>
, 1/0/0;
<date id="FF8C87B4FF80CB0A466E7666FE2CFEAF" box="[318,395,295,322]" pageId="37" pageNumber="38" value="1123-01">I 1123</date>
, 3/1/0;
<emphasis id="B9467D66FF80CB0A46B47669FDF0FEAF" bold="true" box="[484,599,295,321]" pageId="37" pageNumber="38">
<collectingRegion id="49F66F96FF80CB0A46B47669FDF5FEAF" box="[484,594,295,321]" country="Italy" name="Calabria" pageId="37" pageNumber="38">Calabria</collectingRegion>
:
</emphasis>
<date id="FF8C87B4FF80CB0A45317666FD17FEAC" box="[609,688,295,322]" pageId="37" pageNumber="38" value="1085-01">I 1085</date>
, 0/2/0 (0/1/0 slide);
<date id="FF8C87B4FF80CB0A44CE7666FC4BFEAF" box="[926,1004,295,322]" pageId="37" pageNumber="38" value="1088-01">I 1088</date>
, 0/1/0;
<date id="FF8C87B4FF80CB0A43167666FB34FEAF" box="[1094,1171,295,322]" pageId="37" pageNumber="38" value="1100-01">I 1100</date>
, 1/0/0;
<date id="FF8C87B4FF80CB0A43BC7666FA9DFEAF" box="[1260,1338,295,322]" pageId="37" pageNumber="38" value="1108-01">I 1108</date>
, 0/1/0;
<date id="FF8C87B4FF80CB0A42C37666FF77FE8B" pageId="37" pageNumber="38" value="1110-01">I 1110</date>
, 0/1/0;
<date id="FF8C87B4FF80CB0A46767602FED6FE8B" box="[294,369,331,358]" pageId="37" pageNumber="38" value="1114-01">I 1114</date>
, 3/3/1;
<date id="FF8C87B4FF80CB0A46977602FDB4FE8B" box="[455,531,331,358]" pageId="37" pageNumber="38" value="1126-01">I 1126</date>
, 2/3/0;
<date id="FF8C87B4FF80CB0A45397602FD13FE8B" box="[617,692,331,358]" pageId="37" pageNumber="38" value="1134-01">I 1134</date>
, 0/1/0;
<date id="FF8C87B4FF80CB0A445A7602FCF1FE88" box="[778,854,331,358]" pageId="37" pageNumber="38" value="1135-01">I 1135</date>
, 3/0/1;
<emphasis id="B9467D66FF80CB0A44FC7605FC5FFE8B" bold="true" box="[940,1016,331,357]" pageId="37" pageNumber="38">
<collectingRegion id="49F66F96FF80CB0A44FC7605FC54FE8B" box="[940,1011,331,357]" country="Italy" name="Lazio" pageId="37" pageNumber="38">Lazio</collectingRegion>
:
</emphasis>
I 883, 2/1/0 slide; I 887, 0/1/0 slide;
<emphasis id="B9467D66FF80CB0A47C77621FE8FFE67" bold="true" box="[151,296,367,393]" pageId="37" pageNumber="38">
<collectingRegion id="49F66F96FF80CB0A47C77621FE84FE67" box="[151,291,367,393]" country="Italy" name="Lombardia" pageId="37" pageNumber="38">Lombardia</collectingRegion>
:
</emphasis>
I TG 13-02b, 0/3/0;
