609 lines
70 KiB
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609 lines
70 KiB
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<document id="188D783815E4304A5971D9D3F0436377" ID-CLB-Dataset="22297" ID-DOI="10.5852/ejt.2018.455" ID-GBIF-Dataset="44a2fcf6-cc1b-4c07-9554-dd6dd279a76d" ID-Zenodo-Dep="3817707" ID-ZooBank="C445E95B-446A-4601-AAA3-C1CCBAB627F9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1588963536872" checkinUser="valdenar" docAuthor="Páll-Gergely, Barna" docDate="2018" docId="03CC87A06C4C8A42FD91FA194411FB28" docLanguage="en" docName="ejt-455_pall-gergely.pdf.imf" docOrigin="European Journal of Taxonomy 455" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Plectopylis malayana Páll-Gergely, 2018, sp. nov." docType="treatment" docVersion="6" lastPageNumber="102" masterDocId="FFF5FFD86C2C8A27FFF8FFC04709FFA9" masterDocTitle="Systematic revision of the Plectopylinae (Gastropoda, Pulmonata, Plectopylidae)" masterLastPageNumber="114" masterPageNumber="1" pageNumber="97" updateTime="1698839078568" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="8EBEAF23375535452CE745204AB4D366">
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<mods:title id="2728F92A46FFD08743460DDB8B35033D">Systematic revision of the Plectopylinae (Gastropoda, Pulmonata, Plectopylidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="31FF6DE0DB1E7004A19D315DE07D1553" type="personal">
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<mods:roleTerm id="33C8F36ED2A718F9BA77775FF28C9516">Author</mods:roleTerm>
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<mods:namePart id="9DA58C8A6F772027407B2C73EEE98816">Páll-Gergely, Barna</mods:namePart>
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<mods:affiliation id="109F349AF14A5C13FB16D28D9A8AFC27">Centre for Agricultural Research, Hungarian Academy of Sciences (MTA), Herman Ottó Street 15, Budapest, H- 1022, Hungary.</mods:affiliation>
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<mods:nameIdentifier id="28776288CCC2BF0615B95398AE8065DA" type="email">pall-gergely.barna@agrar.mta.hu</mods:nameIdentifier>
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<mods:typeOfResource id="EF70D7E7B245AB4FA4A61EF08F218378">text</mods:typeOfResource>
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<mods:title id="80BEF2BC271BB5C9C153C86A9604E371">European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part id="27B569B5883EE2A59DD813AEA619D72B">
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<mods:date id="C389F9BC9B41AD0134DFE9DB68ED94B4">2018</mods:date>
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<mods:detail id="533C6CF6106CF242E43551F310FD1C5E" type="pubDate">
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<mods:number id="62D58E3260E98D57B2AC861237DDE151">2018-08-16</mods:number>
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<mods:detail id="98CFC568EB8166C2CB08105CA0C1D6B1" type="volume">
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<mods:number id="7B578458E1CD5908AC3B9B575BB2D51A">455</mods:number>
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<mods:start id="2CAA2DF5ED1D6DB4F08AC9359A896023">1</mods:start>
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<mods:identifier id="F56AB6713DEFC246913A31EF120880EE" type="CLB-Dataset">22297</mods:identifier>
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<mods:identifier id="369A8E6C23137635FA183275D59CB992" type="DOI">10.5852/ejt.2018.455</mods:identifier>
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<mods:identifier id="D2D54D3299ACB9FC44ADF6D8FE6D6972" type="GBIF-Dataset">44a2fcf6-cc1b-4c07-9554-dd6dd279a76d</mods:identifier>
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<treatment id="03CC87A06C4C8A42FD91FA194411FB28" ID-DOI="http://doi.org/10.5281/zenodo.3818806" ID-GBIF-Taxon="163993788" ID-Zenodo-Dep="3818806" LSID="urn:lsid:plazi:treatment:03CC87A06C4C8A42FD91FA194411FB28" httpUri="http://treatment.plazi.org/id/03CC87A06C4C8A42FD91FA194411FB28" lastPageId="101" lastPageNumber="102" pageId="96" pageNumber="97">
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<subSubSection id="C37F653D6C4C8A47FD91FA1943C8F9B1" pageId="96" pageNumber="97" type="nomenclature">
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<paragraph id="8BDA36B66C4C8A47FD91FA1944C2FA5D" blockId="96.[371,1217,1497,1595]" box="[617,971,1497,1524]" pageId="96" pageNumber="97">
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<heading id="D09281DA6C4C8A47FD91FA1944C2FA5D" box="[617,971,1497,1524]" centered="true" fontSize="11" level="2" pageId="96" pageNumber="97" reason="2">
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<taxonomicName id="4C654D356C4C8A47FD91FA194460FA5D" ID-CoL="4K3PJ" authorityName="Pall-Gergely" authorityYear="2018" box="[617,873,1497,1524]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="96" pageNumber="97" phylum="Mollusca" rank="species" species="malayana" status="sp. nov.">
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<emphasis id="B911EAA46C4C8A47FD91FA194460FA5D" bold="true" box="[617,873,1497,1524]" italics="true" pageId="96" pageNumber="97">Plectopylis malayana</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22257DF6C4C8A47FC89FA1A44C2FA5D" box="[881,971,1498,1524]" pageId="96" pageNumber="97" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="8BDA36B66C4C8A47FE8BFA3D43C8F9B1" blockId="96.[371,1217,1497,1595]" box="[371,1217,1533,1560]" pageId="96" pageNumber="97">
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<uri id="FFF43AB46C4C8A47FE8BFA3D43C8F9B1" box="[371,1217,1533,1560]" pageId="96" pageNumber="97">
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urn:lsid:zoobank.org:act:
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<uuid id="FFC30C636C4C8A47FD68FA3D43C8F9B1" box="[656,1217,1533,1560]" pageId="96" pageNumber="97">987C6622-4318-4E5E-AE9C-097C21DC546F</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37F653D6C4C8A47FDA0F9E044D2F993" box="[600,987,1568,1595]" pageId="96" pageNumber="97" type="description">
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<paragraph id="8BDA36B66C4C8A47FDA0F9E044D2F993" blockId="96.[371,1217,1497,1595]" box="[600,987,1568,1595]" pageId="96" pageNumber="97">
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<figureCitation id="135E2A336C4C8A47FDA0F9E045A7F992" box="[600,686,1568,1595]" captionStart="Fig" captionStartId="81.[189,232,1598,1624]" captionTargetBox="[198,1377,359,1489]" captionTargetId="figure@81.[190,1399,342,1515]" captionTargetPageId="81" captionText="Fig. 35. Parietal (A–E, G, I–L, N) and palatal (F, H, M, O) plication of species of Plectopylis Benson, 1860. A. P. anguina Gould, 1847, paralectotype (NHMUK 20140812). B. P. repercussa Gould, 1856, lectotype (MCZ 169336). C. P. bensoni var. repercussoides Gude, 1899, holotype (NHMUK 1922.8.29.46). D. P. cairnsi Gude, 1898, holotype (NHMUK 1922.8.29.47). E–F. P. cairnsi (UF 76761). G–H. P. crassilabris sp. nov., paratype (UF 448577). I. P. cyclaspis Benson, 1859 (NHMUK 1906.02.02.148). J. P. karenorum (W. Blanford, 1865) from “Pegu”, Myanmar (NHMUK). K. P. linterae var. fusca Gude, 1898 (NHMUK 1888.12.04.1541–3). L–M. P. malayana sp. nov., paratypes from the type locality (coll PGB). N–O. P. thompsoni sp. nov., paratype (UF 448578). Arrows below the letters indicate the direction of the aperture. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.3817771" httpUri="https://zenodo.org/record/3817771/files/figure.png" pageId="96" pageNumber="97">Figs 35</figureCitation>
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L–M, 44A–E, 45, 46A–B
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</paragraph>
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</subSubSection>
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<subSubSection id="C37F653D6C4C8A47FF45F9A246CEF96A" pageId="96" pageNumber="97" type="reference_group">
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<paragraph id="8BDA36B66C4C8A47FF45F9A246CEF96A" blockId="96.[189,1399,1634,1731]" pageId="96" pageNumber="97">
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<treatmentCitationGroup id="AB7511986C4C8A47FF45F9A246CAF96A" pageId="96" pageNumber="97">
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<taxonomicName id="4C654D356C4C8A47FF45F9A246A1F9D5" ID-CoL="7SR8S" baseAuthorityName="Pfeiffer" baseAuthorityYear="1867" box="[189,424,1634,1660]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="96" pageNumber="97" phylum="Mollusca" rank="species" species="revoluta">
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<emphasis id="B911EAA46C4C8A47FF45F9A246A1F9D5" box="[189,424,1634,1660]" italics="true" pageId="96" pageNumber="97">Plectopylis revoluta</emphasis>
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</taxonomicName>
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– van Benthem Jutting 1960: 17 [“Limestone hill Kaki Bukit, near kampong Wang Tangga,
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<collectingRegion id="49A1F8546C4C8A47FEAAF9454694F909" box="[338,413,1669,1696]" country="Malaysia" name="Perlis" pageId="96" pageNumber="97">Perlis</collectingRegion>
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”]. —
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<treatmentCitation id="0AC410A76C4C8A47FE1DF9464597F909" box="[485,670,1669,1696]" pageId="96" pageNumber="97">
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<bibRefCitation id="EFF44B476C4C8A47FE1DF9464597F909" author="Berry A. J." box="[485,670,1669,1696]" pageId="96" pageNumber="97" pagination="1 - 17" refId="ref51894" refString="Berry A. J. 1963. An introduction to the non-marine molluscs of Malaya. Malayan Nature Journal 17: 1 - 17." type="journal article" year="1963">Berry 1963: 13</bibRefCitation>
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</treatmentCitation>
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, fig. 51 [“abundant in limestone hills in North Malaya”].
