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<document id="AE219DCFA2D03699654FFA54CE587A19" ENCODING="Cp1252" ID-GBIF-Dataset="2a850022-3fbc-4643-a17a-8f7bca314619" ModsDocID="20017" checkinTime="1252781348276" checkinUser="donat" docAuthor="Wilson, E. O." docDate="2003" docId="176FF54F7C2656D5B5B410F0BBFBB71A" docLanguage="en" docName="20017_p_and_more_group_gg1.xml" docOrigin="Cambridge, MA: Harvard University Press" docSource="http://atbi.biosci.ohio-state.edu/HymOnline/reference-full.html?id=20017" docTitle="Pheidole torosa Wilson, 2003, new species" docType="treatment" docVersion="13" lastPageNumber="603" masterDocId="73EDC9C5AB0725CB6B0DF69F187F7798" masterDocTitle="Pheidole in the New World. A dominant, hyperdiverse ant genus." pageNumber="603" updateTime="1701311082057" updateUser="plazi">
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<mods:title id="958E66BA94729A9E02BD15CBF696ABEC">Pheidole in the New World. A dominant, hyperdiverse ant genus.</mods:title>
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<mods:namePart id="5D3B71DB35C5D9FF680445B79AB45EAC">Wilson, E. O.</mods:namePart>
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<treatment id="176FF54F7C2656D5B5B410F0BBFBB71A" ID-DOI="http://doi.org/10.5281/zenodo.6274346" ID-GBIF-Taxon="100119683" ID-Zenodo-Dep="6274346" LSID="urn:lsid:plazi:treatment:176FF54F7C2656D5B5B410F0BBFBB71A" httpUri="http://treatment.plazi.org/id/176FF54F7C2656D5B5B410F0BBFBB71A" lastPageNumber="603" pageNumber="603">
<subSubSection id="A1629838E5723377CB68291596A86F7E" type="nomenclature">
<paragraph id="78ED384F90FC7466DC846C4D00C7716E" pageNumber="603">
<taxonomicName id="5C7D6EB2C58FA2CA7FF045B27B405C06" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181977" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole torosa Wilson" lsidName-HNS="Pheidole torosa Wilson" order="Hymenoptera" pageNumber="603" phylum="Arthropoda" rank="species" species="torosa">Pheidole torosa</taxonomicName>
<taxonomicNameLabel id="5C54C038A980112D05AA024E6A2FA031" rank="species">new species</taxonomicNameLabel>
</paragraph>
<paragraph id="F39EC9580CA30A5A861F9211B73643BA" pageNumber="603">Types Mus. Comp. Zool. Harvard.</paragraph>
</subSubSection>
<subSubSection id="D44D6469CD39BCE7B8C8ABA1EB53D186" type="etymology">
<paragraph id="9EB52B4DA0ED545BEB09D41BA93607EF" pageNumber="603">
Etymology L
<taxonomicName id="FDF0F57F4ACCF100B0580BB0B1251646" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181977" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole torosa Wilson" lsidName-HNS="Pheidole torosa Wilson" order="Hymenoptera" pageNumber="603" phylum="Arthropoda" rank="species" species="torosa">torosa</taxonomicName>
, muscular, bulging, a reference to the major caste.
</paragraph>
</subSubSection>
<subSubSection id="E203338587466F5124A2202AD7587A75" type="diagnosis">
<paragraph id="D0CE48C037313015D9FCE64E95E1CFC7" pageNumber="603">
diagnosis A member of the &quot;
<taxonomicName id="86B244A992D15214CA90DB2D23141FD6" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33930" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole pilifera (Roger)" lsidName-HNS="Pheidole pilifera (Roger)" order="Hymenoptera" pageNumber="603" phylum="Arthropoda" rank="species" species="pilifera">pilifera</taxonomicName>
complex&quot; of the larger
<taxonomicName id="600A9C716DE40E78E7D4EE5135788698" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33930" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole pilifera (Roger)" lsidName-HNS="Pheidole pilifera (Roger)" order="Hymenoptera" pageNumber="603" phylum="Arthropoda" rank="species" species="pilifera">pilifera</taxonomicName>
group, comprising
<taxonomicName id="4461C5F9D5AF28CF200075CCDE343A98" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33642" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole calens Forel" lsidName-HNS="Pheidole calens Forel" order="Hymenoptera" pageNumber="603" phylum="Arthropoda" rank="species" species="calens">calens</taxonomicName>
,
<taxonomicName id="0C69D5429216A3AFF2CD8D3523074A48" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33643" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole californica Mayr" lsidName-HNS="Pheidole californica Mayr" order="Hymenoptera" pageNumber="603" phylum="Arthropoda" rank="species" species="californica">californica</taxonomicName>
,
<taxonomicName id="9BBD7D2C52AF2B03EE74608340C39F82" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33651" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole carrolli Naves" lsidName-HNS="Pheidole carrolli Naves" order="Hymenoptera" pageNumber="603" phylum="Arthropoda" rank="species" species="carrolli">carrolli</taxonomicName>
,
<taxonomicName id="276114AA90E0E4F7ADF423C4BF471A24" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181972" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole cavigenis Wheeler" lsidName-HNS="Pheidole cavigenis Wheeler" order="Hymenoptera" pageNumber="603" phylum="Arthropoda" rank="species" species="cavigenis">cavigenis</taxonomicName>
,
<taxonomicName id="620D815270009A8B3199B8CCC004D2D9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33670" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole clementensis Gregg" lsidName-HNS="Pheidole clementensis Gregg" order="Hymenoptera" pageNumber="603" phylum="Arthropoda" rank="species" species="clementensis">clementensis</taxonomicName>
