treatments-xml/data/00/05/87/000587EFFFADFFC267F7FAC4351CFBC7.xml

<document id="B1FFD1817AF8A01F2FB9E20BAC13BDDC" ID-DOI="10.11646/zootaxa.3990.1.6" ID-GBIF-Dataset="7c9f16b9-348c-4225-8e93-cfc58555f348" ID-ISSN="1175-5326" ID-Zenodo-Dep="238553" ID-ZooBank="915ED168-EDD0-4008-BCC4-9E7ACCE6655A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461267109910" checkinUser="plazi" docAuthor="Craig, Matthew T., Bogorodsky, Sergey V., Randall, John E. &amp; Mal, Ahmad O." docDate="2015" docId="000587EFFFADFFC267F7FAC4351CFBC7" docLanguage="en" docName="zt03990p122.pdf" docOrigin="Zootaxa 3990 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepadichthys erythraeus Briggs &amp; Link 1963" docType="treatment" docVersion="8" lastPageNumber="121" masterDocId="FC3CFF97FFA9FFCA6760FFF83122FFBC" masterDocTitle="Lepadichthys bilineatus, a new species of clingfish from Oman (Teleostei: Gobiesocidae), with a redescription of Lepadichthys erythraeus Briggs and Link from the Red Sea" masterLastPageNumber="122" masterPageNumber="113" pageNumber="117" updateTime="1698610771167" updateUser="plazi">
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<mods:title id="A6CBCCD1666C16C4A1E23A5608ED4C69">Lepadichthys bilineatus, a new species of clingfish from Oman (Teleostei: Gobiesocidae), with a redescription of Lepadichthys erythraeus Briggs and Link from the Red Sea</mods:title>
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<mods:namePart id="6C7DE14E715FBC14D2FCE5704EEEDD48">Craig, Matthew T.</mods:namePart>
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<mods:namePart id="73EC818829CBB95FA8909854107C3888">Bogorodsky, Sergey V.</mods:namePart>
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<treatment id="000587EFFFADFFC267F7FAC4351CFBC7" ID-DOI="http://doi.org/10.5281/zenodo.5677305" ID-GBIF-Taxon="119638094" ID-Zenodo-Dep="5677305" LSID="urn:lsid:plazi:treatment:000587EFFFADFFC267F7FAC4351CFBC7" httpUri="http://treatment.plazi.org/id/000587EFFFADFFC267F7FAC4351CFBC7" lastPageId="8" lastPageNumber="121" pageId="4" pageNumber="117">
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<paragraph id="881336F9FFADFFCE67F7FAC4339CFAEA" blockId="4.[151,702,1340,1399]" box="[151,702,1340,1366]" pageId="4" pageNumber="117">
<heading id="D35B8195FFADFFCE67F7FAC4339CFAEA" bold="true" box="[151,702,1340,1366]" fontSize="11" level="1" pageId="4" pageNumber="117" reason="1">
<taxonomicName id="4FAC4D7AFFADFFCE67F7FAC4339CFAEA" ID-CoL="3T7J9" authority="Briggs &amp; Link 1963" authorityName="Briggs &amp; Link" authorityYear="1963" box="[151,702,1340,1366]" class="Actinopterygii" family="Gobiesocidae" genus="Lepadichthys" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="117" phylum="Chordata" rank="species" species="erythraeus">
<emphasis id="BAD8EAEBFFADFFCE67F7FAC4339CFAEA" bold="true" box="[151,702,1340,1366]" pageId="4" pageNumber="117">
<emphasis id="BAD8EAEBFFADFFCE67F7FAC430E2FAEA" bold="true" box="[151,448,1340,1366]" italics="true" pageId="4" pageNumber="117">Lepadichthys erythraeus</emphasis>
<bibRefCitation id="EC3D4B08FFADFFCE66A7FAC4339CFAEA" author="Briggs" box="[455,702,1340,1366]" pageId="4" pageNumber="117" refString="Briggs, J. C. &amp; Link, G. (1963) New clingfishes of the genus Lepadichthys from the northern Indian Ocean and Red Sea. Senckenbergiana Biologica, 44 (2), 101 - 105." type="journal article" year="1963">Briggs &amp; Link 1963</bibRefCitation>
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</paragraph>
<paragraph id="881336F9FFADFFCE67F7FAA6305EFACB" blockId="4.[151,702,1340,1399]" box="[151,380,1374,1399]" pageId="4" pageNumber="117">
<figureCitation id="10972A7CFFADFFCE67F7FAA6303DFACB" box="[151,287,1374,1399]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="5.[151,250,1447,1469]" captionStartId-1="6.[151,250,1464,1486]" captionStartId-2="7.[151,250,1942,1964]" captionTargetBox-0="[150,1436,481,1424]" captionTargetBox-1="[389,1195,198,1427]" captionTargetBox-2="[179,1407,265,1919]" captionTargetId-0="figure@5.[150,1436,481,1425]" captionTargetId-1="figure@6.[374,1212,193,1443]" captionTargetId-2="figure@7.[179,1408,265,1920]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURE 2 Lepadichthys erythraeus, SMF 5586, holotype, 36.6 mm SL, Ghardaqa, Egypt. Photo by S. Traenkner." captionText-1="FIGURE 3. Sensory head pores of Lepadichthys erythraeus. LC = Lachrymal canal pores, NC = Nasal canal pores, PO = Postocular canal pores, PR = Preopercular canal pores. In L. erythraeus there are 3 PR pores; the third is visible only on the ventral surface and cannot be seen in profile." captionText-2="FIGURE 4. Lepadichthys erythraeus. A: uncatalogued, 23.5 mm SL, Sharm el Moya, Egypt, Red Sea; B: uncatalogued, 36 mm SL, Sharm el Moya Bay, Egypt, Red Sea. C: BPBM 40985, 19.1 mm SL, El Quseir, Egypt, Red Sea. Photos by S. V. Bogorodsky." httpUri-0="https://zenodo.org/record/238555/files/figure.png" httpUri-1="https://zenodo.org/record/238556/files/figure.png" httpUri-2="https://zenodo.org/record/238557/files/figure.png" pageId="4" pageNumber="117">Figures 2–4</figureCitation>
, 
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<taxonomicName id="4FAC4D7AFFADFFCE67F7FA5E33BBFA00" ID-CoL="3T7J9" authority="Briggs &amp; Link 1963:102" authorityName="Briggs &amp; Link" authorityPageNumber="102" authorityYear="1963" box="[151,665,1446,1468]" class="Actinopterygii" family="Gobiesocidae" genus="Lepadichthys" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="117" phylum="Chordata" rank="species" species="erythraeus">
<emphasis id="BAD8EAEBFFADFFCE67F7FA5E30B0FA00" box="[151,402,1446,1468]" italics="true" pageId="4" pageNumber="117">Lepadichthys erythraeus</emphasis>
<treatmentCitation id="090D10E8FFADFFCE66FAFA5E33BBFA00" author="Briggs" box="[410,665,1446,1468]" page="102" pageId="4" pageNumber="117" year="1963">
<bibRefCitation id="EC3D4B08FFADFFCE66FAFA5E334CFA00" author="Briggs" box="[410,622,1446,1468]" pageId="4" pageNumber="117" refString="Briggs, J. C. &amp; Link, G. (1963) New clingfishes of the genus Lepadichthys from the northern Indian Ocean and Red Sea. Senckenbergiana Biologica, 44 (2), 101 - 105." type="journal article" year="1963">Briggs &amp; Link 1963</bibRefCitation>
:102
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(Hurghada (Al-Ghardaqa), Governorate, 
<collectingCountry id="F0BB7669FFADFFCE6325FA5F35A6FA00" box="[1093,1156,1447,1468]" name="Egypt" pageId="4" pageNumber="117">Egypt</collectingCountry>
, Red Sea). 
