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<document id="D9E12BD5661FF622651701F943DCE914" ID-CLB-Dataset="100341" ID-DOI="10.5852/ejt.2022.831.1867" ID-GBIF-Dataset="ad320da4-c372-4775-a5b0-00c299d40c1a" ID-ISSN="2118-9773" ID-Zenodo-Dep="6862200" ID-ZooBank="752825D9-49B3-456C-8692-8DE2844593B1" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658241750918" checkinUser="felipe" docAuthor="Hassan, Muhammad Asghar, Zheng, Yuchen &amp; Liu, Xingyue" docDate="2022" docId="03D887BED71DFFC2FDE0FB95CC5A4E84" docLanguage="en" docName="ejt-831_1-44.pdf" docOrigin="European Journal of Taxonomy 831" docSource="http://dx.doi.org/10.5852/ejt.2022.831.1867" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Myrmeleon hyalinus subsp. hyalinus Olivier 1811" docType="treatment" docVersion="7" lastPageNumber="26" masterDocId="FFE1FFC6D709FFDBFFF2FFE1C9174A36" masterDocTitle="Taxonomic notes on the antlion tribe Myrmeleontini Latreille (Neuroptera, Myrmeleontidae, Myrmeleontinae) from Pakistan, with description of a new species" masterLastPageNumber="44" masterPageNumber="1" pageNumber="21" updateTime="1699360279719" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="D5CAE7C073FEE8C068DE88AE2A476B6E">Taxonomic notes on the antlion tribe Myrmeleontini Latreille (Neuroptera, Myrmeleontidae, Myrmeleontinae) from Pakistan, with description of a new species</mods:title>
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<mods:namePart id="846BB55E3505F7B5E38170D2D4845E74">Hassan, Muhammad Asghar</mods:namePart>
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<mods:affiliation id="B00FB2A1B0C739C5DEA3C0140467B10C">Department of Entomology, China Agricultural University, Beijing 100193, China.</mods:affiliation>
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<mods:namePart id="A7F539AE06738A1DDBDC0BC1BB5D6E7E">Zheng, Yuchen</mods:namePart>
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<mods:affiliation id="3565A9E3420D30FEC2AAE082342FCDB6">Department of Entomology, China Agricultural University, Beijing 100193, China.</mods:affiliation>
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<mods:namePart id="9B78ADC20888BE8C48661F733BC0CC15">Liu, Xingyue</mods:namePart>
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<mods:affiliation id="1CEB2AA3254FFBBF2B174D145F6B17A3">Department of Entomology, China Agricultural University, Beijing 100193, China.</mods:affiliation>
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<taxonomicName id="4C714D2BD71DFFCFFDE0FB95CD354EB8" ID-CoL="7HH89" ID-ENA="2744829" authority="Olivier, 1811" authorityName="Olivier" authorityYear="1811" box="[530,1058,1140,1167]" class="Insecta" family="Myrmeleontidae" genus="Myrmeleon" kingdom="Animalia" order="Neuroptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="subSpecies" species="hyalinus" subSpecies="hyalinus">
<emphasis id="B905EABAD71DFFCFFDE0FB95CA684EB9" bold="true" box="[530,895,1140,1167]" italics="true" pageId="20" pageNumber="21">Myrmeleon hyalinus hyalinus</emphasis>
<bibRefCitation id="EFE04B59D71DFFCFFC75FB95CD354EB8" author="Olivier G. A." box="[903,1058,1140,1166]" pageId="20" pageNumber="21" pagination="175 - 179" refId="ref18604" refString="Olivier G. A. 1811. Encyclopedie methodique. Histoire naturelle. Vol. 8 (Insectes). Paris: 175 - 179." type="book chapter" year="1811">Olivier, 1811</bibRefCitation>
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<figureCitation id="134A2A2DD71DFFCFFD30FB76CA5F4E84" box="[706,840,1175,1202]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="21.[188,231,1963,1989]" captionStartId-1="22.[188,231,1963,1989]" captionStartId-2="23.[189,232,1928,1954]" captionStartId-3="24.[189,232,1892,1918]" captionTargetBox-0="[205,1384,1062,1893]" captionTargetBox-1="[204,1382,873,1884]" captionTargetBox-2="[206,1382,445,1865]" captionTargetBox-3="[189,1387,692,1820]" captionTargetId-0="figure-276@21.[189,1399,1043,1908]" captionTargetId-1="figure-228@22.[189,1399,858,1901]" captionTargetId-2="figure-58@23.[189,1399,439,1880]" captionTargetId-3="figure-143@24.[189,1399,682,1833]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="23" captionTargetPageId-3="24" captionText-0="Fig. 12. Myrmeleon hyalinus hyalinus Olivier, 1811 (CAUPK000024). Male, dorsal habitus. Scale bar = 0.5 mm." captionText-1="Fig. 13. Myrmeleon hyalinus hyalinus Olivier, 1811 (CAUPK000024). Right fore- and hind wing. Scale bar = 5.0 mm" captionText-2="Fig. 14. Myrmeleon hyalinus hyalinus Olivier, 1811 (CAUPK000024). A. Dorsal habitus of head and thorax. B. Head, frontal view. C. Head and pronotum, dorsal view. D. Antenna. E. Lateral view. Scale bars: A = 2.0 mm; B–D = 1.0 mm; E = 3.0 mm." captionText-3="Fig. 15. Myrmeleon hyalinus hyalinus Olivier, 1811 (CAUPK000024). A. Male genitalia, lateral view. B. Same, ventral view. C. Complex of gonocoxites 9 + gonocoxites 11, dorsal view. D. Same, ventral view. E. Same, lateral view. Abbreviations: ect = ectoproct; gst11 = gonostylus 11; gx9 = gonocoxites 9; gx11 = gonocoxites 11; S = sternites; T = tergites. Scale bars = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6862244" figureDoi-1="http://doi.org/10.5281/zenodo.6862248" figureDoi-2="http://doi.org/10.5281/zenodo.6862250" figureDoi-3="http://doi.org/10.5281/zenodo.6862254" httpUri-0="https://zenodo.org/record/6862244/files/figure.png" httpUri-1="https://zenodo.org/record/6862248/files/figure.png" httpUri-2="https://zenodo.org/record/6862250/files/figure.png" httpUri-3="https://zenodo.org/record/6862254/files/figure.png" pageId="20" pageNumber="21">Figs 12–15</figureCitation>
, 
<figureCitation id="134A2A2DD71DFFCFFCA6FB79CA664E84" box="[852,881,1176,1202]" captionStart="Fig" captionStartId="31.[189,232,1963,1989]" captionTargetBox="[197,1387,1073,1905]" captionTargetId="figure-275@31.[189,1399,1067,1913]" captionTargetPageId="31" captionText="Fig. 20. Distribution map of species of Baliga Navás, 1912 and Myrmeleon Linnaeus, 1767 from Pakistan." figureDoi="http://doi.org/10.5281/zenodo.6862268" httpUri="https://zenodo.org/record/6862268/files/figure.png" pageId="20" pageNumber="21">20</figureCitation>
</paragraph>
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<treatmentCitation id="0AD010B9D71DFFCFFF4FFB38CBEB4EC5" author="Olivier G. A." box="[189,764,1241,1267]" page="126" pageId="20" pageNumber="21" year="1811">
<taxonomicName id="4C714D2BD71DFFCFFF4FFB38CBEB4EC5" ID-CoL="7HH89" authority="Olivier, 1811: 126" authorityName="Olivier" authorityPageNumber="126" authorityYear="1811" box="[189,764,1241,1267]" class="Insecta" family="Myrmeleontidae" genus="Myrmeleon" kingdom="Animalia" order="Neuroptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="subSpecies" species="hyalinus" subSpecies="hyalinus">
<emphasis id="B905EABAD71DFFCFFF4FFB38CB0C4EC5" box="[189,539,1241,1267]" italics="true" pageId="20" pageNumber="21">Myrmeleon hyalinus hyalinus</emphasis>
<bibRefCitation id="EFE04B59D71DFFCFFDD1FB38CBEB4EC5" author="Olivier G. A." box="[547,764,1241,1267]" pageId="20" pageNumber="21" pagination="175 - 179" refId="ref18604" refString="Olivier G. A. 1811. Encyclopedie methodique. Histoire naturelle. Vol. 8 (Insectes). Paris: 175 - 179." type="book chapter" year="1811">Olivier, 1811: 126</bibRefCitation>
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. 
