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<document id= "443C6C0BEE4F3D40B273A7B22CEA8224" ID-DOI= "10.1590/S0031-10492012021600001" ID-ISSN= "1807-0205" ID-Zenodo-Dep= "13175196" IM.bibliography_requiresApprovalFor= "GgImagineBatch" IM.illustrations_approvedBy= "felipe" IM.materialsCitations_approvedBy= "felipe" IM.metadata_approvedBy= "carolina" IM.taxonomicNames_approvedBy= "felipe" IM.treatmentCitations_approvedBy= "felipe" IM.treatments_approvedBy= "felipe" checkinTime= "1720046602614" checkinUser= "felipe" docAuthor= "Simone, Luiz Ricardo L." docDate= "2012" docId= "0399C3592B17087AFD11FD166A17FE65" docLanguage= "en" docName= "PapAvZool.52.36.431-439.pdf.imf" docOrigin= "Papéis Avulsos de Zoologia 52 (36)" docSource= "http://www.scielo.br/scielo.php?script=sci_arttext&pid=S0031-10492012021600001&lng=en&tlng=en" docStyle= "DocumentStyle:342E0B2A517D47F211C16A9F07059896.3:PapAvZool.2002-2013.journal_article.type2" docStyleId= "342E0B2A517D47F211C16A9F07059896" docStyleName= "PapAvZool.2002-2013.journal_article.type2" docStyleVersion= "3" docTitle= "Megalobulimus amandus Simone, 2012, new species" docType= "treatment" docVersion= "4" lastPageNumber= "439" masterDocId= "FFA0BB212B12087DFFE0FFA16951FFB5" masterDocTitle= "Taxonomical Study On A Sample Of Pulmonates From Santa Maria Da Vitória, Bahia, Brazil, With Description Of A New Genus And Four New Species (Mollusca: Orthalicidae And Megalobulimidae)" masterLastPageNumber= "439" masterPageNumber= "431" pageNumber= "431" updateTime= "1722623525616" updateUser= "ExternalLinkService" >
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<mods:title id= "8462C4D4F94EB610CA6E974679C4B4AD" > Taxonomical Study On A Sample Of Pulmonates From Santa Maria Da Vitória, Bahia, Brazil, With Description Of A New Genus And Four New Species (Mollusca: Orthalicidae And Megalobulimidae)</mods:title>
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<mods:namePart id= "9CA9BC933BBC972C1FFC6F8F9C55BDA0" > Simone, Luiz Ricardo L.</mods:namePart>
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<mods:title id= "7CD3FF1EBA5180ADE99FBDE93DEC9FAC" > Papéis Avulsos de Zoologia</mods:title>
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<mods:part id= "7FBAA6BAF1346C448CA66C6E7B278F5C" >
<mods:date id= "87FFB909E49C32DE192722CB39F50029" > 2012</mods:date>
<mods:detail id= "DAFB21106E0DF5F7E54EFAF5C8E35881" type= "pubDate" >
<mods:number id= "96D44035605B8DD3EB642DFB9EFCFD9D" > 2012-12-20</mods:number>
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<treatment id= "0399C3592B17087AFD11FD166A17FE65" LSID= "urn:lsid:plazi:treatment:0399C3592B17087AFD11FD166A17FE65" httpUri= "http://treatment.plazi.org/id/0399C3592B17087AFD11FD166A17FE65" lastPageId= "7" pageId= "5" >
<subSubSection id= "C32A21C42B170878FD11FD166DD5FD65" box= "[753,1156,695,720]" pageId= "5" type= "nomenclature" >
<paragraph id= "8B8F724F2B170878FD11FD166DD5FD65" blockId= "5.[753,1156,695,752]" box= "[753,1156,695,720]" pageId= "5" >
<heading id= "D0C7C5232B170878FD11FD166DD5FD65" bold= "true" box= "[753,1156,695,720]" centered= "true" fontSize= "9" level= "2" pageId= "5" reason= "2" >
<emphasis id= "B944AE5D2B170878FD11FD166DD5FD65" bold= "true" box= "[753,1156,695,720]" pageId= "5" >
<taxonomicName id= "4C3009CC2B170878FD11FD166D51FD7A" ID-CoL= "B598P" authorityName= "Simone" authorityYear= "2012" box= "[753,1024,695,719]" class= "Gastropoda" family= "Strophocheilidae" genus= "Megalobulimus" kingdom= "Animalia" order= "Stylommatophora" pageId= "5" phylum= "Mollusca" rank= "species" species= "amandus" status= "sp. nov." >
<emphasis id= "B944AE5D2B170878FD11FD166D51FD7A" bold= "true" box= "[753,1024,695,719]" italics= "true" pageId= "5" > Megalobulimus amandus</emphasis>
</taxonomicName>
<taxonomicNameLabel id= "A27713262B170878FBE7FD196DD5FD65" box= "[1031,1156,696,720]" pageId= "5" rank= "species" > new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id= "C32A21C42B17087AFC95FD796A17FE65" lastPageId= "7" pageId= "5" type= "description" >
<paragraph id= "8B8F724F2B170878FC95FD796D51FD45" blockId= "5.[753,1156,695,752]" box= "[885,1024,728,752]" pageId= "5" >
<emphasis id= "B944AE5D2B170878FC95FD796D51FD45" bold= "true" box= "[885,1024,728,752]" pageId= "5" >
(
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<figureCitation id= "130B6ECA2B170878FC9CFD796AA9FD45" box= "[892,1016,728,752]" captionStart= "FIGURES 15‑ 26" captionStartId= "6.[132,222,1724,1743]" captionTargetBox= "[132,1266,497,1688]" captionTargetId= "figure-99@6.[132,1266,497,1688]" captionTargetPageId= "6" captionText= "FIGURES 15‑ 26: Shell of types and ordinary specimens: 15-20) Anostoma tessa new species; 15) Holotype, apertural view (H 31.1 mm); 16) Same, dorsal view; 17) Same, left view; 18) Same, right view; 19) Same, detail of aperture, apertural-slightly anterior view; 20) Paratype MZSP 103915, apertural view (H 31.1 mm); 21) Rhinus suturalis, MZSP 103924, frontal view (H 23.6 mm); 22-26) Megalobulimus amandus new species; 22) Holotype, apertural view (H 78.9 mm); 23) Same, right view; 24) Same, dorsal view; 25) Paratype MZSP 103919(2), young specimen, apical (larger) and apertural view (H 25.6 mm); 26) Paratype MZSP 103919(1), adult with thin lip, apertural and right views (H 69.8 mm)." figureDoi= "http://doi.org/10.5281/zenodo.13175200" httpUri= "https://zenodo.org/record/13175200/files/figure.png" pageId= "5" > Figs. 22-26</figureCitation>
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)
</emphasis>
</paragraph>
<paragraph id= "8B8F724F2B170878FD48FCB86AA4FC85" blockId= "5.[680,1013,792,816]" box= "[680,1013,792,816]" pageId= "5" >
<emphasis id= "B944AE5D2B170878FD48FCB86BB2FC85" box= "[680,739,793,816]" italics= "true" pageId= "5" > Types:</emphasis>