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, SMF 5586, 
<quantity id="4F549B1CFFADFFCE6666FA3D3042FA67" box="[262,352,1477,1499]" metricMagnitude="-2" metricUnit="m" metricValue="3.66" pageId="4" pageNumber="117" unit="mm" value="36.6">36.6 mm</quantity>
SL (
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).
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</paragraph>
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<treatmentCitationGroup id="A8BC11D7FFADFFCE67F7FA1C31D9F9A4" pageId="4" pageNumber="117">
<taxonomicName id="4FAC4D7AFFADFFCE67F7FA1C330FFA46" ID-CoL="3T7J9" authority="Dor 1984: 54" authorityName="Dor" authorityPageNumber="54" authorityYear="1984" box="[151,557,1508,1530]" class="Actinopterygii" family="Gobiesocidae" genus="Lepadichthys" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="117" phylum="Chordata" rank="species" species="erythraeus">
<emphasis id="BAD8EAEBFFADFFCE67F7FA1C30B0FA46" box="[151,402,1508,1530]" italics="true" pageId="4" pageNumber="117">Lepadichthys erythraeus</emphasis>
: 
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<bibRefCitation id="EC3D4B08FFADFFCE66C2FA1D332BFA46" author="Dor" box="[418,521,1508,1530]" pageId="4" pageNumber="117" refString="Dor, M. (1984) CLOFRES. Checklist of fishes of the Red Sea. The Israeli Academy of Sciences and Humanities, Jerusalem, xxii + 437 pp." type="book" year="1984">Dor 1984</bibRefCitation>
: 54
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(Red Sea); 
<treatmentCitation id="090D10E8FFADFFCE65CBFA1C32B8FA46" author="Goren" box="[683,922,1508,1530]" page="13" pageId="4" pageNumber="117" year="1994">
<bibRefCitation id="EC3D4B08FFADFFCE65CBFA1C3257FA46" author="Goren" box="[683,885,1508,1530]" pageId="4" pageNumber="117" refString="Goren, M. &amp; Dor, M. (1994) An updated checklist of the fishes of the Red Sea. CLOFRES II. The Israel Academy of Sciences and Humanities, Jerusalem, xiii + 120 pp." type="book" year="1994">Goren &amp; Dor 1994</bibRefCitation>
: 13
</treatmentCitation>
(Red Sea); 
<treatmentCitation id="090D10E8FFADFFCE6379FA1C3440FA46" author="Golani" box="[1049,1378,1508,1530]" page="45" pageId="4" pageNumber="117" year="2010">
<bibRefCitation id="EC3D4B08FFADFFCE6379FA1C341CFA46" author="Golani" box="[1049,1342,1508,1530]" pageId="4" pageNumber="117" refString="Golani, D. &amp; Bogorodsky, S. V. (2010) The fishes of the Red Sea - reappraisal and updated checklist. Zootaxa, 2463, 1 - 135." type="journal article" year="2010">Golani &amp; Bogorodsky 2010</bibRefCitation>
: 45
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(Red Sea).
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</paragraph>
</subSubSection>
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<emphasis id="BAD8EAEBFFADFFCE67F7F9BD3033F9E2" bold="true" box="[151,273,1605,1630]" pageId="4" pageNumber="117">Diagnosis.</emphasis>
A species of 
<taxonomicName id="4FAC4D7AFFADFFCE66C7F9BD331EF9E1" box="[423,572,1605,1629]" class="Actinopterygii" family="Gobiesocidae" genus="Lepadichthys" kingdom="Animalia" order="Gobiesociformes" pageId="4" pageNumber="117" phylum="Chordata" rank="genus">
<emphasis id="BAD8EAEBFFADFFCE66C7F9BD331EF9E1" box="[423,572,1605,1629]" italics="true" pageId="4" pageNumber="117">Lepadichthys</emphasis>
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with a membrane joining dorsal and anal fins with the caudal fin; anus closer to anal fin than to disc, about two-thirds the distance from the rear margin of the disc to the origin of anal fin; upper attachment of gill membrane opposite ninth pectoral-fin ray; eight rakers on second gill arch; postdorsal-caudal distance 
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dorsal-fin length. Dorsal-fin rays 15–16; anal-fin rays 13; pectoral-fin rays 26–28 (rarely 28); principle caudal-fin rays 12; total caudal-fin rays 20.
</paragraph>
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<subSubSection id="C0B66572FFADFFCF67A7F901342AF86C" lastPageId="5" lastPageNumber="118" pageId="4" pageNumber="117" type="description">
<paragraph id="881336F9FFADFFCE67A7F90130F7F8C2" blockId="4.[151,1436,1605,2026]" pageId="4" pageNumber="117">
<emphasis id="BAD8EAEBFFADFFCE67A7F901307BF8AE" bold="true" box="[199,345,1785,1810]" pageId="4" pageNumber="117">Description.</emphasis>
Count and proportion for 
<typeStatus id="5717885BFFADFFCE65EBF90133CFF8AE" box="[651,749,1785,1810]" pageId="4" pageNumber="117" type="holotype">holotype</typeStatus>
given in parentheses; data for each examined specimen are presented in 
<tableCitation id="C52E0342FFADFFCE664DF8E530A6F88A" box="[301,388,1821,1846]" captionStart="TABLE 1" captionStartId="3.[125,212,483,504]" captionTargetBox="[125,1863,551,1725]" captionTargetPageId="3" captionText="TABLE 1. Morphometric (expressed as % SL) and meristic data of Lepadichthys bilineatus n. sp. and L. erythraeus. Count of caudal-fin rays given as number of principle rays followed by total rays in parentheses." httpUri="http://table.plazi.org/id/DCD36671FFAAFFC9671DFE1B33B1FDA8" pageId="4" pageNumber="117" tableUuid="DCD36671FFAAFFC9671DFE1B33B1FDA8">Table 1</tableCitation>
. Dorsal-fin rays 15–16 (15); anal-fin rays 13 (13); pectoral-fin rays 26–27 (28); principle caudal-fin rays 12 (12); total caudal-fin rays 20 (20); gill rakers on second gill arch 8 (8); pre-anal vertebrae 21 (21); total vertebrae 34 (34).
