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<document id="0256042DC499DD2078D5965A682A5E88" ID-DOI="10.11646/zootaxa.4028.1.5" ID-GBIF-Dataset="51f4ec88-362b-47c8-92ce-ac81c56f91b1" ID-ISSN="1175-5326" ID-Zenodo-Dep="243995" ID-ZooBank="D762F1C0-CAB4-4163-8E3F-3B17CCFCB593" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461284645030" checkinUser="plazi" docAuthor="Mullen, Logan" docDate="2015" docId="03E1AD4D2E3568086F9AFE19FD9EFEF0" docLanguage="en" docName="zt04028p128.pdf" docOrigin="Zootaxa 4028 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phlaeopterus czerskyi Shavrin 2001, comb. nov." docType="treatment" docVersion="8" lastPageNumber="127" masterDocId="FFD8D5352E37680E6F0DFFC1FFEDFF95" masterDocTitle="Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) — a new genus for the Palaearctic: new combination" masterLastPageNumber="128" masterPageNumber="121" pageNumber="123" updateTime="1698614968450" updateUser="plazi">
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<mods:title id="0BBD18ECA8A58CE8E4241A4463A05334">Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) — a new genus for the Palaearctic: new combination</mods:title>
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<mods:name id="E986B8B18B94DDA030370D777EDA11FD" type="personal">
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<mods:namePart id="B496E91F226F972EE80EC855F4664949">Mullen, Logan</mods:namePart>
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<mods:typeOfResource id="5247CAEFF828F1763187AC891EA8DB3B">text</mods:typeOfResource>
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<mods:title id="7253767A1B4AEE9C5790B92A52BA7874">Zootaxa</mods:title>
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<mods:date id="A01E9203A2AA1E79BC2EDC6DA6004248">2015</mods:date>
<mods:detail id="C8E6CB59EB64F61546717AB3C664C406" type="volume">
<mods:number id="C24CC5F12EE947BBEF2EC3BBB397F3F6">4028</mods:number>
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<mods:classification id="9BD82AB5AA9750199A1841AAEDEF064A">journal article</mods:classification>
<mods:identifier id="2E275A099793B8631199D09BA2E33CE4" type="DOI">10.11646/zootaxa.4028.1.5</mods:identifier>
<mods:identifier id="E1CFA69B795236461285EF387ADBD5FA" type="GBIF-Dataset">51f4ec88-362b-47c8-92ce-ac81c56f91b1</mods:identifier>
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<treatment id="03E1AD4D2E3568086F9AFE19FD9EFEF0" ID-DOI="http://doi.org/10.5281/zenodo.6108662" ID-GBIF-Taxon="119641116" ID-Zenodo-Dep="6108662" LSID="urn:lsid:plazi:treatment:03E1AD4D2E3568086F9AFE19FD9EFEF0" httpUri="http://treatment.plazi.org/id/03E1AD4D2E3568086F9AFE19FD9EFEF0" lastPageId="6" lastPageNumber="127" pageId="2" pageNumber="123">
<subSubSection id="C3524FD02E35680C6F9AFE19FECDFD81" pageId="2" pageNumber="123" type="nomenclature">
<paragraph id="8BF71C5B2E35680C6F9AFE19FD1FFE66" blockId="2.[151,754,472,533]" box="[151,754,472,499]" pageId="2" pageNumber="123">
<heading id="D0BFAB372E35680C6F9AFE19FD1FFE66" bold="true" box="[151,754,472,499]" fontSize="11" level="1" pageId="2" pageNumber="123" reason="1">
<emphasis id="B93CC0492E35680C6F9AFE19FD1FFE66" bold="true" box="[151,754,472,499]" pageId="2" pageNumber="123">
<taxonomicName id="4C4867D82E35680C6F9AFE19FD88FE66" ID-CoL="76ZZV" authority="Shavrin, 2001" authorityName="Shavrin" authorityYear="2001" box="[151,613,472,499]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="123" phylum="Arthropoda" rank="species" species="czerskyi" status="comb. nov.">
<emphasis id="B93CC0492E35680C6F9AFE19FE73FE67" bold="true" box="[151,414,472,498]" italics="true" pageId="2" pageNumber="123">Phlaeopterus czerskyi</emphasis>
(
<bibRefCitation id="EFD961AA2E35680C6EA0FE18FDB1FE66" author="Shavrin" box="[429,604,473,499]" pageId="2" pageNumber="123" refString="Shavrin, A. V. (2001) New and little-known species of Omaliinae from the Baikal-Transbaikal area (Coleoptera: Staphylinidae). Zoosystematica Rossica, 9, 189 - 193." type="journal article" year="2001">Shavrin, 2001</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A20F7D322E35680C6D61FE18FD1FFE66" box="[620,754,473,499]" pageId="2" pageNumber="123">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF71C5B2E35680C6F9AFE3DFECDFD81" blockId="2.[151,754,472,533]" box="[151,288,508,533]" pageId="2" pageNumber="123">
(
<figureCitation id="137300DE2E35680C6F92FE3DFEFAFD81" box="[159,279,508,533]" captionStart-0="FIGURES 1 – 2" captionStart-1="FIGURES 3 – 9" captionStart-2="FIGURES 10 – 11" captionStart-3="FIGURE 12" captionStartId-0="2.[151,264,1863,1886]" captionStartId-1="3.[151,264,1944,1967]" captionStartId-2="4.[151,264,1268,1291]" captionStartId-3="5.[151,250,1914,1936]" captionTargetBox-0="[165,1426,1057,1836]" captionTargetBox-1="[217,1376,831,1929]" captionTargetBox-2="[215,1371,184,1238]" captionTargetBox-3="[209,1378,1273,1892]" captionTargetId-0="figure@2.[151,1436,1050,1840]" captionTargetId-1="figure@3.[205,1382,823,1937]" captionTargetId-2="figure@4.[204,1383,175,1248]" captionTargetId-3="figure@5.