<documentid="AF0AEE53647210157DD9FA38484F8D17"ID-CLB-Dataset="299539"ID-DOI="10.1590/S1679-62252013000100007"ID-GBIF-Dataset="3b46cedf-2daf-4ae3-acdb-85e3c94029e3"ID-ISSN="1982-0224"ID-Zenodo-Dep="12652838"IM.bibliography_approvedBy="felipe"IM.illustrations_approvedBy="juliana"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.tables_approvedBy="juliana"IM.taxonomicNames_approvedBy="juliana"IM.treatments_approvedBy="juliana"checkinTime="1720079298511"checkinUser="felipe"docAuthor="Armbruster, Jonathan W. & Taphorn, Donald C."docDate="2013"docId="03C8165E343B49129BCB86A60B7531BD"docLanguage="en"docName="NeotropIchthyol.11.1.65-72.pdf"docOrigin="Neotropical Ichthyology 11 (1)"docSource="http://www.scielo.br/scielo.php?script=sci_arttext&pid=S1679-62252013000100065&lng=en&tlng=en"docStyle="DocumentStyle:85A40B78B715C4497F837D9D78407CE5.4:NeotropIchthyol.2003-2013.journal_article"docStyleId="85A40B78B715C4497F837D9D78407CE5"docStyleName="NeotropIchthyol.2003-2013.journal_article"docStyleVersion="4"docTitle="Neblinichthys peniculatus Armbruster & Taphorn, 2013, new species"docType="treatment"docVersion="6"lastPageNumber="69"masterDocId="FFF16E26343949169B2385750E263735"masterDocTitle="Description of Neblinichthys peniculatus, a new species of loricariid catfish from the río Paragua drainage of Venezuela"masterLastPageNumber="72"masterPageNumber="65"pageNumber="67"updateTime="1734027831463"updateUser="ExternalLinkService"zenodo-license-document="CC-BY-4.0"zenodo-license-figures="CC-BY-4.0">
<mods:titleid="9EB2C4452BE69951B01DAC469C2F2112">Description of Neblinichthys peniculatus, a new species of loricariid catfish from the río Paragua drainage of Venezuela</mods:title>
<mods:namePartid="BB5CFB8418C19895E699B53126870F42">Armbruster, Jonathan W.</mods:namePart>
<mods:affiliationid="8477AF903C7C25404B499EBE43FFEB2D">Department of Biological Sciences and Auburn University Museum of Natural History, 331 Funchess, Auburn, AL 36849, USA. armbrjw @ auburn. edu</mods:affiliation>
<taxonomicNameid="4C61DCCB343B49149BCB86A60C3334DE"ID-CoL="45YLT"authorityName="Armbruster & Taphorn"authorityYear="2013"box="[232,533,979,1003]"class="Actinopterygii"family="Loricariidae"genus="Neblinichthys"kingdom="Animalia"order="Siluriformes"pageId="2"pageNumber="67"phylum="Chordata"rank="species"species="peniculatus"status="sp. nov.">
<figureCitationid="135ABBCD343B49149A4886810FED3339"box="[363,459,1012,1036]"captionStart-0="Fig"captionStart-1="Fig"captionStart-2="Fig"captionStartId-0="3.[129,167,1695,1719]"captionStartId-1="4.[140,179,687,711]"captionStartId-2="5.[129,168,1709,1733]"captionTargetBox-0="[129,1452,205,1686]"captionTargetBox-1="[136,779,174,662]"captionTargetBox-2="[129,1452,229,1686]"captionTargetId-0="figure-102@3.[129,1452,205,1686]"captionTargetId-1="figure-9@4.[136,779,174,662]"captionTargetId-2="figure-24@5.[129,1452,229,1686]"captionTargetPageId-0="3"captionTargetPageId-1="4"captionTargetPageId-2="5"captionText-0="Fig. 2. Dorsal, lateral, and ventral views of Neblinichthys peniculatus,holotype, male, MBUCVV-35680, 78.8mm SL, Venezuela, Bolivar, río Paragua drainage, río Carapo at first rapids along right bank."captionText-1="Fig. 3. Dorsal view of head of holotype of Neblinichthys peniculatus,MBUCV V-35680, 78.8 mm SL."captionText-2="Fig. 4. Dorsal, lateral, and ventral views of Neblinichthys peniculatus, female, paratypes, AMNH 91020, 78.8 mm SL."figureDoi-0="http://doi.org/10.5281/zenodo.12652842"figureDoi-1="http://doi.org/10.5281/zenodo.12652844"figureDoi-2="http://doi.org/10.5281/zenodo.12652846"httpUri-0="https://zenodo.org/record/12652842/files/figure.png"httpUri-1="https://zenodo.org/record/12652844/files/figure.png"httpUri-2="https://zenodo.org/record/12652846/files/figure.png"pageId="2"pageNumber="67">Figs. 2-4</figureCitation>