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locality: 
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.
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<emphasis id="B905EABAD71DFFCFFF4FFAFBC8214F02" bold="true" box="[189,310,1306,1332]" pageId="20" pageNumber="21">Diagnosis</emphasis>
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It can be distinguished by the distinctive head and thoracic markings: frons black, except for ventral corner yellow (
<figureCitation id="134A2A2DD71DFFCFFE86FA8DC8F64FB0" box="[372,481,1388,1414]" captionStart="Fig" captionStartId="23.[189,232,1928,1954]" captionTargetBox="[206,1382,445,1865]" captionTargetId="figure-58@23.[189,1399,439,1880]" captionTargetPageId="23" captionText="Fig. 14. Myrmeleon hyalinus hyalinus Olivier, 1811 (CAUPK000024). A. Dorsal habitus of head and thorax. B. Head, frontal view. C. Head and pronotum, dorsal view. D. Antenna. E. Lateral view. Scale bars: A = 2.0 mm; B–D = 1.0 mm; E = 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6862250" httpUri="https://zenodo.org/record/6862250/files/figure.png" pageId="20" pageNumber="21">Fig. 14B</figureCitation>
); vertex black, median and posterior portions with yellow markings in dorsal view; pronotum yellow, medially with a longitudinal brownish stripe, anterior transverse furrow dark brown, distally with a pair of well-separated transverse brownish stripes (
<figureCitation id="134A2A2DD71DFFCFFBCAFA53CDB14FFB" box="[1080,1190,1458,1485]" captionStart="Fig" captionStartId="23.[189,232,1928,1954]" captionTargetBox="[206,1382,445,1865]" captionTargetId="figure-58@23.[189,1399,439,1880]" captionTargetPageId="23" captionText="Fig. 14. Myrmeleon hyalinus hyalinus Olivier, 1811 (CAUPK000024). A. Dorsal habitus of head and thorax. B. Head, frontal view. C. Head and pronotum, dorsal view. D. Antenna. E. Lateral view. Scale bars: A = 2.0 mm; B–D = 1.0 mm; E = 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6862250" httpUri="https://zenodo.org/record/6862250/files/figure.png" pageId="20" pageNumber="21">Fig. 14C</figureCitation>
); wings narrowly elongated, acutely pointed at apex, initial branching point of CuA distad Rs origin (
<figureCitation id="134A2A2DD71DFFCFFB62FA37CDF04FC6" box="[1168,1255,1494,1520]" captionStart="Fig" captionStartId="22.[188,231,1963,1989]" captionTargetBox="[204,1382,873,1884]" captionTargetId="figure-228@22.[189,1399,858,1901]" captionTargetPageId="22" captionText="Fig. 13. Myrmeleon hyalinus hyalinus Olivier, 1811 (CAUPK000024). Right fore- and hind wing. Scale bar = 5.0 mm" figureDoi="http://doi.org/10.5281/zenodo.6862248" httpUri="https://zenodo.org/record/6862248/files/figure.png" pageId="20" pageNumber="21">Fig. 13</figureCitation>
). Moreover, male genitalia is distinctive among species of 
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<emphasis id="B905EABAD71DFFCFFD3FFA18CA434C25" box="[717,852,1529,1555]" italics="true" pageId="20" pageNumber="21">Myrmeleon</emphasis>
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in 
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: gonocoxites 11 highly sclerotized, lateral arms elongated; gonostylus 11 rounded in lateral view; gonocoxites 9 narrow and elongated, wider in lateral view with pointed apex.
</paragraph>
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<emphasis id="B905EABAD71DFFCFFF4FF961C8BD4CAC" bold="true" box="[189,426,1664,1690]" pageId="20" pageNumber="21">Material examined</emphasis>
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<materialsCitation id="3B193CF5D71DFFCFFF4FF94ECDD64CDB" ID-GBIF-Occurrence="3864033301" collectingDate="2019-08-24" collectionCode="CAU" collectorName="Hassan M. A." country="Pakistan" elevation="555" latitude="33.75143" location="Shahdarah" longLatPrecision="1" longitude="73.16116" municipality="Quaid-e-Azam University" pageId="20" pageNumber="21" specimenCount="1" specimenCount-male="1" stateProvince="Islamabad Capital Territory">
<collectingCountry id="F3667638D71DFFCFFF4FF94EC85E4CFF" box="[189,329,1711,1737]" name="Pakistan" pageId="20" pageNumber="21">PAKISTAN</collectingCountry>
– 
<emphasis id="B905EABAD71DFFCFFE76F94ECA314CFF" bold="true" box="[388,806,1711,1737]" pageId="20" pageNumber="21">
<collectingRegion id="49B5F84AD71DFFCFFE76F94ECA124CFF" box="[388,773,1711,1737]" country="Pakistan" name="Islamabad Capital Territory" pageId="20" pageNumber="21">Islamabad Capital Territory</collectingRegion>
•
</emphasis>
<specimenCount id="9D77FD21D71DFFCFFCCFF94ECA6C4CFC" box="[829,891,1711,1738]" pageId="20" pageNumber="21" type="male">1 ♂</specimenCount>
; 
<collectingMunicipality id="6BAAACD2D71DFFCFFC64F94ECDCF4CFF" box="[918,1240,1711,1737]" pageId="20" pageNumber="21">Quaid-e-Azam University</collectingMunicipality>
, 
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; 
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, 
<geoCoordinate id="EE45506FD71DFFCFFE60F930CB4F4CDB" box="[402,600,1745,1773]" degrees="73" direction="east" minutes="9" orientation="longitude" pageId="20" pageNumber="21" precision="1" seconds="40.1754" value="73.16116">73°9′40.1754″ E</geoCoordinate>
; 
<quantity id="4C899B4DD71DFFCFFD96F933CBB84CDB" box="[612,687,1746,1773]" metricMagnitude="2" metricUnit="m" metricValue="5.55" pageId="20" pageNumber="21" unit="m" value="555.0">
<elevation id="005CD19BD71DFFCFFD96F933CBB84CDB" box="[612,687,1746,1773]" metricMagnitude="2" metricUnit="m" metricValue="5.55" pageId="20" pageNumber="21" unit="m" value="555.0">555 m</elevation>
</quantity>
a.sl.; 
<date id="FFCF1068D71DFFCFFD07F932CA804CDB" box="[757,919,1747,1773]" pageId="20" pageNumber="21" value="2019-08-24">
<collectingDate id="EF8BE980D71DFFCFFD07F932CA804CDB" box="[757,919,1747,1773]" pageId="20" pageNumber="21" value="2019-08-24">24 Aug. 2019</collectingDate>
</date>
; 
<collectorName id="2684537ED71DFFCFFC51F932CD2A4CDB" box="[931,1085,1747,1773]" pageId="20" pageNumber="21">Hassan M.A.</collectorName>
leg.; 
<collectionCode id="ED60AE6DD71DFFCFFB8DF932CDD64CDB" box="[1151,1217,1747,1773]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="20" pageNumber="21">CAU</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BCE36A8D71DFFCFFF4FF8F5C8624D18" blockId="20.[189,373,1812,1838]" box="[189,373,1812,1838]" pageId="20" pageNumber="21">
<heading id="D08681C4D71DFFCFFF4FF8F5C8624D18" bold="true" box="[189,373,1812,1838]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="B905EABAD71DFFCFFF4FF8F5C8624D18" bold="true" box="[189,373,1812,1838]" pageId="20" pageNumber="21">Re-description</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36B6523D71DFFC2FF4FF8A2CD734B5A" lastPageId="25" lastPageNumber="26" pageId="20" pageNumber="21" type="description">
<paragraph id="8BCE36A8D71DFFCFFF4FF8A2CB254DB6" blockId="20.[189,1399,1858,1920]" pageId="20" pageNumber="21">
MEASUREMENT (
<specimenCount id="9D77FD21D71DFFCFFE87F8A3C89B4D6A" box="[373,396,1858,1884]" pageId="20" pageNumber="21" type="male">♂</specimenCount>
n=1). Forewing: length 
<quantity id="4C899B4DD71DFFCFFD5DF8A3CA0B4D6B" box="[687,796,1858,1885]" metricMagnitude="-2" metricUnit="m" metricValue="2.62" pageId="20" pageNumber="21" unit="mm" value="26.2">26.2 mm</quantity>
, width 
<quantity id="4C899B4DD71DFFCFFC86F8A3CAC54D6B" box="[884,978,1858,1885]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="20" pageNumber="21" unit="mm" value="5.5">5.5 mm</quantity>
; hind wing: length 
<quantity id="4C899B4DD71DFFCFFB48F8A3CC304D6B" box="[1210,1319,1858,1885]" metricMagnitude="-2" metricUnit="m" metricValue="2.62" pageId="20" pageNumber="21" unit="mm" value="26.2">26.2 mm</quantity>
, width 
<quantity id="4C899B4DD71DFFCFFF4CF884C80C4DB6" box="[190,283,1893,1920]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="20" pageNumber="21" unit="mm" value="4.6">4.6 mm</quantity>
; body length: 24.0 mm.
