treatments-xml/data/03/CA/87/03CA87DBFF86841EFF645CAEFFCC25B8.xml

<document id="B4F969EE94C8317FADFDEFDCFA171252" ID-DOI="10.11646/zootaxa.5434.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10954759" ID-ZooBank="7BB35AD7-6AE7-4361-B9EF-520F6C978B14" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1712753665371" checkinUser="plazi" docAuthor="Salata, Sebastian, Demetriou, Jakovos, Georgiadis, Christos &amp; Borowiec, Lech" docDate="2024" docId="03CA87DBFF86841EFF645CAEFFCC25B8" docLanguage="en" docName="zootaxa.5433.5.1.pdf" docOrigin="Zootaxa 5434 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5434.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Temnothorax cerastarum Salata &amp; Demetriou &amp; Georgiadis &amp; Borowiec 2024, n. sp." docType="treatment" docVersion="2" lastPageNumber="41" masterDocId="FFF3FFA3FFA58436FFF35C39FD42244D" masterDocTitle="The genus Temnothorax Mayr, 1861 (Hymenoptera: Formicidae) in Cyprus" masterLastPageNumber="69" masterPageNumber="1" pageNumber="36" updateTime="1712754637106" updateUser="ExternalLinkService">
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<mods:title id="B78C852E16B7777BCBDA17C7074392D4">The genus Temnothorax Mayr, 1861 (Hymenoptera: Formicidae) in Cyprus</mods:title>
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<mods:namePart id="F19C8345BA6BB7F4586AB5EE836F1DA4">Salata, Sebastian</mods:namePart>
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<mods:affiliation id="4C25912CDD3874EBB6C09CD87200F0D9">University of Wrocław, Department of Biodiversity and Evolutionary Taxonomy, Myrmecological Laboratory, Przybyszewskiego 65, 51 - 148 Wrocław, Poland &amp; sebastian. salata @ uwr. edu. pl, https: // orcid. org / 0000 - 0003 - 0811 - 2309</mods:affiliation>
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<mods:namePart id="9F788834145A37EF1192AB0C70462F16">Demetriou, Jakovos</mods:namePart>
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<mods:affiliation id="0DEB30861438CBA7EDCDBB67C28885AE">Department of Ecology and Systematics, Faculty of Biology, National and Kapodistrian University of Athens, 15784 Athens, Greece &amp; Laboratory of Vector Ecology and Applied Entomology, Joint Services Health Unit Cyprus, BFC RAF Akrotiri BFPO 57, Akrotiri, Cyprus &amp; Enalia Physis Environmental Research Centre, Acropoleos 2, Aglantzia 2101, Nicosia, Cyprus jakovosdemetriou @ gmail. com, https: // orcid. org / 0000 - 0001 - 5273 - 7109</mods:affiliation>
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<mods:namePart id="E5171B5595D90BA7D5A87774067224A8">Georgiadis, Christos</mods:namePart>
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<mods:affiliation id="0B80CB97EF5F1CEE7B02974BF3067381">Section of Zoology and Marine Biology, Department of Biology, National and Kapodistrian University of Athens, 15784 Athens, Greece &amp; Museum of Zoology, National and Kapodistrian University of Athens, 15784 Athens, Greece cgeorgia @ biol. uoa. gr, https: // orcid. org / 0000 - 0002 - 2728 - 3122</mods:affiliation>
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<mods:namePart id="D4EE900C6134097ABE9B37989971E60A">Borowiec, Lech</mods:namePart>
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<mods:affiliation id="5DC00C2E54D7AE862A956AA1D0B9BD9C">University of Wrocław, Department of Biodiversity and Evolutionary Taxonomy, Myrmecological Laboratory, Przybyszewskiego 65, 51 - 148 Wrocław, Poland &amp; University of Wrocław, Department of Biodiversity and Evolutionary Taxonomy, Myrmecological Laboratory, Przybyszewskiego 65, 51 - 148 Wrocław, Poland &amp; lech. borowiec @ uwr. edu. pl, https: // orcid. org / 0000 - 0001 - 5668 - 6855 Corresponding author</mods:affiliation>
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<taxonomicName id="4C634D4EFF868415FF645CAEFC8B24FC" authority="Salata &amp; Demetriou &amp; Georgiadis &amp; Borowiec, 2024" authorityName="Salata &amp; Demetriou &amp; Georgiadis &amp; Borowiec" authorityYear="2024" box="[151,457,151,178]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="cerastarum" status="sp. nov.">
<emphasis id="B917EADFFF868415FF645CAEFC8B24FC" bold="true" box="[151,457,151,178]" italics="true" pageId="35" pageNumber="36">Temnothorax cerastarum</emphasis>
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(
<figureCitation id="13582A48FF868415FDD15CAEFFE824FC" box="[546,682,151,177]" captionStart-0="FIGURES 29, 30" captionStart-1="FIGURE 31" captionStart-2="FIGURES 32, 33" captionStart-3="FIGURE 34" captionStartId-0="35.[151,264,1981,2005]" captionStartId-1="36.[151,250,1974,1998]" captionStartId-2="37.[151,264,1447,1471]" captionStartId-3="39.[151,250,1492,1516]" captionTargetBox-0="[151,1436,417,1942]" captionTargetBox-1="[151,1436,741,1951]" captionTargetBox-2="[151,1436,186,1423]" captionTargetBox-3="[151,1436,181,1468]" captionTargetId-0="figure-98@35.[151,1436,417,1958]" captionTargetId-1="figure-303@36.[151,1436,741,1951]" captionTargetId-2="figure-18@37.[151,1436,181,1423]" captionTargetId-3="figure-18@39.[151,1436,181,1468]" captionTargetPageId-0="35" captionTargetPageId-1="36" captionTargetPageId-2="37" captionTargetPageId-3="39" captionText-0="FIGURES 29, 30. Holotype worker of Temnothorax cerastarum n. sp. 29 dorsal 30 lateral (scale bar = 1 mm)." captionText-1="FIGURE 31. Holotype worker head of Temnothorax cerastarum n. sp. (scale bar = 0.5 mm)." captionText-2="FIGURES 32, 33. Gyne of Temnothorax cerastarum n. sp. 32 dorsal 33 lateral (scale bar = 1 mm)." captionText-3="FIGURE 34. Gyne head of Temnothorax cerastarum n. sp. (scale bar = 0.5 mm)." figureDoi-0="http://doi.org/10.5281/zenodo.10954807" figureDoi-1="http://doi.org/10.5281/zenodo.10954809" figureDoi-2="http://doi.org/10.5281/zenodo.10954811" figureDoi-3="http://doi.org/10.5281/zenodo.10954813" httpUri-0="https://zenodo.org/record/10954807/files/figure.png" httpUri-1="https://zenodo.org/record/10954809/files/figure.png" httpUri-2="https://zenodo.org/record/10954811/files/figure.png" httpUri-3="https://zenodo.org/record/10954813/files/figure.png" pageId="35" pageNumber="36">Figs 29–34</figureCitation>
, 
<figureCitation id="13582A48FF868415FD445CAEFF9424FC" box="[695,726,151,177]" captionStart="FIGURE 64" captionStartId="64.[151,250,1944,1968]" captionTargetBox="[179,1409,181,1920]" captionTargetId="figure-23@64.[179,1409,181,1920]" captionTargetPageId="64" captionText="FIGURE 64. Distribution in Cyprus Temnothorax cerastarum n. sp. (upper map) and T. evagorae n. sp. + T. oreades n. sp. (bottom map)" figureDoi="http://doi.org/10.5281/zenodo.10954860" httpUri="https://zenodo.org/record/10954860/files/figure.png" pageId="35" pageNumber="36">64</figureCitation>
)
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3796546FF868415FF645CE6FFBF25C4" pageId="35" pageNumber="36" type="etymology">
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<emphasis id="B917EADFFF868415FF645CE6FC5C24B4" bold="true" box="[151,286,223,249]" pageId="35" pageNumber="36">Etymology.</emphasis>
Named after Cerastes (horned ones) [Greek Κεραστές], a group of men native to 
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, who Aphrodite transformed into a tribe of bull-horned centaurs, as punishment for murdering foreign visitors to the island (probably pilgrims visiting the goddess' famous shrine). They were born of Gaia, the Earth, when Zeus accidentally impregnated her during a failed attempt to couple with the goddess Aphrodite. We gave this name from the propodeum's protruding spines (resembling horns).