<materialsCitationid="3B09AD15343B49149BAB814F0C043390"ID-GBIF-Occurrence="4903017307"collectingDate="1990-02-22"collectionCode="MBUCV, R"collectorName="C. J. Ferraris & Royero"country="Venezuela"latitude="5.6975"location="rio Carapo"longLatPrecision="21"longitude="-63.541668"municipality="rio Paragua"pageId="2"pageNumber="67"specimenCount="1"stateProvince="Bolivar"typeStatus="holotype">
<collectionCodeid="ED703F8D343B49149BD8814F0F73337A"box="[251,341,1082,1103]"country="Venezuela"httpUri="http://grbio.org/cool/7nxg-etxt"name="Museo de Biologia de la Universidad Central de Venezuela"pageId="2"pageNumber="67">MBUCV</collectionCode>
<collectingDateid="EF9B7860343B49149AE881010C6933BC"box="[459,591,1139,1161]"pageId="2"pageNumber="67"value="1990-02-22">22 Feb 1990</collectingDate>
</date>
,
<collectorNameid="2694C29E343B4914997E81010CC033BC"box="[605,742,1139,1161]"pageId="2"pageNumber="67">C. J. Ferraris</collectorName>
, A. Machado-A. &
<collectionCodeid="ED703F8D343B49149A0181E40F133393"box="[290,309,1169,1190]"country="Chile"name="Departamento de Geologia, Universidad de Chile"pageId="2"pageNumber="67">R</collectionCode>
<figureCitationid="135ABBCD343B49149AB781E50FFE3390"box="[404,472,1168,1190]"captionStart="Fig"captionStartId="3.[129,167,1695,1719]"captionTargetBox="[129,1452,205,1686]"captionTargetId="figure-102@3.[129,1452,205,1686]"captionTargetPageId="3"captionText="Fig. 2. Dorsal, lateral, and ventral views of Neblinichthys peniculatus,holotype, male, MBUCVV-35680, 78.8mm SL, Venezuela, Bolivar, río Paragua drainage, río Carapo at first rapids along right bank."figureDoi="http://doi.org/10.5281/zenodo.12652842"httpUri="https://zenodo.org/record/12652842/files/figure.png"pageId="2"pageNumber="67">Figs. 2</figureCitation>
and
<figureCitationid="135ABBCD343B4914992B81E50C303390"box="[520,534,1168,1189]"captionStart="Fig"captionStartId="4.[140,179,687,711]"captionTargetBox="[136,779,174,662]"captionTargetId="figure-9@4.[136,779,174,662]"captionTargetPageId="4"captionText="Fig. 3. Dorsal view of head of holotype of Neblinichthys peniculatus,MBUCV V-35680, 78.8 mm SL."figureDoi="http://doi.org/10.5281/zenodo.12652844"httpUri="https://zenodo.org/record/12652844/files/figure.png"pageId="2"pageNumber="67">3</figureCitation>
<specimenCodeid="DBC70F33343B49149BDE81BF0FBD33D5"box="[253,411,1226,1248]"collectionCode="AMNH"country="USA"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925"lsid="urn:lsid:biocol.org:col:34925"name="American Museum of Natural History"pageId="2"pageNumber="67">AMNH 255488</specimenCode>
,
<specimenCodeid="DBC70F33343B49149A8581BF0F9433EA"box="[422,434,1226,1247]"collectionCode="AMNH"country="USA"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925"lsid="urn:lsid:biocol.org:col:34925"name="American Museum of Natural History"pageId="2"pageNumber="67">2</specimenCode>
<specimenCodeid="DBC70F33343B49149B9381920F1933C9"box="[176,319,1255,1276]"collectionCode="AMNH"country="USA"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925"lsid="urn:lsid:biocol.org:col:34925"name="American Museum of Natural History"pageId="2"pageNumber="67">AMNH 91020</specimenCode>