<emphasis id="B9467D66FF80CB0A45487621FD34FE67" bold="true" box="[536,659,367,393]" pageId="37" pageNumber="38">
<collectingRegion id="49F66F96FF80CB0A45487621FD29FE67" box="[536,654,367,393]" country="Italy" name="Sardegna" pageId="37" pageNumber="38">Sardegna</collectingRegion>
:
</emphasis>
I TG 15-02, 2/5/0; I TG 17-03a, 0/1/0;
<emphasis id="B9467D66FF80CB0A430A7621FB0AFE67" bold="true" box="[1114,1197,367,393]" pageId="37" pageNumber="38">
<collectingRegion id="49F66F96FF80CB0A430A7621FB00FE67" box="[1114,1191,367,393]" country="Italy" name="Sicilia" pageId="37" pageNumber="38">Sicilia</collectingRegion>
:
</emphasis>
All parts. I 59, 0/1/0 slide; I 99, 2/1/0 slide; I 100, 0/2/0 (0/1/0 slide); I 176, 1/0/0 slide; I 212, 1/0/0 slide; I 234, 0/1/0 slide; I 142, 1/0/0 slide; I 242, 1/0/0 slide; I 243, 1/5/0, 1/2/0 slide; I 244, 0/1/0 slide; I 250, 2/2/0 (1/1/0 slide); I 282, 0/2/0 (0/1/0 slide); I 286, 3/4/0 (2/2/0 slide); I 319, 0/1/0 slide; I 399, 0/2/0; I 401, 0/1/0 (0/1/0 slide); I 406, 2/5/0 (1/1/0 slide); I 407, 0/2/0 (0/1/0 slide); I 468, 1/1/1 slide; I 500, 2/0/0 slide; I 501, 1/1/0 slide; I 502, 3/1/0 slide; I 505, 1/0/0 slide; I 506, 0/2/0 (0/1/0 slide); I 517, 0/4/0 (0/1/0 slide); I 519, 0/1/0 slide; I 520, 2/5/0 (1/1/0 slide); I 529, 0/1/0 slide; I 553 Int, 1/0/0 slide; I 557, 1/1/0; I 559 Int, 0/0/1; I 560, 1/7/0; I 568, 0/2/0 (0/1/0 slide); I 575, 1/3/0 (1/1/0 slide); I 587, 2/0/0 slide; I 594, 4/3/0; I 596, 3/8/0 (1/1/0 slide); I 603, 3/3/0; I 604, 1/1/0 slide; I 607, 2/2/0; I 611, 1/0/0 slide; I 615, 1/0/0 slide; I 619, 1/0/0 slide; I 628, 1/0/0 slide; I 642, 2/4/1 (0/1/1 slide); I 647, 0/3/0 slide; I 649, 0/1/0 slide; I 650, 0/4/0; I 651, 0/1/0; I 657, 2/3/0; 659, 2/1/1; I 848, 0/1/0;
<date id="FF8C87B4FF80CB0A440575FAFC07FD23" box="[853,928,691,718]" pageId="37" pageNumber="38" value="1067-01">I 1067</date>
, 1/0/0 slide;
<date id="FF8C87B4FF80CB0A437E75FAFBDEFD23" box="[1070,1145,691,718]" pageId="37" pageNumber="38" value="1068-01">I 1068</date>
, 1/0/0 slide; I dp 12, 0/1/0 slide; I dp 56, 1/1/0 slide; I dp 64, 1/0/0 slide; I dp 66, 0/1/0 slide; I dp 67, 2/10/1 slide. 76 sites,
<specimenCount id="9D346AFDFF80CB0A43817599FADDFD1F" box="[1233,1402,727,753]" pageId="37" pageNumber="38" type="generic">232 specimens</specimenCount>
.