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<emphasis id="B911EAA46C4C8A47FAA4F946427FF936" box="[1372,1398,1670,1695]" italics="true" pageId="96" pageNumber="97">—</emphasis>
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<treatmentCitation id="0AC410A76C4C8A47FF12F96946CAF96A" box="[234,451,1705,1731]" pageId="96" pageNumber="97">
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<bibRefCitation id="EFF44B476C4C8A47FF12F96946CAF96A" author="Maassen W. J. M." box="[234,451,1705,1731]" pageId="96" pageNumber="97" pagination="1 - 155" refId="ref54723" refString="Maassen W. J. M. 2001. A preliminary checklist of the non-marine Molluscs of West-Malaysia. " A handlist ". De Kreukel, Extra Editie [for 2001]: 1 - 155." type="journal article" year="2001">Maassen 2001: 78</bibRefCitation>
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</treatmentCitation>
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</treatmentCitationGroup>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37F653D6C4C8A47FF45F92A46F4F8D3" pageId="96" pageNumber="97" type="diagnosis">
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<paragraph id="8BDA36B66C4C8A47FF45F92A463FF8AD" blockId="96.[189,310,1770,1796]" box="[189,310,1770,1796]" pageId="96" pageNumber="97">
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<heading id="D09281DA6C4C8A47FF45F92A463FF8AD" bold="true" box="[189,310,1770,1796]" fontSize="11" level="3" pageId="96" pageNumber="97" reason="3">
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<emphasis id="B911EAA46C4C8A47FF45F92A463FF8AD" bold="true" box="[189,310,1770,1796]" pageId="96" pageNumber="97">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BDA36B66C4C8A47FF45F8DA46F4F8D3" blockId="96.[189,1399,1817,1914]" pageId="96" pageNumber="97">A medium-sized, sinistral, brown-shelled species having periostracal filaments along the upper keel of the body whorl; apertural fold not in contact with parietal callus; lambda-complex with a short connecting plica anteriorly.</paragraph>
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</subSubSection>
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<subSubSection id="C37F653D6C4C8A47FF45F862435DF842" box="[189,1108,1954,2027]" pageId="96" pageNumber="97" type="etymology">
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<paragraph id="8BDA36B66C4C8A47FF45F862464DF815" blockId="96.[189,324,1954,1980]" box="[189,324,1954,1980]" pageId="96" pageNumber="97">
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<heading id="D09281DA6C4C8A47FF45F862464DF815" bold="true" box="[189,324,1954,1980]" fontSize="11" level="3" pageId="96" pageNumber="97" reason="3">
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<emphasis id="B911EAA46C4C8A47FF45F862464DF815" bold="true" box="[189,324,1954,1980]" pageId="96" pageNumber="97">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BDA36B66C4C8A47FF45F811435DF842" blockId="96.[189,1108,2001,2027]" box="[189,1108,2001,2027]" pageId="96" pageNumber="97">
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The species is named for
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<collectingCountry id="F37276266C4C8A47FE10F811455EF842" box="[488,599,2001,2027]" name="Malaysia" pageId="96" pageNumber="97">Malaysia</collectingCountry>
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, where most of the samples were collected.
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</paragraph>
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</subSubSection>
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<caption id="DF1A663E6C4D8A46FF45F8FC427EF869" ID-DOI="http://doi.org/10.5281/zenodo.3817789" ID-Zenodo-Dep="3817789" httpUri="https://zenodo.org/record/3817789/files/figure.png" pageId="97" pageNumber="98" startId="97.[189,232,1852,1878]" targetBox="[189,1398,265,1805]" targetPageId="97">
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<paragraph id="8BDA36B66C4D8A46FF45F8FC427EF869" blockId="97.[189,1399,1852,1985]" pageId="97" pageNumber="98">
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<emphasis id="B911EAA46C4D8A46FF45F8FC4617F8FF" bold="true" box="[189,286,1852,1878]" pageId="97" pageNumber="98">Fig. 44.</emphasis>
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Shells of species of
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<taxonomicName id="4C654D356C4D8A46FDDBF8FC4450F8FF" authority="Benson, 1860" authorityName="Benson" authorityYear="1860" box="[547,857,1852,1879]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="97" pageNumber="98" phylum="Mollusca" rank="genus">
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<emphasis id="B911EAA46C4D8A46FDDBF8FC45ACF8FF" box="[547,677,1852,1878]" italics="true" pageId="97" pageNumber="98">Plectopylis</emphasis>
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<bibRefCitation id="EFF44B476C4D8A46FD49F8FD4450F8FF" author="Benson W. H." box="[689,857,1852,1879]" pageId="97" pageNumber="98" pagination="243 - 247" refId="ref51737" refString="Benson W. H. 1860. Notes on Plectopylis, a group a Helicidae distinguished by several internal plicate Ephiphragms; with the Characters of a new Species. The Annals and Magazine of Natural History 3 (5): 243 - 247. Available from https: // biodiversitylibrary. org / page / 18658580 [accessed 30 Jul. 2018]." type="journal article" year="1860">Benson, 1860</bibRefCitation>
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</taxonomicName>
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.
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<emphasis id="B911EAA46C4D8A46FC91F8FC4477F8FF" bold="true" box="[873,894,1852,1878]" pageId="97" pageNumber="98">A</emphasis>
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.
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<taxonomicName id="4C654D356C4D8A46FC68F8FD432CF8FF" authorityName="Pall-Gergely" authorityYear="2018" box="[912,1061,1852,1878]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="97" pageNumber="98" phylum="Mollusca" rank="species" species="malayana" status="sp. nov.">
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<emphasis id="B911EAA46C4D8A46FC68F8FD432CF8FF" box="[912,1061,1852,1878]" italics="true" pageId="97" pageNumber="98">P. malayana</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22257DF6C4D8A46FBC8F8FD4398F8FE" box="[1072,1169,1853,1879]" pageId="97" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
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, holotype (HNHM 97469).
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<emphasis id="B911EAA46C4D8A46FED8F89F465EF8D0" bold="true" box="[288,343,1887,1913]" pageId="97" pageNumber="98">B–C</emphasis>
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. Living specimen of
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<taxonomicName id="4C654D356C4D8A46FDAEF8A045E1F8D3" authorityName="Pall-Gergely" authorityYear="2018" box="[598,744,1888,1914]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="97" pageNumber="98" phylum="Mollusca" rank="species" species="malayana" status="sp. nov.">
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<emphasis id="B911EAA46C4D8A46FDAEF8A045E1F8D3" box="[598,744,1888,1914]" italics="true" pageId="97" pageNumber="98">P. malayana</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22257DF6C4D8A46FD08F8A04445F8D3" box="[752,844,1888,1914]" pageId="97" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
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|
|
<emphasis id="B911EAA46C4D8A46FCACF8A04482F8D3" bold="true" box="[852,907,1888,1914]" pageId="97" pageNumber="98">D–E</emphasis>
|
|||
|
|
. SEM images of the periostracal folds of
|
|||
|
|
<taxonomicName id="4C654D356C4D8A46FF45F8444645F834" authorityName="Pall-Gergely" authorityYear="2018" box="[189,332,1923,1949]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="97" pageNumber="98" phylum="Mollusca" rank="species" species="malayana" status="sp. nov.">
|
|||
|
|
<emphasis id="B911EAA46C4D8A46FF45F8444645F834" box="[189,332,1923,1949]" italics="true" pageId="97" pageNumber="98">P. malayana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A22257DF6C4D8A46FEAAF84346A4F834" box="[338,429,1923,1949]" pageId="97" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
, specimens from loc. 2013/14.
|
|||
|
|
<emphasis id="B911EAA46C4D8A46FCEDF843442EF834" bold="true" box="[789,807,1923,1949]" pageId="97" pageNumber="98">F</emphasis>
|
|||
|
|
.
|
|||
|
|
<taxonomicName id="4C654D356C4D8A46FCCCF84444C0F834" authorityName="Pall-Gergely" authorityYear="2018" box="[820,969,1923,1949]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="97" pageNumber="98" phylum="Mollusca" rank="species" species="thompsoni" status="sp. nov.">
|
|||
|
|
<emphasis id="B911EAA46C4D8A46FCCCF84444C0F834" box="[820,969,1923,1949]" italics="true" pageId="97" pageNumber="98">P. thompsoni</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A22257DF6C4D8A46FC37F8434322F834" box="[975,1067,1923,1949]" pageId="97" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
, holotype (UF 77249). Scale bars: A, F (lower scale) = 10 mm. Images: T. Deli (A, F), T.-S. Liew (B–C) and B. Páll-Gergely (D–E).
