treatments-xml/data/03/DC/6C/03DC6C76CC1E4C14F7DFFF78FDC8F820.xml

<document id="1DFC40F730AB8A0FF8A3784DC9024E09" ID-DOI="10.11646/zootaxa.5267.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7840783" ID-ZooBank="9CD59054-8D7D-413F-B9FD-29EAFE7E511D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1681816955386" checkinUser="plazi" docAuthor="Lehmann, Ingo, Zahiri, Reza &amp; Husemann, Martin" docDate="2023" docId="03DC6C76CC1E4C14F7DFFF78FDC8F820" docLanguage="en" docName="zootaxa.5267.1.1.pdf" docOrigin="Zootaxa 5267 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5267.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Lukeniana bergsteni Lehmann, Zahiri &amp; Husemann 2023, sp. nov." docType="treatment" docUuid="DE2DA909-706C-4591-BE9C-11C98A31FBF7" docUuidSource="ZooBank" docVersion="3" lastPageNumber="49" masterDocId="FFE5140ECC314C24F748FFEFFF9EFFB5" masterDocTitle="Revision of the Metarbelodes Strand, 1909 genus-group (Lepidoptera: Cossoidea: Metarbelidae) with descriptions of two new genera and 33 new species from high elevations of eastern and southern Africa" masterLastPageNumber="106" masterPageNumber="1" pageNumber="48" updateTime="1699487052491" updateUser="plazi">
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<mods:title id="D8C665B14F7B45C593201B2F852BE184">Revision of the Metarbelodes Strand, 1909 genus-group (Lepidoptera: Cossoidea: Metarbelidae) with descriptions of two new genera and 33 new species from high elevations of eastern and southern Africa</mods:title>
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<mods:namePart id="545CBF483001169D7B2AC4C9E4A8B4D8">Lehmann, Ingo</mods:namePart>
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<mods:namePart id="FCB00ADCFDFE9D618AB05482DFA1D580">Zahiri, Reza</mods:namePart>
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<mods:namePart id="34529A47C501A034A8ACC502A722E2E7">Husemann, Martin</mods:namePart>
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<taxonomicName id="4C75A6E3CC1E4C0BF7DFFF78FC80FF04" ID-CoL="B8R7N" authority="Lehmann, Zahiri &amp; Husemann" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[151,798,151,178]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="bergsteni" status="sp. nov.">
<emphasis id="B9010172CC1E4C0BF7DFFF78FE0CFF07" bold="true" box="[151,402,151,178]" italics="true" pageId="47" pageNumber="48">Lukeniana bergsteni</emphasis>
Lehmann, Zahiri &amp; Husemann
</taxonomicName>
<taxonomicNameLabel id="A232BC09CC1E4C0BF46EFF78FC1DFF04" box="[806,899,151,177]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="C36F8EEBCC1E4C0BF7DFFF54FB7CFEF4" pageId="47" pageNumber="48" type="description">
<paragraph id="8BCADD60CC1E4C0BF7DFFF54FC53FF4F" blockId="47.[151,973,151,250]" pageId="47" pageNumber="48">
<figureCitation id="134EC1E5CC1E4C0BF7DFFF54FEA1FF60" box="[151,319,187,213]" captionStart="FIGURE 8" captionStartId="51.[151,250,1779,1803]" captionTargetBox="[151,1436,452,1755]" captionTargetId="figure-137@51.[151,1436,452,1755]" captionTargetPageId="51" captionText="FIGURE 8. a. Lukeniana jankiellandi, holotype, male, Tanzania, Katavi Region, Luega village; b. L. bergsteni, paratype, male, Tanzania, Kilimanjaro Region, Mount Kilimanjaro, Kibonoto; c. L. bergsteni, paratype, male, Tanzania, Arusha Region, Usa River; d. L. bergsteni, paratype, female, Tanzania, Kilimanjaro Region, Mount Kilimanjaro, Kibonoto; e. L. carolae, holotype, male, Malawi, Central Region, Mount Ntchisi, Ntchisi Forest Reserve; f. L. stevecollinsi, holotype, male, Malawi, Northern Region, Nyika Plateau, Nyika National Park, near Thazima Gate." figureDoi="http://doi.org/10.5281/zenodo.7840805" httpUri="https://zenodo.org/record/7840805/files/figure.png" pageId="47" pageNumber="48">Figs 8b, 8c, 8d</figureCitation>
, 
<figureCitation id="134EC1E5CC1E4C0BF603FF54FEE8FF60" box="[331,374,187,213]" captionStart="FIGURE 14" captionStartId="65.[151,250,1754,1778]" captionTargetBox="[159,1425,534,1722]" captionTargetId="figure-191@65.[152,1436,525,1730]" captionTargetPageId="65" captionText="FIGURE 14. a. Lukeniana bergsteni, paratype, male, Tanzania, Kilimanjaro Region, Mount Kilimanjaro, Kibonoto; b. L. carolae, holotype, male, Malawi, Central Region, Mount Ntchisi, Ntchisi Forest Reserve; c. L. stevecollinsi, holotype, male, Malawi, Northern Region, Nyika Plateau, Nyika National Park, near Thazima Gate; d. L. butleri, paratype, male, Zimbabwe, Harare Province, Harare, “Fawlty Towers”; e. L. kollhorsti, holotype, male, Zimbabwe, Manicaland Province, Busi Farm; f. Zambezia diredaouaensis, holotype, male, Ethiopia, Harar Plateau, Dire Daua, today: chartered city of Dire Dawa." figureDoi="http://doi.org/10.5281/zenodo.7840822" httpUri="https://zenodo.org/record/7840822/files/figure.png" pageId="47" pageNumber="48">14a</figureCitation>
, 
<figureCitation id="134EC1E5CC1E4C0BF6C9FF54FE32FF63" box="[385,428,187,214]" captionStart="FIGURE 15" captionStartId="66.[151,250,1656,1680]" captionTargetBox="[172,1419,191,1622]" captionTargetId="figure-28@66.[151,1436,181,1632]" captionTargetPageId="66" captionText="FIGURE 15. Examples of male genitalia in lateral view: a. Zambezia madambae, paratype, with extremely small thorn-like process and small weakly-sclerotized projection (red arrows); b. Lukeniana timdavenporti, paratype, with strongly bent ventral edge of valva and well developed thorn-like process (red arrows); c. L. bergsteni, paratype, without strongly bent ventral edge of valva and two thorn-like processes (red arrows); d. L. butleri, paratype, with many long setae on valva that appear to be black in colour and one thorn-like process (red arrows)." figureDoi="http://doi.org/10.5281/zenodo.7840824" httpUri="https://zenodo.org/record/7840824/files/figure.png" pageId="47" pageNumber="48">15c</figureCitation>
, 
<figureCitation id="134EC1E5CC1E4C0BF6F0FF54FE7EFF60" box="[440,480,187,213]" captionStart="FIGURE 18" captionStartId="69.[151,250,1796,1820]" captionTargetBox="[151,1436,731,1773]" captionTargetId="figure-176@69.[151,1436,730,1773]" captionTargetPageId="69" captionText="FIGURE 18. a. shows two fronto-clypeal projections (red arrows) of Lukeniana bergsteni, female, paratype. The projections are close together, that means the distance between the projections is ca. 1.0–1.5× of the width of the pit behind the labial palpus, and they are more pronounced in females than in males of Lukeniana and Zambezia. The two projections are more widely separated in species of Mountelgonia Lehmann, 2013, that means the distance between the projections is ca. 2.5–3.5× of the width of the pit behind the labial palpus. b. shows an enlargement of one fronto-clypeal projection. c. shows four branches of the female antenna. The pictures were taken with a Scanning Electron Microscope at ZFMK by Karin Ulmen in March 2012." figureDoi="http://doi.org/10.5281/zenodo.7840835" httpUri="https://zenodo.org/record/7840835/files/figure.png" pageId="47" pageNumber="48">18a</figureCitation>
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<materialsCitation id="3B1DD73DCC1E4C0BF7DFFEC7FB40FEF4" box="[151,1246,295,322]" collectionCode="NRM" country="Tanzania" location="Swedish Natural History Museum" municipality="Stockholm" pageId="47" pageNumber="48" specimenCount="1" typeStatus="holotype">
<typeStatus id="54CE63C2CC1E4C0BF7DFFEC7FF51FEF7" box="[151,207,296,322]" pageId="47" pageNumber="48">Type</typeStatus>
locality and repository: 
<collectingCountry id="F3629DF0CC1E4C0BF6A8FEC8FDD6FEF4" box="[480,584,295,321]" name="Tanzania" pageId="47" pageNumber="48">Tanzania</collectingCountry>
, the 
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, 
<collectingMunicipality id="6BAE471ACC1E4C0BF340FEC8FB1FFEF4" box="[1032,1153,295,321]" pageId="47" pageNumber="48">Stockholm</collectingMunicipality>
(
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)
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.