<figureCitationid="135ABBCD343B4914990481920C4233C9"box="[551,612,1255,1277]"captionStart="Fig"captionStartId="5.[129,168,1709,1733]"captionTargetBox="[129,1452,229,1686]"captionTargetId="figure-24@5.[129,1452,229,1686]"captionTargetPageId="5"captionText="Fig. 4. Dorsal, lateral, and ventral views of Neblinichthys peniculatus, female, paratypes, AMNH 91020, 78.8 mm SL."figureDoi="http://doi.org/10.5281/zenodo.12652846"httpUri="https://zenodo.org/record/12652846/files/figure.png"pageId="2"pageNumber="67">Fig. 4</figureCitation>
)
</materialsCitation>
,
<materialsCitationid="3B09AD15343B4914995581920C1D322C"ID-GBIF-Occurrence="4903017301"collectingDate="1990-02-22"collectionCode="AUM"collectorName="C. J. Ferraris & Royero"country="Venezuela"latitude="5.6975"location="rio Carapo"longLatPrecision="21"longitude="-63.541668"municipality="rio Paragua"pageId="2"pageNumber="67"specimenCode="AUM 56753, 1"specimenCount="1"stateProvince="Bolivar"typeStatus="paratype">
<specimenCodeid="DBC70F33343B4914995581920CD233C8"box="[630,756,1255,1277]"collectionCode="AUM"country="USA"httpUri="http://grbio.org/cool/8wk2-p21y"name="Auburn University Museum of Natural History"pageId="2"pageNumber="67"type="University or college">AUM 56753</specimenCode>
,
<specimenCodeid="DBC70F33343B491499DC81920D2D33C9"box="[767,779,1255,1276]"collectionCode="AUM"country="USA"httpUri="http://grbio.org/cool/8wk2-p21y"name="Auburn University Museum of Natural History"pageId="2"pageNumber="67"type="University or college">1</specimenCode>
<specimenCodeid="DBC70F33343B4914996180710CF6322C"box="[578,720,1284,1305]"collectionCode="AMNH"country="USA"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925"lsid="urn:lsid:biocol.org:col:34925"name="American Museum of Natural History"pageId="2"pageNumber="67">AMNH 91021</specimenCode>
,
<specimenCodeid="DBC70F33343B491499FA80710CD7322C"box="[729,753,1284,1305]"collectionCode="AMNH"country="USA"httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925"lsid="urn:lsid:biocol.org:col:34925"name="American Museum of Natural History"pageId="2"pageNumber="67">14</specimenCode>
<materialsCitationid="3B09AD15343B49149A6180480D2D32B8"ID-GBIF-Occurrence="4903017306"collectionCode="MBUCV"collectorName="C. J. Ferraris"country="Venezuela"latitude="5.713333"location="rio Carapo"longLatPrecision="21"longitude="-63.533333"municipality="rio Paragua"pageId="2"pageNumber="67"specimenCount="1"stateProvince="Bolivar"typeStatus="paratype">
<collectionCodeid="ED703F8D343B49149A6180480FBC3267"box="[322,410,1341,1362]"country="Venezuela"httpUri="http://grbio.org/cool/7nxg-etxt"name="Museo de Biologia de la Universidad Central de Venezuela"pageId="2"pageNumber="67">MBUCV</collectionCode>
<collectionCodeid="ED703F8D343B49149B8280E10E92329C"box="[161,180,1428,1449]"country="Chile"name="Departamento de Geologia, Universidad de Chile"pageId="2"pageNumber="67">R</collectionCode>
<collectingDateid="EF9B7860343B49149A3080E10FE3329C"box="[275,453,1427,1449]"pageId="2"pageNumber="67"value="1990-02-28">28 February 1990</collectingDate>
<taxonomicNameid="4C61DCCB343B49149A3080A40C6332DC"authorityName="Armbruster & Taphorn"authorityYear="2013"box="[275,581,1489,1513]"class="Actinopterygii"family="Loricariidae"genus="Neblinichthys"kingdom="Animalia"order="Siluriformes"pageId="2"pageNumber="67"phylum="Chordata"rank="species"species="peniculatus">
