treatments-xml/data/03/D7/87/03D787C9FFA4FF95FF1BC825C92EE518.xml

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<mods:title id="4077F73470B86B55EE6097D1BD5AD819">Revision of some types of Philonthus Stephens and Gabrius Stephens from southern South America (Coleoptera: Staphylinidae), with nomenclatural notes</mods:title>
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<subSubSection id="C3646554FFA4FF94FF1BC825CB44E4A1" pageId="8" pageNumber="39" type="nomenclature">
<paragraph id="8BC136DFFFA4FF94FF1BC825C8E5E45C" blockId="8.[151,653,1365,1426]" box="[151,653,1365,1391]" pageId="8" pageNumber="39">
<heading id="D08981B3FFA4FF94FF1BC825C8E5E45C" bold="true" box="[151,653,1365,1391]" fontSize="11" level="1" pageId="8" pageNumber="39" reason="1">
<taxonomicName id="4C7E4D5CFFA4FF94FF1BC825C8E5E45C" ID-CoL="779LY" authority="Bernhauer, 1912" authorityName="Bernhauer" authorityYear="1912" box="[151,653,1365,1391]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="cribriventris">
<emphasis id="B90AEACDFFA4FF94FF1BC825C8E5E45C" bold="true" box="[151,653,1365,1391]" pageId="8" pageNumber="39">
<emphasis id="B90AEACDFFA4FF94FF1BC825CBD0E45C" bold="true" box="[151,440,1365,1391]" italics="true" pageId="8" pageNumber="39">Philonthus cribriventris</emphasis>
<bibRefCitation id="EFEF4B2EFFA4FF94FE33C825C8E5E45C" author="Bernhauer" box="[447,653,1365,1391]" pageId="8" pageNumber="41" refString="Bernhauer, M. (1912) Zur Staphylinidenfauna von Sudamerika (10. Beitrag). Verhandlungen der k. k. zoologisch- Botanischen Gesellschaft in Wien, 62, 26 - 48." type="journal article" year="1912">Bernhauer, 1912</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BC136DFFFA4FF94FF1BC808CB44E4A1" blockId="8.[151,653,1365,1426]" box="[151,300,1400,1426]" pageId="8" pageNumber="39">
(
<figureCitation id="13452A5AFFA4FF94FF13C808CB4BE4A1" box="[159,291,1400,1426]" captionStart="FIGURES 1 – 12" captionStartId="6.[151,275,1634,1658]" captionText="FIGURES 1 – 12. Type specimens: habitus and labels; labels with lectotype designation are not included. (1 a – b) Philonthus argentinus Bernhauer (now in Gabrius), (lectotype with labels *). (2 a – b) Philonthus jujuyensis Bernhauer (now in Gabrius), (lectotype with labels *). (3 a – b) Philonthus tucumanensis Bernhauer (now in Gabrius), (lectotype with labels *). (4) Philonthus nidicola Bernhauer (now in Gabrius), (only labels are shown *). (5 a – b) Philonthus quadraticeps (lectotype with labels *). (6 a – b) P. tenebrosus (= P. quadraticeps), (lectotype with labels *). (7 a – b) Philonthus bonariensis (lectotype with labels *). (8 a – b) Philonthus weiseri (= P. bonariensis), (lectotype with labels *). (9 a – b) Philonthus catamarcanus (= P. bonariensis), (lectotype with labels *). (10 a – b) Philonthus cribriventris (lectotype with labels *). (11) Philonthus catamarcanus var. densior (= P. cribriventris), (lectotype, labels are not shown). (12) G. chiliensis (= G. nigritulus), (paratype, labels are not shown). * Labels with lectotype designation are not included (pictures were taken prior to any designation)." pageId="8" pageNumber="39">Figs 10, 11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3646554FFA4FF94FF1BC8B5CEA2E72F" pageId="8" pageNumber="39" type="reference_group">
<paragraph id="8BC136DFFFA4FF94FF1BC8B5CEF2E4CF" blockId="8.[151,1436,1477,1564]" pageId="8" pageNumber="39">
<treatmentCitationGroup id="AB6E11F1FFA4FF94FF1BC8B5CEF2E4CF" pageId="8" pageNumber="39">
<taxonomicName id="4C7E4D5CFFA4FF94FF1BC8B5C817E4EF" ID-CoL="779LY" authority="Bernhauer, 1912: 43" authorityName="Bernhauer" authorityPageNumber="43" authorityYear="1912" box="[151,639,1477,1500]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="cribriventris">
<emphasis id="B90AEACDFFA4FF94FF1BC8B5CBF0E4EF" box="[151,408,1477,1500]" italics="true" pageId="8" pageNumber="39">Philonthus cribriventris</emphasis>
<treatmentCitation id="0ADF10CEFFA4FF94FE2CC8B5C817E4EF" author="Bernhauer" box="[416,639,1477,1500]" page="43" pageId="8" pageNumber="39" year="1912">
<bibRefCitation id="EFEF4B2EFFA4FF94FE2CC8B5C83EE4EF" author="Bernhauer" box="[416,598,1477,1500]" pageId="8" pageNumber="41" refString="Bernhauer, M. (1912) Zur Staphylinidenfauna von Sudamerika (10. Beitrag). Verhandlungen der k. k. zoologisch- Botanischen Gesellschaft in Wien, 62, 26 - 48." type="journal article" year="1912">Bernhauer, 1912</bibRefCitation>
: 43
</treatmentCitation>
</taxonomicName>
; 
<treatmentCitation id="0ADF10CEFFA4FF94FD00C8B5C992E4EF" author="Bernhauer" box="[652,1018,1477,1500]" page="334" pageId="8" pageNumber="39" year="1914">