</paragraph>
<paragraph id="8B8DA174FF80CB0A479775B5FF47FCD7" blockId="37.[151,1437,259,2014]" pageId="37" pageNumber="38">
Previously published records from
<collectingCountry id="F325E1E4FF80CB0A453575B5FD3AFCFB" box="[613,669,763,789]" name="Italy" pageId="37" pageNumber="38">Italy</collectingCountry>
:
<collectingRegion id="49F66F96FF80CB0A45FD75B5FCB2FCFB" box="[685,789,763,789]" country="Italy" name="Sardegna" pageId="37" pageNumber="38">Sardegna</collectingRegion>
(
<bibRefCitation id="EFA3DC85FF80CB0A447875B5FC79FCF8" author="Gerecke, R." box="[808,990,763,790]" pageId="37" pageNumber="38" pagination="735 - 759" refId="ref74426" refString="Gerecke, R. (2014 b) The water mites of the genus Atractides Koch, 1837 (Acari, Hydrachnidia: Hygrobatidae) in Corsica and Sardinia. Zoosystema, 36 (4), 735 - 759. https: // doi. org / 10.5252 / z 2014 n 4 a 3" type="journal article" year="2014">Gerecke 2014b</bibRefCitation>
);
<collectingRegion id="49F66F96FF80CB0A44A775B5FB47FCFB" box="[1015,1248,763,789]" country="Italy" name="Trentino-Alto Adige" pageId="37" pageNumber="38">Trentino-Alto Adige</collectingRegion>
(
<bibRefCitation id="EFA3DC85FF80CB0A43A475B5FF73FCD7" author="Gerecke, R. &amp; Schatz, H. &amp; Wohltmann, A." pageId="37" pageNumber="38" pagination="303 - 333" refId="ref74836" refString="Gerecke, R., Schatz, H. &amp; Wohltmann, A. (2009) The mites (Chelicerata: Acari) of the CRENODAT project: Faunistic records and ecological data from springs in the Autonomous Province of Trento (Italian Alps). International Journal of Acarology, 35, 303 - 333 https: // doi. org / 10.1080 / 01647950903059452" type="journal article" year="2009">
Gerecke
<emphasis id="B9467D66FF80CB0A420F75B2FA3AFCFB" box="[1375,1437,763,789]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
2009
</bibRefCitation>
).
</paragraph>
<paragraph id="8B8DA174FF80CB0A4797740DFCF1FB93" blockId="37.[151,1437,259,2014]" pageId="37" pageNumber="38">Description: Both sexes: Integument strongly lineated. Glandularia medium sized, in aged specimens glandularia enlarged and with slightly irregular outline, and coxae with secondary sclerotization. I-L-5-6: Fig. 26 G. I-L-5 slender, ventral and dorsal margins in proximal half slightly, in distal half more strongly diverging; S-1 and -2 rather short, little away from each other, similar in size with blunt tips; I-L-6 stout and little curved, maximum H in distal part, ventral margin slightly convex in proximal quarter, slightly concave at basal third, more distally again slightly convex; I-L-claws strong. Palp (Fig. 26 H) without remarkable sexual dimorphism, P-2 ventral margin slightly concave, forming a distoventral angle; P-3 and -4 slender, P-4 with maximum H near proximoventral seta, sword seta closer to distoventral seta insertions, both setae associated with a tiny pointed projection, ventral sectors similar in size, the distal one in general slightly longer than the others.</paragraph>
<paragraph id="8B8DA174FF80CB0A479773C9FCDDF9E7" blockId="37.