</paragraph>
<paragraph id="8BCE36A8D71DFFCEFF4FF849CC7A4BC0" blockId="20.[188,1399,1959,2021]" lastBlockId="21.[188,1401,265,502]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
HEAD (
<figureCitation id="134A2A2DD71DFFCFFEFCF846C88C4DF7" box="[270,411,1959,1985]" captionStart="Fig" captionStartId="23.[189,232,1928,1954]" captionTargetBox="[206,1382,445,1865]" captionTargetId="figure-58@23.[189,1399,439,1880]" captionTargetPageId="23" captionText="Fig. 14. Myrmeleon hyalinus hyalinus Olivier, 1811 (CAUPK000024). A. Dorsal habitus of head and thorax. B. Head, frontal view. C. Head and pronotum, dorsal view. D. Antenna. E. Lateral view. Scale bars: A = 2.0 mm; B–D = 1.0 mm; E = 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6862250" httpUri="https://zenodo.org/record/6862250/files/figure.png" pageId="20" pageNumber="21">Fig. 14B–C</figureCitation>
). Vertex moderately raised; in frontal view black, without yellow markings; in dorsal view black, medially with a pair of transverse and posteriorly with a pair of longitudinal yellow markings; epicranial area black, with longitudinal grooves, covered with short brownish pubescence. Frons black, but ventral corner yellow, covered with short yellowish pubescence. Occiput and postorbital sclerites yellow. Clypeus yellow, medially with two indistinct dark brown markings. Labrum yellow, covered with erected brownish setae at proximal margin. Genae pale yellow. Maxillary and labial palps pale yellow, terminal labial palpomere spindle-shaped, palpimacula brownish, small and circular, with short black setae. Antennae brownish, dorsal ring of scape and pedicel yellow, covered with short black setae, flagellum brownish with proximal and distal margin dark brown. Antennal sclerites yellow (
<figureCitation id="134A2A2DD71CFFCEFB06FE3DCC484BC0" box="[1268,1375,476,502]" captionStart="Fig" captionStartId="23.[189,232,1928,1954]" captionTargetBox="[206,1382,445,1865]" captionTargetId="figure-58@23.[189,1399,439,1880]" captionTargetPageId="23" captionText="Fig. 14. Myrmeleon hyalinus hyalinus Olivier, 1811 (CAUPK000024). A. Dorsal habitus of head and thorax. B. Head, frontal view. C. Head and pronotum, dorsal view. D. Antenna. E. Lateral view. Scale bars: A = 2.0 mm; B–D = 1.0 mm; E = 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6862250" httpUri="https://zenodo.org/record/6862250/files/figure.png" pageId="21" pageNumber="22">Fig. 14B</figureCitation>
).
</paragraph>
<paragraph id="8BCE36A8D71CFFCEFF4FFDFFC82A4943" blockId="21.[187,1401,542,885]" pageId="21" pageNumber="22">
THORAX (
<figureCitation id="134A2A2DD71CFFCEFED9FDFFC88E480E" box="[299,409,542,568]" captionStart="Fig" captionStartId="23.[189,232,1928,1954]" captionTargetBox="[206,1382,445,1865]" captionTargetId="figure-58@23.[189,1399,439,1880]" captionTargetPageId="23" captionText="Fig. 14. Myrmeleon hyalinus hyalinus Olivier, 1811 (CAUPK000024). A. Dorsal habitus of head and thorax. B. Head, frontal view. C. Head and pronotum, dorsal view. D. Antenna. E. Lateral view. Scale bars: A = 2.0 mm; B–D = 1.0 mm; E = 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6862250" httpUri="https://zenodo.org/record/6862250/files/figure.png" pageId="21" pageNumber="22">Fig. 14A</figureCitation>
). Pronotum slightly wider than long, yellow, medially with a longitudinal brownish stripe, slightly interrupted at anterior transverse furrow; dark brown stripe along anterior transverse furrow, not reaching at lateral margins; distally with a pair of well-separated transverse brownish stripes; covered with sparse yellow setae. Mesonotum dark brown; prescutum laterally with a narrow yellow stripe; mesoscutum with yellow markings on median and posterolateral margins; mesoscutellum at distal ⅔ yellow; covered with sparse yellow setae, but prescutum with long dark brown setae. Metanotum dark brown; prescutum medially with faintly brownish longitudinal yellow marking; metascutum distally with a pair of large yellow markings; metascutellum laterally and distally brownish yellow; covered with sparse yellow setae. Pleuron dark brown, with yellow markings, covered with sparse yellow setae (
<figureCitation id="134A2A2DD71CFFCEFF34FCBBC8274943" box="[198,304,858,885]" captionStart="Fig" captionStartId="23.[189,232,1928,1954]" captionTargetBox="[206,1382,445,1865]" captionTargetId="figure-58@23.[189,1399,439,1880]" captionTargetPageId="23" captionText="Fig. 14. Myrmeleon hyalinus hyalinus Olivier, 1811 (CAUPK000024). A. Dorsal habitus of head and thorax. B. Head, frontal view. C. Head and pronotum, dorsal view. D. Antenna. E. Lateral view. Scale bars: A = 2.0 mm; B–D = 1.0 mm; E = 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6862250" httpUri="https://zenodo.org/record/6862250/files/figure.png" pageId="21" pageNumber="22">Fig. 14E</figureCitation>
).