<materialsCitation id= "3B5878122B170878FD0AFCB96AA4FC85" box= "[746,1013,792,816]" collectionCode= "MZSP" pageId= "5" specimenCode= "MZSP 103917" specimenCount= "1" typeStatus= "holotype" >
<typeStatus id= "548BCCED2B170878FD0AFCB96A1AFC85" box= "[746,843,792,816]" pageId= "5" type= "holotype" > Holotype</typeStatus>
<specimenCode id= "DB96DA342B170878FCB2FCB96AA0FC85" box= "[850,1009,792,816]" collectionCode= "MZSP" country= "Brazil" httpUri= "http://grbio.org/cool/9yp6-zxp9" name= "Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId= "5" type= "Museum" > MZSP 103917</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id= "8B8F724F2B170878FD48FCF86A97FC05" blockId= "5.[680,1229,856,944]" pageId= "5" >
<materialsCitation id= "3B5878122B170878FD48FCF86D41FCC5" box= "[680,1040,856,880]" collectionCode= "MZSP" pageId= "5" specimenCode= "MZSP 103919, 3" specimenCount= "1" typeStatus= "paratype" >
<emphasis id= "B944AE5D2B170878FD48FCF86A5BFCC5" box= "[680,778,857,880]" italics= "true" pageId= "5" >
<typeStatus id= "548BCCED2B170878FD48FCF86A55FCC5" box= "[680,772,857,880]" pageId= "5" type= "paratype" > Paratypes</typeStatus>
:
</emphasis>
<specimenCode id= "DB96DA342B170878FCF2FCF96AE2FCC5" box= "[786,947,856,880]" collectionCode= "MZSP" country= "Brazil" httpUri= "http://grbio.org/cool/9yp6-zxp9" name= "Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId= "5" type= "Museum" > MZSP 103919</specimenCode>
,
<specimenCode id= "DB96DA342B170878FC20FCF96A9CFCC5" box= "[960,973,856,880]" collectionCode= "MZSP" country= "Brazil" httpUri= "http://grbio.org/cool/9yp6-zxp9" name= "Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId= "5" type= "Museum" > 3</specimenCode>
shells
</materialsCitation>
;
<collectionCode id= "ED21EA8A2B170878FBFCFCF96D3CFCC5" box= "[1052,1133,856,880]" country= "USA" httpUri= "http://biocol.org/urn:lsid:biocol.org:col:34871" lsid= "urn:lsid:biocol.org:col:34871" name= "Smithsonian Institution, National Museum of Natural History" pageId= "5" type= "Museum" > USNM</collectionCode>
, 1 shells;
<collectionCode id= "ED21EA8A2B170878FD48FCD96BA3FC3A" box= "[680,754,888,911]" country= "Brazil" httpUri= "http://grbio.org/cool/zi1i-a0b5" name= "Museu Nacional/Universidade Federal de Rio de Janeiro" pageId= "5" > MNRJ</collectionCode>
, 1 shells; NMHN, 1 shells;
<materialsCitation id= "3B5878122B170878FBEBFCD96A92FC05" collectionCode= "MZSP" pageId= "5" specimenCode= "MZSP 103918, 10" specimenCount= "1" typeStatus= "paratype" >
<specimenCode id= "DB96DA342B170878FBEBFCD96DF8FC25" box= "[1035,1193,888,912]" collectionCode= "MZSP" country= "Brazil" httpUri= "http://grbio.org/cool/9yp6-zxp9" name= "Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId= "5" type= "Museum" > MZSP 103918</specimenCode>
,
<specimenCode id= "DB96DA342B170878FB52FCD96D9CFC25" box= "[1202,1229,888,912]" collectionCode= "MZSP" country= "Brazil" httpUri= "http://grbio.org/cool/9yp6-zxp9" name= "Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId= "5" type= "Museum" > 10</specimenCode>
shells; all from type locality
</materialsCitation>
.
</paragraph>
<paragraph id= "8B8F724F2B170878FD48FC786BACFB85" blockId= "5.[680,1229,984,1072]" pageId= "5" >
<emphasis id= "B944AE5D2B170878FD48FC786A76FC45" box= "[680,807,984,1008]" italics= "true" pageId= "5" > Type locality:</emphasis>
<materialsCitation id= "3B5878122B170878FCCDFC796BABFB85" collectingDate= "2012-01" collectorName= "Coltro" country= "Brazil" elevation= "460" latitude= "-13.4" location= "Santa Maria da Vitoria" longLatPrecision= "1291" longitude= "-44.2" pageId= "5" specimenCount= "1" stateProvince= "Bahia" >
<collectingCountry id= "F32732DF2B170878FCCDFC796ADDFC45" box= "[813,908,984,1008]" name= "Brazil" pageId= "5" > BRAZIL</collectingCountry>
.
<emphasis id= "B944AE5D2B170878FC74FC796A8BFC45" bold= "true" box= "[916,986,984,1008]" pageId= "5" >
<collectingRegion id= "49F4BCAD2B170878FC74FC796A84FC45" box= "[916,981,984,1008]" country= "Brazil" name= "Bahia" pageId= "5" > Bahia</collectingRegion>
;
</emphasis>
<location id= "8EEF24942B170878FC3FFC796D98FC45" LSID= "urn:lsid:plazi:treatment:0399C3592B17087AFD11FD166A17FE65:8EEF24942B170878FC3FFC796D98FC45" box= "[991,1225,984,1008]" country= "Brazil" latitude= "-13.4" longLatPrecision= "1291" longitude= "-44.2" name= "Santa Maria da Vitoria" pageId= "5" stateProvince= "Bahia" > Santa Maria da Vitória</location>
, ~
<geoCoordinate id= "EE0414882B170878FD57FC596A58FBA5" box= "[695,777,1016,1040]" degrees= "13" direction= "south" minutes= "24" orientation= "latitude" pageId= "5" precision= "925" value= "-13.4" > 13°24’ S</geoCoordinate>
,
<geoCoordinate id= "EE0414882B170878FCF3FC596A21FBA5" box= "[787,880,1016,1040]" degrees= "44" direction= "west" minutes= "12" orientation= "longitude" pageId= "5" precision= "925" value= "-44.2" > 44°12’ W</geoCoordinate>
, ~
<quantity id= "4CC8DFAA2B170878FC67FC596A9AFBBA" box= "[903,971,1016,1040]" metricMagnitude= "2" metricUnit= "m" metricValue= "4.6" pageId= "5" unit= "m" value= "460.0" >
<elevation id= "001D957C2B170878FC67FC596A9AFBBA" box= "[903,971,1016,1040]" metricMagnitude= "2" metricUnit= "m" metricValue= "4.6" pageId= "5" unit= "m" value= "460.0" > 460 m</elevation>
</quantity>
of elevation (
<collectorName id= "26C517992B170878FBB5FC596DCBFBA5" box= "[1109,1178,1016,1040]" pageId= "5" > Coltro</collectorName>
col.,
<date id= "FF8E548F2B170878FD48FBB96BA0FB85" box= "[680,753,1048,1072]" bridgedPair= "/" pageId= "5" value= "2012-01" >
<collectingDate id= "EFCAAD672B170878FD48FBB96BA0FB85" box= "[680,753,1048,1072]" bridgedPair= "/" pageId= "5" value= "2012-01" > i/2012</collectingDate>
</date>
)
</materialsCitation>
.