<bibRefCitation id="EFEF4B2EFFA4FF94FD00C8B5C9AEE4EF" author="Bernhauer" box="[652,966,1477,1500]" pageId="8" pageNumber="41" refString="Bernhauer, M. &amp; Schubert, K. (1914) Staphylinidae IV. In: Schenkling, S. (Ed.), Coleopterorum Catalogus, 5 (57). W. Junk, Berlin, pp. 289 - 408." type="journal article" year="1914">Bernhauer &amp; Schubert, 1914</bibRefCitation>
: 334
</treatmentCitation>
(catalog); 
<treatmentCitation id="0ADF10CEFFA4FF94FBFDC8B5CF45E4EF" author="Bruch" box="[1137,1325,1477,1500]" page="444" pageId="8" pageNumber="39" year="1915">
<bibRefCitation id="EFEF4B2EFFA4FF94FBFDC8B5CE92E4EF" author="Bruch" box="[1137,1274,1477,1500]" pageId="8" pageNumber="41" refString="Bruch, C. (1915) Catalogo Sistematico de los coleopteros de la Republica Argentina. Pars VI. Revista del Museo de La Plata, 19 (Segunda Serie) (6), 235 - 302." type="journal article" year="1915">Bruch, 1915</bibRefCitation>
: 444
</treatmentCitation>
(catalog); 
<treatmentCitation id="0ADF10CEFFA4FF94FF4BC895CBADE4CF" author="Blackwelder" box="[199,453,1509,1532]" page="132" pageId="8" pageNumber="39" year="1944">
<bibRefCitation id="EFEF4B2EFFA4FF94FF4BC895CBFBE4CF" author="Blackwelder" box="[199,403,1509,1532]" pageId="8" pageNumber="41" refString="Blackwelder, R. E. (1944) Checklist of the Coleopterous insects of Mexico, Central America, the West Indies, and South America. Part I. United States National Museum Bulletin, 185, 1 - 188." type="journal article" year="1944">Blackwelder, 1944</bibRefCitation>
: 132
</treatmentCitation>
(list); 
<treatmentCitation id="0ADF10CEFFA4FF94FD80C895C880E4CF" author="Herman" box="[524,744,1509,1532]" page="2793" pageId="8" pageNumber="39" year="2001">
<bibRefCitation id="EFEF4B2EFFA4FF94FD80C895C8C0E4CF" author="Herman" box="[524,680,1509,1532]" pageId="8" pageNumber="42" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Part V. Bulletin of the American Museum of Natural History, 265, 2441 - 3020." type="journal article" year="2001">Herman, 2001</bibRefCitation>
: 2793
</treatmentCitation>
(catalog); Chani-Posse, 2004: 230 (list).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC136DFFFA4FF94FF1BCB75CEA2E72F" blockId="8.[151,1436,1477,1564]" box="[151,1226,1540,1564]" pageId="8" pageNumber="39">
<treatmentCitationGroup id="AB6E11F1FFA4FF94FF1BCB75CEA2E72F" box="[151,1226,1540,1564]" pageId="8" pageNumber="39">
<taxonomicName id="4C7E4D5CFFA4FF94FF1BCB75CBCFE72F" ID-CoL="76XKJ" box="[151,423,1541,1564]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="catamarcanus">
<emphasis id="B90AEACDFFA4FF94FF1BCB75CBCFE72F" box="[151,423,1541,1564]" italics="true" pageId="8" pageNumber="39">Philonthus catamarcanus</emphasis>
</taxonomicName>
<taxonomicName id="4C7E4D5CFFA4FF94FE22CB75C966E72F" authority="Bernhauer, 1916: 36" authorityName="Bernhauer" authorityPageNumber="36" authorityYear="1916" box="[430,782,1541,1564]" pageId="8" pageNumber="39" rank="variety" variety="densior">
var. 
<emphasis id="B90AEACDFFA4FF94FE50CB75C844E72F" box="[476,556,1541,1564]" italics="true" pageId="8" pageNumber="39">densior</emphasis>
<treatmentCitation id="0ADF10CEFFA4FF94FDBECB75C966E72F" author="Bernhauer" box="[562,782,1541,1564]" page="36" pageId="8" pageNumber="39" year="1916">
<bibRefCitation id="EFEF4B2EFFA4FF94FDBECB75C88EE72F" author="Bernhauer" box="[562,742,1541,1564]" pageId="8" pageNumber="41" refString="Bernhauer, M. (1916) Zur Staphylinidenfauna des tropischen Amerika, insbesondere der columbischen Cordilleren. Koleopterologische Rundschau, 5, 25 - 40." type="journal article" year="1916">Bernhauer, 1916</bibRefCitation>
: 36
</treatmentCitation>
</taxonomicName>
; 
<treatmentCitation id="0ADF10CEFFA4FF94FC95CB75CE74E72F" author="Scheerpeltz" box="[793,1052,1541,1564]" page="1335" pageId="8" pageNumber="39" year="1933">
<bibRefCitation id="EFEF4B2EFFA4FF94FC95CB75C9B4E72F" author="Scheerpeltz" box="[793,988,1541,1564]" pageId="8" pageNumber="42" refString="Scheerpeltz, O. (1933) Staphylinidae VII. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Pars 129. W. Junk, Berlin, pp. 989 - 1500." type="journal article" year="1933">Scheerpeltz, 1933</bibRefCitation>
: 1335
</treatmentCitation>
(cat.), 
<emphasis id="B90AEACDFFA4FF94FBEBCB74CEA2E72F" bold="true" box="[1127,1226,1540,1564]" pageId="8" pageNumber="39">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3646554FFA4FF94FF1BCB3BC8FBE6DB" pageId="8" pageNumber="39" type="materials_examined">
<paragraph id="8BC136DFFFA4FF94FF1BCB3BC918E614" blockId="8.[151,1437,1611,2024]" pageId="8" pageNumber="39">
<emphasis id="B90AEACDFFA4FF94FF1BCB3BCBBCE756" bold="true" box="[151,468,1611,1637]" pageId="8" pageNumber="39">
<typeStatus id="54C5887DFFA4FF94FF1BCB3BCABDE756" box="[151,213,1611,1637]" pageId="8" pageNumber="39">Type</typeStatus>
material. 