</paragraph>
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<emphasis id="B917EADFFF868415FF645B84FC182398" bold="true" box="[151,346,1981,2005]" pageId="35" pageNumber="36">FIGURES 29, 30.</emphasis>
Holotype worker of 
<taxonomicName id="4C634D4EFF868415FDDC5B87FE6F2398" authority="Salata &amp; Demetriou &amp; Georgiadis &amp; Borowiec, 2024" authorityName="Salata &amp; Demetriou &amp; Georgiadis &amp; Borowiec" authorityYear="2024" box="[559,813,1982,2006]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="cerastarum" status="sp. nov.">
<emphasis id="B917EADFFF868415FDDC5B87FE6F2398" box="[559,813,1982,2006]" italics="true" pageId="35" pageNumber="36">Temnothorax cerastarum</emphasis>
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<emphasis id="B917EADFFF868415FCC05B87FECE2398" bold="true" box="[819,908,1981,2006]" pageId="35" pageNumber="36">
<taxonomicNameLabel id="A22457A4FF868415FCC05B87FE2F239B" box="[819,877,1982,2006]" pageId="35" pageNumber="36" rank="species">n. sp.</taxonomicNameLabel>
29
</emphasis>
dorsal 
<emphasis id="B917EADFFF868415FC245B84FEB22398" bold="true" box="[983,1008,1981,2005]" pageId="35" pageNumber="36">30</emphasis>
lateral (scale bar = 1 mm).
</paragraph>
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<emphasis id="B917EADFFF818412FF345CAFFCF124FD" bold="true" box="[199,435,150,176]" pageId="36" pageNumber="37">Material examined.</emphasis>
<materialsCitation id="3B0B3C90FF818412FE4F5CAEFFED2498" collectingDate="2022-04-22" collectionCode="MNHW" collectorName="L. Borowiec" country="Cyprus" elevation="1138" latitude="34.99092" location="Cedar Valley" longLatPrecision="1" longitude="32.68846" pageId="36" pageNumber="37" specimenCount="1" stateProvince="Pafos" typeStatus="holotype">
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<typeStatus id="54D8886FFF818412FE4F5CAEFF6F24FC" box="[444,557,151,177]" pageId="36" pageNumber="37" type="holotype">Holotype</typeStatus>
:
</emphasis>
worker (pin) “ 
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, 
<collectingRegion id="49A7F82FFF818412FCAD5CAEFEF024FC" box="[862,946,151,177]" country="Cyprus" name="Pafos" pageId="36" pageNumber="37">Paphos</collectingRegion>
, 
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<elevation id="004ED1FEFF818412FC4C5CAEF95724FC" box="[959,1045,151,177]" metricMagnitude="3" metricUnit="m" metricValue="1.138" pageId="36" pageNumber="37" unit="m" value="1138.0">1138 m</elevation>
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| 
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loc. 2 | 
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/ 
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| 
<date id="FFDD100DFF818412FEE05C82FCD62498" box="[275,404,187,213]" pageId="36" pageNumber="37" value="2022-04-22">
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</date>
, 
<collectorName id="2696531BFF818412FE6D5C82FF6E2498" box="[414,556,187,213]" pageId="36" pageNumber="37">L. Borowiec</collectorName>
” (
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)
</materialsCitation>
. 
<materialsCitation id="3B0B3C90FF818412FD415C82FCE524B4" collectingDate="2022-04-22" collectionCode="MNHW, JDC, ZMUA" collectorName="L. Borowiec" country="Cyprus" elevation="1138" latitude="34.99092" location="Cedar Valley" longLatPrecision="1" longitude="32.68846" pageId="36" pageNumber="37" specimenCount="41" specimenCount-worker="41" stateProvince="Pafos" typeStatus="paratype">
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<typeStatus id="54D8886FFF818412FD415C82FE682498" box="[690,810,187,213]" pageId="36" pageNumber="37" type="paratype">Paratypes</typeStatus>
</emphasis>
(1 gyne, 
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, pin): the same data as for holotype (
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, 
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, 
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)
</materialsCitation>
.
</paragraph>
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<materialsCitation id="3B0B3C90FF818412FF345D3BF91C2550" box="[199,1118,258,285]" collectingDate="2022-04-22" collectionCode="MNHW" collectorName="L. Borowiec" country="Cyprus" elevation="1138" latitude="34.99092" location="Cedar Valley" longLatPrecision="1" longitude="32.68846" pageId="36" pageNumber="37" specimenCount="15" specimenCount-worker="15" stateProvince="Pafos">
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<typeStatus id="54D8886FFF818412FF345D3BFCB42551" box="[199,502,258,284]" isOtherTypeStatus="true" pageId="36" pageNumber="37">Other material examined</typeStatus>
.
</emphasis>
<specimenCount id="9D65FD44FF818412FDF15D3BFF3D2550" box="[514,639,258,285]" count="15" pageId="36" pageNumber="37" type="worker">15 workers</specimenCount>
(EtOH): the same data as for the holotype
</materialsCitation>
.
</paragraph>
<paragraph id="8BDC36CDFF818412FF345D1FF96F2680" blockId="36.[151,1437,150,717]" pageId="36" pageNumber="37">
<emphasis id="B917EADFFF818412FF345D1FFCEA250C" bold="true" box="[199,424,294,321]" pageId="36" pageNumber="37">Comparative note.</emphasis>
<taxonomicName id="4C634D4EFF818412FE415D1EFF8C250D" authorityName="Salata &amp; Demetriou &amp; Georgiadis &amp; Borowiec" authorityYear="2024" box="[434,718,295,321]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="cerastarum">
<emphasis id="B917EADFFF818412FE415D1EFF8C250D" box="[434,718,295,321]" italics="true" pageId="36" pageNumber="37">Temnothorax cerastarum</emphasis>
</taxonomicName>
is a member of the 
<taxonomicName id="4C634D4EFF818412FC485D1EF970250D" baseAuthorityName="Forel" baseAuthorityYear="1911" box="[955,1074,295,320]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="graecus">
<emphasis id="B917EADFFF818412FC485D1EF970250D" box="[955,1074,295,320]" italics="true" pageId="36" pageNumber="37">T. graecus</emphasis>
</taxonomicName>
group. It has uniformly yellow antennae and in this character is similar to 
<taxonomicName id="4C634D4EFF818412FD755D72FE492529" authorityName="Salata &amp; Demetriou &amp; Georgiadis &amp; Borowiec" authorityYear="2024" box="[646,779,331,356]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="evagorae">
<emphasis id="B917EADFFF818412FD755D72FE492529" box="[646,779,331,356]" italics="true" pageId="36" pageNumber="37">T. evagorae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C634D4EFF818412FCB55D72FEF92528" authorityName="Salata &amp; Demetriou &amp; Georgiadis &amp; Borowiec" authorityYear="2024" box="[838,955,331,357]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="oreades">
<emphasis id="B917EADFFF818412FCB55D72FEF92528" box="[838,955,331,357]" italics="true" pageId="36" pageNumber="37">T. oreades</emphasis>
</taxonomicName>
, while the fourth species of this group, 
<taxonomicName id="4C634D4EFF818412FA7B5D72FDAA25C4" baseAuthorityName="Forel" baseAuthorityYear="1911" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="aeolius">
<emphasis id="B917EADFFF818412FA7B5D72FDAA25C4" italics="true" pageId="36" pageNumber="37">T. aeolius</emphasis>
</taxonomicName>
, well differs in partly brown to black antennal club. 
