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140 lines
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.480.9046" ID-GBIF-Dataset="896d3365-3129-48f4-a14a-26039d9fab55" ID-PMC="PMC4319107" ID-Pensoft-Pub="1313-2970-480-41" ID-PubMed="25685022" ID-ZBK="F1DE2C0F1C90468E856B4A0BCEC56A07" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-480-41" ModsDocOrigin="ZooKeys 480" ModsDocTitle="One new species of the genus Belisana Thorell, 1898 (Araneae, Pholcidae) from northern Vietnam" checkinTime="1451244697677" checkinUser="pensoft" docAuthor="Pham, Dinh-Sac" docDate="2015" docId="655A288F545FEBCB743BD95C089EB2AB" docLanguage="en" docName="ZooKeys 480: 41-47" docOrigin="ZooKeys 480" docSource="http://dx.doi.org/10.3897/zookeys.480.9046" docTitle="Belisana denticulata Pham, 2015, sp. n." docType="treatment" docUuid="D82D1866-428A-4F07-B1F9-43E1BD589AC9" docUuidSource="ZooBank" docVersion="4" lastPageNumber="44" masterDocId="2F0AFFDB1422FFF04D17AE2AFFB78C2E" masterDocTitle="One new species of the genus Belisana Thorell, 1898 (Araneae, Pholcidae) from northern Vietnam" masterLastPageNumber="47" masterPageNumber="41" pageNumber="42" updateTime="1668159908903" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>One new species of the genus Belisana Thorell, 1898 (Araneae, Pholcidae) from northern Vietnam</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pham, Dinh-Sac</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="volume">
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<mods:number>480</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>41</mods:start>
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<mods:end>47</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.480.9046</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.480.9046</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-480-41</mods:identifier>
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<mods:identifier type="ZBK">F1DE2C0F1C90468E856B4A0BCEC56A07</mods:identifier>
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<mods:identifier type="ZooBank">F1DE2C0F1C90468E856B4A0BCEC56A07</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152058131" LSID="urn:lsid:zoobank.org:act:D82D1866-428A-4F07-B1F9-43E1BD589AC9" httpUri="http://treatment.plazi.org/id/655A288F545FEBCB743BD95C089EB2AB" lastPageId="3" lastPageNumber="44" pageId="1" pageNumber="42">
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<subSubSection pageId="1" pageNumber="42" type="multiple">
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<paragraph pageId="1" pageNumber="42">Taxon classification Animalia Araneae Pholcidae</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="42" type="nomenclature">
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<paragraph pageId="1" pageNumber="42">
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<taxonomicName LSID="http://zoobank.org/D82D1866-428A-4F07-B1F9-43E1BD589AC9" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana denticulata" order="Araneae" pageId="1" pageNumber="42" phylum="Arthropoda" rank="species" species="denticulata">Belisana denticulata</taxonomicName>
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<taxonomicNameLabel pageId="1" pageNumber="42">sp. n.</taxonomicNameLabel>
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Figs 1, 2, 3, 4
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="42" type="type material">
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<paragraph pageId="1" pageNumber="42">Type material.</paragraph>
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<paragraph pageId="1" pageNumber="42">
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Holotype: ♂, fogging, natural forest (
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="21.525017">21°31.501'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="105.557236">105°33.434'E</geoCoordinate>
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, elevation 1060 m), Tam Dao National Park, Vinh Phuc, Vietnam, 26 July 2008, D.S. Pham leg. Paratype: 1 ♂, same data but disturbed forest (
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="21.472282">21°28.337'N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="105.6349">105°38.094'E</geoCoordinate>
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, elevation 1007 m).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="43" type="etymology">
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<paragraph pageId="2" pageNumber="43">
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<pageBreakToken pageId="2" pageNumber="43" start="start">Etymology</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="2" pageNumber="43">The specific name is derived from the Latin denticulatus (with small teeth), in reference to the small teeth on the prolateral sclerite of the distal procursus; adjective.</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="43" type="diagnosis">
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<paragraph pageId="2" pageNumber="43">Diagnosis.</paragraph>
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<paragraph pageId="2" pageNumber="43">
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This species resembles
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<taxonomicName class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana scharffi" order="Araneae" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="scharffi">Belisana scharffi</taxonomicName>
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Huber, 2005 (see
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<bibRefCitation author="Huber, BA" journalOrPublisher="Zoologica" pageId="3" pageNumber="44" pagination="1 - 126" title="High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae)." volume="155" year="2005">Huber 2005</bibRefCitation>
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: 20, figs 9-10, 65, 136-153), but can be distinguished by relatively long distance between proximal parts of proximo-lateral apophysis and distal apophysis on male chelicerae (Figs 2B, 3C), by presence of a nearly saddle-shaped prolateral sclerite on procursus (arrows in Figs 1A, C, 3A), and by different shape of retrolateral membranous flap on procursus (Fig. 3B).