</paragraph>
<paragraph id="881336F9FFADFFCE67A7F87134BEF856" blockId="4.[151,1436,1605,2026]" pageId="4" pageNumber="117">Body slender, compressed posteriorly, anterior portion being flattened dorsally; body depth 6.9–8.1 (6.9) in SL, body width 5.9–7.6 (6.4) in SL, body narrower in smaller specimens. Caudal peduncle very short, depth 0.5– 0.6 (0.6) in its length. Head narrow and depressed, its length 2.8–3.2 (2.9) in SL, its width 5.4–6.2 (5.2) in SL.</paragraph>
<paragraph id="881336F9FFACFFCF67F7FF603088FE11" blockId="5.[151,1437,151,429]" pageId="5" pageNumber="118">Snout relatively long, giving specimen a “pointed” appearance in lateral view, but rounded in dorsal outline, its length 2.8–4.0 (3.2) in HL, snout shorter in smaller specimens. Mouth slightly inferior, oblique, and moderately large, but not extending below anterior margin of eye, upper-jaw length 3.1–4.3 (4.1) in HL. A single row of teeth on each jaw, those on lower jaw exclusively conical in shape, and those on upper jaw are compressed and have reversed tips. Upper edge of eye slightly protruding above dorsal profile of head, eye diameter 5.0–6.4 (5.7) in HL; interorbital space moderately broad, flat, distance 3.3–5.3 (3.9) in HL, interorbital broader in larger specimens. Anterior nostril lacking dermal flap; posterior nostril located slightly behind anterior edge of orbit and contains pronounced dermal flap.</paragraph>
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<paragraph id="881336F9FFACFFCF67F7FA5F3400FA01" blockId="5.[151,1314,1447,1469]" box="[151,1314,1447,1469]" pageId="5" pageNumber="118">
<emphasis id="BAD8EAEBFFACFFCF67F7FA5F302FFA00" bold="true" box="[151,269,1447,1469]" pageId="5" pageNumber="118">FIGURE 2</emphasis>
<taxonomicName id="4FAC4D7AFFACFFCF6673FA5F332EFA01" box="[275,524,1447,1469]" class="Actinopterygii" family="Gobiesocidae" genus="Lepadichthys" kingdom="Animalia" order="Gobiesociformes" pageId="5" pageNumber="118" phylum="Chordata" rank="species" species="erythraeus">
<emphasis id="BAD8EAEBFFACFFCF6673FA5F332EFA01" box="[275,524,1447,1469]" italics="true" pageId="5" pageNumber="118">Lepadichthys erythraeus</emphasis>
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, SMF 5586, holotype, 36.6 mm SL, Ghardaqa, Egypt. Photo by S. Traenkner.
</paragraph>
</caption>
<paragraph id="881336F9FFACFFCF67A7FA1D32CFF90D" blockId="5.[151,1437,1508,2001]" pageId="5" pageNumber="118">
Upper attachment of gill membrane opposite ninth pectoral-fin ray. Both dorsal and anal fins joined to caudal fin by a thin membrane. Dorsal-fin origin behind vertical at anus and forward to anal-fin origin, predorsal length 1.4–1.7 (1.5) in SL. Postdorsal-caudal distance 
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dorsal-fin length. Dorsal-caudal distance when brought forward reaches about middle of pectoral fin. Disc single and small, its length 6.6–7.6 (6.3) in SL. Disc region A with 1–2 rows of flattened papillae, disc region B with 8 rows of flattened papillae. Anus closer to anal fin than to disc, about two-thirds distance from rear margin of disc to origin of anal fin.
</paragraph>
<paragraph id="881336F9FFACFFCF67A7F94433F1F945" blockId="5.[151,1437,1508,2001]" pageId="5" pageNumber="118">
Cephalic pores systems (
<figureCitation id="10972A7CFFACFFCF6688F9443313F969" box="[488,561,1724,1749]" captionStart="FIGURE 3" captionStartId="6.[151,250,1464,1486]" captionTargetBox="[389,1195,198,1427]" captionTargetId="figure@6.[374,1212,193,1443]" captionTargetPageId="6" captionText="FIGURE 3. Sensory head pores of Lepadichthys erythraeus. LC = Lachrymal canal pores, NC = Nasal canal pores, PO = Postocular canal pores, PR = Preopercular canal pores. In L. erythraeus there are 3 PR pores; the third is visible only on the ventral surface and cannot be seen in profile." httpUri="https://zenodo.org/record/238556/files/figure.png" pageId="5" pageNumber="118">Fig. 3</figureCitation>
): There are 2 pores in nasal canal, 2 pores in postorbital canal, 2 pores in lachrymal canal, and 3 pores in preopercular canal.
</paragraph>
<paragraph id="881336F9FFACFFCF67A7F8FC342AF86C" blockId="5.[151,1437,1508,2001]" pageId="5" pageNumber="118">
Coloration in life (
<figureCitation id="10972A7CFFACFFCF66F5F8FC30C0F8A1" box="[405,482,1796,1821]" captionStart="FIGURE 4" captionStartId="7.[151,250,1942,1964]" captionTargetBox="[179,1407,265,1919]" captionTargetId="figure@7.[179,1408,265,1920]" captionTargetPageId="7" captionText="FIGURE 4. Lepadichthys erythraeus. A: uncatalogued, 23.5 mm SL, Sharm el Moya, Egypt, Red Sea; B: uncatalogued, 36 mm SL, Sharm el Moya Bay, Egypt, Red Sea. C: BPBM 40985, 19.1 mm SL, El Quseir, Egypt, Red Sea. Photos by S. V. Bogorodsky." httpUri="https://zenodo.org/record/238557/files/figure.png" pageId="5" pageNumber="118">Figs. 4</figureCitation>
A–B) generally reddish brown with small black flecks irregularly spaced on body and a distinct dark (black to brown) stripe running from tip of snout, through eye, and terminating about half the distance from rear margin of orbit to gill opening. Some individuals with a pale streak originating on tip of both jaws, running over head between the eyes, and terminating just posterior to them. Base of pectoral fin darker than body color, but not as dark as stripe running through eye. Pupil dark with a distinct yellow ring on its outer margin. Median fins dark with light margins. In preservative, all color fades and specimens are uniformly tan.
</paragraph>
</subSubSection>
<caption id="DCD36671FFAFFFCC67F7FA40337CF9B7" httpUri="https://zenodo.org/record/238556/files/figure.png" pageId="6" pageNumber="119" targetBox="[389,1195,198,1427]" targetPageId="6">
<paragraph id="881336F9FFAFFFCC67F7FA40337CF9B7" blockId="6.[151,1436,1464,1547]" pageId="6" pageNumber="119">
<emphasis id="BAD8EAEBFFAFFFCC67F7FA403032FA71" bold="true" box="[151,272,1464,1486]" pageId="6" pageNumber="119">FIGURE 3</emphasis>
. Sensory head pores of 
<taxonomicName id="4FAC4D7AFFAFFFCC6573FA403232FA72" box="[531,784,1464,1486]" class="Actinopterygii" family="Gobiesocidae" genus="Lepadichthys" kingdom="Animalia" order="Gobiesociformes" pageId="6" pageNumber="119" phylum="Chordata" rank="species" species="erythraeus">
<emphasis id="BAD8EAEBFFAFFFCC6573FA403232FA72" box="[531,784,1464,1486]" italics="true" pageId="6" pageNumber="119">Lepadichthys erythraeus</emphasis>
</taxonomicName>
. LC = Lachrymal canal pores, NC= Nasal canal pores, PO = Postocular canal pores, PR = Preopercular canal pores. In 
<taxonomicName id="4FAC4D7AFFAFFFCC6591FA203258FA51" box="[753,890,1495,1517]" class="Actinopterygii" family="Gobiesocidae" genus="Lepadichthys" kingdom="Animalia" order="Gobiesociformes" pageId="6" pageNumber="119" phylum="Chordata" rank="species" species="erythraeus">
<emphasis id="BAD8EAEBFFAFFFCC6591FA203258FA51" box="[753,890,1495,1517]" italics="true" pageId="6" pageNumber="119">L. erythraeus</emphasis>
</taxonomicName>
there are 3 PR pores; the third is visible only on the ventral surface and cannot be seen in profile.