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<paragraph id="8BCADD60CC1E4C0BF7DFFE80FD79F9BC" blockId="47.[151,1437,367,2014]" pageId="47" pageNumber="48">
<emphasis id="B9010172CC1E4C0BF7DFFE80FE1CFE3C" bold="true" box="[151,386,367,393]" pageId="47" pageNumber="48">Material examined.</emphasis>
<typeStatus id="54CE63C2CC1E4C0BF6C3FE80FE6DFE3C" box="[395,499,367,393]" pageId="47" pageNumber="48" type="holotype">Holotype</typeStatus>
male, 
<collectingCountry id="F3629DF0CC1E4C0BF509FE80FD37FE3C" box="[577,681,367,393]" name="Tanzania" pageId="47" pageNumber="48">Tanzania</collectingCountry>
, 
<collectingRegion id="49B11382CC1E4C0BF5FDFE80FC07FE3C" box="[693,921,367,393]" country="Tanzania" name="Kilimanjaro" pageId="47" pageNumber="48">Kilimanjaro Region</collectingRegion>
, Siha District, “ 
<collectingRegion id="49B11382CC1E4C0BF305FE80FB76FE3C" box="[1101,1256,367,393]" country="Tanzania" name="Kilimanjaro" pageId="47" pageNumber="48">Kilimandjaro</collectingRegion>
, Kibonoto” (today also known as Kibongoto or Kibogoto), 1905–1906, October (written in German as “okt.” for “Oktober” on a small second label), 1905?, Sjöstedt leg., on a third and red label with the number “149 67”, genitalia slide number 15/092009 I. Lehmann (
<collectionCode id="ED6445A5CC1E4C0BF6ECFE34FE74FE40" box="[420,490,475,501]" country="Sweden" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="47" pageNumber="48" type="Museum">NRM</collectionCode>
). 
<emphasis id="B9010172CC1E4C0BF6B4FE33FDF0FE40" box="[508,622,476,501]" italics="true" pageId="47" pageNumber="48">
<typeStatus id="54CE63C2CC1E4C0BF6B4FE33FDF0FE40" box="[508,622,476,501]" pageId="47" pageNumber="48" type="paratype">Paratypes</typeStatus>
</emphasis>
: 
<specimenCount id="9D7316E9CC1E4C0BF534FE33FD64FE40" box="[636,762,475,502]" pageId="47" pageNumber="48" type="female">one female</specimenCount>
, 1905–1906, 
<date id="FFCBFBA0CC1E4C0BF4D8FE34FBD1FE43" box="[912,1103,475,502]" pageId="47" pageNumber="48" value="1905-10-14">14 October 1905</date>
?, Sjöstedt leg., with one additional information on a second label regarding the locality: “Kibonoto cultural zone” (the latter is written in German as “kulturz.” for “Kulturzone”), on a third and red label with the number “150 67”, genitalia slide number 28/022012 I. Lehmann (
<collectionCode id="ED6445A5CC1E4C0BF66EFDA8FEF7FDD4" box="[294,361,583,609]" country="Sweden" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="47" pageNumber="48" type="Museum">NRM</collectionCode>
) and the following 
<specimenCount id="9D7316E9CC1E4C0BF50CFDA8FD39FDD4" box="[580,679,583,610]" pageId="47" pageNumber="48" type="male">25 males</specimenCount>
from the same locality and collector: 
<specimenCount id="9D7316E9CC1E4C0BF307FDA8FB4CFDD4" box="[1103,1234,583,609]" pageId="47" pageNumber="48" type="male">three males</specimenCount>
, 1905–1906, September, 1905? (no genitalia dissection done); 
<specimenCount id="9D7316E9CC1E4C0BF5D4FD83FC98FD30" box="[668,774,619,646]" pageId="47" pageNumber="48" type="male">one male</specimenCount>
, 
<date id="FFCBFBA0CC1E4C0BF45AFD84FC6CFD33" box="[786,1010,619,646]" pageId="47" pageNumber="48" value="1905-09-20">20 September 1905</date>
?, genitalia slide number 23/042013 I. Lehmann (
<collectionCode id="ED6445A5CC1E4C0BF646FD60FECAFD1C" box="[270,340,655,681]" country="Sweden" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="47" pageNumber="48" type="Museum">NRM</collectionCode>
); 
<specimenCount id="9D7316E9CC1E4C0BF62EFD7FFE53FD1C" box="[358,461,655,682]" pageId="47" pageNumber="48" type="male">one male</specimenCount>
, 
<date id="FFCBFBA0CC1E4C0BF69FFD60FD2FFD1F" box="[471,689,655,682]" pageId="47" pageNumber="48" value="1905-09-20">20 September 1905</date>
?, genitalia slide number 08/022016 I. Lehmann (
<collectionCode id="ED6445A5CC1E4C0BF383FD60FA8FFD1C" box="[1227,1297,655,681]" country="Sweden" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="47" pageNumber="48" type="Museum">NRM</collectionCode>
); 
<specimenCount id="9D7316E9CC1E4C0BF26BFD60FA06FD1C" box="[1315,1432,655,681]" pageId="47" pageNumber="48" type="male">five males</specimenCount>
, 1905–1906, 
<date id="FFCBFBA0CC1E4C0BF66CFD5CFD9FFD7B" box="[292,513,691,718]" pageId="47" pageNumber="48" value="1905-09-24">24 September 1905</date>
? (no genitalia dissection done), including 
<specimenCount id="9D7316E9CC1E4C0BF497FD5BFBDBFD78" box="[991,1093,691,718]" pageId="47" pageNumber="48" type="male">one male</specimenCount>
that is determined by Sjöstedt as “ 
<taxonomicName id="4C75A6E3CC1E4C0BF78CFD38FD99FD44" authority="Auriv." authorityName="Auriv." box="[196,519,727,753]" class="Insecta" family="Cossidae" genus="Metarbela" kingdom="Animalia" order="Lepidoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="umtaliana">
<emphasis id="B9010172CC1E4C0BF78CFD38FE2AFD44" box="[196,436,727,753]" italics="true" pageId="47" pageNumber="48">Metarbela umtaliana</emphasis>
Auriv.