<typeStatus id="54C5887DFFA4FF94FEDBCB3BCBBCE756" box="[343,468,1611,1637]" pageId="8" pageNumber="39" type="lectotype">Lectotype</typeStatus>
</emphasis>
of 
<taxonomicName id="4C7E4D5CFFA4FF94FE70CB3CC970E756" box="[508,792,1612,1637]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="cribriventris">
<emphasis id="B90AEACDFFA4FF94FE70CB3CC970E756" box="[508,792,1612,1637]" italics="true" pageId="8" pageNumber="39">Philonthus cribriventris</emphasis>
</taxonomicName>
(here designated; 
<figureCitation id="13452A5AFFA4FF94FC78CB3CCE3CE755" box="[1012,1108,1612,1638]" captionStart="FIGURES 1 – 12" captionStartId="6.[151,275,1634,1658]" captionText="FIGURES 1 – 12. Type specimens: habitus and labels; labels with lectotype designation are not included. (1 a – b) Philonthus argentinus Bernhauer (now in Gabrius), (lectotype with labels *). (2 a – b) Philonthus jujuyensis Bernhauer (now in Gabrius), (lectotype with labels *). (3 a – b) Philonthus tucumanensis Bernhauer (now in Gabrius), (lectotype with labels *). (4) Philonthus nidicola Bernhauer (now in Gabrius), (only labels are shown *). (5 a – b) Philonthus quadraticeps (lectotype with labels *). (6 a – b) P. tenebrosus (= P. quadraticeps), (lectotype with labels *). (7 a – b) Philonthus bonariensis (lectotype with labels *). (8 a – b) Philonthus weiseri (= P. bonariensis), (lectotype with labels *). (9 a – b) Philonthus catamarcanus (= P. bonariensis), (lectotype with labels *). (10 a – b) Philonthus cribriventris (lectotype with labels *). (11) Philonthus catamarcanus var. densior (= P. cribriventris), (lectotype, labels are not shown). (12) G. chiliensis (= G. nigritulus), (paratype, labels are not shown). * Labels with lectotype designation are not included (pictures were taken prior to any designation)." pageId="8" pageNumber="39">Figs. 10</figureCitation>
a–b), 3, with labels: “Rep. 
<collectingCountry id="F369764FFFA4FF94FF1BCB02CB67E7BF" box="[151,271,1650,1676]" name="Argentina" pageId="8" pageNumber="39">Argentina</collectingCountry>
/ Prov. Tucuman / 
<date id="FFC0101FFFA4FF94FE76CB02C806E7BF" box="[506,622,1650,1676]" pageId="8" pageNumber="39" value="1908-01-30">30.I.1908</date>
/ C. Bruch”, “ 
<taxonomicName id="4C7E4D5CFFA4FF94FCADCB03C995E7BF" authority="Brnh" authorityName="Brnh" box="[801,1021,1650,1676]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="cribriventris">
<emphasis id="B90AEACDFFA4FF94FCADCB03C9D1E7BF" box="[801,953,1651,1676]" italics="true" pageId="8" pageNumber="39">cribriventris</emphasis>
Brnh
</taxonomicName>
/ 
<typeStatus id="54C5887DFFA4FF94FB9BCB02CE0EE7BF" box="[1047,1126,1650,1676]" pageId="8" pageNumber="39">Typus</typeStatus>
”, “Chicago NHMus. / M. Bernhauer Collection”, “ 
<typeStatus id="54C5887DFFA4FF94FE4FCBE9C857E780" box="[451,575,1689,1715]" pageId="8" pageNumber="39" type="lectotype">Lectotype</typeStatus>
<taxonomicName id="4C7E4D5CFFA4FF94FDC4CBE9CE54E780" authority="Bernhauer, 1912" authorityName="Bernhauer" authorityYear="1912" box="[584,1084,1689,1715]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="cribriventris">
<emphasis id="B90AEACDFFA4FF94FDC4CBE9C903E781" box="[584,875,1689,1714]" italics="true" pageId="8" pageNumber="39">Philonthus cribriventris</emphasis>
<bibRefCitation id="EFEF4B2EFFA4FF94FCFFCBE9CE54E780" author="Bernhauer" box="[883,1084,1689,1715]" pageId="8" pageNumber="41" refString="Bernhauer, M. (1912) Zur Staphylinidenfauna von Sudamerika (10. Beitrag). Verhandlungen der k. k. zoologisch- Botanischen Gesellschaft in Wien, 62, 26 - 48." type="journal article" year="1912">Bernhauer, 1912</bibRefCitation>
</taxonomicName>
/ Des. Chani-Posse de Maus 2008” (
<collectionCode id="ED6FAE1AFFA4FF94FF62CBB0CB22E7E9" LSID="urn:lsid:biocol.org:col:34795" box="[238,330,1728,1754]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="39">FMNH</collectionCode>
). 