</paragraph>
<paragraph id= "8B8F724F2B170878FD48FBF86A8FFB65" blockId= "5.[680,1229,1112,1232]" pageId= "5" >
<emphasis id= "B944AE5D2B170878FD48FBF86A5FFBC5" box= "[680,782,1113,1136]" italics= "true" pageId= "5" > Diagnosis:</emphasis>
Shell with less than
<quantity id= "4CC8DFAA2B170878FC11FBF96D13FBDA" box= "[1009,1090,1112,1136]" metricMagnitude= "-2" metricUnit= "m" metricValue= "8.0" pageId= "5" unit= "mm" value= "80.0" > 80 mm</quantity>
, tip pointed, spire acuminate. Protoconch bearing series of uniform, delicate axial cords. Shell walls relatively thick. Peristome simple, pink to red.
</paragraph>
<paragraph id= "8B8F724F2B17087BFD48FB586AD3FF45" blockId= "5.[680,1229,1272,1872]" lastBlockId= "6.[718,1266,184,240]" lastPageId= "6" pageId= "5" >
<emphasis id= "B944AE5D2B170878FD48FB586A4EFAA5" box= "[680,799,1273,1296]" italics= "true" pageId= "5" > Description:</emphasis>
Shell up to
<quantity id= "4CC8DFAA2B170878FC4EFB596D50FABA" box= "[942,1025,1272,1296]" metricMagnitude= "-2" metricUnit= "m" metricValue= "8.0" pageId= "5" unit= "mm" value= "80.0" > 80 mm</quantity>
; outline oval; apex acuminate; width ~60 of length, dorso-ventral height ~50% of shell length. Color mostly pale beige in periostracum-lacking specimens (
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<figureCitation id= "130B6ECA2B170878FBE5FAF96DD1FAC5" box= "[1029,1152,1368,1392]" captionStart= "FIGURES 15‑ 26" captionStartId= "6.[132,222,1724,1743]" captionTargetBox= "[132,1266,497,1688]" captionTargetId= "figure-99@6.[132,1266,497,1688]" captionTargetPageId= "6" captionText= "FIGURES 15‑ 26: Shell of types and ordinary specimens: 15-20) Anostoma tessa new species; 15) Holotype, apertural view (H 31.1 mm); 16) Same, dorsal view; 17) Same, left view; 18) Same, right view; 19) Same, detail of aperture, apertural-slightly anterior view; 20) Paratype MZSP 103915, apertural view (H 31.1 mm); 21) Rhinus suturalis, MZSP 103924, frontal view (H 23.6 mm); 22-26) Megalobulimus amandus new species; 22) Holotype, apertural view (H 78.9 mm); 23) Same, right view; 24) Same, dorsal view; 25) Paratype MZSP 103919(2), young specimen, apical (larger) and apertural view (H 25.6 mm); 26) Paratype MZSP 103919(1), adult with thin lip, apertural and right views (H 69.8 mm)." figureDoi= "http://doi.org/10.5281/zenodo.13175200" httpUri= "https://zenodo.org/record/13175200/files/figure.png" pageId= "5" > Figs. 22-25</figureCitation>
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); some few specimens with periostracum (
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<figureCitation id= "130B6ECA2B170878FBECFAD96D06FA25" box= "[1036,1111,1400,1424]" captionStart= "FIGURES 15‑ 26" captionStartId= "6.[132,222,1724,1743]" captionTargetBox= "[132,1266,497,1688]" captionTargetId= "figure-99@6.[132,1266,497,1688]" captionTargetPageId= "6" captionText= "FIGURES 15‑ 26: Shell of types and ordinary specimens: 15-20) Anostoma tessa new species; 15) Holotype, apertural view (H 31.1 mm); 16) Same, dorsal view; 17) Same, left view; 18) Same, right view; 19) Same, detail of aperture, apertural-slightly anterior view; 20) Paratype MZSP 103915, apertural view (H 31.1 mm); 21) Rhinus suturalis, MZSP 103924, frontal view (H 23.6 mm); 22-26) Megalobulimus amandus new species; 22) Holotype, apertural view (H 78.9 mm); 23) Same, right view; 24) Same, dorsal view; 25) Paratype MZSP 103919(2), young specimen, apical (larger) and apertural view (H 25.6 mm); 26) Paratype MZSP 103919(1), adult with thin lip, apertural and right views (H 69.8 mm)." figureDoi= "http://doi.org/10.5281/zenodo.13175200" httpUri= "https://zenodo.org/record/13175200/files/figure.png" pageId= "5" > Fig. 26</figureCitation>
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) somewhat glossy, eroded, presenting mosaic of pale and dark brown bands randomly disposed axially; peristome red to pale pink. Protoconch of ~3 weakly convex whorls, forming a somewhat pointed dome with ~70°; first whorl mostly smooth, opaque, remaining whorls bearing much delicate, uniform, narrow axial cords, ~
<quantity id= "4CC8DFAA2B170878FCD9F9F96AD0F9C5" box= "[825,897,1624,1648]" metricMagnitude= "0" metricUnit= "m" metricValue= "3.048" pageId= "5" unit= "in" value= "120.0" > 120 in</quantity>
last nepionic whorl (
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<figureCitation id= "130B6ECA2B170878FB8FF9F96D91F9C5" box= "[1135,1216,1624,1648]" captionStart= "FIGURES 15‑ 26" captionStartId= "6.[132,222,1724,1743]" captionTargetBox= "[132,1266,497,1688]" captionTargetId= "figure-99@6.[132,1266,497,1688]" captionTargetPageId= "6" captionText= "FIGURES 15‑ 26: Shell of types and ordinary specimens: 15-20) Anostoma tessa new species; 15) Holotype, apertural view (H 31.1 mm); 16) Same, dorsal view; 17) Same, left view; 18) Same, right view; 19) Same, detail of aperture, apertural-slightly anterior view; 20) Paratype MZSP 103915, apertural view (H 31.1 mm); 21) Rhinus suturalis, MZSP 103924, frontal view (H 23.6 mm); 22-26) Megalobulimus amandus new species; 22) Holotype, apertural view (H 78.9 mm); 23) Same, right view; 24) Same, dorsal view; 25) Paratype MZSP 103919(2), young specimen, apical (larger) and apertural view (H 25.6 mm); 26) Paratype MZSP 103919(1), adult with thin lip, apertural and right views (H 69.8 mm)." figureDoi= "http://doi.org/10.5281/zenodo.13175200" httpUri= "https://zenodo.org/record/13175200/files/figure.png" pageId= "5" > Fig. 25</figureCitation>
