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<document id="5FAFF5ABF307BCFA3627247634A91D3B" ID-CLB-Dataset="7895" ID-DOI="10.11646/zootaxa.4938.2.5" ID-GBIF-Dataset="22508435-bb1f-472a-b8ac-c29726076ea4" ID-ISSN="1175-5326" ID-Zenodo-Dep="4563882" ID-ZooBank="1BE3B4A4-8BA0-4267-A89C-2EAD3CCF13C9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614326287498" checkinUser="plazi" docAuthor="Zhang, Wen-Xuan &amp; Yin, Zi-Wei" docDate="2021" docId="03C787F3FFD6E66FB6CF08829D5383D1" docLanguage="en" docName="zootaxa.4938.2.5.pdf" docOrigin="Zootaxa 4938 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Batrisodes grossepunctatus Zhang &amp; Yin 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="248" masterDocId="FFFEFF8BFFD2E66AB6580815985B8548" masterDocTitle="Two new species of Batrisodes Reitter from Shaanxi, China (Coleoptera Staphylinidae: Pselaphinae)" masterLastPageNumber="250" masterPageNumber="243" pageNumber="247" updateTime="1698931162887" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="C98984B62E9B6426FAA3D24083477BF9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="482FBC2F6B66A571A6024E8D04F0F633">
<mods:title id="2C7C0AE32EE7602D8E4FD8A669A8132D">Two new species of Batrisodes Reitter from Shaanxi, China (Coleoptera Staphylinidae: Pselaphinae)</mods:title>
</mods:titleInfo>
<mods:name id="6717064DD6EA2E633867EEBB381EE0FB" type="personal">
<mods:role id="03DA0235562FD839CB0398BB909C8254">
<mods:roleTerm id="38B923E8FC1D983B65BDC20DE054299E">Author</mods:roleTerm>
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<mods:namePart id="13DB720F015A2E431A6EF137C2694A1E">Zhang, Wen-Xuan</mods:namePart>
<mods:nameIdentifier id="C403C065345EE727EA92AEE4C1813491" type="ORCID">0000-0003-0707-3694</mods:nameIdentifier>
<mods:affiliation id="0D8A3CD1AA60AF6BF597F7E389D00881">Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, 100 Guilin Road, Xuhui District, Shanghai 200234, China &amp; zhangwx @ protonmail. com; https: // orcid. org / 0000 - 0003 - 0707 - 3694</mods:affiliation>
<mods:nameIdentifier id="BD79D1A6CA2EFF96F84F280760DEE02B" type="email">zhangwx@protonmail.com</mods:nameIdentifier>
</mods:name>
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<mods:role id="AAF28BB2A93C11F53DE4BA9E2F806595">
<mods:roleTerm id="A1D893C074D2A1199A9A280FE4D44FBA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9CF8CA017D50AEAAAD644809BD02A244">Yin, Zi-Wei</mods:namePart>
<mods:affiliation id="4E41A7132609445C28EB31FA3BDDF499">Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, 100 Guilin Road, Xuhui District, Shanghai 200234, China</mods:affiliation>
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<mods:title id="AA6C38C69D0998FD8B413207D231FBC5">Zootaxa</mods:title>
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<mods:part id="1DC0A5B6AB58D627AFBE7E4EE23C6484">
<mods:date id="7DBCA4AD2A1253CB2CF3BC2E1F9F3591">2021</mods:date>
<mods:detail id="9A58D17207967EF4F83C3022DE835773" type="pubDate">
<mods:number id="6FE97230318A190256B3CD56162BA99A">2021-02-26</mods:number>
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<mods:detail id="AD66484743143095F9461A577979C336" type="volume">
<mods:number id="65DF2C9EE709ACA39254CCDA098EE20E">4938</mods:number>
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<mods:classification id="0D77E6846F4D593EB9B38748F951D7CE">journal article</mods:classification>
<mods:identifier id="DE22B04C1A9D3DFD9896D18D1A4D2C09" type="CLB-Dataset">7895</mods:identifier>
<mods:identifier id="430B4AFC7E6AE6168870B28634AF1C71" type="DOI">10.11646/zootaxa.4938.2.5</mods:identifier>
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<treatment id="03C787F3FFD6E66FB6CF08829D5383D1" ID-DOI="http://doi.org/10.5281/zenodo.4563890" ID-GBIF-Taxon="179988696" ID-Zenodo-Dep="4563890" LSID="urn:lsid:plazi:treatment:03C787F3FFD6E66FB6CF08829D5383D1" httpUri="http://treatment.plazi.org/id/03C787F3FFD6E66FB6CF08829D5383D1" lastPageId="5" lastPageNumber="248" pageId="4" pageNumber="247">
<subSubSection id="C374656EFFD6E66EB6CF08829AA285F9" box="[151,761,151,178]" pageId="4" pageNumber="247" type="nomenclature">
<paragraph id="8BD136E5FFD6E66EB6CF08829AA285F9" blockId="4.[151,761,151,213]" box="[151,761,151,178]" pageId="4" pageNumber="247">
