treatments-xml/data/03/A9/65/03A9654BFFA3077B536AFA1E78E6F7B5.xml

<document id="F2A0ABE3233CCF19F992767CD566E14E" ID-DOI="10.5281/zenodo.5384590" ID-ISSN="2345-7600" ID-Zenodo-Dep="5384590" ID-ZooBank="40BCDD62-D35E-46D1-95A3-2CC0DF219DEE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630299679004" checkinUser="valdenar" docAuthor="Ng, Peter K. L. &amp; Forges, Bertrand Richer De" docDate="2015" docId="03A9654BFFA3077B536AFA1E78E6F7B5" docLanguage="en" docName="63rbz110-225.pdf" docOrigin="Raffles Bulletin of Zoology 63" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Alcomaja Ng &amp; Forges 2015, n. gen." docType="treatment" docUuid="C3EE8C3E-4F10-433C-B1FA-974B8F3E563E" docUuidSource="ZooBank" docVersion="4" lastPageNumber="145" masterDocId="FF901D33FF8107585767FF837962FFA3" masterDocTitle="Revision of the spider crab genus Maja Lamarck, 1801 (Crustacea: Brachyura: Majoidea: Majidae), with descriptions of seven new genera and 17 new species from the Atlantic and Indo-West Pacific" masterLastPageNumber="225" masterPageNumber="110" pageNumber="144" updateTime="1699220132769" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="5BD2921C4700A462C5F467F3BA572059">Revision of the spider crab genus Maja Lamarck, 1801 (Crustacea: Brachyura: Majoidea: Majidae), with descriptions of seven new genera and 17 new species from the Atlantic and Indo-West Pacific</mods:title>
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<mods:namePart id="02A3A2043C08C79629CE70F907D6CABE">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="DB0BE23ADDE9F864C061157A1CBA2C49">Forges, Bertrand Richer De</mods:namePart>
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<taxonomicName id="4C00AFDEFFA3077A536AFA1E7D14FA16" ID-CoL="7NNHK" authority="Ng &amp; Forges, 2015" authorityName="Ng &amp; Forges" authorityYear="2015" box="[1037,1142,1437,1461]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="144" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B974084FFFA3077A536AFA1E7D14FA16" bold="true" box="[1037,1142,1437,1461]" italics="true" pageId="34" pageNumber="144">Alcomaja</emphasis>
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<taxonomicNameLabel id="A247B534FFA3077A5319FA1E7DA9FA16" box="[1150,1227,1437,1461]" pageId="34" pageNumber="144" rank="genus">n. gen.</taxonomicNameLabel>
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<paragraph id="8BBFD45DFFA3077B5448FA5E7A60FA76" blockId="34.[811,1454,1501,2101]" lastBlockId="35.[134,776,189,1493]" lastPageId="35" lastPageNumber="145" pageId="34" pageNumber="144">
<emphasis id="B974084FFFA3077A5448FA5E7AC1FA56" bold="true" box="[815,931,1501,1525]" pageId="34" pageNumber="144">Diagnosis.</emphasis>
Carapace ovate; dorsal surface convex, covered by granules or tubercles; gastric and branchial regions delimited by distinct grooves (
<figureCitation id="133BC8D8FFA3077A530AF99E7DA7F996" box="[1133,1221,1565,1589]" captionStart="Fig" captionStartId="39.[136,171,1902,1924]" captionTargetBox="[136,1474,291,1887]" captionTargetId="figure-8@39.[136,1474,291,1887]" captionTargetPageId="39" captionText="Fig. 16. General habitus, Alcomaja species. A, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; B, A. gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; C, A. gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; D, A. confragosa, female (21.5 × 18.2 mm) (NMV J53978), Australia; E, A. confragosa, holotype male (as Maja confragosa, after Griffin &amp; Tranter 1986: pl. 16a)." figureDoi="http://doi.org/10.5281/zenodo.5384628" httpUri="https://zenodo.org/record/5384628/files/figure.png" pageId="34" pageNumber="144">Figs. 16</figureCitation>
, 
<figureCitation id="133BC8D8FFA3077A53A8F99E7D8EF996" box="[1231,1260,1565,1589]" captionStart="Fig" captionStartId="40.[113,149,1934,1956]" captionTargetBox="[113,1451,257,1918]" captionTargetId="figure-15@40.[113,1451,257,1920]" captionTargetPageId="40" captionText="Fig. 17. General habitus, Alcomaja species. A, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; B, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM-ZC9928), Japan; C, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Japan; D, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; E, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; F, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384630" httpUri="https://zenodo.org/record/5384630/files/figure.png" pageId="34" pageNumber="144">17</figureCitation>
). Intestinal region without distinct median spine (
<figureCitation id="133BC8D8FFA3077A531AF9BE7DBBF9F6" box="[1149,1241,1597,1621]" captionStart="Fig" captionStartId="39.[136,171,1902,1924]" captionTargetBox="[136,1474,291,1887]" captionTargetId="figure-8@39.[136,1474,291,1887]" captionTargetPageId="39" captionText="Fig. 16. General habitus, Alcomaja species. A, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; B, A. gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; C, A. gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; D, A. confragosa, female (21.5 × 18.2 mm) (NMV J53978), Australia; E, A. confragosa, holotype male (as Maja confragosa, after Griffin &amp; Tranter 1986: pl. 16a)." figureDoi="http://doi.org/10.5281/zenodo.5384628" httpUri="https://zenodo.org/record/5384628/files/figure.png" pageId="34" pageNumber="144">Figs. 16</figureCitation>
, 
<figureCitation id="133BC8D8FFA3077A5381F9BE7C61F9F6" box="[1254,1283,1597,1621]" captionStart="Fig" captionStartId="40.[113,149,1934,1956]" captionTargetBox="[113,1451,257,1918]" captionTargetId="figure-15@40.[113,1451,257,1920]" captionTargetPageId="40" captionText="Fig. 17. General habitus, Alcomaja species. A, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; B, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM-ZC9928), Japan; C, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Japan; D, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; E, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; F, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384630" httpUri="https://zenodo.org/record/5384630/files/figure.png" pageId="34" pageNumber="144">17</figureCitation>
). Pseudorostral spines long, diverging, forming a V, (
<figureCitation id="133BC8D8FFA3077A5385F9DE7C23F9D6" box="[1250,1345,1629,1653]" captionStart="Fig" captionStartId="41.[136,171,1906,1928]" captionTargetBox="[136,1474,257,1891]" captionTargetId="figure-8@41.[136,1474,257,1891]" captionTargetPageId="41" captionText="Fig. 18. Frontal regions of carapace showing different structures of intercalated teeth, Alcomaja species. A, A. confragosa, holotype male (as Maja confragosa, after Griffin &amp; Tranter, 1986: pl. 16a); B, A. confragosa, female (21.5 × 18.2 mm) (NMV J53978), Australia; C, A. gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; D, A. gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; E, A. irrorata n. sp., paratype male (44.7 × 39.1 mm) (ZRC 2013.1229), Philippines; F, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384632" httpUri="https://zenodo.org/record/5384632/files/figure.png" pageId="34" pageNumber="144">Figs. 18</figureCitation>