</paragraph>
<paragraph id="8BCE36A8D71CFFCDFF4FFC7DCD4148B5" blockId="21.[189,1399,924,986]" lastBlockId="22.[188,1400,263,643]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
LEGS (
<figureCitation id="134A2A2DD71CFFCEFEFEFC7DC86D4980" box="[268,378,924,950]" captionStart="Fig" captionStartId="23.[189,232,1928,1954]" captionTargetBox="[206,1382,445,1865]" captionTargetId="figure-58@23.[189,1399,439,1880]" captionTargetPageId="23" captionText="Fig. 14. Myrmeleon hyalinus hyalinus Olivier, 1811 (CAUPK000024). A. Dorsal habitus of head and thorax. B. Head, frontal view. C. Head and pronotum, dorsal view. D. Antenna. E. Lateral view. Scale bars: A = 2.0 mm; B–D = 1.0 mm; E = 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6862250" httpUri="https://zenodo.org/record/6862250/files/figure.png" pageId="21" pageNumber="22">Fig. 14E</figureCitation>
). Foreleg: coxa and trochanter yellow, with short yellow setae. Femur yellow, light brownish at distal ½, with short black setae, but posterolaterally with a few long black setae at proximal half; femoral sense hair shorter than proximal ½ of profemora. Tibia yellow, with mixed, short and long black setae at proximal half, antennal cleaning setae yellow; tibial spurs brownish, straight, as long as Ta1. Tarsomeres yellow, with short black setae; Ta1 equal to combined length of Ta2–Ta3; Ta2, Ta3 and Ta4 nearly equal in size; Ta5 equal to combined length of Ta1–Ta3. Pretarsal claws brownish, curved. Mid leg: coxa yellow, slightly brownish at proximal ½, with yellow setae. Trochanter yellow, with short black setae. Femur yellow, light brownish at distal ½, covered with short black setae, laterally with a few long black setae at proximal half; femoral sense hair shorter than proximal half of mid femora. Tibia yellow, covered with mixed, short and long black setae; tibial spurs similar to foreleg. Tarsomeres and pretarsal claws similar to foreleg. Hind leg: coxa and trochanter similar to mid leg. Femur yellow, light brownish at distal ⅓, covered with short black setae, proximal half with a few long black setae; femoral sense hair absent. Tibia, tarsomeres, and pretarsal claws are similar to mid leg.
</paragraph>
<caption id="DF0E6620D71CFFCEFF4EF84AC8744DDF" ID-DOI="http://doi.org/10.5281/zenodo.6862244" ID-Zenodo-Dep="6862244" httpUri="https://zenodo.org/record/6862244/files/figure.png" pageId="21" pageNumber="22" startId="21.[188,231,1963,1989]" targetBox="[205,1384,1062,1893]" targetPageId="21">
<paragraph id="8BCE36A8D71CFFCEFF4EF84AC8744DDF" blockId="21.[188,1399,1963,2025]" pageId="21" pageNumber="22">
<emphasis id="B905EABAD71CFFCEFF4EF84AC8094DF3" bold="true" box="[188,286,1963,1989]" pageId="21" pageNumber="22">Fig. 12.</emphasis>
<taxonomicName id="4C714D2BD71CFFCEFED9F84ACA294DF3" authority="Olivier, 1811" authorityName="Olivier" authorityYear="1811" box="[299,830,1963,1989]" class="Insecta" family="Myrmeleontidae" genus="Myrmeleon" kingdom="Animalia" order="Neuroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subSpecies" species="hyalinus" subSpecies="hyalinus">
<emphasis id="B905EABAD71CFFCEFED9F84ACB854DF3" box="[299,658,1963,1989]" italics="true" pageId="21" pageNumber="22">Myrmeleon hyalinus hyalinus</emphasis>
<bibRefCitation id="EFE04B59D71CFFCEFD6DF84ACA294DF3" author="Olivier G. A." box="[671,830,1963,1989]" pageId="21" pageNumber="22" pagination="175 - 179" refId="ref18604" refString="Olivier G. A. 1811. Encyclopedie methodique. Histoire naturelle. Vol. 8 (Insectes). Paris: 175 - 179." type="book chapter" year="1811">Olivier, 1811</bibRefCitation>
</taxonomicName>
(CAUPK000024). Male, dorsal habitus. Scale bar = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BCE36A8D71FFFC3FF4FFD4ACBF24BBB" blockId="22.[189,1399,682,744]" lastBlockId="24.[189,1400,265,397]" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23">
WINGS (
<figureCitation id="134A2A2DD71FFFCDFEE9FD4BC86548F3" box="[283,370,682,709]" captionStart="Fig" captionStartId="22.[188,231,1963,1989]" captionTargetBox="[204,1382,873,1884]" captionTargetId="figure-228@22.[189,1399,858,1901]" captionTargetPageId="22" captionText="Fig. 13. Myrmeleon hyalinus hyalinus Olivier, 1811 (CAUPK000024). Right fore- and hind wing. Scale bar = 5.0 mm" figureDoi="http://doi.org/10.5281/zenodo.6862248" httpUri="https://zenodo.org/record/6862248/files/figure.png" pageId="22" pageNumber="23">Fig. 13</figureCitation>
). Forewing as long as hind wing, acute at apex; membrane hyaline; costal area slightly narrow at proximal region; venation brownish yellow, covered with sparse short black setae; poststigmal area with a few interconnected crossveins; nine presectoral crossveins; initial branching point of CuA proximal to Rs origin; Rs with 11 branches; CuP origin at the same level to basal crossveins, fused with 1A after a short free base; pterostigma indistinct; anterior Banksian line absent; posterior Banksian present. Hind wing: relatively narrow, acute at apex; membrane hyaline; venation similar to forewing; five presectoral crossveins; median fork at proximal to Rs origin; Rs with 12 branches; pterostigma indistinct; anterior Banksian line absent; posterior Banksian indistinct; pilula axillaris small, with rounded knob covered with dense brown setae.
</paragraph>
<caption id="DF0E6620D71FFFCDFF4EF84AC84D4DDF" ID-DOI="http://doi.org/10.5281/zenodo.6862248" ID-Zenodo-Dep="6862248" httpUri="https://zenodo.org/record/6862248/files/figure.png" pageId="22" pageNumber="23" startId="22.[188,231,1963,1989]" targetBox="[204,1382,873,1884]" targetPageId="22">
<paragraph id="8BCE36A8D71FFFCDFF4EF84AC84D4DDF" blockId="22.[188,1399,1963,2025]" pageId="22" pageNumber="23">
<emphasis id="B905EABAD71FFFCDFF4EF84AC80F4DF3" bold="true" box="[188,280,1963,1989]" pageId="22" pageNumber="23">Fig. 13.</emphasis>
<taxonomicName id="4C714D2BD71FFFCDFEEDF84ACA0C4DF3" authority="Olivier, 1811" authorityName="Olivier" authorityYear="1811" box="[287,795,1963,1989]" class="Insecta" family="Myrmeleontidae" genus="Myrmeleon" kingdom="Animalia" order="Neuroptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="subSpecies" species="hyalinus" subSpecies="hyalinus">
<emphasis id="B905EABAD71FFFCDFEEDF84ACB6C4DF3" box="[287,635,1963,1989]" italics="true" pageId="22" pageNumber="23">Myrmeleon hyalinus hyalinus</emphasis>
<bibRefCitation id="EFE04B59D71FFFCDFD70F84ACA0C4DF3" author="Olivier G. A." box="[642,795,1963,1989]" pageId="22" pageNumber="23" pagination="175 - 179" refId="ref18604" refString="Olivier G. A. 1811. Encyclopedie methodique. Histoire naturelle. Vol. 8 (Insectes). Paris: 175 - 179." type="book chapter" year="1811">Olivier, 1811</bibRefCitation>
</taxonomicName>
(CAUPK000024). Right fore- and hind wing. Scale bar = 5.0 mm
</paragraph>
</caption>
<caption id="DF0E6620D71EFFCCFF4FF869CBFD4DDF" ID-DOI="http://doi.org/10.5281/zenodo.6862250" ID-Zenodo-Dep="6862250" httpUri="https://zenodo.org/record/6862250/files/figure.png" pageId="23" pageNumber="24" startId="23.[189,232,1928,1954]" targetBox="[206,1382,445,1865]" targetPageId="23">
<paragraph id="8BCE36A8D71EFFCCFF4FF869CBFD4DDF" blockId="23.[189,1400,1928,2025]" pageId="23" pageNumber="24">
<emphasis id="B905EABAD71EFFCCFF4FF869C80B4D94" bold="true" box="[189,284,1928,1954]" pageId="23" pageNumber="24">Fig. 14.</emphasis>
<taxonomicName id="4C714D2BD71EFFCCFED6F869CA3D4D94" authority="Olivier, 1811" authorityName="Olivier" authorityYear="1811" box="[292,810,1928,1954]" class="Insecta" family="Myrmeleontidae" genus="Myrmeleon" kingdom="Animalia" order="Neuroptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="subSpecies" species="hyalinus" subSpecies="hyalinus">
<emphasis id="B905EABAD71EFFCCFED6F869CB924D94" box="[292,645,1928,1954]" italics="true" pageId="23" pageNumber="24">Myrmeleon hyalinus hyalinus</emphasis>
<bibRefCitation id="EFE04B59D71EFFCCFD7DF869CA3D4D94" author="Olivier G. A." box="[655,810,1928,1954]" pageId="23" pageNumber="24" pagination="175 - 179" refId="ref18604" refString="Olivier G. A. 1811. Encyclopedie methodique. Histoire naturelle. Vol. 8 (Insectes). Paris: 175 - 179." type="book chapter" year="1811">Olivier, 1811</bibRefCitation>
</taxonomicName>
(CAUPK000024). 