</taxonomicName>
”; 
<specimenCount id="9D7316E9CC1E4C0BF556FD37FD17FD44" box="[542,649,727,754]" pageId="47" pageNumber="48" type="male">one male</specimenCount>
, 1905–1906, 
<date id="FFCBFBA0CC1E4C0BF46CFD38FB98FD47" box="[804,1030,727,754]" pageId="47" pageNumber="48" value="1905-09-30">30 September 1905</date>
?, genitalia slide number 01/072012 I. Lehmann (
<collectionCode id="ED6445A5CC1E4C0BF665FD14FEEDFCA0" box="[301,371,763,789]" country="Sweden" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="47" pageNumber="48" type="Museum">NRM</collectionCode>
); 
<specimenCount id="9D7316E9CC1E4C0BF6C1FD13FE6AFCA0" box="[393,500,763,790]" pageId="47" pageNumber="48" type="male">one male</specimenCount>
, 1905–1906, 
<date id="FFCBFBA0CC1E4C0BF5D9FD14FCEBFCA3" box="[657,885,763,790]" pageId="47" pageNumber="48" value="1905-09-30">30 September 1905</date>
?, genitalia slide number 20/122009 I. Lehmann (
<collectionCode id="ED6445A5CC1E4C0BF7D7FCF0FF7BFC8C" box="[159,229,799,825]" country="Sweden" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="47" pageNumber="48" type="Museum">NRM</collectionCode>
). No genitalia dissections were done for the following specimens: 
<specimenCount id="9D7316E9CC1E4C0BF4BBFCCFFBC1FC8C" box="[1011,1119,799,826]" pageId="47" pageNumber="48" type="male">one male</specimenCount>
, 1905–1906, 
<date id="FFCBFBA0CC1E4C0BF3B6FCF0FF4DFCEB" pageId="47" pageNumber="48" value="1905-09-30">30 September 1905</date>
?; 
<specimenCount id="9D7316E9CC1E4C0BF7A4FCACFEC6FCE8" box="[236,344,835,861]" pageId="47" pageNumber="48" type="male">six males</specimenCount>
, 1905–1906, 
<date id="FFCBFBA0CC1E4C0BF6BAFCACFD0EFCEB" box="[498,656,835,862]" pageId="47" pageNumber="48" value="1905-10">October 1905</date>
?; 
<specimenCount id="9D7316E9CC1E4C0BF5E1FCABFC8CFCE8" box="[681,786,835,862]" pageId="47" pageNumber="48" type="male">one male</specimenCount>
, same date with one additional information regarding the locality “Kibonoto cultural zone” (the latter is written in German as “kulturz.” for “Kulturzone”); 
<specimenCount id="9D7316E9CC1E4C0BF3AAFC87FAD5FC34" box="[1250,1355,871,898]" pageId="47" pageNumber="48" type="male">one male</specimenCount>
, 1905– 1906, 
<date id="FFCBFBA0CC1E4C0BF795FC64FE00FC13" box="[221,414,907,934]" pageId="47" pageNumber="48" value="1905-10-03">03 October 1905</date>
?; 
<specimenCount id="9D7316E9CC1E4C0BF6FEFC63FDB5FC10" box="[438,555,907,934]" pageId="47" pageNumber="48" type="male">two males</specimenCount>
, 1905–1906, 
<date id="FFCBFBA0CC1E4C0BF58BFC64FC1DFC13" box="[707,899,907,934]" pageId="47" pageNumber="48" value="1905-10-04">04.October 1905</date>
?; 
<specimenCount id="9D7316E9CC1E4C0BF4D3FC63FB9AFC10" box="[923,1028,907,934]" pageId="47" pageNumber="48" type="male">one male</specimenCount>
, 1905–1906, 
<date id="FFCBFBA0CC1E4C0BF3D4FC64FAC2FC13" box="[1180,1372,907,934]" pageId="47" pageNumber="48" value="1905-10-12">12 October 1905</date>
?; 
<specimenCount id="9D7316E9CC1E4C0BF23CFC63FF4EFC7C" pageId="47" pageNumber="48" type="male">one male</specimenCount>
, 1905–1906, 
<date id="FFCBFBA0CC1E4C0BF62FFC40FDDEFC7F" box="[359,576,943,970]" pageId="47" pageNumber="48" value="1905-11-03">03 November 1905</date>
? One male, 
<collectingCountry id="F3629DF0CC1E4C0BF582FC40FCACFC7C" box="[714,818,943,969]" name="Tanzania" pageId="47" pageNumber="48">Tanzania</collectingCountry>
, 
<collectingRegion id="49B11382CC1E4C0BF474FC40FB80FC7C" box="[828,1054,943,969]" country="Tanzania" name="Kilimanjaro" pageId="47" pageNumber="48">Kilimanjaro Region</collectingRegion>
, Hai District, Makoa, 
<date id="FFCBFBA0CC1E4C0BF25DFC40FF4CFC5B" pageId="47" pageNumber="48" value="1952-02-14">14 February 1952</date>
, no collector mentioned, a second label states “ 
<taxonomicName id="4C75A6E3CC1E4C0BF5B9FC3CFBA0FC58" authority="Aur." authorityName="Aurivillius" authorityYear="1901" box="[753,1086,979,1005]" class="Insecta" family="Cossidae" genus="Metarbelodes" kingdom="Animalia" order="Lepidoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="umtaliana">
<emphasis id="B9010172CC1E4C0BF5B9FC3CFB98FC58" box="[753,1030,979,1005]" italics="true" pageId="47" pageNumber="48">Metarbelodes umtaliana</emphasis>
Aur.
</taxonomicName>
det. F. Daniel 1955”, genitalia slide number 21/052015 I. Lehmann (
<collectionCode id="ED6445A5CC1E4C0BF50BFC18FD0BFBA4" box="[579,661,1015,1041]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="47" pageNumber="48">SMNS</collectionCode>
); 
<specimenCount id="9D7316E9CC1E4C0BF5E1FC17FC8DFBA4" box="[681,787,1015,1042]" pageId="47" pageNumber="48" type="male">one male</specimenCount>
, “ 
<collectionCode id="ED6445A5CC1E4C0BF461FC18FCA5FBA4" box="[809,827,1015,1041]" name="Tavera, Department of Geology and Geophysics" pageId="47" pageNumber="48">T</collectionCode>
. 