, 
<figureCitation id="133BC8D8FFA3077A5236F9DE7CC0F9D6" box="[1361,1442,1629,1653]" captionStart="Fig" captionStartId="72.[113,148,1742,1764]" captionTargetBox="[113,1451,281,1727]" captionTargetId="figure-15@72.[113,1451,281,1728]" captionTargetPageId="72" captionText="Fig. 36. Frontal regions of carapaces. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja brachydactyla, male (161.2 × 140.1 mm) (ZRC 2008.0179), Spain; D, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; E, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; F, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; G, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; H, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; I, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; J, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; K, Paramaja turgida n. sp., paratype male (67.9 × 60.5 mm) (MNHN), Philippines; L, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; M, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; N, Alcomaja latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; O, Alcomaja miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar." figureDoi="http://doi.org/10.5281/zenodo.5384670" httpUri="https://zenodo.org/record/5384670/files/figure.png" pageId="34" pageNumber="144">36L–O</figureCitation>
). Supraorbital eave with anterior part longitudinally narrow, rectangular, not prominently expanded; antorbital spine sharp (
<figureCitation id="133BC8D8FFA3077A5454F93E7AF1F976" box="[819,915,1725,1749]" captionStart="Fig" captionStartId="41.[136,171,1906,1928]" captionTargetBox="[136,1474,257,1891]" captionTargetId="figure-8@41.[136,1474,257,1891]" captionTargetPageId="41" captionText="Fig. 18. Frontal regions of carapace showing different structures of intercalated teeth, Alcomaja species. A, A. confragosa, holotype male (as Maja confragosa, after Griffin &amp; Tranter, 1986: pl. 16a); B, A. confragosa, female (21.5 × 18.2 mm) (NMV J53978), Australia; C, A. gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; D, A. gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; E, A. irrorata n. sp., paratype male (44.7 × 39.1 mm) (ZRC 2013.1229), Philippines; F, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384632" httpUri="https://zenodo.org/record/5384632/files/figure.png" pageId="34" pageNumber="144">Figs. 18</figureCitation>
, 
<figureCitation id="133BC8D8FFA3077A54C4F93E7A97F976" box="[931,1013,1725,1749]" captionStart="Fig" captionStartId="72.[113,148,1742,1764]" captionTargetBox="[113,1451,281,1727]" captionTargetId="figure-15@72.[113,1451,281,1728]" captionTargetPageId="72" captionText="Fig. 36. Frontal regions of carapaces. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja brachydactyla, male (161.2 × 140.1 mm) (ZRC 2008.0179), Spain; D, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; E, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; F, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; G, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; H, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; I, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; J, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; K, Paramaja turgida n. sp., paratype male (67.9 × 60.5 mm) (MNHN), Philippines; L, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; M, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; N, Alcomaja latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; O, Alcomaja miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar." figureDoi="http://doi.org/10.5281/zenodo.5384670" httpUri="https://zenodo.org/record/5384670/files/figure.png" pageId="34" pageNumber="144">36L–O</figureCitation>
). Intercalated spine distinct, separated from supraorbital eave and postorbital spine by distinct gaps; postorbital spine strong, lobiform; hepatic region with 1 strong spine, much shorter than postorbital spine; 1 or more smaller spines below (
<figureCitation id="133BC8D8FFA3077A53F3F8BE7D8DF8F6" box="[1172,1263,1853,1877]" captionStart="Fig" captionStartId="41.[136,171,1906,1928]" captionTargetBox="[136,1474,257,1891]" captionTargetId="figure-8@41.[136,1474,257,1891]" captionTargetPageId="41" captionText="Fig. 18. Frontal regions of carapace showing different structures of intercalated teeth, Alcomaja species. A, A. confragosa, holotype male (as Maja confragosa, after Griffin &amp; Tranter, 1986: pl. 16a); B, A. confragosa, female (21.5 × 18.2 mm) (NMV J53978), Australia; C, A. gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; D, A. gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; E, A. irrorata n. sp., paratype male (44.7 × 39.1 mm) (ZRC 2013.1229), Philippines; F, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384632" httpUri="https://zenodo.org/record/5384632/files/figure.png" pageId="34" pageNumber="144">Figs. 18</figureCitation>
, 
<figureCitation id="133BC8D8FFA3077A539BF8BE7C2FF8F6" box="[1276,1357,1853,1877]" captionStart="Fig" captionStartId="72.[113,148,1742,1764]" captionTargetBox="[113,1451,281,1727]" captionTargetId="figure-15@72.[113,1451,281,1728]" captionTargetPageId="72" captionText="Fig. 36. Frontal regions of carapaces. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja brachydactyla, male (161.2 × 140.1 mm) (ZRC 2008.0179), Spain; D, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; E, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; F, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; G, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; H, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; I, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; J, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; K, Paramaja turgida n. sp., paratype male (67.9 × 60.5 mm) (MNHN), Philippines; L, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; M, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; N, Alcomaja latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; O, Alcomaja miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar." figureDoi="http://doi.org/10.5281/zenodo.5384670" httpUri="https://zenodo.org/record/5384670/files/figure.png" pageId="34" pageNumber="144">36L–O</figureCitation>
). Lateral carapace margin with 3 spines and numerous tubercles or granules, 1 branchial spine or tubercle usually discernible, may be low in large specimens (
