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<document id="7B1498423FD0EF84765801EA5FA92370" ID-DOI="10.5852/ejt.2018.399" ID-GBIF-Dataset="b389e1f3-82be-4258-9b09-b7ef9d839f99" ID-ISSN="2118-9773" ID-Zenodo-Dep="1163422" ID-ZooBank="E836E138-D6E2-4F62-B4B3-CE2E073F2B24" IM.bibliography_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="existingObjects,operationResults,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1517404992146" checkinUser="plazi" docAuthor="Zonstein, Sergei, Kunt, Kadir B. &amp; Yağmur, Ersen A." docDate="2018" docId="03B9B44C56520C00517B8354FD834D54" docLanguage="en" docName="ejt-399_zonstein_kunt.pdf.imf" docOrigin="European Journal of Taxonomy 399" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Raveniola vonwicki Zonstein 2000" docType="treatment" docVersion="15" lastPageNumber="40" masterDocId="FF80CC3456770C27533C8513FFDA4805" masterDocTitle="A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia" masterLastPageNumber="93" masterPageNumber="1" pageNumber="38" updateTime="1734610394516" updateUser="guilherme">
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<mods:title id="665D9AAE09FB4E614B43FB0C6AE5E831">A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia</mods:title>
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<mods:namePart id="A1370C3D96E8653F01CF4C465F558F48">Zonstein, Sergei</mods:namePart>
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<mods:namePart id="C7AB9F70AFD2AAA31AC12F934E3C359A">Kunt, Kadir B.</mods:namePart>
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<treatment id="03B9B44C56520C00517B8354FD834D54" ID-DOI="http://doi.org/10.5281/zenodo.5980308" ID-GBIF-Taxon="140561352" ID-Zenodo-Dep="5980308" LSID="urn:lsid:plazi:treatment:03B9B44C56520C00517B8354FD834D54" httpUri="http://treatment.plazi.org/id/03B9B44C56520C00517B8354FD834D54" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38" scope_family="Nemesiidae" scope_order="Araneae">
<subSubSection id="C30A56D156520C02517B8354FC364E67" box="[583,1004,1607,1634]" pageId="37" pageNumber="38" type="nomenclature">
<paragraph id="8BAF055A56520C02517B8354FC364E67" blockId="37.[450,1137,1607,1670]" box="[583,1004,1607,1634]" pageId="37" pageNumber="38">
<heading id="D0E7B23656520C02517B8354FC364E67" box="[583,1004,1607,1634]" centered="true" fontSize="11" level="2" pageId="37" pageNumber="38" reason="2">
<taxonomicName id="4C107ED956520C02517B8354FC364E67" ID-CoL="79599" authority="Zonstein, 2000" authorityName="Zonstein" authorityYear="2000" box="[583,1004,1607,1634]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="vonwicki">
<emphasis id="B964D94856520C02517B8354FCEF4E67" bold="true" box="[583,821,1607,1634]" italics="true" pageId="37" pageNumber="38">Raveniola vonwicki</emphasis>
<bibRefCitation id="EF8178AB56520C0250008354FC364E67" author="Zonstein S. L." box="[828,1004,1607,1634]" journalOrPublisher="Tethys Entomological Research" pageId="37" pageNumber="38" pagination="49 - 52" part="2" refString="Zonstein S. L. 2000. Two new species of the mygalomorph spider genus Raveniola Zonstein, 1987 (Araneae, Nemesiidae) from southwestern Asia. Tethys Entomological Research 2: 49 - 52." title="Two new species of the mygalomorph spider genus Raveniola Zonstein, 1987 (Araneae, Nemesiidae) from southwestern Asia" type="journal article" year="2000">Zonstein, 2000</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156520C0252FE8378FBAB4E80" box="[450,1137,1643,1670]" pageId="37" pageNumber="38" type="description">
<paragraph id="8BAF055A56520C0252FE8378FBAB4E80" blockId="37.[450,1137,1607,1670]" box="[450,1137,1643,1670]" pageId="37" pageNumber="38">
<figureCitation id="132B19DF56520C0252FE8378FD924E80" box="[450,584,1643,1669]" captionStart="Figs 59–67" captionStartId="59.[189,241,1662,1688]" captionTargetBox="[189,1398,265,1617]" captionTargetId="figure@59.[189,1398,265,1617]" captionText="Figs 59–67. Raveniola spp., ♀♀ (59, 63, 66) and ♂♂ (60–62, 64–65, 67). Left chelicera (59–61; white arrow indicates male intercheliceral organ), sternum, labium and maxillae (62–64), and spinnerets (65– 67); prolateral (59–61,65, 67)and ventral (62–64,66) views.— 59–60. R.pontica (Spassky, 1937). 61, 63, 66. R. zaizevi (Charitonov, 1948). 62, 65. R. hyrcanica Dunin, 1988. 64, 67. R. vonwicki Zonstein, 2000. Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/1163442/files/figure.png" pageId="37" pageNumber="38">Figs 64, 67</figureCitation>
, 
<figureCitation id="132B19DF56520C0251688378FDAE4E80" box="[596,628,1643,1669]" captionStart="Figs 79–83" captionStartId="62.[188,240,1783,1809]" captionTargetBox="[193,1394,269,1675]" captionTargetId="figure@62.[189,1402,265,1694]" captionText="Figs 79–83. Raveniola spp., ♂♂. Carapace and chelicerae, dorsal views. — 79. R. mazandaranica Marusik,Zamani&amp;Mirshamsi,2014(holotype).80.R.zaitzevi(Charitonov,1948)(Car).81.R.marusiki sp. nov. (holotype). 82. R. vonwicki (Zonstein, 2000) (holotype). 83. R. niedermeyeri (Brignoli, 1972) (holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163448/files/figure.png" pageId="37" pageNumber="38">82</figureCitation>
, 
<figureCitation id="132B19DF56520C0251BC8378FD7A4E80" box="[640,672,1643,1669]" captionStart="Figs 90–95" captionStartId="64.[189,241,1891,1917]" captionTargetBox="[189,1398,265,1846]" captionTargetId="figure@64.[189,1398,265,1849]" captionText="Figs 90–95. Raveniola spp., ♀♀. Carapace and chelicerae, dorsal views. — 90. R. turcica sp. nov. (paratype). 91. R. micropa (Ausserer, 1871) (holotype). 92. Same species as preceding (Uludağ). 93. R. zaitzevi(Charitonov, 1948)(Lagodekhi).94.R.vonwickiZonstein, 2000(paratype).95.R.niedermeyeri (Brignoli, 1972) (paratype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163452/files/figure.png" pageId="37" pageNumber="38">94</figureCitation>
, 