<taxonomicName id="4C634D4EFF818412FCC05D56F90E25C5" authorityName="Salata &amp; Demetriou &amp; Georgiadis &amp; Borowiec" authorityYear="2024" box="[819,1100,367,393]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="cerastarum">
<emphasis id="B917EADFFF818412FCC05D56F90E25C5" box="[819,1100,367,393]" italics="true" pageId="36" pageNumber="37">Temnothorax cerastarum</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C634D4EFF818412FB2C5D56F81025C4" authorityName="Salata &amp; Demetriou &amp; Georgiadis &amp; Borowiec" authorityYear="2024" box="[1247,1362,367,393]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="oreades">
<emphasis id="B917EADFFF818412FB2C5D56F81025C4" box="[1247,1362,367,393]" italics="true" pageId="36" pageNumber="37">T. oreades</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C634D4EFF818412FA7B5D56FDBD25E1" authorityName="Salata &amp; Demetriou &amp; Georgiadis &amp; Borowiec" authorityYear="2024" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="evagorae">
<emphasis id="B917EADFFF818412FA7B5D56FDBD25E1" italics="true" pageId="36" pageNumber="37">T. evagorae</emphasis>
</taxonomicName>
in the less sculptured head with a great part of frontal and occipital area smooth and shiny, longer, needle-shaped propodeal spines with mean PSLI 0.297, and longer antennal scapus with mean SI1 0.933 and SI2 0.785. The last yellow Cypriot species, 
<taxonomicName id="4C634D4EFF818412FE245DE2FFC225B8" authorityName="Salata &amp; Demetriou &amp; Georgiadis &amp; Borowiec" authorityYear="2024" box="[471,640,475,501]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="akrotiriensis">
<emphasis id="B917EADFFF818412FE245DE2FFC225B8" box="[471,640,475,501]" italics="true" pageId="36" pageNumber="37">T. akrotiriensis</emphasis>
</taxonomicName>
differs from the pale form of 
<taxonomicName id="4C634D4EFF818412FC275DE2F93225B9" authorityName="Salata &amp; Demetriou &amp; Georgiadis &amp; Borowiec" authorityYear="2024" box="[980,1136,475,500]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="cerastarum">
<emphasis id="B917EADFFF818412FC275DE2F93225B9" box="[980,1136,475,500]" italics="true" pageId="36" pageNumber="37">T. cerastarum</emphasis>
</taxonomicName>
in a more sculptured head with a distinctly microreticulate background between the prominent sculpture and thus dull frontal part of the head while in 
<taxonomicName id="4C634D4EFF818412FF045E1AFCD32671" authorityName="Salata &amp; Demetriou &amp; Georgiadis &amp; Borowiec" authorityYear="2024" box="[247,401,547,572]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="cerastarum">
<emphasis id="B917EADFFF818412FF045E1AFCD32671" box="[247,401,547,572]" italics="true" pageId="36" pageNumber="37">T. cerastarum</emphasis>
</taxonomicName>
this background sculpture is diffused or absent thus frontal part of the head is distinctly shiny. 
<taxonomicName id="4C634D4EFF818412FF645E7EFCFF262C" authorityName="Salata &amp; Demetriou &amp; Georgiadis &amp; Borowiec" authorityYear="2024" box="[151,445,583,609]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="akrotiriensis">
<emphasis id="B917EADFFF818412FF645E7EFCFF262C" box="[151,445,583,609]" italics="true" pageId="36" pageNumber="37">Temnothorax akrotiriensis</emphasis>
</taxonomicName>
has a very narrow dark band at the end of the first gastral tergite, which occupies at most its ¼ posterior surface, while in 
<taxonomicName id="4C634D4EFF818412FDF75E52FFE226C9" authorityName="Salata &amp; Demetriou &amp; Georgiadis &amp; Borowiec" authorityYear="2024" box="[516,672,619,644]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="cerastarum">
<emphasis id="B917EADFFF818412FDF75E52FFE226C9" box="[516,672,619,644]" italics="true" pageId="36" pageNumber="37">T. cerastarum</emphasis>
</taxonomicName>
this band usually occupies more than half the surface of the tergite. 
<taxonomicName id="4C634D4EFF818412FF645EB6FCF226E5" authorityName="Salata &amp; Demetriou &amp; Georgiadis &amp; Borowiec" authorityYear="2024" box="[151,432,655,681]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="cerastarum">
<emphasis id="B917EADFFF818412FF645EB6FCF226E5" box="[151,432,655,681]" italics="true" pageId="36" pageNumber="37">Temnothorax cerastarum</emphasis>
</taxonomicName>
was collected in a mountain pine forest at an altitude of 
<quantity id="4C9B9B28FF818412FBD75EB6F93A26E4" box="[1060,1144,655,681]" metricMagnitude="3" metricUnit="m" metricValue="1.138" pageId="36" pageNumber="37" unit="m" value="1138.0">1138 m</quantity>
while 
<taxonomicName id="4C634D4EFF818412FB305EB6F82926E4" authorityName="Salata &amp; Demetriou &amp; Georgiadis &amp; Borowiec" authorityYear="2024" box="[1219,1387,655,681]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="akrotiriensis">
<emphasis id="B917EADFFF818412FB305EB6F82926E4" box="[1219,1387,655,681]" italics="true" pageId="36" pageNumber="37">T. akrotiriensis</emphasis>
</taxonomicName>
was collected only in lowland open habitats from the sea coast to an altitude of 
<specimenCount id="9D65FD44FF818412FC2C5E8BF96F2680" box="[991,1069,690,717]" count="156" pageId="36" pageNumber="37" type="generic">
<quantity id="4C9B9B28FF818412FC2C5E8BF96B2680" box="[991,1065,690,717]" metricMagnitude="2" metricUnit="m" metricValue="1.56" pageId="36" pageNumber="37" unit="m" value="156.0">156 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<caption id="DF1C6645FF818412FF645B8FF90A2382" ID-DOI="http://doi.org/10.5281/zenodo.10954809" ID-Zenodo-Dep="10954809" box="[151,1096,1974,1999]" httpUri="https://zenodo.org/record/10954809/files/figure.png" pageId="36" pageNumber="37" startId="36.[151,250,1974,1998]" targetBox="[151,1436,741,1951]" targetPageId="36" targetType="figure">
<paragraph id="8BDC36CDFF818412FF645B8FF90A2382" blockId="36.[151,1096,1974,1999]" box="[151,1096,1974,1999]" pageId="36" pageNumber="37">
<emphasis id="B917EADFFF818412FF645B8FFC622383" bold="true" box="[151,288,1974,1998]" pageId="36" pageNumber="37">FIGURE 31.</emphasis>
Holotype worker head of 
<taxonomicName id="4C634D4EFF818412FDD95B8FFE6A2383" authority="Salata &amp; Demetriou &amp; Georgiadis &amp; Borowiec, 2024" authorityName="Salata &amp; Demetriou &amp; Georgiadis &amp; Borowiec" authorityYear="2024" box="[554,808,1974,1998]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="cerastarum" status="sp. nov.">
<emphasis id="B917EADFFF818412FDD95B8FFE6A2383" box="[554,808,1974,1998]" italics="true" pageId="36" pageNumber="37">Temnothorax cerastarum</emphasis>
</taxonomicName>
<emphasis id="B917EADFFF818412FCDC5B8EFE2A2382" bold="true" box="[815,872,1975,1999]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="A22457A4FF818412FCDC5B8EFE2A2382" box="[815,872,1975,1999]" pageId="36" pageNumber="37" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(scale bar = 0.5 mm).