<collectionCode id="ED6445A5CC1E4C0BF476FC18FCCEFBA4" box="[830,848,1015,1041]" name="Tavera, Department of Geology and Geophysics" pageId="47" pageNumber="48">T</collectionCode>
.O.Afrika” (written in German for “Tanganyika Territorium Ost-Afrika”), today 
<collectingCountry id="F3629DF0CC1E4C0BF691FBF4FDDFFB80" box="[473,577,1051,1077]" name="Tanzania" pageId="47" pageNumber="48">Tanzania</collectingCountry>
, 
<collectingRegion id="49B11382CC1E4C0BF502FBF4FCB5FB80" box="[586,811,1051,1077]" country="Tanzania" name="Kilimanjaro" pageId="47" pageNumber="48">Kilimanjaro Region</collectingRegion>
, Hai District, Makoa, 
<date id="FFCBFBA0CC1E4C0BF357FBF4FA9DFB83" box="[1055,1283,1051,1078]" pageId="47" pageNumber="48" value="1959-01-07" valueMax="1959-01-27" valueMin="1959-01-07">07–27 January 1959</date>
, Lindner leg., genitalia slide number 19/102015 I. Lehmann (
<collectionCode id="ED6445A5CC1E4C0BF5FEFBD0FC96FBEC" box="[694,776,1087,1113]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="47" pageNumber="48">SMNS</collectionCode>
); 
<specimenCount id="9D7316E9CC1E4C0BF456FBD0FC09FBEC" box="[798,919,1087,1113]" pageId="47" pageNumber="48" type="male">five males</specimenCount>
, same locality, 
<date id="FFCBFBA0CC1E4C0BF303FBD0FAA8FBEF" box="[1099,1334,1087,1114]" pageId="47" pageNumber="48" value="1959-01-07" valueMax="1959-01-27" valueMin="1959-01-07">07–27 January 1959</date>
, Lindner leg., no genitalia dissection done, det. I. Lehmann (
<collectionCode id="ED6445A5CC1E4C0BF59DFB8CFCB9FBC8" box="[725,807,1123,1149]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="47" pageNumber="48">SMNS</collectionCode>
); 
<specimenCount id="9D7316E9CC1E4C0BF473FB8BFC3AFBC8" box="[827,932,1123,1150]" pageId="47" pageNumber="48" type="male">one male</specimenCount>
, 
<collectingCountry id="F3629DF0CC1E4C0BF4E6FB8CFB88FBC8" box="[942,1046,1123,1149]" name="Tanzania" pageId="47" pageNumber="48">Tanzania</collectingCountry>
, 
<collectingRegion id="49B11382CC1E4C0BF368FB8CFB52FBC8" box="[1056,1228,1123,1149]" country="Tanzania" name="Arusha" pageId="47" pageNumber="48">Arusha Region</collectingRegion>
, Arumeru District, Usa River, 
<quantity id="4C8D7085CC1E4C0BF65CFB68FEECFB17" box="[276,370,1159,1186]" metricMagnitude="0" metricUnit="m" metricValue="1.17" pageId="47" pageNumber="48" unit="m" value="1.17">1.170 m</quantity>
, 
<date id="FFCBFBA0CC1E4C0BF635FB68FDD8FB14" box="[381,582,1159,1185]" pageId="47" pageNumber="48" value="1992-02-03">03 February 1992</date>
, L. Aarvik leg., genitalia slide number 24/042012 I. Lehmann (
<collectionCode id="ED6445A5CC1E4C0BF24DFB68FAFFFB14" box="[1285,1377,1159,1185]" country="Norway" httpUri="http://grbio.org/cool/2knt-7f1r" name="Natural History Museum, University of Oslo" pageId="47" pageNumber="48" type="Museum">NHMO</collectionCode>
); 
<specimenCount id="9D7316E9CC1E4C0BF23CFB67FF4EFB70" pageId="47" pageNumber="48" type="male">one male</specimenCount>
, 
<collectingCountry id="F3629DF0CC1E4C0BF793FB44FEDDFB70" box="[219,323,1195,1221]" name="Tanzania" pageId="47" pageNumber="48">Tanzania</collectingCountry>
, same locality and elevation, 
<date id="FFCBFBA0CC1E4C0BF5C6FB44FCC6FB70" box="[654,856,1195,1221]" pageId="47" pageNumber="48" value="1992-02-04">04 February 1992</date>
, L. Aarvik leg., genitalia slide number 22/042012 I. Lehmann (
<collectionCode id="ED6445A5CC1E4C0BF658FB20FEF2FB5C" box="[272,364,1231,1257]" country="Norway" httpUri="http://grbio.org/cool/2knt-7f1r" name="Natural History Museum, University of Oslo" pageId="47" pageNumber="48" type="Museum">NHMO</collectionCode>
). Note: 
<taxonomicName id="4C75A6E3CC1E4C0BF68BFB20FD39FB5C" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[451,679,1231,1257]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="bergsteni">
<emphasis id="B9010172CC1E4C0BF68BFB20FD39FB5C" box="[451,679,1231,1257]" italics="true" pageId="47" pageNumber="48">Lukeniana bergsteni</emphasis>
</taxonomicName>
was determined and published by 
<bibRefCitation id="EFE4A091CC1E4C0BF360FB20FB6DFB5C" author="Aurivillius, C." box="[1064,1267,1231,1257]" pageId="47" pageNumber="48" pagination="49 - 50" refId="ref75086" refString="Aurivillius, C. (1910) Lepidoptera. Arbelidae. In: Sjostedt, Y. (Ed.), Wissenschaftliche Ergebnisse der schwedischen zoologischen Expedition nach dem Kilimandjaro, dem Meru und den umgebenen Massaisteppen Deutsch-Ostafrikas 1905 - 1906, Part 9, pp. 49 - 50, 1 b / w fig." type="journal article" year="1910">Aurivillius (1910)</bibRefCitation>
as 
<emphasis id="B9010172CC1E4C0BF250FB20FE81FAB8" italics="true" pageId="47" pageNumber="48">
“ 
<taxonomicName id="4C75A6E3CC1E4C0BF26AFB20FE8FFAB8" class="Insecta" family="Cossidae" genus="Metarbela" kingdom="Animalia" order="Lepidoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="umtaliana">Metarbela umtaliana</taxonomicName>
.”
</emphasis>
Aurivillius stated in the same context that “ 
<taxonomicName id="4C75A6E3CC1E4C0BF451FB1BFB82FAB8" authority="Baker" authorityName="Baker" box="[793,1052,1267,1293]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="obliqualinea">
<emphasis id="B9010172CC1E4C0BF451FB1BFC50FAB8" box="[793,974,1267,1293]" italics="true" pageId="47" pageNumber="48">M. obliqualinea</emphasis>
Baker
</taxonomicName>
” (for Bethune-Baker) is probably identical with 
<taxonomicName id="4C75A6E3CC1E4C0BF676FAF8FE2EFA84" box="[318,432,1303,1329]" class="Insecta" family="Cossidae" genus="Metarbela" kingdom="Animalia" order="Lepidoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="umtaliana">
<emphasis id="B9010172CC1E4C0BF676FAF8FE2EFA84" box="[318,432,1303,1329]" italics="true" pageId="47" pageNumber="48">umtaliana</emphasis>
</taxonomicName>
(both are treated here as separate species and in different genera). The original text is written in German as follows (
<bibRefCitation id="EFE4A091CC1E4C0BF6BFFAD4FD28FAE0" author="Aurivillius, C." box="[503,694,1339,1365]" pageId="47" pageNumber="48" pagination="49 - 50" refId="ref75086" refString="Aurivillius, C. (1910) Lepidoptera. Arbelidae. In: Sjostedt, Y. (Ed.), Wissenschaftliche Ergebnisse der schwedischen zoologischen Expedition nach dem Kilimandjaro, dem Meru und den umgebenen Massaisteppen Deutsch-Ostafrikas 1905 - 1906, Part 9, pp. 49 - 50, 1 b / w fig." type="journal article" year="1910">Aurivillius 1910</bibRefCitation>
, p. 49): “ 
<taxonomicName id="4C75A6E3CC1E4C0BF46AFAD4FC09FAE0" box="[802,919,1339,1365]" class="Insecta" family="Arbelidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="family">Arbelidae</taxonomicName>
. 
<taxonomicName id="4C75A6E3CC1E4C0BF4EBFAD4FB68FAE3" authority="AURIV." authorityName="AURIV." box="[931,1270,1339,1366]" class="Insecta" family="Cossidae" genus="Metarbela" kingdom="Animalia" order="Lepidoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="umtaliana">
<emphasis id="B9010172CC1E4C0BF4EBFAD4FB0DFAE0" box="[931,1171,1339,1365]" italics="true" pageId="47" pageNumber="48">Metarbela umtaliana</emphasis>
AURIV.