[151,1437,259,2014]" pageId="37" pageNumber="38">Males (n = 11): Idiosoma L/W 400-650/300-500, coxal field L/W 240-370/320-380; Cx-I+II mL 100-140, lL 185-240, W 240-300. I-L-5 dL/vL 132-156/86-112, ratio 1.3-1.5, distal sector 28-46 %; HA 28-33, HB 33-40, HC 45-53, ratio dL/HB 3.6-4.1; S-1 L 53-63, L/W ratio 9,6-12,5; S-2 L 48-60, L/W ratio 7.0-9.0; distance S-1-2, 6-8, L ratio S-1/-2, 1.0-1.2; I-L-6 dL 108-128, HA 25-29, HB 24-28, HC 31-36; ratio dL/HB 4.2-4.9, dL/HC 3.3-3.7; dL ratio I-L-5/6, 1.10-1.26. Genital field L/W 88-128/96-142, rounded, anterior margin in juveniles equally convex, in aged specimens medially strongly extended by secondary sclerotization, posterior margin with a shallow medial indentation; gonopore L 50-65, Ac in a curved line, surrounded by 16-18 pairs of setae; maximum diameter Ac-1, 24-34; Ac-2, 22-32; Ac-3, 24-32. Gnathosoma L 116-148, chelicera L 204-244, L/H ratio 4.8-5.4, basal segment/ claw 2.2-2.4. Palp total L 254-308; L/H (ratio, rel. L [%]) P-1, 25-30/21-28 (1.0-1.2, 9-10); P-2, 56-65/40-50 (1.3- 1.5, 21-24); P-3, 60-75/33-40 (1.7-2.0, 23-25); P-4, 83-108/28-35 (2.9-3.1, 32-36); P-5, 25-30/10-14 (2.2-2.7, 9-11); L ratio P-2/P-3, 0.85-1.02; P-2/P-4, 0.58-0.74; P-3/P-4, 0.67-0.73.</paragraph>
<paragraph id="8B8DA174FF80CB0A4797715DFD38F854" blockId="37.[151,1437,259,2014]" pageId="37" pageNumber="38">Females (n = 3): Idiosoma L/W 580-750/480-600, coxal field L/W 350-380/420-510; Cx-I+II mL 130-140, lL 260-300, W 340-370. I-L-5 dL/vL 188-196/132-144, ratio 1.4-1.7, distal sector 33-36 %; HA 43-45, HB 50-53, HC 66-69, ratio dL/HB 3.7-3.8; S-1 L 74-80, L/W ratio 9.4-11.4; S-2 L 65-73, L/W ratio 6.5-7.3; distance S-1-2, 13-15, L ratio S-1/-2, 1.1-1.2; I-L-6 dL 156-164, HA 33-35, HB 30-34, HC 43; ratio dL/HB 4.9-5.3, dL/HC 3.7-3.9; dL ratio I-L-5/6, 1.20-1.21. Genital field L 130-180, pregenital sclerites crescent-shaped, with fine tips, Postgenital sclerite half disk shaped, with straight anterior margin, genital plates L/W 124-132/44-48, weakly curved, anteriorly forming a blunt tip, posteriorly rounded, bearing 10-12 pairs of fine, hair-like setae; Ac similar in size, maximum diameter Ac-1, 44; Ac-2, 38-46; Ac-3, 38-42. Gnathosoma L 180-184, chelicera L 320-328, L/H ratio 5.5-5.7, basal segment/claw 2.3. Palp total L 359-378; L/H (ratio, rel. L [%]) P-1, 35-38/30-33 (1.2, 10); P-2, 78-85/60-63 (1.3- 1.4, 22-23); P-3, 95-100/48-50 (2.0, 26); P-4, 115-118/33-35 (3.4-3.5, 31-32); P-5, 36-38/14-15 (2.5-2.6, 10); L ratio P-2/P-3, 0.82-0.85; P-2/P-4, 0.67-0.72; P-3/P-4, 0.83-0.85. P-2 and P-3 with straight ventral margins, P-4 slenderer than in males, fine dorsal setae less numerous.</paragraph>
<paragraph id="8B8DA174FF80CB094797708DFEFDFE67" blockId="37.[151,1437,259,2014]" lastBlockId="38.[151,1437,151,610]" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
Remarks: At a first glance, specimens of
<taxonomicName id="4C32DAF7FF80CB0A45DE708DFC80F833" authorityName="K. O. Viets" authorityYear="1955" box="[654,807,1987,2013]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="protendens">
<emphasis id="B9467D66FF80CB0A45DE708DFC80F833" box="[654,807,1987,2013]" italics="true" pageId="37" pageNumber="38">A. protendens</emphasis>
</taxonomicName>
resemble
<taxonomicName id="4C32DAF7FF80CB0A44C9708DFBB7F833" authorityName="K. Viets" authorityYear="1950" box="[921,1040,1987,2013]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="graecus">
<emphasis id="B9467D66FF80CB0A44C9708DFBB7F833" box="[921,1040,1987,2013]" italics="true" pageId="37" pageNumber="38">A. graecus</emphasis>
</taxonomicName>
. Occasionally, a key difference, the lineation of the dorsal integument (in
<taxonomicName id="4C32DAF7FF83CB09451477D9FD1AFF5E" authorityName="K. Viets" authorityYear="1950" box="[580,701,151,177]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="graecus">
<emphasis id="B9467D66FF83CB09451477D9FD1AFF5E" box="[580,701,151,177]" italics="true" pageId="38" pageNumber="39">A. graecus</emphasis>
</taxonomicName>