</paragraph>
</caption>
<caption id="DF1C6645FF808413FF64599EF9CF21F2" ID-DOI="http://doi.org/10.5281/zenodo.10954811" ID-Zenodo-Dep="10954811" box="[151,1165,1447,1471]" httpUri="https://zenodo.org/record/10954811/files/figure.png" pageId="37" pageNumber="38" startId="37.[151,264,1447,1471]" targetBox="[151,1436,186,1423]" targetPageId="37" targetType="figure">
<paragraph id="8BDC36CDFF808413FF64599EF9CF21F2" blockId="37.[151,1165,1447,1471]" box="[151,1165,1447,1471]" pageId="37" pageNumber="38">
<emphasis id="B917EADFFF808413FF64599EFC1821F2" bold="true" box="[151,346,1447,1471]" pageId="37" pageNumber="38">FIGURES 32, 33.</emphasis>
Gyne of 
<taxonomicName id="4C634D4EFF808413FE4A599EFFF521F2" authority="Salata &amp; Demetriou &amp; Georgiadis &amp; Borowiec, 2024" authorityName="Salata &amp; Demetriou &amp; Georgiadis &amp; Borowiec" authorityYear="2024" box="[441,695,1447,1471]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="cerastarum" status="sp. nov.">
<emphasis id="B917EADFFF808413FE4A599EFFF521F2" box="[441,695,1447,1471]" italics="true" pageId="37" pageNumber="38">Temnothorax cerastarum</emphasis>
</taxonomicName>
<emphasis id="B917EADFFF808413FD4D599EFE5421F2" bold="true" box="[702,790,1447,1471]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="A22457A4FF808413FD4D599EFFB521F2" box="[702,759,1447,1471]" pageId="37" pageNumber="38" rank="species">n. sp.</taxonomicNameLabel>
32
</emphasis>
dorsal 
<emphasis id="B917EADFFF808413FC92599EFE3821F2" bold="true" box="[865,890,1447,1471]" pageId="37" pageNumber="38">33</emphasis>
lateral (scale bar = 1 mm).
</paragraph>
</caption>
<subSubSection id="C3796546FF80841EFF3459D4FFCC25B8" lastPageId="40" lastPageNumber="41" pageId="37" pageNumber="38" type="description">
<paragraph id="8BDC36CDFF808413FF3459D4FF6E22F6" blockId="37.[151,1437,1517,2012]" pageId="37" pageNumber="38">
<emphasis id="B917EADFFF808413FF3459D4FC1B224A" bold="true" box="[199,345,1517,1543]" pageId="37" pageNumber="38">Description.</emphasis>
Worker (n = 10): HL: 
<geoCoordinate id="EE57500AFF808413FDA959D4FFDB2245" box="[602,665,1517,1544]" degrees="0.527" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="55" value="0.527">0.527</geoCoordinate>
<geoCoordinate id="EE57500AFF808413FD6B59D4FFA42245" box="[664,742,1517,1544]" degrees="0.686" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="55" value="-0.686">–0.686</geoCoordinate>
(0.635); HW: 
<geoCoordinate id="EE57500AFF808413FC7859D4FE882245" box="[907,970,1517,1544]" degrees="0.437" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="55" value="0.437">0.437</geoCoordinate>
<geoCoordinate id="EE57500AFF808413FC3959D4F9552245" box="[970,1047,1517,1544]" degrees="0.587" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="55" value="-0.587">–0.587</geoCoordinate>
(0.581); SL: 
<geoCoordinate id="EE57500AFF808413FB5C59D4F9AF2245" box="[1199,1261,1517,1544]" degrees="0.421" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="55" value="0.421">0.421</geoCoordinate>
<geoCoordinate id="EE57500AFF808413FB1E59D4F8792245" box="[1261,1339,1517,1544]" degrees="0.552" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="55" value="-0.552">–0.552</geoCoordinate>
(0.498); EL: 
<geoCoordinate id="EE57500AFF808413FF395A28FC452261" box="[202,263,1553,1580]" degrees="0.111" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="55" value="0.111">0.111</geoCoordinate>
<geoCoordinate id="EE57500AFF808413FEF45A28FC162261" box="[263,340,1553,1580]" degrees="0.158" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="55" value="-0.158">–0.158</geoCoordinate>
(0.143); EW: 
<geoCoordinate id="EE57500AFF808413FE0A5A28FF752266" box="[505,567,1553,1579]" degrees="0.087" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="55" value="0.087">0.087</geoCoordinate>
<geoCoordinate id="EE57500AFF808413FDC45A28FFC72266" box="[567,645,1553,1579]" degrees="0.124" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="55" value="-0.124">–0.124</geoCoordinate>
(0.111); PNW: 
<geoCoordinate id="EE57500AFF808413FCC85A28FE382266" box="[827,890,1553,1579]" degrees="0.294" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="55" value="0.294">0.294</geoCoordinate>
<geoCoordinate id="EE57500AFF808413FC895A28FE842266" box="[890,966,1553,1579]" degrees="0.398" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="55" value="-0.398">–0.398</geoCoordinate>
(0.370); WL: 
<geoCoordinate id="EE57500AFF808413FB9F5A28F9E92261" box="[1132,1195,1553,1580]" degrees="0.603" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="55" value="0.603">0.603</geoCoordinate>
<geoCoordinate id="EE57500AFF808413FB585A28F9BA2261" box="[1195,1272,1553,1580]" degrees="0.825" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="55" value="-0.825">–0.825</geoCoordinate>
(0.754); PEL: 
<geoCoordinate id="EE57500AFF808413FF645A0CFD942202" box="[151,214,1589,1615]" degrees="0.190" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="55" value="0.19">0.190</geoCoordinate>
<geoCoordinate id="EE57500AFF808413FF255A0CFC612202" box="[214,291,1589,1615]" degrees="0.302" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="55" value="-0.302">–0.302</geoCoordinate>
(0.258); PEH: 
<geoCoordinate id="EE57500AFF808413FE2E5A0CFF592202" box="[477,539,1589,1615]" degrees="0.143" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="55" value="0.143">0.143</geoCoordinate>
<geoCoordinate id="EE57500AFF808413FDE85A0CFF2A2202" box="[539,616,1589,1615]" degrees="0.222" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="55" value="-0.222">–0.222</geoCoordinate>
(0.207); PPL: 
<geoCoordinate id="EE57500AFF808413FCED5A0CFE1F2202" box="[798,861,1589,1615]" degrees="0.144" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="55" value="0.144">0.144</geoCoordinate>