[209,1378,1273,1893]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURES 1 – 2. Habitus, female (Fig. 1) and habitat (Fig. 2) in upper of Babkha River (Eastern Siberia, Khamar-Daban Mts.) of Phlaeopterus czerskyi (Shavrin, 2001). Scale bar: 1.0 mm." captionText-1="FIGURES 3 – 9. Phlaeopterus czerskyi (Shavrin, 2001): 3 — labrum (dorsal view), 4 — left and right mandibles (dorsal view), 5 — labium (dorsal view), 6 — mentum (dorsal view), 7 — right maxilla (dorsal view), 8 — male abdominal sternite VIII (dorsal view), 9 — male abdominal tergite VIII (dorsal view). Scale bar: 0.1 mm." captionText-2="FIGURES 10 – 11. Aedeagus of Phlaeopterus czerskyi (Shavrin, 2001): 10 — parameral view, 11 — lateral view. Scale bar: 0.1 mm." captionText-3="FIGURE 12. Distribution of Phlaeopterus czerskyi (Shavrin, 2001)." httpUri-0="https://zenodo.org/record/243996/files/figure.png" httpUri-1="https://zenodo.org/record/243997/files/figure.png" httpUri-2="https://zenodo.org/record/243998/files/figure.png" httpUri-3="https://zenodo.org/record/243999/files/figure.png" pageId="2" pageNumber="123">Figs. 1–12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3524FD02E35680C6F9AFD84FDE9FDCF" box="[151,516,580,602]" pageId="2" pageNumber="123" type="reference_group">
<paragraph id="8BF71C5B2E35680C6F9AFD84FDE9FDCF" blockId="2.[151,516,580,602]" box="[151,516,580,602]" pageId="2" pageNumber="123">
<treatmentCitationGroup id="AB583B752E35680C6F9AFD84FDE9FDCF" box="[151,516,580,602]" pageId="2" pageNumber="123">
<taxonomicName id="4C4867D82E35680C6F9AFD84FDE9FDCF" ID-CoL="72358" authority="Shavrin, 2001: 190" authorityName="Shavrin" authorityPageNumber="190" authorityYear="2001" box="[151,516,580,602]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="123" phylum="Arthropoda" rank="species" species="czerskyi">
<emphasis id="B93CC0492E35680C6F9AFD84FED1FDCF" box="[151,316,580,602]" italics="true" pageId="2" pageNumber="123">Lesteva czerskyi</emphasis>
<treatmentCitation id="0AE93A4A2E35680C6E4FFD85FDE9FDCF" author="Shavrin" box="[322,516,580,602]" page="190" pageId="2" pageNumber="123" year="2001">
<bibRefCitation id="EFD961AA2E35680C6E4FFD85FE39FDCF" author="Shavrin" box="[322,468,580,602]" pageId="2" pageNumber="123" refString="Shavrin, A. V. (2001) New and little-known species of Omaliinae from the Baikal-Transbaikal area (Coleoptera: Staphylinidae). Zoosystematica Rossica, 9, 189 - 193." type="journal article" year="2001">Shavrin, 2001</bibRefCitation>
: 190
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3524FD02E35680C6F9AFD47FBF2FC76" pageId="2" pageNumber="123" type="materials_examined">
<paragraph id="8BF71C5B2E35680C6F9AFD47FC24FCE2" blockId="2.[151,1437,646,995]" pageId="2" pageNumber="123">
<emphasis id="B93CC0492E35680C6F9AFD47FE28FD0A" bold="true" box="[151,453,646,671]" pageId="2" pageNumber="123">
<typeStatus id="54F3A2F92E35680C6F9AFD47FF3FFD0A" box="[151,210,646,671]" pageId="2" pageNumber="123">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="54F3A2F92E35680C6EC2FD47FDDAFD0A" box="[463,567,646,671]" pageId="2" pageNumber="123" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C4867D82E35680C6D69FD46FC25FD0A" authority="Shavrin, 2000" authorityName="Shavrin" authorityYear="2000" box="[612,968,646,671]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="123" phylum="Arthropoda" rank="species" species="czerskyi">
<emphasis id="B93CC0492E35680C6D69FD46FCF0FD0A" box="[612,797,646,671]" italics="true" pageId="2" pageNumber="123">Lesteva czerskyi</emphasis>
Shavrin, 2000
</taxonomicName>
♂: ‘
<date id="FFF63A9B2E35680C6CF2FD47FBA5FD0A" box="[1023,1096,646,671]" pageId="2" pageNumber="123">14.VI.</date>
[19]96 [printed on the side] | Хамар-Дабан [Khamar-Daban Mts.] | Комаринский [Komarinskiy] | xp. [Mt.] h=1900 | Øàâðèí À. [Shavrin A. leg.]’, ‘
<typeStatus id="54F3A2F92E35680C6FEBFD0FFEBEFD72" box="[230,339,718,743]" pageId="2" pageNumber="123" type="holotype">Holotype</typeStatus>
|
<taxonomicName id="4C4867D82E35680C6E67FD0EFE2DFD73" box="[362,448,719,742]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="123" phylum="Arthropoda" rank="genus">
<emphasis id="B93CC0492E35680C6E67FD0EFE2DFD73" box="[362,448,719,742]" italics="true" pageId="2" pageNumber="123">Lesteva</emphasis>
</taxonomicName>
|
<taxonomicName id="4C4867D82E35680C6EDBFD0FFDDDFD72" box="[470,560,718,743]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="123" phylum="Arthropoda" rank="species" species="czerskyi">
<emphasis id="B93CC0492E35680C6EDBFD0FFDDDFD72" box="[470,560,718,743]" italics="true" pageId="2" pageNumber="123">czerskyi</emphasis>
</taxonomicName>
| Shavrin A. 2000’ &lt;red printed label&gt; (
<collectionCode id="ED59849E2E35680C6CF3FD0EFBDFFD72" box="[1022,1074,719,743]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="2" pageNumber="123">ZIN</collectionCode>
). The
<typeStatus id="54F3A2F92E35680C6B77FD0FFB31FD72" box="[1146,1244,718,743]" pageId="2" pageNumber="123" type="holotype">holotype</typeStatus>
was collected in upper flow of Slyudyanka River (Irkutsk Area, Slyudyanskiy District, Khamar-Daban Mts., Komarinskiy Mt.;
<geoCoordinate id="EE7C7A9C2E35680C6F9AFCD7FF16FCBB" box="[151,251,790,815]" direction="north" orientation="latitude" pageId="2" pageNumber="123" precision="925" value="51.85">51º51' N</geoCoordinate>
,
<geoCoordinate id="EE7C7A9C2E35680C6E05FCD7FE94FCBA" box="[264,377,790,815]" direction="east" orientation="longitude" pageId="2" pageNumber="123" precision="925" value="104.01667">103º61' E</geoCoordinate>
).