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); each cord running from suture to suture since second whorl, interval between cords very narrow; limit between protoconch and teleoconch barely clear, orthocline. Teleoconch of 2.2 to 2.5 whorls, first whorl with almost straight profile, last whorl more convex than preceding ones. Spite ~60% of shell length. Sculpture similar to that of protoconch, with cords becoming delicate, but well-marked undulations, ~
<quantity id= "4CC8DFAA2B14087BFD6EFF1969C8FF45" metricMagnitude= "0" metricUnit= "m" metricValue= "1.524" pageId= "6" unit= "in" value= "60.0" > 60 in</quantity>
penultimate whorl; ~30% of specimens possessing hammer-like marks in last whorl mid-region (
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<figureCitation id= "130B6ECA2B14087BFDB7FF596BF0FEA5" box= "[599,673,248,272]" captionStart= "FIGURES 15‑ 26" captionStartId= "6.[132,222,1724,1743]" captionTargetBox= "[132,1266,497,1688]" captionTargetId= "figure-99@6.[132,1266,497,1688]" captionTargetPageId= "6" captionText= "FIGURES 15‑ 26: Shell of types and ordinary specimens: 15-20) Anostoma tessa new species; 15) Holotype, apertural view (H 31.1 mm); 16) Same, dorsal view; 17) Same, left view; 18) Same, right view; 19) Same, detail of aperture, apertural-slightly anterior view; 20) Paratype MZSP 103915, apertural view (H 31.1 mm); 21) Rhinus suturalis, MZSP 103924, frontal view (H 23.6 mm); 22-26) Megalobulimus amandus new species; 22) Holotype, apertural view (H 78.9 mm); 23) Same, right view; 24) Same, dorsal view; 25) Paratype MZSP 103919(2), young specimen, apical (larger) and apertural view (H 25.6 mm); 26) Paratype MZSP 103919(1), adult with thin lip, apertural and right views (H 69.8 mm)." figureDoi= "http://doi.org/10.5281/zenodo.13175200" httpUri= "https://zenodo.org/record/13175200/files/figure.png" pageId= "6" > Fig. 24</figureCitation>
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) and region preceding peristome. Peristome complete, normally thick (
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<figureCitation id= "130B6ECA2B14087BFECAFE9968F9FEE5" box= "[298,424,312,336]" captionStart= "FIGURES 15‑ 26" captionStartId= "6.[132,222,1724,1743]" captionTargetBox= "[132,1266,497,1688]" captionTargetId= "figure-99@6.[132,1266,497,1688]" captionTargetPageId= "6" captionText= "FIGURES 15‑ 26: Shell of types and ordinary specimens: 15-20) Anostoma tessa new species; 15) Holotype, apertural view (H 31.1 mm); 16) Same, dorsal view; 17) Same, left view; 18) Same, right view; 19) Same, detail of aperture, apertural-slightly anterior view; 20) Paratype MZSP 103915, apertural view (H 31.1 mm); 21) Rhinus suturalis, MZSP 103924, frontal view (H 23.6 mm); 22-26) Megalobulimus amandus new species; 22) Holotype, apertural view (H 78.9 mm); 23) Same, right view; 24) Same, dorsal view; 25) Paratype MZSP 103919(2), young specimen, apical (larger) and apertural view (H 25.6 mm); 26) Paratype MZSP 103919(1), adult with thin lip, apertural and right views (H 69.8 mm)." figureDoi= "http://doi.org/10.5281/zenodo.13175200" httpUri= "https://zenodo.org/record/13175200/files/figure.png" pageId= "6" > Figs. 22, 23</figureCitation>
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), glossy, lacking tooth of folds; aperture elliptic, ~54% of shell width, ~50% of shell length. Outer lip simple and rounded. Inner lip feebly concave; no clear separation with callus and parietal region (
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<figureCitation id= "130B6ECA2B14087BFC9AFF196D50FF65" box= "[890,1025,184,208]" captionStart= "FIGURES 15‑ 26" captionStartId= "6.[132,222,1724,1743]" captionTargetBox= "[132,1266,497,1688]" captionTargetId= "figure-99@6.[132,1266,497,1688]" captionTargetPageId= "6" captionText= "FIGURES 15‑ 26: Shell of types and ordinary specimens: 15-20) Anostoma tessa new species; 15) Holotype, apertural view (H 31.1 mm); 16) Same, dorsal view; 17) Same, left view; 18) Same, right view; 19) Same, detail of aperture, apertural-slightly anterior view; 20) Paratype MZSP 103915, apertural view (H 31.1 mm); 21) Rhinus suturalis, MZSP 103924, frontal view (H 23.6 mm); 22-26) Megalobulimus amandus new species; 22) Holotype, apertural view (H 78.9 mm); 23) Same, right view; 24) Same, dorsal view; 25) Paratype MZSP 103919(2), young specimen, apical (larger) and apertural view (H 25.6 mm); 26) Paratype MZSP 103919(1), adult with thin lip, apertural and right views (H 69.8 mm)." figureDoi= "http://doi.org/10.5281/zenodo.13175200" httpUri= "https://zenodo.org/record/13175200/files/figure.png" pageId= "6" > Figs. 22, 26</figureCitation>
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). Umbilicus extremely narrow to absent.
</paragraph>
<paragraph id= "8B8F724F2B14087BFD2EFEB86A55FEC5" blockId= "6.[718,1266,280,368]" pageId= "6" >
<emphasis id= "B944AE5D2B14087BFD2EFEB86AE8FE9A" box= "[718,953,280,304]" italics= "true" pageId= "6" > Measurements (in mm):</emphasis>
<typeStatus id= "548BCCED2B14087BFC5FFEB96D75FE85" box= "[959,1060,280,304]" pageId= "6" type= "holotype" > Holotype</typeStatus>
: 78.9 by 50.0;
<typeStatus id= "548BCCED2B14087BFB5EFEB96BABFEE5" pageId= "6" type= "paratype" > paratype</typeStatus>
<collectionCode id= "ED21EA8A2B14087BFCE4FE996A18FEE5" box= "[772,841,312,336]" country= "Brazil" httpUri= "http://grbio.org/cool/9yp6-zxp9" name= "Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId= "6" type= "Museum" > MZSP</collectionCode>
103919: (1) 69.8 by 40.2; (2) 25.6 by 17.6.
</paragraph>
<paragraph id= "8B8F724F2B14087BFD2EFE396DD5FE05" blockId= "6.[718,1156,408,432]" box= "[718,1156,408,432]" pageId= "6" >
<emphasis id= "B944AE5D2B14087BFD2EFE396A01FE05" box= "[718,848,408,432]" italics= "true" pageId= "6" > Distribution:</emphasis>
Known only for
<typeStatus id= "548BCCED2B14087BFBE1FE396D7CFE05" box= "[1025,1069,408,432]" pageId= "6" > type</typeStatus>
locality.