<figureCitation id="132B19DF56520C0251908378FD034E83" box="[684,729,1643,1670]" captionStart="Figs 102–116" captionStartId="66.[189,241,1755,1781]" captionTargetBox="[204,1382,265,1739]" captionTargetId="figure@66.[187,1396,250,1752]" captionText="Figs 102–116. Raveniola spp., ♂♂. Eyes and clypeus, dorsal views. — 102. R. adjarica sp. nov. (holotype).103. R.duninisp.nov.(holotype).104. R.hyrcanicaDunin, 1988(holotype).105.R.pontica (Spassky, 1937) (Lazarevskoe). 106.R.birecikensis sp. nov.(holotype).107. Samespecies as preceding (dwarf conspecific male?). 108. R. anadolu sp. nov. (holotype). 109. R. arthuri Kunt &amp; Yağmur, 2010 (holotype). 110. R. turcica sp. nov. (holotype). 111. R. micropa (Ausserer, 1871) (Uludağ). 112. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014 (holotype). 113. R. zaitzevi (Charitonov, 1948) (Car). 114. R. marusiki sp. nov. (holotype). 115. R. vonwicki (Zonstein, 2000) (holotype). 116. R. niedermeyeri (Brignoli, 1972) (holotype). Scalebar: 0.5 mm." httpUri="https://zenodo.org/record/1163456/files/figure.png" pageId="37" pageNumber="38">115</figureCitation>
, 
<figureCitation id="132B19DF56520C0251D98378FCC94E80" box="[741,787,1643,1669]" captionStart="Figs 117–128" captionStartId="67.[189,241,1621,1647]" captionTargetBox="[193,1382,278,1581]" captionTargetId="figure@67.[189,1398,265,1581]" captionText="Figs 117–128. Raveniola spp., ♀♀. Eyes and clypeus, dorsal views. — 117. R. adjarica sp. nov. (paratype).118. R.dunini sp. nov.(paratype).119.R.hyrcanica Dunin, 1988(paratype).120. R.pontica (Spassky, 1937) (Lazarevskoe). 121. R. sinani sp. nov. (holotype). 122. R. nana sp. nov. (holotype). 123. R. turcica sp. nov. (paratype). 124. R. micropa (Ausserer, 1871) (holotype). 125. Same species as preceding (Uludağ). 126. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 127. R. vonwicki Zonstein, 2000 (paratype). 128. R. niedermeyeri (Brignoli, 1972) (paratype). Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/1163458/files/figure.png" pageId="37" pageNumber="38">127</figureCitation>
, 
<figureCitation id="132B19DF56520C02501C8378FC944E80" box="[800,846,1643,1669]" captionStart="Figs 141–144" captionStartId="71.[189,241,1858,1884]" captionTargetBox="[239,1352,266,1791]" captionTargetId="figure@71.[206,1378,265,1793]" captionText="Figs 141–144. Raveniola spp., ♂♂. Tibia and metatarsus I, retrolateral views. — 141. R. zaitzevi (Charitonov, 1948) (Car). 142. R. marusiki sp. nov. (holotype). 143. R. vonwicki Zonstein, 2000 (holotype). 144. R. niedermeyeri (Brignoli, 1972) (holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163466/files/figure.png" pageId="37" pageNumber="38">143</figureCitation>
, 
<figureCitation id="132B19DF56520C0250678378FC534E80" box="[859,905,1643,1669]" captionStart="Figs 161–164" captionStartId="76.[189,241,1877,1903]" captionTargetBox="[197,1406,265,1871]" captionTargetId="figure@76.[197,1406,265,1875]" captionText="Figs 161–164. Raveniola spp., ♂♂. Palpal tibia, cymbium and palpal organ, retrolateral views. — 161.R.zaitzevi(Charitonov,1948)(Car).162.R.marusikisp.nov.(holotype).163.R.vonwickiZonstein, 2000 (holotype). 164. R. niedermeyeri (Brignoli, 1972) (holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163476/files/figure.png" pageId="37" pageNumber="38">163</figureCitation>
, 
<figureCitation id="132B19DF56520C0250A98378FC254E83" box="[917,1023,1643,1670]" captionStart="Figs 189–197" captionStartId="80.[189,241,1823,1849]" captionTargetBox="[200,1386,265,1801]" captionTargetId="figure@80.[200,1386,265,1801]" captionText="Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.1163484" httpUri="https://zenodo.org/record/1163484/files/figure.png" pageId="37" pageNumber="38">195–196</figureCitation>
, 
<figureCitation id="132B19DF56520C0257378378FBE04E80" box="[1035,1082,1643,1669]" captionStart="Figs 204–212" captionStartId="82.[189,241,1656,1682]" captionTargetBox="[198,1407,277,1637]" captionTargetId="figure@82.[198,1407,265,1652]" captionText="Figs 204–212. Raveniola spp., ♀♀. Vulva, dorsal views. — 204. R. nana sp. nov. (holotype). 205. R. turcica sp. nov. (paratype). 206. R. micropa (Ausserer, 1871) (Uludağ). 207. Same aspecies as preceding (holotype). 208. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 209. Same species as preceding (Pirkuli). 210. R. vonwicki Zonstein, 2000 (paratype). 211. R. niedermeyeri (Brignoli, 1972) (paratype, Gorgan). 212. Same species as preceding (Mazandaran). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163488/files/figure.png" pageId="37" pageNumber="38">210</figureCitation>
, 
<figureCitation id="132B19DF56520C0257798378FBAB4E80" box="[1093,1137,1643,1669]" captionStart="Figure 222" captionStartId="87.[189,272,1855,1881]" captionTargetBox="[189,1398,1034,1819]" captionTargetId="figure@87.[189,1398,1033,1819]" captionText="Figure 222. Localities of Raveniola spp. in Iran: R. dunini sp. nov. (), R. marusiki sp. nov. (). R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014 (), R. niedermeyeri (Brignoli, 1972) () andR.vonwickiZonstein,2000().Abbreviations:AJ=Azerbaijan;AM =Armenia;UZ =Uzbekistan." httpUri="https://zenodo.org/record/1163500/files/figure.png" pageId="37" pageNumber="38">222</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156520C02538183BEFC994EC2" box="[189,835,1709,1735]" pageId="37" pageNumber="38" type="reference_group">
<paragraph id="8BAF055A56520C02538183BEFC994EC2" blockId="37.[189,835,1709,1735]" box="[189,835,1709,1735]" pageId="37" pageNumber="38">
<taxonomicName id="4C107ED956520C02538183BEFD504EC2" ID-CoL="79599" authority="Zonstein, 2000: 49" authorityName="Zonstein" authorityPageNumber="49" authorityYear="2000" box="[189,650,1709,1735]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="vonwicki">
<emphasis id="B964D94856520C02538183BEFE794EC2" box="[189,419,1709,1735]" italics="true" pageId="37" pageNumber="38">Raveniola vonwicki</emphasis>
<bibRefCitation id="EF8178AB56520C02529783BEFD854EC2" author="Zonstein S. L." box="[427,607,1709,1735]" journalOrPublisher="Tethys Entomological Research" pageId="37" pageNumber="38" pagination="49 - 52" part="2" refString="Zonstein S. L. 2000. Two new species of the mygalomorph spider genus Raveniola Zonstein, 1987 (Araneae, Nemesiidae) from southwestern Asia. Tethys Entomological Research 2: 49 - 52." title="Two new species of the mygalomorph spider genus Raveniola Zonstein, 1987 (Araneae, Nemesiidae) from southwestern Asia" type="journal article" year="2000">Zonstein, 2000</bibRefCitation>
: 49
</taxonomicName>
, figs 1–3 (♂♀).