</paragraph>
</caption>
<subSubSection id="C0B66572FFAFFFCC67A7F9C0321AF9C9" pageId="6" pageNumber="119" type="distribution">
<paragraph id="881336F9FFAFFFCC67A7F9C0321AF9C9" blockId="6.[151,1437,1592,2013]" pageId="6" pageNumber="119">
<emphasis id="BAD8EAEBFFAFFFCC67A7F9C03042F9ED" bold="true" box="[199,352,1592,1617]" pageId="6" pageNumber="119">Distribution.</emphasis>
<taxonomicName id="4FAC4D7AFFAFFFCC6607F9C03359F9EC" box="[359,635,1592,1616]" class="Actinopterygii" family="Gobiesocidae" genus="Lepadichthys" kingdom="Animalia" order="Gobiesociformes" pageId="6" pageNumber="119" phylum="Chordata" rank="species" species="erythraeus">
<emphasis id="BAD8EAEBFFAFFFCC6607F9C03359F9EC" box="[359,635,1592,1616]" italics="true" pageId="6" pageNumber="119">Lepadichthys erythraeus</emphasis>
</taxonomicName>
is presently known only from the Red Sea, from the Gulf of Aqaba and Gulf of Suez, south to Al Wajh (northeastern 
<collectingCountry id="F0BB7669FFAFFFCC65F4F9A43209F9C9" box="[660,811,1628,1653]" name="Saudi Arabia" pageId="6" pageNumber="119">Saudi Arabia</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C0B66572FFAFFFC267A7F9783335FE54" lastPageId="8" lastPageNumber="121" pageId="6" pageNumber="119" type="discussion">
<paragraph id="881336F9FFAFFFCD67A7F97830E9FF69" blockId="6.[151,1437,1592,2013]" lastBlockId="7.[151,1436,151,213]" lastPageId="7" lastPageNumber="120" pageId="6" pageNumber="119">
<emphasis id="BAD8EAEBFFAFFFCC67A7F9783019F925" bold="true" box="[199,315,1664,1689]" pageId="6" pageNumber="119">Remarks.</emphasis>
A specimen of 
<taxonomicName id="4FAC4D7AFFAFFFCC668BF97833DDF924" box="[491,767,1664,1688]" class="Actinopterygii" family="Gobiesocidae" genus="Lepadichthys" kingdom="Animalia" order="Gobiesociformes" pageId="6" pageNumber="119" phylum="Chordata" rank="species" species="erythraeus">
<emphasis id="BAD8EAEBFFAFFFCC668BF97833DDF924" box="[491,767,1664,1688]" italics="true" pageId="6" pageNumber="119">Lepadichthys erythraeus</emphasis>
</taxonomicName>
(BPBM 40985) was collected by the second author in 2008 and is the first specimen of this species to be photographed in color (
<figureCitation id="10972A7CFFAFFFCC64F2F95C32F4F901" box="[914,982,1700,1725]" captionStart="FIGURE 4" captionStartId="7.[151,250,1942,1964]" captionTargetBox="[179,1407,265,1919]" captionTargetId="figure@7.[179,1408,265,1920]" captionTargetPageId="7" captionText="FIGURE 4. Lepadichthys erythraeus. A: uncatalogued, 23.5 mm SL, Sharm el Moya, Egypt, Red Sea; B: uncatalogued, 36 mm SL, Sharm el Moya Bay, Egypt, Red Sea. C: BPBM 40985, 19.1 mm SL, El Quseir, Egypt, Red Sea. Photos by S. V. Bogorodsky." httpUri="https://zenodo.org/record/238557/files/figure.png" pageId="6" pageNumber="119">Fig. 4</figureCitation>
C). The specimen was collected from the base of a coral block (about 
<quantity id="4F549B1CFFAFFFCC66BDF930333CF95C" box="[477,542,1736,1761]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="119" unit="m" value="1.5">1.5 m</quantity>
in diameter) on a reef flat with mixed sand and flat rock with patches of dead corals. The specimen was collected from within this live coral head in a depth of about 
<quantity id="4F549B1CFFAFFFCC63CEF91535FCF8B8" box="[1198,1246,1773,1797]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="119" unit="m" value="1.0">1 m</quantity>
in the sheltered Mangrove Bay (El Quseir, 
<collectingCountry id="F0BB7669FFAFFFCC66A8F8E93332F895" box="[456,528,1809,1833]" name="Egypt" pageId="6" pageNumber="119">Egypt</collectingCountry>
). Later, in 2011 one specimen was collected by the second author from Sharm el Moya area (Sharm el Sheikh, 
<collectingCountry id="F0BB7669FFAFFFCC6683F8CD330AF8F1" box="[483,552,1845,1869]" name="Egypt" pageId="6" pageNumber="119">Egypt</collectingCountry>
) from a fringing reef at a depth of 
<quantity id="4F549B1CFFAFFFCC64C9F8CD32D2F8F0" box="[937,1008,1845,1869]" metricMagnitude="0" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="6" pageNumber="119" unit="m" value="2.0" valueMax="3.0" valueMin="1.0">1–3 m</quantity>
and three specimens were collected at the entrance of Sharm el Moya Bay (
<collectingCountry id="F0BB7669FFAFFFCC6522F8A133AAF8CD" box="[578,648,1881,1905]" name="Egypt" pageId="6" pageNumber="119">Egypt</collectingCountry>
) from an isolated coral block. All collected specimens were in close association with sea urchins of the genus 
<taxonomicName id="4FAC4D7AFFAFFFCC6505F88533E9F829" box="[613,715,1917,1941]" class="Echinoidea" family="Diadematidae" genus="Diadema" kingdom="Animalia" order="Diadematoida" pageId="6" pageNumber="119" phylum="Echinodermata" rank="genus">
<emphasis id="BAD8EAEBFFAFFFCC6505F88533E9F829" box="[613,715,1917,1941]" italics="true" pageId="6" pageNumber="119">Diadema</emphasis>
</taxonomicName>
, utilizing the long spines for shelter. During recent field work in 
<collectingCountry id="F0BB7669FFAFFFCC67F7F8583008F805" box="[151,298,1952,1977]" name="Saudi Arabia" pageId="6" pageNumber="119">Saudi Arabia</collectingCountry>
in June of 2013 additional specimens were collected by the second author in the northern part of the Red Sea (Maqna, Al Khuraybah, and Al Wajh). There are several lots in USNM collected by V. G. Springer and team in the Gulf of Aqaba and Gulf of Suez in 1969, we examined one lot (USNM 314225) from the Gulf of Aqaba that contains 
<specimenCount id="9EAAFD70FFAEFFCD664CFF4430E5FF69" box="[300,455,188,213]" pageId="7" pageNumber="120" type="generic">14 specimens</specimenCount>
.