<figureCitation id="133BC8D8FFA3077A53C4F81E7C60F816" box="[1187,1282,1949,1973]" captionStart="Fig" captionStartId="39.[136,171,1902,1924]" captionTargetBox="[136,1474,291,1887]" captionTargetId="figure-8@39.[136,1474,291,1887]" captionTargetPageId="39" captionText="Fig. 16. General habitus, Alcomaja species. A, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; B, A. gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; C, A. gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; D, A. confragosa, female (21.5 × 18.2 mm) (NMV J53978), Australia; E, A. confragosa, holotype male (as Maja confragosa, after Griffin &amp; Tranter 1986: pl. 16a)." figureDoi="http://doi.org/10.5281/zenodo.5384628" httpUri="https://zenodo.org/record/5384628/files/figure.png" pageId="34" pageNumber="144">Figs. 16</figureCitation>
, 
<figureCitation id="133BC8D8FFA3077A5275F81E7C52F816" box="[1298,1328,1949,1973]" captionStart="Fig" captionStartId="40.[113,149,1934,1956]" captionTargetBox="[113,1451,257,1918]" captionTargetId="figure-15@40.[113,1451,257,1920]" captionTargetPageId="40" captionText="Fig. 17. General habitus, Alcomaja species. A, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; B, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM-ZC9928), Japan; C, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Japan; D, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; E, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; F, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384630" httpUri="https://zenodo.org/record/5384630/files/figure.png" pageId="34" pageNumber="144">17</figureCitation>
). Posterior carapace margin with 2 short median spines (
<figureCitation id="133BC8D8FFA3077A527DF83E7C17F876" box="[1306,1397,1981,2005]" captionStart="Fig" captionStartId="39.[136,171,1902,1924]" captionTargetBox="[136,1474,291,1887]" captionTargetId="figure-8@39.[136,1474,291,1887]" captionTargetPageId="39" captionText="Fig. 16. General habitus, Alcomaja species. A, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; B, A. gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; C, A. gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; D, A. confragosa, female (21.5 × 18.2 mm) (NMV J53978), Australia; E, A. confragosa, holotype male (as Maja confragosa, after Griffin &amp; Tranter 1986: pl. 16a)." figureDoi="http://doi.org/10.5281/zenodo.5384628" httpUri="https://zenodo.org/record/5384628/files/figure.png" pageId="34" pageNumber="144">Figs. 16</figureCitation>
, 
<figureCitation id="133BC8D8FFA3077A52E6F83E7CFCF876" box="[1409,1438,1981,2005]" captionStart="Fig" captionStartId="40.[113,149,1934,1956]" captionTargetBox="[113,1451,257,1918]" captionTargetId="figure-15@40.[113,1451,257,1920]" captionTargetPageId="40" captionText="Fig. 17. General habitus, Alcomaja species. A, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; B, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM-ZC9928), Japan; C, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Japan; D, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; E, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; F, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384630" httpUri="https://zenodo.org/record/5384630/files/figure.png" pageId="34" pageNumber="144">17</figureCitation>
). Eyes relatively long, slender, with ovoid cornea (
<figureCitation id="133BC8D8FFA3077A522AF85E7CCBF856" box="[1357,1449,2013,2037]" captionStart="Fig" captionStartId="41.[136,171,1906,1928]" captionTargetBox="[136,1474,257,1891]" captionTargetId="figure-8@41.[136,1474,257,1891]" captionTargetPageId="41" captionText="Fig. 18. Frontal regions of carapace showing different structures of intercalated teeth, Alcomaja species. A, A. confragosa, holotype male (as Maja confragosa, after Griffin &amp; Tranter, 1986: pl. 16a); B, A. confragosa, female (21.5 × 18.2 mm) (NMV J53978), Australia; C, A. gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; D, A. gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; E, A. irrorata n. sp., paratype male (44.7 × 39.1 mm) (ZRC 2013.1229), Philippines; F, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384632" httpUri="https://zenodo.org/record/5384632/files/figure.png" pageId="34" pageNumber="144">Figs. 18</figureCitation>
, 
<figureCitation id="133BC8D8FFA3077A544AF87E7A1FF7B6" box="[813,893,2045,2069]" captionStart="Fig" captionStartId="72.[113,148,1742,1764]" captionTargetBox="[113,1451,281,1727]" captionTargetId="figure-15@72.[113,1451,281,1728]" captionTargetPageId="72" captionText="Fig. 36. Frontal regions of carapaces. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja brachydactyla, male (161.2 × 140.1 mm) (ZRC 2008.0179), Spain; D, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; E, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; F, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; G, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; H, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; I, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; J, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; K, Paramaja turgida n. sp., paratype male (67.9 × 60.5 mm) (MNHN), Philippines; L, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; M, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; N, Alcomaja latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; O, Alcomaja miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar." figureDoi="http://doi.org/10.5281/zenodo.5384670" httpUri="https://zenodo.org/record/5384670/files/figure.png" pageId="34" pageNumber="144">36L–O</figureCitation>
). Antennal flagellum short, slender. Basal antennal article longer than broad, rectangular; surface with several tubercles or granules, with 2 spines distally; inner and outer lateral margins may have low granules or lobes, or entire; proximal outer angle rounded; antero-external crested rim of antennular fossa touches does not substantially overlap basal antennal article (
<figureCitation id="133BC8D8FFA2077B5625FEBE78ABFEF6" box="[322,457,317,341]" captionStart="Fig" captionStartId="75.[136,172,1760,1782]" captionTargetBox="[138,1471,262,1745]" captionTargetId="figure-8@75.[138,1471,261,1745]" captionTargetPageId="75" captionText="Fig. 39. Antennae, antennules and epistomes. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; D, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; E, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN- IU-2013-4046), Congo; F, Neomaja goltziana, male (90.2 × 83.1 mm) (MNHN-IU-2013-4044), Gabon; G, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; H, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; I, Paramaja gibba, lectotype female (25.1 × 21.2 mm) (NHM 1896.5.14.9), India; J, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; K, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; L, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; M, Alcomaja gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; N, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; O, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384676" httpUri="https://zenodo.org/record/5384676/files/figure.png" pageId="35" pageNumber="145">Fig. 39L–O</figureCitation>