<heading id="D0998189FFD6E66EB6CF08829AA285F9" bold="true" box="[151,761,151,178]" fontSize="11" level="1" pageId="4" pageNumber="247" reason="1">
<emphasis id="B91AEAF7FFD6E66EB6CF08829AA285F9" bold="true" box="[151,761,151,178]" pageId="4" pageNumber="247">
<taxonomicName id="4C6E4D66FFD6E66EB6CF08829AD585F9" ID-CoL="8XVMZ" authority="Zhang &amp; Yin" authorityName="Zhang &amp; Yin" authorityYear="2021" box="[151,654,151,178]" class="Insecta" family="Staphylinidae" genus="Batrisodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="247" phylum="Arthropoda" rank="species" species="grossepunctatus" status="sp. nov.">
<emphasis id="B91AEAF7FFD6E66EB6CF088299B985F9" bold="true" box="[151,482,151,178]" italics="true" pageId="4" pageNumber="247">Batrisodes grossepunctatus</emphasis>
Zhang &amp; Yin
</taxonomicName>
,
<taxonomicNameLabel id="A229578CFFD6E66EB4C308829AA285F9" box="[667,761,151,177]" pageId="4" pageNumber="247" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C374656EFFD6E66EB6CF08AE9950859D" box="[151,267,187,213]" pageId="4" pageNumber="247" type="description">
<paragraph id="8BD136E5FFD6E66EB6CF08AE9950859D" blockId="4.[151,761,151,213]" box="[151,267,187,213]" pageId="4" pageNumber="247">
(
<figureCitation id="13552A60FFD6E66EB6C708AE98BC859D" box="[159,231,187,213]" captionStart="FIGURE 3" captionStartId="4.[151,250,1852,1876]" captionTargetBox="[155,1430,423,1821]" captionTargetId="figure-92@4.[151,1435,417,1829]" captionTargetPageId="4" captionText="FIGURE 3. Dorsal habitus of Batrisodes grossepunctatus sp. nov. A. Male. B. Female. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4563892" httpUri="https://zenodo.org/record/4563892/files/figure.png" pageId="4" pageNumber="247">Figs 3</figureCitation>
,
<figureCitation id="13552A60FFD6E66EB6AC08AE9958859D" box="[244,259,187,213]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[263,1324,181,1913]" captionTargetId="figure-20@6.[263,1324,181,1920]" captionTargetPageId="6" captionText="FIGURE 4. Diagnostic characters of Batrisodes grossepunctatus sp. nov. (A, C, D–G. Male. B, H. Female) A, B. Head and pronotum. C. Mesofemur and mesotibia. D. Tergite VIII. E. Sternite VIII. F, G. Aedeagus, in lateral (F) and ventral (G) view. H. Female genitalia. Scale bars: 0.3 mm in A–C; 0.2 mm in F, G; 0.1 mm in D, E, H." figureDoi="http://doi.org/10.5281/zenodo.4563894" httpUri="https://zenodo.org/record/4563894/files/figure.png" pageId="4" pageNumber="247">4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C374656EFFD6E66EB6CF09169B4084C1" pageId="4" pageNumber="247" type="materials_examined">
<paragraph id="8BD136E5FFD6E66EB6CF09169B4084C1" blockId="4.[151,1437,259,394]" pageId="4" pageNumber="247">
<emphasis id="B91AEAF7FFD6E66EB6CF0916991D8455" bold="true" box="[151,326,259,285]" pageId="4" pageNumber="247">Type material.</emphasis>
<materialsCitation id="3B063CB8FFD6E66EB715091698A3842D" ID-GBIF-Occurrence="3045077303" collectedFrom="mixed leaf litter" collectingDate="2012-07-15" collectingMethod="sifted" collectionCode="SNUC" collectorName="Y. - H. Pan" country="China" county="Nanzheng County" elevation="1500" latitude="32.838886" location="Liping Forest Park" longLatPrecision="20" longitude="106.604996" municipality="Liping Forest Park" pageId="4" pageNumber="247" specimenCount="1" stateProvince="Shaanxi" typeStatus="holotype">
<emphasis id="B91AEAF7FFD6E66EB71509169A388455" bold="true" box="[333,611,259,285]" pageId="4" pageNumber="247">
<typeStatus id="54D58847FFD6E66EB715091699AB8455" box="[333,496,259,285]" pageId="4" pageNumber="247" type="holotype">HOLOTYPE</typeStatus>
:
<collectingCountry id="F3797675FFD6E66EB7A509169A018455" box="[509,602,259,285]" name="China" pageId="4" pageNumber="247">CHINA</collectingCountry>
:
</emphasis>
<specimenCount id="9D68FD6CFFD6E66EB43309109A268468" box="[619,637,261,288]" pageId="4" pageNumber="247" type="male">♂</specimenCount>
, ‘
<collectingCountry id="F3797675FFD6E66EB4CB09169A848455" box="[659,735,259,285]" pageId="4" pageNumber="247">China</collectingCountry>
:
<collectingRegion id="49AAF807FFD6E66EB4B209169B1C8455" box="[746,839,259,285]" country="China" name="Shaanxi" pageId="4" pageNumber="247">Shaanxi</collectingRegion>
,