finely striated as the palm of a human finger), is to be seen with difficulty only in undissected specimens or after treatment with chemical agents (e.g., after enzymatical digestion). The new measurements show that
<taxonomicName id="4C32DAF7FF83CB0945797791FD61FF17" authorityName="K. O. Viets" authorityYear="1955" box="[553,710,223,249]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="protendens">
<emphasis id="B9467D66FF83CB0945797791FD61FF17" box="[553,710,223,249]" italics="true" pageId="38" pageNumber="39">A. protendens</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C32DAF7FF83CB0943557791FB27FF16" authorityName="K. Viets" authorityYear="1950" box="[1029,1152,223,249]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="graecus">
<emphasis id="B9467D66FF83CB0943557791FB27FF16" box="[1029,1152,223,249]" italics="true" pageId="38" pageNumber="39">A. graecus</emphasis>
</taxonomicName>
(in parentheses) also by generally larger acetabula - maximum diameter
<specimenCount id="9D346AFDFF83CB0945FA764DFD73FEF3" box="[682,724,259,285]" pageId="38" pageNumber="39" type="male">♂♂</specimenCount>
22-32 (16-21),
<specimenCount id="9D346AFDFF83CB0944D2764DFC0BFEF3" box="[898,940,259,285]" pageId="38" pageNumber="39" type="female">♀♀</specimenCount>
38-46 (19-28), and several details of the I-L, e.g. I-L-5 longer, but stouter,
<specimenCount id="9D346AFDFF83CB0946B07669FDADFEAF" box="[480,522,295,321]" pageId="38" pageNumber="39" type="male">♂♂</specimenCount>
dL&gt; 130 (&lt;125),
<specimenCount id="9D346AFDFF83CB09458D7669FCA0FEAF" box="[733,775,295,321]" pageId="38" pageNumber="39" type="female">♀♀</specimenCount>
&gt; 180 (&lt;160), in both sexes dL/HB ratio 3.6-4.1 (4.1-4.5), setae S-1/-2 longer,&gt; 45 (&lt;45) and more away from each other, distance 8-14 (3-4). At site I 647, both species were found coexisting.
</paragraph>
<paragraph id="8B8DA174FF83CB09479776DDFA21FE1B" blockId="38.[151,1437,151,610]" pageId="38" pageNumber="39">
Habitat. Crenobiont – more than 80 % of the records in the study area from springs, mostly weakly seeping rheohelocrenes. In exceptional cases in low order streams, in two occasions taken in interstitial digs. Most records from middle to higher elevations (mean 930, maximum
<quantity id="4CCA0C91FF83CB09445B7695FCC4FE1B" box="[779,867,475,501]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="38" pageNumber="39" unit="m" value="1800.0">1800 m</quantity>
), but also in the macchia belt (minimum
<quantity id="4CCA0C91FF83CB09427E7694FADEFE1B" box="[1326,1401,474,501]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="38" pageNumber="39" unit="m" value="150.0">150 m</quantity>
).
</paragraph>
<paragraph id="8B8DA174FF83CB09479776B1FBCDFD8F" blockId="38.[151,1437,151,610]" pageId="38" pageNumber="39">
Distribution: Central and Southern Europe. The numerous new records suggest that
<taxonomicName id="4C32DAF7FF83CB09432E76B1FABEFDF7" authorityName="K. O. Viets" authorityYear="1955" box="[1150,1305,511,537]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="protendens">
<emphasis id="B9467D66FF83CB09432E76B1FABEFDF7" box="[1150,1305,511,537]" italics="true" pageId="38" pageNumber="39">A. protendens</emphasis>
</taxonomicName>
is a typical Mediterranean species that has extended its northern distribution margin under favourable conditions (sun-exposed open land springs) to the Alps and scattered parts of central European mountain ranges.
</paragraph>
</subSubSection>
</treatment>
</document>
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