). Epistome as wide as long, anterior margin with low lobes or low swellings; posterior margin composed of 4 rectangular plates separated by shallow fissures (
<figureCitation id="133BC8D8FFA2077B5784FE1E780EFE16" box="[227,364,413,437]" captionStart="Fig" captionStartId="75.[136,172,1760,1782]" captionTargetBox="[138,1471,262,1745]" captionTargetId="figure-8@75.[138,1471,261,1745]" captionTargetPageId="75" captionText="Fig. 39. Antennae, antennules and epistomes. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; D, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; E, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN- IU-2013-4046), Congo; F, Neomaja goltziana, male (90.2 × 83.1 mm) (MNHN-IU-2013-4044), Gabon; G, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; H, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; I, Paramaja gibba, lectotype female (25.1 × 21.2 mm) (NHM 1896.5.14.9), India; J, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; K, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; L, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; M, Alcomaja gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; N, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; O, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384676" httpUri="https://zenodo.org/record/5384676/files/figure.png" pageId="35" pageNumber="145">Figs. 39L–O</figureCitation>
, 
<figureCitation id="133BC8D8FFA2077B5610FE1E78AAFE16" box="[375,456,413,437]" captionStart="Fig" captionStartId="77.[136,171,1977,1999]" captionTargetBox="[136,1475,673,1962]" captionTargetId="figure-8@77.[135,1475,673,1962]" captionTargetPageId="77" captionText="Fig. 41. Epistomes. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; D, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; E, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; F, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; G, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; H, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384680" httpUri="https://zenodo.org/record/5384680/files/figure.png" pageId="35" pageNumber="145">41G, H</figureCitation>
). Suborbital margin separated from basal antennal article by short fissure, confluent with margin of postorbital tooth (
<figureCitation id="133BC8D8FFA2077B56AAFE5E7B34FE56" box="[461,598,477,501]" captionStart="Fig" captionStartId="75.[136,172,1760,1782]" captionTargetBox="[138,1471,262,1745]" captionTargetId="figure-8@75.[138,1471,261,1745]" captionTargetPageId="75" captionText="Fig. 39. Antennae, antennules and epistomes. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; D, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; E, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN- IU-2013-4046), Congo; F, Neomaja goltziana, male (90.2 × 83.1 mm) (MNHN-IU-2013-4044), Gabon; G, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; H, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; I, Paramaja gibba, lectotype female (25.1 × 21.2 mm) (NHM 1896.5.14.9), India; J, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; K, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; L, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; M, Alcomaja gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; N, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; O, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384676" httpUri="https://zenodo.org/record/5384676/files/figure.png" pageId="35" pageNumber="145">Fig. 39L–O</figureCitation>
). Outer surface of third maxilliped covered by very short setae in adults; ischium subrectangular, distinctly longer than broad; posteroexternal angle of merus relatively broad, “inserted” into shallower concavity on outer margin of ischium; anterointernal part of ischium rounded, auriculiform (
<figureCitation id="133BC8D8FFA2077B551FFDFE7B99FD36" box="[632,763,637,661]" captionStart="Fig" captionStartId="81.[136,171,1723,1745]" captionTargetBox="[137,1475,328,1708]" captionTargetId="figure-8@81.[136,1475,328,1708]" captionTargetPageId="81" captionText="Fig. 45. Third maxillipeds. A, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; B, Alcomaja gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea [third max laterally inverted to match]; C, Alcomaja gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; D, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; E, Alcomaja nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; F, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; G, Alcomaja latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; H, Alcomaja miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; I, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; J, Paramaya ouch n. sp., holotype male (76.8 × 60.0 mm) (NMCR), Philippines; K, Paramaya ouch n. sp., paratype male (39.7 × 29.6 mm) (ZRC 2011.0045), Philippines; L, Paramaya coccinea n. sp., holotype male (69.0 × 55.6 mm) (MNHN), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.5384688" httpUri="https://zenodo.org/record/5384688/files/figure.png" pageId="35" pageNumber="145">Fig. 45A–H</figureCitation>
). Male chelipeds relatively short in adult males, surfaces of merus and carpus almost smooth to covered with very small granules, low longitudinal ridges with low tubercles; carpus short, with low, uneven longitudinal ridge; propodus of palm elongated, curved, smooth, without lateral cristae in young, short, inflated in adults, palm longer than fingers; fingers long, slender, gently curved, with distinct basal gape when closed (
<figureCitation id="133BC8D8FFA2077B5787FCFE785FFC36" box="[224,317,893,917]" captionStart="Fig" captionStartId="39.[136,171,1902,1924]" captionTargetBox="[136,1474,291,1887]" captionTargetId="figure-8@39.[136,1474,291,1887]" captionTargetPageId="39" captionText="Fig. 16. General habitus, Alcomaja species. A, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; B, A. gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; C, A. gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; D, A. confragosa, female (21.5 × 18.2 mm) (NMV J53978), Australia; E, A. confragosa, holotype male (as Maja confragosa, after Griffin &amp; Tranter 1986: pl. 16a)." figureDoi="http://doi.org/10.5281/zenodo.5384628" httpUri="https://zenodo.org/record/5384628/files/figure.png" pageId="35" pageNumber="145">Figs. 16</figureCitation>