<collectingCounty id="62B04E69FFD6E66EB50B09169C448455" box="[851,1055,259,285]" pageId="4" pageNumber="247">Nanzheng County</collectingCounty>
(
<emphasis id="B91AEAF7FFD6E66EB26809119CDF8453" bold="true" box="[1072,1156,260,283]" pageId="4" pageNumber="247">Ɯfflfl</emphasis>
),
<collectingMunicipality id="6BB5AC9FFFD6E66EB2C409169D348455" box="[1180,1391,259,286]" pageId="4" pageNumber="247">Liping Forest Park</collectingMunicipality>
(
<emphasis id="B91AEAF7FFD6E66EB3D8091199008477" bold="true" pageId="4" pageNumber="247">= ḢÑẎƙfin'ẘ</emphasis>
),
<geoCoordinate id="EE5A5022FFD6E66EB72D093C99A6840C" box="[373,509,297,324]" degrees="32" direction="north" minutes="50" orientation="latitude" pageId="4" pageNumber="247" precision="15" seconds="20" value="32.838886">32°50′20″N</geoCoordinate>
,
<geoCoordinate id="EE5A5022FFD6E66EB452093C9AC6840C" box="[522,669,297,324]" degrees="106" direction="east" minutes="36" orientation="longitude" pageId="4" pageNumber="247" precision="15" seconds="18" value="106.604996">106°36′18″E</geoCoordinate>
,
<date id="FFD01025FFD6E66EB4F309329B77840A" box="[683,812,295,322]" pageId="4" pageNumber="247" value="2012-07-15">
<collectingDate id="EF94E9CDFFD6E66EB4F309329B77840A" box="[683,812,295,322]" pageId="4" pageNumber="247" value="2012-07-15">15.vii.2012</collectingDate>
</date>
,
<quantity id="4C969B00FFD6E66EB56309329B8A8409" box="[827,977,295,321]" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="4" pageNumber="247" unit="m" value="1500.0" valueMax="1600.0" valueMin="1400.0">
<elevation id="0043D1D6FFD6E66EB56309329B8A8409" box="[827,977,295,321]" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="4" pageNumber="247" unit="m" value="1500.0" valueMax="1600.0" valueMin="1400.0">1400–1600m</elevation>
</quantity>
,
<collectedFrom id="38727518FFD6E66EB58709329CC38409" box="[991,1176,295,321]" pageId="4" pageNumber="247">mixed leaf litter</collectedFrom>
,
<collectingMethod id="522F4EF2FFD6E66EB2FD09329CBD8409" box="[1189,1254,295,321]" pageId="4" pageNumber="247">sifted</collectingMethod>
,
<collectorName id="269B5333FFD6E66EB2AB093D9D39840A" box="[1267,1378,296,322]" pageId="4" pageNumber="247">Y.-H. Pan</collectorName>
leg.’ (
<collectionCode id="ED7FAE20FFD6E66EB6C7095E98AB842D" box="[159,240,331,357]" pageId="4" pageNumber="247">SNUC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B063CB8FFD6E66EB75A095E9C36842D" ID-GBIF-Occurrence="3045077304" box="[258,1133,331,360]" collectedFrom="mixed leaf litter" collectingDate="2012-07-16" collectingMethod="sifted" collectionCode="SNUC" collectorName="Li Li-Zheng" country="China" county="Nanzheng County" elevation="1500" latitude="32.838886" location="CHINA" longLatPrecision="20" longitude="106.604996" municipality="Liping Forest Park" pageId="4" pageNumber="247" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Shaanxi" typeStatus="paratype">
<emphasis id="B91AEAF7FFD6E66EB75A095E9A47842D" bold="true" box="[258,540,331,357]" pageId="4" pageNumber="247">
<typeStatus id="54D58847FFD6E66EB75A095E99F1842D" box="[258,426,331,357]" pageId="4" pageNumber="247" type="paratype">PARATYPES</typeStatus>
:
<collectingCountry id="F3797675FFD6E66EB7EE095E9A48842D" box="[438,531,331,357]" name="China" pageId="4" pageNumber="247">CHINA</collectingCountry>
:
</emphasis>
<specimenCount id="9D68FD6CFFD6E66EB47B095E9A118420" box="[547,586,331,360]" pageId="4" pageNumber="247" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D68FD6CFFD6E66EB403095E9AC1842E" box="[603,666,331,358]" pageId="4" pageNumber="247" type="female">3 ♀♀</specimenCount>
, same label data as for holotype (
<collectionCode id="ED7FAE20FFD6E66EB248095E9C3A842D" box="[1040,1121,331,357]" collectionName="China, Shanghai, Shanghai Normal University, Department of Biology, insect Collection" pageId="4" pageNumber="247">SNUC</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B063CB8FFD6E66EB22C095E9B4C84C1" ID-GBIF-Occurrence="3045077302" collectedFrom="mixed leaf litter" collectingDate="2012-07-16" collectingMethod="sifted" collectionCode="SNUC" collectorName="Li Li-Zheng" country="China" county="Nanzheng County" elevation="1500" latitude="32.838886" location="CHINA" longLatPrecision="20" longitude="106.604996" municipality="Liping Forest Park" pageId="4" pageNumber="247" specimenCount="1" specimenCount-female="3" specimenCount-male="1" stateProvince="Shaanxi" typeStatus="paratype">