</taxonomicName>
<collectingRegion id="49B11382CC1E4C0BF3B7FAD4FA07FAE0" box="[1279,1433,1339,1365]" country="Tanzania" name="Kilimanjaro" pageId="47" pageNumber="48">Kilimandjaro</collectingRegion>
: 
<quantity id="4C8D7085CC1E4C0BF7DFFAB0FEDAFACF" box="[151,324,1375,1402]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.9" metricValueMin="1.0" pageId="47" pageNumber="48" unit="m" value="1450.0" valueMax="1900.0" valueMin="1000.0">
<geoCoordinate id="EE41BBA7CC1E4C0BF7DFFAB0FF48FACC" box="[151,214,1375,1401]" degrees="1,000" direction="north" orientation="latitude" pageId="47" pageNumber="48" precision="55" value="1.0">1,000</geoCoordinate>
<geoCoordinate id="EE41BBA7CC1E4C0BF79EFAB0FEBAFACC" box="[214,292,1375,1401]" degrees="1,900" direction="west" orientation="longitude" pageId="47" pageNumber="48" precision="55" value="-1.9">‒1,900</geoCoordinate>
m
</quantity>
. 44 ♁, 
<specimenCount id="9D7316E9CC1E4C0BF6DEFAB0FE5FFACF" box="[406,449,1375,1402]" pageId="47" pageNumber="48" type="female">1 ♀</specimenCount>
. Sept.—Dez. Flug schnell, aber unstätig. Die ♁ kamen im Zelte an′s Licht. Die ♁ stim- men gut mit typischen Stücken aus dem Mashunalande überein. Das bisher unbekannte Weibchen ist dem ♁ ähnlich gefärbt und gezeichnet, hat aber eine Flügelspannung von 
<quantity id="4C8D7085CC1E4C0BF47BFA48FC1DFA77" box="[819,899,1447,1474]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" pageId="47" pageNumber="48" unit="mm" value="42.0">42 mm</quantity>
und sehr kurze Kammzähne der Fühler. Die ♁ wechseln in der Grösse zwischen 20 und 
<quantity id="4C8D7085CC1E4C0BF52AFA24FD2BFA53" box="[610,693,1483,1510]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="47" pageNumber="48" unit="mm" value="30.0">30 mm</quantity>
. Nach der Beschreibung scheint 
<taxonomicName id="4C75A6E3CC1E4C0BF36EFA23FABAFA50" authority="Baker" authorityName="Baker" box="[1062,1316,1483,1509]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="obliqualinea">
<emphasis id="B9010172CC1E4C0BF36EFA23FB44FA50" box="[1062,1242,1483,1509]" italics="true" pageId="47" pageNumber="48">M. obliqualinea</emphasis>
Baker
</taxonomicName>
(An. N.H. (8) 
<specimenCount id="9D7316E9CC1E4C0BF7F7FA00FF77F9BF" box="[191,233,1519,1546]" pageId="47" pageNumber="48" type="generic">3 p.</specimenCount>
425, 1909) identisch mit 
<taxonomicName id="4C75A6E3CC1E4C0BF541FA00FDE5F9BC" box="[521,635,1519,1545]" class="Insecta" family="Cossidae" genus="Metarbela" kingdom="Animalia" order="Lepidoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="umtaliana">
<emphasis id="B9010172CC1E4C0BF541FA00FDE5F9BC" box="[521,635,1519,1545]" italics="true" pageId="47" pageNumber="48">umtaliana</emphasis>
</taxonomicName>
zu sein. 
<emphasis id="B9010172CC1E4C0BF59FFA00FD79F9BC" box="[727,743,1519,1545]" italics="true" pageId="47" pageNumber="48">”</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC1E4C14F78FF9FCFB5FFC58" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48" type="description">
<paragraph id="8BCADD60CC1E4C0BF78FF9FCFA08F9C0" blockId="47.[151,1437,367,2014]" pageId="47" pageNumber="48">
<emphasis id="B9010172CC1E4C0BF78FF9FCFEC7F998" bold="true" box="[199,345,1555,1581]" pageId="47" pageNumber="48">Description.</emphasis>
Male. 
<emphasis id="B9010172CC1E4C0BF6EDF9FCFE7CF998" box="[421,482,1555,1581]" italics="true" pageId="47" pageNumber="48">Head</emphasis>
: Yellow ochre; eyes brown with black spots; antenna 0.50× length of forewing, bipectinate, branches 7× width of shaft, covered with few ivory-yellow scales laterally, shaft covered with ivory-yellow scales dorsally; antennal tips slightly spatulate and with two long scales, bending towards apex; labial palpi ochre.
</paragraph>
<paragraph id="8BCADD60CC1E4C0BF78FF990FB86F80C" blockId="47.[151,1437,367,2014]" pageId="47" pageNumber="48">
<emphasis id="B9010172CC1E4C0BF78FF990FE86F92C" box="[199,280,1663,1689]" italics="true" pageId="47" pageNumber="48">Thorax</emphasis>
: Patagia and tegulae with long hair-like scales of ochre mixed with yellow ochre. A small crest of ivoryyellow on metathorax. Hindlegs ochre with fine hair-like scales, shiny; a pair of narrow, long tibial spurs of unequal length, outer spur 
<emphasis id="B9010172CC1E4C0BF62CF927FE18F954" box="[356,390,1736,1761]" italics="true" pageId="47" pageNumber="48">ca.</emphasis>
<quantity id="4C8D7085CC1E4C0BF6C5F928FE78F957" box="[397,486,1735,1762]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="47" pageNumber="48" unit="mm" value="1.1">1.1 mm</quantity>
, inner spur 
<emphasis id="B9010172CC1E4C0BF523F927FD12F954" box="[619,652,1736,1761]" italics="true" pageId="47" pageNumber="48">ca.</emphasis>
1.0 mm. Forewing length in 
<typeStatus id="54CE63C2CC1E4C0BF49EF928FBA6F954" box="[982,1080,1735,1761]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
14.0 mm (wingspan 
<quantity id="4C8D7085CC1E4C0BF26FF928FA0EF954" box="[1319,1424,1735,1762]" metricMagnitude="-2" metricUnit="m" metricValue="3.15" pageId="47" pageNumber="48" unit="mm" value="31.5">31.5 mm</quantity>
); upperside ochre; costal margin sepia with few sepia coloured striae; termen with striae of sepia; bent terminal line and subterminal line of sepia, more or less parallel, from near apex to end of CuA 
<subScript id="17F1DF25CC1E4C0BF37AF8CDFBA4F885" attach="left" box="[1074,1082,1826,1840]" fontSize="6" pageId="47" pageNumber="48">2</subScript>
and to middle of CuA 
<subScript id="17F1DF25CC1E4C0BF27DF8CDFAA3F885" attach="left" box="[1333,1341,1826,1840]" fontSize="6" pageId="47" pageNumber="48">1</subScript>
, respectively; CuA 
<subScript id="17F1DF25CC1E4C0BF652F8A9FEBCF8E1" attach="left" box="[282,290,1862,1876]" fontSize="6" pageId="47" pageNumber="48">2</subScript>
more narrow, pure white, edged sepia above; all remaining veins not distinctly coloured; cilia long, 
<quantity id="4C8D7085CC1E4C0BF231F8DCFF5BF8C7" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="47" pageNumber="48" unit="mm" value="1.8">1.8 mm</quantity>
, yellow ochre, glossy,. Underside of forewing roughly scaled, pale ochre, glossy, costal margin sepia; subterminal line reduced to few faded spots of sepia. Hindwing upperside ivory-yellow or pure white, glossy; cilia same colour as in forewing, 
<quantity id="4C8D7085CC1E4C0BF6DCF870FE74F80F" box="[404,490,1951,1978]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="47" pageNumber="48" unit="mm" value="1.8">1.8 mm</quantity>
long; underside as in forewing but without striae.