<emphasis id="B90AEACDFFA4FF94FED1CBCFC87BE7EA" bold="true" box="[349,531,1727,1753]" pageId="8" pageNumber="39">
<typeStatus id="54C5887DFFA4FF94FED1CBCFC866E7EA" box="[349,526,1727,1753]" pageId="8" pageNumber="39" type="paralectotype">Paralectotype</typeStatus>
:
</emphasis>
3, with labels: “Rep. 
<collectingCountry id="F369764FFFA4FF94FC97CBB0C9E7E7E9" box="[795,911,1728,1754]" name="Argentina" pageId="8" pageNumber="39">Argentina</collectingCountry>
/ Prov. Tucuman / 
<date id="FFC0101FFFA4FF94FBFCCBB0CE8CE7E9" box="[1136,1252,1728,1754]" pageId="8" pageNumber="39" value="1908-01-30">30.I.1908</date>
”, “ 
<taxonomicName id="4C7E4D5CFFA4FF94FA85CBB0CABCE633" authority="Brnh" authorityName="Brnh" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="cribriventris">
<emphasis id="B90AEACDFFA4FF94FA85CBB0CFF3E7EA" box="[1289,1435,1728,1753]" italics="true" pageId="8" pageNumber="39">cribriventris</emphasis>
Brnh
</taxonomicName>
/ Cotypus”, “Chicago NHMus. / M. Bernhauer Collection”, “ 
<typeStatus id="54C5887DFFA4FF94FC4ACB96CE07E633" box="[966,1135,1766,1792]" pageId="8" pageNumber="39" type="paralectotype">Paralectotype</typeStatus>
<taxonomicName id="4C7E4D5CFFA4FF94FBFBCB97CB31E614" authority="Bernhauer, 1912" authorityName="Bernhauer" authorityYear="1912" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="cribriventris">
<emphasis id="B90AEACDFFA4FF94FBFBCB97CFF5E633" box="[1143,1437,1767,1792]" italics="true" pageId="8" pageNumber="39">Philonthus cribriventris</emphasis>
<bibRefCitation id="EFEF4B2EFFA4FF94FF1BCA7DCB31E614" author="Bernhauer" box="[151,345,1805,1831]" pageId="8" pageNumber="41" refString="Bernhauer, M. (1912) Zur Staphylinidenfauna von Sudamerika (10. Beitrag). Verhandlungen der k. k. zoologisch- Botanischen Gesellschaft in Wien, 62, 26 - 48." type="journal article" year="1912">Bernhauer, 1912</bibRefCitation>
</taxonomicName>
/ Des. Chani-Posse de Maus 2008” (
<collectionCode id="ED6FAE1AFFA4FF94FC84CA7DC90CE614" LSID="urn:lsid:biocol.org:col:34795" box="[776,868,1805,1831]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="39">FMNH</collectionCode>
).
</paragraph>
<paragraph id="8BC136DFFFA4FF94FF4ACA43CF77E647" blockId="8.[151,1437,1611,2024]" pageId="8" pageNumber="39">
<emphasis id="B90AEACDFFA4FF94FF4ACA43CB50E67E" bold="true" box="[198,312,1843,1869]" pageId="8" pageNumber="39">
<typeStatus id="54C5887DFFA4FF94FF4ACA43CB50E67E" box="[198,312,1843,1869]" pageId="8" pageNumber="39" type="holotype">Holotype</typeStatus>
</emphasis>
of 
<taxonomicName id="4C7E4D5CFFA4FF94FEEECA44C8E6E67E" box="[354,654,1844,1869]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="catamarcanus">
<emphasis id="B90AEACDFFA4FF94FEEECA44C8E6E67E" box="[354,654,1844,1869]" italics="true" pageId="8" pageNumber="39">Philonthus catamarcanus</emphasis>
</taxonomicName>
<taxonomicName id="4C7E4D5CFFA4FF94FD1ACA44C94AE67E" box="[662,802,1844,1870]" pageId="8" pageNumber="39" rank="variety" variety="densior">
var. 
<emphasis id="B90AEACDFFA4FF94FD46CA44C94AE67E" box="[714,802,1844,1869]" italics="true" pageId="8" pageNumber="39">densior</emphasis>
</taxonomicName>
(
<figureCitation id="13452A5AFFA4FF94FCBFCA44C9E3E67D" box="[819,907,1844,1870]" captionStart="FIGURES 1 – 12" captionStartId="6.[151,275,1634,1658]" captionText="FIGURES 1 – 12. Type specimens: habitus and labels; labels with lectotype designation are not included. (1 a – b) Philonthus argentinus Bernhauer (now in Gabrius), (lectotype with labels *). (2 a – b) Philonthus jujuyensis Bernhauer (now in Gabrius), (lectotype with labels *). (3 a – b) Philonthus tucumanensis Bernhauer (now in Gabrius), (lectotype with labels *). (4) Philonthus nidicola Bernhauer (now in Gabrius), (only labels are shown *). (5 a – b) Philonthus quadraticeps (lectotype with labels *). (6 a – b) P. tenebrosus (= P. quadraticeps), (lectotype with labels *). (7 a – b) Philonthus bonariensis (lectotype with labels *). (8 a – b) Philonthus weiseri (= P. bonariensis), (lectotype with labels *). (9 a – b) Philonthus catamarcanus (= P. bonariensis), (lectotype with labels *). (10 a – b) Philonthus cribriventris (lectotype with labels *). (11) Philonthus catamarcanus var. densior (= P. cribriventris), (lectotype, labels are not shown). (12) G. chiliensis (= G. nigritulus), (paratype, labels are not shown). * Labels with lectotype designation are not included (pictures were taken prior to any designation)." pageId="8" pageNumber="39">Fig. 11</figureCitation>
): Ƥ, “Rep. 