<specimenCount id="9D68FD6CFFD6E66EB22C095E9CC18420" box="[1140,1178,331,360]" pageId="4" pageNumber="247" type="male">1 ♂</specimenCount>
, same label data as for holotype, except ‘
<date id="FFD01025FFD6E66EB704097A99B984C1" box="[348,482,367,393]" pageId="4" pageNumber="247" value="2012-07-16">
<collectingDate id="EF94E9CDFFD6E66EB704097A99B984C1" box="[348,482,367,393]" pageId="4" pageNumber="247" value="2012-07-16">16.vii.2012</collectingDate>
</date>
,
<collectorName id="269B5333FFD6E66EB7B6097A9A2384C1" box="[494,632,367,393]" pageId="4" pageNumber="247">Li Li-Zheng</collectorName>
leg.’ (
<collectionCode id="ED7FAE20FFD6E66EB4E6097A9B5484C1" box="[702,783,367,393]" collectionName="China, Shanghai, Shanghai Normal University, Department of Biology, insect Collection" pageId="4" pageNumber="247">SNUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF11666DFFD6E66EB6CF0F299CBB821C" ID-DOI="http://doi.org/10.5281/zenodo.4563892" ID-Zenodo-Dep="4563892" box="[151,1248,1852,1877]" httpUri="https://zenodo.org/record/4563892/files/figure.png" pageId="4" pageNumber="247" startId="4.[151,250,1852,1876]" targetBox="[155,1430,423,1821]" targetPageId="4">
<paragraph id="8BD136E5FFD6E66EB6CF0F299CBB821C" blockId="4.[151,1248,1852,1877]" box="[151,1248,1852,1877]" pageId="4" pageNumber="247">
<emphasis id="B91AEAF7FFD6E66EB6CF0F299948821D" bold="true" box="[151,275,1852,1877]" pageId="4" pageNumber="247">FIGURE 3.</emphasis>
Dorsal habitus of
<taxonomicName id="4C6E4D66FFD6E66EB7960F289ABE821C" authority="Zhang &amp; Yin, 2021" authorityName="Zhang &amp; Yin" authorityYear="2021" box="[462,741,1853,1877]" class="Insecta" family="Staphylinidae" genus="Batrisodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="247" phylum="Arthropoda" rank="species" species="grossepunctatus" status="sp. nov.">
<emphasis id="B91AEAF7FFD6E66EB7960F289ABE821C" box="[462,741,1853,1877]" italics="true" pageId="4" pageNumber="247">Batrisodes grossepunctatus</emphasis>
</taxonomicName>
<emphasis id="B91AEAF7FFD6E66EB4B30F289B00821C" bold="true" box="[747,859,1852,1877]" pageId="4" pageNumber="247">
<taxonomicNameLabel id="A229578CFFD6E66EB4B30F289B60821D" box="[747,827,1853,1877]" pageId="4" pageNumber="247" rank="species">sp. nov.</taxonomicNameLabel>
A.
</emphasis>
Male.
<emphasis id="B91AEAF7FFD6E66EB5FA0F289BE2821D" bold="true" box="[930,953,1853,1877]" pageId="4" pageNumber="247">B.</emphasis>
Female. Scale bars: 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C374656EFFD6E66FB69F0F969A9C8455" lastPageId="5" lastPageNumber="248" pageId="4" pageNumber="247" type="diagnosis">
<paragraph id="8BD136E5FFD6E66FB69F0F969A9C8455" blockId="4.[151,1437,1923,2021]" lastBlockId="5.[151,1437,151,1689]" lastPageId="5" lastPageNumber="248" pageId="4" pageNumber="247">
<emphasis id="B91AEAF7FFD6E66EB69F0F96991982D5" bold="true" box="[199,322,1923,1949]" pageId="4" pageNumber="247">Diagnosis.</emphasis>
Male. Body length
<quantity id="4C969B00FFD6E66EB47B0F969A9282D5" box="[547,713,1923,1949]" metricMagnitude="-3" metricUnit="m" metricValue="2.445" metricValueMax="2.48" metricValueMin="2.41" pageId="4" pageNumber="247" unit="mm" value="2.445" valueMax="2.48" valueMin="2.41">2.41–2.48 mm</quantity>
. Head base rounded, wider than pronotum; frons with anterolateral margin greatly protruding anteriorly, deeply and roundly concave at middle; clypeus with large projection at middle, sides of projection narrowing posteriorly from apical 1/
<quantity id="4C969B00FFD6E66EB53D0FDE9BCA82AD" box="[869,913,1995,2021]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="4" pageNumber="247" unit="in" value="3.0">3 in</quantity>
dorsal view. Pronotum roughly punctate, with four distinct discal teeth. Mesofemur with long ventral spine near middle, mesotibia expanded for apical 1/3 of tibial length, with blunt projection before middle. Aedeagus dorso-ventrally symmetric, with symmetric, well-sclerotized endophallus. Female. Similar to male in external morphology, lacking modifications on head and legs; may be identified only by geographical association with male.