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<subSubSection id="C37F653D6C4E8A45FF45FECB4451F842" pageId="98" pageNumber="99" type="materials_examined">
|
|||
|
|
<paragraph id="8BDA36B66C4E8A45FF45FECB46A3FE8C" blockId="98.[189,426,267,293]" box="[189,426,267,293]" pageId="98" pageNumber="99">
|
|||
|
|
<heading id="D09281DA6C4E8A45FF45FECB46A3FE8C" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="98" pageNumber="99" reason="3">
|
|||
|
|
<emphasis id="B911EAA46C4E8A45FF45FECB46A3FE8C" bold="true" box="[189,426,267,293]" pageId="98" pageNumber="99">Material examined</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<materialsCitation id="3B0D3CEB6C4E8A45FF12FEFB46BCFE08" ID-GBIF-Occurrence="2611489308" collectingDate="1998-10" collectionCode="HNHM" collectorName="J. and Ch. Hemmen" country="Malaysia" location="Kaki Bukit" pageId="98" pageNumber="99" specimenCode="HNHM 97469" specimenCount="1" stateProvince="Perlis" typeStatus="holotype">
|
|||
|
|
<paragraph id="8BDA36B66C4E8A45FF12FEFB4655FEFC" blockId="98.[189,1399,315,417]" box="[234,348,315,341]" pageId="98" pageNumber="99">
|
|||
|
|
<emphasis id="B911EAA46C4E8A45FF12FEFB4655FEFC" bold="true" box="[234,348,315,341]" pageId="98" pageNumber="99">
|
|||
|
|
<typeStatus id="54DE88146C4E8A45FF12FEFB4655FEFC" box="[234,348,315,341]" pageId="98" pageNumber="99" type="holotype">Holotype</typeStatus>
|
|||
|
|
</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDA36B66C4E8A45FF45FEA046BCFE08" blockId="98.[189,1399,315,417]" pageId="98" pageNumber="99">
|
|||
|
|
<collectingCountry id="F37276266C4E8A45FF45FEA04653FED3" box="[189,346,352,378]" name="Malaysia" pageId="98" pageNumber="99">MALAYSIA</collectingCountry>
|
|||
|
|
: 1 shell (D =
|
|||
|
|
<quantity id="4C9D9B536C4E8A45FE01FEA0456FFED2" box="[505,614,352,379]" metricMagnitude="-2" metricUnit="m" metricValue="1.81" pageId="98" pageNumber="99" unit="mm" value="18.1">18.1 mm</quantity>
|
|||
|
|
),
|
|||
|
|
<collectingRegion id="49A1F8546C4E8A45FD82FEA045C9FED3" box="[634,704,352,378]" country="Malaysia" name="Perlis" pageId="98" pageNumber="99">Perlis</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<location id="8EBA606D6C4E8A45FD34FEA04444FED3" LSID="urn:lsid:plazi:treatment:03CC87A06C4C8A42FD91FA194411FB28:8EBA606D6C4E8A45FD34FEA04444FED3" box="[716,845,352,378]" country="Malaysia" name="Kaki Bukit" pageId="98" pageNumber="99" stateProvince="Perlis">Kaki Bukit</location>
|
|||
|
|
(
|
|||
|
|
<locationDeviation id="9A682F946C4E8A45FCA5FEA044FEFED2" box="[861,1015,352,379]" location="Kangar" pageId="98" pageNumber="99">N of Kangar</locationDeviation>
|
|||
|
|
),
|
|||
|
|
<date id="FFDB10766C4E8A45FBF3FEA04388FED3" box="[1035,1153,352,378]" pageId="98" pageNumber="99" value="1998-10">
|
|||
|
|
<collectingDate id="EF9FE99E6C4E8A45FBF3FEA04388FED3" box="[1035,1153,352,378]" pageId="98" pageNumber="99" value="1998-10">Oct. 1998</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
,
|
|||
|
|
<collectorName id="269053606C4E8A45FB76FEA1427EFED2" box="[1166,1399,352,379]" pageId="98" pageNumber="99">J. and Ch. Hemmen</collectorName>
|
|||
|
|
leg. (
|
|||
|
|
<specimenCode id="DBC39ECD6C4E8A45FF0FFE4646A0FE08" box="[247,425,390,417]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="98" pageNumber="99">HNHM 97469</specimenCode>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</materialsCitation>
|
|||
|
|
<paragraph id="8BDA36B66C4E8A45FF12FE0A466EFE4D" blockId="98.[189,1399,458,1510]" box="[234,359,458,484]" pageId="98" pageNumber="99">
|
|||
|
|
<emphasis id="B911EAA46C4E8A45FF12FE0A466EFE4D" bold="true" box="[234,359,458,484]" pageId="98" pageNumber="99">
|
|||
|
|
<typeStatus id="54DE88146C4E8A45FF12FE0A466EFE4D" box="[234,359,458,484]" pageId="98" pageNumber="99" type="paratype">Paratypes</typeStatus>
|
|||
|
|
</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDA36B66C4E8A45FF45FE304541FA4F" blockId="98.[189,1399,458,1510]" pageId="98" pageNumber="99">
|
|||
|
|
<collectingCountry id="F37276266C4E8A45FF45FE304653FDA3" box="[189,346,496,522]" name="Malaysia" pageId="98" pageNumber="99">MALAYSIA</collectingCountry>
|
|||
|
|
: 11 shells, same data as for
|
|||
|
|
<typeStatus id="54DE88146C4E8A45FD5FFE304407FDA3" box="[679,782,496,522]" pageId="98" pageNumber="99" type="holotype">holotype</typeStatus>
|
|||
|
|
(coll.
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FCA4FE304488FDA3" box="[860,897,496,522]" pageId="98" pageNumber="99">JG</collectionCode>
|
|||
|
|
); 9 shells, same data as for
|
|||
|
|
<typeStatus id="54DE88146C4E8A45FB32FE304238FDA3" box="[1226,1329,496,522]" pageId="98" pageNumber="99" type="holotype">holotype</typeStatus>
|
|||
|
|
(coll.
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FF45FDD647F2FD99" box="[189,251,534,560]" pageId="98" pageNumber="99">PGB</collectionCode>
|
|||
|
|
); 20 shells, same data as for
|
|||
|
|
<typeStatus id="54DE88146C4E8A45FD98FDD645CEFD99" box="[608,711,534,560]" pageId="98" pageNumber="99" type="holotype">holotype</typeStatus>
|
|||
|
|
(coll.
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FCE1FDD7444CFD98" box="[793,837,535,561]" pageId="98" pageNumber="99">HE</collectionCode>
|
|||
|
|
); 2 shells, same data as for
|
|||
|
|
<typeStatus id="54DE88146C4E8A45FB64FDD6420AFD99" box="[1180,1283,534,560]" pageId="98" pageNumber="99" type="holotype">holotype</typeStatus>
|
|||
|
|
(
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FAEDFDD6427EFD99" box="[1301,1399,534,560]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="98" pageNumber="99" type="Museum">NHMW</collectionCode>
|
|||
|
|
108117); 5 shells,
|
|||
|
|
<collectingRegion id="49A1F8546C4E8A45FE6BFDFC46D0FDFF" box="[403,473,572,598]" country="Malaysia" name="Perlis" pageId="98" pageNumber="99">Perlis</collectingRegion>
|
|||
|
|
, Bukit Kubu (NE of Kuala
|
|||
|
|
<collectingRegion id="49A1F8546C4E8A45FCEEFDFC4454FDFE" box="[790,861,572,599]" country="Malaysia" name="Perlis" pageId="98" pageNumber="99">Perlis</collectingRegion>
|
|||
|
|
),
|
|||
|
|
<date id="FFDB10766C4E8A45FC89FDFC4306FDFF" box="[881,1039,572,599]" pageId="98" pageNumber="99" value="1998-03-21">21 Mar. 1998</date>
|
|||
|
|
, J. and Ch. Hemmen leg. (coll.
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FF45FDA247EBFDD5" box="[189,226,610,636]" pageId="98" pageNumber="99">JG</collectionCode>
|
|||
|
|
); 4 shells, same data as for preceding (coll.
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FD1EFDA2442DFDD5" box="[742,804,610,636]" pageId="98" pageNumber="99">PGB</collectionCode>
|
|||
|
|
); 1 shell,
|
|||
|
|
<collectingRegion id="49A1F8546C4E8A45FC6EFDA244D5FDD5" box="[918,988,610,636]" country="Malaysia" name="Perlis" pageId="98" pageNumber="99">Perlis</collectingRegion>
|
|||
|
|
, Wangga Tangga Valley, near Kaki Bukit,
|
|||
|
|
<date id="FFDB10766C4E8A45FEF7FD4846BBFD0B" box="[271,434,648,675]" pageId="98" pageNumber="99" value="1998-03-23">23 Mar. 1998</date>
|
|||
|
|
, J. and Ch. Hemmen leg. (coll.
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FCC9FD484469FD0B" box="[817,864,648,674]" pageId="98" pageNumber="99">HA</collectionCode>
|
|||
|
|
); 10 shells, same data as for preceding (coll.
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FF45FD6E47EBFD61" box="[189,226,686,712]" pageId="98" pageNumber="99">JG</collectionCode>
|
|||
|
|
); 9 shells, same data as for preceding (coll.
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FCF8FD6E4437FD61" box="[768,830,686,712]" pageId="98" pageNumber="99">PGB</collectionCode>
|
|||
|
|
); 3 shells, Nha-Krabi, south,
|
|||
|
|
<date id="FFDB10766C4E8A45FB5BFD6E4214FD61" box="[1187,1309,686,712]" pageId="98" pageNumber="99" value="2003-04">Apr. 2003</date>
|
|||
|
|
, Dieter Hörner leg. (coll.
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FE77FD1446BDFD47" box="[399,436,724,750]" pageId="98" pageNumber="99">JG</collectionCode>
|
|||
|
|
); 1 shell,
|
|||
|
|
<collectingRegion id="49A1F8546C4E8A45FDDFFD144564FD47" box="[551,621,724,750]" country="Malaysia" name="Perlis" pageId="98" pageNumber="99">Perlis</collectingRegion>
|
|||
|
|
, about
|
|||
|
|
<quantity id="4C9D9B536C4E8A45FD39FD144402FD47" box="[705,779,724,750]" metricMagnitude="4" metricUnit="m" metricValue="2.1" pageId="98" pageNumber="99" unit="km" value="21.0">21 km</quantity>
|
|||
|
|
N of Kangar, max.
|
|||
|
|
<quantity id="4C9D9B536C4E8A45FC0EFD14433BFD47" box="[1014,1074,724,750]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="98" pageNumber="99" unit="km" value="3.0">3 km</quantity>
|
|||
|
|
from the Thai border, Kaki Bukit, near Kampung Wang Tangga,
|
|||
|
|
<geoCoordinate id="EE5150716C4E8A45FD88FD3845FEFCBC" box="[624,759,760,789]" degrees="6" direction="north" minutes="38" orientation="latitude" pageId="98" pageNumber="99" precision="15" seconds="28" value="6.641111">6°38′28″ N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE5150716C4E8A45FCFBFD3844AAFCBC" box="[771,931,760,789]" degrees="100" direction="east" minutes="12" orientation="longitude" pageId="98" pageNumber="99" precision="15" seconds="08" value="100.20222">100°12′08″ E</geoCoordinate>
|
|||
|
|
, in leaf litter in a secondary forest, base of limestone hill,
|
|||
|
|
<date id="FFDB10766C4E8A45FE6EFCE04526FC92" box="[406,559,800,827]" pageId="98" pageNumber="99" value="1995-03-17">17 Mar.1995</date>
|
|||
|
|
, V. Kessner leg. (coll.
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FCC2FCE04471FC93" box="[826,888,800,826]" pageId="98" pageNumber="99">PGB</collectionCode>
|
|||
|
|
); 3 shells, same data as for preceding (coll.