<emphasis id="B905EABAD71EFFCCFBE0F869CD304D94" bold="true" box="[1042,1063,1928,1954]" pageId="23" pageNumber="24">A</emphasis>
. Dorsal habitus of head and thorax. 
<emphasis id="B905EABAD71EFFCCFEE5F84AC83C4DF3" bold="true" box="[279,299,1963,1989]" pageId="23" pageNumber="24">B</emphasis>
. Head, frontal view. 
<emphasis id="B905EABAD71EFFCCFDD4F84ACB2D4DF3" bold="true" box="[550,570,1963,1989]" pageId="23" pageNumber="24">C</emphasis>
. Head and pronotum, dorsal view. 
<emphasis id="B905EABAD71EFFCCFC29F84ACAF84DF3" bold="true" box="[987,1007,1963,1989]" pageId="23" pageNumber="24">D</emphasis>
. Antenna. 
<emphasis id="B905EABAD71EFFCCFB80F84ACD914DF3" bold="true" box="[1138,1158,1963,1989]" pageId="23" pageNumber="24">E</emphasis>
. Lateral view. Scale bars: A = 2.0 mm; B–D = 1.0 mm; E = 3.0 mm.
</paragraph>
</caption>
<paragraph id="8BCE36A8D711FFC3FF4FFE55CA614823" blockId="24.[189,1398,436,533]" pageId="24" pageNumber="25">
ABDOMEN (
<figureCitation id="134A2A2DD711FFC3FEB0FE55C8894BF8" box="[322,414,436,463]" captionStart="Fig" captionStartId="21.[188,231,1963,1989]" captionTargetBox="[205,1384,1062,1893]" captionTargetId="figure-276@21.[189,1399,1043,1908]" captionTargetPageId="21" captionText="Fig. 12. Myrmeleon hyalinus hyalinus Olivier, 1811 (CAUPK000024). Male, dorsal habitus. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6862244" httpUri="https://zenodo.org/record/6862244/files/figure.png" pageId="24" pageNumber="25">Fig. 12</figureCitation>
). Tergites dark brown, distally with a narrow yellow stripe on terga 1–8, lateral margins yellow, covered with short brownish yellow setae. Sternites dark brown, distally with a narrow yellowish stripe, covered with short brownish yellow setae.
</paragraph>
<paragraph id="8BCE36A8D711FFC2FF4FFDDDCD734B5A" blockId="24.[189,1399,572,634]" lastBlockId="25.[188,1399,265,364]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
MALE GENITALIA (
<figureCitation id="134A2A2DD711FFC3FE78FDDDC8F54861" box="[394,482,572,599]" captionStart="Fig" captionStartId="24.[189,232,1892,1918]" captionTargetBox="[189,1387,692,1820]" captionTargetId="figure-143@24.[189,1399,682,1833]" captionTargetPageId="24" captionText="Fig. 15. Myrmeleon hyalinus hyalinus Olivier, 1811 (CAUPK000024). A. Male genitalia, lateral view. B. Same, ventral view. C. Complex of gonocoxites 9 + gonocoxites 11, dorsal view. D. Same, ventral view. E. Same, lateral view. Abbreviations: ect = ectoproct; gst11 = gonostylus 11; gx9 = gonocoxites 9; gx11 = gonocoxites 11; S = sternites; T = tergites. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6862254" httpUri="https://zenodo.org/record/6862254/files/figure.png" pageId="24" pageNumber="25">Fig. 15</figureCitation>
). Tergum 9 subtrapezoidal in lateral view. Sternum 9 ovate-shaped, covered with long black setae at distal ⅓. Ectoproct nearly rectangular in lateral view, posteroventral corner slightly prominent, covered with long thick black setae. Gonocoxites 11 highly sclerotized, lateral arms elongated, apex broad and curved ventrad in dorsal view. Gonostylus 11 rounded in lateral view. Gonocoxites 9 narrow and elongated, wider in lateral view with pointed apex.
</paragraph>
</subSubSection>
<caption id="DF0E6620D711FFC3FF4FF885CD144DDF" ID-DOI="http://doi.org/10.5281/zenodo.6862254" ID-Zenodo-Dep="6862254" httpUri="https://zenodo.org/record/6862254/files/figure.png" pageId="24" pageNumber="25" startId="24.[189,232,1892,1918]" targetBox="[189,1387,692,1820]" targetPageId="24">
<paragraph id="8BCE36A8D711FFC3FF4FF885CD144DDF" blockId="24.[188,1399,1892,2025]" pageId="24" pageNumber="25">
<emphasis id="B905EABAD711FFC3FF4FF885C80C4D49" bold="true" box="[189,283,1892,1919]" pageId="24" pageNumber="25">Fig. 15.</emphasis>
<taxonomicName id="4C714D2BD711FFC3FED1F884CA324D49" authority="Olivier, 1811" authorityName="Olivier" authorityYear="1811" box="[291,805,1893,1919]" class="Insecta" family="Myrmeleontidae" genus="Myrmeleon" kingdom="Animalia" order="Neuroptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="hyalinus" subSpecies="hyalinus">
<emphasis id="B905EABAD711FFC3FED1F884CB954D49" box="[291,642,1893,1919]" italics="true" pageId="24" pageNumber="25">Myrmeleon hyalinus hyalinus</emphasis>
<bibRefCitation id="EFE04B59D711FFC3FD78F884CA324D49" author="Olivier G. A." box="[650,805,1893,1919]" pageId="24" pageNumber="25" pagination="175 - 179" refId="ref18604" refString="Olivier G. A. 1811. Encyclopedie methodique. Histoire naturelle. Vol. 8 (Insectes). Paris: 175 - 179." type="book chapter" year="1811">Olivier, 1811</bibRefCitation>
</taxonomicName>
(CAUPK000024). 
<emphasis id="B905EABAD711FFC3FBF9F885CD374D48" bold="true" box="[1035,1056,1892,1918]" pageId="24" pageNumber="25">A</emphasis>
. Male genitalia, lateral view. 
<emphasis id="B905EABAD711FFC3FF4FF869C9C74D94" bold="true" box="[189,208,1928,1954]" pageId="24" pageNumber="25">B</emphasis>
. Same, ventral view. 
<emphasis id="B905EABAD711FFC3FE26F869C8FF4D94" bold="true" box="[468,488,1928,1954]" pageId="24" pageNumber="25">C</emphasis>
. Complex of gonocoxites 9 + gonocoxites 11, dorsal view. 
<emphasis id="B905EABAD711FFC3FB42F869CDD34D94" bold="true" box="[1200,1220,1928,1954]" pageId="24" pageNumber="25">D</emphasis>
. Same, ventral view. 