<collectingCountry id="F369764FFFA4FF94FB98CA44CEE1E67D" box="[1044,1161,1844,1870]" name="Argentina" pageId="8" pageNumber="39">Argentina</collectingCountry>
/ Prov. Tucuman / 19 / Richter”, “ 
<taxonomicName id="4C7E4D5CFFA4FF94FE99CA2ACB88E647" authority="Bernh." authorityName="Bernh." box="[277,480,1882,1908]" pageId="8" pageNumber="39" rank="variety" variety="densior">
v. 
<emphasis id="B90AEACDFFA4FF94FEBDCA2BCBE1E647" box="[305,393,1883,1908]" italics="true" pageId="8" pageNumber="39">densior</emphasis>
Bernh.
</taxonomicName>
/ 
<typeStatus id="54C5887DFFA4FF94FE7ACA2AC855E647" box="[502,573,1882,1908]" pageId="8" pageNumber="39">Typus</typeStatus>
unicus”, “Chicago NHMus. / Bernhauer Collection” (
<collectionCode id="ED6FAE1AFFA4FF94FB3BCA2ACF7BE647" LSID="urn:lsid:biocol.org:col:34795" box="[1207,1299,1882,1908]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="39">FMNH</collectionCode>
).
</paragraph>
<paragraph id="8BC136DFFFA4FF94FF49CAF0C8FBE6DB" blockId="8.[151,1437,1611,2024]" pageId="8" pageNumber="39">
<emphasis id="B90AEACDFFA4FF94FF49CAF0C964E6A9" bold="true" box="[197,780,1920,1946]" pageId="8" pageNumber="39">
Additional material. 
<collectingCountry id="F369764FFFA4FF94FE45CAF0C8E9E6A9" box="[457,641,1920,1946]" name="Argentina" pageId="8" pageNumber="39">ARGENTINA</collectingCountry>
: Tucumán:
</emphasis>
3, “Rep. 
<collectingCountry id="F369764FFFA4FF94FC0ECAF1C99EE6A8" box="[898,1014,1921,1947]" name="Argentina" pageId="8" pageNumber="39">Argentina</collectingCountry>
/ Prov. Tucuman / 19 / Richter”, “in Bernhauer handwriting &quot; 
<taxonomicName id="4C7E4D5CFFA4FF94FE4CCAD8C811E6F1" authority="Gr." authorityName="Gr." box="[448,633,1960,1986]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="39" phylum="Arthropoda" rank="species" species="longicornis">
<emphasis id="B90AEACDFFA4FF94FE4CCAD8C820E6F2" box="[448,584,1960,1985]" italics="true" pageId="8" pageNumber="39">longicornis</emphasis>
Gr.
</taxonomicName>
&quot; / det. Bernhauer”, “ 
<taxonomicName id="4C7E4D5CFFA4FF94FCF9CAD8CE2DE6F1" authority="Bernh." authorityName="Bernh." box="[885,1093,1960,1986]" pageId="8" pageNumber="39" rank="variety" variety="densior">
v. 
<emphasis id="B90AEACDFFA4FF94FC1FCAD8C985E6F2" box="[915,1005,1960,1985]" italics="true" pageId="8" pageNumber="39">densior</emphasis>
Bernh.
</taxonomicName>
/ det. Bernhauer”, “Chicago NHMus. / Bernhauer Collection” (
<collectionCode id="ED6FAE1AFFA4FF94FDA7CABEC8EFE6DB" LSID="urn:lsid:biocol.org:col:34795" box="[555,647,1998,2024]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="39">FMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3646554FFA5FF95FF4ACDE7CB4EE259" pageId="9" pageNumber="40" type="diagnosis">
<paragraph id="8BC136DFFFA5FF95FF4ACDE7C8EAE1CC" blockId="9.[151,1437,151,1067]" pageId="9" pageNumber="40">
<emphasis id="B90AEACDFFA5FF95FF4ACDE7CB2EE182" bold="true" box="[198,326,151,177]" pageId="9" pageNumber="40">Diagnosis.</emphasis>
<taxonomicName id="4C7E4D5CFFA5FF95FEC2CDE8C801E182" box="[334,617,152,177]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="cribriventris">
<emphasis id="B90AEACDFFA5FF95FEC2CDE8C801E182" box="[334,617,152,177]" italics="true" pageId="9" pageNumber="40">Philonthus cribriventris</emphasis>
</taxonomicName>
may be identified by the eyes shorter than temples, the first segment of metatarsus longer than last segment, the basal area of abdominal terga with coarse punctuation, and the apex of tergum 10 subacute in both sexes.