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156520C02538183FCFCD74FEE" pageId="37" pageNumber="38" type="diagnosis">
<paragraph id="8BAF055A56520C02538183FCFEEC4F0C" blockId="37.[189,310,1775,1801]" box="[189,310,1775,1801]" pageId="37" pageNumber="38">
<heading id="D0E7B23656520C02538183FCFEEC4F0C" bold="true" box="[189,310,1775,1801]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
<emphasis id="B964D94856520C02538183FCFEEC4F0C" bold="true" box="[189,310,1775,1801]" pageId="37" pageNumber="38">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56520C025381820CFCD74FEE" blockId="37.[189,1399,1822,2027]" pageId="37" pageNumber="38">
In the shape of the bulb this species resembles 
<taxonomicName id="4C107ED956520C025039820CFC4B4F3D" authority="Zonstein &amp; Kunt &amp; Yağmur, 2018" authorityName="Zonstein &amp; Kunt &amp; Yağmur" authorityYear="2018" box="[773,913,1822,1848]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="marusiki" status="sp. nov.">
<emphasis id="B964D94856520C025039820CFC4B4F3D" box="[773,913,1822,1848]" italics="true" pageId="37" pageNumber="38">R. marusiki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257643356520C0250A0820CFC204F3C" box="[924,1018,1823,1849]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C107ED956520C025706820CFB6D4F3D" authorityName="Charitonov" authorityYear="1948" box="[1082,1207,1823,1848]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="zaitzevi">
<emphasis id="B964D94856520C025706820CFB6D4F3D" box="[1082,1207,1823,1848]" italics="true" pageId="37" pageNumber="38">R. zaitzevi</emphasis>
</taxonomicName>
. It differs from both species by a noticeably longer apical segment of the PLS (
<figureCitation id="132B19DF56520C0250938251FBD24F59" box="[943,1032,1858,1884]" captionStart="Figs 59–67" captionStartId="59.[189,241,1662,1688]" captionTargetBox="[189,1398,265,1617]" captionTargetId="figure@59.[189,1398,265,1617]" captionText="Figs 59–67. Raveniola spp., ♀♀ (59, 63, 66) and ♂♂ (60–62, 64–65, 67). Left chelicera (59–61; white arrow indicates male intercheliceral organ), sternum, labium and maxillae (62–64), and spinnerets (65– 67); prolateral (59–61,65, 67)and ventral (62–64,66) views.— 59–60. R.pontica (Spassky, 1937). 61, 63, 66. R. zaizevi (Charitonov, 1948). 62, 65. R. hyrcanica Dunin, 1988. 64, 67. R. vonwicki Zonstein, 2000. Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/1163442/files/figure.png" pageId="37" pageNumber="38">Fig. 67</figureCitation>
; cf. 
<figureCitation id="132B19DF56520C0257078251FB484F58" box="[1083,1170,1858,1885]" captionStart="Figs 59–67" captionStartId="59.[189,241,1662,1688]" captionTargetBox="[189,1398,265,1617]" captionTargetId="figure@59.[189,1398,265,1617]" captionText="Figs 59–67. Raveniola spp., ♀♀ (59, 63, 66) and ♂♂ (60–62, 64–65, 67). Left chelicera (59–61; white arrow indicates male intercheliceral organ), sternum, labium and maxillae (62–64), and spinnerets (65– 67); prolateral (59–61,65, 67)and ventral (62–64,66) views.— 59–60. R.pontica (Spassky, 1937). 61, 63, 66. R. zaizevi (Charitonov, 1948). 62, 65. R. hyrcanica Dunin, 1988. 64, 67. R. vonwicki Zonstein, 2000. Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/1163442/files/figure.png" pageId="37" pageNumber="38">Fig. 66</figureCitation>
). The males can be distinguished in having a slightly longer embolus lacking dentate keels (
<figureCitation id="132B19DF56520C02571E8276FB104F85" box="[1058,1226,1893,1920]" captionStart="Figs 189–197" captionStartId="80.[189,241,1823,1849]" captionTargetBox="[200,1386,265,1801]" captionTargetId="figure@80.[200,1386,265,1801]" captionText="Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.1163484" httpUri="https://zenodo.org/record/1163484/files/figure.png" pageId="37" pageNumber="38">Figs 195–196</figureCitation>
; cf. 
<figureCitation id="132B19DF56520C02563C8276FF304FA6" captionStart="Figs 189–197" captionStartId="80.[189,241,1823,1849]" captionTargetBox="[200,1386,265,1801]" captionTargetId="figure@80.[200,1386,265,1801]" captionText="Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.1163484" httpUri="https://zenodo.org/record/1163484/files/figure.png" pageId="37" pageNumber="38">Figs 191– 194</figureCitation>
) and a more slender palpal tibia (
<figureCitation id="132B19DF56520C0251B9829AFD374FA6" box="[645,749,1929,1955]" captionStart="Figs 161–164" captionStartId="76.[189,241,1877,1903]" captionTargetBox="[197,1406,265,1871]" captionTargetId="figure@76.[197,1406,265,1875]" captionText="Figs 161–164. Raveniola spp., ♂♂. Palpal tibia, cymbium and palpal organ, retrolateral views. — 161.R.zaitzevi(Charitonov,1948)(Car).162.R.marusikisp.nov.(holotype).163.R.vonwickiZonstein, 2000 (holotype). 164. R. niedermeyeri (Brignoli, 1972) (holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163476/files/figure.png" pageId="37" pageNumber="38">Fig. 163</figureCitation>
; cf. 
<figureCitation id="132B19DF56520C025018829AFC114FA1" box="[804,971,1929,1956]" captionStart="Figs 161–164" captionStartId="76.[189,241,1877,1903]" captionTargetBox="[197,1406,265,1871]" captionTargetId="figure@76.[197,1406,265,1875]" captionText="Figs 161–164. Raveniola spp., ♂♂. Palpal tibia, cymbium and palpal organ, retrolateral views. — 161.R.zaitzevi(Charitonov,1948)(Car).162.R.marusikisp.nov.(holotype).163.R.vonwickiZonstein, 2000 (holotype). 164. R. niedermeyeri (Brignoli, 1972) (holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163476/files/figure.png" pageId="37" pageNumber="38">Figs 161–162</figureCitation>
). The female of 
<taxonomicName id="4C107ED956520C0257A98299FAFA4FA6" authorityName="Zonstein" authorityYear="2000" box="[1173,1312,1929,1955]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="vonwicki">
<emphasis id="B964D94856520C0257A98299FAFA4FA6" box="[1173,1312,1929,1955]" italics="true" pageId="37" pageNumber="38">R. vonwicki</emphasis>
</taxonomicName>
differs from those in the same species group in having longer spermathecae with stalked, not sessile, lateral receptacles (
<figureCitation id="132B19DF56520C02527382C2FE6C4FEE" box="[335,438,2001,2027]" captionStart="Figs 204–212" captionStartId="82.[189,241,1656,1682]" captionTargetBox="[198,1407,277,1637]" captionTargetId="figure@82.[198,1407,265,1652]" captionText="Figs 204–212. Raveniola spp., ♀♀. Vulva, dorsal views. — 204. R. nana sp. nov. (holotype). 205. R. turcica sp. nov. (paratype). 206. R. micropa (Ausserer, 1871) (Uludağ). 207. Same aspecies as preceding (holotype). 208. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 209. Same species as preceding (Pirkuli). 210. R. vonwicki Zonstein, 2000 (paratype). 211. R. niedermeyeri (Brignoli, 1972) (paratype, Gorgan). 212. Same species as preceding (Mazandaran). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163488/files/figure.png" pageId="37" pageNumber="38">Fig. 210</figureCitation>
; cf. 