</paragraph>
</subSubSection>
<subSubSection id="C374656EFFD7E66FB69F09339B068319" pageId="5" pageNumber="248" type="description">
<paragraph id="8BD136E5FFD7E66FB69F09339D208111" blockId="5.[151,1437,151,1689]" pageId="5" pageNumber="248">
<emphasis id="B91AEAF7FFD7E66FB69F093399028408" bold="true" box="[199,345,294,320]" pageId="5" pageNumber="248">Description.</emphasis>
Male (
<figureCitation id="13552A60FFD7E66FB7F009329A598409" box="[424,514,295,321]" captionStart="FIGURE 3" captionStartId="4.[151,250,1852,1876]" captionTargetBox="[155,1430,423,1821]" captionTargetId="figure-92@4.[151,1435,417,1829]" captionTargetPageId="4" captionText="FIGURE 3. Dorsal habitus of Batrisodes grossepunctatus sp. nov. A. Male. B. Female. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4563892" httpUri="https://zenodo.org/record/4563892/files/figure.png" pageId="5" pageNumber="248">Fig. 3A</figureCitation>
). Body reddish brown, length from anterior margin of clypeus to apex of abdomen
<quantity id="4C969B00FFD7E66FB6CF095E9966842D" box="[151,317,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="2.445" metricValueMax="2.48" metricValueMin="2.41" pageId="5" pageNumber="248" unit="mm" value="2.445" valueMax="2.48" valueMin="2.41">2.41–2.48 mm</quantity>
. Head (
<figureCitation id="13552A60FFD7E66FB7C9095E99B3842D" box="[401,488,331,357]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[263,1324,181,1913]" captionTargetId="figure-20@6.[263,1324,181,1920]" captionTargetPageId="6" captionText="FIGURE 4. Diagnostic characters of Batrisodes grossepunctatus sp. nov. (A, C, D–G. Male. B, H. Female) A, B. Head and pronotum. C. Mesofemur and mesotibia. D. Tergite VIII. E. Sternite VIII. F, G. Aedeagus, in lateral (F) and ventral (G) view. H. Female genitalia. Scale bars: 0.3 mm in A–C; 0.2 mm in F, G; 0.1 mm in D, E, H." figureDoi="http://doi.org/10.5281/zenodo.4563894" httpUri="https://zenodo.org/record/4563894/files/figure.png" pageId="5" pageNumber="248">Fig. 4A</figureCitation>
) slight wider than long, length from anterior margin of clypeus to head base (excluding occipital constriction)
<quantity id="4C969B00FFD7E66FB79B097B9A3284C1" box="[451,617,366,393]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" metricValueMax="5.4" metricValueMin="5.2" pageId="5" pageNumber="248" unit="mm" value="0.53" valueMax="0.54" valueMin="0.52">0.52–0.54 mm</quantity>
, width across eyes
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; surface roughly punctate, with large, nude vertexal foveae; frons with anterolateral margin greatly protruding anteriorly, deeply and roundly concave at middle; clypeus with large projection at middle, margins of projection narrowing posteriorly from apical 1/
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dorsal view. Clypeal anterior margin rounded; lateral vertexal carina faint, extending from occipital constriction to level of posterior margin of eyes, mediobasal carina weakly extending from head base to level of anterior margin of eyes. Eyes relatively small, each composed of approximately 40 facets. Antenna lacking distinct club, antennomere 1 (scape) thick, subcylindrical, antennomeres 2–10 moniliform, antennomeres 2–8 of similar width, antennomere 9 wider than 8, antennomeres 9 and 10 successively broader, each strongly transverse, antennomere 11 largest, subconical, lacking hook-like spine. Pronotum approximately as long as wide, length along midline
<quantity id="4C969B00FFD7E66FB3760A9B9DC387E1" box="[1326,1432,654,681]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="248" unit="mm" value="0.5">0.50 mm</quantity>
, maximum width
<quantity id="4C969B00FFD7E66FB70E0AA699E68785" box="[342,445,691,717]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" pageId="5" pageNumber="248" unit="mm" value="0.49">0.49 mm</quantity>
; sides rounded, widest near the middle, disc slightly convex, roughly punctate, with four distinct teeth; median longitudinal sulci weekly present; lateral longitudinal sulci distinct; with small nude median and setose lateral antebasal foveae; outer and inner pair of basolateral foveae distinct. Elytra wider than long, length along suture