</paragraph>
<caption id="DCD36671FFAEFFCD67F7F86E303AF855" httpUri="https://zenodo.org/record/238557/files/figure.png" pageId="7" pageNumber="120" targetBox="[179,1407,265,1919]" targetPageId="7">
<paragraph id="881336F9FFAEFFCD67F7F86E303AF855" blockId="7.[151,1436,1942,2025]" pageId="7" pageNumber="120">
<emphasis id="BAD8EAEBFFAEFFCD67F7F86E302DF817" bold="true" box="[151,271,1942,1964]" pageId="7" pageNumber="120">FIGURE 4</emphasis>
. 
<taxonomicName id="4FAC4D7AFFAEFFCD667DF86E333AF810" box="[285,536,1942,1964]" class="Actinopterygii" family="Gobiesocidae" genus="Lepadichthys" kingdom="Animalia" order="Gobiesociformes" pageId="7" pageNumber="120" phylum="Chordata" rank="species" species="erythraeus">
<emphasis id="BAD8EAEBFFAEFFCD667DF86E333AF810" box="[285,536,1942,1964]" italics="true" pageId="7" pageNumber="120">Lepadichthys erythraeus</emphasis>
</taxonomicName>
. 
<emphasis id="BAD8EAEBFFAEFFCD6547F86E331BF817" bold="true" box="[551,569,1942,1963]" pageId="7" pageNumber="120">A</emphasis>
: uncatalogued, 23.5 mm SL, Sharm el Moya, Egypt, Red Sea; 
<emphasis id="BAD8EAEBFFAEFFCD63ADF86E35FCF817" bold="true" box="[1229,1246,1942,1963]" pageId="7" pageNumber="120">B</emphasis>
: uncatalogued, 36 mm SL, Sharm el Moya Bay, Egypt, Red Sea. 
<emphasis id="BAD8EAEBFFAEFFCD65E9F84D33B9F877" bold="true" box="[649,667,1973,1995]" pageId="7" pageNumber="120">C</emphasis>
: BPBM 40985, 19.1 mm SL, El Quseir, Egypt, Red Sea. Photos by S.V. Bogorodsky.
</paragraph>
</caption>
<paragraph id="881336F9FFA1FFC267A7FF60356FFE3C" blockId="8.[151,1437,151,1147]" pageId="8" pageNumber="121">
We noted two characters in our examination that differed from those reported in the original description, the first being the number of dorsal-fin rays. 
<bibRefCitation id="EC3D4B08FFA1FFC2651EFF42325FFF6F" author="Briggs" box="[638,893,186,211]" pageId="8" pageNumber="121" refString="Briggs, J. C. &amp; Link, G. (1963) New clingfishes of the genus Lepadichthys from the northern Indian Ocean and Red Sea. Senckenbergiana Biologica, 44 (2), 101 - 105." type="journal article" year="1963">Briggs &amp; Link (1963)</bibRefCitation>
did not use x-rays to count these elements (J. Briggs, pers. comm.), and this difference is readily explained by the difficulty in accurately counting the smallest of these rays, particularly at the origin of the dorsal fin. Second, the position of the anus was described as being “much closer” to the origin of the anal fin than to the rear margin of the disc. This subjective term makes interpretation difficult, and we herein report a semi-quantitative measure for the position of the anus as roughly two-thirds the distance from the anterior edge of the disc to the origin of the anal fin.
</paragraph>
<paragraph id="881336F9FFA1FFC267A7FE723335FE54" blockId="8.[151,1437,151,1147]" pageId="8" pageNumber="121">
The position and number of the sensory head pores in 
<taxonomicName id="4FAC4D7AFFA1FFC26449FE73329CFE1F" box="[809,958,395,419]" class="Actinopterygii" family="Gobiesocidae" genus="Lepadichthys" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="121" phylum="Chordata" rank="species" species="erythraeus">
<emphasis id="BAD8EAEBFFA1FFC26449FE73329CFE1F" box="[809,958,395,419]" italics="true" pageId="8" pageNumber="121">L. erythraeus</emphasis>
</taxonomicName>
matches well with other known species of 
<taxonomicName id="4FAC4D7AFFA1FFC267F7FE553372FE7A" authority="Shiogaki &amp; Dotsu 1983" authorityName="Shiogaki &amp; Dotsu" authorityYear="1983" box="[151,592,429,454]" class="Actinopterygii" family="Gobiesocidae" genus="Lepadichthys" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="121" phylum="Chordata" rank="genus">
<emphasis id="BAD8EAEBFFA1FFC267F7FE55300EFE79" box="[151,300,429,453]" italics="true" pageId="8" pageNumber="121">Lepadichthys</emphasis>
(
<bibRefCitation id="EC3D4B08FFA1FFC2665AFE55336AFE7A" author="Shiogaki" box="[314,584,429,454]" pageId="8" pageNumber="121" refString="Shiogaki, M. &amp; Dotsu, Y. (1983) Two new genera and two new species of clingfishes from Japan, with comments on head sensory canals of the Gobiesocidae. Japanese Journal of Ichthyology, 30 (2), 111 - 121." type="journal article" year="1983">Shiogaki &amp; Dotsu 1983</bibRefCitation>
)
</taxonomicName>
. We thus agree with 
<bibRefCitation id="EC3D4B08FFA1FFC26456FE553572FE7A" author="Shiogaki" box="[822,1104,429,454]" pageId="8" pageNumber="121" refString="Shiogaki, M. &amp; Dotsu, Y. (1983) Two new genera and two new species of clingfishes from Japan, with comments on head sensory canals of the Gobiesocidae. Japanese Journal of Ichthyology, 30 (2), 111 - 121." type="journal article" year="1983">Shiogaki &amp; Dotsu (1983)</bibRefCitation>
that these canals may be well suited to identify clingfish genera.
</paragraph>
</subSubSection>
<subSubSection id="C0B66572FFA1FFC267A7FE0B356EFC16" pageId="8" pageNumber="121" type="reference_group">
<paragraph id="881336F9FFA1FFC267A7FE0B3289FCFE" blockId="8.[151,1437,151,1147]" pageId="8" pageNumber="121">
<taxonomicName id="4FAC4D7AFFA1FFC267A7FE0B30FEFDB7" ID-CoL="3T7J9" box="[199,476,499,523]" class="Actinopterygii" family="Gobiesocidae" genus="Lepadichthys" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="121" phylum="Chordata" rank="species" species="erythraeus">
<emphasis id="BAD8EAEBFFA1FFC267A7FE0B30FEFDB7" box="[199,476,499,523]" italics="true" pageId="8" pageNumber="121">Lepadichthys erythraeus</emphasis>
</taxonomicName>
superficially resembles 
<taxonomicName id="4FAC4D7AFFA1FFC2658FFE0B3518FDB7" ID-CoL="3T7J7" authority="Regan 1921" authorityName="Regan" authorityYear="1921" box="[751,1082,499,523]" class="Actinopterygii" family="Gobiesocidae" genus="Lepadichthys" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="121" phylum="Chordata" rank="species" species="coccinotaenia">
<emphasis id="BAD8EAEBFFA1FFC2658FFE0B3288FDB6" box="[751,938,499,522]" italics="true" pageId="8" pageNumber="121">L. coccinotaenia</emphasis>
<bibRefCitation id="EC3D4B08FFA1FFC264D3FE0B3518FDB7" author="Regan" box="[947,1082,499,523]" pageId="8" pageNumber="121" refString="Regan, C. T. (1921) Three new fishes from South Africa, collected by Mr. H. W. Bell Marley. Annals of the Durban Museum, 3, 1 - 2." type="journal article" year="1921">Regan 1921</bibRefCitation>
</taxonomicName>
known from the Indian Ocean. Both species possess a dark band extending from the snout, through the eye, and terminating on the operculum. 