<figureCitation id="132B19DF56520C0252DB82C2FCD84FEE" box="[487,770,2001,2027]" captionStart="Figs 204–212" captionStartId="82.[189,241,1656,1682]" captionTargetBox="[198,1407,277,1637]" captionTargetId="figure@82.[198,1407,265,1652]" captionText="Figs 204–212. Raveniola spp., ♀♀. Vulva, dorsal views. — 204. R. nana sp. nov. (holotype). 205. R. turcica sp. nov. (paratype). 206. R. micropa (Ausserer, 1871) (Uludağ). 207. Same aspecies as preceding (holotype). 208. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 209. Same species as preceding (Pirkuli). 210. R. vonwicki Zonstein, 2000 (paratype). 211. R. niedermeyeri (Brignoli, 1972) (paratype, Gorgan). 212. Same species as preceding (Mazandaran). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163488/files/figure.png" pageId="37" pageNumber="38">Figs 208–209, 211–212</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156510C0153818418FD274A0C" pageId="38" pageNumber="39" type="materials_examined">
<paragraph id="8BAF055A56510C0153818418FE704920" blockId="38.[189,426,267,293]" box="[189,426,267,293]" pageId="38" pageNumber="39">
<heading id="D0E7B23656510C0153818418FE704920" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="B964D94856510C0153818418FE704920" bold="true" box="[189,426,267,293]" pageId="38" pageNumber="39">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56510C0153D6842FFE6E49A4" blockId="38.[189,1398,316,417]" pageId="38" pageNumber="39">
<materialsCitation id="3B780F0756510C0153D6842FFE6E49A4" ID-GBIF-Occurrence="1806244351" collectingDate="1914-04-07" collectionCode="ZISP" collectorName="N. S. von Wick" country="Iran" location="E slope of Bogrovdagh Mts" municipality="Hishtpar District" pageId="38" pageNumber="39" specimenCode="ZISP 454-914" specimenCount="1" specimenCount-male="1" stateProvince="Gilan Province" typeStatus="holotype">
<typeStatus id="54ABBBF856510C0153D6842FFE864953" box="[234,348,316,342]" pageId="38" pageNumber="39" type="holotype">
<emphasis id="B964D94856510C0153D6842FFE864953" bold="true" box="[234,348,316,342]" pageId="38" pageNumber="39">Holotype</emphasis>
</typeStatus>
<collectingCountry id="F30745CA56510C0153818472FED3497E" box="[189,265,353,379]" name="Iran" pageId="38" pageNumber="39">IRAN</collectingCountry>
: 
<specimenCount id="9D16CED356510C0152298472FEF7497E" box="[277,301,353,379]" count="1" pageId="38" pageNumber="39" type="male">♂</specimenCount>
, 
<collectingRegion id="49D4CBB856510C0152078472FE2A497E" box="[315,496,353,379]" country="Iran" name="Gilan" pageId="38" pageNumber="39">Gilan Province</collectingRegion>
, 
<collectingMunicipality id="6BCB9F2056510C0152C18472FD1C497E" box="[509,710,353,379]" pageId="38" pageNumber="39">Hishtpar District</collectingMunicipality>
, 
<location id="8ECF538156510C0151EE8472FBC4497E" LSID="urn:lsid:plazi:treatment:03B9B44C56520C00517B8354FD834D54:8ECF538156510C0151EE8472FBC4497E" box="[722,1054,353,379]" country="Iran" municipality="Hishtpar District" name="E slope of Bogrovdagh Mts" pageId="38" pageNumber="39" stateProvince="Gilan Province">E slope of Bogrovdagh Mts</location>
, 
<collectingDate id="EFEADA7256510C0157178472FB60497E" box="[1067,1210,353,379]" pageId="38" pageNumber="39" value="1914-04-07">
<date id="FFAE239A56510C0157178472FB60497E" box="[1067,1210,353,379]" pageId="38" pageNumber="39" value="1914-04-07">7 Apr. 1914</date>
</collectingDate>
, 
<collectorName id="26E5608C56510C0157FB8472FAAC497E" box="[1223,1398,353,379]" pageId="38" pageNumber="39">N.S. von Wick</collectorName>
leg. (
<specimenCode id="DBB6AD2156510C0153CB8495FE7249A4" box="[247,424,390,417]" pageId="38" pageNumber="39">ZISP 454–914</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BAF055A56510C0153D684D9FD274A0C" blockId="38.[189,765,458,522]" pageId="38" pageNumber="39">
<materialsCitation id="3B780F0756510C0153D684D9FD274A0C" ID-GBIF-Occurrence="1806244380" collectingDate="1914-04-07" collectionCode="ZISP" collectorName="N. S. von Wick" country="Iran" location="IRAN" municipality="Hishtpar District" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1" stateProvince="Gilan Province" typeStatus="paratype">
<emphasis id="B964D94856510C0153D684D9FE8649E1" bold="true" box="[234,348,458,484]" pageId="38" pageNumber="39">
<typeStatus id="54ABBBF856510C0153D684D9FE8649E1" box="[234,348,458,484]" pageId="38" pageNumber="39">Paratype</typeStatus>
</emphasis>
<collectingCountry id="F30745CA56510C01538184E3FED34A0F" box="[189,265,496,522]" name="Iran" pageId="38" pageNumber="39">IRAN</collectingCountry>
: 
<specimenCount id="9D16CED356510C01522884FCFE9A4A0F" box="[276,320,495,522]" count="1" pageId="38" pageNumber="39" type="female">1 ♀</specimenCount>
, collected together with the holotype.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156510C0053818720FB904C69" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39" type="description">
<paragraph id="8BAF055A56510C0153818720FEB14A48" blockId="38.[189,363,563,589]" box="[189,363,563,589]" pageId="38" pageNumber="39">