<geoCoordinate id="EE57500AFF808413FCAE5A0CFEEB2202" box="[861,937,1589,1615]" degrees="0.198" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="55" value="-0.198">–0.198</geoCoordinate>
(0.185); PPW: 
<geoCoordinate id="EE57500AFF808413FB945A0CF9E42202" box="[1127,1190,1589,1615]" degrees="0.167" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="55" value="0.167">0.167</geoCoordinate>
<geoCoordinate id="EE57500AFF808413FB555A0CF9B12202" box="[1190,1267,1589,1615]" degrees="0.237" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="55" value="-0.237">–0.237</geoCoordinate>
(0.218); PSL: 
<geoCoordinate id="EE57500AFF808413FF645A60FD97223E" box="[151,213,1625,1651]" degrees="0.127" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="55" value="0.127">0.127</geoCoordinate>
<geoCoordinate id="EE57500AFF808413FF255A60FC61223E" box="[214,291,1625,1651]" degrees="0.192" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="55" value="-0.192">–0.192</geoCoordinate>
(0.168); CI: 
<geoCoordinate id="EE57500AFF808413FE4E5A60FCBE223E" box="[445,508,1625,1651]" degrees="1.149" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="55" value="1.149">1.149</geoCoordinate>
<geoCoordinate id="EE57500AFF808413FE0F5A60FF0A223E" box="[508,584,1625,1651]" degrees="1.231" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="55" value="-1.231">–1.231</geoCoordinate>
(1.190); EI1: 
<geoCoordinate id="EE57500AFF808413FD1D5A60FE6F223E" box="[750,813,1625,1651]" degrees="1.269" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="55" value="1.269">1.269</geoCoordinate>
<geoCoordinate id="EE57500AFF808413FCDE5A60FE38223E" box="[813,890,1625,1651]" degrees="1.377" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="55" value="-1.377">–1.377</geoCoordinate>
(1.293); EI2: 
<geoCoordinate id="EE57500AFF808413FBD35A60F91C223E" box="[1056,1118,1625,1651]" degrees="0.211" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="55" value="0.211">0.211</geoCoordinate>
<geoCoordinate id="EE57500AFF808413FBAD5A60F9E9223E" box="[1118,1195,1625,1651]" degrees="0.243" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="55" value="-0.243">–0.243</geoCoordinate>
(0.225); SI1: 
<geoCoordinate id="EE57500AFF808413FABC5A60F8CC223E" box="[1359,1422,1625,1651]" degrees="0.766" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="55" value="0.766">0.766</geoCoordinate>
<geoCoordinate id="EE57500AFF808413FA7D5A60FD9422DA" degrees="0.963" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="55" value="-0.963">– 0.963</geoCoordinate>
(0.933); SI2: 
<geoCoordinate id="EE57500AFF808413FE735A44FCFD22DA" box="[384,447,1661,1687]" degrees="0.766" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="55" value="0.766">0.766</geoCoordinate>
<geoCoordinate id="EE57500AFF808413FE4C5A44FF4E22DA" box="[447,524,1661,1687]" degrees="0.804" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="55" value="-0.804">–0.804</geoCoordinate>
(0.785); MI: 
<geoCoordinate id="EE57500AFF808413FD425A44FFAC22D5" box="[689,750,1661,1688]" degrees="1.955" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="55" value="1.955">1.955</geoCoordinate>
<geoCoordinate id="EE57500AFF808413FD1D5A44FE7F22D5" box="[750,829,1661,1688]" degrees="2.094" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="55" value="-2.094">–2.094</geoCoordinate>
(2.037); PI: 
<geoCoordinate id="EE57500AFF808413FC2A5A44F95A22DA" box="[985,1048,1661,1687]" degrees="1.072" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="55" value="1.072">1.072</geoCoordinate>
<geoCoordinate id="EE57500AFF808413FBEB5A44F92722DA" box="[1048,1125,1661,1687]" degrees="1.379" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="55" value="-1.379">–1.379</geoCoordinate>
(1.253); PPI: 
<geoCoordinate id="EE57500AFF808413FAE35A44F80C22DA" box="[1296,1358,1661,1687]" degrees="1.083" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="55" value="1.083">1.083</geoCoordinate>
<geoCoordinate id="EE57500AFF808413FABD5A44F8DE22DA" box="[1358,1436,1661,1687]" degrees="1.267" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="55" value="-1.267">–1.267</geoCoordinate>
(1.183); PSLI: 
<geoCoordinate id="EE57500AFF808413FEB35A98FC3D22F6" box="[320,383,1697,1723]" degrees="0.224" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="55" value="0.224">0.224</geoCoordinate>
<geoCoordinate id="EE57500AFF808413FE8C5A98FC8E22F6" box="[383,460,1697,1723]" degrees="0.329" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="55" value="-0.329">–0.329</geoCoordinate>
(0.297).
</paragraph>
<paragraph id="8BDC36CDFF808410FF345AFCF8DE2014" blockId="37.[151,1437,1517,2012]" lastBlockId="38.[151,1437,151,2013]" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
<emphasis id="B917EADFFF808413FF345AFCFC522292" bold="true" box="[199,272,1733,1759]" pageId="37" pageNumber="38">Color.</emphasis>
Almost whole body yellow, only first gastral tergite with ⅔ of space yellowish-brown band except yellow basal spot and posterior margin.Antennae and legs uniformly yellow, femora never infuscate in the middle (
<figureCitation id="13582A48FF808413FAB25AD0FDF5236A" captionStart="FIGURES 29, 30" captionStartId="35.[151,264,1981,2005]" captionTargetBox="[151,1436,417,1942]" captionTargetId="figure-98@35.[151,1436,417,1958]" captionTargetPageId="35" captionText="FIGURES 29, 30. Holotype worker of Temnothorax cerastarum n. sp. 29 dorsal 30 lateral (scale bar = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.10954807" httpUri="https://zenodo.org/record/10954807/files/figure.png" pageId="37" pageNumber="38">Figs 29, 30</figureCitation>
). 