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FF45FC8747F8FCC8" box="[189,241,839,865]" pageId="98" pageNumber="99">REI</collectionCode>
|
|||
|
|
); 2 shells,
|
|||
|
|
<collectingRegion id="49A1F8546C4E8A45FE8BFC8646B0FCC9" box="[371,441,838,864]" country="Malaysia" name="Perlis" pageId="98" pageNumber="99">Perlis</collectingRegion>
|
|||
|
|
, Titi Tinggi Mukim, Kaki Bukit, Gua Kelam, towards Wang Tangga, near fossil locality site, live/dead collected among disturbed limestone rocks and litters near exit of cave,
|
|||
|
|
<date id="FFDB10766C4E8A45FAE7FCAC47F3FC05" pageId="98" pageNumber="99" value="1996-04-26">26 Apr. 1996</date>
|
|||
|
|
, Chan Sow-Yan and Leo Nguang leg., ex Chan 1997 (
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FC8BFC5244A6FC05" box="[883,943,914,940]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="98" pageNumber="99">SMF</collectionCode>
|
|||
|
|
311305); 35 shells,
|
|||
|
|
<collectingRegion id="49A1F8546C4E8A45FB65FC5243EAFC05" box="[1181,1251,914,940]" country="Malaysia" name="Perlis" pageId="98" pageNumber="99">Perlis</collectingRegion>
|
|||
|
|
, Kaki Bukit, Gua Kelam,
|
|||
|
|
<geoCoordinate id="EE5150716C4E8A45FEADFC7646E1FC7A" box="[341,488,950,979]" degrees="06" direction="north" minutes="38.68" orientation="latitude" pageId="98" pageNumber="99" precision="9" value="6.6446667">06°38.68′ N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE5150716C4E8A45FE0DFC76459BFC7A" box="[501,658,950,979]" degrees="100" direction="east" minutes="12.23" orientation="longitude" pageId="98" pageNumber="99" precision="9" value="100.203835">100°12.23′ E</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<date id="FFDB10766C4E8A45FD58FC78442EFC7B" box="[672,807,952,978]" pageId="98" pageNumber="99" value="1999-07-09">9 Jul. 1999</date>
|
|||
|
|
, W. Maassen leg. (coll.
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FBBFFC79438BFC7A" box="[1095,1154,953,979]" pageId="98" pageNumber="99">WM</collectionCode>
|
|||
|
|
); 4 shells (preserved in ethanol: PL009, PL010, PL019, PL020 used for molecular study),
|
|||
|
|
<collectingRegion id="49A1F8546C4E8A45FBF9FC1E434EFC51" box="[1025,1095,990,1016]" country="Malaysia" name="Perlis" pageId="98" pageNumber="99">Perlis</collectingRegion>
|
|||
|
|
, Kaki Bukit, around Gua Kelam,
|
|||
|
|
<geoCoordinate id="EE5150716C4E8A45FEE1FBC2468DFBB6" box="[281,388,1026,1055]" degrees="06" direction="north" minutes="38" orientation="latitude" pageId="98" pageNumber="99" precision="925" value="6.633333">06°38′ N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE5150716C4E8A45FE77FBC2450DFBB6" box="[399,516,1026,1055]" degrees="100" direction="east" minutes="11" orientation="longitude" pageId="98" pageNumber="99" precision="925" value="100.183334">100°11′ E</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<quantity id="4C9D9B536C4E8A45FDE8FBC44542FBB6" box="[528,587,1028,1055]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="98" pageNumber="99" unit="m" value="50.0">50 m</quantity>
|
|||
|
|
a.s.l.,
|
|||
|
|
<date id="FFDB10766C4E8A45FD6DFBC44423FBB7" box="[661,810,1028,1055]" pageId="98" pageNumber="99" value="2013-01-11">11 Jan. 2013</date>
|
|||
|
|
, A. Hunyadi leg. (coll.
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FBCFFBC4436FFBB7" box="[1079,1126,1028,1054]" pageId="98" pageNumber="99">HA</collectionCode>
|
|||
|
|
);
|
|||
|
|
<specimenCount id="9D63FD3F6C4E8A45FB82FBC443FDFBB6" box="[1146,1268,1028,1055]" pageId="98" pageNumber="99" type="generic" typeStatus="paratype">1 paratype</specimenCount>
|
|||
|
|
(preserved in ethanol, anatomically examined), same data as for preceding (
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FC44FBEA4324FBED" box="[956,1069,1066,1092]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="98" pageNumber="99" type="Museum">NHMUK</collectionCode>
|
|||
|
|
20170153); 44 shells, West Malaysia,
|
|||
|
|
<collectingRegion id="49A1F8546C4E8A45FEC1FB904676FBC3" box="[313,383,1104,1130]" country="Malaysia" name="Perlis" pageId="98" pageNumber="99">Perlis</collectingRegion>
|
|||
|
|
, Wang Kelian State Park near Thai Border,
|
|||
|
|
<geoCoordinate id="EE5150716C4E8A45FC77FB8E432BFBC2" box="[911,1058,1102,1131]" degrees="06" direction="north" minutes="41.98" orientation="latitude" pageId="98" pageNumber="99" precision="9" value="6.6996665">06°41.98′ N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE5150716C4E8A45FBC8FB8E43C5FBC2" box="[1072,1228,1102,1131]" degrees="100" direction="east" minutes="11.80" orientation="longitude" pageId="98" pageNumber="99" precision="9" value="100.19666">100°11.80′ E</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<date id="FFDB10766C4E8A45FB23FB904278FBC3" box="[1243,1393,1104,1130]" pageId="98" pageNumber="99" value="1999-07-10">10 Jul. 1999</date>
|
|||
|
|
, W. Maassen leg. (coll.
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FE1AFBB74514FB38" box="[482,541,1143,1169]" pageId="98" pageNumber="99">WM</collectionCode>
|
|||
|
|
); 10 shells, West Malaysia,
|
|||
|
|
<collectingRegion id="49A1F8546C4E8A45FC8BFBB644B0FB39" box="[883,953,1142,1168]" country="Malaysia" name="Perlis" pageId="98" pageNumber="99">Perlis</collectingRegion>
|
|||
|
|
, limestone hill,
|
|||
|
|
<quantity id="4C9D9B536C4E8A45FB86FBB643C2FB39" box="[1150,1227,1142,1168]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="98" pageNumber="99" unit="km" value="30.0">30 km</quantity>
|
|||
|
|
N of Kangar,
|
|||
|
|
<geoCoordinate id="EE5150716C4E8A45FF45FB5A4646FB1E" box="[189,335,1178,1207]" degrees="06" direction="north" minutes="41.00" orientation="latitude" pageId="98" pageNumber="99" precision="9" value="6.6833334">06°41.00′ N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE5150716C4E8A45FEA4FB5A46FEFB1E" box="[348,503,1178,1207]" degrees="100" direction="east" minutes="11.21" orientation="longitude" pageId="98" pageNumber="99" precision="9" value="100.18684">100°11.21′ E</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<date id="FFDB10766C4E8A45FDFDFB5C4590FB1F" box="[517,665,1180,1206]" pageId="98" pageNumber="99" value="1999-07-10">10 Jul. 1999</date>
|
|||
|
|
, W. Maassen leg. (coll.
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FC4EFB5D44F8FB1E" box="[950,1009,1181,1207]" pageId="98" pageNumber="99">WM</collectionCode>
|
|||
|
|
); 5 shells, West Malaysia,
|
|||
|
|
<collectingRegion id="49A1F8546C4E8A45FAD4FB5C427BFB1F" box="[1324,1394,1180,1206]" country="Malaysia" name="Perlis" pageId="98" pageNumber="99">Perlis</collectingRegion>
|
|||
|
|
, Bukit Air Park, N of Kangar,
|
|||
|
|
<date id="FFDB10766C4E8A45FDEDFB0245AFFB75" box="[533,678,1218,1244]" pageId="98" pageNumber="99" value="1999-07-11">11 Jul. 1999</date>
|
|||
|
|
, W. Maassen leg. (coll.
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FC44FB0344FEFB74" box="[956,1015,1219,1245]" pageId="98" pageNumber="99">WM</collectionCode>
|
|||
|
|
); 4 shells, West Malaysia,
|
|||
|
|
<collectingRegion id="49A1F8546C4E8A45FAD4FB02427BFB75" box="[1324,1394,1218,1244]" country="Malaysia" name="Perlis" pageId="98" pageNumber="99">Perlis</collectingRegion>
|
|||
|
|
, Kaki Bukit, near Kampung Wang Tangga,
|
|||
|
|
<date id="FFDB10766C4E8A45FD3DFB284457FAAA" box="[709,862,1256,1283]" pageId="98" pageNumber="99" value="1995-07-11">11 Jul. 1995</date>
|
|||
|
|
, V. Kessner leg. (coll.