<heading id="D0E7B23656510C0153818720FEB14A48" bold="true" box="[189,363,563,589]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="B964D94856510C0153818720FEB14A48" bold="true" box="[189,363,563,589]" pageId="38" pageNumber="39">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56510C0153D68777FE714A7A" blockId="38.[189,1191,612,676]" box="[234,427,612,639]" pageId="38" pageNumber="39">
<emphasis id="B964D94856510C0153D68777FEF04A7B" bold="true" box="[234,298,612,638]" pageId="38" pageNumber="39">Male</emphasis>
(holotype)
</paragraph>
<paragraph id="8BAF055A56510C0153818799FB7D4AA6" blockId="38.[189,1191,612,676]" box="[189,1191,649,676]" pageId="38" pageNumber="39">MEASUREMENTS. TBL 12.00, CL 5.53, CW 4.15, LL 0.43, LW 0.87, SL 2.53, SW 2.27.</paragraph>
<paragraph id="8BAF055A56510C01538187DEFD254B37" blockId="38.[189,1398,717,818]" pageId="38" pageNumber="39">COLOUR. Carapace, palps and legs dorsally dark brownish rufous; chelicerae reddish brown; sternum, labium, maxillae, palps and legs ventrally light brown; abdomen dorsally brown, with dense light brown spots; ventral surface of abdomen brownish grey.</paragraph>
<paragraph id="8BAF055A56510C015381864FFA9C4BE3" blockId="38.[189,1399,859,998]" pageId="38" pageNumber="39">
PROSOMA. Carapace and chelicerae as shown in 
<figureCitation id="132B19DF56510C01503E8648FC864B70" box="[770,860,859,885]" captionStart="Figs 79–83" captionStartId="62.[188,240,1783,1809]" captionTargetBox="[193,1394,269,1675]" captionTargetId="figure@62.[189,1402,265,1694]" captionText="Figs 79–83. Raveniola spp., ♂♂. Carapace and chelicerae, dorsal views. — 79. R. mazandaranica Marusik,Zamani&amp;Mirshamsi,2014(holotype).80.R.zaitzevi(Charitonov,1948)(Car).81.R.marusiki sp. nov. (holotype). 82. R. vonwicki (Zonstein, 2000) (holotype). 83. R. niedermeyeri (Brignoli, 1972) (holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163448/files/figure.png" pageId="38" pageNumber="39">Fig. 82</figureCitation>
. Clypeus and eye group as in 
<figureCitation id="132B19DF56510C0157EF8648FAE04B73" box="[1235,1338,859,886]" captionStart="Figs 102–116" captionStartId="66.[189,241,1755,1781]" captionTargetBox="[204,1382,265,1739]" captionTargetId="figure@66.[187,1396,250,1752]" captionText="Figs 102–116. Raveniola spp., ♂♂. Eyes and clypeus, dorsal views. — 102. R. adjarica sp. nov. (holotype).103. R.duninisp.nov.(holotype).104. R.hyrcanicaDunin, 1988(holotype).105.R.pontica (Spassky, 1937) (Lazarevskoe). 106.R.birecikensis sp. nov.(holotype).107. Samespecies as preceding (dwarf conspecific male?). 108. R. anadolu sp. nov. (holotype). 109. R. arthuri Kunt &amp; Yağmur, 2010 (holotype). 110. R. turcica sp. nov. (holotype). 111. R. micropa (Ausserer, 1871) (Uludağ). 112. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014 (holotype). 113. R. zaitzevi (Charitonov, 1948) (Car). 114. R. marusiki sp. nov. (holotype). 115. R. vonwicki (Zonstein, 2000) (holotype). 116. R. niedermeyeri (Brignoli, 1972) (holotype). Scalebar: 0.5 mm." httpUri="https://zenodo.org/record/1163456/files/figure.png" pageId="38" pageNumber="39">Fig. 115</figureCitation>
. Eye diameters and interdistances: AME 0.16(0.21), ALE 0.27, PLE 0.13, PME 0.14, AME–AME 0.11(0.06), ALE–AME 0.11(0.09), ALE–PLE 0.09, PLE–PME 0.05, PME–PME 0.35. Each cheliceral furrow with 9–10 promarginal teeth and 4–5 mesobasal denticles. Maxillae with 20–24 cuspules each (
<figureCitation id="132B19DF56510C0157DD86D8FAE34BE3" box="[1249,1337,971,998]" captionStart="Figs 59–67" captionStartId="59.[189,241,1662,1688]" captionTargetBox="[189,1398,265,1617]" captionTargetId="figure@59.[189,1398,265,1617]" captionText="Figs 59–67. Raveniola spp., ♀♀ (59, 63, 66) and ♂♂ (60–62, 64–65, 67). Left chelicera (59–61; white arrow indicates male intercheliceral organ), sternum, labium and maxillae (62–64), and spinnerets (65– 67); prolateral (59–61,65, 67)and ventral (62–64,66) views.— 59–60. R.pontica (Spassky, 1937). 61, 63, 66. R. zaizevi (Charitonov, 1948). 62, 65. R. hyrcanica Dunin, 1988. 64, 67. R. vonwicki Zonstein, 2000. Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/1163442/files/figure.png" pageId="38" pageNumber="39">Fig. 64</figureCitation>
).
</paragraph>
<paragraph id="8BAF055A56510C015381811CFE814C9C" blockId="38.[189,1399,1039,1177]" pageId="38" pageNumber="39">
LEGS. Tibia and metatarsus I as shown in 
<figureCitation id="132B19DF56510C015182811CFCFC4C2C" box="[702,806,1039,1065]" captionStart="Figs 141–144" captionStartId="71.[189,241,1858,1884]" captionTargetBox="[239,1352,266,1791]" captionTargetId="figure@71.[206,1378,265,1793]" captionText="Figs 141–144. Raveniola spp., ♂♂. Tibia and metatarsus I, retrolateral views. — 141. R. zaitzevi (Charitonov, 1948) (Car). 142. R. marusiki sp. nov. (holotype). 143. R. vonwicki Zonstein, 2000 (holotype). 144. R. niedermeyeri (Brignoli, 1972) (holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163466/files/figure.png" pageId="38" pageNumber="39">Fig. 143</figureCitation>
. Scopula: distal on metatarsi I–II, entire on tarsi I–II, vestigial on tarsi III, absent on tarsi IV. Trichobothria: 2 rows of 7–8 each on tibiae, 11–14 on metatarsi, 11–12 on tarsi, 9 on cymbium. Paired tarsal claws: outer margin with 8–9 teeth, 6–9 teeth on inner margin.