</paragraph>
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<caption id= "DF4F22C72B14087BFF64F91D682FF8FA" ID-DOI= "http://doi.org/10.5281/zenodo.13175200" ID-Zenodo-Dep= "13175200" httpUri= "https://zenodo.org/record/13175200/files/figure.png" pageId= "6" startId= "6.[132,222,1724,1743]" targetBox= "[132,1266,497,1688]" targetPageId= "6" targetType= "figure" >
2024-07-12 03:17:58 +02:00
<paragraph id= "8B8F724F2B14087BFF64F91D682FF8FA" blockId= "6.[132,1267,1724,1871]" pageId= "6" >
<emphasis id= "B944AE5D2B14087BFF64F91D684CF97A" bold= "true" box= "[132,285,1724,1743]" pageId= "6" > FIGURES 15‑ 26:</emphasis>
Shell of types and ordinary specimens:
<emphasis id= "B944AE5D2B14087BFD84F91D6BCFF97A" bold= "true" box= "[612,670,1724,1743]" pageId= "6" > 15-20)</emphasis>
<taxonomicName id= "4C3009CC2B14087BFD44F91D6A49F97B" authorityName= "Simone" authorityYear= "2012" box= "[676,792,1724,1743]" class= "Gastropoda" family= "Odontostomidae" genus= "Anostoma" kingdom= "Animalia" order= "Stylommatophora" pageId= "6" phylum= "Mollusca" rank= "species" species= "tessa" status= "sp. nov." >
<emphasis id= "B944AE5D2B14087BFD44F91D6A49F97B" box= "[676,792,1724,1743]" italics= "true" pageId= "6" > Anostoma tessa</emphasis>
</taxonomicName>
<taxonomicNameLabel id= "A27713262B14087BFCFEF91D6A2EF97A" box= "[798,895,1724,1743]" pageId= "6" rank= "species" > new species</taxonomicNameLabel>
;
<emphasis id= "B944AE5D2B14087BFC67F91D6AF4F97A" bold= "true" box= "[903,933,1724,1743]" pageId= "6" > 15)</emphasis>
Holotype, apertural view (H 31.1 mm);
<emphasis id= "B944AE5D2B14087BFF64F97469F3F95D" bold= "true" box= "[132,162,1749,1768]" pageId= "6" > 16)</emphasis>
Same, dorsal view;
<emphasis id= "B944AE5D2B14087BFEA6F9746835F95D" bold= "true" box= "[326,356,1749,1768]" pageId= "6" > 17)</emphasis>
Same, left view;
<emphasis id= "B944AE5D2B14087BFE11F9746B5EF95D" bold= "true" box= "[497,527,1749,1768]" pageId= "6" > 18)</emphasis>
Same, right view;
<emphasis id= "B944AE5D2B14087BFD4AF9746B96F95D" bold= "true" box= "[682,711,1749,1768]" pageId= "6" > 19)</emphasis>
Same, detail of aperture, apertural-slightly anterior view;
<emphasis id= "B944AE5D2B14087BFB45F9746D93F95D" bold= "true" box= "[1189,1218,1749,1768]" pageId= "6" > 20)</emphasis>
Paratype MZSP 103915, apertural view (H 31.1 mm);
<emphasis id= "B944AE5D2B14087BFDC7F94E6B15F8B7" bold= "true" box= "[551,580,1775,1794]" pageId= "6" > 21)</emphasis>
<taxonomicName id= "4C3009CC2B14087BFDA9F94E6A58F8B7" authority= ", MZSP" authorityName= "MZSP" box= "[585,777,1775,1794]" class= "Gastropoda" family= "Simpulopsidae" genus= "Rhinus" kingdom= "Animalia" order= "Stylommatophora" pageId= "6" phylum= "Mollusca" rank= "species" species= "suturalis" >
<emphasis id= "B944AE5D2B14087BFDA9F94E6B9FF8B7" box= "[585,718,1775,1794]" italics= "true" pageId= "6" > Rhinus suturalis,</emphasis>
MZSP
</taxonomicName>
103924, frontal view (H 23.6 mm);
<emphasis id= "B944AE5D2B14087BFBDAF94E6D25F8B7" bold= "true" box= "[1082,1140,1775,1794]" pageId= "6" > 22-26)</emphasis>
<taxonomicName id= "4C3009CC2B14087BFB9AF94E699AF8A9" authorityName= "Simone" authorityYear= "2012" class= "Gastropoda" family= "Strophocheilidae" genus= "Megalobulimus" kingdom= "Animalia" order= "Stylommatophora" pageId= "6" phylum= "Mollusca" rank= "species" species= "amandus" status= "sp. nov." >
<emphasis id= "B944AE5D2B14087BFB9AF94E699AF8A9" italics= "true" pageId= "6" > Megalobulimus amandus</emphasis>
</taxonomicName>
<taxonomicNameLabel id= "A27713262B14087BFF33F8A86866F8A9" box= "[211,311,1801,1820]" pageId= "6" rank= "species" > new species</taxonomicNameLabel>
;
<emphasis id= "B944AE5D2B14087BFEA1F8A9680FF8AE" bold= "true" box= "[321,350,1800,1819]" pageId= "6" > 22)</emphasis>
Holotype, apertural view (H 78.9 mm);
<emphasis id= "B944AE5D2B14087BFD20F8A96B8CF8AE" bold= "true" box= "[704,733,1800,1819]" pageId= "6" > 23)</emphasis>
Same, right view;
<emphasis id= "B944AE5D2B14087BFC9FF8A96ACCF8AE" bold= "true" box= "[895,925,1800,1819]" pageId= "6" > 24)</emphasis>
Same, dorsal view;
<emphasis id= "B944AE5D2B14087BFBA8F8A96D34F8AE" bold= "true" box= "[1096,1125,1800,1819]" pageId= "6" > 25)</emphasis>
Paratype MZSP 103919(2), young specimen, apical (larger) and apertural view (H 25.6 mm);
<emphasis id= "B944AE5D2B14087BFD1EF8836A4DF880" bold= "true" box= "[766,796,1826,1845]" pageId= "6" > 26)</emphasis>
Paratype MZSP 103919(1), adult with thin lip, apertural and right views (H 69.8 mm).
</paragraph>
</caption>
<paragraph id= "8B8F724F2B15087AFFBEFF1968F8FF65" blockId= "7.[94,425,184,208]" box= "[94,425,184,208]" pageId= "7" >
<emphasis id= "B944AE5D2B15087AFFBEFF1969E5FF65" box= "[94,180,184,208]" italics= "true" pageId= "7" > Habitat:</emphasis>
Caatinga environment.
</paragraph>
<paragraph id= "8B8F724F2B15087AFFBEFF596838FEA5" blockId= "7.[94,361,248,272]" box= "[94,361,248,272]" pageId= "7" >
<emphasis id= "B944AE5D2B15087AFFBEFF596870FEA5" box= "[94,289,248,272]" italics= "true" pageId= "7" > Material examined:</emphasis>
<typeStatus id= "548BCCED2B15087AFEC6FF596834FEA5" box= "[294,357,248,272]" pageId= "7" > Types</typeStatus>
.
</paragraph>
<paragraph id= "8B8F724F2B15087AFFBEFE9968F0FE05" blockId= "7.[94,643,312,432]" pageId= "7" >
<emphasis id= "B944AE5D2B15087AFFBEFE99699BFEE5" box= "[94,202,312,336]" italics= "true" pageId= "7" > Etymology:</emphasis>
The specific epithet refers to the outline of the shell, resembling a drop. From the Tupi native language from South America,
<emphasis id= "B944AE5D2B15087AFE2AFED96B4AFE25" box= "[458,539,376,400]" italics= "true" pageId= "7" > amanda</emphasis>
or
<emphasis id= "B944AE5D2B15087AFDDEFED86BD3FE25" box= "[574,642,377,400]" italics= "true" pageId= "7" > amana</emphasis>
meaning rain or related to rain.