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FB8FFB2943BBFAAA" box="[1143,1202,1257,1283]" pageId="98" pageNumber="99">WM</collectionCode>
|
|||
|
|
); 1 shell,
|
|||
|
|
<collectingRegion id="49A1F8546C4E8A45FAD4FB28427BFAAB" box="[1324,1394,1256,1282]" country="Malaysia" name="Perlis" pageId="98" pageNumber="99">Perlis</collectingRegion>
|
|||
|
|
, Kaki Bukit, near Kampung Wang Tangga, in leaf litter, secondary forest at the base of limestone hill,
|
|||
|
|
<date id="FFDB10766C4E8A45FF45FAF44656FAE6" box="[189,351,1332,1359]" pageId="98" pageNumber="99" value="1995-03-17">17 Mar. 1995</date>
|
|||
|
|
, V. Kessner leg. (
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FDC9FAF44584FAE7" box="[561,653,1332,1358]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="98" pageNumber="99">HNHM</collectionCode>
|
|||
|
|
42264); 2 shells,
|
|||
|
|
<collectingRegion id="49A1F8546C4E8A45FC9CFAF444A3FAE7" box="[868,938,1332,1358]" country="Malaysia" name="Perlis" pageId="98" pageNumber="99">Perlis</collectingRegion>
|
|||
|
|
, Kaki Bukit, Kampunk, Wang Tangga, coll. L. Drimmer (
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FE59FA9A4509FADD" box="[417,512,1370,1396]" pageId="98" pageNumber="99">MMGY</collectionCode>
|
|||
|
|
69644); 1 shell,
|
|||
|
|
<collectingRegion id="49A1F8546C4E8A45FD22FA9A4429FADD" box="[730,800,1370,1396]" country="Malaysia" name="Perlis" pageId="98" pageNumber="99">Perlis</collectingRegion>
|
|||
|
|
, Kampung Tebing Tinggi,
|
|||
|
|
<quantity id="4C9D9B536C4E8A45FB94FA9A43CBFADD" box="[1132,1218,1370,1397]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="98" pageNumber="99" unit="km" value="3.5">3.5 km</quantity>
|
|||
|
|
W of Kangar,
|
|||
|
|
<geoCoordinate id="EE5150716C4E8A45FF45FABE4657FA32" box="[189,350,1406,1435]" degrees="06" direction="north" minutes="26.707" orientation="latitude" pageId="98" pageNumber="99" precision="1" value="6.4451165">06°26.707′ N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE5150716C4E8A45FE93FABE452CFA32" box="[363,549,1406,1435]" degrees="100" direction="east" minutes="09.1931" orientation="longitude" pageId="98" pageNumber="99" precision="1" value="100.15322">100°09.1931′ E</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<quantity id="4C9D9B536C4E8A45FDCAFA404566FA32" box="[562,623,1408,1435]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="98" pageNumber="99" unit="m" value="20.0">20 m</quantity>
|
|||
|
|
a.s.l.,
|
|||
|
|
<date id="FFDB10766C4E8A45FD46FA404451FA33" box="[702,856,1408,1435]" pageId="98" pageNumber="99" value="2013-01-12">12 Jan. 2013</date>
|
|||
|
|
, A. Hunyadi leg. (coll.
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FB97FA404397FA33" box="[1135,1182,1408,1434]" pageId="98" pageNumber="99">HA</collectionCode>
|
|||
|
|
); 46 shells,
|
|||
|
|
<collectingRegion id="49A1F8546C4E8A45FAD4FA40427BFA33" box="[1324,1394,1408,1434]" country="Malaysia" name="Perlis" pageId="98" pageNumber="99">Perlis</collectingRegion>
|
|||
|
|
, Kaki Bukit, around Gua Kelam, loc. 2013/14,
|
|||
|
|
<geoCoordinate id="EE5150716C4E8A45FD0BFA64449FFA68" box="[755,918,1444,1473]" degrees="06" direction="north" minutes="38.674" orientation="latitude" pageId="98" pageNumber="99" precision="1" value="6.6445665">06°38.674′ N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE5150716C4E8A45FC5DFA64435BFA68" box="[933,1106,1444,1473]" degrees="100" direction="east" minutes="12.170" orientation="longitude" pageId="98" pageNumber="99" precision="1" value="100.202835">100°12.170′ E</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<quantity id="4C9D9B536C4E8A45FB9AFA6643C4FA68" box="[1122,1229,1446,1473]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" pageId="98" pageNumber="99" unit="m" value="50.0" valueMax="55.0" valueMin="45.0">45–55 m</quantity>
|
|||
|
|
a.s.l.,
|
|||
|
|
<date id="FFDB10766C4E8A45FAD8FA6647F3FA4F" pageId="98" pageNumber="99" value="2013-01-13">13 Jan. 2013</date>
|
|||
|
|
, A. Hunyadi leg. (coll.
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FDF4FA0C4532FA4F" box="[524,571,1484,1510]" pageId="98" pageNumber="99">HA</collectionCode>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDA36B66C4E8A45FF45F9D146E0F89C" blockId="98.[189,1399,1552,1845]" pageId="98" pageNumber="99">
|
|||
|
|
<collectingCountry id="F37276266C4E8A45FF45F9D14650F982" box="[189,345,1553,1579]" name="Thailand" pageId="98" pageNumber="99">THAILAND</collectingCountry>
|
|||
|
|
: 3 shells,
|
|||
|
|
<collectingRegion id="49A1F8546C4E8A45FE33F9D14589F983" box="[459,640,1552,1579]" country="Thailand" name="Satun" pageId="98" pageNumber="99">Satun Province</collectingRegion>
|
|||
|
|
, Ban Rae, coll. Brandt, 1966 (
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FC19F9D14348F982" box="[993,1089,1553,1579]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="98" pageNumber="99">MNHN</collectionCode>
|
|||
|
|
); 4 shells,
|
|||
|
|
<collectingRegion id="49A1F8546C4E8A45FB45F9D1427BF983" box="[1213,1394,1552,1579]" country="Thailand" name="Satun" pageId="98" pageNumber="99">Satun Province</collectingRegion>
|
|||
|
|
, Ban Rae,
|
|||
|
|
<date id="FFDB10766C4E8A45FECAF9F746B7F9F9" box="[306,446,1590,1617]" pageId="98" pageNumber="99" value="1966-01-04">4 Jan. 1966</date>
|
|||
|
|
, Brandt leg. (
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FDA7F9F745ABF9F8" box="[607,674,1591,1617]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="98" pageNumber="99" type="Museum">ZMH</collectionCode>
|
|||
|
|
45914); 3 shells, same data as for preceding (
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FB3FF9F7420AF9F8" box="[1223,1283,1591,1617]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="98" pageNumber="99">SMF</collectionCode>
|
|||
|
|
207667); 2 shells, same data as for preceding except coll. Klemm (nr. 54168) (
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FC0BF99D435CF9DE" box="[1011,1109,1629,1655]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="98" pageNumber="99" type="Museum">NHMW</collectionCode>
|
|||
|
|
108223); 3 shells, same data as for preceding except coll. Oberwimmer (
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FD0FF9434450F934" box="[759,857,1667,1693]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="98" pageNumber="99" type="Museum">NHMW</collectionCode>
|
|||
|
|
108224); 4 shells, same data as for preceding except coll. Biggs (
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FE5AF969451AF96A" box="[418,531,1705,1731]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="98" pageNumber="99" type="Museum">NHMUK</collectionCode>
|
|||
|
|
20170159); 3 shells, same data as for preceding except coll. Brand 1966 (
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FF3DF90F4628F940" box="[197,289,1743,1769]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="98" pageNumber="99">MNHN</collectionCode>
|
|||
|
|
IM-
|
|||
|
|
<date id="FFDB10766C4E8A45FEA4F90E46D6F940" box="[348,479,1742,1769]" pageId="98" pageNumber="99" value="2012" valueMax="2519">2012-2519</date>
|
|||
|
|
);
|
|||
|
|
<specimenCount id="9D63FD3F6C4E8A45FE00F90F4571F940" box="[504,632,1743,1769]" pageId="98" pageNumber="99" type="generic" typeStatus="paratype">1 paratype</specimenCount>
|
|||
|
|
(preserved in ethanol),
|
|||
|
|
<collectingRegion id="49A1F8546C4E8A45FC5DF90F4356F941" box="[933,1119,1742,1769]" country="Thailand" name="Satun" pageId="98" pageNumber="99">Satun Province</collectingRegion>
|
|||
|
|
, Thale Ban Nat. Park, lowland rain forest,
|
|||
|
|
<geoCoordinate id="EE5150716C4E8A45FE5FF933450DF8A6" box="[423,516,1779,1807]" degrees="6" direction="north" minutes="42" orientation="latitude" pageId="98" pageNumber="99" precision="925" value="6.7">6°42′ N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE5150716C4E8A45FDF7F933458CF8A6" box="[527,645,1779,1807]" degrees="100" direction="east" minutes="10" orientation="longitude" pageId="98" pageNumber="99" precision="925" value="100.166664">100°10′ E</geoCoordinate>
|
|||
|
|
, ca
|
|||
|
|
<quantity id="4C9D9B536C4E8A45FD4AF93545F4F8A6" box="[690,765,1781,1807]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="98" pageNumber="99" unit="m" value="400.0">400 m</quantity>
|
|||
|
|
,
|
|||
|
|
<date id="FFDB10766C4E8A45FCF1F93544ABF8A6" box="[777,930,1781,1807]" pageId="98" pageNumber="99" value="1991-10-20">20 Oct. 1991</date>
|
|||
|
|
, M. Andersen, O. Martin and N. Scharff leg. (ZMUC-GAS-1834).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDA36B66C4E8A45FF12F89E46E8F8D1" blockId="98.[189,1406,1886,2027]" box="[234,481,1886,1912]" pageId="98" pageNumber="99">
|
|||
|
|
<emphasis id="B911EAA46C4E8A45FF12F89E46E8F8D1" bold="true" box="[234,481,1886,1912]" pageId="98" pageNumber="99">Additional material</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDA36B66C4E8A45FF45F8454451F842" blockId="98.[189,1406,1886,2027]" pageId="98" pageNumber="99">
|
|||
|
|
<collectingCountry id="F37276266C4E8A45FF45F8454653F836" box="[189,346,1925,1951]" name="Malaysia" pageId="98" pageNumber="99">MALAYSIA</collectingCountry>
|
|||
|
|
:
|
|||
|
|
<specimenCount id="9D63FD3F6C4E8A45FE9CF84546ADF836" box="[356,420,1925,1951]" pageId="98" pageNumber="99" type="generic">1 juv.</specimenCount>
|
|||
|
|
shell,
|
|||
|
|
<collectingRegion id="49A1F8546C4E8A45FE08F845453FF836" box="[496,566,1925,1951]" country="Malaysia" name="Perlis" pageId="98" pageNumber="99">Perlis</collectingRegion>
|
|||
|
|
, about
|
|||
|
|
<quantity id="4C9D9B536C4E8A45FD70F84545D8F836" box="[648,721,1925,1951]" metricMagnitude="4" metricUnit="m" metricValue="2.1" pageId="98" pageNumber="99" unit="km" value="21.0">21 km</quantity>
|
|||
|
|
N of Kangar, max.