caudal peduncle all about the same color or light below and dark above;
<figureCitationid="135ABBCD343B491499F683440EB3315C"captionStart="Fig"captionStartId="3.[129,167,1695,1719]"captionTargetBox="[129,1452,205,1686]"captionTargetId="figure-102@3.[129,1452,205,1686]"captionTargetPageId="3"captionText="Fig. 2. Dorsal, lateral, and ventral views of Neblinichthys peniculatus,holotype, male, MBUCVV-35680, 78.8mm SL, Venezuela, Bolivar, río Paragua drainage, río Carapo at first rapids along right bank."figureDoi="http://doi.org/10.5281/zenodo.12652842"httpUri="https://zenodo.org/record/12652842/files/figure.png"pageId="2"pageNumber="67">Figs. 2</figureCitation>
and
<figureCitationid="135ABBCD343B49149BFF83240EC8315C"box="[220,238,1617,1641]"captionStart="Fig"captionStartId="5.[129,168,1709,1733]"captionTargetBox="[129,1452,229,1686]"captionTargetId="figure-24@5.[129,1452,229,1686]"captionTargetPageId="5"captionText="Fig. 4. Dorsal, lateral, and ventral views of Neblinichthys peniculatus, female, paratypes, AMNH 91020, 78.8 mm SL."figureDoi="http://doi.org/10.5281/zenodo.12652846"httpUri="https://zenodo.org/record/12652846/files/figure.png"pageId="2"pageNumber="67">4</figureCitation>
), by having spots on the head (spots less conspicuous in nuptial males as the head is darker;
<figureCitationid="135ABBCD343B49149BF183E40F39319C"box="[210,287,1681,1705]"captionStart="Fig"captionStartId="3.[129,167,1695,1719]"captionTargetBox="[129,1452,205,1686]"captionTargetId="figure-102@3.[129,1452,205,1686]"captionTargetPageId="3"captionText="Fig. 2. Dorsal, lateral, and ventral views of Neblinichthys peniculatus,holotype, male, MBUCVV-35680, 78.8mm SL, Venezuela, Bolivar, río Paragua drainage, río Carapo at first rapids along right bank."figureDoi="http://doi.org/10.5281/zenodo.12652842"httpUri="https://zenodo.org/record/12652842/files/figure.png"pageId="2"pageNumber="67">Figs. 2</figureCitation>
and
<figureCitationid="135ABBCD343B49149A7983E40F4C319C"box="[346,362,1681,1705]"captionStart="Fig"captionStartId="5.[129,168,1709,1733]"captionTargetBox="[129,1452,229,1686]"captionTargetId="figure-24@5.[129,1452,229,1686]"captionTargetPageId="5"captionText="Fig. 4. Dorsal, lateral, and ventral views of Neblinichthys peniculatus, female, paratypes, AMNH 91020, 78.8 mm SL."figureDoi="http://doi.org/10.5281/zenodo.12652846"httpUri="https://zenodo.org/record/12652846/files/figure.png"pageId="2"pageNumber="67">4</figureCitation>
), and by having hypertrophied snout odontodes smaller than orbit diameter (
<figureCitationid="135ABBCD343B49149A2483840F63303C"box="[263,325,1777,1801]"captionStart="Fig"captionStartId="4.[140,179,687,711]"captionTargetBox="[136,779,174,662]"captionTargetId="figure-9@4.[136,779,174,662]"captionTargetPageId="4"captionText="Fig. 3. Dorsal view of head of holotype of Neblinichthys peniculatus,MBUCV V-35680, 78.8 mm SL."figureDoi="http://doi.org/10.5281/zenodo.12652844"httpUri="https://zenodo.org/record/12652844/files/figure.png"pageId="2"pageNumber="67">Fig. 3</figureCitation>
<taxonomicNameid="4C61DCCB343B49149AB283840C73303C"authorityName="Taphorn, Armbruster, Lopez-Fernandez & Bernard"authorityYear="2010"box="[401,597,1777,1801]"class="Actinopterygii"family="Loricariidae"genus="Neblinichthys"kingdom="Animalia"order="Siluriformes"pageId="2"pageNumber="67"phylum="Chordata"rank="species"species="brevibracchium">