</paragraph>
<paragraph id="8BAF055A56510C01538181D0FBEB4D9D" blockId="38.[189,1399,1218,1432]" pageId="38" pageNumber="39">SPINATION. Palp: femur d1–1–1, pd1, rd1; patella r1; tibia d1–1–1, p1–1–1, v7(8); tarsus d5. Leg I: femur d1–1–b–b, pd1–1–1, rd1(0); tibia p1–1, pv1, rv1–1–1–m–m; metatarsus v2. Leg II: femur d1–1–b–b, pd1–1–1; patella p1; tibia p1–1–1, v2–2–3; metatarsus p1, v1–2–3. Leg III: femur d1–1–b–b, pd1–1–1, rd1–1–1; patella p1, r1; tibia d1–1, p1–1, r1–1–1, v2–2–2; metatarsus d1–1–1, pd1–1–1, p1–1–1, r1– 1–1, v2–2–3. Leg IV: femur d1–1–b–b, pd1–1–1, pr1–1–1; patella p1, r1; tibia d1–1, p1–1–1, r1–1–1, v2–2–3; metatarsus d1–1–1, p1–1–1, r1–1–1, v3–2–2–2. Patella I aspinose.</paragraph>
<paragraph id="8BAF055A56510C01538180D1FEA34E22" blockId="38.[189,1399,1473,1575]" pageId="38" pageNumber="39">
PALP. Tibia, cymbium and palpal organ as shown in 
<figureCitation id="132B19DF56510C01502080D2FC5A4DDE" box="[796,896,1473,1499]" captionStart="Figs 161–164" captionStartId="76.[189,241,1877,1903]" captionTargetBox="[197,1406,265,1871]" captionTargetId="figure@76.[197,1406,265,1875]" captionText="Figs 161–164. Raveniola spp., ♂♂. Palpal tibia, cymbium and palpal organ, retrolateral views. — 161.R.zaitzevi(Charitonov,1948)(Car).162.R.marusikisp.nov.(holotype).163.R.vonwickiZonstein, 2000 (holotype). 164. R. niedermeyeri (Brignoli, 1972) (holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163476/files/figure.png" pageId="38" pageNumber="39">Fig. 163</figureCitation>
. Embolus consisting of almost straight and slightly dilated proximal part lacking keels and of sharply narrowed, spiraled and flattened apical part (
<figureCitation id="132B19DF56510C0153F9831FFEB64E22" box="[197,364,1548,1575]" captionStart="Figs 189–197" captionStartId="80.[189,241,1823,1849]" captionTargetBox="[200,1386,265,1801]" captionTargetId="figure@80.[200,1386,265,1801]" captionText="Figs 189–197. Raveniola niedermeyeri species group, holotypic (189–192, 195–197) and conspecific (193–194) ♂♂. Details of palpal organ; retrolateral (189, 191, 194–195, 197) and ventral (190, 192–193, 196) views. — 189–190. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014. 191, 192.R.marusikisp.nov.193–194. R.zaitzevi(Charitonov, 1948).195–196. R.vonwickiZonstein,2000. 197. R. niedermeyeri (Brignoli, 1972). White arrows indicate low tibial mound with sensilla. Black arrows and contour lines show a dentate vestige of the embolickeel. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.1163484" httpUri="https://zenodo.org/record/1163484/files/figure.png" pageId="38" pageNumber="39">Figs 195–196</figureCitation>
).
</paragraph>
<paragraph id="8BAF055A56510C0153818343FB094E8A" blockId="38.[189,1398,1615,1680]" pageId="38" pageNumber="39">
SPINNERETS. See 
<figureCitation id="132B19DF56510C0152478343FE0B4E6F" box="[379,465,1616,1642]" captionStart="Figs 59–67" captionStartId="59.[189,241,1662,1688]" captionTargetBox="[189,1398,265,1617]" captionTargetId="figure@59.[189,1398,265,1617]" captionText="Figs 59–67. Raveniola spp., ♀♀ (59, 63, 66) and ♂♂ (60–62, 64–65, 67). Left chelicera (59–61; white arrow indicates male intercheliceral organ), sternum, labium and maxillae (62–64), and spinnerets (65– 67); prolateral (59–61,65, 67)and ventral (62–64,66) views.— 59–60. R.pontica (Spassky, 1937). 61, 63, 66. R. zaizevi (Charitonov, 1948). 62, 65. R. hyrcanica Dunin, 1988. 64, 67. R. vonwicki Zonstein, 2000. Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/1163442/files/figure.png" pageId="38" pageNumber="39">Fig. 67</figureCitation>
. PMS: length 0.53, diameter 0.17. PLS: maximal diameter 0.45; length of basal, medial and apical segments 0.70, 0.63, 0.77; total length 2.10; apical segment digitiform.
</paragraph>
<paragraph id="8BAF055A56510C00538183AAFE1F4920" blockId="38.[189,487,1721,1747]" lastBlockId="39.[189,1192,267,331]" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">
LEG MEASUREMENTS. ♂(♀) 
<emphasis id="B964D94856500C0053D6841FFE9F4923" bold="true" box="[234,325,268,294]" pageId="39" pageNumber="40">Female</emphasis>
(paratype)
</paragraph>
<paragraph id="8BAF055A56510C01525B83E6FABD4FDA" pageId="38" pageNumber="39">
<table id="F910F7FA5651F3D853F483E6FABD4FE5" box="[200,1383,1781,2016]" gridcols="7" gridrows="6" inLine="true" pageId="38" pageNumber="39">
<tr id="352007185651F3D853F483E6FABD4F0A" box="[200,1383,1781,1807]" gridrow="0" pageId="38" pageNumber="39" rowspan-0="1">
<th id="76F16E645651F3D8527083E6FE174F0A" box="[332,461,1781,1807]" gridcol="1" gridrow="0" pageId="38" pageNumber="39">Femur</th>
<th id="76F16E645651F3D8513D83E6FD594F0A" box="[513,643,1781,1807]" gridcol="2" gridrow="0" pageId="38" pageNumber="39">Patella</th>
<th id="76F16E645651F3D8518B83E6FCE24F0A" box="[695,824,1781,1807]" gridcol="3" gridrow="0" pageId="38" pageNumber="39">Tibia</th>
<th id="76F16E645651F3D8505083E6FC344F0A" box="[876,1006,1781,1807]" gridcol="4" gridrow="0" pageId="38" pageNumber="39">Metatarsus</th>
<th id="76F16E645651F3D8571E83E6FB794F0A" box="[1058,1187,1781,1807]" gridcol="5" gridrow="0" pageId="38" pageNumber="39">Tarsus</th>
<th id="76F16E645651F3D857F583E6FABD4F0A" box="[1225,1383,1781,1807]" gridcol="6" gridrow="0" pageId="38" pageNumber="39">Total</th>
</tr>
<tr id="352007185651F3D853F4820DFABD4F3C" box="[200,1383,1822,1849]" gridrow="1" pageId="38" pageNumber="39">
<th id="76F16E645651F3D853F4820DFEC64F3C" box="[200,284,1822,1849]" gridcol="0" gridrow="1" pageId="38" pageNumber="39">Palp</th>
<td id="76F16E645651F3D85270820DFE174F3C" box="[332,461,1822,1849]" gridcol="1" gridrow="1" pageId="38" pageNumber="39">2.75 (3.40)</td>
<td id="76F16E645651F3D8513D820DFD594F3C" box="[513,643,1822,1849]" gridcol="2" gridrow="1" pageId="38" pageNumber="39">1.82 (2.23)</td>