<emphasis id="B917EADFFF808413FF385B34FC50236A" bold="true" box="[203,274,1805,1831]" pageId="37" pageNumber="38">Head.</emphasis>
Slightly elongate, 1.14–1.23 × as long as wide, sides almost parallel or softly converging behind eyes, occipital corners regularly rounded, occipital margin of head straight (
<figureCitation id="13582A48FF808413FC455B08F9492306" box="[950,1035,1841,1867]" captionStart="FIGURE 31" captionStartId="36.[151,250,1974,1998]" captionTargetBox="[151,1436,741,1951]" captionTargetId="figure-303@36.[151,1436,741,1951]" captionTargetPageId="36" captionText="FIGURE 31. Holotype worker head of Temnothorax cerastarum n. sp. (scale bar = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.10954809" httpUri="https://zenodo.org/record/10954809/files/figure.png" pageId="37" pageNumber="38">Fig. 31</figureCitation>
). Anterior margin of clypeus softly convex, without central angulation, with 4–6 short setae, medial notch absent. Eyes moderate, short oval, 1.3–1.4 × as long as wide, 0.23 × as long as head length. Antennal scape long, in lateral view slightly curved, approximately 0.9 × as long as width of the head, in apex gradually widened with well-marked preapical constriction, its base with angulate dorsal and ventral angle. Funiculus distinctly longer than scape, first segment 2.7 × as long as wide at apex, 2.6–2.7 × longer than second segment, segments 2–6 transverse, segment 7 only slightly wider than long, club large, 1.2 × as long as segments 1–8 combined, last segment of club elongate, 1.4 × as long as segments 8 and 9 combined (
<figureCitation id="13582A48FF838410FF6C5CE6FDB624B4" box="[159,244,223,249]" captionStart="FIGURE 31" captionStartId="36.[151,250,1974,1998]" captionTargetBox="[151,1436,741,1951]" captionTargetId="figure-303@36.[151,1436,741,1951]" captionTargetPageId="36" captionText="FIGURE 31. Holotype worker head of Temnothorax cerastarum n. sp. (scale bar = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.10954809" httpUri="https://zenodo.org/record/10954809/files/figure.png" pageId="38" pageNumber="39">Fig. 31</figureCitation>
). Surface of scape with fine microreticulation, shiny, covered with thin, moderate dense, appressed hairs. Mandibles with thick sparse, longitudinal striae, shiny, covered with short appressed hairs. Clypeus with median keel and two keels on each side, interspaces diffusely microreticulate, shiny. Frons narrow, approximately 0.42 × as wide as head width. Frontal carinae short, slightly extending beyond frontal lobes. Antennal fossa deep, margined with sharp circular striae with smooth interspaces. Frontal lobes narrow, placed only slightly upwards (
<figureCitation id="13582A48FF838410FACA5D56F8CD25C4" box="[1337,1423,367,393]" captionStart="FIGURE 31" captionStartId="36.[151,250,1974,1998]" captionTargetBox="[151,1436,741,1951]" captionTargetId="figure-303@36.[151,1436,741,1951]" captionTargetPageId="36" captionText="FIGURE 31. Holotype worker head of Temnothorax cerastarum n. sp. (scale bar = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.10954809" httpUri="https://zenodo.org/record/10954809/files/figure.png" pageId="38" pageNumber="39">Fig. 31</figureCitation>
). Gena, malar area and frons laterally with sharp longitudinal rugae, postocular area and occipital corners with fine striation, central part of frons, vertex and occipitum smooth and shiny, frons centrally without or with extremely small setose punctation. Surface of head without appressed pubescence, frons, vertex and occipital area with erect, pale, short and thick setae, the longest on occipitum 0.46 × as long as eye width (
<figureCitation id="13582A48FF838410FBC95DC6F9D22654" box="[1082,1168,511,537]" captionStart="FIGURES 29, 30" captionStartId="35.[151,264,1981,2005]" captionTargetBox="[151,1436,417,1942]" captionTargetId="figure-98@35.[151,1436,417,1958]" captionTargetPageId="35" captionText="FIGURES 29, 30. Holotype worker of Temnothorax cerastarum n. sp. 29 dorsal 30 lateral (scale bar = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.10954807" httpUri="https://zenodo.org/record/10954807/files/figure.png" pageId="38" pageNumber="39">Fig. 30</figureCitation>
). 
<emphasis id="B917EADFFF838410FB575DC6F86B2654" bold="true" box="[1188,1321,511,537]" pageId="38" pageNumber="39">Mesosoma.</emphasis>
Elongate, approximately twice as long as wide, slightly arched in anterior third, flat in posterior ⅔ length, without metanotal groove or impression. Pronotum convex on sides. Anterior slope of pronotum with transverse rugae, dorsum with fine longitudinal rugae and microreticulate interspaces, shiny. Sides of pronotum with sharp longitudinal rugae and diffusely microreticulate interspaces. Dorsum of mesonotum microreticulate, on sides or on whole surface with few fine longitudinal rugae, sides of mesonotum predominantly microgranulate but in upper ⅓ of space with more or less marked striation or rugae, occasionally also lower space with rudiments of rugae. Dorsum of propodeum centrally microreticulate, on sides with longitudinal rugae, posterior face microreticulate without or with few transverse rugae, sides of propodeum predominantly with sharp longitudinal rugae, especially on metapleural lobe, surface appears irregular but shiny. Propodeal spines long, mean PSL/HW 0.3, running strongly upwards, in form of elongate spines with slightly widened base, straight or with only slightly curved apex, sharply acute apically (
<figureCitation id="13582A48FF838410FF6C5FB2FDB627E8" box="[159,244,907,933]" captionStart="FIGURES 29, 30" captionStartId="35.[151,264,1981,2005]" captionTargetBox="[151,1436,417,1942]" captionTargetId="figure-98@35.[151,1436,417,1958]" captionTargetPageId="35" captionText="FIGURES 29, 30. Holotype worker of Temnothorax cerastarum n. sp. 29 dorsal 30 lateral (scale bar = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.10954807" httpUri="https://zenodo.org/record/10954807/files/figure.png" pageId="38" pageNumber="39">Fig. 30</figureCitation>
). Entire mesosoma bearing erect, yellow, moderately long setae, the longest on pronotum 0.6 × as long as eye diameter (
<figureCitation id="13582A48FF838410FEC55F96FCC92784" box="[310,395,943,969]" captionStart="FIGURES 29, 30" captionStartId="35.[151,264,1981,2005]" captionTargetBox="[151,1436,417,1942]" captionTargetId="figure-98@35.[151,1436,417,1958]" captionTargetPageId="35" captionText="FIGURES 29, 30. Holotype worker of Temnothorax cerastarum n. sp. 29 dorsal 30 lateral (scale bar = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.10954807" httpUri="https://zenodo.org/record/10954807/files/figure.png" pageId="38" pageNumber="39">Fig. 30</figureCitation>
). 
<emphasis id="B917EADFFF838410FE6D5F97FCB42785" bold="true" box="[414,502,942,968]" pageId="38" pageNumber="39">Petiole.</emphasis>
Moderately elongate, mean PEL/PEH 1.25, anterior face straight or very shallowly concave, ventral margin anteriorly with sharp spine, node in lateral view obtusely angulate with sharp lateral carinae, with few short rugae on top, whole surface microreticulate, shiny. 
<emphasis id="B917EADFFF838410FC675FCFF95C205D" bold="true" box="[916,1054,1014,1040]" pageId="38" pageNumber="39">Postpetiole.</emphasis>
In dorsal view almost as long as wide, approximately 1.3 × as wide as petiole, surface microreticulate with longitudinal striation, appears irregular but shiny. Dorsal surface of petiole and postpetiole with long erect setae, as long as setae on pronotum (
<figureCitation id="13582A48FF838410FAC95806F8D22014" box="[1338,1424,1087,1113]" captionStart="FIGURES 29, 30" captionStartId="35.[151,264,1981,2005]" captionTargetBox="[151,1436,417,1942]" captionTargetId="figure-98@35.[151,1436,417,1958]" captionTargetPageId="35" captionText="FIGURES 29, 30. Holotype worker of Temnothorax cerastarum n. sp. 29 dorsal 30 lateral (scale bar = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.10954807" httpUri="https://zenodo.org/record/10954807/files/figure.png" pageId="38" pageNumber="39">Fig. 30</figureCitation>
).
</paragraph>
<paragraph id="8BDC36CDFF838410FF64585BFE3F2088" blockId="38.[151,1437,151,2013]" pageId="38" pageNumber="39">
<emphasis id="B917EADFFF838410FF64585BFDAF2031" bold="true" box="[151,237,1122,1148]" pageId="38" pageNumber="39">Gaster.</emphasis>
Smooth and shiny, bearing erect, thin, pale setae, twice shorter than setae on mesosoma (
<figureCitation id="13582A48FF838410FB01585AF80A2030" box="[1266,1352,1123,1149]" captionStart="FIGURES 29, 30" captionStartId="35.[151,264,1981,2005]" captionTargetBox="[151,1436,417,1942]" captionTargetId="figure-98@35.[151,1436,417,1958]" captionTargetPageId="35" captionText="FIGURES 29, 30. Holotype worker of Temnothorax cerastarum n. sp. 29 dorsal 30 lateral (scale bar = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.10954807" httpUri="https://zenodo.org/record/10954807/files/figure.png" pageId="38" pageNumber="39">Fig. 29</figureCitation>
). 