|
|||
|
|
<quantity id="4C9D9B536C4E8A45FC4EF84544F9F836" box="[950,1008,1925,1951]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="98" pageNumber="99" unit="km" value="3.0">3 km</quantity>
|
|||
|
|
from the Thai border, Kaki Bukit near Kampung Wang Tangga,
|
|||
|
|
<geoCoordinate id="EE5150716C4E8A45FDD4F86945BFF86C" box="[556,694,1961,1989]" degrees="6" direction="north" minutes="38" orientation="latitude" pageId="98" pageNumber="99" precision="15" seconds="28" value="6.641111">6°38′28″ N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE5150716C4E8A45FD3DF8694460F86C" box="[709,873,1961,1989]" degrees="100" direction="east" minutes="12" orientation="longitude" pageId="98" pageNumber="99" precision="15" seconds="08" value="100.20222">100°12′08″ E</geoCoordinate>
|
|||
|
|
, in leaf litter in a secondary forest, base of limestone hill,
|
|||
|
|
<date id="FFDB10766C4E8A45FE94F811450CF842" box="[364,517,2000,2027]" pageId="98" pageNumber="99" value="1995-03-17">17 Mar.1995</date>
|
|||
|
|
, V. Kessner leg. (coll.
|
|||
|
|
<collectionCode id="ED74AE736C4E8A45FCF5F8114442F842" box="[781,843,2001,2027]" pageId="98" pageNumber="99">PGB</collectionCode>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C37F653D6C4F8A43FF45FECB4501FD5E" lastPageId="100" lastPageNumber="101" pageId="99" pageNumber="100" type="description">
|
|||
|
|
<paragraph id="8BDA36B66C4F8A44FF45FECB4647FE8C" blockId="99.[189,334,267,293]" box="[189,334,267,293]" pageId="99" pageNumber="100">
|
|||
|
|
<heading id="D09281DA6C4F8A44FF45FECB4647FE8C" bold="true" box="[189,334,267,293]" fontSize="11" level="3" pageId="99" pageNumber="100" reason="3">
|
|||
|
|
<emphasis id="B911EAA46C4F8A44FF45FECB4647FE8C" bold="true" box="[189,334,267,293]" pageId="99" pageNumber="100">Description</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDA36B66C4F8A44FF45FEFD4600FDC0" blockId="99.[189,1399,317,617]" pageId="99" pageNumber="100">SHELL. Sinistral, chocolate brown (or yellowish, if weathered), nearly flat, strongly depressed conical or flat with protoconch slightly protruding above dorsal surface; protoconch consists of 3.25–3.5 whorls, very finely granular, with inconspicuous growth lines on last 1.5–2 whorls; ventral surface of protoconch roughly ribbed; teleoconch with a reticulated surface consisting of very fine, dense ribs and widely-spaced spiral lines; periostracum of fresh specimens with deciduous folds on upper keel of body whorl; whorls 6–6.5, separated by a relatively shallow suture; aperture rounded; peristome thickened and expanded; parietal callus sharp; apertural fold usually short, white and never in contact with parietal callus.</paragraph>
|
|||
|
|
<paragraph id="8BDA36B66C4F8A44FF45FD55443CFC69" blockId="99.[189,1399,660,961]" pageId="99" pageNumber="100">Four shells were opened. Lambda-complex compete; left leg and upper branch relatively short; middle portion of right leg with a short horizontal, attached plica, which is rarely in contact with main plica; short lower plica situated below lambda-complex; in some specimens main plica connected to apertural fold, whereas in others they are free (this trait is probably age-dependent). Palatal wall with six plicae; first three straight with sinuate (S-shaped) posterior ends; last plica slightly curved with its convex side pointing upwards; additional short plica visible above posterior end of last plica; fourth and fifth plicae vertical and fused together, forming a single, vertical plate, which is usually bifurcated below and possesses 1–2 additional horizontal plicae posteriorly.</paragraph>
|
|||
|
|
<paragraph id="8BDA36B66C4F8A44FF45FC2C448CFBAF" blockId="99.[189,901,1003,1030]" box="[189,901,1003,1030]" pageId="99" pageNumber="100">MEASUREMENTS (in mm). D = 17.6–19.0, H = 6.6–7.2 (n = 4).</paragraph>
|
|||
|
|
<caption id="DF1A663E6C4F8A44FF45F84A43CAF842" ID-DOI="http://doi.org/10.5281/zenodo.3817791" ID-Zenodo-Dep="3817791" httpUri="https://zenodo.org/record/3817791/files/figure.png" pageId="99" pageNumber="100" startId="99.[189,232,1930,1956]" targetBox="[206,1399,1077,1890]" targetPageId="99">
|
|||
|
|
<paragraph id="8BDA36B66C4F8A44FF45F84A43CAF842" blockId="99.[189,1399,1930,2027]" pageId="99" pageNumber="100">
|
|||
|
|
<emphasis id="B911EAA46C4F8A44FF45F84A4612F80D" bold="true" box="[189,283,1930,1956]" pageId="99" pageNumber="100">Fig. 45.</emphasis>
|
|||
|
|
Genitalia of
|
|||
|
|
<taxonomicName id="4C654D356C4F8A44FE4FF84A45BFF80D" authorityName="Pall-Gergely" authorityYear="2018" box="[439,694,1930,1956]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="99" pageNumber="100" phylum="Mollusca" rank="species" species="malayana" status="sp. nov.">
|
|||
|
|
<emphasis id="B911EAA46C4F8A44FE4FF84A45BFF80D" box="[439,694,1930,1956]" italics="true" pageId="99" pageNumber="100">Plectopylis malayana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A22257DF6C4F8A44FD46F84B4415F80C" box="[702,796,1931,1957]" pageId="99" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
, PL010 (NHMUK 20170153).
|
|||
|
|
<emphasis id="B911EAA46C4F8A44FB77F84A43ADF80D" bold="true" box="[1167,1188,1930,1956]" pageId="99" pageNumber="100">A</emphasis>
|
|||
|
|
. Whole genitalia.
|
|||
|
|
<emphasis id="B911EAA46C4F8A44FF45F86E47FAF861" bold="true" box="[189,243,1966,1992]" pageId="99" pageNumber="100">B–E</emphasis>
|
|||
|
|
. Inner structure of genital organs (B = penis, C = epiphallus, D = vagina, E = diverticulum). Scale bars: A = 5 mm; B–E = 1 mm. For abbreviations see Fig. 2. All photos: B. Páll-Gergely.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BDA36B66C488A43FF45FECB437EFDFF" blockId="100.[189,1399,267,599]" pageId="100" pageNumber="101">
|
|||
|
|
CHARACTERS OF THE GENITAL SYSTEM. One specimen was examined. Locality information:
|
|||
|
|
<collectingCountry id="F37276266C488A43FAFCFECB427AFE8C" box="[1284,1395,267,293]" name="Malaysia" pageId="100" pageNumber="101">Malaysia</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<collectingRegion id="49A1F8546C488A43FF45FEF7460AFEF8" box="[189,259,311,337]" country="Malaysia" name="Perlis" pageId="100" pageNumber="101">Perlis</collectingRegion>
|
|||
|
|
, Kaki Bukit, around Gua Kelam,
|
|||
|
|
<geoCoordinate id="EE5150716C488A43FD68FEF545F4FEF8" box="[656,765,309,337]" degrees="06" direction="north" minutes="38" orientation="latitude" pageId="100" pageNumber="101" precision="925" value="6.633333">06°38′ N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE5150716C488A43FCF1FEF54489FEF8" box="[777,896,309,337]" degrees="100" direction="east" minutes="11" orientation="longitude" pageId="100" pageNumber="101" precision="925" value="100.183334">100°11′ E</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<quantity id="4C9D9B536C488A43FC75FEF744C0FEF8" box="[909,969,311,338]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="100" pageNumber="101" unit="m" value="50.0">50 m</quantity>
|
|||
|
|
a.s.l., A. Hunyadi leg.,
|
|||
|
|
<date id="FFDB10766C488A43FB19FEF7427EFEF8" box="[1249,1399,311,337]" pageId="100" pageNumber="101" value="2013-01-11">11 Jan. 2013</date>
|
|||
|
|
(NHMUK 20170153). Left ommatophoral retractor runs between penis and vagina; penis relatively long and slender, internally with parallel, serrate folds; epiphallus and vas deferens attached to penis with weak muscle fibres; no penial caecum observed; retractor muscle thick and short, inserts on proximal part of penis; vagina approx. as long as penis, but thicker; bursa copulatrix with a very long, slender stalk, bursa elongate ovoid; diverticulum short, blunt, ovoid; four developing embryos were found in uterus, the most distal one was the most well-developed, with a shell of 3 whorls.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDA36B66C488A43FF45FD464501FD5E" blockId="100.[189,1399,646,759]" pageId="100" pageNumber="101">RADULA. Centrals with small, triangular cusps; endocones of laterals rhomboid; ectocones of first laterals larger than centrals; ectocones of marginals undivided, endocones of marginals mostly undivided, or divided by shallow incision.</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<caption id="DF1A663E6C488A43FF45F84A45A8F842" ID-DOI="http://doi.org/10.5281/zenodo.3817793" ID-Zenodo-Dep="3817793" httpUri="https://zenodo.org/record/3817793/files/figure.png" pageId="100" pageNumber="101" startId="100.[189,232,1930,1956]" targetBox="[189,1398,847,1889]" targetPageId="100">
|
|||
|
|
<paragraph id="8BDA36B66C488A43FF45F84A45A8F842" blockId="100.[189,1399,1930,2028]" pageId="100" pageNumber="101">
|
|||
|
|
<emphasis id="B911EAA46C488A43FF45F84A4610F80D" bold="true" box="[189,281,1930,1956]" pageId="100" pageNumber="101">Fig. 46.</emphasis>
|
|||
|
|
Radula of
|
|||
|
|
<taxonomicName id="4C654D356C488A43FE6FF84A45C9F80D" authority="Benson, 1860" authorityName="Benson" authorityYear="1860" box="[407,704,1930,1957]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="100" pageNumber="101" phylum="Mollusca" rank="genus">
|
|||
|
|
<emphasis id="B911EAA46C488A43FE6FF84A4510F80D" box="[407,537,1930,1956]" italics="true" pageId="100" pageNumber="101">Plectopylis</emphasis>
|
|||
|
|
<bibRefCitation id="EFF44B476C488A43FDD8F84B45C9F80D" author="Benson W. H." box="[544,704,1930,1957]" pageId="100" pageNumber="101" pagination="243 - 247" refId="ref51737" refString="Benson W. H. 1860. Notes on Plectopylis, a group a Helicidae distinguished by several internal plicate Ephiphragms; with the Characters of a new Species. The Annals and Magazine of Natural History 3 (5): 243 - 247. Available from https: // biodiversitylibrary. org / page / 18658580 [accessed 30 Jul. 2018]." type="journal article" year="1860">Benson, 1860</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species.