<td id="76F16E645651F3D8518B820DFCE24F3C" box="[695,824,1822,1849]" gridcol="3" gridrow="1" pageId="38" pageNumber="39">2.53 (2.65)</td>
<td id="76F16E645651F3D85050820DFC344F3C" box="[876,1006,1822,1849]" gridcol="4" gridrow="1" pageId="38" pageNumber="39">–</td>
<td id="76F16E645651F3D8571E820DFB794F3C" box="[1058,1187,1822,1849]" gridcol="5" gridrow="1" pageId="38" pageNumber="39">0.97 (2.17)</td>
<td id="76F16E645651F3D857F5820DFABD4F3C" box="[1225,1383,1822,1849]" gridcol="6" gridrow="1" pageId="38" pageNumber="39">8.08 (10.45)</td>
</tr>
<tr id="352007185651F3D853F4825BFABD4F66" box="[200,1383,1864,1891]" gridrow="2" pageId="38" pageNumber="39">
<th id="76F16E645651F3D853F4825BFEC64F66" box="[200,284,1864,1891]" gridcol="0" gridrow="2" pageId="38" pageNumber="39">Leg I</th>
<td id="76F16E645651F3D85270825BFE174F66" box="[332,461,1864,1891]" gridcol="1" gridrow="2" pageId="38" pageNumber="39">4.47 (4.47)</td>
<td id="76F16E645651F3D8513D825BFD594F66" box="[513,643,1864,1891]" gridcol="2" gridrow="2" pageId="38" pageNumber="39">2.85 (3.07)</td>
<td id="76F16E645651F3D8518B825BFCE24F66" box="[695,824,1864,1891]" gridcol="3" gridrow="2" pageId="38" pageNumber="39">3.93 (3.05)</td>
<td id="76F16E645651F3D85050825BFC344F66" box="[876,1006,1864,1891]" gridcol="4" gridrow="2" pageId="38" pageNumber="39">3.55 (2.65)</td>
<td id="76F16E645651F3D8571E825BFB794F66" box="[1058,1187,1864,1891]" gridcol="5" gridrow="2" pageId="38" pageNumber="39">2.20 (2.03)</td>
<td id="76F16E645651F3D857F5825BFABD4F66" box="[1225,1383,1864,1891]" gridcol="6" gridrow="2" pageId="38" pageNumber="39">17.00 (15.27)</td>
</tr>
<tr id="352007185651F3D853F48262FABD4F89" box="[200,1383,1905,1932]" gridrow="3" pageId="38" pageNumber="39">
<th id="76F16E645651F3D853F48262FEC64F89" box="[200,284,1905,1932]" gridcol="0" gridrow="3" pageId="38" pageNumber="39">Leg II</th>
<td id="76F16E645651F3D852708262FE174F89" box="[332,461,1905,1932]" gridcol="1" gridrow="3" pageId="38" pageNumber="39">4.10 (4.05)</td>
<td id="76F16E645651F3D8513D8262FD594F89" box="[513,643,1905,1932]" gridcol="2" gridrow="3" pageId="38" pageNumber="39">2.43 (2.85)</td>
<td id="76F16E645651F3D8518B8262FCE24F89" box="[695,824,1905,1932]" gridcol="3" gridrow="3" pageId="38" pageNumber="39">3.30 (3.13)</td>
<td id="76F16E645651F3D850508262FC344F89" box="[876,1006,1905,1932]" gridcol="4" gridrow="3" pageId="38" pageNumber="39">3.25 (2.70)</td>
<td id="76F16E645651F3D8571E8262FB794F89" box="[1058,1187,1905,1932]" gridcol="5" gridrow="3" pageId="38" pageNumber="39">2.13 (2.07)</td>
<td id="76F16E645651F3D857F58262FABD4F89" box="[1225,1383,1905,1932]" gridcol="6" gridrow="3" pageId="38" pageNumber="39">15.21 (14.80)</td>
</tr>
<tr id="352007185651F3D853F48288FABD4FB3" box="[200,1383,1947,1974]" gridrow="4" pageId="38" pageNumber="39">
<th id="76F16E645651F3D853F48288FEC64FB3" box="[200,284,1947,1974]" gridcol="0" gridrow="4" pageId="38" pageNumber="39">Leg III</th>
<td id="76F16E645651F3D852708288FE174FB3" box="[332,461,1947,1974]" gridcol="1" gridrow="4" pageId="38" pageNumber="39">3.65 (3.73)</td>
<td id="76F16E645651F3D8513D8288FD594FB3" box="[513,643,1947,1974]" gridcol="2" gridrow="4" pageId="38" pageNumber="39">2.10 (2.15)</td>
<td id="76F16E645651F3D8518B8288FCE24FB3" box="[695,824,1947,1974]" gridcol="3" gridrow="4" pageId="38" pageNumber="39">2.25 (2.63)</td>
<td id="76F16E645651F3D850508288FC344FB3" box="[876,1006,1947,1974]" gridcol="4" gridrow="4" pageId="38" pageNumber="39">3.77 (3.50)</td>
<td id="76F16E645651F3D8571E8288FB794FB3" box="[1058,1187,1947,1974]" gridcol="5" gridrow="4" pageId="38" pageNumber="39">2.20 (2.07)</td>
<td id="76F16E645651F3D857F58288FABD4FB3" box="[1225,1383,1947,1974]" gridcol="6" gridrow="4" pageId="38" pageNumber="39">13.97 (14.08)</td>
</tr>
<tr id="352007185651F3D853F482D6FABD4FE5" box="[200,1383,1989,2016]" gridrow="5" pageId="38" pageNumber="39">
<th id="76F16E645651F3D853F482D6FEC64FE5" box="[200,284,1989,2016]" gridcol="0" gridrow="5" pageId="38" pageNumber="39">Leg IV</th>
<td id="76F16E645651F3D8527082D6FE174FE5" box="[332,461,1989,2016]" gridcol="1" gridrow="5" pageId="38" pageNumber="39">4.57 (4.70)</td>
<td id="76F16E645651F3D8513D82D6FD594FE5" box="[513,643,1989,2016]" gridcol="2" gridrow="5" pageId="38" pageNumber="39">2.17 (2.63)</td>
<td id="76F16E645651F3D8518B82D6FCE24FE5" box="[695,824,1989,2016]" gridcol="3" gridrow="5" pageId="38" pageNumber="39">3.97 (3.87)</td>
<td id="76F16E645651F3D8505082D6FC344FE5" box="[876,1006,1989,2016]" gridcol="4" gridrow="5" pageId="38" pageNumber="39">5.85 (4.95)</td>
<td id="76F16E645651F3D8571E82D6FB794FE5" box="[1058,1187,1989,2016]" gridcol="5" gridrow="5" pageId="38" pageNumber="39">2.43 (2.27)</td>
<td id="76F16E645651F3D857F582D6FABD4FE5" box="[1225,1383,1989,2016]" gridcol="6" gridrow="5" pageId="38" pageNumber="39">17.99 (18.42)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BAF055A56500C0053818423FB72494F" blockId="39.[189,1192,267,331]" box="[189,1192,304,331]" pageId="39" pageNumber="40">MEASUREMENTS. TBL 16.50, CL 6.45, CW 5.67, LL 0.57, LW 1.10, SL 3.25, SW 2.87.</paragraph>
<paragraph id="8BAF055A56500C0053818460FE714989" blockId="39.[189,427,370,397]" box="[189,427,370,397]" pageId="39" pageNumber="40">COLOUR. As in male.</paragraph>
<paragraph id="8BAF055A56500C00538184A5FB614A38" blockId="39.[189,1399,437,573]" pageId="39" pageNumber="40">
PROSOMA. Carapace and chelicerae as shown in 