</paragraph>
<paragraph id="8BCADD60CC1E4C14F78FF82CFBDCFE40" blockId="47.[151,1437,367,2014]" lastBlockId="48.[151,1437,150,1941]" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48">
<emphasis id="B9010172CC1E4C0BF78FF82CFEAFF868" box="[199,305,1987,2013]" italics="true" pageId="47" pageNumber="48">Abdomen</emphasis>
: Ochre with ivory-yellow, glossy; abdominal tuft one-fifth of abdomen length. Genitalia (
<figureCitation id="134EC1E5CC1E4C0BF27AF82CFA06F868" box="[1330,1432,1987,2013]" captionStart="FIGURE 14" captionStartId="65.[151,250,1754,1778]" captionTargetBox="[159,1425,534,1722]" captionTargetId="figure-191@65.[152,1436,525,1730]" captionTargetPageId="65" captionText="FIGURE 14. a. Lukeniana bergsteni, paratype, male, Tanzania, Kilimanjaro Region, Mount Kilimanjaro, Kibonoto; b. L. carolae, holotype, male, Malawi, Central Region, Mount Ntchisi, Ntchisi Forest Reserve; c. L. stevecollinsi, holotype, male, Malawi, Northern Region, Nyika Plateau, Nyika National Park, near Thazima Gate; d. L. butleri, paratype, male, Zimbabwe, Harare Province, Harare, “Fawlty Towers”; e. L. kollhorsti, holotype, male, Zimbabwe, Manicaland Province, Busi Farm; f. Zambezia diredaouaensis, holotype, male, Ethiopia, Harar Plateau, Dire Daua, today: chartered city of Dire Dawa." figureDoi="http://doi.org/10.5281/zenodo.7840822" httpUri="https://zenodo.org/record/7840822/files/figure.png" pageId="47" pageNumber="48">Figs 14a</figureCitation>
; 
<figureCitation id="134EC1E5CC014C14F7DFFF79FF5EFF04" box="[151,192,150,177]" captionStart="FIGURE 15" captionStartId="66.[151,250,1656,1680]" captionTargetBox="[172,1419,191,1622]" captionTargetId="figure-28@66.[151,1436,181,1632]" captionTargetPageId="66" captionText="FIGURE 15. Examples of male genitalia in lateral view: a. Zambezia madambae, paratype, with extremely small thorn-like process and small weakly-sclerotized projection (red arrows); b. Lukeniana timdavenporti, paratype, with strongly bent ventral edge of valva and well developed thorn-like process (red arrows); c. L. bergsteni, paratype, without strongly bent ventral edge of valva and two thorn-like processes (red arrows); d. L. butleri, paratype, with many long setae on valva that appear to be black in colour and one thorn-like process (red arrows)." figureDoi="http://doi.org/10.5281/zenodo.7840824" httpUri="https://zenodo.org/record/7840824/files/figure.png" pageId="48" pageNumber="49">15c</figureCitation>
) with uncus ovoid with narrowly rounded tips, outer edge slightly C-shaped, inner edge rounded at base, lobes densely covered with short setae ventrally, basal edge of uncus strongly bent at middle; gnathos arms long (1.3× basal width of valva), touching coastal margin of valva; valva narrow and elongate ovoid, costa with few short setae; sacculus with few long setae; weakly-sclerotized projection setose, rectangular, shorter in length than thick, short thorn-like process below, latter hollow with a rounded tip, without setae, a second short thorn-like process at its base; median sector of valva with few scattered setae on inner side; short, broad, ovoid emargination extending up to 35% of length of valva between weakly-sclerotized projection and thorn-like process; ventral side of valva not bent at middle; saccus triangular with a rounded tip. Juxta larger than saccus, broad with two acuminate tips, each bearing triangular thorn-like process, emargination between tips 90% of length of juxta. Phallus long, 1.2× length of valva, not trumpet-like, strongly bent near middle, bilobed with cleft at each end.
</paragraph>
<paragraph id="8BCADD60CC014C14F78FFE10FC22FD78" blockId="48.[151,1437,150,1941]" pageId="48" pageNumber="49">
<emphasis id="B9010172CC014C14F78FFE10FEBBFDAC" bold="true" box="[199,293,511,537]" pageId="48" pageNumber="49">Female.</emphasis>
<emphasis id="B9010172CC014C14F666FE10FEF5FDAC" box="[302,363,511,537]" italics="true" pageId="48" pageNumber="49">Head</emphasis>
and 
<emphasis id="B9010172CC014C14F6EDFE10FE68FDAC" box="[421,502,511,537]" italics="true" pageId="48" pageNumber="49">Thorax</emphasis>
: Essentially as described for male, except antenna 0.36× length of forewing, unipectinate, with branches 1.0× width of shaft, antennal tips very short, acuminate, without hair-like scales bending towards apex. Forewing length 
<quantity id="4C8D7085CC014C14F54AFDA9FDF6FDD4" box="[514,616,582,609]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" pageId="48" pageNumber="49" unit="mm" value="21.5">21.5 mm</quantity>
(wingspan 45.0 mm). Forewing very broad, almost 1.5× width of male forewing; outer half of forewing faded, with reticulate pattern absent, terminal striae faint; CuA 
<subScript id="17F1DF25CC014C14F38BFD92FB55FD3E" attach="left" box="[1219,1227,637,651]" fontSize="6" pageId="48" pageNumber="49">2</subScript>
very broad, ivoryyellow mixed with pure white, edged by a broad band of sepia above. More narrow tibial spurs, and hence, less pronounced than in male, outer spur 
<emphasis id="B9010172CC014C14F57BFD5CFDCAFD79" box="[563,596,691,716]" italics="true" pageId="48" pageNumber="49">ca.</emphasis>
<quantity id="4C8D7085CC014C14F513FD5CFD2DFD78" box="[603,691,691,717]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="48" pageNumber="49" unit="mm" value="0.9">0.9 mm</quantity>
, inner spur 
<emphasis id="B9010172CC014C14F47EFD5CFCC6FD79" box="[822,856,691,716]" italics="true" pageId="48" pageNumber="49">ca.</emphasis>
<quantity id="4C8D7085CC014C14F417FD5CFC26FD78" box="[863,952,691,717]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="48" pageNumber="49" unit="mm" value="0.8">0.8 mm</quantity>
.