<emphasis id="B905EABAD711FFC3FEF0F84AC8014DF3" bold="true" box="[258,278,1963,1989]" pageId="24" pageNumber="25">E</emphasis>
. Same, lateral view. Abbreviations: ect = ectoproct; gst11 = gonostylus 11; gx9 = gonocoxites 9; gx11 = gonocoxites 11; S = sternites; T = tergites. Scale bars = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C36B6523D710FFC2FF4FFE72CBDD49CF" pageId="25" pageNumber="26" type="discussion">
<paragraph id="8BCE36A8D710FFC2FF4FFE72C9EF4B9B" blockId="25.[189,248,403,429]" box="[189,248,403,429]" pageId="25" pageNumber="26">
<heading id="D08681C4D710FFC2FF4FFE72C9EF4B9B" bold="true" box="[189,248,403,429]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="B905EABAD710FFC2FF4FFE72C9EF4B9B" bold="true" box="[189,248,403,429]" pageId="25" pageNumber="26">Note</emphasis>
</heading>
</paragraph>
<paragraph id="8BCE36A8D710FFC2FF4FFE23CBDD49CF" blockId="25.[189,1400,450,1017]" pageId="25" pageNumber="26">
<taxonomicName id="4C714D2BD710FFC2FF4FFE23C8A44BEA" authorityName="Olivier" authorityYear="1811" box="[189,435,450,476]" class="Insecta" family="Myrmeleontidae" genus="Myrmeleon" kingdom="Animalia" order="Neuroptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="hyalinus">
<emphasis id="B905EABAD710FFC2FF4FFE23C8A44BEA" box="[189,435,450,476]" italics="true" pageId="25" pageNumber="26">Myrmeleon hyalinus</emphasis>
</taxonomicName>
currently includes five subspecies: 
<emphasis id="B905EABAD710FFC2FC94FE22CAB24BEA" box="[870,933,450,476]" italics="true" pageId="25" pageNumber="26">M. h.</emphasis>
<taxonomicName id="4C714D2BD710FFC2FC5CFE23CDAC4BEB" authority="Olivier, 1811" authorityName="Olivier" authorityYear="1811" box="[942,1211,450,477]" class="Insecta" family="Myrmeleontidae" genus="Myrmeleon" kingdom="Animalia" order="Neuroptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="hyalinus">
<emphasis id="B905EABAD710FFC2FC5CFE23CD044BEA" box="[942,1043,450,476]" italics="true" pageId="25" pageNumber="26">hyalinus</emphasis>
<bibRefCitation id="EFE04B59D710FFC2FBEEFE23CDAC4BEB" author="Olivier G. A." box="[1052,1211,450,477]" pageId="25" pageNumber="26" pagination="175 - 179" refId="ref18604" refString="Olivier G. A. 1811. Encyclopedie methodique. Histoire naturelle. Vol. 8 (Insectes). Paris: 175 - 179." type="book chapter" year="1811">Olivier, 1811</bibRefCitation>
</taxonomicName>
(widespread in Northern Africa and the Middle East, Atlantic islands, 
<collectingCountry id="F3667638D710FFC2FCAEFE07CA8A4836" box="[860,925,486,512]" name="India" pageId="25" pageNumber="26">India</collectingCountry>
, and 
<collectingCountry id="F3667638D710FFC2FC2DFE07CD5D4836" box="[991,1098,486,512]" name="Pakistan" pageId="25" pageNumber="26">Pakistan</collectingCountry>
), 
<emphasis id="B905EABAD710FFC2FB92FE06CC0F4836" box="[1120,1304,486,512]" italics="true" pageId="25" pageNumber="26">M. h. afghanus</emphasis>
<bibRefCitation id="EFE04B59D710FFC2FAEDFE07C9EC4813" author="Holzel H." pageId="25" pageNumber="26" pagination="78 - 88" refId="ref16717" refString="Holzel H. 1987. Myrmeleon hyalinus Olivier - eine chorologisch-taxonomische Analyse (Neuropteroidea: Planipennia: Myrmeleonidae). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen 38: 78 - 88." type="journal article" year="1987">Hölzel, 1987</bibRefCitation>
(
<collectingCountry id="F3667638D710FFC2FEFFFDEBC8B14812" box="[269,422,522,548]" name="Afghanistan" pageId="25" pageNumber="26">Afghanistan</collectingCountry>
), 
<emphasis id="B905EABAD710FFC2FE49FDEACBB24812" box="[443,677,522,548]" italics="true" pageId="25" pageNumber="26">M. h. caboverdicus</emphasis>
<bibRefCitation id="EFE04B59D710FFC2FD5DFDEBCA5A4813" author="Holzel H." box="[687,845,522,549]" pageId="25" pageNumber="26" pagination="78 - 88" refId="ref16717" refString="Holzel H. 1987. Myrmeleon hyalinus Olivier - eine chorologisch-taxonomische Analyse (Neuropteroidea: Planipennia: Myrmeleonidae). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen 38: 78 - 88." type="journal article" year="1987">Hölzel, 1987</bibRefCitation>
(
<collectingCountry id="F3667638D710FFC2FCADFDEACAF94812" box="[863,1006,522,549]" name="Cape Verde" pageId="25" pageNumber="26">Cape Verde</collectingCountry>
Islands), 
<emphasis id="B905EABAD710FFC2FB9BFDEACC0D4812" box="[1129,1306,522,548]" italics="true" pageId="25" pageNumber="26">M. h. cabrerai</emphasis>
Navás, 1912 (Canary Islands), and 
<emphasis id="B905EABAD710FFC2FDE7FDCECA10487E" box="[533,775,558,584]" italics="true" pageId="25" pageNumber="26">M. h. distinguendus</emphasis>
<bibRefCitation id="EFE04B59D710FFC2FCE0FDCFCAA8487F" author="Rambur J. P." box="[786,959,558,585]" pageId="25" pageNumber="26" refId="ref18757" refString="Rambur J. P. 1842. Histoire naturelle des Insectes. Nevropteres. Librairie encyclopedique de Roret. Fain et Thunot, Paris." type="book" year="1842">Rambur, 1842</bibRefCitation>
(widespread in Southern Europe to the Middle East), which can be distinguished in adult morphology and geographical distribution (
<bibRefCitation id="EFE04B59D710FFC2FF35FD96C84948A7" author="Holzel H." box="[199,350,631,657]" pageId="25" pageNumber="26" pagination="78 - 88" refId="ref16717" refString="Holzel H. 1987. Myrmeleon hyalinus Olivier - eine chorologisch-taxonomische Analyse (Neuropteroidea: Planipennia: Myrmeleonidae). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen 38: 78 - 88." type="journal article" year="1987">Hölzel 1987</bibRefCitation>
). The morphological characters to delimit the subspecies of 
<taxonomicName id="4C714D2BD710FFC2FBA5FD96CDFF48A7" authorityName="Olivier" authorityYear="1811" box="[1111,1256,631,657]" class="Insecta" family="Myrmeleontidae" genus="Myrmeleon" kingdom="Animalia" order="Neuroptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="hyalinus">
<emphasis id="B905EABAD710FFC2FBA5FD96CDFF48A7" box="[1111,1256,631,657]" italics="true" pageId="25" pageNumber="26">M. hyalinus</emphasis>
</taxonomicName>
are largely unreliable, particularly the frontal and prothoracic marking patterns (
<figureCitation id="134A2A2DD710FFC2FC01FD7ACD934883" box="[1011,1156,667,693]" captionStart="Fig" captionStartId="23.[189,232,1928,1954]" captionTargetBox="[206,1382,445,1865]" captionTargetId="figure-58@23.[189,1399,439,1880]" captionTargetPageId="23" captionText="Fig. 14. Myrmeleon hyalinus hyalinus Olivier, 1811 (CAUPK000024). A. Dorsal habitus of head and thorax. B. Head, frontal view. C. Head and pronotum, dorsal view. D. Antenna. E. Lateral view. Scale bars: A = 2.0 mm; B–D = 1.0 mm; E = 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6862250" httpUri="https://zenodo.org/record/6862250/files/figure.png" pageId="25" pageNumber="26">Fig. 14A–C</figureCitation>