<taxonomicName id="4FAC4D7AFFA1FFC267F7FDC03089FDEC" ID-CoL="3T7J9" box="[151,427,568,592]" class="Actinopterygii" family="Gobiesocidae" genus="Lepadichthys" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="121" phylum="Chordata" rank="species" species="erythraeus">
<emphasis id="BAD8EAEBFFA1FFC267F7FDC03089FDEC" box="[151,427,568,592]" italics="true" pageId="8" pageNumber="121">Lepadichthys erythraeus</emphasis>
</taxonomicName>
has more dorsal-fin and anal-fin rays, 15–16 and 13, respectively 
<emphasis id="BAD8EAEBFFA1FFC263ECFDC13581FDEC" box="[1164,1187,569,592]" italics="true" pageId="8" pageNumber="121">vs</emphasis>
. 
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11–13 and 
<quantity id="4F549B1CFFA1FFC2624BFDC03440FDEC" box="[1323,1378,568,592]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" metricValueMax="2.794" metricValueMin="2.286" pageId="8" pageNumber="121" unit="in" value="10.0" valueMax="11.0" valueMin="9.0">9–11</quantity>
</date>
in 
<taxonomicName id="4FAC4D7AFFA1FFC262E5FDC13016FDCE" ID-CoL="3T7J7" class="Actinopterygii" family="Gobiesocidae" genus="Lepadichthys" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="121" phylum="Chordata" rank="species" species="coccinotaenia">
<emphasis id="BAD8EAEBFFA1FFC262E5FDC13016FDCE" italics="true" pageId="8" pageNumber="121">L. coccinotaenia</emphasis>
</taxonomicName>
; the dorsal and anal fins joined with the caudal by a thin membrane 
<emphasis id="BAD8EAEBFFA1FFC263C9FDA335E5FDCE" box="[1193,1223,603,626]" italics="true" pageId="8" pageNumber="121">vs.</emphasis>
not joined in 
<taxonomicName id="4FAC4D7AFFA1FFC262E5FDA33016FD29" ID-CoL="3T7J7" class="Actinopterygii" family="Gobiesocidae" genus="Lepadichthys" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="121" phylum="Chordata" rank="species" species="coccinotaenia">
<emphasis id="BAD8EAEBFFA1FFC262E5FDA33016FD29" italics="true" pageId="8" pageNumber="121">L. coccinotaenia</emphasis>
</taxonomicName>
; the postdorsal-caudal distance is shorter, 
<quantity id="4F549B1CFFA1FFC2646EFD85325DFD2A" box="[782,895,637,662]" metricMagnitude="-1" metricUnit="m" metricValue="1.7399" metricValueMax="1.8541999999999998" metricValueMin="1.6256" pageId="8" pageNumber="121" unit="in" value="6.85" valueMax="7.3" valueMin="6.4">6.4–7.3 in</quantity>
dorsal-fin length 
<emphasis id="BAD8EAEBFFA1FFC26326FD863546FD29" box="[1094,1124,638,661]" italics="true" pageId="8" pageNumber="121">vs.</emphasis>
<quantity id="4F549B1CFFA1FFC2630BFD8635FEFD2A" box="[1131,1244,637,662]" metricMagnitude="-2" metricUnit="m" metricValue="6.985" metricValueMax="7.111999999999999" metricValueMin="6.858" pageId="8" pageNumber="121" unit="in" value="2.75" valueMax="2.8" valueMin="2.7">2.7–2.8 in</quantity>
dorsal-fin length in 
<taxonomicName id="4FAC4D7AFFA1FFC267DAFD593058FD0B" ID-CoL="3T7J7" box="[186,378,672,696]" class="Actinopterygii" family="Gobiesocidae" genus="Lepadichthys" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="121" phylum="Chordata" rank="species" species="coccinotaenia">
<emphasis id="BAD8EAEBFFA1FFC267DAFD593058FD0B" box="[186,378,672,696]" italics="true" pageId="8" pageNumber="121">L. coccinotaenia</emphasis>
</taxonomicName>
; and the caudal peduncle is shorter, 
<quantity id="4F549B1CFFA1FFC2645CFD673291FD04" box="[828,947,671,696]" metricMagnitude="-2" metricUnit="m" metricValue="1.397" metricValueMax="1.524" metricValueMin="1.27" pageId="8" pageNumber="121" unit="in" value="0.55" valueMax="0.6" valueMin="0.5">0.5–0.6 in</quantity>
its length 
<emphasis id="BAD8EAEBFFA1FFC26359FD59357AFD04" box="[1081,1112,673,696]" italics="true" pageId="8" pageNumber="121">vs.</emphasis>
<quantity id="4F549B1CFFA1FFC26305FD5835F9FD04" box="[1125,1243,671,696]" metricMagnitude="-2" metricUnit="m" metricValue="1.905" metricValueMax="2.032" metricValueMin="1.7779999999999998" pageId="8" pageNumber="121" unit="in" value="0.75" valueMax="0.8" valueMin="0.7">0.7–0.8 in</quantity>
its length in 
<taxonomicName id="4FAC4D7AFFA1FFC262E5FD593016FD66" ID-CoL="3T7J7" class="Actinopterygii" family="Gobiesocidae" genus="Lepadichthys" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="121" phylum="Chordata" rank="species" species="coccinotaenia">
<emphasis id="BAD8EAEBFFA1FFC262E5FD593016FD66" italics="true" pageId="8" pageNumber="121">L. coccinotaenia</emphasis>
</taxonomicName>
. 