<typeStatus id="54F3A2F92E35680C6E80FCD6FE02FCBA" box="[397,495,791,815]" pageId="2" pageNumber="123" type="paratype">Paratype</typeStatus>
of
<taxonomicName id="4C4867D82E35680C6D1AFCD6FC98FCBA" authority="Shavrin, 2000" authorityName="Shavrin" authorityYear="2000" box="[535,885,790,815]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="123" phylum="Arthropoda" rank="species" species="czerskyi">
<emphasis id="B93CC0492E35680C6D1AFCD6FD23FCBA" box="[535,718,790,815]" italics="true" pageId="2" pageNumber="123">Lesteva czerskyi</emphasis>
Shavrin, 2000
</taxonomicName>
♀: ‘Хамар-Дабан [Khamar-Daban Mts.] | ср. т. Бабхи [middle flow of Babkha River] [
<geoCoordinate id="EE7C7A9C2E35680C6D60FCFBFD34FCC7" box="[621,729,826,851]" direction="north" orientation="latitude" pageId="2" pageNumber="123" precision="925" value="51.766666">51º46' N</geoCoordinate>
,
<geoCoordinate id="EE7C7A9C2E35680C6DE4FCFBFCB1FCC7" box="[745,860,826,851]" direction="east" orientation="longitude" pageId="2" pageNumber="123" precision="925" value="104.583336">103º95' E</geoCoordinate>
] |
<date id="FFF63A9B2E35680C6C8FFCFAFBFCFCC6" box="[898,1041,827,851]" pageId="2" pageNumber="123" value="1999-05-08" valueMax="1999-05-14" valueMin="1999-05-08">8–14.V.1999</date>
| Шаврин А. [Shavrin A. leg.]’, ‘
<typeStatus id="54F3A2F92E35680C6F96FC9EFEEFFCE2" box="[155,258,863,887]" pageId="2" pageNumber="123" type="paratype">Paratype</typeStatus>
|
<taxonomicName id="4C4867D82E35680C6E1BFC9EFE86FCE3" box="[278,363,863,886]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="123" phylum="Arthropoda" rank="genus">
<emphasis id="B93CC0492E35680C6E1BFC9EFE86FCE3" box="[278,363,863,886]" italics="true" pageId="2" pageNumber="123">Lesteva</emphasis>
</taxonomicName>
|
<taxonomicName id="4C4867D82E35680C6E72FC9FFE34FCE2" box="[383,473,862,887]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="123" phylum="Arthropoda" rank="species" species="czerskyi">
<emphasis id="B93CC0492E35680C6E72FC9FFE34FCE2" box="[383,473,862,887]" italics="true" pageId="2" pageNumber="123">czerskyi</emphasis>
</taxonomicName>
| Shavrin A. 2000’ &lt;red printed label&gt; (cS).
</paragraph>
<paragraph id="8BF71C5B2E35680C6FCAFC43FBF2FC76" blockId="2.[151,1437,646,995]" pageId="2" pageNumber="123">
<emphasis id="B93CC0492E35680C6FCAFC43FE57FC0E" bold="true" box="[199,442,898,923]" pageId="2" pageNumber="123">Additional material.</emphasis>
1 ♀: E Siberia, Khamar-Daban Mts., upper of Babkha River, Babkha mountain. [
<geoCoordinate id="EE7C7A9C2E35680C6A59FC43FF43FC2B" direction="north" orientation="latitude" pageId="2" pageNumber="123" precision="925" value="51.7">51º42' N</geoCoordinate>
, 103º.89' E] h=
<quantity id="4CB0B1BE2E35680C6E54FC67FE5CFC2B" box="[345,433,934,959]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="123" unit="m" value="1600.0">1600 m</quantity>
.
<date id="FFF63A9B2E35680C6EB1FC67FDD0FC2A" box="[444,573,934,959]" pageId="2" pageNumber="123" value="2006-06-27">27.06.2006</date>
. Shavrin A. leg. (cS);
<specimenCount id="9D4ED7D22E35680C6C3BFC66FC8FFC2B" box="[822,866,935,959]" pageId="2" pageNumber="123" type="generic">1 ♂</specimenCount>
: East Siberia, Khamar-Daban Mts., middle flow of Babkha River. [
<geoCoordinate id="EE7C7A9C2E35680C6E48FC0BFE46FC77" box="[325,427,970,995]" direction="north" orientation="latitude" pageId="2" pageNumber="123" precision="925" value="51.783333">51º47' N</geoCoordinate>
,
<geoCoordinate id="EE7C7A9C2E35680C6EBBFC0BFDC8FC77" box="[438,549,970,995]" direction="east" orientation="longitude" pageId="2" pageNumber="123" precision="925" value="104.6">103º96' E</geoCoordinate>
; h=
<quantity id="4CB0B1BE2E35680C6D43FC0BFD76FC77" box="[590,667,970,995]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="123" unit="m" value="500.0">500 m</quantity>
].
<date id="FFF63A9B2E35680C6DA0FC0BFCC3FC76" box="[685,814,970,995]" pageId="2" pageNumber="123" value="2006-06-27">27.06.2006</date>
. Shavrin A. leg. (cS).
</paragraph>
</subSubSection>
<caption id="DF374CD32E35680C6F9AF886FCEFF8E8" httpUri="https://zenodo.org/record/243996/files/figure.png" pageId="2" pageNumber="123" targetBox="[165,1426,1057,1836]" targetPageId="2">
<paragraph id="8BF71C5B2E35680C6F9AF886FCEFF8E8" blockId="2.[151,1435,1863,1917]" pageId="2" pageNumber="123">
<emphasis id="B93CC0492E35680C6F9AF886FED7F8C8" bold="true" box="[151,314,1863,1886]" pageId="2" pageNumber="123">FIGURES 1–2.</emphasis>
Habitus, female (Fig. 1) and habitat (Fig. 2) in upper of Babkha River (Eastern Siberia, Khamar-Daban Mts.) of
<taxonomicName id="4C4867D82E35680C6FBFF8A6FDD4F8E8" authority="Shavrin, 2001" authorityName="Shavrin" authorityYear="2001" box="[178,569,1895,1917]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="123" phylum="Arthropoda" rank="species" species="czerskyi">
<emphasis id="B93CC0492E35680C6FBFF8A6FE7CF8E8" box="[178,401,1895,1917]" italics="true" pageId="2" pageNumber="123">Phlaeopterus czerskyi</emphasis>
(Shavrin, 2001)
</taxonomicName>
. Scale bar: 1.0 mm.