</paragraph>
<paragraph id= "8B8F724F2B15087AFFBEFE7869ABFCC5" blockId= "7.[94,643,472,1872]" pageId= "7" >
<emphasis id= "B944AE5D2B15087AFFBEFE786882FE45" box= "[94,467,472,496]" italics= "true" pageId= "7" >
Discussion:
<taxonomicName id= "4C3009CC2B15087AFF37FE796882FE45" authorityName= "Simone" authorityYear= "2012" box= "[215,467,472,496]" class= "Gastropoda" family= "Strophocheilidae" genus= "Megalobulimus" kingdom= "Animalia" order= "Stylommatophora" pageId= "7" phylum= "Mollusca" rank= "species" species= "amandus" > Megalobulimus amandus</taxonomicName>
</emphasis>
clear belong to the informal complex
<taxonomicName id= "4C3009CC2B15087AFEA9FE596992FD85" authority= "(Muller, 1774)" baseAuthorityName= "Muller" baseAuthorityYear= "1774" class= "Gastropoda" family= "Strophocheilidae" genus= "Megalobulimus" kingdom= "Animalia" order= "Stylommatophora" pageId= "7" phylum= "Mollusca" rank= "species" species= "oblongus" >
<emphasis id= "B944AE5D2B15087AFEA9FE596B6CFDA5" box= "[329,573,504,528]" italics= "true" pageId= "7" > Megalobulimus oblongus</emphasis>
(Müller, 1774)
</taxonomicName>
as introduced by
<bibRefCitation id= "EFA10FBE2B15087AFE61FDB96B2EFD85" author= "SIMONE, L. R. L. & LEME, J. L. M." box= "[385,639,536,560]" pageId= "7" pagination= "189 - 203" refId= "ref6060" refString= "SIMONE, L. R. L. & LEME, J. L. M. 1998. Two new species of Megalobulimidae (Gastropoda, Strophocheiloidea) from north Sao Paulo, Brazil. Iheringia serie Zoologia, 85: 189 - 203." type= "journal article" year= "1998" > Simone & Leme (1998)</bibRefCitation>
. This complex includes species with deciduous periostracum, peristome normally reddish, sculpture almost exclusively axial strong undulations, and a protoconch also only sculptured by clear axial, uniform undulations or narrow cords. This set of characters can be regarded as the definition of the genus
<taxonomicName id= "4C3009CC2B15087AFDD0FD796854FCA5" authority= "Morretes, 1952" authorityName= "Morretes" authorityYear= "1952" class= "Gastropoda" family= "Strophocheilidae" genus= "Psiloicus" kingdom= "Animalia" order= "Stylommatophora" pageId= "7" phylum= "Mollusca" rank= "genus" >
<emphasis id= "B944AE5D2B15087AFDD0FD796BD3FD45" box= "[560,642,728,752]" italics= "true" pageId= "7" > Psiloicus</emphasis>
<bibRefCitation id= "EFA10FBE2B15087AFFBEFD596854FCA5" author= "MORRETES, F. L." box= "[94,261,760,784]" pageId= "7" pagination= "109 - 126" refId= "ref5865" refString= "MORRETES, F. L. 1952. Novas especies brasileiras da familia Strophocheilidae. Arquivos de Zoologia, 8 (4): 109 - 126." type= "journal article" year= "1952" > Morretes, 1952</bibRefCitation>
</taxonomicName>
, in such
<taxonomicName id= "4C3009CC2B15087AFE91FD5868A5FCA5" authorityName= "Simone" authorityYear= "2012" box= "[369,500,760,784]" class= "Gastropoda" family= "Strophocheilidae" genus= "Megalobulimus" kingdom= "Animalia" order= "Stylommatophora" pageId= "7" phylum= "Mollusca" rank= "species" species= "amandus" >
<emphasis id= "B944AE5D2B15087AFE91FD5868A5FCA5" box= "[369,500,760,784]" italics= "true" pageId= "7" > M. amandus</emphasis>
</taxonomicName>
and remaining
<taxonomicName id= "4C3009CC2B15087AFF67FCB86850FC85" baseAuthorityName= "Muller" baseAuthorityYear= "1774" box= "[135,257,792,816]" class= "Gastropoda" family= "Strophocheilidae" genus= "Megalobulimus" kingdom= "Animalia" order= "Stylommatophora" pageId= "7" phylum= "Mollusca" rank= "species" species= "oblongus" >
<emphasis id= "B944AE5D2B15087AFF67FCB86850FC85" box= "[135,257,792,816]" italics= "true" pageId= "7" > M. oblongus</emphasis>
</taxonomicName>
complex can someday belong after a deeper revision of the group. A conservative approach is given herein.
</paragraph>
<paragraph id= "8B8F724F2B15087AFF77FCD968D0F9A5" blockId= "7.[94,643,472,1872]" pageId= "7" >
Only few of the 62 valid species of
<taxonomicName id= "4C3009CC2B15087AFDEBFCD969D7FC05" authorityName= "Miller" authorityYear= "1878" class= "Gastropoda" family= "Strophocheilidae" genus= "Megalobulimus" kingdom= "Animalia" order= "Stylommatophora" pageId= "7" phylum= "Mollusca" rank= "genus" >
<emphasis id= "B944AE5D2B15087AFDEBFCD969D7FC05" italics= "true" pageId= "7" > Megalobulimus</emphasis>
</taxonomicName>
have pointed shell apex (
<bibRefCitation id= "EFA10FBE2B15087AFE72FC396B66FC05" author= "BEQUAERT, J. C." box= "[402,567,920,944]" pageId= "7" pagination= "1 - 210" refId= "ref5647" refString= "BEQUAERT, J. C. 1948. Monograph of the Strophocheilidae, a neotropical family of terrestrial mollusks. Bulletin of the Museum of Comparative Zoology, 100 (1): 1 - 210, 32 pls." type= "book chapter" year= "1948" > Bequaert, 1948</bibRefCitation>
;
<bibRefCitation id= "EFA10FBE2B15087AFDA3FC3969C7FC65" author= "LEME, J. L. M." pageId= "7" pagination= "295 - 337" refId= "ref5795" refString= "LEME, J. L. M. 1973. Anatomy and systematics of the Neotropical Strophecheiloidea (Gastropoda, Pulmonata) with the description of a new family. Arquivos de Zoologia, 23 (5): 295 - 337." type= "journal article" year= "1973" > Leme, 1973</bibRefCitation>
;
<bibRefCitation id= "EFA10FBE2B15087AFF43FC19686AFC65" author= "SIMONE, L. R. L." box= "[163,315,952,976]" pageId= "7" refId= "ref6033" refString= "SIMONE, L. R. L. 2006. Land and freshwater molluscs of Brazil. EGB, FAPESP. Sao Paulo. 390 p." type= "book" year= "2006" > Simone, 2006</bibRefCitation>
). This is one of the main characters of
<emphasis id= "B944AE5D2B15087AFF5DFC786812FC45" box= "[189,323,984,1008]" italics= "true" pageId= "7" >