<taxonomicName id="4FAC4D7AFFA1FFC26625FD3B337FFD67" ID-CoL="3T7J9" box="[325,605,707,731]" class="Actinopterygii" family="Gobiesocidae" genus="Lepadichthys" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="121" phylum="Chordata" rank="species" species="erythraeus">
<emphasis id="BAD8EAEBFFA1FFC26625FD3B337FFD67" box="[325,605,707,731]" italics="true" pageId="8" pageNumber="121">Lepadichthys erythraeus</emphasis>
</taxonomicName>
is also similar in morphology to 
<taxonomicName id="4FAC4D7AFFA1FFC26487FD3B35BDFD67" ID-CoL="6P8BW" box="[999,1183,707,731]" class="Actinopterygii" family="Gobiesocidae" genus="Lepadichthys" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="121" phylum="Chordata" rank="species" species="sandaracatus">
<emphasis id="BAD8EAEBFFA1FFC26487FD3B35BDFD67" box="[999,1183,707,731]" italics="true" pageId="8" pageNumber="121">L. sandaracatus</emphasis>
</taxonomicName>
known from Western 
<collectingCountry id="F0BB7669FFA1FFC267F7FD1D3023FD42" box="[151,257,741,766]" name="Australia" pageId="8" pageNumber="121">Australia</collectingCountry>
, but 
<taxonomicName id="4FAC4D7AFFA1FFC26659FD1E30F2FD41" ID-CoL="3T7J9" box="[313,464,741,765]" class="Actinopterygii" family="Gobiesocidae" genus="Lepadichthys" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="121" phylum="Chordata" rank="species" species="erythraeus">
<emphasis id="BAD8EAEBFFA1FFC26659FD1E30F2FD41" box="[313,464,741,765]" italics="true" pageId="8" pageNumber="121">L. erythraeus</emphasis>
</taxonomicName>
has the upper attachment of the gill membrane opposite the ninth pectoral-fin ray 
<emphasis id="BAD8EAEBFFA1FFC2621DFD1E34BEFD41" box="[1405,1436,742,765]" italics="true" pageId="8" pageNumber="121">vs.</emphasis>
opposite the fourth to seventh pectoral-fin ray in 
<taxonomicName id="4FAC4D7AFFA1FFC265A4FCF1325BFC9C" ID-CoL="6P8BW" box="[708,889,776,800]" class="Actinopterygii" family="Gobiesocidae" genus="Lepadichthys" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="121" phylum="Chordata" rank="species" species="sandaracatus">
<emphasis id="BAD8EAEBFFA1FFC265A4FCF1325BFC9C" box="[708,889,776,800]" italics="true" pageId="8" pageNumber="121">L. sandaracatus</emphasis>
</taxonomicName>
; and shorter postdorsal-caudal distance, 
<quantity id="4F549B1CFFA1FFC26228FCFF318FFCFF" metricMagnitude="-1" metricUnit="m" metricValue="1.7399" metricValueMax="1.8541999999999998" metricValueMin="1.6256" pageId="8" pageNumber="121" unit="in" value="6.85" valueMax="7.3" valueMin="6.4">6.4–7.3 in</quantity>
dorsal-fin length 
<emphasis id="BAD8EAEBFFA1FFC26614FCD330B1FCFE" box="[372,403,811,834]" italics="true" pageId="8" pageNumber="121">vs.</emphasis>
<quantity id="4F549B1CFFA1FFC266FAFCD33329FCFF" box="[410,523,810,835]" metricMagnitude="-2" metricUnit="m" metricValue="9.143999999999998" metricValueMax="9.652" metricValueMin="8.636" pageId="8" pageNumber="121" unit="in" value="3.5999999999999996" valueMax="3.8" valueMin="3.4">3.4–3.8 in</quantity>
dorsal-fin length in 
<taxonomicName id="4FAC4D7AFFA1FFC2658FFCD33286FCFF" ID-CoL="6P8BW" box="[751,932,811,835]" class="Actinopterygii" family="Gobiesocidae" genus="Lepadichthys" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="121" phylum="Chordata" rank="species" species="sandaracatus">
<emphasis id="BAD8EAEBFFA1FFC2658FFCD33286FCFF" box="[751,932,811,835]" italics="true" pageId="8" pageNumber="121">L. sandaracatus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="881336F9FFA1FFC267A7FCB5356EFC16" blockId="8.[151,1437,151,1147]" pageId="8" pageNumber="121">
Only one congeneric species occurs in the Red Sea, 
<taxonomicName id="4FAC4D7AFFA1FFC26471FCB632AAFCD9" ID-CoL="3T7JC" box="[785,904,845,869]" class="Actinopterygii" family="Gobiesocidae" genus="Lepadichthys" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="121" phylum="Chordata" rank="species" species="lineatus">
<emphasis id="BAD8EAEBFFA1FFC26471FCB632AAFCD9" box="[785,904,845,869]" italics="true" pageId="8" pageNumber="121">L. lineatus</emphasis>
</taxonomicName>
, which clearly differs from 
<taxonomicName id="4FAC4D7AFFA1FFC263A2FCB63475FCD9" ID-CoL="3T7J9" box="[1218,1367,845,869]" class="Actinopterygii" family="Gobiesocidae" genus="Lepadichthys" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="121" phylum="Chordata" rank="species" species="erythraeus">
<emphasis id="BAD8EAEBFFA1FFC263A2FCB63475FCD9" box="[1218,1367,845,869]" italics="true" pageId="8" pageNumber="121">L. erythraeus</emphasis>
</taxonomicName>
by its striped color pattern and close association with crinoids. Some authors have placed this species in the genus 
<taxonomicName id="4FAC4D7AFFA1FFC267F7FC6B303BFC16" ID-CoL="46TY" box="[151,281,915,938]" class="Actinopterygii" family="Gobiesocidae" genus="Discotrema" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="121" phylum="Chordata" rank="genus">
<emphasis id="BAD8EAEBFFA1FFC267F7FC6B303BFC16" box="[151,281,915,938]" italics="true" pageId="8" pageNumber="121">Discotrema</emphasis>
</taxonomicName>
, but 
<bibRefCitation id="EC3D4B08FFA1FFC26631FC6A30CFFC17" author="Briggs" box="[337,493,914,939]" pageId="8" pageNumber="121" refString="Briggs, J. C. (2001) New species of Lepadichthys from the Philippine Islands. Copeia, 2001 (2), 499 - 500. http: // dx. doi. org / 10.1643 / 0045 - 8511 (2001) 001 [0499: NSOLFT] 2.0. CO; 2" type="journal article" year="2001">Briggs (2001)</bibRefCitation>
and 
<bibRefCitation id="EC3D4B08FFA1FFC26544FC6A320DFC17" author="Craig" box="[548,815,914,939]" pageId="8" pageNumber="121" refString="Craig, M. T. &amp; Randall, J. E. (2008) Two new species of the Indo-Pacific clingfish genus Discotrema (Gobiesocidae). Copeia, 2008 (1), 68 - 74. http: // dx. doi. org / 10.1643 / CI- 07 - 025" type="journal article" year="2008">Craig &amp; Randall (2008)</bibRefCitation>
retain it in 
<taxonomicName id="4FAC4D7AFFA1FFC264D0FC6B3567FC17" ID-CoL="6398T" box="[944,1093,915,939]" class="Actinopterygii" family="Gobiesocidae" genus="Lepadichthys" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="121" phylum="Chordata" rank="genus">