</paragraph>
</caption>
<subSubSection id="C3524FD02E3568086FCAF865FD9EFEF0" lastPageId="6" lastPageNumber="127" pageId="2" pageNumber="123" type="description">
<paragraph id="8BF71C5B2E35680D6FCAF865FC93FE1D" blockId="2.[151,1436,1956,2017]" lastBlockId="3.[151,1437,151,789]" lastPageId="3" lastPageNumber="124" pageId="2" pageNumber="123">
<emphasis id="B93CC0492E35680C6FCAF865FE99F828" bold="true" box="[199,372,1956,1981]" pageId="2" pageNumber="123">Redescription.</emphasis>
Measurements (n=4): Maximum width of head including eyes: 0.74–0.75; length of head (from base of labrum to neck constriction along the head midline): 0.50–0.60; length of antenna: 1.50; longitudinal length of eye: 0.20–0.21; length of temple (from posterior margin of eye to neck constriction): 0.50–0.60; length of pronotum: 0.90–1.01; maximal width of pronotum: 1.25–1.40; minimal width of pronotum at the base: 0.85; length/width of segment 3 of maxillary palpi: 0.075 / 0.050; length/width of segments 4 of maxillary palpi: 0.175 / 0.05; sutural length of elytra (length of elytra from apex of scutellum to posterior margin of sutural angle): 1.25– 1.40; maximal width of elytra: 1.55–1.65; width of segment IV of abdomen: 1.55–1.65; length of hind tarsus (without length of tarsal claws): 0.45; length of segment 5 of metatarsus (without length of tarsal claws): 0.125; length of aedeagus: 0.56–0.57; body length: 4.00 (
<typeStatus id="54F3A2F92E34680D6DC7FEAEFCDDFE1D" box="[714,816,367,392]" pageId="3" pageNumber="124" type="holotype">holotype</typeStatus>
)–4.30.
</paragraph>
<paragraph id="8BF71C5B2E34680D6FCAFE55FCC6FC81" blockId="3.[151,1437,151,789]" pageId="3" pageNumber="124">
Forebody reddish brown (sometimes head slightly darker, brown with lateral margins of pronotum in posterior third and epicranial borders of head lighter, yellowish brown); abdomen reddish brown to black; clypeus and legs reddish brown to yellowish brown; antennomeres IV–IX and ocelli brown; antennal segments I–III and maxillary palpi yellow (sometimes segment IV of maxillary palpi darker, yellowish brown). Punctation of head dense, punctures deep and coarse, separated by one puncture diameter; punctation on temples and near neck smaller and denser, separated by one or two puncture diameters; pronotal punctures coarser than on head, on disc separated by one or two puncture diameters; scutellum with fine dense punctures; punctation of elytra irregular, punctures large and densely distributed, separated by one or two puncture diameters; punctation of abdominal tergites regular, fine and dense; surface of abdominal tergite IV slightly rugose between fine indistinct punctures.. Body without microsculpture, glossy. Pubescence of forebody decumbent, long, yellow; pubescence of tibiae dense and long; pubescence of abdomen short, yellow. Habitus as in
<figureCitation id="137300DE2E34680D6DEFFD3DFCC8FC81" box="[738,805,764,789]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,1863,1886]" captionTargetBox="[165,1426,1057,1836]" captionTargetId="figure@2.[151,1436,1050,1840]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Habitus, female (Fig. 1) and habitat (Fig. 2) in upper of Babkha River (Eastern Siberia, Khamar-Daban Mts.) of Phlaeopterus czerskyi (Shavrin, 2001). Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/243996/files/figure.png" pageId="3" pageNumber="124">Fig. 1</figureCitation>
.
</paragraph>
<caption id="DF374CD32E34680D6F9AF859FC9BF879" httpUri="https://zenodo.org/record/243997/files/figure.png" pageId="3" pageNumber="124" targetBox="[217,1376,831,1929]" targetPageId="3">
<paragraph id="8BF71C5B2E34680D6F9AF859FC9BF879" blockId="3.[151,1436,1944,2028]" pageId="3" pageNumber="124">
<emphasis id="B93CC0492E34680D6F9AF859FED0F838" bold="true" box="[151,317,1944,1967]" pageId="3" pageNumber="124">FIGURES 3–9.</emphasis>
<taxonomicName id="4C4867D82E34680D6E48F859FD38F83B" authority="Shavrin, 2001" authorityName="Shavrin" authorityYear="2001" box="[325,725,1944,1966]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="124" phylum="Arthropoda" rank="species" species="czerskyi">
<emphasis id="B93CC0492E34680D6E48F859FDCAF83B" box="[325,551,1944,1966]" italics="true" pageId="3" pageNumber="124">Phlaeopterus czerskyi</emphasis>
(Shavrin, 2001)
</taxonomicName>
: 3—labrum (dorsal view), 4—left and right mandibles(dorsal view), 5—labium (dorsal view), 6—mentum (dorsal view), 7—right maxilla (dorsal view), 8—male abdominal sternite VIII(dorsal view), 9—male abdominal tergite VIII (dorsal view). Scale bar: 0.1 mm.