; 
<bibRefCitation id="EFE04B59D710FFC2FB7DFD7ACC354883" author="Holzel H." box="[1167,1314,667,693]" pageId="25" pageNumber="26" pagination="78 - 88" refId="ref16717" refString="Holzel H. 1987. Myrmeleon hyalinus Olivier - eine chorologisch-taxonomische Analyse (Neuropteroidea: Planipennia: Myrmeleonidae). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen 38: 78 - 88." type="journal article" year="1987">Hölzel 1987</bibRefCitation>
: fig. 2; 
<bibRefCitation id="EFE04B59D710FFC2FF4FFD5EC88C48EF" author="Akhtar S. &amp; Ashfaq M. &amp; Zia A. &amp; Ali S. &amp; Ali G. M. &amp; Farhatullah &amp; Zafar Y." box="[189,411,703,729]" pageId="25" pageNumber="26" pagination="180 - 190" refId="ref15228" refString="Akhtar S., Ashfaq M., Zia A., Ali S., Ali G. M., Farhatullah &amp; Zafar Y. 2018. First report and redescription of five species of genus Myrmeleon (Neuroptera: Myrmeleontidae) from Pakistan. Journal of Biodiversity and Environmental Sciences 13: 180 - 190." type="journal article" year="2018">
Akhtar 
<emphasis id="B905EABAD710FFC2FEEBFD5EC84148EF" box="[281,342,703,729]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
2018
</bibRefCitation>
: fig. 3a; 
<bibRefCitation id="EFE04B59D710FFC2FDFBFD5ECA5F48EF" author="Hajiesmaeilian A. &amp; Shoushtari R. V. &amp; Mozaffarian F. &amp; Ebrahimi E." box="[521,840,703,729]" pageId="25" pageNumber="26" pagination="153 - 160" refId="ref16303" refString="Hajiesmaeilian A., Shoushtari R. V., Mozaffarian F. &amp; Ebrahimi E. 2020. Tribe Myrmeleontini (Neuroptera: Planipennia: Myrmeleontidae) in Iran. Zootaxa 4751: 153 - 160. https: // doi. org / 10.11646 / zootaxa. 4751.1.9" type="journal article" year="2020">
Hajiesmaeilian 
<emphasis id="B905EABAD710FFC2FD34FD5ECA1448EF" box="[710,771,703,729]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
2020
</bibRefCitation>
: fig. 11). For example, the marking pattern of frons in our examined specimen is similar to 
<emphasis id="B905EABAD710FFC2FD06FD02CAB148CB" box="[756,934,739,765]" italics="true" pageId="25" pageNumber="26">M. h. cabrerai</emphasis>
with lower margin of frons narrowly yellow (
<figureCitation id="134A2A2DD710FFC2FED2FCE6C8984917" box="[288,399,775,801]" captionStart="Fig" captionStartId="22.[188,231,1963,1989]" captionTargetBox="[204,1382,873,1884]" captionTargetId="figure-228@22.[189,1399,858,1901]" captionTargetPageId="22" captionText="Fig. 13. Myrmeleon hyalinus hyalinus Olivier, 1811 (CAUPK000024). Right fore- and hind wing. Scale bar = 5.0 mm" figureDoi="http://doi.org/10.5281/zenodo.6862248" httpUri="https://zenodo.org/record/6862248/files/figure.png" pageId="25" pageNumber="26">Fig. 13B</figureCitation>
; 
<bibRefCitation id="EFE04B59D710FFC2FE6DFCE6CB254917" author="Holzel H." box="[415,562,775,801]" pageId="25" pageNumber="26" pagination="78 - 88" refId="ref16717" refString="Holzel H. 1987. Myrmeleon hyalinus Olivier - eine chorologisch-taxonomische Analyse (Neuropteroidea: Planipennia: Myrmeleonidae). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen 38: 78 - 88." type="journal article" year="1987">Hölzel 1987</bibRefCitation>
: fig. 10), but different from 
<emphasis id="B905EABAD710FFC2FC7DFCE6CD2A4917" box="[911,1085,775,801]" italics="true" pageId="25" pageNumber="26">
M. h. 
<taxonomicName id="4C714D2BD710FFC2FC2AFCE6CD2A4917" authorityName="Olivier" authorityYear="1811" box="[984,1085,775,801]" class="Insecta" family="Myrmeleontidae" genus="Myrmeleon" kingdom="Animalia" order="Neuroptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="hyalinus">hyalinus</taxonomicName>
</emphasis>
that is distinguished by a median yellow marking on frons at lower margin (
<figureCitation id="134A2A2DD710FFC2FCD1FCCACA864973" box="[803,913,811,837]" captionStart="Fig" captionStartId="23.[189,232,1928,1954]" captionTargetBox="[206,1382,445,1865]" captionTargetId="figure-58@23.[189,1399,439,1880]" captionTargetPageId="23" captionText="Fig. 14. Myrmeleon hyalinus hyalinus Olivier, 1811 (CAUPK000024). A. Dorsal habitus of head and thorax. B. Head, frontal view. C. Head and pronotum, dorsal view. D. Antenna. E. Lateral view. Scale bars: A = 2.0 mm; B–D = 1.0 mm; E = 3.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6862250" httpUri="https://zenodo.org/record/6862250/files/figure.png" pageId="25" pageNumber="26">Fig. 14B</figureCitation>
; 
<bibRefCitation id="EFE04B59D710FFC2FC6DFCCACD254973" author="Holzel H." box="[927,1074,811,837]" pageId="25" pageNumber="26" pagination="78 - 88" refId="ref16717" refString="Holzel H. 1987. Myrmeleon hyalinus Olivier - eine chorologisch-taxonomische Analyse (Neuropteroidea: Planipennia: Myrmeleonidae). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen 38: 78 - 88." type="journal article" year="1987">Hölzel 1987</bibRefCitation>
: fig. 8; 
<bibRefCitation id="EFE04B59D710FFC2FB63FCCACC794973" author="Akhtar S. &amp; Ashfaq M. &amp; Zia A. &amp; Ali S. &amp; Ali G. M. &amp; Farhatullah &amp; Zafar Y." box="[1169,1390,811,837]" pageId="25" pageNumber="26" pagination="180 - 190" refId="ref15228" refString="Akhtar S., Ashfaq M., Zia A., Ali S., Ali G. M., Farhatullah &amp; Zafar Y. 2018. First report and redescription of five species of genus Myrmeleon (Neuroptera: Myrmeleontidae) from Pakistan. Journal of Biodiversity and Environmental Sciences 13: 180 - 190." type="journal article" year="2018">
Akhtar 
<emphasis id="B905EABAD710FFC2FB1FFCCACC3D4973" box="[1261,1322,811,837]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
2018
</bibRefCitation>
: fig. 7a; 
<bibRefCitation id="EFE04B59D710FFC2FEECFCAECB80495F" author="Hajiesmaeilian A. &amp; Shoushtari R. V. &amp; Mozaffarian F. &amp; Ebrahimi E." box="[286,663,847,873]" pageId="25" pageNumber="26" pagination="153 - 160" refId="ref16303" refString="Hajiesmaeilian A., Shoushtari R. V., Mozaffarian F. &amp; Ebrahimi E. 2020. Tribe Myrmeleontini (Neuroptera: Planipennia: Myrmeleontidae) in Iran. Zootaxa 4751: 153 - 160. https: // doi. org / 10.11646 / zootaxa. 4751.1.9" type="journal article" year="2020">
Hajiesmaeilian 
<emphasis id="B905EABAD710FFC2FE2EFCAECB0C495F" box="[476,539,847,873]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
2020: 20
</bibRefCitation>
). However, the intraspecific divergence herein observed for 
<emphasis id="B905EABAD710FFC2FF4FFC92C87C49BB" box="[189,363,883,909]" italics="true" pageId="25" pageNumber="26">