, 
<figureCitation id="133BC8D8FFA2077B562BFCFE780BFC36" box="[332,361,893,917]" captionStart="Fig" captionStartId="40.[113,149,1934,1956]" captionTargetBox="[113,1451,257,1918]" captionTargetId="figure-15@40.[113,1451,257,1920]" captionTargetPageId="40" captionText="Fig. 17. General habitus, Alcomaja species. A, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; B, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM-ZC9928), Japan; C, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Japan; D, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; E, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; F, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384630" httpUri="https://zenodo.org/record/5384630/files/figure.png" pageId="35" pageNumber="145">17</figureCitation>
, 
<figureCitation id="133BC8D8FFA2077B5610FCFE78A9FC36" box="[375,459,893,917]" captionStart="Fig" captionStartId="89.[135,170,1806,1828]" captionTargetBox="[136,1474,216,1791]" captionTargetId="figure-8@89.[136,1474,215,1791]" captionTargetPageId="89" captionText="Fig. 53. Chelae. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja brachydactyla, male (161.2 × 140.1 mm), (ZRC 2008.0179), Spain; D, Maja cornuta, male (88.8 × 86.8 mm) (NHM 1928.12.1.177–) (lectotype of Mamaia queketti Stebbing, 1908), South Africa; E, Maja cornuta, male (115.3 × 103.4 mm) (ZRC 2013.1184), South Africa; F, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; G, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; H, Neomaja goltziana, male (90.2 × 83.1 mm) (MNHN-IU-2013-4044), Gabon; I, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; J, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; K, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; L, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; M, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; N, Alcomaja gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; O, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; P, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384708" httpUri="https://zenodo.org/record/5384708/files/figure.png" pageId="35" pageNumber="145">53M–P</figureCitation>
). Ambulatory legs relatively long, slender; merus without dorsal subdistal spine; dactylus elongate, curved, covered with long setae when young except for corneous tip, almost completely smooth in adults (
<figureCitation id="133BC8D8FFA2077B57E8FC7E7988FBB6" box="[143,234,1021,1045]" captionStart="Fig" captionStartId="39.[136,171,1902,1924]" captionTargetBox="[136,1474,291,1887]" captionTargetId="figure-8@39.[136,1474,291,1887]" captionTargetPageId="39" captionText="Fig. 16. General habitus, Alcomaja species. A, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; B, A. gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; C, A. gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; D, A. confragosa, female (21.5 × 18.2 mm) (NMV J53978), Australia; E, A. confragosa, holotype male (as Maja confragosa, after Griffin &amp; Tranter 1986: pl. 16a)." figureDoi="http://doi.org/10.5281/zenodo.5384628" httpUri="https://zenodo.org/record/5384628/files/figure.png" pageId="35" pageNumber="145">Figs. 16</figureCitation>
, 
<figureCitation id="133BC8D8FFA2077B5790FC7E7876FBB6" box="[247,276,1021,1045]" captionStart="Fig" captionStartId="40.[113,149,1934,1956]" captionTargetBox="[113,1451,257,1918]" captionTargetId="figure-15@40.[113,1451,257,1920]" captionTargetPageId="40" captionText="Fig. 17. General habitus, Alcomaja species. A, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; B, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM-ZC9928), Japan; C, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Japan; D, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; E, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; F, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384630" httpUri="https://zenodo.org/record/5384630/files/figure.png" pageId="35" pageNumber="145">17</figureCitation>
, 
<figureCitation id="133BC8D8FFA2077B5646FC7E7816FBB6" box="[289,372,1021,1045]" captionStart="Fig" captionStartId="91.[135,171,1862,1884]" captionTargetBox="[170,1440,189,1847]" captionTargetId="figure-8@91.[170,1440,189,1847]" captionTargetPageId="91" captionText="Fig. 55. First ambulatory dactyli and propodi. A, Maja squinado, neotype male (147.1 × 126.3 mm) (SMF-4548), Croatia; B, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; C, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; D, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; E, Paramaja kominatoensis, dried male (62.6 × 56.1 mm) (KPM NH0104298), Japan; F, Paramaja kominatoensis, dried male (73.0 × 66.0 mm) (PCM), Taiwan; G, Paramaja kominatoensis, dried male (60.4 × 52.5 mm) (KPM NH124171), Japan; H, Paramaja gibba, male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; I, Paramaja turgida n. sp., holotype male (74.1 × 66.8 mm) (NMCR), Philippines; J, Paramaja turgida n. sp., paratype female (62.6 × 57.2 mm) (ZRC 2013.1241), Philippines; K, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; L, Alcomaja irrorata, male (43.0 × 37.2 mm) (ZRC 2013.1223), Philippines; M, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; N, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384712" httpUri="https://zenodo.org/record/5384712/files/figure.png" pageId="35" pageNumber="145">55K–N</figureCitation>
). Thoracic sternum wide; surfaces of somites 5–8 with numerous prominent rounded tubercles and granules; sternites 3 and 4 distinctly depressed; margin between sternites 2 and 3 demarcated by deep notch; anterior margin of sterno-abdominal cavity not forming complete rim (
<figureCitation id="133BC8D8FFA2077B57DBFB1E7874FB16" box="[188,278,1181,1205]" captionStart="Fig" captionStartId="85.[135,170,1745,1767]" captionTargetBox="[136,1474,392,1730]" captionTargetId="figure-8@85.[136,1474,391,1732]" captionTargetPageId="85" captionText="Fig. 49. Male anterior thoracic sternums and abdomens, Alcomaja species. A, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; B, A.gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; C, A. gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; D, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; E, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM-ZC9928), Japan; F, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; G, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; H, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384700" httpUri="https://zenodo.org/record/5384700/files/figure.png" pageId="35" pageNumber="145">
Figs. 