</paragraph>
<paragraph id="8BC136DFFFA5FF95FF4ACC7BCF1FE33D" blockId="9.[151,1437,151,1067]" pageId="9" pageNumber="40">
<emphasis id="B90AEACDFFA5FF95FF4ACC7BCB7CE016" bold="true" box="[198,276,267,293]" pageId="9" pageNumber="40">Notes.</emphasis>
In the original description of 
<taxonomicName id="4C7E4D5CFFA5FF95FD0BCC7CCE7DE015" authority="Benhauer (1912)" authorityName="Benhauer" authorityYear="1912" box="[647,1045,268,294]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="cribriventris">
<emphasis id="B90AEACDFFA5FF95FD0BCC7CC952E016" box="[647,826,268,293]" italics="true" pageId="9" pageNumber="40">P. cribriventris</emphasis>
, 
<bibRefCitation id="EFEF4B2EFFA5FF95FCC0CC7CCE7DE015" author="Benhauer" box="[844,1045,268,294]" pageId="9" pageNumber="41" refString="Bernhauer, M. (1912) Zur Staphylinidenfauna von Sudamerika (10. Beitrag). Verhandlungen der k. k. zoologisch- Botanischen Gesellschaft in Wien, 62, 26 - 48." type="journal article" year="1912">Benhauer (1912)</bibRefCitation>
</taxonomicName>
stated that two specimens were available for the description and gave two different 
<typeStatus id="54C5887DFFA5FF95FCC9CC42C915E07F" box="[837,893,306,332]" pageId="9" pageNumber="40">type</typeStatus>
localities with different collector names: “ 
<collectingCountry id="F369764FFFA5FF95FF2DCC29CB5AE040" box="[161,306,345,371]" name="Argentina" pageId="9" pageNumber="40">Argentinien</collectingCountry>
, Prov. Tucumán, 
<date id="FFC0101FFFA5FF95FE70CC29C8E1E040" box="[508,649,345,371]" pageId="9" pageNumber="40" value="1908-01-19">19. / I. 1908</date>
(C. Bruch, H. Richter), Chaunar Region (Jensen)”. Both specimens listed above agree with the description by 
<bibRefCitation id="EFEF4B2EFFA5FF95FD27CCF0C9E2E0A9" author="Bernhauer" box="[683,906,384,410]" pageId="9" pageNumber="41" refString="Bernhauer, M. (1912) Zur Staphylinidenfauna von Sudamerika (10. Beitrag). Verhandlungen der k. k. zoologisch- Botanischen Gesellschaft in Wien, 62, 26 - 48." type="journal article" year="1912">Bernhauer (1912)</bibRefCitation>
, and both have one same locality data and collection date. One of them includes the name of one of the collectors for the given locality. I have no way to decide which specimen may not belong to the original series. Therefore I have decided to consider both of them part of the 
<typeStatus id="54C5887DFFA5FF95FED4CC84CBE3E33D" box="[344,395,500,526]" pageId="9" pageNumber="40">type</typeStatus>
series, and designate the specimen with the collector´s name on its label as 
<typeStatus id="54C5887DFFA5FF95FA88CC84CF1BE33D" box="[1284,1395,500,526]" pageId="9" pageNumber="40" type="lectotype">lectotype</typeStatus>
.
</paragraph>
<paragraph id="8BC136DFFFA5FF95FF4ACF6ACB4EE259" blockId="9.[151,1437,151,1067]" pageId="9" pageNumber="40">
<bibRefCitation id="EFEF4B2EFFA5FF95FF4ACF6ACBFEE307" author="Bernhauer" box="[198,406,538,564]" pageId="9" pageNumber="41" refString="Bernhauer, M. (1916) Zur Staphylinidenfauna des tropischen Amerika, insbesondere der columbischen Cordilleren. Koleopterologische Rundschau, 5, 25 - 40." type="journal article" year="1916">Bernhauer (1916)</bibRefCitation>
described 
<taxonomicName id="4C7E4D5CFFA5FF95FD9ACF6AC8B0E300" box="[534,728,538,563]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="catamarcanus">
<emphasis id="B90AEACDFFA5FF95FD9ACF6AC8B0E300" box="[534,728,538,563]" italics="true" pageId="9" pageNumber="40">P. catamarcanus</emphasis>
</taxonomicName>
<taxonomicName id="4C7E4D5CFFA5FF95FD6CCF6AC903E300" box="[736,875,538,564]" pageId="9" pageNumber="40" rank="variety" variety="densior">
var. 
<emphasis id="B90AEACDFFA5FF95FC9FCF6AC903E300" box="[787,875,538,563]" italics="true" pageId="9" pageNumber="40">densior</emphasis>
</taxonomicName>
as a variety (“Abart”) from Tucuman province, while the 
<typeStatus id="54C5887DFFA5FF95FE80CF31CB56E368" box="[268,318,577,603]" pageId="9" pageNumber="40">type</typeStatus>
locality of 
<taxonomicName id="4C7E4D5CFFA5FF95FE4ACF31C8E0E369" box="[454,648,577,602]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="catamarcanus">
<emphasis id="B90AEACDFFA5FF95FE4ACF31C8E0E369" box="[454,648,577,602]" italics="true" pageId="9" pageNumber="40">P. catamarcanus</emphasis>
</taxonomicName>
was Catamarca and San Luis provinces. According to Article 45.6 (
<bibRefCitation id="EFEF4B2EFFA5FF95FF13CF18CB41E3B1" author="ICZN" box="[159,297,616,642]" pageId="9" pageNumber="42" refString="ICZN (1999) International Code of Zoological Nomenclatur e. Fourth Edition. The International Trust for Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
) the name 
<taxonomicName id="4C7E4D5CFFA5FF95FE20CF18C86CE3B2" box="[428,516,616,641]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="densior">
<emphasis id="B90AEACDFFA5FF95FE20CF18C86CE3B2" box="[428,516,616,641]" italics="true" pageId="9" pageNumber="40">densior</emphasis>
</taxonomicName>
is of subspecific rank and available. Bernhauer explicitely stated that the 
<typeStatus id="54C5887DFFA5FF95FAE5CF18CFF4E3B1" box="[1385,1436,616,642]" pageId="9" pageNumber="40">type</typeStatus>
series of 
<taxonomicName id="4C7E4D5CFFA5FF95FE89CFFFCBA5E39B" box="[261,461,655,680]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="catamarcanus">
<emphasis id="B90AEACDFFA5FF95FE89CFFFCBA5E39B" box="[261,461,655,680]" italics="true" pageId="9" pageNumber="40">P. catamarcanus</emphasis>
</taxonomicName>
<taxonomicName id="4C7E4D5CFFA5FF95FE59CFFEC80CE39B" box="[469,612,654,680]" pageId="9" pageNumber="40" rank="variety" variety="densior">
var. 