</paragraph>
</caption>
<caption id="DF374CD32E33680A6F9AFB35FF2BFABF" httpUri="https://zenodo.org/record/243998/files/figure.png" pageId="4" pageNumber="125" targetBox="[215,1371,184,1238]" targetPageId="4">
<paragraph id="8BF71C5B2E33680A6F9AFB35FF2BFABF" blockId="4.[151,1436,1268,1322]" pageId="4" pageNumber="125">
<emphasis id="B93CC0492E33680A6F9AFB35FEB9FA9F" bold="true" box="[151,340,1268,1291]" pageId="4" pageNumber="125">FIGURES 10–11.</emphasis>
Aedeagus of
<taxonomicName id="4C4867D82E33680A6EE9FB34FC9DFA9E" authority="(Shavrin, 2001)" authorityName="Shavrin" authorityYear="2001" baseAuthorityName="Shavrin" baseAuthorityYear="2001" box="[484,880,1269,1291]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="czerskyi">
<emphasis id="B93CC0492E33680A6EE9FB34FD29FA9E" box="[484,708,1269,1291]" italics="true" pageId="4" pageNumber="125">Phlaeopterus czerskyi</emphasis>
(Shavrin, 2001)
</taxonomicName>
: 10—parameral view, 11—lateral view. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BF71C5B2E33680A6FCAFA90FACFF9F3" blockId="4.[151,1436,1361,2034]" pageId="4" pageNumber="125">
Head 1.6–1.8 times as wide as long, with prominent small eyes and short temples; eyes four times as long as temples. Vertex convex; surface in front of ocelli without longitudinal furrows, but with deeply impressed anteocellar foveae (two small impressions between anterior margins of eyes). Ocelli large, as large as diameter of two nearest punctures together; distance between ocelli subequal to distance between ocellus and posterior margin of eye. Neck (nuchal) constriction very smooth, vaguely marked. Labrum as in
<figureCitation id="137300DE2E33680A6B23FA20FB99FA6F" box="[1070,1140,1505,1530]" captionStart="FIGURES 3 – 9" captionStartId="3.[151,264,1944,1967]" captionTargetBox="[217,1376,831,1929]" captionTargetId="figure@3.[205,1382,823,1937]" captionTargetPageId="3" captionText="FIGURES 3 – 9. Phlaeopterus czerskyi (Shavrin, 2001): 3 — labrum (dorsal view), 4 — left and right mandibles (dorsal view), 5 — labium (dorsal view), 6 — mentum (dorsal view), 7 — right maxilla (dorsal view), 8 — male abdominal sternite VIII (dorsal view), 9 — male abdominal tergite VIII (dorsal view). Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/243997/files/figure.png" pageId="4" pageNumber="125">Fig. 3</figureCitation>
. Mandibulae as in
<figureCitation id="137300DE2E33680A6A5FFA20FA75FA6F" box="[1362,1432,1505,1530]" captionStart="FIGURES 3 – 9" captionStartId="3.[151,264,1944,1967]" captionTargetBox="[217,1376,831,1929]" captionTargetId="figure@3.[205,1382,823,1937]" captionTargetPageId="3" captionText="FIGURES 3 – 9. Phlaeopterus czerskyi (Shavrin, 2001): 3 — labrum (dorsal view), 4 — left and right mandibles (dorsal view), 5 — labium (dorsal view), 6 — mentum (dorsal view), 7 — right maxilla (dorsal view), 8 — male abdominal sternite VIII (dorsal view), 9 — male abdominal tergite VIII (dorsal view). Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/243997/files/figure.png" pageId="4" pageNumber="125">Fig. 4</figureCitation>
. Labium as in
<figureCitation id="137300DE2E33680A6E30F9C4FE6BF98B" box="[317,390,1541,1566]" captionStart="FIGURES 3 – 9" captionStartId="3.[151,264,1944,1967]" captionTargetBox="[217,1376,831,1929]" captionTargetId="figure@3.[205,1382,823,1937]" captionTargetPageId="3" captionText="FIGURES 3 – 9. Phlaeopterus czerskyi (Shavrin, 2001): 3 — labrum (dorsal view), 4 — left and right mandibles (dorsal view), 5 — labium (dorsal view), 6 — mentum (dorsal view), 7 — right maxilla (dorsal view), 8 — male abdominal sternite VIII (dorsal view), 9 — male abdominal tergite VIII (dorsal view). Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/243997/files/figure.png" pageId="4" pageNumber="125">Fig. 5</figureCitation>
. Mentum as in
<figureCitation id="137300DE2E33680A6D4EF9C4FD61F98B" box="[579,652,1541,1566]" captionStart="FIGURES 3 – 9" captionStartId="3.[151,264,1944,1967]" captionTargetBox="[217,1376,831,1929]" captionTargetId="figure@3.[205,1382,823,1937]" captionTargetPageId="3" captionText="FIGURES 3 – 9. Phlaeopterus czerskyi (Shavrin, 2001): 3 — labrum (dorsal view), 4 — left and right mandibles (dorsal view), 5 — labium (dorsal view), 6 — mentum (dorsal view), 7 — right maxilla (dorsal view), 8 — male abdominal sternite VIII (dorsal view), 9 — male abdominal tergite VIII (dorsal view). Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/243997/files/figure.png" pageId="4" pageNumber="125">Fig. 6</figureCitation>
. Maxillary palpus as in
<figureCitation id="137300DE2E33680A6CA3F9C4FC1BF988" box="[942,1014,1541,1566]" captionStart="FIGURES 3 – 9" captionStartId="3.[151,264,1944,1967]" captionTargetBox="[217,1376,831,1929]" captionTargetId="figure@3.[205,1382,823,1937]" captionTargetPageId="3" captionText="FIGURES 3 – 9. Phlaeopterus czerskyi (Shavrin, 2001): 3 — labrum (dorsal view), 4 — left and right mandibles (dorsal view), 5 — labium (dorsal view), 6 — mentum (dorsal view), 7 — right maxilla (dorsal view), 8 — male abdominal sternite VIII (dorsal view), 9 — male abdominal tergite VIII (dorsal view). Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/243997/files/figure.png" pageId="4" pageNumber="125">Fig. 7</figureCitation>
. Antennae relatively short, reaching shoulders of elytra; all antennomeres elongate, with lengths × widths: 1: 0.20 × 0.075; 2: 0.15 × 0.045; 3: 0.15 × 0.057; 4: 0.125 × 0.074; 5–6: 0.115 × 0.074; 7–8: 0.104 × 0.074; 9–10: 0.104 × 0.087; 11: 0.225 × 0.10.