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, maximum width
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; each elytron with three large, nude basal foveae, inner two close; discal striae shallow and short, extending posteriorly for 1/3 of elytral length. Mesofemur (
<figureCitation id="13552A60FFD7E66FB3650B569DCF8615" box="[1341,1428,835,861]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[263,1324,181,1913]" captionTargetId="figure-20@6.[263,1324,181,1920]" captionTargetPageId="6" captionText="FIGURE 4. Diagnostic characters of Batrisodes grossepunctatus sp. nov. (A, C, D–G. Male. B, H. Female) A, B. Head and pronotum. C. Mesofemur and mesotibia. D. Tergite VIII. E. Sternite VIII. F, G. Aedeagus, in lateral (F) and ventral (G) view. H. Female genitalia. Scale bars: 0.3 mm in A–C; 0.2 mm in F, G; 0.1 mm in D, E, H." figureDoi="http://doi.org/10.5281/zenodo.4563894" httpUri="https://zenodo.org/record/4563894/files/figure.png" pageId="5" pageNumber="248">Fig. 4C</figureCitation>
) with thin but distinct ventral spine near middle, mesotibia expanded for apical 1/3 of tibial length, with blunt ventral projection at apical 2/5, apex of projection rounded, with dense short setae. Abdomen widest at lateral margins of tergite 1 (IV), length of dorsally exposed part along midline
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, maximum width
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; tergite 1 (IV) with strongly oblique marginal carinae, discal carinae thick, tergite 5 (VIII) (
<figureCitation id="13552A60FFD7E66FB2100BC69CFB86A5" box="[1096,1184,979,1005]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[263,1324,181,1913]" captionTargetId="figure-20@6.[263,1324,181,1920]" captionTargetPageId="6" captionText="FIGURE 4. Diagnostic characters of Batrisodes grossepunctatus sp. nov. (A, C, D–G. Male. B, H. Female) A, B. Head and pronotum. C. Mesofemur and mesotibia. D. Tergite VIII. E. Sternite VIII. F, G. Aedeagus, in lateral (F) and ventral (G) view. H. Female genitalia. Scale bars: 0.3 mm in A–C; 0.2 mm in F, G; 0.1 mm in D, E, H." figureDoi="http://doi.org/10.5281/zenodo.4563894" httpUri="https://zenodo.org/record/4563894/files/figure.png" pageId="5" pageNumber="248">Fig. 4D</figureCitation>
) semicircular; sternite 6 (VIII) (
<figureCitation id="13552A60FFD7E66FB7590BE2990C8159" box="[257,343,1015,1041]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[263,1324,181,1913]" captionTargetId="figure-20@6.[263,1324,181,1920]" captionTargetPageId="6" captionText="FIGURE 4. Diagnostic characters of Batrisodes grossepunctatus sp. nov. (A, C, D–G. Male. B, H. Female) A, B. Head and pronotum. C. Mesofemur and mesotibia. D. Tergite VIII. E. Sternite VIII. F, G. Aedeagus, in lateral (F) and ventral (G) view. H. Female genitalia. Scale bars: 0.3 mm in A–C; 0.2 mm in F, G; 0.1 mm in D, E, H." figureDoi="http://doi.org/10.5281/zenodo.4563894" httpUri="https://zenodo.org/record/4563894/files/figure.png" pageId="5" pageNumber="248">Fig. 4E</figureCitation>
) transverse. Aedeagus (
<figureCitation id="13552A60FFD7E66FB43E0BE29A808159" box="[614,731,1015,1041]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[263,1324,181,1913]" captionTargetId="figure-20@6.[263,1324,181,1920]" captionTargetPageId="6" captionText="FIGURE 4. Diagnostic characters of Batrisodes grossepunctatus sp. nov. (A, C, D–G. Male. B, H. Female) A, B. Head and pronotum. C. Mesofemur and mesotibia. D. Tergite VIII. E. Sternite VIII. F, G. Aedeagus, in lateral (F) and ventral (G) view. H. Female genitalia. Scale bars: 0.3 mm in A–C; 0.2 mm in F, G; 0.1 mm in D, E, H." figureDoi="http://doi.org/10.5281/zenodo.4563894" httpUri="https://zenodo.org/record/4563894/files/figure.png" pageId="5" pageNumber="248">Fig. 4F, G</figureCitation>
)
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long, median lobe dorso-ventrally symmetrical and flattened, broadest near apex, apical part narrowing toward apex and curved ventrally in lateral view; endophallus well-sclerotized, lateral lobes broad at bases, abruptly constricted at middle and then evenly narrowing apically.