<emphasis id="B90AEACDFFA5FF95FD86CFFFC80CE39B" box="[522,612,655,680]" italics="true" pageId="9" pageNumber="40">densior</emphasis>
</taxonomicName>
included a single specimen, but did not mention whether this was a male or a female. Considering that the female specimen of 
<taxonomicName id="4C7E4D5CFFA5FF95FCF5CFC5CE2DE3FD" box="[889,1093,693,718]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="catamarcanus">
<emphasis id="B90AEACDFFA5FF95FCF5CFC5CE2DE3FD" box="[889,1093,693,718]" italics="true" pageId="9" pageNumber="40">P. catamarcanus</emphasis>
</taxonomicName>
<taxonomicName id="4C7E4D5CFFA5FF95FBC2CFC5CE89E3FD" box="[1102,1249,693,719]" pageId="9" pageNumber="40" rank="variety" variety="densior">
var. 
<emphasis id="B90AEACDFFA5FF95FB08CFC5CE89E3FD" box="[1156,1249,693,718]" italics="true" pageId="9" pageNumber="40">densior</emphasis>
</taxonomicName>
listed above is labeled as “ 
<typeStatus id="54C5887DFFA5FF95FEAECFACCB06E3C5" box="[290,366,732,758]" pageId="9" pageNumber="40">Typus</typeStatus>
unicus”, and locality label corresponds to the locality mentioned in the description, this female specimen is regarded as the 
<typeStatus id="54C5887DFFA5FF95FDB2CE72C8C1E22F" box="[574,681,770,796]" pageId="9" pageNumber="40" type="holotype">holotype</typeStatus>
. The male specimen from the same locality listed in Additional material was not labeled by Bernhauer as the 
<typeStatus id="54C5887DFFA5FF95FD34CE59C886E270" box="[696,750,809,835]" pageId="9" pageNumber="40">type</typeStatus>
, and therefore cannot be the 
<typeStatus id="54C5887DFFA5FF95FBCBCE59CEC6E270" box="[1095,1198,809,835]" pageId="9" pageNumber="40" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C7E4D5CFFA5FF95FB54CE59CFF4E271" box="[1240,1436,809,834]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="catamarcanus">
<emphasis id="B90AEACDFFA5FF95FB54CE59CFF4E271" box="[1240,1436,809,834]" italics="true" pageId="9" pageNumber="40">P. catamarcanus</emphasis>
</taxonomicName>
<taxonomicName id="4C7E4D5CFFA5FF95FF1BCE20CB77E25A" box="[151,287,848,874]" pageId="9" pageNumber="40" rank="variety" variety="densior">
var. 
<emphasis id="B90AEACDFFA5FF95FF45CE20CB77E25A" box="[201,287,848,873]" italics="true" pageId="9" pageNumber="40">densior</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3646554FFA5FF95FF4ACE06C92EE518" pageId="9" pageNumber="40" type="discussion">
<paragraph id="8BC136DFFFA5FF95FF4ACE06C92EE518" blockId="9.[151,1437,151,1067]" pageId="9" pageNumber="40">
<emphasis id="B90AEACDFFA5FF95FF4ACE06CB3DE2A3" bold="true" box="[198,341,886,912]" pageId="9" pageNumber="40">Comments.</emphasis>
<bibRefCitation id="EFEF4B2EFFA5FF95FED0CE06C83FE2A3" author="Bernhauer" box="[348,599,886,912]" pageId="9" pageNumber="41" refString="Bernhauer, M. (1916) Zur Staphylinidenfauna des tropischen Amerika, insbesondere der columbischen Cordilleren. Koleopterologische Rundschau, 5, 25 - 40." type="journal article" year="1916">Bernhauer (1916: 36)</bibRefCitation>
described 
<taxonomicName id="4C7E4D5CFFA5FF95FD5ACE07C922E2A3" box="[726,842,887,912]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="densior">
<emphasis id="B90AEACDFFA5FF95FD5ACE07C922E2A3" box="[726,842,887,912]" italics="true" pageId="9" pageNumber="40">P. densior</emphasis>
</taxonomicName>
as a variety of 
<taxonomicName id="4C7E4D5CFFA5FF95FC71CE07CECAE2A3" box="[1021,1186,887,912]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="catamarcanus">
<emphasis id="B90AEACDFFA5FF95FC71CE07CECAE2A3" box="[1021,1186,887,912]" italics="true" pageId="9" pageNumber="40">catamarcanus</emphasis>
</taxonomicName>
and since then it was a valid taxon at the subspecific rank. The 
<typeStatus id="54C5887DFFA5FF95FD0ACEEDC8D0E284" box="[646,696,925,951]" pageId="9" pageNumber="40">type</typeStatus>
material of 
<taxonomicName id="4C7E4D5CFFA5FF95FCC4CEEDC9D6E285" box="[840,958,925,950]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="densior">
<emphasis id="B90AEACDFFA5FF95FCC4CEEDC9D6E285" box="[840,958,925,950]" italics="true" pageId="9" pageNumber="40">P. densior</emphasis>
</taxonomicName>
(
<typeStatus id="54C5887DFFA5FF95FC41CEEDCE52E284" box="[973,1082,925,951]" pageId="9" pageNumber="40" type="lectotype">lectotype</typeStatus>
Ƥ, 
<typeStatus id="54C5887DFFA5FF95FBE4CEEDCF6EE284" box="[1128,1286,925,951]" pageId="9" pageNumber="40" type="paralectotype">paralectotype</typeStatus>
3) (