<taxonomicNameid="4C61DCCB343B49149BC782E40F97309C"authorityName="Taphorn, Armbruster, Lopez-Fernandez & Bernard"authorityYear="2010"box="[228,433,1937,1961]"class="Actinopterygii"family="Loricariidae"genus="Neblinichthys"kingdom="Animalia"order="Siluriformes"pageId="2"pageNumber="67"phylum="Chordata"rank="species"species="brevibracchium">
<figureCitationid="135ABBCD343B49149E1A84440BA2367C"box="[1337,1412,305,329]"captionStart="Fig"captionStartId="3.[129,167,1695,1719]"captionTargetBox="[129,1452,205,1686]"captionTargetId="figure-102@3.[129,1452,205,1686]"captionTargetPageId="3"captionText="Fig. 2. Dorsal, lateral, and ventral views of Neblinichthys peniculatus,holotype, male, MBUCVV-35680, 78.8mm SL, Venezuela, Bolivar, río Paragua drainage, río Carapo at first rapids along right bank."figureDoi="http://doi.org/10.5281/zenodo.12652842"httpUri="https://zenodo.org/record/12652842/files/figure.png"pageId="2"pageNumber="67">Figs. 2</figureCitation>
and
<figureCitationid="135ABBCD343B4914981384240D66365C"box="[816,832,337,361]"captionStart="Fig"captionStartId="5.[129,168,1709,1733]"captionTargetBox="[129,1452,229,1686]"captionTargetId="figure-24@5.[129,1452,229,1686]"captionTargetPageId="5"captionText="Fig. 4. Dorsal, lateral, and ventral views of Neblinichthys peniculatus, female, paratypes, AMNH 91020, 78.8 mm SL."figureDoi="http://doi.org/10.5281/zenodo.12652846"httpUri="https://zenodo.org/record/12652846/files/figure.png"pageId="2"pageNumber="67">4</figureCitation>
<figureCitationid="135ABBCD343B4914981387C40D5B35FC"box="[816,893,689,713]"captionStart="Fig"captionStartId="3.[129,167,1695,1719]"captionTargetBox="[129,1452,205,1686]"captionTargetId="figure-102@3.[129,1452,205,1686]"captionTargetPageId="3"captionText="Fig. 2. Dorsal, lateral, and ventral views of Neblinichthys peniculatus,holotype, male, MBUCVV-35680, 78.8mm SL, Venezuela, Bolivar, río Paragua drainage, río Carapo at first rapids along right bank."figureDoi="http://doi.org/10.5281/zenodo.12652842"httpUri="https://zenodo.org/record/12652842/files/figure.png"pageId="2"pageNumber="67">Figs. 2</figureCitation>
and
<figureCitationid="135ABBCD343B4914989B87C40DEE35FC"box="[952,968,689,713]"captionStart="Fig"captionStartId="5.[129,168,1709,1733]"captionTargetBox="[129,1452,229,1686]"captionTargetId="figure-24@5.[129,1452,229,1686]"captionTargetPageId="5"captionText="Fig. 4. Dorsal, lateral, and ventral views of Neblinichthys peniculatus, female, paratypes, AMNH 91020, 78.8 mm SL."figureDoi="http://doi.org/10.5281/zenodo.12652846"httpUri="https://zenodo.org/record/12652846/files/figure.png"pageId="2"pageNumber="67">4</figureCitation>
), four to five bands or rows of spots in the caudal fin (
<figureCitationid="135ABBCD343B49149FA387A40AEB35DC"box="[1152,1229,721,745]"captionStart="Fig"captionStartId="3.[129,167,1695,1719]"captionTargetBox="[129,1452,205,1686]"captionTargetId="figure-102@3.[129,1452,205,1686]"captionTargetPageId="3"captionText="Fig. 2. Dorsal, lateral, and ventral views of Neblinichthys peniculatus,holotype, male, MBUCVV-35680, 78.8mm SL, Venezuela, Bolivar, río Paragua drainage, río Carapo at first rapids along right bank."figureDoi="http://doi.org/10.5281/zenodo.12652842"httpUri="https://zenodo.org/record/12652842/files/figure.png"pageId="2"pageNumber="67">Figs. 2</figureCitation>
and