,
<taxonomicName id="8731284E247568C9AACA43B12FFFD6FD" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33691" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole creightoni Gregg" lsidName-HNS="Pheidole creightoni Gregg" order="Hymenoptera" pageNumber="603" phylum="Arthropoda" rank="species" species="creightoni">creightoni</taxonomicName>
,
<taxonomicName id="7DC946B9AAE46B9C14162F3DC65FEEA8" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181973" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole hoplitica Wilson" lsidName-HNS="Pheidole hoplitica Wilson" order="Hymenoptera" pageNumber="603" phylum="Arthropoda" rank="species" species="hoplitica">hoplitica</taxonomicName>
,
<taxonomicName id="8BED525CD85413552E90623BF8773DE5" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33833" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole littoralis Cole" lsidName-HNS="Pheidole littoralis Cole" order="Hymenoptera" pageNumber="603" phylum="Arthropoda" rank="species" species="littoralis">littoralis</taxonomicName>
,
<taxonomicName id="542843CE0C6415C7BD5FA6AA4C58D1FD" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33868" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole micula Wheeler" lsidName-HNS="Pheidole micula Wheeler" order="Hymenoptera" pageNumber="603" phylum="Arthropoda" rank="species" species="micula">micula</taxonomicName>
,
<taxonomicName id="1593EAF60E7A2C0D18A183D4B2E86BF7" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33930" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole pilifera (Roger)" lsidName-HNS="Pheidole pilifera (Roger)" order="Hymenoptera" pageNumber="603" phylum="Arthropoda" rank="species" species="pilifera">pilifera</taxonomicName>
,
<taxonomicName id="67B3AEFADEFD0DFA70880C49D27B192E" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181975" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole polymorpha Wilson" lsidName-HNS="Pheidole polymorpha Wilson" order="Hymenoptera" pageNumber="603" phylum="Arthropoda" rank="species" species="polymorpha">polymorpha</taxonomicName>
,
<taxonomicName id="EB1C5A8A85915C2059BE60BE6AEF6B14" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33980" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole rugulosa Gregg" lsidName-HNS="Pheidole rugulosa Gregg" order="Hymenoptera" pageNumber="603" phylum="Arthropoda" rank="species" species="rugulosa">rugulosa</taxonomicName>
,
<taxonomicName id="A7D8D58E255569877C13E8D13C525F17" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33998" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole senex Gregg" lsidName-HNS="Pheidole senex Gregg" order="Hymenoptera" pageNumber="603" phylum="Arthropoda" rank="species" species="senex">senex</taxonomicName>
,
<taxonomicName id="1291ED3DE02915FF7C6382AE190247A7" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142583" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole soritis Wheeler" lsidName-HNS="Pheidole soritis Wheeler" order="Hymenoptera" pageNumber="603" phylum="Arthropoda" rank="species" species="soritis">soritis</taxonomicName>
,
<taxonomicName id="F17D8B9A51F02A1E7AC92A9E0C5A4763" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:34049" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole tepicana Pergande" lsidName-HNS="Pheidole tepicana Pergande" order="Hymenoptera" pageNumber="603" phylum="Arthropoda" rank="species" species="tepicana">tepicana</taxonomicName>
, and
<taxonomicName id="9A17FF48ACE2E5DCBE16DB71BEDD2F68" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181977" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole torosa Wilson" lsidName-HNS="Pheidole torosa Wilson" order="Hymenoptera" pageNumber="603" phylum="Arthropoda" rank="species" species="torosa">torosa</taxonomicName>
, which complex is distinguished by the following traits. Major: dorsal head surface extensively sculptured; occipital lobe transversely rugulose (or, in
<taxonomicName id="408F34D6BFD9B2D66585A8D480EBD1C7" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33651" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole carrolli Naves" lsidName-HNS="Pheidole carrolli Naves" order="Hymenoptera" pageNumber="603" phylum="Arthropoda" rank="species" species="carrolli">carrolli</taxonomicName>
smooth, in
<taxonomicName id="5207025B6FA52B3339BD5488A4F5191C" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33833" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole littoralis Cole" lsidName-HNS="Pheidole littoralis Cole" order="Hymenoptera" pageNumber="603" phylum="Arthropoda" rank="species" species="littoralis">littoralis</taxonomicName>
foveate, and in
<taxonomicName id="1294E6846C60E62EF60CDF7E12367649" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33868" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole micula Wheeler" lsidName-HNS="Pheidole micula Wheeler" order="Hymenoptera" pageNumber="603" phylum="Arthropoda" rank="species" species="micula">micula</taxonomicName>
and
<taxonomicName id="F7FDDBFE2BD0BB44A2EED7D238EF7558" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142583" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole soritis Wheeler" lsidName-HNS="Pheidole soritis Wheeler" order="Hymenoptera" pageNumber="603" phylum="Arthropoda" rank="species" species="soritis">soritis</taxonomicName>
carinulate); postpetiole from above diamond-shaped, trapezoidal, or spinose. Minor: eye medium-sized to large.