|
|||
|
|
<emphasis id="B911EAA46C488A43FCD1F84A4468F80D" bold="true" box="[809,865,1930,1956]" pageId="100" pageNumber="101">A, C</emphasis>
|
|||
|
|
. Central and first lateral teeth.
|
|||
|
|
<emphasis id="B911EAA46C488A43FB3FF84A43F4F80D" bold="true" box="[1223,1277,1930,1956]" pageId="100" pageNumber="101">B, D</emphasis>
|
|||
|
|
. Marginal teeth.
|
|||
|
|
<emphasis id="B911EAA46C488A43FEFBF86D4633F86E" bold="true" box="[259,314,1965,1991]" pageId="100" pageNumber="101">A–B</emphasis>
|
|||
|
|
.
|
|||
|
|
<taxonomicName id="4C654D356C488A43FEB0F86D454CF86E" authorityName="Pall-Gergely" authorityYear="2018" box="[328,581,1965,1991]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="100" pageNumber="101" phylum="Mollusca" rank="species" species="malayana" status="sp. nov.">
|
|||
|
|
<emphasis id="B911EAA46C488A43FEB0F86D454CF86E" box="[328,581,1965,1991]" italics="true" pageId="100" pageNumber="101">Plectopylis malayana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A22257DF6C488A43FDB3F86E45ADF861" box="[587,676,1966,1992]" pageId="100" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
(loc. 2013/14, coll. HA).
|
|||
|
|
<emphasis id="B911EAA46C488A43FC36F86D430EF86E" bold="true" box="[974,1031,1965,1991]" pageId="100" pageNumber="101">C–D</emphasis>
|
|||
|
|
.
|
|||
|
|
<taxonomicName id="4C654D356C488A43FBECF86D421EF86E" authorityName="Pall-Gergely" authorityYear="2018" box="[1044,1303,1965,1991]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="100" pageNumber="101" phylum="Mollusca" rank="species" species="thompsoni" status="sp. nov.">
|
|||
|
|
<emphasis id="B911EAA46C488A43FBECF86D421EF86E" box="[1044,1303,1965,1991]" italics="true" pageId="100" pageNumber="101">Plectopylis thompsoni</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A22257DF6C488A43FAE5F86E427EF861" box="[1309,1399,1966,1992]" pageId="100" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
(UF 76482). All images: B. Páll-Gergely.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<subSubSection id="C37F653D6C498A42FF45FECB47F5FDE1" pageId="101" pageNumber="102" type="diagnosis">
|
|||
|
|
<paragraph id="8BDA36B66C498A42FF45FECB46C0FE8C" blockId="101.[189,457,267,293]" box="[189,457,267,293]" pageId="101" pageNumber="102">
|
|||
|
|
<heading id="D09281DA6C498A42FF45FECB46C0FE8C" bold="true" box="[189,457,267,293]" fontSize="11" level="3" pageId="101" pageNumber="102" reason="3">
|
|||
|
|
<emphasis id="B911EAA46C498A42FF45FECB46C0FE8C" bold="true" box="[189,457,267,293]" pageId="101" pageNumber="102">Differential diagnosis</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDA36B66C498A42FF45FEF947F5FDE1" blockId="101.[189,1399,313,584]" pageId="101" pageNumber="102">
|
|||
|
|
<taxonomicName id="4C654D356C498A42FF45FEF946B2FEFA" authorityName="Pall-Gergely" authorityYear="2018" box="[189,443,313,339]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="101" pageNumber="102" phylum="Mollusca" rank="species" species="malayana" status="sp. nov.">
|
|||
|
|
<emphasis id="B911EAA46C498A42FF45FEF946B2FEFA" box="[189,443,313,339]" italics="true" pageId="101" pageNumber="102">Plectopylis malayana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A22257DF6C498A42FE3BFEFA4516FEFD" box="[451,543,314,340]" pageId="101" pageNumber="102" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
differs from
|
|||
|
|
<taxonomicName id="4C654D356C498A42FD44FEFA4423FEFA" authorityName="Gude" authorityYear="1898" box="[700,810,314,339]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="101" pageNumber="102" phylum="Mollusca" rank="species" species="cairnsi">
|
|||
|
|
<emphasis id="B911EAA46C498A42FD44FEFA4423FEFA" box="[700,810,314,339]" italics="true" pageId="101" pageNumber="102">P. cairnsi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by its darker shell, thinner lip, less rounded body whorl, several additional plicae on the palatal wall, shorter left leg, and shorter upper branch of the lambda-complex. Furthermore, the lambda-complex of
|
|||
|
|
<taxonomicName id="4C654D356C498A42FC9DFE4044D0FE30" authorityName="Gude" authorityYear="1898" box="[869,985,384,409]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="101" pageNumber="102" phylum="Mollusca" rank="species" species="cairnsi">
|
|||
|
|
<emphasis id="B911EAA46C498A42FC9DFE4044D0FE30" box="[869,985,384,409]" italics="true" pageId="101" pageNumber="102">P. cairnsi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is never fused to the main plica (sometimes fused in
|
|||
|
|
<taxonomicName id="4C654D356C498A42FE49FE63454BFE15" authorityName="Pall-Gergely" authorityYear="2018" box="[433,578,418,444]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="101" pageNumber="102" phylum="Mollusca" rank="species" species="malayana" status="sp. nov.">
|
|||
|
|
<emphasis id="B911EAA46C498A42FE49FE63454BFE15" box="[433,578,418,444]" italics="true" pageId="101" pageNumber="102">P. malayana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A22257DF6C498A42FDB1FE6345AEFE14" box="[585,679,419,445]" pageId="101" pageNumber="102" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
), and
|
|||
|
|
<taxonomicName id="4C654D356C498A42FD16FE634477FE15" authorityName="Pall-Gergely" authorityYear="2018" box="[750,894,418,444]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="101" pageNumber="102" phylum="Mollusca" rank="species" species="malayana" status="sp. nov.">
|
|||
|
|
<emphasis id="B911EAA46C498A42FD16FE634477FE15" box="[750,894,418,444]" italics="true" pageId="101" pageNumber="102">P. malayana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A22257DF6C498A42FC7EFE6344E9FE14" box="[902,992,419,445]" pageId="101" pageNumber="102" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
has a small fold that is attached to the central part of the right leg.
|
|||
|
|
<taxonomicName id="4C654D356C498A42FDD5FE054418FE76" authorityName=", Gude" authorityYear="1914" box="[557,785,453,479]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="101" pageNumber="102" phylum="Mollusca" rank="species" species="bensoni">
|
|||
|
|
<emphasis id="B911EAA46C498A42FDD5FE054418FE76" box="[557,785,453,479]" italics="true" pageId="101" pageNumber="102">Plectopylis bensoni</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is usually larger than
|
|||
|
|
<taxonomicName id="4C654D356C498A42FBEBFE0643AAFE76" authorityName="Pall-Gergely" authorityYear="2018" box="[1043,1187,453,479]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="101" pageNumber="102" phylum="Mollusca" rank="species" species="malayana" status="sp. nov.">
|
|||
|
|
<emphasis id="B911EAA46C498A42FBEBFE0643AAFE76" box="[1043,1187,453,479]" italics="true" pageId="101" pageNumber="102">P. malayana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A22257DF6C498A42FB52FE06420DFE49" box="[1194,1284,454,480]" pageId="101" pageNumber="102" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
and has a rounded body whorl, whereas the body whorl of
|
|||
|
|
<taxonomicName id="4C654D356C498A42FD0FFE29448EFDAB" authorityName="Pall-Gergely" authorityYear="2018" box="[759,903,488,514]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="101" pageNumber="102" phylum="Mollusca" rank="species" species="malayana" status="sp. nov.">
|
|||
|
|
<emphasis id="B911EAA46C498A42FD0FFE29448EFDAB" box="[759,903,488,514]" italics="true" pageId="101" pageNumber="102">P. malayana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A22257DF6C498A42FC76FE2944E1FDAA" box="[910,1000,489,515]" pageId="101" pageNumber="102" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
is angled. Moreover,
|
|||
|
|
<taxonomicName id="4C654D356C498A42FB1EFE29427EFDAB" authorityName="Pall-Gergely" authorityYear="2018" box="[1254,1399,488,514]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="101" pageNumber="102" phylum="Mollusca" rank="species" species="malayana" status="sp. nov.">
|
|||
|
|
<emphasis id="B911EAA46C498A42FB1EFE29427EFDAB" box="[1254,1399,488,514]" italics="true" pageId="101" pageNumber="102">P. malayana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A22257DF6C498A42FF45FDCC461EFD8F" box="[189,279,524,550]" pageId="101" pageNumber="102" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
has a finely reticulated shell surface, which is nearly smooth in
|
|||
|
|
<taxonomicName id="4C654D356C498A42FC07FDCC437FFD8C" authorityName=", Gude" authorityYear="1914" box="[1023,1142,523,549]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="101" pageNumber="102" phylum="Mollusca" rank="species" species="bensoni">
|
|||
|
|
<emphasis id="B911EAA46C498A42FC07FDCC437FFD8C" box="[1023,1142,523,549]" italics="true" pageId="101" pageNumber="102">P. bensoni</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
with irregular growth lines.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C37F653D6C498A42FF45FDAF437BFD72" pageId="101" pageNumber="102" type="distribution">
|
|||
|
|
<paragraph id="8BDA36B66C498A42FF45FDAF465FFD20" blockId="101.[189,342,623,649]" box="[189,342,623,649]" pageId="101" pageNumber="102">
|
|||
|
|
<heading id="D09281DA6C498A42FF45FDAF465FFD20" bold="true" box="[189,342,623,649]" fontSize="11" level="3" pageId="101" pageNumber="102" reason="3">