<figureCitationid="135ABBCD343B49149E2487A40B3135DC"box="[1287,1303,721,745]"captionStart="Fig"captionStartId="5.[129,168,1709,1733]"captionTargetBox="[129,1452,229,1686]"captionTargetId="figure-24@5.[129,1452,229,1686]"captionTargetPageId="5"captionText="Fig. 4. Dorsal, lateral, and ventral views of Neblinichthys peniculatus, female, paratypes, AMNH 91020, 78.8 mm SL."figureDoi="http://doi.org/10.5281/zenodo.12652846"httpUri="https://zenodo.org/record/12652846/files/figure.png"pageId="2"pageNumber="67">4</figureCitation>
), and a larger predorsal length/postanal length ratio (150.1-164.6%
<tableCitationid="C6E392F3343B49149E6E86240B8B345C"box="[1357,1453,849,873]"captionStart="Table 1"captionStartId="1.[129,192,1083,1107]"captionTargetPageId="1"captionText="Table 1. Morphometric features of the heads as percentage of HL for Neblinichthys brevibracchium (N = 23, except SL, N = 29), N. echinasus (N = 3), N. pilosus (N = 6), N. roraima (N = 4), and N. yaravi (N = 8, except anal-fin spine L. and interorbital W., N = 7). Landmarks are the numbered landmarks inArmbruster (2003). Some N. roraima have completely adnate barbels, giving a measurement of zero. Revised from Taphorn et al. (2010) where these measurements were inadvertently published as ratios of predorsal length."httpUri="http://table.plazi.org/id/DF1EF7C0343849179BA2814E0FB133E6"pageId="2"pageNumber="67"tableUuid="DF1EF7C0343849179BA2814E0FB133E6">Table 1</tableCitation>
<paragraphid="8BDEA748343B4914987586C40A75321C"blockId="2.[816,1460,848,1961]"pageId="2"pageNumber="67">Body dorsoventrally flattened. Caudal peduncle laterally compressed. Profile from tip of snout to dorsal-fin origin convex. Orbits slightly elevated above head profile. Deepest point of body just anterior to dorsal fin. Profile straight or gently concave to adipose-fin origin, then slightly concave to base of upper caudal-fin spine. Ventral surface flat or slightly concave from just behind oral disk to anal-fin origin then straight to base of lower caudal spine. Eyes widely separated, laterally oriented, not visible from below. Snout broad and rounded. Body widest at pectoral-fin origin, then narrowing gently to origin of pelvic fins, then narrowing more quickly to caudal fin.</paragraph>
<paragraphid="8BDEA748343B4914987580440AD532DC"blockId="2.[816,1460,848,1961]"pageId="2"pageNumber="67">Anterior margin and dorsal surface of snout with small odontodes except in nuptial males (see Sexual dimorphism). Cheek plates strongly evertible (to 90° from head), cheek odontodes relatively long and numerous (~45-80), longest reaches to level of pectoral-fin spine insertion. Iris operculum absent, making dorsal rim of pupil round.</paragraph>
<paragraphid="8BDEA748343B4914987580840A31305C"blockId="2.[816,1460,848,1961]"pageId="2"pageNumber="67">Mouth moderate in size with narrow premaxillary and dentary tooth cups forming gentle arcs; dentigerous surfaces of dentary slightly smaller than premaxillary. Premaxillary teeth 49-90; dentary teeth 53-80. Teeth villiform and bicuspid with very short cusps (medial cusp longer than lateral). Edge of oral disk largely smooth, extending to vertical through anterior margin of eye, not extending beyond lateral margins of head. Enlarged central papilla present in buccal cavity. Maxillary barbel short, just reaching base of evertible cheek plates. Ventral surface of lips papillose. Papillae largest in center of lower lip, decreasing radially. No enlarged papillae located behind dentary teeth.</paragraph>