<emphasis id="B917EADFFF838410FAAE585AF8DE2030" bold="true" box="[1373,1436,1123,1149]" pageId="38" pageNumber="39">Legs.</emphasis>
Moderately elongate, femora swollen in the middle, tibiae widened from base to ¾ length, surface of legs covered with extremely sparse, appressed hairs, appears smooth and shiny.
</paragraph>
<paragraph id="8BDC36CDFF838410FF3458F6FF2C217C" blockId="38.[151,1437,151,2013]" pageId="38" pageNumber="39">Gyne (n = 1): HL: 0.732; HW: 0.683; SL: 0.540; EL: 0.214; EW: 0.175; PNW: 0.849; WL: 1.373; PEL: 0.397; PEH: 0.273; PPL: 0.278; PPW: 0.317; PSL: 0.246; CI: 1.072; EI1: 1.223; EI2: 0.292; SI1: 0.791; SI2: 0.738; MI: 1.617; PI: 1.454; PPI: 1.140; PSLI: 0.360.</paragraph>
<paragraph id="8BDC36CDFF83841EFF345903FCFD25E0" blockId="38.[151,1437,151,2013]" lastBlockId="40.[151,1437,150,501]" lastPageId="40" lastPageNumber="41" pageId="38" pageNumber="39">
<emphasis id="B917EADFFF838410FF345903FC522119" bold="true" box="[199,272,1338,1364]" pageId="38" pageNumber="39">Color.</emphasis>
Head predominantly yellowish-brown, with yellow postocular area and ventral surface, mesosoma predominantly yellow, only scutellum brown centrally and yellowish-brown laterally, petiole and postpetiole yellow, gaster predominantly brown, first tergite with large, yellow basal spot, second tergite with yellow basal and posterior margins. Antennae uniformly yellow, legs yellow with infuscate femora (
<figureCitation id="13582A48FF838410FB99599EF9AF218C" box="[1130,1261,1447,1473]" captionStart="FIGURES 32, 33" captionStartId="37.[151,264,1447,1471]" captionTargetBox="[151,1436,186,1423]" captionTargetId="figure-18@37.[151,1436,181,1423]" captionTargetPageId="37" captionText="FIGURES 32, 33. Gyne of Temnothorax cerastarum n. sp. 32 dorsal 33 lateral (scale bar = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.10954811" httpUri="https://zenodo.org/record/10954811/files/figure.png" pageId="38" pageNumber="39">Figs 32, 33</figureCitation>
). 
<emphasis id="B917EADFFF838410FAF1599EF80B218C" bold="true" box="[1282,1353,1447,1473]" pageId="38" pageNumber="39">Head.</emphasis>
Broad, approximately 1.1 × as long as wide, widest behind eyes, gena parallel, behind eyes softly convergent, occipital corners rounded, occipital margin slightly convex (
<figureCitation id="13582A48FF838410FD1B59D6FE022244" box="[744,832,1519,1545]" captionStart="FIGURE 34" captionStartId="39.[151,250,1492,1516]" captionTargetBox="[151,1436,181,1468]" captionTargetId="figure-18@39.[151,1436,181,1468]" captionTargetPageId="39" captionText="FIGURE 34. Gyne head of Temnothorax cerastarum n. sp. (scale bar = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.10954813" httpUri="https://zenodo.org/record/10954813/files/figure.png" pageId="38" pageNumber="39">Fig. 34</figureCitation>
). Anterior margin of clypeus softly convex, without central angulation with minute median notch, with 2 long and 4 short setae. Eyes large, short oval, 1.2 × as long as wide, 0.23 × as long as head length. Antennal scape moderately long, in lateral view slightly curved, approximately 0.8 × as long as width of the head, in apex only slightly widened with very shallow preapical constriction, its basal corners obtusely angulate. Funiculus distinctly longer than scape, first segment 2.3 × as long as wide at apex, 2.3 × longer than second segment, segments 2–6 transverse, segment 7 only slightly wider than long, club large, almost as long as segments 1–8 combined, last segment of club elongate, 1.3 × as long as segments 8 and 9 combined (
<figureCitation id="13582A48FF838410FEE65AD2FC2B2348" box="[277,361,1771,1797]" captionStart="FIGURE 34" captionStartId="39.[151,250,1492,1516]" captionTargetBox="[151,1436,181,1468]" captionTargetId="figure-18@39.[151,1436,181,1468]" captionTargetPageId="39" captionText="FIGURE 34. Gyne head of Temnothorax cerastarum n. sp. (scale bar = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.10954813" httpUri="https://zenodo.org/record/10954813/files/figure.png" pageId="38" pageNumber="39">Fig. 34</figureCitation>
). Surface of scape with fine microreticulation, shiny, covered with thin, moderate dense, basally appressed apically subdecumbent hairs. Mandibles with thick, sparse, longitudinal striae, shiny, covered with short to moderate subdecumbent to decumbent hairs. Clypeus with median keel and two keels on each side, interspaces smooth, shiny. Frons narrow, approximately 0.35 × as wide as head width. Frontal carinae short, extending to ⅓ length of eyes. Antennal fossa deep, margined with sharp circular striae with smooth interspaces. Frontal lobes very narrow, placed only slightly upwards (
<figureCitation id="13582A48FF838410FDB95BA6FFE223F4" box="[586,672,1951,1977]" captionStart="FIGURE 34" captionStartId="39.[151,250,1492,1516]" captionTargetBox="[151,1436,181,1468]" captionTargetId="figure-18@39.[151,1436,181,1468]" captionTargetPageId="39" captionText="FIGURE 34. Gyne head of Temnothorax cerastarum n. sp. (scale bar = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.10954813" httpUri="https://zenodo.org/record/10954813/files/figure.png" pageId="38" pageNumber="39">Fig. 34</figureCitation>
). Whole surface of head with sharp longitudinal rugae extending to occipital margin, interspaces smooth or with diffused microreticulation, shiny, frons centrally without punctation. Surface of head without appressed pubescence, only sides with short decumbent hairs, frons, vertex and occipital area with erect, pale, short and thick setae, the longest on occipitum 0.4 × as long as eye width. 