</paragraph>
<paragraph id="8BCADD60CC014C14F78FFD38FB5FFC58" blockId="48.[151,1437,150,1941]" pageId="48" pageNumber="49">
<emphasis id="B9010172CC014C14F78FFD38FEAFFD44" box="[199,305,727,753]" italics="true" pageId="48" pageNumber="49">Abdomen</emphasis>
: Papillae anales broad, 8-shaped in posterior view, densely setose with long and short setae. On segment 8 only very few long setae along posterior margin; one very broad latero-ventral sclerotized band, triangular with acuminate end towards base of anterior apophysis. This band is covering 60% of ventral side of segment 8 with a broader second band behind it; both only partly fused along posterior margin, not along anterior margin. Dorso-anterior margin of abdominal plate without emargination. Posterior end of posterior apophysis narrow, stick-like, 
<emphasis id="B9010172CC014C14F233FC88FA02FC35" box="[1403,1436,871,896]" italics="true" pageId="48" pageNumber="49">ca.</emphasis>
40% length of apophysis, slightly C-shaped and sclerotized; remaining part of posterior apophysis broader towards tip, slightly bent towards ventral side of segment 8, tip rectangular. Anterior apophysis with rounded tip slightly bent towards ventral side of segment 8. Posterior apophyses slightly longer than anterior apophyses.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC014C14F78FFC19FA03F9BC" pageId="48" pageNumber="49" type="diagnosis">
<paragraph id="8BCADD60CC014C14F78FFC19FA0CFBEC" blockId="48.[151,1437,150,1941]" pageId="48" pageNumber="49">
<emphasis id="B9010172CC014C14F78FFC19FEDCFBA5" bold="true" box="[199,322,1014,1040]" pageId="48" pageNumber="49">Diagnosis.</emphasis>
<taxonomicName id="4C75A6E3CC014C14F600FC18FDB5FBA4" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[328,555,1015,1041]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="bergsteni">
<emphasis id="B9010172CC014C14F600FC18FDB5FBA4" box="[328,555,1015,1041]" italics="true" pageId="48" pageNumber="49">Lukeniana bergsteni</emphasis>
</taxonomicName>
resembles 
<taxonomicName id="4C75A6E3CC014C14F5E0FC18FCD5FBA4" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[680,843,1015,1041]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="jankiellandi">
<emphasis id="B9010172CC014C14F5E0FC18FCD5FBA4" box="[680,843,1015,1041]" italics="true" pageId="48" pageNumber="49">L. jankiellandi</emphasis>
</taxonomicName>
(see diagnosis of the latter species) but can be distinguished by three characters: (i) the lobes of the uncus are narrowly ovoid (rounded in 
<taxonomicName id="4C75A6E3CC014C14F314FBF4FA9EFB80" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[1116,1280,1051,1077]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="jankiellandi">
<emphasis id="B9010172CC014C14F314FBF4FA9EFB80" box="[1116,1280,1051,1077]" italics="true" pageId="48" pageNumber="49">L. jankiellandi</emphasis>
</taxonomicName>
); (ii) the gnathos arms are long (touching the costa of valva), with (iii) their horizontal part as long as the basal width of valva.
</paragraph>
<paragraph id="8BCADD60CC014C14F78FFB8CFDCFFACC" blockId="48.[151,1437,150,1941]" pageId="48" pageNumber="49">
Regarding the statements of 
<bibRefCitation id="EFE4A091CC014C14F54BFB8CFD66FBC8" author="Aurivillius, C." box="[515,760,1123,1149]" pageId="48" pageNumber="49" pagination="49 - 50" refId="ref75086" refString="Aurivillius, C. (1910) Lepidoptera. Arbelidae. In: Sjostedt, Y. (Ed.), Wissenschaftliche Ergebnisse der schwedischen zoologischen Expedition nach dem Kilimandjaro, dem Meru und den umgebenen Massaisteppen Deutsch-Ostafrikas 1905 - 1906, Part 9, pp. 49 - 50, 1 b / w fig." type="journal article" year="1910">Aurivillius (1910: 49)</bibRefCitation>
that specimens from 
<collectingRegion id="49B11382CC014C14F4AFFB8CFB21FBC8" box="[999,1215,1123,1149]" country="Tanzania" name="Kilimanjaro" pageId="48" pageNumber="49">Mount Kilimanjaro</collectingRegion>
belong to 
<taxonomicName id="4C75A6E3CC014C14F27CFB8CFED5FB14" authorityName="Aurivillius" authorityYear="1901" class="Insecta" family="Cossidae" genus="Metarbelodes" kingdom="Animalia" order="Lepidoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="umtaliana">
<emphasis id="B9010172CC014C14F27CFB8CFED5FB14" italics="true" pageId="48" pageNumber="49">Metarbelodes umtaliana</emphasis>
</taxonomicName>
, it has to be noted that when compared to 
<taxonomicName id="4C75A6E3CC014C14F46DFB68FC34FB14" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[805,938,1159,1185]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="bergsteni">
<emphasis id="B9010172CC014C14F46DFB68FC34FB14" box="[805,938,1159,1185]" italics="true" pageId="48" pageNumber="49">L. bergsteni</emphasis>
</taxonomicName>
, the male genitalia of 
<taxonomicName id="4C75A6E3CC014C14F3ECFB68FAA5FB14" authorityName="Aurivillius" authorityYear="1901" box="[1188,1339,1159,1185]" class="Insecta" family="Cossidae" genus="Metarbelodes" kingdom="Animalia" order="Lepidoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="umtaliana">
<emphasis id="B9010172CC014C14F3ECFB68FAA5FB14" box="[1188,1339,1159,1185]" italics="true" pageId="48" pageNumber="49">M. umtaliana</emphasis>
</taxonomicName>
differ in the following features: (i) a narrow, finger-shaped saccus; (ii) a shorter emargination of the valva; (iii) no thorn-like appendices at the base of the thorn-like process; (iv) a more curved, narrower, and longer thorn-like process on the valva that is bent downwards from the costa and (v) shorter gnathos arms. Additionally, the forewing upperside shows two remarkable differences: (i) the terminal line is straight in 
<emphasis id="B9010172CC014C14F4D5FAF8FBA2FA84" box="[925,1084,1303,1329]" italics="true" pageId="48" pageNumber="49">
<taxonomicName id="4C75A6E3CC014C14F4D5FAF8FBA9FA84" authorityName="Aurivillius" authorityYear="1901" box="[925,1079,1303,1329]" class="Insecta" family="Cossidae" genus="Metarbelodes" kingdom="Animalia" order="Lepidoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="umtaliana">M. umtaliana</taxonomicName>
,
</emphasis>
but curved (running parallel to the termen) in 
<taxonomicName id="4C75A6E3CC014C14F677FAD4FE5BFAE0" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[319,453,1339,1365]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="bergsteni">
<emphasis id="B9010172CC014C14F677FAD4FE5BFAE0" box="[319,453,1339,1365]" italics="true" pageId="48" pageNumber="49">L. bergsteni</emphasis>
</taxonomicName>
and (ii) the marking of vein CuA 
<subScript id="17F1DF25CC014C14F40EFAA2FCD0FAEE" attach="left" box="[838,846,1357,1371]" fontSize="6" pageId="48" pageNumber="49">2</subScript>
is broader, particularly the ivory-yellow band, in 
<taxonomicName id="4C75A6E3CC014C14F2CEFAD4FF60FACC" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="bergsteni">
<emphasis id="B9010172CC014C14F2CEFAD4FF60FACC" italics="true" pageId="48" pageNumber="49">L. bergsteni</emphasis>
</taxonomicName>
compared with 
<taxonomicName id="4C75A6E3CC014C14F6FCFAB0FDD5FACC" authorityName="Aurivillius" authorityYear="1901" box="[436,587,1375,1401]" class="Insecta" family="Cossidae" genus="Metarbelodes" kingdom="Animalia" order="Lepidoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="umtaliana">
<emphasis id="B9010172CC014C14F6FCFAB0FDD5FACC" box="[436,587,1375,1401]" italics="true" pageId="48" pageNumber="49">M. umtaliana</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BCADD60CC014C14F78FFA6CFA03F9BC" blockId="48.[151,1437,150,1941]" pageId="48" pageNumber="49">
In the female post-abdominal structure of 
<taxonomicName id="4C75A6E3CC014C14F5FAFA6CFCA4FA28" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[690,826,1411,1437]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="bergsteni">
<emphasis id="B9010172CC014C14F5FAFA6CFCA4FA28" box="[690,826,1411,1437]" italics="true" pageId="48" pageNumber="49">L. bergsteni</emphasis>
</taxonomicName>
there are two diagnostic characters: (i) the posterior apophyses have a narrow, stick-like posterior end that occupies 
<emphasis id="B9010172CC014C14F425FA48FC10FA75" box="[877,910,1447,1472]" italics="true" pageId="48" pageNumber="49">ca.</emphasis>
40% of their length and is much more sclerotized, while the remaining 60% of the posterior apophysis is broader than the width of the anterior apophyses; (ii) the band-like plate covers 60% of the ventral side of segment 8 with a slightly broader band behind the outer band.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC014C14F78FF9FDFE77F8B0" pageId="48" pageNumber="49" type="distribution">
<paragraph id="8BCADD60CC014C14F78FF9FDFACAF955" blockId="48.[151,1437,150,1941]" pageId="48" pageNumber="49">
<emphasis id="B9010172CC014C14F78FF9FDFEFFF999" bold="true" box="[199,353,1554,1580]" pageId="48" pageNumber="49">Distribution.</emphasis>
<taxonomicName id="4C75A6E3CC014C14F62FF9FCFDD2F998" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[359,588,1555,1581]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="bergsteni">
<emphasis id="B9010172CC014C14F62FF9FCFDD2F998" box="[359,588,1555,1581]" italics="true" pageId="48" pageNumber="49">Lukeniana bergsteni</emphasis>
</taxonomicName>
is known from the southwestern and southern slopes of Mount Kilimanjaro extending 
<emphasis id="B9010172CC014C14F645F9D8FEB0F9E5" box="[269,302,1591,1616]" italics="true" pageId="48" pageNumber="49">ca.</emphasis>
<quantity id="4C8D7085CC014C14F67DF9D9FEE2F9E4" box="[309,380,1590,1617]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="48" pageNumber="49" unit="km" value="50.0">50 km</quantity>
southwest to Usa River (
<collectingCountry id="F3629DF0CC014C14F5DEF9D8FC9CF9E4" box="[662,770,1591,1617]" name="Tanzania" pageId="48" pageNumber="49">Tanzania</collectingCountry>
). 