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</paragraph>
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<caption pageId="2" pageNumber="43">
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<paragraph pageId="2" pageNumber="43">
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Figure 1.
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<taxonomicName class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana denticulata" order="Araneae" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="denticulata">Belisana denticulata</taxonomicName>
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sp. n., holotype male.
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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Pedipalp (A Prolateral view, arrow points at nearly saddle-shaped sclerite B Retrolateral view)
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<normalizedToken originalValue="C–D">C-D</normalizedToken>
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Distal part of procursus (C Prolateral view, arrow points at nearly saddle-shaped sclerite D Retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus.
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</paragraph>
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</caption>
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<caption pageId="2" pageNumber="43">
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<paragraph pageId="2" pageNumber="43">
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Figure 2.
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<taxonomicName class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana denticulata" order="Araneae" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="denticulata">Belisana denticulata</taxonomicName>
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sp. n., holotype (
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<normalizedToken originalValue="B–E">B-E</normalizedToken>
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) and paratype (A) males. A Bulb, prolateral view B Chelicerae, frontal view
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<normalizedToken originalValue="C–E">C-E</normalizedToken>
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Habitus (C Dorsal view D Ventral view E Lateral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, pa = proximo-lateral apophysis.
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</paragraph>
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</caption>
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<caption pageId="2" pageNumber="43">
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<paragraph pageId="2" pageNumber="43">
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Figure 3.
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<taxonomicName class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana denticulata" order="Araneae" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="denticulata">Belisana denticulata</taxonomicName>
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sp. n., holotype (
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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) and paratype (D) males.
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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Pedipalp (A Prolateral view, arrow points at nearly saddle-shaped sclerite B Retrolateral view) C Chelicerae, frontal view D Bulb, prolateral view. b = bulb, ba = bulbal apophysis, e = embolus, f = membranous flap, pr = procursus.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="44" pageId="2" pageNumber="43" type="description">
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<paragraph pageId="2" pageNumber="43">Description.</paragraph>
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<paragraph lastPageId="3" lastPageNumber="44" pageId="2" pageNumber="43">
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Male (holotype): Total length 1.97 (2.09 with clypeus), prosoma 0.74 long, 0.80 wide, opisthosoma 1.23 long, 0.84 wide. Leg I: - (3.72 + 0.36 + - + - + -), leg II: 10.78 (2.95 + 0.34 + 2.62 + 3.89 + 0.98), leg III: 6.96 (1.87 + 0.30 + 1.72 + 2.41 + 0.66), leg IV: 9.22 (2.60 + 0.31 + 2.34 + 3.25 + 0.72). Habitus as in
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<pageBreakToken pageId="3" pageNumber="44" start="start">Fig</pageBreakToken>
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. 2
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<normalizedToken originalValue="C–E">C-E</normalizedToken>
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. Dorsal shield of prosoma yellowish, with brown lateral margins; sternum yellowish, without marks. Legs yellowish, without darker rings. Opisthosoma yellowish, without spots. Distance PME-PME 0.21, diameter PME 0.09, distance PME-ALE 0.02, AME absent. Ocular area not elevated. Thoracic furrow absent. Clypeus unmodified. Sternum wider than long (0.61/0.57). Chelicerae (Figs 2B, 3C) with a pair of thumb-shaped proximo-lateral apophyses and a pair of long, curved distal apophyses (distance between tips: 0.42). Pedipalps as in Figs 1
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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, 3
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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; trochanter with a short retrolatero-ventral apophysis; femur with a small dorsal apophysis; procursus simple proximally but complex distally, with a prolateral sclerite provided with small teeth, a small membranous flap retrolaterally and a curved spine; bulb with a hooked apophysis and a simple embolus. Legs with short vertical hairs on metatarsi, without spines and curved hairs.
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</paragraph>
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<paragraph pageId="3" pageNumber="44">Variation. In another male: Tibia I: 3.37; tibia I L/d: 45. Retrolateral trichobothrium of tibia I at 13% proximally; tarsus I with about 12 distinct pseudosegments.</paragraph>
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<paragraph pageId="3" pageNumber="44">Female. Unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="44" type="distribution">
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<paragraph pageId="3" pageNumber="44">Distribution.</paragraph>
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<paragraph pageId="3" pageNumber="44">Known from two nearby localities in Tam Dao National Park (Fig. 4).</paragraph>
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<caption pageId="3" pageNumber="44">
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<paragraph pageId="3" pageNumber="44">
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Figure 4. Distribution records of
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<taxonomicName class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Belisana denticulata" order="Araneae" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="denticulata">Belisana denticulata</taxonomicName>
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sp. n. from northern Vietnam.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document>
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