<taxonomicName id= "4C3009CC2B15087AFF5DFC78686EFC45" authorityName= "Simone" authorityYear= "2012" box= "[189,319,984,1008]" class= "Gastropoda" family= "Strophocheilidae" genus= "Megalobulimus" kingdom= "Animalia" order= "Stylommatophora" pageId= "7" phylum= "Mollusca" rank= "species" species= "amandus" > M. amandus</taxonomicName>
,
</emphasis>
as most
<taxonomicName id= "4C3009CC2B15087AFE41FC796B69FC45" authorityName= "Miller" authorityYear= "1878" box= "[417,568,984,1008]" class= "Gastropoda" family= "Strophocheilidae" genus= "Megalobulimus" kingdom= "Animalia" order= "Stylommatophora" pageId= "7" phylum= "Mollusca" rank= "genus" >
<emphasis id= "B944AE5D2B15087AFE41FC796B69FC45" box= "[417,568,984,1008]" italics= "true" pageId= "7" > Megalobulimus</emphasis>
</taxonomicName>
in fact possess a more rounded, dome-shaped apex. For this reason,
<taxonomicName id= "4C3009CC2B15087AFF4EFBB8687CFB85" authorityName= "Simone" authorityYear= "2012" box= "[174,301,1048,1072]" class= "Gastropoda" family= "Strophocheilidae" genus= "Megalobulimus" kingdom= "Animalia" order= "Stylommatophora" pageId= "7" phylum= "Mollusca" rank= "species" species= "amandus" >
<emphasis id= "B944AE5D2B15087AFF4EFBB8687CFB85" box= "[174,301,1048,1072]" italics= "true" pageId= "7" > M. amandus</emphasis>
</taxonomicName>
only needs to be compared with species from
<taxonomicName id= "4C3009CC2B15087AFF04FB98680CFBE5" baseAuthorityName= "Muller" baseAuthorityYear= "1774" box= "[228,349,1080,1104]" class= "Gastropoda" family= "Strophocheilidae" genus= "Megalobulimus" kingdom= "Animalia" order= "Stylommatophora" pageId= "7" phylum= "Mollusca" rank= "species" species= "oblongus" >
<emphasis id= "B944AE5D2B15087AFF04FB98680CFBE5" box= "[228,349,1080,1104]" italics= "true" pageId= "7" > M. oblongus</emphasis>
</taxonomicName>
complex with pointed apex.
<taxonomicName id= "4C3009CC2B15087AFFBEFBF8698EFBC5" authorityName= "Simone" authorityYear= "2012" box= "[94,223,1112,1136]" class= "Gastropoda" family= "Strophocheilidae" genus= "Megalobulimus" kingdom= "Animalia" order= "Stylommatophora" pageId= "7" phylum= "Mollusca" rank= "species" species= "amandus" >
<emphasis id= "B944AE5D2B15087AFFBEFBF8698EFBC5" box= "[94,223,1112,1136]" italics= "true" pageId= "7" > M. amandus</emphasis>
</taxonomicName>
differs from
<taxonomicName id= "4C3009CC2B15087AFE90FBF86841FB25" authority= "(Barattini & Ledon, 1949)" baseAuthorityName= "Barattini & Ledon" baseAuthorityYear= "1949" class= "Gastropoda" family= "Strophocheilidae" genus= "Megalobulimus" kingdom= "Animalia" order= "Stylommatophora" pageId= "7" phylum= "Mollusca" rank= "species" species= "formicacorsii" >
<emphasis id= "B944AE5D2B15087AFE90FBF86B47FBC5" box= "[368,534,1112,1136]" italics= "true" pageId= "7" > M. formicacorsii</emphasis>
(Barattini & Ledón, 1949)
</taxonomicName>
, from
<collectingCountry id= "F32732DF2B15087AFEBAFBD968E5FB25" box= "[346,436,1144,1168]" name= "Uruguay" pageId= "7" > Uruguay</collectingCountry>
, in having aperture wider and longer, sculpture shallower, and umbilicus narrower or absent. It differs from
<taxonomicName id= "4C3009CC2B15087AFE22FB18698AFB45" authority= "(Sowerby, 1825)" baseAuthorityName= "Sowerby" baseAuthorityYear= "1825" class= "Gastropoda" family= "Strophocheilidae" genus= "Megalobulimus" kingdom= "Animalia" order= "Stylommatophora" pageId= "7" phylum= "Mollusca" rank= "species" species= "maximus" >
<emphasis id= "B944AE5D2B15087AFE22FB186B11FB65" box= "[450,576,1209,1232]" italics= "true" pageId= "7" > M. maximus</emphasis>
(Sowerby, 1825)
</taxonomicName>
, from Amazon, in being much shorter and smaller, by shallower suture, and by reddish peristome. It differs from
<taxonomicName id= "4C3009CC2B15087AFEDDFAB869C7FAE5" authority= "Simone & Leme, 1998" authorityName= "Simone & Leme" authorityYear= "1998" class= "Gastropoda" family= "Strophocheilidae" genus= "Megalobulimus" kingdom= "Animalia" order= "Stylommatophora" pageId= "7" phylum= "Mollusca" rank= "species" species= "riopretensis" >
<emphasis id= "B944AE5D2B15087AFEDDFAB8689FFA85" box= "[317,462,1305,1328]" italics= "true" pageId= "7" > M. riopretensis</emphasis>
<bibRefCitation id= "EFA10FBE2B15087AFE33FAB969C7FAE5" author= "SIMONE, L. R. L. & LEME, J. L. M." pageId= "7" pagination= "189 - 203" refId= "ref6060" refString= "SIMONE, L. R. L. & LEME, J. L. M. 1998. Two new species of Megalobulimidae (Gastropoda, Strophocheiloidea) from north Sao Paulo, Brazil. Iheringia serie Zoologia, 85: 189 - 203." type= "journal article" year= "1998" > Simone & Leme, 1998</bibRefCitation>
</taxonomicName>
, from São Paulo, in having narrower aperture, narrower shell width, by more delicate protoconch and teleoconch sculptures, and by less developed umbilicus. It differs from
<taxonomicName id= "4C3009CC2B15087AFEB5FA386B2EFA05" authority= "Morretes, 1952" authorityName= "Morretes" authorityYear= "1952" box= "[341,639,1432,1456]" class= "Gastropoda" family= "Strophocheilidae" genus= "Megalobulimus" kingdom= "Animalia" order= "Stylommatophora" pageId= "7" phylum= "Mollusca" rank= "species" species= "wohlersi" >
<emphasis id= "B944AE5D2B15087AFEB5FA38689FFA05" box= "[341,462,1432,1456]" italics= "true" pageId= "7" > M. wohlersi</emphasis>
<bibRefCitation id= "EFA10FBE2B15087AFE39FA396B2EFA05" author= "MORRETES, F. L." box= "[473,639,1432,1456]" pageId= "7" pagination= "109 - 126" refId= "ref5865" refString= "MORRETES, F. L. 1952. Novas especies brasileiras da familia Strophocheilidae. Arquivos de Zoologia, 8 (4): 109 - 126." type= "journal article" year= "1952" > Morretes, 1952</bibRefCitation>
</taxonomicName>
, from Mato Grosso do Sul, by narrower shape, more elongated aperture, shallower suture, broader spire, and more delicate sculpture.