<geoCoordinate id="EE34B29AFFA2077B579EFB1E7874FB16" box="[249,278,1181,1205]" degrees="49, 52" direction="east" orientation="longitude" pageId="35" pageNumber="145" precision="555" value="49.52">49</geoCoordinate>
</figureCitation>
, 
<figureCitation id="133BC8D8FFA2077B5646FB1E7810FB16" box="[289,370,1181,1205]" captionStart="Fig" captionStartId="88.[113,148,1712,1734]" captionTargetBox="[113,1452,369,1696]" captionTargetId="figure-170@88.[113,1452,369,1696]" captionTargetPageId="88" captionText="Fig. 52. Male sterno-abdominal cavities. A, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; B, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; C, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; D, Paramaja gibba n. sp., male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; E, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; F, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; G, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; H, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; I, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; J, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; K, Planaja plana n. gen. n. sp., holotype male (43.7 × 37.1 mm) (NMCR), Philippines; L, Ovimaja compressipes, male (49.6 × 39.6 mm) (ZRC 2008.1318), Taiwan." figureDoi="http://doi.org/10.5281/zenodo.5384706" httpUri="https://zenodo.org/record/5384706/files/figure.png" pageId="35" pageNumber="145">52E–G</figureCitation>
). Male abdomen subrectangular, with 6 free somites and telson; somites 3 and 4 subequal to or slightly wider than somites 5, 6 and telson (
<figureCitation id="133BC8D8FFA2077B550FFB5E7BD8FB56" box="[616,698,1245,1269]" captionStart="Fig" captionStartId="85.[135,170,1745,1767]" captionTargetBox="[136,1474,392,1730]" captionTargetId="figure-8@85.[136,1474,391,1732]" captionTargetPageId="85" captionText="Fig. 49. Male anterior thoracic sternums and abdomens, Alcomaja species. A, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; B, A.gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; C, A. gracilipes, male (41.4 × 35.4 mm) (ZRC 2013.1225), Philippines; D, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; E, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM-ZC9928), Japan; F, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; G, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; H, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines." figureDoi="http://doi.org/10.5281/zenodo.5384700" httpUri="https://zenodo.org/record/5384700/files/figure.png" pageId="35" pageNumber="145">Fig. 49</figureCitation>
). Male press-button abdominal locking mechanism submedian in position on sterno-abdominal cavity (
<figureCitation id="133BC8D8FFA2077B5578FA9E7BC6FA96" box="[543,676,1309,1333]" captionStart="Fig" captionStartId="88.[113,148,1712,1734]" captionTargetBox="[113,1452,369,1696]" captionTargetId="figure-170@88.[113,1452,369,1696]" captionTargetPageId="88" captionText="Fig. 52. Male sterno-abdominal cavities. A, Maja brachydactyla, male (98.4 × 89.0 mm) (ZRC 2009.1130), U.K.; B, Maja crispata, male (63.1 × 51.9 mm) (MNHN-IU-2013-4042), Italy; C, Neomaja goltziana, male (73.4 × 65.0 mm) (MNHN-IU-2013-4046), Congo; D, Paramaja gibba n. sp., male (79.5 × 77.9 mm) (ZRC 2013.1232), Bay of Bengal; E, Alcomaja irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; F, Alcomaja desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines; G, Alcomaja nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; H, Paramaya spinigera, male (85.0 × 66.4 mm) (ZRC 1999.738), Taiwan; I, Holthuija miersi, male (32.6 × 25.6 mm) (ZRC 2000.1497), Singapore; J, Sakaija japonica, male (22.3 × 17.8 mm) (ZRC 2013.1267), Taiwan; K, Planaja plana n. gen. n. sp., holotype male (43.7 × 37.1 mm) (NMCR), Philippines; L, Ovimaja compressipes, male (49.6 × 39.6 mm) (ZRC 2008.1318), Taiwan." figureDoi="http://doi.org/10.5281/zenodo.5384706" httpUri="https://zenodo.org/record/5384706/files/figure.png" pageId="35" pageNumber="145">Fig. 52E–G</figureCitation>
). Female abdomen dome-shaped, covering most of thoracic sternum. G1 long, slender, gently curved, distal part subconical with rounded tip, dorsal part folding downwards to form ventral projection, with scattered setae on ventral surface and numerous short setae on dorsal surface (
<figureCitation id="133BC8D8FFA2077B5508FA3E7BA8FA76" box="[623,714,1469,1493]" captionStart="Fig" captionStartId="42.[113,148,794,816]" captionTargetBox="[114,1451,189,779]" captionTargetId="figure-15@42.[104,1473,179,782]" captionTargetPageId="42" captionText="Fig. 19. Left G1s, Alcomaja species. A–C, A. irrorata n. sp., holotype male (52.6 × 45.7 mm) (NMCR), Philippines; D–F, A. gracilipes, holotype male (45.4 × 43.5 mm) (IOCAS K33B-34), South China Sea; G, A. confragosa (as Maja confragosa, after Griffin &amp; Tranter, 1986: fig. 73a). Scale bars = 1.0 mm [A–C]." figureDoi="http://doi.org/10.5281/zenodo.5384634" httpUri="https://zenodo.org/record/5384634/files/figure.png" pageId="35" pageNumber="145">Figs. 19</figureCitation>
, 