</paragraph>
<paragraph id="8BF71C5B2E33680A6FCAF9B3FF3DF963" blockId="4.[151,1436,1361,2034]" pageId="4" pageNumber="125">Pronotum transverse (width/length ratio: 1.38), very convex, 1.6–1.8 times wider than head, widest near middle, strongly narrowed from widest point to base and apex, with deep long lateral impressions, hind angles about 90º; both lateral margins crenulate, strongly behind widest point, transitioning gradually to indistinct near base.</paragraph>
<paragraph id="8BF71C5B2E33680A6FCAF8C0FD10F88C" blockId="4.[151,1436,1361,2034]" box="[199,765,1793,1818]" pageId="4" pageNumber="125">Scutellum large, subtriangular, with rounded apex.</paragraph>
<paragraph id="8BF71C5B2E33680A6FCAF8E4FBBAF813" blockId="4.[151,1436,1361,2034]" pageId="4" pageNumber="125">Elytra moderately short and wide (width/length ratio: 1.2 on average), wider than maximal width of pronotum (maximal width of elytra/maximal width of pronotum ratio: 1.2 on average), posterior angles broadly rounded, hind margins of elytra obliquely truncate towards the suture. Wings reduced to short lobes.</paragraph>
<paragraph id="8BF71C5B2E33680A6FCAF850FED9F867" blockId="4.[151,1436,1361,2034]" pageId="4" pageNumber="125">Apical part of protibiae and inner and outer parts of meso- and metatibiae with several small long spines between setae. Metatarsomere 1 slightly shorter than apical metatarsomere; metatarsomere 5 equal to combined length of 3–4.</paragraph>
<paragraph id="8BF71C5B2E32680B6FCAFF56FDA6FF41" blockId="5.[151,1437,151,1221]" pageId="5" pageNumber="126">Abdomen very convex, slightly wider than elytra; abdominal tergites without wing-folding patches (tomentose spots) or palisade fringe on tergite VII.</paragraph>
<paragraph id="8BF71C5B2E32680B6FCAFF1EFF55FE1D" blockId="5.[151,1437,151,1221]" pageId="5" pageNumber="126">
Male. Pronotum slightly narrower; lateral tooth in middle of pronotum smoother, not well defined. First four protarsomeres weakly dilated, with longer and thicker ventral setae. Abdominal sternite VIII (
<figureCitation id="137300DE2E32680B6BC1FEC5FAFCFE89" box="[1228,1297,260,285]" captionStart="FIGURES 3 – 9" captionStartId="3.[151,264,1944,1967]" captionTargetBox="[217,1376,831,1929]" captionTargetId="figure@3.[205,1382,823,1937]" captionTargetPageId="3" captionText="FIGURES 3 – 9. Phlaeopterus czerskyi (Shavrin, 2001): 3 — labrum (dorsal view), 4 — left and right mandibles (dorsal view), 5 — labium (dorsal view), 6 — mentum (dorsal view), 7 — right maxilla (dorsal view), 8 — male abdominal sternite VIII (dorsal view), 9 — male abdominal tergite VIII (dorsal view). Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/243997/files/figure.png" pageId="5" pageNumber="126">Fig. 8</figureCitation>
) and tergite VIII (
<figureCitation id="137300DE2E32680B6FD7FEE6FECCFED5" box="[218,289,295,320]" captionStart="FIGURES 3 – 9" captionStartId="3.[151,264,1944,1967]" captionTargetBox="[217,1376,831,1929]" captionTargetId="figure@3.[205,1382,823,1937]" captionTargetPageId="3" captionText="FIGURES 3 – 9. Phlaeopterus czerskyi (Shavrin, 2001): 3 — labrum (dorsal view), 4 — left and right mandibles (dorsal view), 5 — labium (dorsal view), 6 — mentum (dorsal view), 7 — right maxilla (dorsal view), 8 — male abdominal sternite VIII (dorsal view), 9 — male abdominal tergite VIII (dorsal view). Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/243997/files/figure.png" pageId="5" pageNumber="126">Fig. 9</figureCitation>
) with weakly emarginate posterior margin. Aedeagus (
<figureCitation id="137300DE2E32680B6CA8FEE6FC17FED5" box="[933,1018,295,320]" captionStart="FIGURES 10 – 11" captionStartId="4.[151,264,1268,1291]" captionTargetBox="[215,1371,184,1238]" captionTargetId="figure@4.[204,1383,175,1248]" captionTargetPageId="4" captionText="FIGURES 10 – 11. Aedeagus of Phlaeopterus czerskyi (Shavrin, 2001): 10 — parameral view, 11 — lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/243998/files/figure.png" pageId="5" pageNumber="126">Fig. 10</figureCitation>
) wide and short, gradually tapering towards apex; parameres wide, exceeding apex of aedeagus, slightly curved medially. Aedeagus laterally as in
<figureCitation id="137300DE2E32680B6A7DFE8DFF59FE1D" captionStart="FIGURES 10 – 11" captionStartId="4.[151,264,1268,1291]" captionTargetBox="[215,1371,184,1238]" captionTargetId="figure@4.[204,1383,175,1248]" captionTargetPageId="4" captionText="FIGURES 10 – 11. Aedeagus of Phlaeopterus czerskyi (Shavrin, 2001): 10 — parameral view, 11 — lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/243998/files/figure.png" pageId="5" pageNumber="126">Fig. 11</figureCitation>
.
</paragraph>
<paragraph id="8BF71C5B2E32680B6FCAFE55FF1BFE60" blockId="5.[151,1437,151,1221]" pageId="5" pageNumber="126">Female. Pronotum distinctly transverse, with well defined lateral tooth at middle. Protarsomeres not dilated, without longer and thicker ventral setae. Abdominal sternite VIII and tergite VIII without emargination, straight apically.</paragraph>
<paragraph id="8BF71C5B2E32680B6FCAFE3EFE46FC81" blockId="5.[151,1437,151,1221]" pageId="5" pageNumber="126">
<emphasis id="B93CC0492E32680B6FCAFE3EFE42FD8D" bold="true" box="[199,431,511,536]" pageId="5" pageNumber="126">Comparative notes.</emphasis>
<taxonomicName id="4C4867D82E32680B6EBBFE3EFD46FD8D" box="[438,683,511,536]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="czerskyi">
<emphasis id="B93CC0492E32680B6EBBFE3EFD46FD8D" box="[438,683,511,536]" italics="true" pageId="5" pageNumber="126">Phlaeopterus czerskyi</emphasis>
</taxonomicName>
specimens were compared to specimens of all described (as well as some undescribed) North American
<taxonomicName id="4C4867D82E32680B6D21FDE5FD2CFDA8" box="[556,705,548,573]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="B93CC0492E32680B6D21FDE5FD2CFDA8" box="[556,705,548,573]" italics="true" pageId="5" pageNumber="126">Phlaeopterus</emphasis>
</taxonomicName>
species by the second author.
<taxonomicName id="4C4867D82E32680B6B1DFDE5FAEBFDA8" box="[1040,1286,548,573]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="czerskyi">
<emphasis id="B93CC0492E32680B6B1DFDE5FAEBFDA8" box="[1040,1286,548,573]" italics="true" pageId="5" pageNumber="126">Phlaeopterus czerskyi</emphasis>
</taxonomicName>
can easily be distinguished from all other known congeners by the crenulate sides of the pronotum, which are not present in any other species of the genus. It most closely resambles the Nearctic
<taxonomicName id="4C4867D82E32680B6C75FDACFB41FD10" authority="Hatch, 1957" authorityName="Hatch" authorityYear="1957" box="[888,1196,620,645]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="lagrandeuri">
<emphasis id="B93CC0492E32680B6C75FDACFBF7FD10" box="[888,1050,620,645]" italics="true" pageId="5" pageNumber="126">P. lagrandeuri</emphasis>
<bibRefCitation id="EFD961AA2E32680B6B2DFDADFB41FD10" author="Hatch" box="[1056,1196,620,645]" pageId="5" pageNumber="126" refString="Hatch, M. H. (1957) The beetles of the Pacific Northwest. Part II: Staphyliniformia. University of Washington Publications in Biology, 16 (2), i - ix + 1 - 384." type="journal article" year="1957">Hatch, 1957</bibRefCitation>
</taxonomicName>
, known from
<collectingCountry id="F35F5CCB2E32680B6A45FDADFA71FD10" box="[1352,1436,620,645]" name="Canada" pageId="5" pageNumber="126">Canada</collectingCountry>
and the
<collectingCountry id="F35F5CCB2E32680B6FF5FD51FEDDFD3D" box="[248,304,656,680]" name="United States of America" pageId="5" pageNumber="126">USA</collectingCountry>
, but can be distinguished from
<taxonomicName id="4C4867D82E32680B6DA1FD50FCBCFD3D" box="[684,849,655,680]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="lagrandeuri">
<emphasis id="B93CC0492E32680B6DA1FD50FCBCFD3D" box="[684,849,655,680]" italics="true" pageId="5" pageNumber="126">P. lagrandeuri</emphasis>
</taxonomicName>
by the head with anteocellar foveae more deeply impressed, pronotum and elytra with punctation coarser and deeper, metatrochanter lacking a large tooth on the apical margin, elytra shorter (see previous paragraph) and wider, and heavily sclerotized aedeagus with apex of median lobe more acute.