</paragraph>
<paragraph id= "8B8F724F2B15087AFF77F9B96D79FF45" blockId= "7.[94,643,472,1872]" lastBlockId= "7.[680,1229,184,464]" pageId= "7" >
The megalobulimids from Northeastern
<collectingCountry id= "F32732DF2B15087AFDA8F9B96BD2F985" box= "[584,643,1560,1584]" name= "Brazil" pageId= "7" > Brazil</collectingCountry>
normally possess a purple pigmentation in the protoconch and spire first whorls. This is not the case of
<emphasis id= "B944AE5D2B15087AFFBEF9D869B7F925" box= "[94,230,1656,1680]" italics= "true" pageId= "7" >
<taxonomicName id= "4C3009CC2B15087AFFBEF9D869B0F925" authorityName= "Simone" authorityYear= "2012" box= "[94,225,1656,1680]" class= "Gastropoda" family= "Strophocheilidae" genus= "Megalobulimus" kingdom= "Animalia" order= "Stylommatophora" pageId= "7" phylum= "Mollusca" rank= "species" species= "amandus" > M. amandus</taxonomicName>
,
</emphasis>
which has uniform coloration. On the other hand, the species of that region are normally of small size for a
<emphasis id= "B944AE5D2B15087AFEFAF91968B5F965" box= "[282,484,1720,1744]" italics= "true" pageId= "7" >
<taxonomicName id= "4C3009CC2B15087AFEFAF91968E5F965" authorityName= "Miller" authorityYear= "1878" box= "[282,436,1720,1744]" class= "Gastropoda" family= "Strophocheilidae" genus= "Megalobulimus" kingdom= "Animalia" order= "Stylommatophora" pageId= "7" phylum= "Mollusca" rank= "genus" > Megalobulimus</taxonomicName>
, i.e.,
</emphasis>
below
<quantity id= "4CC8DFAA2B15087AFDD0F9196B2EF97A" box= "[560,639,1720,1744]" metricMagnitude= "-2" metricUnit= "m" metricValue= "8.0" pageId= "7" unit= "mm" value= "80.0" > 80 mm</quantity>
, which is the case of the new species. Another interesting feature of
<taxonomicName id= "4C3009CC2B15087AFEF0F95868C1F8A5" authorityName= "Simone" authorityYear= "2012" box= "[272,400,1784,1808]" class= "Gastropoda" family= "Strophocheilidae" genus= "Megalobulimus" kingdom= "Animalia" order= "Stylommatophora" pageId= "7" phylum= "Mollusca" rank= "species" species= "amandus" >
<emphasis id= "B944AE5D2B15087AFEF0F95868C1F8A5" box= "[272,400,1784,1808]" italics= "true" pageId= "7" > M. amandus</emphasis>
</taxonomicName>
is the absence of folds and teeth in the peristome, even in more thickened specimens (
2024-08-02 20:36:26 +02:00
<figureCitation id= "130B6ECA2B15087AFF36F8996806F8E5" box= "[214,343,1848,1872]" captionStart= "FIGURES 15‑ 26" captionStartId= "6.[132,222,1724,1743]" captionTargetBox= "[132,1266,497,1688]" captionTargetId= "figure-99@6.[132,1266,497,1688]" captionTargetPageId= "6" captionText= "FIGURES 15‑ 26: Shell of types and ordinary specimens: 15-20) Anostoma tessa new species; 15) Holotype, apertural view (H 31.1 mm); 16) Same, dorsal view; 17) Same, left view; 18) Same, right view; 19) Same, detail of aperture, apertural-slightly anterior view; 20) Paratype MZSP 103915, apertural view (H 31.1 mm); 21) Rhinus suturalis, MZSP 103924, frontal view (H 23.6 mm); 22-26) Megalobulimus amandus new species; 22) Holotype, apertural view (H 78.9 mm); 23) Same, right view; 24) Same, dorsal view; 25) Paratype MZSP 103919(2), young specimen, apical (larger) and apertural view (H 25.6 mm); 26) Paratype MZSP 103919(1), adult with thin lip, apertural and right views (H 69.8 mm)." figureDoi= "http://doi.org/10.5281/zenodo.13175200" httpUri= "https://zenodo.org/record/13175200/files/figure.png" pageId= "7" > Figs. 23, 24</figureCitation>
2024-07-12 03:17:58 +02:00
). Tooth and folds in parietal callus or in middle level of outer lip are common occurrences in species from that region.
</paragraph>
<paragraph id= "8B8F724F2B15087AFD01FF596A17FE65" blockId= "7.[680,1229,184,464]" pageId= "7" >
A conservative approach is given here considering the genus
<taxonomicName id= "4C3009CC2B15087AFCB8FEB96ABEFE85" authorityName= "Miller" authorityYear= "1878" box= "[856,1007,280,304]" class= "Gastropoda" family= "Strophocheilidae" genus= "Megalobulimus" kingdom= "Animalia" order= "Stylommatophora" pageId= "7" phylum= "Mollusca" rank= "genus" >
<emphasis id= "B944AE5D2B15087AFCB8FEB96ABEFE85" box= "[856,1007,280,304]" italics= "true" pageId= "7" > Megalobulimus</emphasis>
</taxonomicName>
in
<taxonomicName id= "4C3009CC2B15087AFBF9FEB96A79FEE5" authority= "Leme, 1973" authorityName= "Leme" authorityYear= "1973" class= "Gastropoda" family= "Megalobulimidae" higherTaxonomySource= "GBIF" kingdom= "Animalia" order= "Stylommatophora" pageId= "7" phylum= "Mollusca" rank= "family" >
Megalobulimidae
<bibRefCitation id= "EFA10FBE2B15087AFD48FE996A79FEE5" author= "LEME, J. L. M." box= "[680,808,312,336]" pageId= "7" pagination= "295 - 337" refId= "ref5795" refString= "LEME, J. L. M. 1973. Anatomy and systematics of the Neotropical Strophecheiloidea (Gastropoda, Pulmonata) with the description of a new family. Arquivos de Zoologia, 23 (5): 295 - 337." type= "journal article" year= "1973" > Leme, 1973</bibRefCitation>
</taxonomicName>
. However, it is recognized that the taxon can possibly be a special branch of
<taxonomicName id= "4C3009CC2B15087AFBDAFEF96B9FFE25" authorityName= "Pilsbry" authorityYear= "1902" class= "Gastropoda" family= "Strophocheilidae" kingdom= "Animalia" order= "Stylommatophora" pageId= "7" phylum= "Mollusca" rank= "family" > Strophocheilidae</taxonomicName>
, and even
<taxonomicName id= "4C3009CC2B15087AFCDEFED96ACCFE25" authorityName= "Pilsbry" authorityYear= "1895" box= "[830,925,376,400]" class= "Gastropoda" family= "Acavidae" kingdom= "Animalia" order= "Stylommatophora" pageId= "7" phylum= "Mollusca" rank= "family" > Acavidae</taxonomicName>
. A project on this matter has been developed, mainly considering phylogenetic methodologies.
</paragraph>
</subSubSection>
</treatment>
</document>