</paragraph>
<paragraph id="8BD136E5FFD7E66FB69F0C769DD081A1" blockId="5.[151,1437,151,1689]" pageId="5" pageNumber="248">
Female (
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). General morphology similar to male, lacking modifications on head (
<figureCitation id="13552A60FFD7E66FB2C10C769CB48135" box="[1177,1263,1123,1149]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[263,1324,181,1913]" captionTargetId="figure-20@6.[263,1324,181,1920]" captionTargetPageId="6" captionText="FIGURE 4. Diagnostic characters of Batrisodes grossepunctatus sp. nov. (A, C, D–G. Male. B, H. Female) A, B. Head and pronotum. C. Mesofemur and mesotibia. D. Tergite VIII. E. Sternite VIII. F, G. Aedeagus, in lateral (F) and ventral (G) view. H. Female genitalia. Scale bars: 0.3 mm in A–C; 0.2 mm in F, G; 0.1 mm in D, E, H." figureDoi="http://doi.org/10.5281/zenodo.4563894" httpUri="https://zenodo.org/record/4563894/files/figure.png" pageId="5" pageNumber="248">Fig. 4B</figureCitation>
) and legs; each eye composed of about 20 facets; genital complex (
<figureCitation id="13552A60FFD7E66FB48D0C929B7681E9" box="[725,813,1159,1185]" captionStart="FIGURE 4" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[263,1324,181,1913]" captionTargetId="figure-20@6.[263,1324,181,1920]" captionTargetPageId="6" captionText="FIGURE 4. Diagnostic characters of Batrisodes grossepunctatus sp. nov. (A, C, D–G. Male. B, H. Female) A, B. Head and pronotum. C. Mesofemur and mesotibia. D. Tergite VIII. E. Sternite VIII. F, G. Aedeagus, in lateral (F) and ventral (G) view. H. Female genitalia. Scale bars: 0.3 mm in A–C; 0.2 mm in F, G; 0.1 mm in D, E, H." figureDoi="http://doi.org/10.5281/zenodo.4563894" httpUri="https://zenodo.org/record/4563894/files/figure.png" pageId="5" pageNumber="248">Fig. 4H</figureCitation>
) symmetrical, weakly sclerotized,
<quantity id="4C969B00FFD7E66FB2EB0C929D4C81E9" box="[1203,1303,1159,1185]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="5" pageNumber="248" unit="mm" value="0.27">0.27 mm</quantity>
wide. Measurements of body parts (as for male): body length
<quantity id="4C969B00FFD7E66FB4870CBF9BDD818D" box="[735,902,1194,1221]" metricMagnitude="-3" metricUnit="m" metricValue="2.525" metricValueMax="2.53" metricValueMin="2.52" pageId="5" pageNumber="248" unit="mm" value="2.525" valueMax="2.53" valueMin="2.52">2.52–2.53 mm</quantity>
, length/width of head
<quantity id="4C969B00FFD7E66FB2DD0CBF9D72818D" box="[1157,1321,1194,1221]" metricMagnitude="-4" metricUnit="m" metricValue="5.15" metricValueMax="5.2" metricValueMin="5.1" pageId="5" pageNumber="248" unit="mm" value="0.515" valueMax="0.52" valueMin="0.51">0.51–0.52 mm</quantity>
/
<quantity id="4C969B00FFD7E66FB36A0CBF9DC3818D" box="[1330,1432,1194,1221]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="5" pageNumber="248" unit="mm" value="0.57">0.57 mm</quantity>
, pronotum
<quantity id="4C969B00FFD7E66FB7530CDB99F581A1" box="[267,430,1230,1257]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" metricValueMax="5.3" metricValueMin="5.2" pageId="5" pageNumber="248" unit="mm" value="0.525" valueMax="0.53" valueMin="0.52">0.52–0.53 mm</quantity>
/
<quantity id="4C969B00FFD7E66FB7EE0CDB9A0181A1" box="[438,602,1230,1257]" metricMagnitude="-4" metricUnit="m" metricValue="4.95" metricValueMax="5.0" metricValueMin="4.9" pageId="5" pageNumber="248" unit="mm" value="0.495" valueMax="0.5" valueMin="0.49">0.49–0.50 mm</quantity>
, elytra
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/
<quantity id="4C969B00FFD7E66FB5010CDA9BA781A1" box="[857,1020,1231,1257]" metricMagnitude="-4" metricUnit="m" metricValue="8.05" metricValueMax="8.1" metricValueMin="8.0" pageId="5" pageNumber="248" unit="mm" value="0.805" valueMax="0.81" valueMin="0.8">0.80–0.81 mm</quantity>
, abdomen
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/
<quantity id="4C969B00FFD7E66FB3780CDA9DDC81A1" box="[1312,1415,1231,1257]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="248" unit="mm" value="0.8">0.80 mm</quantity>
.
</paragraph>
<paragraph id="8BD136E5FFD7E66FB69F0CE798AB8089" blockId="5.[151,1437,151,1689]" pageId="5" pageNumber="248">
<emphasis id="B91AEAF7FFD7E66FB69F0CE799EF8045" bold="true" box="[199,436,1266,1293]" pageId="5" pageNumber="248">Comparative notes.</emphasis>
The new species is morphologically most similar to
<taxonomicName id="4C6E4D66FFD7E66FB2790CE69CE48045" authority="Zhang &amp; Yin, 2021" authorityName="Zhang &amp; Yin" authorityYear="2021" box="[1057,1215,1267,1293]" class="Insecta" family="Staphylinidae" genus="Batrisodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="248" phylum="Arthropoda" rank="species" species="grandiceps" status="sp. nov.">
<emphasis id="B91AEAF7FFD7E66FB2790CE69C698044" box="[1057,1074,1267,1292]" italics="true" pageId="5" pageNumber="248">B</emphasis>
.
<emphasis id="B91AEAF7FFD7E66FB21B0CE69CE48045" box="[1091,1215,1267,1293]" italics="true" pageId="5" pageNumber="248">grandiceps</emphasis>
</taxonomicName>
<emphasis id="B91AEAF7FFD7E66FB2920CE69D7C8045" bold="true" box="[1226,1319,1267,1293]" pageId="5" pageNumber="248">
<taxonomicNameLabel id="A229578CFFD7E66FB2920CE69D7C8045" box="[1226,1319,1267,1293]" pageId="5" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
described above by the male having a similar position and shape of the cephalic modifications and spination of the legs, as well as the presence of a well-sclerotized endophallus of the aedeagus. They can be clearly separated by the larger body size (
<quantity id="4C969B00FFD7E66FB74A0D4A99EC8031" box="[274,439,1375,1401]" metricMagnitude="-3" metricUnit="m" metricValue="2.445" metricValueMax="2.48" metricValueMin="2.41" pageId="5" pageNumber="248" unit="mm" value="2.445" valueMax="2.48" valueMin="2.41">2.41–2.48 mm</quantity>
vs.