</paragraph>
<paragraph id="8BF71C5B2E32680B6FCAFCDEFB56FBE8" blockId="5.[151,1437,151,1221]" pageId="5" pageNumber="126">
Using the key to the Nearctic
<taxonomicName id="4C4867D82E32680B6D38FCDEFD22FCAD" box="[565,719,799,824]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="tribe" tribe="Anthophagini">Anthophagini</taxonomicName>
of
<bibRefCitation id="EFD961AA2E32680B6DF2FCE1FC14FCAD" box="[767,1017,799,824]" pageId="5" pageNumber="126" refString="Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2000) 22. Staphylinidae Latreille, 1802. In: Arnett, R. H. &amp; Thomas, M. C. (Ed.), American Beetles, Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia, pp. 272 - 418." type="book chapter">Newton et al. (2000)</bibRefCitation>
, the most recent, authoritative, and comprehensive keys to the North American staphylinid genera,
<taxonomicName id="4C4867D82E32680B6C9DFC84FBE6FCC8" box="[912,1035,836,861]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="czerskyi">
<emphasis id="B93CC0492E32680B6C9DFC84FBE6FCC8" box="[912,1035,836,861]" italics="true" pageId="5" pageNumber="126">P. czerskyi</emphasis>
</taxonomicName>
keys to
<taxonomicName id="4C4867D82E32680B6B76FC85FAFDFCC8" box="[1147,1296,836,861]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="B93CC0492E32680B6B76FC85FAFDFCC8" box="[1147,1296,836,861]" italics="true" pageId="5" pageNumber="126">Phlaeopterus</emphasis>
</taxonomicName>
rather than
<taxonomicName id="4C4867D82E32680B6F9AFCA8FF00FC15" box="[151,237,873,896]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="B93CC0492E32680B6F9AFCA8FF00FC15" box="[151,237,873,896]" italics="true" pageId="5" pageNumber="126">Lesteva</emphasis>
</taxonomicName>
by its pronotum shape (not cordate and with explanate edges and lateral impressions), and by its head with neck (nuchal) constriction very smooth, vaguely marked. However, the exceptionally short elytra of
<taxonomicName id="4C4867D82E32680B6A0FFC4CFA95FC30" box="[1282,1400,908,933]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="czerskyi">
<emphasis id="B93CC0492E32680B6A0FFC4CFA95FC30" box="[1282,1400,908,933]" italics="true" pageId="5" pageNumber="126">P. czerskyi</emphasis>
</taxonomicName>
do not match the diagnosis of
<taxonomicName id="4C4867D82E32680B6EC1FC6EFD8CFC5D" box="[460,609,943,968]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="B93CC0492E32680B6EC1FC6EFD8CFC5D" box="[460,609,943,968]" italics="true" pageId="5" pageNumber="126">Phlaeopterus</emphasis>
</taxonomicName>
given by
<bibRefCitation id="EFD961AA2E32680B6DD8FC71FC29FC5D" box="[725,964,943,968]" pageId="5" pageNumber="126" refString="Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2000) 22. Staphylinidae Latreille, 1802. In: Arnett, R. H. &amp; Thomas, M. C. (Ed.), American Beetles, Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia, pp. 272 - 418." type="book chapter">Newton et al. (2000)</bibRefCitation>
. The ratio of elytra to pronotum length is used in the key of
<bibRefCitation id="EFD961AA2E32680B6E6AFC14FDBCFC79" box="[359,593,980,1005]" pageId="5" pageNumber="126" refString="Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2000) 22. Staphylinidae Latreille, 1802. In: Arnett, R. H. &amp; Thomas, M. C. (Ed.), American Beetles, Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia, pp. 272 - 418." type="book chapter">Newton et al. (2000)</bibRefCitation>
to differentiate
<taxonomicName id="4C4867D82E32680B6C08FC15FC77FC78" box="[773,922,980,1005]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="B93CC0492E32680B6C08FC15FC77FC78" box="[773,922,980,1005]" italics="true" pageId="5" pageNumber="126">Phlaeopterus</emphasis>
</taxonomicName>
from the genus
<taxonomicName id="4C4867D82E32680B6B5EFC14FB47FC79" box="[1107,1194,981,1004]" class="Insecta" family="Staphylinidae" genus="Unamis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="B93CC0492E32680B6B5EFC14FB47FC79" box="[1107,1194,981,1004]" italics="true" pageId="5" pageNumber="126">Unamis</emphasis>
</taxonomicName>
: “elytral length more than 2, usually more than 2.2, times that of pronotum” for
<taxonomicName id="4C4867D82E32680B6C31FC36FC3CFB85" box="[828,977,1015,1040]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="B93CC0492E32680B6C31FC36FC3CFB85" box="[828,977,1015,1040]" italics="true" pageId="5" pageNumber="126">Phlaeopterus</emphasis>
</taxonomicName>
and “elytral length seldom more than 2 times that of pronotum” for
<taxonomicName id="4C4867D82E32680B6EEFFBDCFDD4FBA1" box="[482,569,1053,1076]" class="Insecta" family="Staphylinidae" genus="Unamis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="B93CC0492E32680B6EEFFBDCFDD4FBA1" box="[482,569,1053,1076]" italics="true" pageId="5" pageNumber="126">Unamis</emphasis>
</taxonomicName>
. The elytra of
<taxonomicName id="4C4867D82E32680B6DE6FBDCFC89FBA0" box="[747,868,1052,1077]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="czerskyi">
<emphasis id="B93CC0492E32680B6DE6FBDCFC89FBA0" box="[747,868,1052,1077]" italics="true" pageId="5" pageNumber="126">P. czerskyi</emphasis>
</taxonomicName>
are less than twice as long as the pronotum, the shortest of any
<taxonomicName id="4C4867D82E32680B6E4EFBFEFE35FBCD" box="[323,472,1087,1112]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="B93CC0492E32680B6E4EFBFEFE35FBCD" box="[323,472,1087,1112]" italics="true" pageId="5" pageNumber="126">Phlaeopterus</emphasis>
</taxonomicName>
species. Nonetheless,
<taxonomicName id="4C4867D82E32680B6DD8FB80FCA6FBCD" box="[725,843,1087,1112]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="czerskyi">
<emphasis id="B93CC0492E32680B6DD8FB80FCA6FBCD" box="[725,843,1087,1112]" italics="true" pageId="5" pageNumber="126">P. czerskyi</emphasis>
</taxonomicName>
matches the diagnosis of
<taxonomicName id="4C4867D82E32680B6B60FBFEFAEFFBCD" box="[1133,1282,1087,1112]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="B93CC0492E32680B6B60FBFEFAEFFBCD" box="[1133,1282,1087,1112]" italics="true" pageId="5" pageNumber="126">Phlaeopterus</emphasis>
</taxonomicName>
of
<bibRefCitation id="EFD961AA2E32680B6A25FB81FEE9FBE9" pageId="5" pageNumber="126" refString="Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2000) 22. Staphylinidae Latreille, 1802. In: Arnett, R. H. &amp; Thomas, M. C. (Ed.), American Beetles, Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia, pp. 272 - 418." type="book chapter">Newton et al. (2000)</bibRefCitation>
in all other characters, as well as the revised diagnosis of
<taxonomicName id="4C4867D82E32680B6C83FBA5FBCEFBE8" box="[910,1059,1124,1149]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="B93CC0492E32680B6C83FBA5FBCEFBE8" box="[910,1059,1124,1149]" italics="true" pageId="5" pageNumber="126">Phlaeopterus</emphasis>
</taxonomicName>
given above.