M. h. 
<taxonomicName id="4C714D2BD710FFC2FEF4FC92C87C49BB" authorityName="Olivier" authorityYear="1811" box="[262,363,883,909]" class="Insecta" family="Myrmeleontidae" genus="Myrmeleon" kingdom="Animalia" order="Neuroptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="hyalinus">hyalinus</taxonomicName>
</emphasis>
and 
<emphasis id="B905EABAD710FFC2FE58FC92CB8E49BB" box="[426,665,883,909]" italics="true" pageId="25" pageNumber="26">M. h. distinguendus</emphasis>
was 
<geoCoordinate id="EE45506FD710FFC2FD28FC92CA0B49BB" box="[730,796,883,909]" degrees="0.021" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="55" value="0.021">0.021</geoCoordinate>
<geoCoordinate id="EE45506FD710FFC2FCEEFC92CA6749BB" box="[796,880,883,909]" degrees="0.049" direction="west" orientation="longitude" pageId="25" pageNumber="26" precision="55" value="-0.049">–0.049</geoCoordinate>
based on COI gene data. Furthermore, the monophyly of 
<emphasis id="B905EABAD710FFC2FE86FC76CB324987" box="[372,549,919,945]" italics="true" pageId="25" pageNumber="26">
M. h. 
<taxonomicName id="4C714D2BD710FFC2FE32FC76CB324987" authorityName="Olivier" authorityYear="1811" box="[448,549,919,945]" class="Insecta" family="Myrmeleontidae" genus="Myrmeleon" kingdom="Animalia" order="Neuroptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="hyalinus">hyalinus</taxonomicName>
</emphasis>
was not recovered based on our analysis (
<figureCitation id="134A2A2DD710FFC2FBC9FC76CD8F4987" box="[1083,1176,919,945]" captionStart="Fig" captionStartId="32.[189,232,1892,1918]" captionTargetBox="[190,1393,648,1838]" captionTargetId="figure-107@32.[189,1399,634,1846]" captionTargetPageId="32" captionText="Fig. 21. Phylogenetic relationships among the species of Baliga Navás, 1912 and Myrmeleon Linnaeus, 1767 from Pakistan, China, Egypt, Greece, and Japan, based on COI (A) and 16S rRNA (B) genes. The numbers at each node show the posterior probabilities and bootstrap values more than 50% in BI/ML/ NJ analysis." figureDoi="http://doi.org/10.5281/zenodo.6862270" httpUri="https://zenodo.org/record/6862270/files/figure.png" pageId="25" pageNumber="26">Fig. 21</figureCitation>
). In conclusion, a combined morphological and molecular data should be applied for all subspecies to resolve the status of these geographically isolated subspecies.
</paragraph>
</subSubSection>
<subSubSection id="C36B6523D710FFC2FF4FFBC0CC5A4E84" pageId="25" pageNumber="26" type="distribution">
<paragraph id="8BCE36A8D710FFC2FF4FFBC0C8414E0D" blockId="25.[189,342,1057,1083]" box="[189,342,1057,1083]" pageId="25" pageNumber="26">
<heading id="D08681C4D710FFC2FF4FFBC0C8414E0D" bold="true" box="[189,342,1057,1083]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="B905EABAD710FFC2FF4FFBC0C8414E0D" bold="true" box="[189,342,1057,1083]" pageId="25" pageNumber="26">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BCE36A8D710FFC2FF4FFBB1CC5A4E84" blockId="25.[189,1398,1104,1205]" pageId="25" pageNumber="26">
<collectingCountry id="F3667638D710FFC2FF4FFBB1C83F4E5C" box="[189,296,1104,1130]" name="Pakistan" pageId="25" pageNumber="26">Pakistan</collectingCountry>
: (
<collectingRegion id="49B5F84AD710FFC2FEC9FBB1CB1E4E5C" box="[315,521,1104,1130]" country="India" name="Punjab" pageId="25" pageNumber="26">Punjab Province</collectingRegion>
: District Bahawalpur, Lal Sohanra National Park); widespread: Southern Europe, Northern Africa, Middle East, East to Western 
<collectingCountry id="F3667638D710FFC2FC92FB95CA894EB8" box="[864,926,1140,1166]" name="India" pageId="25" pageNumber="26">India</collectingCountry>
and Macaronesia (
<bibRefCitation id="EFE04B59D710FFC2FB75FB95CC794EB8" author="Aspock H. &amp; Holzel H. &amp; Aspock U." box="[1159,1390,1140,1166]" pageId="25" pageNumber="26" pagination="1 - 606" refId="ref15384" refString="Aspock H., Holzel H. &amp; Aspock U. 2001. Kommentierter Katalog der Neuropterida (Insecta: Raphidioptera, Megaloptera, Neuroptera) der Westpalaarktis. Denisia 2: 1 - 606." type="journal article" year="2001">
Aspöck 
<emphasis id="B905EABAD710FFC2FB1EFB94CC3E4EB8" box="[1260,1321,1140,1166]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
2001
</bibRefCitation>
; 
<bibRefCitation id="EFE04B59D710FFC2FF4FFB79C8444E84" author="Stange L. A." box="[189,339,1176,1202]" pageId="25" pageNumber="26" pagination="1 - 565" refId="ref19159" refString="Stange L. A. 2004. A systematic catalog, bibliography and classification of the world antlions (Insecta: Neuroptera: Myrmeleontidae). Memoirs of the American Entomological Institute 74: 1 - 565." type="journal article" year="2004">Stange 2004</bibRefCitation>
; 
<bibRefCitation id="EFE04B59D710FFC2FE96FB78CB0F4E84" author="Abraham L." box="[356,536,1176,1205]" pageId="25" pageNumber="26" pagination="177 - 191" refId="ref15039" refString="Abraham L. 2010. Short report on the fauna of ant-lion and owl-fly (Neuroptera) from Socotra Archipelago. Natura Somogyiensis 17: 177 - 191." type="journal article" year="2010">Ábrahám 2010</bibRefCitation>
, 2011, 2017; 
<bibRefCitation id="EFE04B59D710FFC2FD4CFB79CA8D4E84" author="Akhtar S. &amp; Ashfaq M. &amp; Zia A. &amp; Ali S. &amp; Ali G. M. &amp; Farhatullah &amp; Zafar Y." box="[702,922,1176,1202]" pageId="25" pageNumber="26" pagination="180 - 190" refId="ref15228" refString="Akhtar S., Ashfaq M., Zia A., Ali S., Ali G. M., Farhatullah &amp; Zafar Y. 2018. First report and redescription of five species of genus Myrmeleon (Neuroptera: Myrmeleontidae) from Pakistan. Journal of Biodiversity and Environmental Sciences 13: 180 - 190." type="journal article" year="2018">
Akhtar 
<emphasis id="B905EABAD710FFC2FCEBFB78CA414E84" box="[793,854,1176,1202]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
2018
</bibRefCitation>
; 
<bibRefCitation id="EFE04B59D710FFC2FC59FB78CD9D4E84" author="Hassan M. A. &amp; Oswald J. D. &amp; Zia A. &amp; Liu X. Y." box="[939,1162,1176,1203]" pageId="25" pageNumber="26" pagination="497 - 541" refId="ref16399" refString="Hassan M. A., Oswald J. D., Zia A. &amp; Liu X. Y. 2019. Neuropterida (Insecta: Megaloptera, Raphidioptera, Neuroptera) of Pakistan: a catalogue and faunistic review. Zootaxa 4686: 497 - 541. https: // doi. org / 10.11646 / zootaxa. 4686.4.3" type="journal article" year="2019">
Hassan 
<emphasis id="B905EABAD710FFC2FBF8FB78CD514E84" box="[1034,1094,1176,1202]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
2019
</bibRefCitation>
; 
<bibRefCitation id="EFE04B59D710FFC2FB69FB79CC2C4E84" author="Oswald J. D." box="[1179,1339,1176,1202]" pageId="25" pageNumber="26" pagination="1 - 94" refId="ref18630" refString="Oswald J. D. 2020. Lacewing Digital Library. Available from: https: // lacewing. tamu. edu / SpeciesCatalog / Main (accessed 13 Jul. 2020) Oswald J. D. &amp; Penny N. D. 1991. Genus-group names of the Neuroptera, Megaloptera and Raphidioptera of the world. Occasional Papers of the California Academy of Sciences 147: 1 - 94. https: // doi. org / 10.5962 / bhl. part. 3428" type="journal article" year="2020">Oswald 2020</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>