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), the posteriorly narrowing sides of the projection on the clypeus, the roughly punctate pronotal disc, and the different shape of the lateral lobes of the aedeagal endophallus of the new species.
</paragraph>
<paragraph id="8BD136E5FFD7E66FB69F0DDE9B068319" blockId="5.[151,1437,151,1689]" pageId="5" pageNumber="248">
The modified male head is not an uncommon character state in Asian
<taxonomicName id="4C6E4D66FFD7E66FB5A40DDE9C2F80AD" box="[1020,1140,1483,1509]" class="Insecta" family="Staphylinidae" genus="Batrisodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="248" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAF7FFD7E66FB5A40DDE9C2F80AD" box="[1020,1140,1483,1509]" italics="true" pageId="5" pageNumber="248">Batrisodes</emphasis>
</taxonomicName>
. However, a roundly and deeply excavated anterior margin of the frons occurs only in
<taxonomicName id="4C6E4D66FFD7E66FB5100DFA9C1B8341" authority="Nomura" authorityName="Nomura" authorityYear="2007" box="[840,1088,1519,1545]" class="Insecta" family="Staphylinidae" genus="Batrisodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="248" phylum="Arthropoda" rank="species" species="babaianus">
<emphasis id="B91AEAF7FFD7E66FB5100DFA9B028340" box="[840,857,1519,1544]" italics="true" pageId="5" pageNumber="248">B</emphasis>
.
<emphasis id="B91AEAF7FFD7E66FB5300DFA9B868341" box="[872,989,1519,1545]" italics="true" pageId="5" pageNumber="248">babaianus</emphasis>
Nomura
</taxonomicName>
distributed in Meifeng (
<emphasis id="B91AEAF7FFD7E66FB30C0DFA9DD7834E" bold="true" box="[1364,1420,1519,1542]" pageId="5" pageNumber="248">Ṙữ</emphasis>
),
<collectingCountry id="F3797675FFD7E66FB6CF0E0698B68365" box="[151,237,1555,1581]" name="Taiwan" pageId="5" pageNumber="248">Taiwan</collectingCountry>
. The two new species described here can be readily separated from
<taxonomicName id="4C6E4D66FFD7E66FB5BB0E069C2C8365" authorityName="Nomura" authorityYear="2007" box="[995,1143,1555,1581]" class="Insecta" family="Staphylinidae" genus="Batrisodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="248" phylum="Arthropoda" rank="species" species="babaianus">
<emphasis id="B91AEAF7FFD7E66FB5BB0E069BAF8364" box="[995,1012,1555,1580]" italics="true" pageId="5" pageNumber="248">B</emphasis>
.
<emphasis id="B91AEAF7FFD7E66FB25A0E069C2C8365" box="[1026,1143,1555,1581]" italics="true" pageId="5" pageNumber="248">babaianus</emphasis>
</taxonomicName>
by the presence of a large projection on the clypeus, which is lacking in the latter species.
</paragraph>
</subSubSection>
<subSubSection id="C374656EFFD7E66FB69F0E4F9A40833D" box="[199,539,1626,1653]" pageId="5" pageNumber="248" type="distribution">
<paragraph id="8BD136E5FFD7E66FB69F0E4F9A40833D" blockId="5.[151,1437,151,1689]" box="[199,539,1626,1653]" pageId="5" pageNumber="248">
<emphasis id="B91AEAF7FFD7E66FB69F0E4F993A833C" bold="true" box="[199,353,1626,1652]" pageId="5" pageNumber="248">Distribution.</emphasis>
<collectingCountry id="F3797675FFD7E66FB73F0E4E99F5833D" box="[359,430,1627,1653]" name="China" pageId="5" pageNumber="248">China</collectingCountry>
:
<collectingRegion id="49AAF807FFD7E66FB7E10E4E9A4C833D" box="[441,535,1627,1653]" country="China" name="Shaanxi" pageId="5" pageNumber="248">Shaanxi</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C374656EFFD7E66FB69F0E6A9D5383D1" box="[199,1288,1663,1689]" pageId="5" pageNumber="248" type="etymology">
<paragraph id="8BD136E5FFD7E66FB69F0E6A9D5383D1" blockId="5.[151,1437,151,1689]" box="[199,1288,1663,1689]" pageId="5" pageNumber="248">
<emphasis id="B91AEAF7FFD7E66FB69F0E6A991583D1" bold="true" box="[199,334,1663,1689]" pageId="5" pageNumber="248">Etymology.</emphasis>
The new specific epithet refers to the roughly punctate pronotum of the new species.
</paragraph>
</subSubSection>
</treatment>
</document>
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