<taxonomicName id="7A42224C71C16D08DAD864B2D12E8A7E" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:181977" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole torosa Wilson" lsidName-HNS="Pheidole torosa Wilson" order="Hymenoptera" pageNumber="603" phylum="Arthropoda" rank="species" species="torosa">P. torosa</taxonomicName>
is distinguished within the complex as follows.
</paragraph>
</subSubSection>
<subSubSection id="52EA1CF7EF6DEC99124051250D0C6843" type="description">
<paragraph id="640CAA27C2DEA6B73BF296B4277E6FCE" pageNumber="603">Major: bicolorous; transverse rugulae cover posterior fifth of dorsal surface of head in full-face view, and longitudinal carinulae cover all of the remainder of the dorsal surface except the frontal triangle; humerus in dorsal-oblique view prominent and lobose; in side view, mesonotum drops almost vertically to metanotum; propodeal spines very long, stout, and vertical to propodeal basal face; seen from above, postpetiole very wide relative to petiole, and spinose.</paragraph>
<paragraph id="C02691A5DF608B88629F28F9789D0C57" pageNumber="603">Minor: all of head and mesosoma foveolate and opaque; pilosity along promesonotal dorsum consists of evenly spaced pairs of setae.</paragraph>
<paragraph id="8BB4ED38A7EBC1F510BD0315787A0AB8" pageNumber="603">
This species is very close to and possibly synonymous with
<taxonomicName id="B185AB6B449C37B39F9CB80359753E26" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33998" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole senex Gregg" lsidName-HNS="Pheidole senex Gregg" order="Hymenoptera" pageNumber="603" phylum="Arthropoda" rank="species" species="senex">senex</taxonomicName>
.
</paragraph>
<paragraph id="A2E08DB57C257EEDC6CD06D62313030F" pageNumber="603">Measurements (mm) Holotype major: HW 1.56, HL 1.84, SL 0.76, EL 0.20, PW 0.68.</paragraph>
<paragraph id="9C6443091A52AA8F04C133D9765D7118" pageNumber="603">Paratype minor: HW 0.58, HL 0.64, SL 0.62, EL 0.14, PW 0.32.</paragraph>
<paragraph id="28AB7FC42C811A3AB9B6278E64C44D41" pageNumber="603">color Major: bicolorous, with light reddish brown head and appendages contrasting with dark brown mesosoma and gaster.</paragraph>
<paragraph id="D9BD34059D2866DD0DE1949802840D2F" pageNumber="603">Minor: concolorous light brown.</paragraph>
</subSubSection>
<subSubSection id="72364161F75FE0EAFBDFA5859D8B7C96" type="distribution">
<paragraph id="F73D51085B655136186BF37FC1EFD5CA" pageNumber="603">Range Known only from the type locality.</paragraph>
</subSubSection>
<subSubSection id="643AAB3DED3AED7DF055C6DC74D55E74" type="biology_ecology">
<paragraph id="F95031434627B18615BEE2974EEFD966" pageNumber="603">Biology Unknown.</paragraph>
</subSubSection>
<subSubSection id="5D9BDAA1B25E8DED359B5BAB5F725E9E" type="materials_examined">
<paragraph id="0E3556D243921D92CE773D5100936A5E" pageNumber="603">Figure Upper: holotype, major. Lower: paratype, minor. MEXICO: El Sueco, Chihuahua. Scale bars = 1 mm.</paragraph>
</subSubSection>
</treatment>
</document>