<figureCitation id="13452A5AFFA5FF95FAB2CEEDCFFBE284" box="[1342,1427,925,951]" captionStart="FIGURES 1 – 12" captionStartId="6.[151,275,1634,1658]" captionText="FIGURES 1 – 12. Type specimens: habitus and labels; labels with lectotype designation are not included. (1 a – b) Philonthus argentinus Bernhauer (now in Gabrius), (lectotype with labels *). (2 a – b) Philonthus jujuyensis Bernhauer (now in Gabrius), (lectotype with labels *). (3 a – b) Philonthus tucumanensis Bernhauer (now in Gabrius), (lectotype with labels *). (4) Philonthus nidicola Bernhauer (now in Gabrius), (only labels are shown *). (5 a – b) Philonthus quadraticeps (lectotype with labels *). (6 a – b) P. tenebrosus (= P. quadraticeps), (lectotype with labels *). (7 a – b) Philonthus bonariensis (lectotype with labels *). (8 a – b) Philonthus weiseri (= P. bonariensis), (lectotype with labels *). (9 a – b) Philonthus catamarcanus (= P. bonariensis), (lectotype with labels *). (10 a – b) Philonthus cribriventris (lectotype with labels *). (11) Philonthus catamarcanus var. densior (= P. cribriventris), (lectotype, labels are not shown). (12) G. chiliensis (= G. nigritulus), (paratype, labels are not shown). * Labels with lectotype designation are not included (pictures were taken prior to any designation)." pageId="9" pageNumber="40">Fig. 11</figureCitation>
) agrees with the 
<typeStatus id="54C5887DFFA5FF95FED0CEB4CBA5E2ED" box="[348,461,964,990]" pageId="9" pageNumber="40" type="lectotype">lectotype</typeStatus>
of 
<taxonomicName id="4C7E4D5CFFA5FF95FE7BCEB4C8DAE2EE" box="[503,690,964,989]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="cribriventris">
<emphasis id="B90AEACDFFA5FF95FE7BCEB4C8DAE2EE" box="[503,690,964,989]" italics="true" pageId="9" pageNumber="40">P. cribriventris</emphasis>
</taxonomicName>
(
<figureCitation id="13452A5AFFA5FF95FD4FCEB4C940E2ED" box="[707,808,964,990]" captionStart="FIGURES 1 – 12" captionStartId="6.[151,275,1634,1658]" captionText="FIGURES 1 – 12. Type specimens: habitus and labels; labels with lectotype designation are not included. (1 a – b) Philonthus argentinus Bernhauer (now in Gabrius), (lectotype with labels *). (2 a – b) Philonthus jujuyensis Bernhauer (now in Gabrius), (lectotype with labels *). (3 a – b) Philonthus tucumanensis Bernhauer (now in Gabrius), (lectotype with labels *). (4) Philonthus nidicola Bernhauer (now in Gabrius), (only labels are shown *). (5 a – b) Philonthus quadraticeps (lectotype with labels *). (6 a – b) P. tenebrosus (= P. quadraticeps), (lectotype with labels *). (7 a – b) Philonthus bonariensis (lectotype with labels *). (8 a – b) Philonthus weiseri (= P. bonariensis), (lectotype with labels *). (9 a – b) Philonthus catamarcanus (= P. bonariensis), (lectotype with labels *). (10 a – b) Philonthus cribriventris (lectotype with labels *). (11) Philonthus catamarcanus var. densior (= P. cribriventris), (lectotype, labels are not shown). (12) G. chiliensis (= G. nigritulus), (paratype, labels are not shown). * Labels with lectotype designation are not included (pictures were taken prior to any designation)." pageId="9" pageNumber="40">Figs. 10</figureCitation>
a–b) in external characters and in the shape of the aedeagus. Therefore, 
<emphasis id="B90AEACDFFA5FF95FE1FCE9AC86CE530" box="[403,516,1002,1027]" italics="true" pageId="9" pageNumber="40">P. d e n s i o r</emphasis>
is removed from synonymy with 
<taxonomicName id="4C7E4D5CFFA5FF95FC1ACE9ACE30E530" box="[918,1112,1002,1027]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="catamarcanus">
<emphasis id="B90AEACDFFA5FF95FC1ACE9ACE30E530" box="[918,1112,1002,1027]" italics="true" pageId="9" pageNumber="40">P. catamarcanus</emphasis>
</taxonomicName>
(at the rank of species) and placed in synonymy with 
<taxonomicName id="4C7E4D5CFFA5FF95FE49C961C929E518" authority="Bernhauer, 1912" authorityName="Bernhauer" authorityYear="1912" box="[453,833,1041,1067]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="40" phylum="Arthropoda" rank="species" species="cribriventris">
<emphasis id="B90AEACDFFA5FF95FE49C961C81CE519" box="[453,628,1041,1066]" italics="true" pageId="9" pageNumber="40">P. cribriventris</emphasis>
<bibRefCitation id="EFEF4B2EFFA5FF95FDF0C961C929E518" author="Bernhauer" box="[636,833,1041,1067]" pageId="9" pageNumber="41" refString="Bernhauer, M. (1912) Zur Staphylinidenfauna von Sudamerika (10. Beitrag). Verhandlungen der k. k. zoologisch- Botanischen Gesellschaft in Wien, 62, 26 - 48." type="journal article" year="1912">Bernhauer, 1912</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>