|
|||
|
|
<emphasis id="B911EAA46C498A42FF45FDAF465FFD20" bold="true" box="[189,342,623,649]" pageId="101" pageNumber="102">Distribution</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDA36B66C498A42FF45FD5E437BFD72" blockId="101.[189,1399,670,731]" pageId="101" pageNumber="102">
|
|||
|
|
<taxonomicName id="4C654D356C498A42FF45FD5E46B0FD11" authorityName="Pall-Gergely" authorityYear="2018" box="[189,441,670,696]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="101" pageNumber="102" phylum="Mollusca" rank="species" species="malayana" status="sp. nov.">
|
|||
|
|
<emphasis id="B911EAA46C498A42FF45FD5E46B0FD11" box="[189,441,670,696]" italics="true" pageId="101" pageNumber="102">Plectopylis malayana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A22257DF6C498A42FE47FD5F4511FD10" box="[447,536,671,697]" pageId="101" pageNumber="102" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
is known from
|
|||
|
|
<collectingRegion id="49A1F8546C498A42FD33FD5E4475FD11" box="[715,892,670,696]" country="Malaysia" name="Perlis" pageId="101" pageNumber="102">Perlis Province</collectingRegion>
|
|||
|
|
(near the Thai border) in northern
|
|||
|
|
<collectingCountry id="F37276266C498A42FAF3FD5E427EFD11" box="[1291,1399,670,696]" name="Malaysia" pageId="101" pageNumber="102">Malaysia</collectingCountry>
|
|||
|
|
and the
|
|||
|
|
<collectingRegion id="49A1F8546C498A42FEE2FD0146C5FD72" box="[282,460,705,731]" country="Thailand" name="Satun" pageId="101" pageNumber="102">Satun Province</collectingRegion>
|
|||
|
|
in southern
|
|||
|
|
<collectingCountry id="F37276266C498A42FDA6FD0145CFFD72" box="[606,710,705,731]" name="Thailand" pageId="101" pageNumber="102">Thailand</collectingCountry>
|
|||
|
|
only. See also Remarks and
|
|||
|
|
<figureCitation id="135E2A336C498A42FBE0FD014367FD72" box="[1048,1134,705,731]" captionStart="Fig" captionStartId="87.[189,232,1930,1956]" captionTargetBox="[189,1398,1044,1890]" captionTargetId="figure@87.[189,1398,1044,1890]" captionTargetPageId="87" captionText="Fig. 40. Distribution of species of Plectopylis Benson, 1860 in Southern Thailand and northern Malaysia. Legends: empty triangle = P. thompsoni sp. nov.; square = P. cairnsi Gude, 1898; filled triangle = sympatric locality of P. thompsoni sp. nov. and P. cairnsi; circle = P. malayana sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3817781" httpUri="https://zenodo.org/record/3817781/files/figure.png" pageId="101" pageNumber="102">Fig. 40</figureCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C37F653D6C498A42FF45FCC34411FB28" pageId="101" pageNumber="102" type="discussion">
|
|||
|
|
<paragraph id="8BDA36B66C498A42FF45FCC34626FCB4" blockId="101.[189,303,771,797]" box="[189,303,771,797]" pageId="101" pageNumber="102">
|
|||
|
|
<heading id="D09281DA6C498A42FF45FCC34626FCB4" bold="true" box="[189,303,771,797]" fontSize="11" level="3" pageId="101" pageNumber="102" reason="3">
|
|||
|
|
<emphasis id="B911EAA46C498A42FF45FCC34626FCB4" bold="true" box="[189,303,771,797]" pageId="101" pageNumber="102">Remarks</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDA36B66C498A42FF45FCF143CDFC53" blockId="101.[189,1399,817,1018]" pageId="101" pageNumber="102">
|
|||
|
|
The species collected from northern
|
|||
|
|
<collectingCountry id="F37276266C498A42FD91FCF145DCFCE2" box="[617,725,817,843]" name="Malaysia" pageId="101" pageNumber="102">Malaysia</collectingCountry>
|
|||
|
|
was mentioned as
|
|||
|
|
<taxonomicName id="4C654D356C498A42FC4BFCF14251FCE5" authority="(Pfeiffer, 1867)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1867" box="[947,1368,817,844]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="101" pageNumber="102" phylum="Mollusca" rank="species" species="revoluta">
|
|||
|
|
<emphasis id="B911EAA46C498A42FC4BFCF14394FCE2" box="[947,1181,817,843]" italics="true" pageId="101" pageNumber="102">Plectopylis revoluta</emphasis>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation id="EFF44B476C498A42FB54FCF14259FCE5" author="Pfeiffer L." box="[1196,1360,817,844]" pageId="101" pageNumber="102" pagination="64" refId="ref55566" refString="Pfeiffer L. 1867. Eine neue Helix. Malakologische Blatter 14: 64. Available from https: // biodiversitylibrary. org / page / 15855865 [accessed 30 Jul. 2018]." type="journal article" year="1867">Pfeiffer, 1867</bibRefCitation>
|
|||
|
|
)
|
|||
|
|
</taxonomicName>
|
|||
|
|
in the literature (see above).
|
|||
|
|
<taxonomicName id="4C654D356C498A42FE0FFC9445EDFCC7" baseAuthorityName="Pfeiffer" baseAuthorityYear="1867" box="[503,740,852,878]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="101" pageNumber="102" phylum="Mollusca" rank="species" species="revoluta">
|
|||
|
|
<emphasis id="B911EAA46C498A42FE0FFC9445EDFCC7" box="[503,740,852,878]" italics="true" pageId="101" pageNumber="102">Plectopylis revoluta</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, however, was originally described from the Andaman Islands.
|
|||
|
|
<bibRefCitation id="EFF44B476C498A42FEE5FCB746B6FC38" author="Gude G. K." box="[285,447,887,913]" pageId="101" pageNumber="102" refId="ref54417" refString="Gude G. K. 1914 b. The Fauna of British India including Ceylon and Burma. Mollusca. ‾ II. (Trochomorphidae- - Janellidae). Taylor & Francis, London. Available from https: // biodiversitylibrary. org / page / 12910698 [accessed 30 Jul. 2018]." type="url" year="1914">Gude (1914b)</bibRefCitation>
|
|||
|
|
discussed the taxonomic status of
|
|||
|
|
<taxonomicName id="4C654D356C498A42FCB3FCB844CEFC38" baseAuthorityName="Pfeiffer" baseAuthorityYear="1867" box="[843,967,887,913]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="101" pageNumber="102" phylum="Mollusca" rank="species" species="revoluta">
|
|||
|
|
<emphasis id="B911EAA46C498A42FCB3FCB844CEFC38" box="[843,967,887,913]" italics="true" pageId="101" pageNumber="102">P. revoluta</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in detail, and concluded that
|
|||
|
|
<taxonomicName id="4C654D356C498A42FAEEFCB7427EFC38" baseAuthorityName="Pfeiffer" baseAuthorityYear="1867" box="[1302,1399,887,913]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="101" pageNumber="102" phylum="Mollusca" rank="species" species="revoluta">
|
|||
|
|
<emphasis id="B911EAA46C498A42FAEEFCB7427EFC38" box="[1302,1399,887,913]" italics="true" pageId="101" pageNumber="102">revoluta</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is possibly a synonym of
|
|||
|
|
<taxonomicName id="4C654D356C498A42FE1AFC5B4563FC1D" baseAuthorityName="Benson" baseAuthorityYear="1859" box="[482,618,922,948]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="101" pageNumber="102" phylum="Mollusca" rank="species" species="cyclaspis">
|
|||
|
|
<emphasis id="B911EAA46C498A42FE1AFC5B4563FC1D" box="[482,618,922,948]" italics="true" pageId="101" pageNumber="102">P. cyclaspis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. The absence of the
|
|||
|
|
<typeStatus id="54DE88146C498A42FCAEFC5A449DFC1D" box="[854,916,922,948]" pageId="101" pageNumber="102">types</typeStatus>
|
|||
|
|
of
|
|||
|
|
<taxonomicName id="4C654D356C498A42FC41FC5B433CFC1D" baseAuthorityName="Pfeiffer" baseAuthorityYear="1867" box="[953,1077,922,948]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="101" pageNumber="102" phylum="Mollusca" rank="species" species="revoluta">
|
|||
|
|
<emphasis id="B911EAA46C498A42FC41FC5B433CFC1D" box="[953,1077,922,948]" italics="true" pageId="101" pageNumber="102">P. revoluta</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
does not allow clarification of this question, but the original description of
|
|||
|
|
<taxonomicName id="4C654D356C498A42FD0CFC7E447AFC7E" baseAuthorityName="Pfeiffer" baseAuthorityYear="1867" box="[756,883,957,983]" class="Gastropoda" family="Plectopylidae" genus="Plectopylis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="101" pageNumber="102" phylum="Mollusca" rank="species" species="revoluta">
|
|||
|
|
<emphasis id="B911EAA46C498A42FD0CFC7E447AFC7E" box="[756,883,957,983]" italics="true" pageId="101" pageNumber="102">P. revoluta</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(e.g., body whorl acutely keeled) certainly does not fit the Malaysian species. Therefore, a new name must be given to this species.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDA36B66C498A42FF45FBE14411FB28" blockId="101.[189,1398,1057,1154]" pageId="101" pageNumber="102">
|
|||
|
|
The sample collected by Brandt in 1966 was distributed to several museums (ZMH, NHMW, SMF, MNHN). According to the label the locality is “
|
|||
|
|
<collectingRegion id="49A1F8546C498A42FD1AFB844422FBF7" box="[738,811,1092,1118]" country="Thailand" name="Satun" pageId="101" pageNumber="102">Satun</collectingRegion>
|
|||
|
|
: Ban Rae”. The locality name “Ban Rae” probably refers to Ban Rae Kot, northwest of Mueang
|
|||
|
|
<collectingRegion id="49A1F8546C498A42FD36FBA7441CFB28" box="[718,789,1127,1153]" country="Thailand" name="Satun" pageId="101" pageNumber="102">Satun</collectingRegion>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|