<figureCitation id="132B19DF56500C00503E84A6FC8649CA" box="[770,860,437,463]" captionStart="Figs 90–95" captionStartId="64.[189,241,1891,1917]" captionTargetBox="[189,1398,265,1846]" captionTargetId="figure@64.[189,1398,265,1849]" captionText="Figs 90–95. Raveniola spp., ♀♀. Carapace and chelicerae, dorsal views. — 90. R. turcica sp. nov. (paratype). 91. R. micropa (Ausserer, 1871) (holotype). 92. Same species as preceding (Uludağ). 93. R. zaitzevi(Charitonov, 1948)(Lagodekhi).94.R.vonwickiZonstein, 2000(paratype).95.R.niedermeyeri (Brignoli, 1972) (paratype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163452/files/figure.png" pageId="39" pageNumber="40">Fig. 94</figureCitation>
. Clypeus and eye group as in 
<figureCitation id="132B19DF56500C0057EE84A6FAE049CA" box="[1234,1338,437,463]" captionStart="Figs 117–128" captionStartId="67.[189,241,1621,1647]" captionTargetBox="[193,1382,278,1581]" captionTargetId="figure@67.[189,1398,265,1581]" captionText="Figs 117–128. Raveniola spp., ♀♀. Eyes and clypeus, dorsal views. — 117. R. adjarica sp. nov. (paratype).118. R.dunini sp. nov.(paratype).119.R.hyrcanica Dunin, 1988(paratype).120. R.pontica (Spassky, 1937) (Lazarevskoe). 121. R. sinani sp. nov. (holotype). 122. R. nana sp. nov. (holotype). 123. R. turcica sp. nov. (paratype). 124. R. micropa (Ausserer, 1871) (holotype). 125. Same species as preceding (Uludağ). 126. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 127. R. vonwicki Zonstein, 2000 (paratype). 128. R. niedermeyeri (Brignoli, 1972) (paratype). Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/1163458/files/figure.png" pageId="39" pageNumber="40">Fig. 127</figureCitation>
. Eye diameters and interdistances: AME 0.18(0.23), ALE 0.31, PLE 0.20, PME 0.18, AME–AME 0.22(0.17), ALE–AME 0.12(0.09), ALE–PLE 0.13, PLE–PME 0.05, PME–PME 0.54. Each cheliceral furrow with 10 promarginal teeth and 6–8 mesobasal denticles. Maxillae with 22–23 cuspules each.
</paragraph>
<paragraph id="8BAF055A56500C0053818775FB184ACD" blockId="39.[189,1399,613,713]" pageId="39" pageNumber="40">LEGS. Scopula: distal on metatarsi I–II, entire on palpal tarsus and tarsi I–II, absent on tarsi III–IV. Trichobothria: 2 rows of 5–7 each on tibiae, 7–11 on metatarsi, 8–10 on tarsi. Paired tarsal claws: outer margin with 5–6 teeth, 5–7 teeth on inner margin. Palpal claw with 5 promarginal teeth.</paragraph>
<paragraph id="8BAF055A56500C00538187E2FC6D4BC7" blockId="39.[189,1399,753,962]" pageId="39" pageNumber="40">SPINATION. All femora with one basodorsal spine and 3–4 stiff dorsal bristles (weak spines) located medially and distally; patella I aspinose. Palp: femur pd1; patella p1; tibia v2–2–3; tarsus v3(8). Leg I: femur pd1; tibia p1–1, v2–2–3; metatarsus v1–2–2(2–2–3). Leg II: femur pd1; patella p1; tibia p1–1, v2–2–3; metatarsus v2–2–3. Leg III: femur pd1, rd1–1; patella p1, r1; tibia d1(0), p1–1, r1–1, v2–2–3; metatarsus d1–1, p1–1–1–1, r1–1–1–1, v2–2–2–3. Leg IV: femur pd1, rd1; patella p1, r1; tibia d1, p1–1, r1–1, v2–2–3; metatarsus d1–1, p1–1–1–1, r1–1–1–1, v2–2–2–3.</paragraph>
<paragraph id="8BAF055A56500C00538186F8FB704C00" blockId="39.[189,1194,1002,1029]" box="[189,1194,1002,1029]" pageId="39" pageNumber="40">
SPERMATHECAE. Narrow and relatively long, with stalked lateral receptacles (
<figureCitation id="132B19DF56500C00570A86F9FB444C00" box="[1078,1182,1002,1029]" captionStart="Figs 204–212" captionStartId="82.[189,241,1656,1682]" captionTargetBox="[198,1407,277,1637]" captionTargetId="figure@82.[198,1407,265,1652]" captionText="Figs 204–212. Raveniola spp., ♀♀. Vulva, dorsal views. — 204. R. nana sp. nov. (holotype). 205. R. turcica sp. nov. (paratype). 206. R. micropa (Ausserer, 1871) (Uludağ). 207. Same aspecies as preceding (holotype). 208. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 209. Same species as preceding (Pirkuli). 210. R. vonwicki Zonstein, 2000 (paratype). 211. R. niedermeyeri (Brignoli, 1972) (paratype, Gorgan). 212. Same species as preceding (Mazandaran). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163488/files/figure.png" pageId="39" pageNumber="40">Fig. 210</figureCitation>
).
</paragraph>
<paragraph id="8BAF055A56500C005381813EFB904C69" blockId="39.[189,1398,1069,1132]" pageId="39" pageNumber="40">SPINNERETS. PMS: length 0.73, diameter 0.25. PLS: maximal diameter 0.70; length of basal, medial and apical segments 1.15, 0.85, 1.10; total length 3.10; apical segment digitiform.</paragraph>
</subSubSection>
<subSubSection id="C30A56D156500C0053818186FEE34CDB" box="[189,313,1173,1246]" pageId="39" pageNumber="40" type="biology_ecology">
<paragraph id="8BAF055A56500C0053818186FEFA4CAA" blockId="39.[189,288,1173,1199]" box="[189,288,1173,1199]" pageId="39" pageNumber="40">
<heading id="D0E7B23656500C0053818186FEFA4CAA" bold="true" box="[189,288,1173,1199]" fontSize="11" level="3" pageId="39" pageNumber="40" reason="3">
<emphasis id="B964D94856500C0053818186FEFA4CAA" bold="true" box="[189,288,1173,1199]" pageId="39" pageNumber="40">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56500C00538181D7FEE34CDB" blockId="39.[189,313,1220,1246]" box="[189,313,1220,1246]" pageId="39" pageNumber="40">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C30A56D156500C0053818014FD834D54" box="[189,601,1287,1361]" pageId="39" pageNumber="40" type="distribution">
<paragraph id="8BAF055A56500C0053818014FE8C4D24" blockId="39.[189,342,1287,1313]" box="[189,342,1287,1313]" pageId="39" pageNumber="40">
<heading id="D0E7B23656500C0053818014FE8C4D24" bold="true" box="[189,342,1287,1313]" fontSize="11" level="3" pageId="39" pageNumber="40" reason="3">
<emphasis id="B964D94856500C0053818014FE8C4D24" bold="true" box="[189,342,1287,1313]" pageId="39" pageNumber="40">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56500C0053818024FD834D54" blockId="39.[189,601,1335,1361]" box="[189,601,1335,1361]" pageId="39" pageNumber="40">Known only from the type locality.</paragraph>
</subSubSection>
</treatment>
</document>