<collectingRegion id="49B11382CC014C14F45CF9D8FC72F9E4" box="[788,1004,1591,1617]" country="Tanzania" name="Kilimanjaro" pageId="48" pageNumber="49">Mount Kilimanjaro</collectingRegion>
is the highest mountain in Africa (elevation 
<quantity id="4C8D7085CC014C14F7ABF9B5FEA1F9C0" box="[227,319,1626,1653]" metricMagnitude="3" metricUnit="m" metricValue="5.895" pageId="48" pageNumber="49" unit="m" value="5895.0">5,895 m</quantity>
) and is located on the frontier with 
<collectingCountry id="F3629DF0CC014C14F58DF9B4FC90F9C0" box="[709,782,1627,1653]" name="Kenya" pageId="48" pageNumber="49">Kenya</collectingCountry>
in north-central 
<collectingCountry id="F3629DF0CC014C14F48DF9B4FBB3F9C0" box="[965,1069,1627,1653]" name="Tanzania" pageId="48" pageNumber="49">Tanzania</collectingCountry>
. The collecting locality “Makoa” is situated on the southern slopes of 
<collectingRegion id="49B11382CC014C14F57BF990FC91F92C" box="[563,783,1663,1689]" country="Tanzania" name="Kilimanjaro" pageId="48" pageNumber="49">Mount Kilimanjaro</collectingRegion>
, just south of Uswaa, and within the “cultural zone” at an elevation of between 1,110 and 
<quantity id="4C8D7085CC014C14F6B3F94CFDC4F908" box="[507,602,1699,1725]" metricMagnitude="3" metricUnit="m" metricValue="1.16" pageId="48" pageNumber="49" unit="m" value="1160.0">1,160 m</quantity>
. The locality “Usa River” is located 
<quantity id="4C8D7085CC014C14F4BCF94CFBA5F908" box="[1012,1083,1699,1725]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="48" pageNumber="49" unit="km" value="20.0">20 km</quantity>
east of 
<collectingRegion id="49B11382CC014C14F3D9F94CFB7CF908" box="[1169,1250,1699,1725]" country="Tanzania" name="Arusha" pageId="48" pageNumber="49">Arusha</collectingRegion>
(
<collectingCountry id="F3629DF0CC014C14F3B8F94CFAC6F908" box="[1264,1368,1699,1725]" name="Tanzania" pageId="48" pageNumber="49">Tanzania</collectingCountry>
) at an elevation of between 1,198 and 
<quantity id="4C8D7085CC014C14F6B5F929FDC2F954" box="[509,604,1734,1761]" metricMagnitude="3" metricUnit="m" metricValue="1.225" pageId="48" pageNumber="49" unit="m" value="1225.0">1,225 m</quantity>
. This species is classified as an Afromontane near-endemic species.
</paragraph>
<paragraph id="8BCADD60CC014C14F78FF905FE77F8B0" blockId="48.[151,1437,150,1941]" box="[199,489,1770,1797]" pageId="48" pageNumber="49">
<emphasis id="B9010172CC014C14F78FF905FEB4F8B1" bold="true" box="[199,298,1770,1796]" pageId="48" pageNumber="49">Habitat.</emphasis>
See Appendix 1.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC014C14F78FF8E0FDC8F820" pageId="48" pageNumber="49" type="etymology">
<paragraph id="8BCADD60CC014C14F78FF8E0FDC8F820" blockId="48.[151,1437,150,1941]" pageId="48" pageNumber="49">
<emphasis id="B9010172CC014C14F78FF8E0FED0F89C" bold="true" box="[199,334,1807,1833]" pageId="48" pageNumber="49">Etymology.</emphasis>
<taxonomicName id="4C75A6E3CC014C14F61CF8E0FDA6F89C" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[340,568,1807,1833]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="bergsteni">
<emphasis id="B9010172CC014C14F61CF8E0FDA6F89C" box="[340,568,1807,1833]" italics="true" pageId="48" pageNumber="49">Lukeniana bergsteni</emphasis>
</taxonomicName>
is named in honour of the senior curator at the NRM and specialist of aquatic Adephagan families (Coleoptera), Dr. Johannes Bergsten (
<collectingRegion id="49B11382CC014C14F474F8DCFC24F8F8" box="[828,954,1843,1869]" country="Sweden" name="Stockholm" pageId="48" pageNumber="49">Stockholm</collectingRegion>
, 
<collectingCountry id="F3629DF0CC014C14F481F8DCFBB8F8F8" box="[969,1062,1843,1869]" name="Sweden" pageId="48" pageNumber="49">Sweden</collectingCountry>
), whom we thank for providing samples of 
<taxonomicName id="4C75A6E3CC014C14F655F8B8FE2FF8C4" baseAuthorityName="Lehmann" baseAuthorityYear="2019" box="[285,433,1879,1905]" class="Insecta" family="Metarbelidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="family">Metarbelidae</taxonomicName>
(particularly various types) from the Swedish Museum of Natural History collections, including the 
<typeStatus id="54CE63C2CC014C14F679F894FE0DF820" box="[305,403,1915,1941]" pageId="48" pageNumber="49" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C75A6E3CC014C14F6F0F894FDCEF820" authorityName="Aurivillius" authorityYear="1901" box="[440,592,1915,1941]" class="Insecta" family="Cossidae" genus="Metarbelodes" kingdom="Animalia" order="Lepidoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="umtaliana">
<emphasis id="B9010172CC014C14F6F0F894FDCEF820" box="[440,592,1915,1941]" italics="true" pageId="48" pageNumber="49">M. umtaliana</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>