</paragraph>
<paragraph id="8BF71C5B2E32680B6FCAFB46FC03FB51" blockId="5.[151,1437,151,1221]" pageId="5" pageNumber="126">
<emphasis id="B93CC0492E32680B6FCAFB46FEA7FB35" bold="true" box="[199,330,1159,1184]" pageId="5" pageNumber="126">Bionomics.</emphasis>
This species was collected in alpine and forest zones of the Khamar-Daban Mts. (
<figureCitation id="137300DE2E32680B6BE0FB46FAADFB35" box="[1261,1344,1159,1184]" captionStart="FIGURE 12" captionStartId="5.[151,250,1914,1936]" captionTargetBox="[209,1378,1273,1892]" captionTargetId="figure@5.[209,1378,1273,1893]" captionTargetPageId="5" captionText="FIGURE 12. Distribution of Phlaeopterus czerskyi (Shavrin, 2001)." httpUri="https://zenodo.org/record/243999/files/figure.png" pageId="5" pageNumber="126">Fig. 12</figureCitation>
), in wet moss on the banks of small streams (
<figureCitation id="137300DE2E32680B6D3EFB6DFD9BFB51" box="[563,630,1196,1221]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,1863,1886]" captionTargetBox="[165,1426,1057,1836]" captionTargetId="figure@2.[151,1436,1050,1840]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Habitus, female (Fig. 1) and habitat (Fig. 2) in upper of Babkha River (Eastern Siberia, Khamar-Daban Mts.) of Phlaeopterus czerskyi (Shavrin, 2001). Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/243996/files/figure.png" pageId="5" pageNumber="126">Fig. 2</figureCitation>
) at altitudes from
<quantity id="4CB0B1BE2E32680B6C49FB6DFC07FB51" box="[836,1002,1196,1220]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.9" metricValueMin="0.9" pageId="5" pageNumber="126" unit="m" value="1400.0" valueMax="1900.0" valueMin="900.0">900 to 1900 m</quantity>
.
</paragraph>
<caption id="DF374CD32E32680B6F9AF8BBFCA0F805" box="[151,845,1914,1936]" httpUri="https://zenodo.org/record/243999/files/figure.png" pageId="5" pageNumber="126" targetBox="[209,1378,1273,1892]" targetPageId="5">
<paragraph id="8BF71C5B2E32680B6F9AF8BBFCA0F805" blockId="5.[151,845,1914,1936]" box="[151,845,1914,1936]" pageId="5" pageNumber="126">
<emphasis id="B93CC0492E32680B6F9AF8BBFEF2F81A" bold="true" box="[151,287,1914,1936]" pageId="5" pageNumber="126">FIGURE 12.</emphasis>
Distribution of
<taxonomicName id="4C4867D82E32680B6ECFF8BBFCA4F805" authority="(Shavrin, 2001)" authorityName="Shavrin" authorityYear="2001" baseAuthorityName="Shavrin" baseAuthorityYear="2001" box="[450,841,1914,1936]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="czerskyi">
<emphasis id="B93CC0492E32680B6ECFF8BBFD4DF805" box="[450,672,1914,1936]" italics="true" pageId="5" pageNumber="126">Phlaeopterus czerskyi</emphasis>
(Shavrin, 2001)
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BF71C5B2E3168086FCAFF56FD9EFEF0" blockId="6.[151,1437,151,357]" pageId="6" pageNumber="127">
Nearctic
<taxonomicName id="4C4867D82E3168086E23FF56FE2EFF25" box="[302,451,151,176]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="B93CC0492E3168086E23FF56FE2EFF25" box="[302,451,151,176]" italics="true" pageId="6" pageNumber="127">Phlaeopterus</emphasis>
</taxonomicName>
species have been collected from sea level to
<quantity id="4CB0B1BE2E3168086CD4FF56FBDDFF3A" box="[985,1072,151,176]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="127" unit="m" value="3500.0">3500 m</quantity>
elevation, and are found on the surface of alpine snowfields, as well as under rocks and in moss at the edges of permanent or long lasting alpine snowfields and cold, cascading streams and waterfalls. Adults of Nearctic
<taxonomicName id="4C4867D82E3168086B1FFF1EFB4AFF6D" box="[1042,1191,223,248]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="B93CC0492E3168086B1FFF1EFB4AFF6D" box="[1042,1191,223,248]" italics="true" pageId="6" pageNumber="127">Phlaeopterus</emphasis>
</taxonomicName>
have been observed foraging on arthropod fallout (mostly flying insects, windblown from lower altitudes and often lethargic or frozen) on the surface of alpine snowfields (at night at lower latitudes but during the day in Alaska), and this feeding strategy is likely widely used in the genus.
</paragraph>
</subSubSection>
</treatment>
</document>
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