Dorsal fin II,7; exposed portion of dorsal-fin spinelet flat with few, short odontodes. Dorsal fin short, not reaching preadipose plate when adpressed. First dorsal-fin ray slightly longer than dorsal-fin spine. Pectoral fin I,6. Pectoral-fin spine short (about equal to pelvic-fin spine), reaching insertion of first branched pelvic-fin ray when adpressed ventral to pelvic fin; narrow but strong, numerous odontodes of equal size present along entire length in females, but odontodes becoming larger distally in nuptial males. Anterior pectoral-fin rays longer than pectoral-fin spine, decreasing to about half of length of spine posteriorly, covered with odontodes, largest on dorsal surface. Pelvic fin I,5; spine thickened and covered with odontodes, reaching into anterior third of anal-fin base when adpressed; anterior pelvic-fin rays longer than pelvic-fin spine with posterior margin of fin curving out beyond posterior tip of spine. Anal fin I,5; all branched anal-fin rays slightly longer than unbranched ray; unbranched ray with odontodes to about three quarters of its length. First anal-fin pterygiophore not exposed to form plate-like structure. Adipose-fin spine straight to slightly curved posteriorly with adipose membrane extending beyond posterior extent of spine, but some specimens with shortened spine and very small membrane; usually one (
with two) preadipose plates. Caudal fin I,14,I; lower caudal-fin spine slightly longer than upper, edge of caudal fin straight and oblique. Dorsal procurrent caudal-fin rays four or five, ventral procurrent caudal-fin rays three or four. Rays of all fins supporting small odontodes.
<taxonomicNameid="4C61DCCB343A4915991783EA0D183182"authorityName="Armbruster & Taphorn"authorityYear="2013"box="[564,830,1695,1719]"class="Actinopterygii"family="Loricariidae"genus="Neblinichthys"kingdom="Animalia"order="Siluriformes"pageId="3"pageNumber="68"phylum="Chordata"rank="species"species="peniculatus">
<paragraphid="8BDEA748343D49129B8D80A40F6731FC"blockId="4.[136,779,849,1737]"pageId="4"pageNumber="69">Median plate series with 22-24 (mode 23) plates, not counting triangular plates on base of caudal fin. Ventral plates forming gentle arc on caudal peduncle and not forming strong rounded keel. Plates in mid-ventral row weakly arched submedially forming low ridge from cleithrum to posterior insertion of pelvic fin. Three rows of plates on caudal peduncle (mid-dorsal plate series ending at level of adipose fin). Abdomen naked.</paragraph>
<emphasisid="B9157B5A343D49129BAB83850FA7303C"bold="true"box="[136,385,1776,1801]"pageId="4"pageNumber="69">Coloration in alcohol.</emphasis>
See
<figureCitationid="135ABBCD343D49129A9083840C32303C"box="[435,532,1777,1801]"captionStart-0="Fig"captionStart-1="Fig"captionStart-2="Fig"captionStartId-0="3.[129,167,1695,1719]"captionStartId-1="4.[140,179,687,711]"captionStartId-2="5.[129,168,1709,1733]"captionTargetBox-0="[129,1452,205,1686]"captionTargetBox-1="[136,779,174,662]"captionTargetBox-2="[129,1452,229,1686]"captionTargetId-0="figure-102@3.[129,1452,205,1686]"captionTargetId-1="figure-9@4.[136,779,174,662]"captionTargetId-2="figure-24@5.