<emphasis id="B917EADFFF828411FB545A07F86F2215" bold="true" box="[1191,1325,1598,1624]" pageId="39" pageNumber="40">Mesosoma.</emphasis>
Elongate, approximately 1.6 × as long as wide. Pronotum not visible from above, scutum and scutellum flat dorsally. Anterior slope of pronotum with granulate sculpture, sides with longitudinal rugae and diffusely microreticulate interspaces, appears shiny. Dorsum of scutum with longitudinal striation, slightly diffused on anterior slope and lateral lobes, interspaces smooth to diffusely microreticulate, shiny. Dorsum of scutellum predominantly smooth and shiny, only sides and close to basal margin with diffused striation (
<figureCitation id="13582A48FF828411FCBE5ACBFEE62341" box="[845,932,1778,1804]" captionStart="FIGURES 32, 33" captionStartId="37.[151,264,1447,1471]" captionTargetBox="[151,1436,186,1423]" captionTargetId="figure-18@37.[151,1436,181,1423]" captionTargetPageId="37" captionText="FIGURES 32, 33. Gyne of Temnothorax cerastarum n. sp. 32 dorsal 33 lateral (scale bar = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.10954811" httpUri="https://zenodo.org/record/10954811/files/figure.png" pageId="39" pageNumber="40">Fig. 32</figureCitation>
). Anepisternum anteriorly and dorsally with longitudinal striation, ventrally and posteriorly predominantly smooth, shiny, katepisternum predominantly smooth and shiny only close to posterior margin with short longitudinal rugae. Dorsum of propodeum with oblique and longitudinal rugae, between spines and posteriorly with diffused irregular rugae, shiny, sides of propodeum with sharp longitudinal rugae, interspaces smooth or diffusely microreticulate, shiny. Propodeal spines moderately long, mean PSL/HW 0.36, in form of triangulate spines with broadly widened base, sharply acute apically (
<figureCitation id="13582A48FF828411FACB5B9FF8CD238D" box="[1336,1423,1958,1984]" captionStart="FIGURES 32, 33" captionStartId="37.[151,264,1447,1471]" captionTargetBox="[151,1436,186,1423]" captionTargetId="figure-18@37.[151,1436,181,1423]" captionTargetPageId="37" captionText="FIGURES 32, 33. Gyne of Temnothorax cerastarum n. sp. 32 dorsal 33 lateral (scale bar = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.10954811" httpUri="https://zenodo.org/record/10954811/files/figure.png" pageId="39" pageNumber="40">Fig. 33</figureCitation>
). Entire mesosoma bearing erect, yellow, moderately long setae, the longest 0.4 × as long as eye diameter (
<figureCitation id="13582A48FF828411FAC85BF3F8CD23A9" box="[1339,1423,1994,2020]" captionStart="FIGURES 32, 33" captionStartId="37.[151,264,1447,1471]" captionTargetBox="[151,1436,186,1423]" captionTargetId="figure-18@37.[151,1436,181,1423]" captionTargetPageId="37" captionText="FIGURES 32, 33. Gyne of Temnothorax cerastarum n. sp. 32 dorsal 33 lateral (scale bar = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.10954811" httpUri="https://zenodo.org/record/10954811/files/figure.png" pageId="39" pageNumber="40">Fig. 33</figureCitation>
). 
<emphasis id="B917EADFFF8D841EFF645CAFFDAD24FD" bold="true" box="[151,239,150,176]" pageId="40" pageNumber="41">Petiole.</emphasis>
Elongate, mean PEL/PEH 1.45, anterior face straight, ventral margin anteriorly with small, sharp spine, node in lateral view sharply angulate, anterior slope smooth to diffusely microreticulate, sides with longitudinal rugae, ventral part with microreticulation but surface appears shiny. 
<emphasis id="B917EADFFF8D841EFC6E5CE7F96524B5" bold="true" box="[925,1063,222,248]" pageId="40" pageNumber="41">Postpetiole.</emphasis>
Transverse, approximately 1.3 × as wide as petiole, surface microreticulate with longitudinal striation, appears shiny. Dorsal surface of petiole and postpetiole with erect setae, slightly longer than setae on pronotum (
<figureCitation id="13582A48FF8D841EFC575D1EFEBB250C" box="[932,1017,295,321]" captionStart="FIGURES 32, 33" captionStartId="37.[151,264,1447,1471]" captionTargetBox="[151,1436,186,1423]" captionTargetId="figure-18@37.[151,1436,181,1423]" captionTargetPageId="37" captionText="FIGURES 32, 33. Gyne of Temnothorax cerastarum n. sp. 32 dorsal 33 lateral (scale bar = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.10954811" httpUri="https://zenodo.org/record/10954811/files/figure.png" pageId="40" pageNumber="41">Fig. 33</figureCitation>
). 
<emphasis id="B917EADFFF8D841EFBFC5D1FF927250D" bold="true" box="[1039,1125,294,320]" pageId="40" pageNumber="41">Gaster.</emphasis>
Smooth and shiny, bearing erect, thin, pale setae, ⅓ shorter than setae on mesosoma (
<figureCitation id="13582A48FF8D841EFCD55D72FEC22528" box="[806,896,331,357]" captionStart="FIGURES 32, 33" captionStartId="37.[151,264,1447,1471]" captionTargetBox="[151,1436,186,1423]" captionTargetId="figure-18@37.[151,1436,181,1423]" captionTargetPageId="37" captionText="FIGURES 32, 33. Gyne of Temnothorax cerastarum n. sp. 32 dorsal 33 lateral (scale bar = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.10954811" httpUri="https://zenodo.org/record/10954811/files/figure.png" pageId="40" pageNumber="41">Figs 32</figureCitation>
). 
<emphasis id="B917EADFFF8D841EFC675D72FE912528" bold="true" box="[916,979,331,357]" pageId="40" pageNumber="41">Legs.</emphasis>
Moderately elongate, femora swollen in the middle, tibiae widened from base to ¾ length, surface of legs covered with extremely sparse, appressed hairs, appears smooth and shiny.
</paragraph>
<caption id="DF1C6645FF828411FF6459EDFE9121A1" ID-DOI="http://doi.org/10.5281/zenodo.10954813" ID-Zenodo-Dep="10954813" box="[151,979,1492,1516]" httpUri="https://zenodo.org/record/10954813/files/figure.png" pageId="39" pageNumber="40" startId="39.[151,250,1492,1516]" targetBox="[151,1436,181,1468]" targetPageId="39" targetType="figure">
<paragraph id="8BDC36CDFF828411FF6459EDFE9121A1" blockId="39.[151,979,1492,1516]" box="[151,979,1492,1516]" pageId="39" pageNumber="40">
<emphasis id="B917EADFFF828411FF6459EDFC6221A1" bold="true" box="[151,288,1492,1516]" pageId="39" pageNumber="40">FIGURE 34.</emphasis>
Gyne head of 
<taxonomicName id="4C634D4EFF828411FE4659EDFFF121A1" authority="Salata &amp; Demetriou &amp; Georgiadis &amp; Borowiec, 2024" authorityName="Salata &amp; Demetriou &amp; Georgiadis &amp; Borowiec" authorityYear="2024" box="[437,691,1492,1516]" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="cerastarum" status="sp. nov.">
<emphasis id="B917EADFFF828411FE4659EDFFF121A1" box="[437,691,1492,1516]" italics="true" pageId="39" pageNumber="40">Temnothorax cerastarum</emphasis>
</taxonomicName>
<emphasis id="B917EADFFF828411FD4959EDFFB121A1" bold="true" box="[698,755,1492,1516]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="A22457A4FF828411FD4959EDFFB121A1" box="[698,755,1492,1516]" pageId="39" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(scale bar = 0.5 mm).
</paragraph>
</caption>
<paragraph id="8BDC36CDFF8D841EFF345D8FFFCC25B8" blockId="40.[151,1437,150,501]" pageId="40" pageNumber="41">
<emphasis id="B917EADFFF8D841EFF345D8FFCC0259C" bold="true" box="[199,386,438,465]" pageId="40" pageNumber="41">Biological note.</emphasis>
A single nest was observed under a moderately large stone inside a mixed pine and cedar forest, in mountains at an altitude of 
<quantity id="4C9B9B28FF8D841EFDC05DE2FFC825B8" box="[563,650,475,501]" metricMagnitude="3" metricUnit="m" metricValue="1.138" pageId="40" pageNumber="41" unit="m" value="1138.0">1138 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>