<figureCitation id="133BC8D8FFA2077B55B0FA3E7B97FA76" box="[727,757,1469,1493]" captionStart="Fig" captionStartId="43.[136,171,1934,1956]" captionTargetBox="[138,1474,203,1912]" captionTargetId="figure-8@43.[132,1494,190,1915]" captionTargetPageId="43" captionText="Fig. 20. Left G1s, Alcomaja species. A, B, A. nagashimaensis (as Maja nagashimaensis, after T. Sakai, 1976: text-fig. 129); C, D, A. nagashimaensis, holotype male (34.0 × 28.5 mm) (USNM 125886), Japan; E–G, A. nagashimaensis, male (37.2 × 32.4 mm) (CBM- ZC9928), Japan; H–J, A. nagashimaensis, male (30.5 × 26.0 mm) (ZRC 2001.430), Philippines; K–N, A. latens n. sp., holotype male (25.3 × 21.4 mm) (MNHN-IU-2013-4050), Solomon Islands; O–Q, A. miriky n. sp., holotype male (26.4 × 20.9 mm) (MNHN-IU-2010-929), Madagascar; R–T, A. desmondi n. sp., holotype male (35.4 × 28.3 mm) (NMCR), Philippines. Scale bars = 1.0 mm [H, K–R]; 2.0 mm [E]; 0.5 mm [C, F, G, I, J, S, T]; 0.25 mm [D]." figureDoi="http://doi.org/10.5281/zenodo.5384636" httpUri="https://zenodo.org/record/5384636/files/figure.png" pageId="35" pageNumber="145">20</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31A87D6FFA2077B57E0FA7E7A65F9B6" box="[135,775,1533,1557]" pageId="35" pageNumber="145" type="type_taxon">
<paragraph id="8BBFD45DFFA2077B57E0FA7E7A65F9B6" blockId="35.[135,775,1533,1557]" box="[135,775,1533,1557]" pageId="35" pageNumber="145">
<emphasis id="B974084FFFA2077B57E0FA7E787AF9B6" bold="true" box="[135,280,1533,1557]" pageId="35" pageNumber="145">
<typeStatus id="54BB6AFFFFA2077B57E0FA7E79A2F9B6" box="[135,192,1533,1557]" pageId="35" pageNumber="145">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C00AFDEFFA2077B567AFA7E78BBF9B6" ID-CoL="BKVV" authority="Ng &amp; Forges, 2015" authorityName="Ng &amp; Forges" authorityYear="2015" box="[285,473,1533,1557]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="145" phylum="Arthropoda" rank="species" species="irrorata" status="sp. nov.">
<emphasis id="B974084FFFA2077B567AFA7E78BBF9B6" box="[285,473,1533,1557]" italics="true" pageId="35" pageNumber="145">Alcomaja irrorata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B534FFA2077B56B9FA7D7B76F9B6" box="[478,532,1534,1557]" pageId="35" pageNumber="145" rank="species">n. sp.</taxonomicNameLabel>
by present designation.
</paragraph>
</subSubSection>
<subSubSection id="C31A87D6FFA2077B57E1F9BE78A6F936" pageId="35" pageNumber="145" type="etymology">
<paragraph id="8BBFD45DFFA2077B57E1F9BE78A6F936" blockId="35.[134,775,1597,1685]" pageId="35" pageNumber="145">
<emphasis id="B974084FFFA2077B57E1F9BE7867F9F6" bold="true" box="[134,261,1597,1621]" pageId="35" pageNumber="145">Etymology.</emphasis>
The genus is named after Alfred William Alcock. The name is an arbitrary combination of his family name with 
<taxonomicName id="4C00AFDEFFA2077B57D8F9FD7995F936" authorityName="Lamarck" authorityYear="1801" box="[191,247,1662,1685]" class="Malacostraca" family="Majidae" genus="Maja" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFFA2077B57D8F9FD7995F936" box="[191,247,1662,1685]" italics="true" pageId="35" pageNumber="145">Maja</emphasis>
</taxonomicName>
. Gender feminine.
</paragraph>
</subSubSection>
<subSubSection id="C31A87D6FFA2077B57E0F93E78E6F7B5" pageId="35" pageNumber="145" type="discussion">
<paragraph id="8BBFD45DFFA2077B57E0F93E7869F8F6" blockId="35.[135,775,1725,1877]" pageId="35" pageNumber="145">
<emphasis id="B974084FFFA2077B57E0F93E7995F976" bold="true" box="[135,247,1725,1749]" pageId="35" pageNumber="145">Remarks.</emphasis>
<taxonomicName id="4C00AFDEFFA2077B5666F93E780AF976" authority="Ng &amp; Forges, 2015" authorityName="Ng &amp; Forges" authorityYear="2015" box="[257,360,1725,1749]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="145" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B974084FFFA2077B5666F93E780AF976" box="[257,360,1725,1749]" italics="true" pageId="35" pageNumber="145">Alcomaja</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B534FFA2077B5615F93D78DDF975" box="[370,447,1726,1750]" pageId="35" pageNumber="145" rank="genus">n. gen.</taxonomicNameLabel>
is established to accomodate 
<taxonomicName id="4C00AFDEFFA2077B57E0F95D799DF956" authority="Ng &amp; Forges, 2015" authorityName="Ng &amp; Forges" authorityYear="2015" box="[135,255,1758,1781]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="145" phylum="Arthropoda" rank="species" species="irrorata" status="sp. nov.">
<emphasis id="B974084FFFA2077B57E0F95D799DF956" box="[135,255,1758,1781]" italics="true" pageId="35" pageNumber="145">A. irrorata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B534FFA2077B566EF95D782AF956" box="[265,328,1758,1781]" pageId="35" pageNumber="145" rank="species">n. sp.</taxonomicNameLabel>
, 
<taxonomicName id="4C00AFDEFFA2077B5630F95D7BB8F956" authority="Chen &amp; Ng, 1999" authorityName="Chen &amp; Ng" authorityYear="1999" box="[343,730,1757,1782]" class="Malacostraca" family="Majidae" genus="Maja" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="145" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="B974084FFFA2077B5630F95D7B67F956" box="[343,517,1757,1781]" italics="true" pageId="35" pageNumber="145">Maja gracilipes</emphasis>