[129,1452,229,1686]"captionTargetPageId-0="3"captionTargetPageId-1="4"captionTargetPageId-2="5"captionText-0="Fig. 2. Dorsal, lateral, and ventral views of Neblinichthys peniculatus,holotype, male, MBUCVV-35680, 78.8mm SL, Venezuela, Bolivar, río Paragua drainage, río Carapo at first rapids along right bank."captionText-1="Fig. 3. Dorsal view of head of holotype of Neblinichthys peniculatus,MBUCV V-35680, 78.8 mm SL."captionText-2="Fig. 4. Dorsal, lateral, and ventral views of Neblinichthys peniculatus, female, paratypes, AMNH 91020, 78.8 mm SL."figureDoi-0="http://doi.org/10.5281/zenodo.12652842"figureDoi-1="http://doi.org/10.5281/zenodo.12652844"figureDoi-2="http://doi.org/10.5281/zenodo.12652846"httpUri-0="https://zenodo.org/record/12652842/files/figure.png"httpUri-1="https://zenodo.org/record/12652844/files/figure.png"httpUri-2="https://zenodo.org/record/12652846/files/figure.png"pageId="4"pageNumber="69">Figs. 2-4</figureCitation>
. Head, dorsum of body and sides tan with irregular, darker brown or black spots and blotches; mottling most intense on caudal peduncle; spots best formed on head, although faded in nuptial males. No individuals with regular rows of spots. Ventrum light colored, whitish or light tan with few faint dark spots on periphery in larger specimens. Oral disk with inner papillated surface pale tan; outer anterior margin of upper lip gray-brown. All fin spines and rays with spots centered on rays, but extending onto membranes. Occasionally, spots on fins combine to form bands with three to four bands in caudal fin (may be regular or irregular) and three to four bands in dorsal fin. Rectangular dark blotch present on base of caudal fin. Dark spot present at base of anteriormost dorsal-fin membrane. Dorsal surface with five saddles, usually faint), first under anterior part of dorsal fin, second under posterior part of dorsal fin, third and fourth very narrow on either side of adipose fin, and fifth dorsal extension of rectangular blotch on base of caudal fin.
<figureCitationid="135ABBCD343D49129F4D87850AF1343D"box="[1134,1239,752,776]"captionStart-0="Fig"captionStart-1="Fig"captionStartId-0="3.[129,167,1695,1719]"captionStartId-1="4.[140,179,687,711]"captionTargetBox-0="[129,1452,205,1686]"captionTargetBox-1="[136,779,174,662]"captionTargetId-0="figure-102@3.[129,1452,205,1686]"captionTargetId-1="figure-9@4.[136,779,174,662]"captionTargetPageId-0="3"captionTargetPageId-1="4"captionText-0="Fig. 2. Dorsal, lateral, and ventral views of Neblinichthys peniculatus,holotype, male, MBUCVV-35680, 78.8mm SL, Venezuela, Bolivar, río Paragua drainage, río Carapo at first rapids along right bank."captionText-1="Fig. 3. Dorsal view of head of holotype of Neblinichthys peniculatus,MBUCV V-35680, 78.8 mm SL."figureDoi-0="http://doi.org/10.5281/zenodo.12652842"figureDoi-1="http://doi.org/10.5281/zenodo.12652844"httpUri-0="https://zenodo.org/record/12652842/files/figure.png"httpUri-1="https://zenodo.org/record/12652844/files/figure.png"pageId="4"pageNumber="69">Figs. 2-3</figureCitation>
4. Males with elongated odontodes on dorsal portion of head and cleithrum anteriorly and ventrally from an arc extending from exposed dorsal part of cleithrum, to exposed opercle, to anterior of eye to nares; odontodes longest just anterior to base of evertible cheek plates, anteriorly above center of snout, and on opercle; longest odontode slightly smaller than orbit diameter. Hypertrophied odontodes also present on opercle, and exposed portion of cleithrum. Snout skin appears to thicken prior to emergence of hypertrophied odontodes as some individuals have thickened skin and shorter odontodes. Size of odontodes in
appear to be maximum size (see Discussion). Odontodes on head of female not elongated. Males also have odontodes on pectoral-fin spines becoming longer distally (