<emphasis id="BAD8EAEBFFA1FFC264D0FC6B3567FC17" box="[944,1093,915,939]" italics="true" pageId="8" pageNumber="121">Lepadichthys</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C0B66572FFA1FFC267A7FC4D351CFBC7" pageId="8" pageNumber="121" type="materials_examined">
<paragraph id="881336F9FFA1FFC267A7FC4D351CFBC7" blockId="8.[151,1437,151,1147]" pageId="8" pageNumber="121">
<emphasis id="BAD8EAEBFFA1FFC267A7FC4D3090FC72" bold="true" box="[199,434,949,974]" pageId="8" pageNumber="121">Material examined.</emphasis>
<collectionCode id="EEBDAE3CFFA1FFC266DCFC4E30D6FC72" LSID="urn:lsid:biocol.org:col:34838" box="[444,500,950,974]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="8" pageNumber="121">SMF</collectionCode>
5586, 
<typeStatus id="5717885BFFA1FFC26525FC4D3388FC72" box="[581,682,949,974]" pageId="8" pageNumber="121" type="holotype">holotype</typeStatus>
, 
<quantity id="4F549B1CFFA1FFC265D7FC4D323DFC71" box="[695,799,949,974]" metricMagnitude="-2" metricUnit="m" metricValue="3.66" pageId="8" pageNumber="121" unit="mm" value="36.6">36.6 mm</quantity>
, 
<collectingCountry id="F0BB7669FFA1FFC2644DFC4E3251FC72" box="[813,883,950,974]" name="Egypt" pageId="8" pageNumber="121">Egypt</collectingCountry>
, Hurghada (Al-Ghardaqa); 
<collectionCode id="EEBDAE3CFFA1FFC263CDFC4E35DCFC72" LSID="urn:lsid:biocol.org:col:34871" box="[1197,1278,950,974]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="121">USNM</collectionCode>
314225, 
<quantity id="4F549B1CFFA1FFC2620BFC4D31E7FC53" metricMagnitude="-2" metricUnit="m" metricValue="2.85" pageId="8" pageNumber="121" unit="mm" value="28.5">28.5 mm</quantity>
, 
<collectingCountry id="F0BB7669FFA1FFC267B2FC2F3031FC4C" box="[210,275,983,1008]" name="Israel" pageId="8" pageNumber="121">Israel</collectingCountry>
, El Himeira, 
<date id="FC121039FFA1FFC266CBFC203360FC4C" box="[427,578,983,1008]" pageId="8" pageNumber="121" value="1969-07-19">19 July 1969</date>
; 
<collectionCode id="EEBDAE3CFFA1FFC2652FFC2033BFFC53" LSID="urn:lsid:biocol.org:col:34697" box="[591,669,984,1007]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="8" pageNumber="121">BPBM</collectionCode>
40985, 
<quantity id="4F549B1CFFA1FFC2659AFC203243FC53" box="[762,865,984,1008]" metricMagnitude="-2" metricUnit="m" metricValue="1.91" pageId="8" pageNumber="121" unit="mm" value="19.1">19.1 mm</quantity>
, 
<collectingCountry id="F0BB7669FFA1FFC2640EFC203296FC4C" box="[878,948,984,1008]" name="Egypt" pageId="8" pageNumber="121">Egypt</collectingCountry>
, El Quseir, 
<date id="FC121039FFA1FFC2635AFC2035F7FC4C" box="[1082,1237,984,1008]" pageId="8" pageNumber="121" value="2008-06-12">12 June 2008</date>
; uncatalogued, 1, 
<quantity id="4F549B1CFFA1FFC267F7FC0231DCFBAE" box="[151,254,1018,1043]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" pageId="8" pageNumber="121" unit="mm" value="23.5">23.5 mm</quantity>
, 
<collectingCountry id="F0BB7669FFA1FFC2666CFC033073FBAF" box="[268,337,1019,1043]" name="Egypt" pageId="8" pageNumber="121">Egypt</collectingCountry>
, Sharm el Moya, 
<date id="FC121039FFA1FFC2657DFC02339AFBAF" box="[541,696,1018,1043]" pageId="8" pageNumber="121" value="2011-06-25">25 June 2011</date>
; uncatalogued, 3, 33.0–42.0 mm, 
<collectingCountry id="F0BB7669FFA1FFC2635BFC0335A3FBAF" box="[1083,1153,1019,1043]" name="Egypt" pageId="8" pageNumber="121">Egypt</collectingCountry>
, Sharm el Moya Bay, 
<date id="FC121039FFA1FFC262E0FC033024FB8A" pageId="8" pageNumber="121" value="2011-07-13">13 July 2011</date>
; 
<collectionCode id="EEBDAE3CFFA1FFC26672FBE63068FB8A" LSID="urn:lsid:biocol.org:col:34838" box="[274,330,1054,1078]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="8" pageNumber="121">SMF</collectionCode>
35410, 2, 26.6–36.0 mm, 
<collectingCountry id="F0BB7669FFA1FFC26512FBE53225FB8A" box="[626,775,1053,1078]" name="Saudi Arabia" pageId="8" pageNumber="121">Saudi Arabia</collectingCountry>
, south of Al Wajh, 
<date id="FC121039FFA1FFC26480FBE6355AFB8A" box="[992,1144,1054,1078]" pageId="8" pageNumber="121" value="2013-06-14">14 June 2013</date>
; 
<collectionCode id="EEBDAE3CFFA1FFC263E4FBE6359EFB8A" LSID="urn:lsid:biocol.org:col:34838" box="[1156,1212,1054,1078]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="8" pageNumber="121">SMF</collectionCode>
35411, 1, 
<quantity id="4F549B1CFFA1FFC26252FBE634BAFB89" box="[1330,1432,1054,1078]" metricMagnitude="-2" metricUnit="m" metricValue="3.79" pageId="8" pageNumber="121" unit="mm" value="37.9">37.9 mm</quantity>
, 
<collectingCountry id="F0BB7669FFA1FFC267F7FBC7300FFBE4" box="[151,301,1087,1112]" name="Saudi Arabia" pageId="8" pageNumber="121">Saudi Arabia</collectingCountry>
, Al Khuraybah, 
<date id="FC121039FFA1FFC26688FBB833A1FBE4" box="[488,643,1088,1112]" pageId="8" pageNumber="121" value="2013-06-24">24 June 2013</date>
; 
<collectionCode id="EEBDAE3CFFA1FFC265EFFBB833DFFBE4" box="[655,765,1088,1112]" pageId="8" pageNumber="121">KAUMM</collectionCode>
318, 2, 
<quantity id="4F549B1CFFA1FFC2643BFBC73522FBEB" box="[859,1024,1087,1112]" metricMagnitude="-2" metricUnit="m" metricValue="3.105" metricValueMax="3.62" metricValueMin="2.59" pageId="8" pageNumber="121" unit="mm" value="31.05" valueMax="36.2" valueMin="25.9">25.9–36.2 mm</quantity>
, 
<collectingCountry id="F0BB7669FFA1FFC2636DFBC73586FBE4" box="[1037,1188,1087,1112]" name="Saudi Arabia" pageId="8" pageNumber="121">Saudi Arabia</collectingCountry>
, south of Al Wajh, 
<date id="FC121039FFA1FFC262E0FBB8302FFBC7" pageId="8" pageNumber="121" value="2013-06-14">14 June 2013</date>
; 
<collectionCode id="EEBDAE3CFFA1FFC26678FB9B30A4FBC7" box="[280,390,1123,1147]" pageId="8" pageNumber="121">KAUMM</collectionCode>
319, 1, 
<quantity id="4F549B1CFFA1FFC26681FB9A3365FBC6" box="[481,583,1122,1147]" metricMagnitude="-2" metricUnit="m" metricValue="3.53" pageId="8" pageNumber="121" unit="mm" value="35.3">35.3 mm</quantity>
, 
<collectingCountry id="F0BB7669FFA1FFC26533FB9A33CBFBC7" box="[595,745,1122,1147]" name="Saudi Arabia" pageId="8" pageNumber="121">Saudi Arabia</collectingCountry>
, Al Khuraybah, 
<date id="FC121039FFA1FFC264C2FB9B3519FBC7" box="[930,1083,1123,1147]" pageId="8" pageNumber="121" value="2013-06-24">24 June 2013</date>
.
</paragraph>
</subSubSection>
</treatment>
</document>