<bibRefCitation id="EF91A9ACFFA2077B5568F95E7BB8F956" author="Chen HL &amp; Ng PKL" box="[527,730,1757,1782]" pageId="35" pageNumber="145" pagination="754 - 758" refId="ref83546" refString="Chen HL &amp; Ng PKL (1999) Description of a new spider crab, Maja gracilipes, from the South China Sea, with notes on the taxonomic validity of M. brevispinosis Dai, 1981 (Crustacea: Decapoda: Brachyura: Majidae). Proceedings of the Biological Society of Washington, 112 (4): 754 - 758, Figs. 1, 2." type="journal article" year="1999">Chen &amp; Ng, 1999</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C00AFDEFFA2077B558EF95D7B69F8B6" authority="Griffin &amp; Tranter, 1986" authorityName="Griffin &amp; Tranter" authorityYear="1986" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="145" phylum="Arthropoda" rank="species" species="confragosa">
<emphasis id="B974084FFFA2077B558EF95D799DF8B6" italics="true" pageId="35" pageNumber="145">M. confragosa</emphasis>
<bibRefCitation id="EF91A9ACFFA2077B5660F97E7B69F8B6" author="Griffin DJG &amp; Tranter HA" box="[263,523,1789,1813]" pageId="35" pageNumber="145" pagination="1 - 335" refId="ref84094" refString="Griffin DJG &amp; Tranter HA (1986) The Decapoda Brachyura of the Siboga Expedition. Part VIII. Majidae. Siboga-Expeditie, 39 C 4: 1 - 335, 22 pls." type="journal article" year="1986">Griffin &amp; Tranter, 1986</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C00AFDEFFA2077B5570F97D786EF896" authority="T. Sakai, 1969" authorityName="T. Sakai" authorityYear="1969" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="145" phylum="Arthropoda" rank="species" species="nagashimaensis">
<emphasis id="B974084FFFA2077B5570F97D7B84F8B6" box="[535,742,1789,1813]" italics="true" pageId="35" pageNumber="145">M. nagashimaensis</emphasis>
T. 
<bibRefCitation id="EF91A9ACFFA2077B57E0F89E786EF896" author="Sakai T" box="[135,268,1821,1845]" pageId="35" pageNumber="145" pagination="243 - 280" refId="ref88407" refString="Sakai T (1969) Two new genera and twenty-two new species of crabs from Japan. Proceedings of the Biological Society of Washington, 82: 243 - 280, figs. 1 - 19, pls. 1, 2." type="journal article" year="1969">Sakai, 1969</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C00AFDEFFA2077B567BF89D78C7F896" authority="Ng &amp; Forges, 2015" authorityName="Ng &amp; Forges" authorityYear="2015" box="[284,421,1821,1845]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="145" phylum="Arthropoda" rank="species" species="desmondi" status="sp. nov.">
<emphasis id="B974084FFFA2077B567BF89D78C7F896" box="[284,421,1821,1845]" italics="true" pageId="35" pageNumber="145">A. desmondi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B534FFA2077B56C8F89D788CF896" box="[431,494,1822,1845]" pageId="35" pageNumber="145" rank="species">n. sp.</taxonomicNameLabel>
, 
<taxonomicName id="4C00AFDEFFA2077B5699F89D7B06F896" authority="Ng &amp; Forges, 2015" authorityName="Ng &amp; Forges" authorityYear="2015" box="[510,612,1821,1845]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="145" phylum="Arthropoda" rank="species" species="miriky" status="sp. nov.">
<emphasis id="B974084FFFA2077B5699F89D7B06F896" box="[510,612,1821,1845]" italics="true" pageId="35" pageNumber="145">A. miriky</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B534FFA2077B5508F89D7BCCF896" box="[623,686,1822,1845]" pageId="35" pageNumber="145" rank="species">n. sp.</taxonomicNameLabel>
, and 
<taxonomicName id="4C00AFDEFFA2077B5588F89D79A5F8F6" authority="Ng &amp; Forges, 2015" authorityName="Ng &amp; Forges" authorityYear="2015" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="145" phylum="Arthropoda" rank="species" species="latens" status="sp. nov.">
<emphasis id="B974084FFFA2077B5588F89D79A5F8F6" italics="true" pageId="35" pageNumber="145">A. latens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247B534FFA2077B57B7F8BD7869F8F6" box="[208,267,1854,1877]" pageId="35" pageNumber="145" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BBFD45DFFA2077B57E0F8FD78E6F7B5" blockId="35.[135,776,1917,2070]" pageId="35" pageNumber="145">
In the form of the suborbital margin, basal antennal article, epistome, third maxilliped, pereiopods, thoracic sternum and male abdomen, 
<taxonomicName id="4C00AFDEFFA2077B5600F83E78AFF876" authorityName="Ng &amp; Forges" authorityYear="2015" box="[359,461,1981,2005]" class="Malacostraca" family="Majidae" genus="Alcomaja" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFFA2077B5600F83E78AFF876" box="[359,461,1981,2005]" italics="true" pageId="35" pageNumber="145">Alcomaja</emphasis>
</taxonomicName>
is most similar to 
<taxonomicName id="4C00AFDEFFA2077B55FAF83D786BF856" authority="Kubo, 1936" authorityName="Kubo" authorityYear="1936" class="Malacostraca" family="Majidae" genus="Paramaja" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="B974084FFFA2077B55FAF83D7A6AF876" box="[669,776,1982,2005]" italics="true" pageId="35" pageNumber="145">Paramaja</emphasis>
<bibRefCitation id="EF91A9ACFFA2077B57E0F85E786BF856" author="Kubo I" box="[135,265,2013,2037]" pageId="35" pageNumber="145" pagination="361 - 364" refId="ref85511" refString="Kubo I (1936) A new decapod crustacean from Japan, Paramaja kominatoensis, gen. et sp. nov. Bulletin of the Japanese Society of Scientific Fisheries, 4 (6): 361 - 364." type="journal article" year="1936">Kubo, 1936</bibRefCitation>
</taxonomicName>
. The differences have been discussed under the remarks for that genus.
</paragraph>
</subSubSection>
</treatment>
</document>